860:. For instance, tularaemia is endemic in the region, and outbreaks had previously occurred in 1938 and the winter of 1941-1942; the disease was already present among the civilian population by the time German troops arrived. Infected rodents — rampant at the front, as sanitation systems had completely broken down and the uncollected grain harvest provided plentiful food for the rodent population — were the key to the large-scale outbreaks, which spread through inhalation of dust from contaminated straw in mattresses or from consumption of tainted food and water. Crucially, Geissler notes that there are no contemporary accounts by neither the German or Soviet armies nor intelligence services regarding the use of
895:, Commander of the Kwantung Army had already ordered the destruction and evacuation of these facilities. The NKVD now switched its focus to apprehending any personnel associated with Units 100 and 731 and began a process of filtration of the 560,000-760,000 Japanese prisoners of war. In December 1949, the military figures identified by the Soviets as participating in the Japanese BW programme were put on trial in Khabarovsk. The defendants were found guilty and sentenced to terms ranging from two to twenty-five years in Soviet labour correction camps. However, the officers, doctors and other personnel from Unit 731 were in fact transferred to the comparative comfort of the
3536:
1568:. A research ship of the Aral fleet had come within 15 km from the island (it was forbidden to come any closer than 40 km). The lab technician of this ship took samples of plankton twice a day from the top deck. The smallpox formulation— 400 gr. of which was exploded on the island—”got her”, and she became infected. After returning home to Aralsk, she infected several people, including children. All of them died. I suspected the reason for this and called the General Chief of Staff at the Ministry of Defense and requested to forbid the
926:, the Soviet minister of internal affairs, maintained control of the Soviet BW programme and further developed its offensive capabilities. The key hub of the Soviet BW programme at this time was the Scientific-Research Institute of Epidemiology and Hygiene located in Kirov. It continued to utilise the biological weapons test site on Vozrozhdeniya Island in the Aral Sea. During the period 1947-1949 a new military biological weapons facility, the USSR Ministry of Defence's Scientific-Research Institute of Hygiene, was established in
969:), the prime task of which was to create covert dual-use BW facilities at a number of pharmaceutical and microbiological enterprises. Over the next decade or so, dual-use BW production plants were created at Berdsk, Omutninsk, Penza and Kurgan. It is therefore apparent that previous perceptions by Western scholars of the Khrushchev era as contributing little to the development of the Soviet Union's biological warfare capabilities are incorrect. Rimmington argues that this "
340:
1650:
1229:
930:, It occupied the site of the former Cherkassk-Sverdlovsk Infantry Academy on Ulitsa Zvezdnaya, 1. The new facility became operational in July 1949. Its core staff were sourced from the Kirov BW facility. The first group to arrive from Kirov included the new director of the Institute of Hygiene, Major General Nikolai Fillipovich Kopylov. The Sverdlovsk facility launched a scientific programme in 1951 which focused on botulinum toxin.
33:
1174:, the first deputy chief of Biopreparat from 1988 to 1992, confirmed that the program had been massive and that it still existed. On 11 April 1992, Yeltsin decreed "the termination of research on offensive biological weapons, the dismantlement of experimental technological lines for the production of biological agents and the closure of biological weapons testing facilities", and in September 1992 Yeltsin agreed in a
775:
1186:(2012), state flatly that "In March 1992...Yeltsin acknowledged the existence of an illegal bioweapons program in the former Soviet Union and ordered it to be dissolved. His decree was, however, not obeyed." They conclude that "In hindsight, we know that with the ultimate failure of the... process and the continued Russian refusal to open the... facilities to the present day, neither the Yeltsin or
981:(BWC). However, citing doubts concerning the United States’ compliance with the BWC, they subsequently augmented their biowarfare programs. The Soviet bioweapon effort became a huge program rivaling its considerable investment in nuclear arms. It comprised various institutions operating under an array of different ministries and departments including the Soviet
699:. Around 100 personnel from Velikanov's Biotechnical Institute participated in the experiments. In July 1937, while planning for a second expedition to the island, Velikanov was arrested by the Soviet security organs and subsequently shot. Later that same summer, Leonid Moiseevich Khatanever, the new director of the Biotechnical Institute and an expert on
758:(the causative organism of plague) and other dangerous pathogens. Zlatogorov was in fact one of the world's leading authorities on pneumonic plague and had studied 40 strains of plague bacilli from around the world. He had been a leading participant of a Russian team despatched to combat the October 1910 to February 1911 outbreak of
1528:
biologics plant is the best-documented of the agricultural BW mobilisation facilities. A UK/US inspection team visited the facility in 1993 and identified five underground reinforced concrete bunkers holding hundreds of thousands of hen’s eggs being used to grow massive quantities of virus, allegedly
1178:
with the United States and the United
Kingdom that the two Western nations would "have a blanket invitation to visit facilities of concern in Russia under ground rules that guarantee unprecedented access, including access to the entire facility, the ability to take samples, the right to interview the
1636:
under the leadership of Dr. Ustinov who was accidentally killed by the virus. The samples of
Marburg taken from Ustinov's organs were more powerful than the original strain. The new strain, called "Variant U", had been successfully weaponized and approved by the Soviet Ministry of Defense in 1990.
933:
In the early 1950s the Soviet leadership became concerned that the USSR was vulnerable to attack by a new generation of virus-based biological weapons. At this time the country only possessed a single facility focused on viruses, the Moscow-based D.I. Ivanovskii
Institute of Virology. In 1952, as a
533:
confirmed that the program had been massive and still existed. In 1992, a
Trilateral Agreement was signed with the United States and the United Kingdom promising to end biological weapons programs and convert facilities to benevolent purposes, but compliance with the agreement—and the fate of the
2375:
United States. Congress. House. Committee on Armed
Services. Subcommittee on Intelligence, Emerging Threats and Capabilities. (11 October 2013). Biodefense: Worldwide Threats and Countermeasure Efforts for the Department of Defense : Hearing Before the Subcommittee on Intelligence, Emerging
798:
were implemented to counter the perceived threat of biological sabotage by the German and
Japanese intelligence services. As a result there was a tightening-up of state control over personnel working on microbial pathogens and an emphasis on the gathering of intelligence from foreign legations
938:) in Zagorsk. It was transferred to the control of the USSR Ministry of Defence in March 1954. The new military institute later pursued major programmes focused on variola virus and viral haemorrhagic fevers. In his uncorroborated account, Alibek claims that capacity for the production of
766:, located at Kronstadt. Here they aimed to develop strains of plague that remained viable when loaded into artillery shells, aerial bombs and other means of dispersal. German intelligence independently identified the secret BW programme allegedly managed by Zlatogorov and Maslakovets.
914:). The leniency with which the Japanese BW specialists were treated - the longest sentence any served was seven years - has led a number of scholars to conclude that some sort of deal was struck between the Soviet authorities and the Unit 731 personnel held captive in the USSR.
1215:
As of 2021, the United States "assesses that the
Russian Federation (Russia) maintains an offensive BW program and is in violation of its obligation under Articles I and II of the BWC. The issue of compliance by Russia with the BWC has been of concern for many years".
1067:
prohibited development, testing, and production efforts... cites several
Gorbachev decrees stepping up the pace of work within the biological weapons complex, and directing the creation of mobile production facilities so that inspectors could not uncover the program.
971:
was in fact a pivotal period in the Soviet programme, when BW production technology was being transferred from the military to facilities concealed within civil manufacturing plants. This was later to manifest itself as a key feature of the subsequent
Biopreparat
1409:"Pokrov, Berdsk and Omutninsk all revealed evidence of biological activity since 1975, such as large-scale production in hardened facilities, aerosol test chambers, excessive containment levels for current activity and accommodation for weapons-filling lines."
3599:
2006:
2796:
752:(SIS) - commonly known as MI6. A total of fourteen highly detailed reports on the Soviet BW programme were issued in the period 1924-1927. SIS identifies Petr Petrovich Maslakovets and Semen Ivanovich Zlatogorov as the lead BW scientists working on
680:). Velikanov was placed in command of the Gorodomlya Island facility which was named as the Biotechnical Institute, also known by the code designation V/2-1094. German intelligence reported that the institute was engaged in experiments focused on
655:
1275:
1278:, Kol'tsovo (1974-1991): "At Koltsova access was again difficult and problematic. The most serious incident was when senior officials contradicted an admission by technical staff that research on smallpox was being conducted there."
