114:
170:
997:
142:
1488:
Gauthier JA, Kearney M, Maisano JA, Rieppel O, Behlke AD (2012). "Assembling the
Squamate Tree of Life: Perspectives from the Phenotype and the Fossil Record".
1952:
1214:
Fry BG, Vidal N, Norman JA, Vonk FJ, Scheib H, Ramjan SF, et al. (February 2006). "Early evolution of the venom system in lizards and snakes".
3182:
3208:
837:
598:
298:
412:
of
Australia and Southeast Asia, and various other species with a similarly distinctive appearance. Their closest living relatives are the
1738:
466:, and all descendants of their last common ancestor. Using one of these definitions leads to the inclusion of the earless monitor lizard (
3298:
3011:
1779:
787:
584:
290:
2894:
3156:
1111:"A spectacular new Philippine monitor lizard reveals a hidden biogeographic boundary and a novel flagship species for conservation"
1056:, being larger in larger-bodied species, although the feet become smaller as compared with the lengths of the other limb segments.
3195:
3293:
3200:
2646:
2325:
2520:
1441:"Molecular evidence for an Asian origin of monitor lizards followed by Tertiary dispersals to Africa and Australasia"
1274:
3260:
3213:
1772:
1611:
1265:
de
Queiroz K, Gauthier J (1988). "Phylogenetic Relationships within Squamata". In Estes RJ, Pregill GK (eds.).
1026:
widely and have large home ranges, and many have high stamina. Although most species are carnivorous, three
3104:
3091:
2834:
2628:
1592:
3109:
2746:
1765:
1664:
Diet and
Arboreality in the Emerald Monitor, Varanus prasinus, with Comments on the Study of Adaptation
3252:
169:
3265:
2619:
2379:
3288:
2829:
2409:
2318:
417:
473:
Lee (1997) created a different definition of the
Varanidae, defining them as the clade containing
2839:
1698:
113:
3239:
3044:
2800:
2728:
2719:
2665:
2583:
2460:
486:
462:
413:
385:
1439:
Vidal N, Marin J, Sassi J, Battistuzzi FU, Donnellan S, Fitch AJ, et al. (October 2012).
3234:
3187:
2427:
2418:
2400:
1694:
1588:
353:
3226:
3130:
2755:
2565:
2556:
2502:
1627:"Metabolic rate and endurance capacity in Australian varanid lizards (Squamata; Varanidae;
1541:
1354:
1223:
454:(2008) (also using morphological data), who defined the Varanidae as the clade containing
8:
3082:
2937:
2683:
2529:
2484:
2311:
1298:
Conrad J (2008). "Phylogeny and systematics of
Squamata (Reptilia) based on morphology".
1109:
Welton LJ, Siler CD, Bennett D, Diesmos A, Duya MR, Dugay R, et al. (October 2010).
1545:
1358:
1227:
1171:
Philosophical
Transactions of the Royal Society of London. Series B, Biological Sciences
493:
is included in or excluded from the
Varanidae depends on the author; for example, Vidal
2538:
2493:
1721:
1593:"Lizard home ranges revisited: effects of sex, body size, diet, habitat, and phylogeny"
1564:
1529:
1505:
1465:
1440:
1416:
1391:
1375:
1342:
1323:
1267:
Phylogenetic
Relationships of the Lizard Families: Essays Commemorating Charles L. Camp
1247:
1191:
1166:
1135:
1110:
164:
1752:
1052:) are primarily frugivores. Among species of living varanids, the limbs show positive
3221:
3117:
2764:
2737:
2574:
1883:
1667:
1647:
1626:
1569:
1470:
1421:
1280:
1270:
1239:
1196:
1140:
1036:
1000:
401:
1509:
1327:
3122:
2782:
2610:
2190:
1965:
1873:
1858:
1713:
1642:
1607:
1559:
1549:
1497:
1460:
1452:
1411:
1403:
1370:
1362:
1315:
1307:
1251:
1231:
1186:
1178:
1130:
1122:
1048:
1042:
1022:
Monitor lizards are reputed to be among the most intelligent lizards. Most species
437:
433:
405:
345:
3096:
2901:
2878:
2710:
2656:
2233:
2200:
2143:
1863:
1675:
1554:
1343:"Pectoral Girdles vs. Hyobranchia in the Snake Genera Liotyphlops and Anomalepis"
1072:
889:
421:
29:
3067:
2791:
2692:
2547:
2511:
2475:
2273:
2210:
2180:
2123:
2098:
2093:
2070:
2050:
1988:
1960:
1906:
1167:"A new stem-varanid lizard (Reptilia, Squamata) from the early Eocene of China"
1077:
914:
658:
314:
246:
3282:
2887:
2809:
2592:
2228:
2218:
2185:
2172:
2148:
2113:
2103:
2040:
2035:
2013:
1868:
1087:
855:
612:
542:
498:
397:
306:
122:
71:
1671:
1284:
497:(2012) classify the earless monitor lizard as a member of a separate family
3004:
2983:
2955:
2701:
2637:
2601:
2436:
2288:
2253:
2153:
2128:
2055:
1980:
1739:
Unidirectional
Airflow In The Lungs Of Birds, Crocs And Now Monitor Lizards
1679:
1573:
1530:"Earliest example of a giant monitor lizard (Varanus, Varanidae, Squamata)"
1474:
1456:
1425:
1407:
1366:
1243:
1200:
1182:
1144:
1126:
1082:
456:
432:
The Varanidae were defined (using morphological characteristics) by Estes,
450:
and all of its descendants. A similar definition was formulated by Conrad
3169:
3076:
2997:
2990:
2969:
2962:
2773:
2451:
2278:
2263:
2243:
2195:
2133:
2108:
2065:
2027:
2008:
1930:
1916:
1878