703:(the causative agent of tularaemia), led a second expedition to Vozrozhdeniya. Two special ships and two aircraft were assigned to Khatanever for use in tests focused on the dissemination of tularaemia bacteria. Germany launched
811:) - on Gorodomlya Island, was under immediate threat of capture. At some point around the 25 September, the facility was evacuated and the buildings partially destroyed. There are various accounts regarding the relocation of
1681:
1480:
A Soviet-era agricultural biowarfare programme was pursued from 1958 through to the collapse of the USSR in 1991. This program focused on anti-crop and anti-livestock biological weapons, with Soviet efforts starting with
1312:
1179:
workers and scientists, and the right to record the visits on video and audio tape." Yeltsin promised to end the
Russian bio-weapons program and to convert its facilities for benevolent scientific and medical purposes.
961:" (Ecology), the new network incorporated three virology facilities. These worked in close collaboration with Soviet military virologists and focused on both animal pathogens such as FMD and exotic zoonotic infections.
3621:
2835:
1154:
aimed to secure and dismantle weapons of mass destruction, including the Soviet biological weapons program. The threat reduction assisted post-Soviet states in containing and destroying the pathogens in Soviet labs.
905:
815:, with official Russian sources indicating that it was initially transferred to Saratov. In the later summer of 1942, in the face of the German offensive to capture Stalingrad, there was a second evacuation of
670:, where around nineteen leading plague and tularaemia specialists were forced to work on the development of biological weapons. By 1936, scientists working on BW at both Vlasikha and Suzdal were transferred to
2239:
Eric Croddy, Sarka Krčálová, Tularemia, Biological Warfare, and the Battle for Stalingrad (1942–1943), Military Medicine, Volume 166, Issue 10, October 2001, Pages 837–838. Retrieved February 2 2024, from
806:
and invaded the Soviet Union along a 2,900-kilometre front. Such was the rapidity and depth of penetration of the attack that, by September, the Red Army's BW facility - the Sanitary Technical Institute
1029:
The Soviet Union continued the development and mass production of offensive biological weapons, despite having signed the 1972 BWC. The development and production were conducted by a main directorate ("
964:
In 1953 the management of the Soviet BW programme was assigned to the USSR Ministry of Defence's Fifteenth Administration. In August 1958, the latter created a new Scientific-Research Technical Bureau
786:" had actively undertaken BW preparations and that the use of such weapons, in case of emergency, was a foregone conclusion. Stalin in response ordered an acceleration of BW preparations and appointed
641:
Running parallel to the work underway at Vlasikha, BW research was also being pursued in an institution controlled by the state security apparatus. In July 1931, the Joint State Political Directorate (
1475:
1287:
1175:
1387:
1701:
401:(BWC), the Soviet authorities subsequently augmented their biowarfare programs. Over the course of its history, the Soviet program is known to have weaponized and stockpiled the following
1098:
in 1990. The development of genetically altered strains of smallpox was presumably conducted in the Institute under the leadership of Dr. Sergei Netyosov in the mid-1990s, according to
618:, emerged as the lead scientist in the early Soviet biological weapons program. In 1930, Velikanov was placed in command of a new facility, the Red Army's Vaccine-Sera Laboratory in
1497:. Another innovation was the "mobilization production facilities"—ostensibly civil manufacturing plants—which incorporated capacity for production of weapons in wartime emergency."
1087:
to kill chicken. These agents were prepared to be sprayed down on enemy fields from tanks attached to airplanes over hundreds of miles. The secret program was code-named "Ecology".
3496:
2376:
Threats and Capabilities of the Committee on Armed Services, House of Representatives, One Hundred Thirteenth Congress, First Session, Hearing Held October 11, 2013. p. 34.
1505:
1466:
programmes was one reason why Alibek defected to the United States in 1992. Journal articles by scientists suggest that in 1999 the experiments were still being continued.
555:
regarding its applicability and use in retaliation. The Soviet Union was one, when it deposited its ratification notice. Due to these reservations, it was in practice a "
1803:
3501:
1633:
1317:
1095:
2609:
1303:
551:, which prohibits the use but not the possession or development of chemical and biological weapons. Upon ratification of the Geneva Protocol, several countries made
3202:
Re-Evaluating Russia's Biological Weapons Policy, as Reflected in the Criminal Code and Official Admissions: Insubordination Leading to a President's Subordination
326:
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883:
in the summer of 1942, and at a later unspecified date, again by Unit 731. On the 9 August 1945, the Soviet Union launched its invasion of Japanese-controlled
650:
1182:
Compliance with the agreement, as well as the fate of the former Soviet bio-agents and facilities, is still mostly undocumented. Leitenberg and Zilinskas, in
517:, for example, was 90 to 100 tons. In the 1980s and 1990s, many of these agents were genetically altered to resist heat, cold, and antibiotics. In the 1990s,
3260:
1362:
369:
2520:
934:
response to this perceived area of weakness, the Soviet government issued a special decree for the creation of the Scientific-Research Sanitary Institute (
1608:
in April 1979. The death toll was at least 66, but no one knows the precise number, because all hospital records and other evidence were destroyed by the
1728:
630:
institute, subordinate to the People's Commissariat of Health, were transferred to the military facility. Early programs at the military lab focused on
513:
These programs became immense and were conducted at dozens of secret sites employing up to 65,000 people. Annualized production capacity for weaponized
1686:
736:. The Leningrad Military Medical Academy began cultivating typhus in chicken embryos. He also alleges that human experimentation occurred with typhus,
674:
where they occupied an institute for the study of foot-and-mouth disease which had been built originally for the People’s Commissariat of Agriculture (
386:
covertly operated the world's largest, longest, and most sophisticated biological weapons program, thereby violating its obligations as a party to the
397:
By 1960, numerous military-purposed biological research facilities existed throughout the Soviet Union. Although the former USSR also signed the 1972
3594:
3470:
1738:
1417:
1911:
1431:
was the codename for the budget to develop microbial weapons with new properties of high virulence, improved stability, and new clinical syndromes.
3616:
3475:
3394:
1723:
1428:
732:
under the control of OGPU (see above). According to Alibek, in 1928, the Revolutionary Military Council signed a decree about the weaponization of
3553:
3531:
1733:
1403:
1038:
954:
748:. Another, possibly more reliable source, regarding the Soviet BW programme in Leningrad are a series of secret reports generated by the British
676:
3211:"The Memoirs of an Inconvenient Man: Revelations About Biological Weapons Research in the Soviet Union" by Igor V. Domaradskij and Wendy Orent,
3526:
1270:
270:
71:
1961:
1934:
1548:
An outbreak of weaponized smallpox occurred during testing in 1971. General Professor Peter Burgasov, former Chief Sanitary Physician of the
1190:
administrations ever carried out 'a visible campaign to dismantle once and for all' the residual elements of the Soviet bioweapons program".
275:
3201:
728:, describes a quite separate strand of early BW research being pursued in Leningrad. This work was allegedly initiated in the 1920s at the
322:
230:
225:
3000:
2695:
2129:
Biohazard: The Chilling True Story of the Largest Covert Biological Weapons Program in the World - Told from Inside by the Man Who Ran it.
1151:
250:
235:
195:
185:
180:
1204:
In the 2000s, the academician, "A.S.", proposed a new biological warfare program, called the "Biological Shield of Russia" to president
3711:
900:
280:
3253:
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362:
95:
90:
3135:
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205:
190:
105:
1564:, the strongest formulations of smallpox were tested. Suddenly, I was informed that there were mysterious cases of mortalities in
3690:
695:
In the summer of 1936, Ivan Mikhailovich Velikanov led the Red Army's first expedition to conduct tests of biological weapons on
265:
210:
175:
130:
85:
80:
2377:
2227:
Geißler, Erhard. (2005). Alibek, Tularaemia and The Battle of Stalingrad. CBW Convention Bull. Retrieved February 2 2024, from
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125:
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100:
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2070:
1895:
1198:
255:
165:
160:
135:
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115:
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1368:
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of 1972. The Soviet program began in the 1920s and lasted until at least September 1992 but has possibly been continued by
355:
343:
145:
140:
1657:
1236:
1159:
762:. The SIS reports indicate that Zlatogorov and Maslakovets conducted some of their research on a so-called Plague Fort -
949:
Just four years after the creation of the Zagorsk facility, the CPSU and the USSR Council of Ministers issued a decree "
1034:
982:
2797:"From Offence to Defence? Russia's Reform of its Biological Weapons Complex and the Implications for Western Security"
3506:
3449:
3389:
3232:
3120:
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1787:(pp. 698-712). Cambridge, Massachusetts; London, England: Harvard University Press. Retrieved February 7, 2021, from
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763:
642:
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2864:
1871:
1804:"2021 Adherence to and Compliance With Arms Control, Nonproliferation, and Disarmament Agreements and Commitments"
1135:
to pressure Gorbachev into opening for inspection several of his facilities. The visits occurred in January 1991.
1102:. (aka Kanatjan Alibekov). In 1998, Alibek reported that "there was significant discussion of the possible use of
3548:
3038:
1517:
155:
1576:, who at that time was the Chief of the KGB, and informed him of the unique formulation of smallpox obtained on
794:, in overall command of the country's biological warfare programme. In May 1941, a number of measures codenamed
3222:
2966:
1494:
599:
3069:
1193:
In the 1990s, specimens of deadly bacteria and viruses were stolen from western laboratories and delivered by
1021:
research. It conducted its secret activities at numerous sites across the USSR and employed 30-40,000 people.
3685:
3670:
3589:
1750:
1712:
978:
398:
387:
296:
2684:
Biological Espionage: Special Operations of the Soviet and Russian Foreign Intelligence Services in the West
1476:
Ministry of Agriculture and Food (Soviet Union) § Role in offensive Soviet biological warfare programme
2340:
The Soviet Union's Invisible Weapons of Mass Destruction: Biopreparat's Covert Biological Warfare Programme
1661:
1482:
1240:
849:
842:, suggests that in the late summer of 1942 the Red Army engaged in the deliberate aerosol dissemination of
492:
2627:
Adherence To and Compliance With Arms Control, Nonproliferation and Disarmament Agreements and Commitments
2007:"The Shortcomings of Indeterminacy in Arms Control Regimes: The Case of the Biological Weapons Convention"
3644:
3584:
3364:
3196:
3048:
2165:
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607:
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of 1979, which had resulted in the deaths of at least 64 people. Defecting Soviet bioweaponeers such as
306:
2281:
3569:
3409:
3369:
3043:
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train from stopping in Aralsk. As a result, an epidemic throughout the country was prevented. I called
24:
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of 1979, which had resulted in the deaths of at least 64 people. Soviet defectors, including Colonel
782:
By 1939, with the USSR on a war footing, the Soviet leadership is reported to have believed that the "
3095:
1266:
606:" and proposed a plan for the organisation of Soviet military bacteriology. It was at this time that
2652:
2648:
1628:
The Soviet Union reportedly had a large biological weapons program enhancing the usefulness of the
1046:
994:
896:
563:
3639:
2742:
1590:
1525:
1382:
1334:
1167:
868:
522:
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Shoham D, Wolfson Z (2004). "The Russian biological weapons program: vanished or disappeared?".
1983:
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3419:
3191:
2699:
2631:
2625:
2148:
1543:
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1072:
867:
In the Far East the Soviet Union had been subject of a BW attack in 1939 during the battles of
844:
682:
434:
856:. However, a number of prominent scholars have disputed Alibek's version of events, including
3675:
3146:
Statement by Dr. Kenneth Alibek before the Joint Economic Committee of United States Congress
1552:, and a senior researcher within the program of biological weapons described this incident:
1498:
595:
301:
3166:
1890:. London: Verification Research, Training and Information Centre (VERTIC). pp. 93–109.
3543:
2404:
1830:
Preventing Proliferation of Biological Weapons: U.S. Assistance to the Former Soviet States
1577:
1557:
1354:
1197:
planes to support the Russian biological weapons program. At least one of the pilots was a
853:
803:
711:
in October, the BW facility on Gorodomlya Island was evacuated and eventually relocated to
704:
696:
62:
3221:, a useful page about non-state weapons transfers with a lot of links to information from
3176:
3090:
2591:
1394:
Today Virological Center NIIM (Scientific research institute) Russian Defense Ministry in
8:
2549:
1509:
1372:
1080:
820:
745:
712:
521:
admitted to an offensive biological weapons program as well as to the true nature of the
446:
2408:
1604:(the causative agent of anthrax) were accidentally released from a military facility in
3665:
3579:
3343:
3318:
3238:
2914:
2816:
2501:
2034:
1697:
1600:
1110:
591:
526:
410:
391:
3145:
2836:"The Soviet Union's Offensive Program: The Implications for Contemporary Arms Control"
562:
The principal architect of the Soviet Union's first military biological programme was
3434:
3186:
3180:
3116:
2941:
2918:
2904:
2843:
2820:
2775:
2748:
2725:
2687:
2613:
2493:
2344:
2314:
2287:
2260:
2228:
2205:
2171:
2132:
2093:
2066:
2026:
1891:
1691:
1490:
1486:
1208:. The program reportedly includes institutes of the Russian Academy of Sciences from
1171:
1132:
1128:
1061:
1014:
888:
759:
671:
552:
530:
470:
2505:
1882:
1166:, admitted to an offensive bio-weapons program as well as to the true nature of the
1071:
In the 1980s, the Soviet Ministry of Agriculture successfully developed variants of
951:
for strengthening scientific-research work in the field of microbiology and virology
857:
3660:
3611:
2896:
2808:
2485:
2412:
2038:
2018:
1912:"What happened after an explosion at a Russian disease research lab called VECTOR?"
1463:
1420:
was the codename for the budget to develop antibiotic-resistant microbial strains.
828:
427:
47:
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1828:
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3374:
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1929:
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923:
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611:
548:
487:
422:
52:
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officer". At least two agents died, presumably from the transported pathogens.
892:
3414:
3293:
3139:
3113:
1878:
1617:
1395:
1331:
All-Union Scientific-Research Institute of Veterinary Virology and Microbiology
1205:
1187:
1139:
1099:
911:
602:) on the Soviet Union's preparedness for biological warfare. It asserted that "
547:
International restrictions on biological warfare began only with the June 1925
240:
57:
2900:
2812:
2489:
2022:
876:
3705:
3333:
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2030:
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1605:
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1378:
1220:
List of Soviet/Russian biological weapons institutions, programs and projects
1163:
1018:
1005:, was created under the Main Administration of the Microbiological Industry (
927:
518:
507:
502:
2893:
The Soviet Union's Agricultural Biowarfare Programme: Ploughshares to Swords
2606:
Working Paper: Biological Weapons in the 20th Century: A Review and Analysis
2310:
The Soviet Union's Agricultural Biowarfare Programme: Ploughshares to Swords
2241:
1956:
1001:. In April 1974, a new agency, the All-Union Science Production Association
3454:
3384:
3348:
2562:
2497:
1451:
556:
383:
3491:
3424:
3404:
3399:
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2974:
2741:
Leitenberg, Milton; Zilinskas, Raymond A.; Kuhn, Jens H. (29 June 2012).
2521:"Ex-Soviet Bioweapons Labs Are Fighting COVID-19. Moscow Doesn't Like It"
2280:
Leitenberg, Milton; Zilinskas, Raymond A.; Kuhn, Jens H. (29 June 2012).
1613:
1549:
1459:
1345:
1325:
1260:
1124:
1084:
1030:
946:. It was reported to use chicken eggs for the cultivation of the virus.
838:
741:
724:
718:
In his account of the history of the Soviet BW programme, Alibek, who as
463:
953:". This decree established a secret network of BW institutes within the
823:, located some 896 kilometres north-east of Moscow on the Vyatka river.
3574:
3439:
1872:"The Trilateral Agreement: lessons for biological weapons verification"
1502:
1076:
833:
719:
635:
2416:
1282:
All-Union Scientific-Research Institute of Highly Pure Biopreparations
534:
former Soviet bio-agents and facilities—is still mostly undocumented.
3338:
3323:
2768:
Geissler, Erhard; Gazsó, Lajos G.; Buder, Ernst (30 September 1998).
1783:
Leitenberg, M., Zilinskas, R., & Kuhn, J. (2012). Conclusion. In
1297:
1209:
884:
666:
439:
402:
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and then the following year created a special prison laboratory, or
3308:
3298:
3283:
3170:
3001:"Anthrax genome reveals secrets about a Soviet bioweapons accident"
2666:
1569:
1561:
1532:
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1358:
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939:
880:
872:
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514:
497:
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451:
1377:
Sverdlovsk bioweapons production facility (Military Compound 19),
3521:
3313:
3288:
2840:
Biological Warfare and Disarmament: New Problems/New Perspectives
1391:
943:
774:
587:
475:
415:
3157:
The Russian Biological Weapons Program: Vanished or Disappeared?
2696:"Interview: Alexander Kouzminov, Author of Biological Espionage"
2256:
Stalin's Secret Weapon: The Origins of Soviet Biological Warfare
2201:
Stalin's Secret Weapon: The Origins of Soviet Biological Warfare
2167:
Stalin's Secret Weapon: The Origins of Soviet Biological Warfare
2089:
Stalin's Secret Weapon: The Origins of Soviet Biological Warfare
2062:
Stalin's Secret Weapon: The Origins of Soviet Biological Warfare
1271:
State Research Centre for Applied Microbiology and Biotechnology
32:
3516:
3444:
3163:, Volume 30, Number 4, October–December 2004, pp. 241–261.
1595:
1565:
733:
661:
623:
2722:
The Perversion Of Knowledge: The True Story of Soviet Science.
3429:
3110:
The Perversion Of Knowledge: The True Story of Soviet Science
1833:, (Congressional Research Service Report for Congress), pg 3.
1462:
and smallpox at the Vector Institute. The existence of these
891:
and Changchun were overrun by the Red Army. However, General
879:. Further attacks were initiated against the Soviet Union by
3026:
Biowarrior: Inside the Soviet/Russian Biological War Machine
1276:
All-Union Scientific-Research Institute of Molecular Biology
1138:
It has been reported that Russia made smallpox available to
1513:
1013:
pursued offensive research, development, and production of
799:
relating to the feasibility and use of biological weapons.
791:
708:
3177:
An Obscure Weapon of the Cold War Edges Into the Limelight
1984:"Russian Federation: Ratification of 1925 Geneva Protocol"
1318:
Vector State Research Center of Virology and Biotechnology
3226:
2610:
Center for International and Security Studies at Maryland
1609:
1444:
1050:
998:
778:
Lavrentiy Beria, commander of Soviet BW programme in 1939
566:. In August 1925, he was appointed the first head of the
3268:
2865:"Next Generation Bioweapons: Genetic Engineering and BW"
2740:
2567:
Institute for the Study of Conflict Ideology and Policy
2279:
1827:
Cook, Michelle Stem and Amy F. Woolf (April 10, 2002),
1640:
1443:
attempted in the late 1980s and early 1990s to combine
1267:
All-Union Scientific-Research Institute of Microbiology
590:). In February 1928, Fishman prepared a key report for
2458:
Preston, Richard (9 March 1998). "THE BIOWEAPONEERS".
1361:, this BW test site was built here and on neighboring
1281:
604:
the bacterial option could be successfully used in war
3197:
Memories of bioweapons developer Domaradsky (Russian)
2223:
2221:
1957:"Disarmament Treaties Database: 1925 Geneva Protocol"
1330:
2834:
Rimmington, Anthony; Wright, S. (1 September 2002).
2643:
Leitenberg, Milton and Raymond A. Zilinskas (2012),
1522:
The Soviet Union’s Agricultural Biowarfare Programme
3215:, Volume 27, Issue 4 October 2001, pages 239 - 266.
2475:
2332:
2330:
1313:
Stepnogorsk Scientific Experimental-Industrial Base
887:. Two major Japanese offensive BW installations at
848:(the causative agent of tularaemia) against German
405:(and to have pursued basic research on many more):
2767:
2614:http://www.cissm.umd.edu/documents/bw%2020th%c.pdf
2592:"Russia Commits to End Biological Weapons Program"
2546:Germs: Biological Weapons and America’s Secret War
2229:https://www.researchgate.net/publication/242160520
2218:
1009:) to spearhead the Soviet offensive BW programme.
899:in a tsarist-era red-brick manor house located in
598:for Military and Navy Affairs and Chairman of the
3167:Red Lies: Biological warfare and the Soviet Union
1529:in order to sustain a strategic weapons system."
707:in June 1941 and following the capture of nearby
3703:
3152:Post-World War II Programs of Biological Weapons
2833:
2744:The Soviet Biological Weapons Program: a history
2645:The Soviet Biological Weapons Program: A History
2327:
2283:The Soviet Biological Weapons Program: a history
1533:Notable biological agent outbreaks and accidents
1184:The Soviet Biological Weapons Program: A History
819:, which was eventually permanently relocated to
578:). In 1926, at a small laboratory controlled by
3208:, Volume 32, Issue 1 January 2006, pages 1 – 13
2886:
2884:
1986:. United Nations Office for Disarmament Affairs
1404:Poison laboratory of the Soviet secret services
2612:, University of Maryland; 2001. Available at:
1263:(18 labs, test sites, and production centers)
16:Russian bioweapons program from 1920s to 1990s
3254:
3024:Domaradskij, Igor V. and Wendy Orent (2003),
2390:
2388:
2386:
2193:
2191:
2189:
2187:
1962:United Nations Office for Disarmament Affairs
1935:United Nations Office for Disarmament Affairs
1269:, Obolensk (1974-1991); later renamed as the
1106:as a biological weapon instead of smallpox."
363:
2881:
2598:
1485:, for which an institute was established on
802:On the 22 June 1941, Nazi Germany commenced
722:had been a biological weapons scientist for
3497:Scientific Experimental and Production Base
3204:by Jan T. Knoph; Kristina S. Westerdahl.
3136:Soviet Defector Warns of Biological Weapons
3091:"Interviews With Biowarriors: Sergei Popov"
3072:. London: Daily Telegraph. 29 November 2001
2856:
2678:
2676:
2544:J Miller, S Engelberg, and W Broad (2001),
1113:convinced the British that the Soviets had
1094:on an industrial scale was launched in the
1056:Alibek maintains that "Soviet leaders from
826:In his account of the Soviet BW programme,
3261:
3247:
2890:
2794:
2383:
2336:
2306:
2252:
2197:
2184:
2163:
2085:
2058:
1865:
1863:
1861:
1859:
1789:http://www.jstor.org/stable/j.ctt2jbscf.30
370:
356:
31:
3233:Bioweapons from Russia: Stemming the Flow
3192:Soviet Army used 'rat weapon' during WWII
2862:
2774:. Springer Science & Business Media.
2471:
2469:
2395:Smithson, Amy (1999). "A bio nightmare".
2242:https://doi.org/10.1093/milmed/166.10.837
2123:
2121:
2119:
2117:
2115:
2113:
2111:
2109:
1857:
1855:
1853:
1851:
1849:
1847:
1845:
1843:
1841:
1839:
2714:
2673:
2585:
2583:
2453:
2451:
2394:
2086:Rimmington, Anthony (15 November 2018).
2059:Rimmington, Anthony (15 November 2018).
1761:United States biological weapons program
1702:Institute of Highly Pure Biopreparations
1024:
773:
686:(the causative agent of tularaemia) and
3691:Sanitary epidemiological reconnaissance
3219:Russian Biological and Chemical Weapons
3062:
2931:
2590:DYBVIK, RUSSELL E (14 September 1992).
2457:
2337:Rimmington, Anthony (3 November 2021).
2198:Rimmington, Anthony (15 October 2018).
2164:Rimmington, Anthony (15 October 2018).
1340:Pokrov Factory of Biologics (1967-1991)
1152:Nunn–Lugar Cooperative Threat Reduction
626:. Buildings at the site belonging to a
3704:
3187:History of Biowarfare and Bioterrorism
2801:The Journal of Slavic Military Studies
2589:
2466:
2431:"Monkeypox could be used as bioweapon"
2106:
1976:
1836:
1756:Russia and weapons of mass destruction
1320:(VECTOR), a weaponized smallpox center
1168:Sverdlovsk biological weapons accident
1145:
864:as a biological weapon at Stalingrad.
523:Sverdlovsk biological weapons accident
3242:
2580:
2448:
2343:. Springer International Publishing.
2054:
2052:
2050:
2048:
2011:American Journal of International Law
2004:
1922:
1869:
1785:The Soviet Biological Weapons Program
1176:Joint Statement on Biological Weapons
784:imperialistic and fascistic countries
600:USSR's Revolutionary Military Council
3270:Soviet program of biological weapons
3159:by Dany Shoham and Ze'ev Wolfson,
2795:Rimmington, Anthony (1 March 2003).
2594:. Federation of American Scientists.
2518:
2307:Rimmington, Anthony (20 July 2021).
1949:
1798:
1796:
1779:
1777:
1775:
1644:
1641:List of Soviet/Russian bioweaponeers
1469:
1388:Zagorsk smallpox production facility
1369:Kirov bioweapons production facility
1223:
1199:Russian Foreign Intelligence Service
1017:under the guise of legitimate civil
482:Venezuelan equine encephalitis virus
2771:Conversion of Former BTW Facilities
1632:. The development was conducted in
1512:to authoritarian neighbors such as
1381:, a weaponized anthrax center, see
1160:President of the Russian Federation
542:
13:
2563:"Perspective, Volume IX, Number 1"
2153:. UK National Archives. 1924–1926.
2045:
1998:
1930:"Text of the 1925 Geneva Protocol"
1434:
1412:
1355:Vozrozhdeniya (Renaissance) Island
922:In the immediate post-war period,
730:Leningrad Military Medical Academy
570:'s Military-Chemical Directorate (
14:
3728:
3712:Soviet biological weapons program
3507:Institute of Applied Biochemistry
3129:
3039:"Interview: Dr Kanatjan Alibekov"
2634:; 2005. Retrieved August 9, 2006.
2397:Bulletin of the Atomic Scientists
1916:Bulletin of the Atomic Scientists
1808:United States Department of State
1793:
1772:
1423:
1294:Institute of Applied Biochemistry
1090:A production line to manufacture
1053:, and other state organizations.
977:The USSR was a signatory of 1972
692:(the causative agent of plague).
622:, around 30 miles to the west of
3213:Critical Reviews in Microbiology
3206:Critical Reviews in Microbiology
3161:Critical Reviews in Microbiology
1648:
1623:
1516:during the decade following the
1489:. From the 1970s, it focused on
1227:
957:. Operating under the codename "
582:, Fishman initiated research on
339:
338:
3681:U.S. biological weapons program
3549:Institute of Virus Preparations
3102:
3084:
3031:
3018:
2993:
2971:Interview with General Burgasov
2959:
2925:
2827:
2788:
2761:
2734:
2658:
2637:
2619:
2555:
2538:
2512:
2423:
2369:
2357:
2300:
2273:
2246:
2233:
2157:
2141:
2079:
1720:, died of Marburg virus disease
1518:dissolution of the Soviet Union
1142:in the beginning of the 1990s.
917:
769:
3183:, Vol. 302, pp. 222 – 223
3070:"Obituary: Vladimir Pasechnik"
2934:"Russia's Poorly Guarded Past"
2863:Ainscough, Michael J. (2004).
1904:
1821:
1495:genetically modified organisms
1039:Soviet Ministry of Agriculture
720:Kanatzhan Baizakovich Alibekov
572:Voenno-khimicheskoe upravlenie
1:
3686:History of biological warfare
3671:Biological Weapons Convention
2967:"Smallpox - not a bad weapon"
2616:. Retrieved January 18, 2006.
2127:Alibek, K. and S. Handelman.
2005:Beard, Jack M. (April 2007).
1766:
1751:History of biological warfare
1508:program successfully averted
979:Biological Weapons Convention
836:, a former senior manager of
760:pneumonic plague in Manchuria
592:Kliment Efremovich Voroshilov
399:Biological Weapons Convention
388:Biological Weapons Convention
3319:Staphylococcal Enterotoxin B
2932:Warrick, Jo (17 June 2002).
2891:Rimmington, Anthony (2021).
2747:. Harvard University Press.
2286:. Harvard University Press.
2253:Rimmington, Anthony (2018).
1326:USSR Ministry of Agriculture
955:USSR Ministry of Agriculture
493:Staphylococcal enterotoxin B
7:
3717:Weapons of the Soviet Union
3108:Birstein, Vadim J. (2004),
2259:. Oxford University Press.
2204:. Oxford University Press.
2170:. Oxford University Press.
2092:. Oxford University Press.
2065:. Oxford University Press.
1744:
1537:
1506:Biological Threat Reduction
852:troops during the critical
750:Secret Intelligence Service
608:Ivan Mikhailovich Velikanov
25:Weapons of mass destruction
10:
3733:
3570:Nikolay Timofeev-Ressovsky
2664:Leitenberg and Zilinskas,
1884:Verification Yearbook 2002
1694:, also known as Ken Alibek
1588:
1584:
1541:
1473:
1309:Omutninsk Chemical Factory
1035:Soviet Ministry of Defense
537:
3653:
3632:
3562:
3485:Facilities and test sites
3484:
3463:
3357:
3276:
3173:Online, February 18, 2004
2901:10.1007/978-3-030-73843-3
2813:10.1080/13518040308430546
2686:, Greenhill Books, 2006,
2490:10.1080/10408410490468812
2150:Bacterial warfare: Russia
2023:10.1017/S0002930000030098
1043:Soviet Ministry of Health
942:virus was established in
788:Lavrentiy Pavlovich Beria
2653:Harvard University Press
2649:Cambridge, Massachusetts
2380:Retrieved 13 April 2020.
1881:; Meier, Oliver (eds.).
1346:USSR Ministry of Defence
1284:, Leningrad (1974-1991);
1047:USSR Academy of Sciences
995:USSR Academy of Sciences
897:NKVD special camp No. 48
649:, seized control of the
634:(the causative agent of
586:(the causative agent of
564:Yakov Moiseevich Fishman
3666:Biological warfare (BW)
3640:Sverdlovsk anthrax leak
3464:Operations and projects
3235:, by Jonathan B. Tucker
1591:Sverdlovsk anthrax leak
1493:and the development of
1383:Sverdlovsk anthrax leak
1304:Institute of Immunology
1288:Berdsk Chemical Factory
987:Ministry of Agriculture
645:), a forerunner of the
3645:Aral smallpox incident
2895:. Palgrave Macmillan.
2724:Westview Press (2004)
2632:US Department of State
1612:, according to former
1544:Aral smallpox incident
1306:, Chekhov (1978-1991);
1115:genetically engineered
1073:foot-and-mouth disease
871:(Nomonhan) by Japan's
845:Francisella tularensis
779:
701:Francisella tularensis
683:Francisella tularensis
632:Francisella tularensis
435:Francisella tularensis
3676:Entomological warfare
3179:, by Gretchen Vogel,
2604:M Leitenberg (2001),
1870:Kelly, David (2002).
1499:Counter-proliferation
1109:In 1989 the defector
1065:personally authorized
1025:Post-BWC developments
777:
3142:, February 25, 1998]
2682:Alexander Kouzminov
2608:, College Park, Md:
2550:Simon & Schuster
2478:Crit. Rev. Microbiol
2378:Google Books website
1662:adding missing items
1578:Vozrozhdeniya Island
1558:Vozrozhdeniya Island
1241:adding missing items
854:Battle of Stalingrad
804:Operation Barbarossa
705:Operation Barbarossa
697:Vozrozhdeniya Island
651:Intercession Convent
3028:; Prometheus Books.
2720:Vadim J. Birstein.
2409:1999BuAtS..55d..69S
2313:. Springer Nature.
2131:1999. Delta (2000)
1918:. 27 November 2019.
1510:technology transfer
1373:Kirov, Kirov Oblast
1363:Komsomolskiy Island
1146:The post-Soviet era
1081:African swine fever
983:Ministry of Defense
447:Burkholderia mallei
3580:Vladimir Pasechnik
2981:on 14 October 2007
2630:, Washington, DC:
1698:Vladimir Pasechnik
1692:Kanatzhan Alibekov
1660:; you can help by
1601:Bacillus anthracis
1239:; you can help by
1158:In the 1990s, the
1111:Vladimir Pasechnik
1033:") along with the
991:Ministry of Health
790:, the head of the
780:
596:People's Commissar
584:Bacillus anthracis
559:" agreement only.
527:Vladimir Pasechnik
411:Bacillus anthracis
392:Russian Federation
3699:
3698:
3575:Kanatjan Alibekov
3358:Researched agents
3277:Weaponized agents
2910:978-3-030-73842-6
2849:978-0-7425-2468-2
2781:978-0-7923-5249-5
2754:978-0-674-06526-0
2462:. The New Yorker.
2417:10.2968/055004019
2350:978-3-030-82881-3
2320:978-3-030-73843-3
2293:978-0-674-06526-0
2266:978-0-19-094314-1
2211:978-0-19-005023-8
2177:978-0-19-005034-4
2099:978-0-19-092885-8
2072:978-0-19-092885-8
1897:978-1-899548-35-4
1729:Sergey Rumyantsev
1678:
1677:
1491:molecular biology
1487:Gorodomlya Island
1470:Project Ekologiya
1458:at Obolensk, and
1257:
1256:
1172:Kanatjan Alibekov
1133:Margaret Thatcher
1129:George H. W. Bush
1127:. This triggered
1062:Mikhail Gorbachev
1015:biological agents
672:Gorodomlya Island
574:, abbreviated to
531:Kanatjan Alibekov
471:Coxiella burnetii
380:
379:
3724:
3661:Biological agent
3625:
3612:Alexander Catsch
3603:
3590:Yuri Ovchinnikov
3554:Laboratory No. 1
3540:
3263:
3256:
3249:
3240:
3239:
3124:
3106:
3100:
3088:
3082:
3081:
3079:
3077:
3066:
3060:
3059:
3057:
3055:
3035:
3029:
3022:
3016:
3015:
3013:
3011:
2997:
2991:
2990:
2988:
2986:
2977:. Archived from
2963:
2957:
2956:
2954:
2952:
2929:
2923:
2922:
2888:
2879:
2878:
2876:
2874:
2869:
2860:
2854:
2853:
2831:
2825:
2824:
2792:
2786:
2785:
2765:
2759:
2758:
2738:
2732:
2718:
2712:
2711:
2709:
2707:
2702:on 25 April 2005
2698:. Archived from
2680:
2671:
2662:
2656:
2641:
2635:
2623:
2617:
2602:
2596:
2595:
2587:
2578:
2577:
2575:
2573:
2559:
2553:
2548:, New York, NY:
2542:
2536:
2535:
2533:
2531:
2519:Stronski, Paul.
2516:
2510:
2509:
2473:
2464:
2463:
2455:
2446:
2445:
2443:
2441:
2427:
2421:
2420:
2392:
2381:
2373:
2367:
2361:
2355:
2354:
2334:
2325:
2324:
2304:
2298:
2297:
2277:
2271:
2270:
2250:
2244:
2237:
2231:
2225:
2216:
2215:
2195:
2182:
2181:
2161:
2155:
2154:
2145:
2139:
2125:
2104:
2103:
2083:
2077:
2076:
2056:
2043:
2042:
2002:
1996:
1995:
1993:
1991:
1980:
1974:
1973:
1971:
1969:
1953:
1947:
1946:
1944:
1942:
1926:
1920:
1919:
1908:
1902:
1901:
1889:
1876:
1867:
1834:
1825:
1819:
1818:
1816:
1814:
1800:
1791:
1781:
1713:Yuri Ovchinnikov
1687:Igor Domaradskiy
1673:
1670:
1652:
1651:
1645:
1634:Vector Institute
1616:deputy director
1464:chimeric viruses
1447:from Venezuelan
1252:
1249:
1231:
1230:
1224:
1096:Vector Institute
1007:Glavmikrobioprom
909:
858:Erhardt Geissler
764:Fort Alexander 1
659:
543:Pre-World War II
372:
365:
358:
342:
341:
323:List of treaties
35:
21:
20:
3732:
3731:
3727:
3726:
3725:
3723:
3722:
3721:
3702:
3701:
3700:
3695:
3649:
3628:
3619:
3607:Nikolai Ustinov
3597:
3595:Igor Domaradsky
3558:
3534:
3480:
3471:Project Bonfire
3459:
3353:
3339:Wheat stem rust
3272:
3267:
3138:By Tim Werner,
3132:
3127:
3107:
3103:
3089:
3085:
3075:
3073:
3068:
3067:
3063:
3053:
3051:
3037:
3036:
3032:
3023:
3019:
3009:
3007:
3005:www.science.org
2999:
2998:
2994:
2984:
2982:
2965:
2964:
2960:
2950:
2948:
2938:Washington Post
2930:
2926:
2911:
2889:
2882:
2872:
2870:
2867:
2861:
2857:
2850:
2832:
2828:
2793:
2789:
2782:
2766:
2762:
2755:
2739:
2735:
2719:
2715:
2705:
2703:
2694:
2681:
2674:
2663:
2659:
2642:
2638:
2624:
2620:
2603:
2599:
2588:
2581:
2571:
2569:
2561:
2560:
2556:
2543:
2539:
2529:
2527:
2517:
2513:
2474:
2467:
2456:
2449:
2439:
2437:
2429:
2428:
2424:
2393:
2384:
2374:
2370:
2362:
2358:
2351:
2335:
2328:
2321:
2305:
2301:
2294:
2278:
2274:
2267:
2251:
2247:
2238:
2234:
2226:
2219:
2212:
2196:
2185:
2178:
2162:
2158:
2147:
2146:
2142:
2126:
2107:
2100:
2084:
2080:
2073:
2057:
2046:
2003:
1999:
1989:
1987:
1982:
1981:
1977:
1967:
1965:
1955:
1954:
1950:
1940:
1938:
1928:
1927:
1923:
1910:
1909:
1905:
1898:
1887:
1879:Findlay, Trevor
1874:
1868:
1837:
1826:
1822:
1812:
1810:
1802:
1801:
1794:
1782:
1773:
1769:
1747:
1739:Tsyren Khanduev
1718:Nikolai Ustinov
1674:
1668:
1665:
1649:
1643:
1626:
1593:
1587:
1546:
1540:
1535:
1520:. According to
1501:efforts of the
1478:
1472:
1441:Chimera Project
1437:
1435:Chimera Project
1426:
1418:Project Bonfire
1415:
1413:Project Bonfire
1253:
1247:
1244:
1228:
1222:
1148:
1120:Yersinia pestis
1058:Leonid Brezhnev
1027:
920:
903:
772:
755:Yersinia pestis
746:Solovetsky camp
689:Yersinia pestis
653:
612:botulinum toxin
610:, an expert on
549:Geneva Protocol
545:
540:
488:Botulinum toxin
423:Yersinia pestis
376:
17:
12:
11:
5:
3730:
3720:
3719:
3714:
3697:
3696:
3694:
3693:
3688:
3683:
3678:
3673:
3668:
3663:
3657:
3655:
3654:Related topics
3651:
3650:
3648:
3647:
3642:
3636:
3634:
3630:
3629:
3627:
3626:
3617:Pyotr Burgasov
3614:
3609:
3604:
3592:
3587:
3582:
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3514:
3509:
3504:
3499:
3494:
3488:
3486:
3482:
3481:
3479:
3478:
3476:Project Factor
3473:
3467:
3465:
3461:
3460:
3458:
3457:
3452:
3447:
3442:
3437:
3432:
3427:
3422:
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3407:
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3397:
3392:
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3359:
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3306:
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3296:
3291:
3286:
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3274:
3273:
3266:
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3258:
3251:
3243:
3237:
3236:
3230:
3216:
3209:
3199:
3194:
3189:
3184:
3174:
3164:
3154:
3149:
3148:, May 20, 1998
3143:
3140:New York Times
3131:
3130:External links
3128:
3126:
3125:
3114:Westview Press
3101:
3083:
3061:
3030:
3017:
2992:
2973:(in Russian).
2958:
2924:
2909:
2880:
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2787:
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2760:
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2733:
2713:
2672:
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2579:
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2525:Foreign Policy
2511:
2465:
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2368:
2356:
2349:
2326:
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2299:
2292:
2272:
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2183:
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2098:
2078:
2071:
2044:
2017:(2): 277–284.
1997:
1975:
1948:
1921:
1903:
1896:
1835:
1820:
1792:
1770:
1768:
1765:
1764:
1763:
1758:
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1742:
1741:
1736:
1731:
1726:
1724:Pyotr Burgasov
1721:
1715:
1710:
1705:
1695:
1689:
1684:
1682:Ivan Velikanov
1676:
1675:
1655:
1653:
1642:
1639:
1625:
1622:
1618:Kenneth Alibek
1589:Main article:
1586:
1583:
1582:
1581:
1542:Main article:
1539:
1536:
1534:
1531:
1524:(2021), "The
1471:
1468:
1436:
1433:
1429:Project Factor
1425:
1424:Project Factor
1422:
1414:
1411:
1407:
1406:
1401:
1400:
1399:
1396:Sergiyev Posad
1385:
1375:
1366:
1343:
1342:
1341:
1338:
1323:
1322:
1321:
1315:
1310:
1307:
1301:
1291:
1285:
1279:
1273:
1255:
1254:
1234:
1232:
1221:
1218:
1206:Vladimir Putin
1147:
1144:
1140:Saddam Hussein
1100:Kenneth Alibek
1083:for pigs, and
1079:against cows,
1026:
1023:
924:Lavrenty Beria
919:
916:
912:Ivanovo Oblast
771:
768:
544:
541:
539:
536:
511:
510:
505:
500:
495:
490:
485:
479:
467:
455:
443:
431:
419:
378:
377:
375:
374:
367:
360:
352:
349:
348:
347:
346:
333:
332:
331:
330:
327:treaty parties
317:
316:
312:
311:
310:
309:
304:
299:
291:
290:
286:
285:
284:
283:
278:
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271:United Kingdom
268:
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27:
15:
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3638:
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3635:
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3613:
3610:
3608:
3605:
3601:
3596:
3593:
3591:
3588:
3586:
3583:
3581:
3578:
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3573:
3571:
3568:
3567:
3565:
3561:
3555:
3552:
3550:
3547:
3545:
3542:
3538:
3533:
3532:Sverdlovsk-19
3530:
3528:
3527:NPO Biosintez
3525:
3523:
3520:
3518:
3515:
3513:
3510:
3508:
3505:
3503:
3500:
3498:
3495:
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3483:
3477:
3474:
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3448:
3446:
3443:
3441:
3438:
3436:
3433:
3431:
3428:
3426:
3423:
3421:
3420:Potato blight
3418:
3416:
3413:
3411:
3408:
3406:
3403:
3401:
3398:
3396:
3393:
3391:
3388:
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3297:
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3150:
3147:
3144:
3141:
3137:
3134:
3133:
3122:
3121:0-8133-4280-5
3118:
3115:
3111:
3105:
3098:
3097:
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2928:
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2898:
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2887:
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2866:
2859:
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2837:
2830:
2822:
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2810:
2806:
2802:
2798:
2791:
2783:
2777:
2773:
2772:
2764:
2756:
2750:
2746:
2745:
2737:
2731:
2730:0-8133-4280-5
2727:
2723:
2717:
2701:
2697:
2693:
2692:1-85367-646-2
2689:
2685:
2679:
2677:
2670:, pp 643-644.
2669:
2668:
2661:
2654:
2650:
2646:
2640:
2633:
2629:
2628:
2622:
2615:
2611:
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2558:
2551:
2547:
2541:
2526:
2522:
2515:
2507:
2503:
2499:
2495:
2491:
2487:
2484:(4): 241–61.
2483:
2479:
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2454:
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2436:
2432:
2426:
2418:
2414:
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2222:
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2207:
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2194:
2192:
2190:
2188:
2179:
2173:
2169:
2168:
2160:
2152:
2151:
2144:
2138:
2137:0-385-33496-6
2134:
2130:
2124:
2122:
2120:
2118:
2116:
2114:
2112:
2110:
2101:
2095:
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2090:
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2074:
2068:
2064:
2063:
2055:
2053:
2051:
2049:
2040:
2036:
2032:
2028:
2024:
2020:
2016:
2012:
2008:
2001:
1985:
1979:
1964:
1963:
1958:
1952:
1937:
1936:
1931:
1925:
1917:
1913:
1907:
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1771:
1762:
1759:
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1754:
1752:
1749:
1748:
1740:
1737:
1735:
1734:Boris Kopylev
1732:
1730:
1727:
1725:
1722:
1719:
1716:
1714:
1711:
1709:
1706:
1703:
1699:
1696:
1693:
1690:
1688:
1685:
1683:
1680:
1679:
1672:
1663:
1659:
1656:This list is
1654:
1647:
1646:
1638:
1635:
1631:
1630:Marburg virus
1624:Marburg virus
1621:
1619:
1615:
1611:
1607:
1603:
1602:
1597:
1592:
1579:
1575:
1571:
1567:
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1402:
1397:
1393:
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1360:
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1348:
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1339:
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1319:
1316:
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1311:
1308:
1305:
1302:
1299:
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1292:
1289:
1286:
1283:
1280:
1277:
1274:
1272:
1268:
1265:
1264:
1262:
1259:
1258:
1251:
1242:
1238:
1235:This list is
1233:
1226:
1225:
1217:
1213:
1211:
1207:
1202:
1200:
1196:
1191:
1189:
1185:
1180:
1177:
1173:
1169:
1165:
1164:Boris Yeltsin
1161:
1156:
1153:
1143:
1141:
1136:
1134:
1130:
1126:
1122:
1121:
1116:
1112:
1107:
1105:
1101:
1097:
1093:
1088:
1086:
1082:
1078:
1074:
1069:
1066:
1063:
1059:
1054:
1052:
1048:
1044:
1040:
1036:
1032:
1022:
1020:
1019:biotechnology
1016:
1012:
1008:
1004:
1000:
996:
992:
988:
984:
980:
975:
973:
968:
962:
960:
956:
952:
947:
945:
941:
937:
931:
929:
925:
915:
913:
907:
902:
898:
894:
890:
886:
882:
878:
874:
870:
865:
863:
862:F. tularensis
859:
855:
851:
847:
846:
841:
840:
835:
831:
830:
824:
822:
818:
814:
810:
805:
800:
797:
793:
789:
785:
776:
767:
765:
761:
757:
756:
751:
747:
743:
739:
735:
731:
727:
726:
721:
716:
714:
710:
706:
702:
698:
693:
691:
690:
685:
684:
679:
678:
673:
669:
668:
663:
657:
652:
648:
644:
639:
637:
633:
629:
625:
621:
617:
613:
609:
605:
601:
597:
593:
589:
585:
581:
577:
573:
569:
565:
560:
558:
554:
550:
535:
532:
528:
524:
520:
519:Boris Yeltsin
516:
509:
508:Orthopoxvirus
506:
504:
503:Marburg virus
501:
499:
496:
494:
491:
489:
486:
483:
480:
477:
473:
472:
468:
465:
461:
460:
456:
453:
449:
448:
444:
441:
437:
436:
432:
429:
425:
424:
420:
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413:
412:
408:
407:
406:
404:
400:
395:
393:
389:
385:
373:
368:
366:
361:
359:
354:
353:
351:
350:
345:
337:
336:
335:
334:
328:
324:
321:
320:
319:
318:
314:
313:
308:
305:
303:
300:
298:
295:
294:
293:
292:
289:Proliferation
288:
287:
282:
279:
277:
276:United States
274:
272:
269:
267:
264:
262:
259:
257:
254:
252:
249:
247:
244:
242:
239:
237:
234:
232:
229:
227:
224:
221:
217:
214:
212:
209:
207:
204:
202:
199:
197:
194:
192:
189:
187:
184:
182:
179:
177:
174:
172:
169:
167:
164:
162:
159:
157:
154:
152:
149:
147:
144:
142:
139:
137:
134:
132:
129:
127:
124:
122:
119:
117:
114:
112:
109:
107:
104:
102:
99:
97:
94:
92:
89:
87:
84:
82:
79:
78:
77:
76:
73:
70:
69:
64:
61:
59:
56:
54:
51:
49:
46:
45:
44:
43:
39:
38:
34:
30:
29:
26:
23:
22:
19:
3585:Sergei Popov
3512:Laboratory B
3455:Yellow fever
3385:Dengue fever
3269:
3212:
3205:
3160:
3109:
3104:
3094:
3086:
3074:. Retrieved
3064:
3052:. Retrieved
3042:
3033:
3025:
3020:
3008:. Retrieved
3004:
2995:
2983:. Retrieved
2979:the original
2970:
2961:
2949:. Retrieved
2937:
2927:
2892:
2871:. Retrieved
2858:
2839:
2829:
2804:
2800:
2790:
2770:
2763:
2743:
2736:
2721:
2716:
2704:. Retrieved
2700:the original
2683:
2665:
2660:
2644:
2639:
2626:
2621:
2605:
2600:
2570:. Retrieved
2566:
2557:
2545:
2540:
2528:. Retrieved
2524:
2514:
2481:
2477:
2459:
2438:. Retrieved
2434:
2425:
2403:(4): 69–71.
2400:
2396:
2371:
2364:
2359:
2339:
2309:
2302:
2282:
2275:
2255:
2248:
2235:
2200:
2166:
2159:
2149:
2143:
2128:
2088:
2081:
2061:
2014:
2010:
2000:
1988:. Retrieved
1978:
1966:. Retrieved
1960:
1951:
1939:. Retrieved
1933:
1924:
1915:
1906:
1883:
1829:
1823:
1811:. Retrieved
1807:
1784:
1708:Sergei Popov
1704:in Leningrad
1666:
1627:
1599:
1594:
1547:
1521:
1479:
1452:encephalitis
1440:
1438:
1427:
1416:
1408:
1245:
1214:
1203:
1192:
1183:
1181:
1157:
1149:
1137:
1118:
1117:a strain of
1108:
1089:
1070:
1064:
1055:
1028:
1010:
1006:
1002:
976:
970:
966:
963:
958:
950:
948:
935:
932:
921:
918:The Cold War
893:Otozō Yamada
866:
861:
843:
837:
827:
825:
816:
812:
808:
801:
795:
783:
781:
770:World War II
753:
723:
717:
700:
694:
687:
681:
675:
665:
646:
640:
631:
603:
583:
579:
575:
571:
561:
557:no-first-use
553:reservations
546:
529:and Colonel
512:
469:
457:
445:
433:
421:
409:
396:
394:after that.
384:Soviet Union
381:
231:South Africa
226:Saudi Arabia
220:Soviet Union
219:
63:Radiological
18:
3620: [
3598: [
3535: [
3492:BIOPREPARAT
3425:Psittacosis
3405:Melioidosis
3400:Lassa fever
3304:Brucellosis
3096:NOVA Online
2975:Moscow News
2873:9 September
2807:(1): 1–43.
1968:16 February
1941:16 February
1614:Biopreparat
1550:Soviet Army
1460:Ebola virus
1337:(1958-1991)
1261:Biopreparat
1125:antibiotics
1085:psittacosis
1031:Biopreparat
1011:Biopreparat
1003:Biopreparat
904: [
877:Shirō Ishii
869:Khalkin Gol
839:Biopreparat
742:melioidosis
725:Biopreparat
654: [
464:brucellosis
251:Switzerland
236:South Korea
196:Philippines
186:North Korea
181:Netherlands
3706:Categories
3563:Scientists
3440:Rinderpest
3395:Hantavirus
3344:Rice blast
2706:5 December
1813:30 October
1767:References
1658:incomplete
1606:Sverdlovsk
1503:Nunn-Lugar
1474:See also:
1379:Sverdlovsk
1237:incomplete
1123:to resist
1077:rinderpest
972:programme"
928:Sverdlovsk
636:tularaemia
403:bio-agents
281:Yugoslavia
72:By country
48:Biological
3633:Accidents
3324:Tularemia
3229:and NGOs.
3093:, (2001)
3044:Frontline
2946:0190-8286
2919:240880486
2821:144757207
2460:pp. 52-65
2031:0002-9300
1483:FMD virus
1298:Omutninsk
1210:Pushchino
1104:monkeypox
959:Ekologiya
885:Manchuria
829:Biohazard
677:Narkomzem
667:sharashka
440:tularemia
96:Australia
91:Argentina
3544:Aralsk-7
3375:Bird flu
3309:Glanders
3299:Botulism
3284:Smallpox
3171:CBC News
2951:22 April
2667:Op. cit.
2655:, pg 14.
2530:18 March
2506:30487628
2498:15646399
2365:Op. cit.
2363:Alibek,
1745:See also
1669:May 2011
1574:Andropov
1570:Alma-Ata
1562:Aral Sea
1538:Smallpox
1456:smallpox
1359:Aral Sea
1351:Aralsk-7
1248:May 2011
1195:Aeroflot
1092:smallpox
940:smallpox
901:Cherntsy
889:Pingfang
881:Unit 100
873:Unit 731
738:glanders
628:smallpox
620:Vlasikha
616:botulism
568:Red Army
515:smallpox
498:Smallpox
459:Brucella
452:glanders
344:Category
315:Treaties
307:Missiles
297:Chemical
206:Rhodesia
191:Pakistan
106:Bulgaria
53:Chemical
3522:Zagorsk
3349:Polonia
3334:Marburg
3314:Q fever
3289:Anthrax
3181:Science
3076:8 March
3054:8 March
2985:18 June
2572:8 April
2405:Bibcode
2039:8354600
1990:3 April
1700:at the
1585:Anthrax
1560:in the
1392:Zagorsk
1365:in 1954
1300:(1994);
1290:(1993);
944:Zagorsk
744:in the
709:Kalinin
588:anthrax
580:VOKhIMU
576:VOKhIMU
538:History
476:Q-fever
416:anthrax
302:Nuclear
266:Ukraine
211:Romania
176:Myanmar
131:Germany
86:Algeria
81:Albania
58:Nuclear
40:By type
3517:Berdsk
3502:VECTOR
3445:Typhus
3415:Plague
3294:Plague
3225:, the
3119:
3010:20 May
2944:
2917:
2907:
2846:
2819:
2778:
2751:
2728:
2690:
2504:
2496:
2440:20 May
2347:
2317:
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2263:
2208:
2174:
2135:
2096:
2069:
2037:
2029:
1894:
1596:Spores
1566:Aralsk
1526:Pokrov
1449:equine
1335:Pokrov
1049:, the
1045:, the
1041:, the
1037:, the
997:, and
875:under
850:panzer
834:Alibek
734:typhus
662:Suzdal
624:Moscow
428:plague
261:Taiwan
246:Sweden
216:Russia
201:Poland
171:Mexico
151:Israel
126:France
111:Canada
101:Brazil
3624:]
3602:]
3539:]
3430:Ricin
3380:CHIKV
2915:S2CID
2868:(PDF)
2817:S2CID
2502:S2CID
2035:S2CID
1888:(PDF)
1877:. In
1875:(PDF)
1188:Putin
908:]
821:Kirov
796:Yurta
713:Kirov
658:]
594:(the
484:(VEE)
462:sp. (
256:Syria
241:Spain
166:Libya
161:Japan
156:Italy
136:India
121:Egypt
116:China
3117:ISBN
3078:2010
3056:2010
3012:2022
2987:2007
2953:2022
2942:ISSN
2905:ISBN
2875:2020
2844:ISBN
2776:ISBN
2749:ISBN
2726:ISBN
2708:2007
2688:ISBN
2574:2015
2532:2022
2494:PMID
2442:2022
2345:ISBN
2315:ISBN
2288:ISBN
2261:ISBN
2206:ISBN
2172:ISBN
2133:ISBN
2094:ISBN
2067:ISBN
2027:ISSN
1992:2021
1970:2021
1943:2021
1892:ISBN
1815:2021
1556:“On
1514:Iran
1454:and
1439:The
1150:The
1131:and
1075:and
967:NITB
936:NIIS
792:NKVD
740:and
647:NKVD
643:OGPU
614:and
382:The
146:Iraq
141:Iran
3450:WEE
3435:RVF
3410:VND
3390:EEE
3370:BHF
3365:AHF
3329:VEE
3227:GAO
3223:CRS
3049:PBS
2897:doi
2809:doi
2486:doi
2435:UPI
2413:doi
2019:doi
2015:101
1664:.
1610:KGB
1598:of
1445:DNA
1243:.
1060:to
1051:KGB
999:KGB
817:STI
813:STI
809:STI
660:in
638:).
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3622:ru
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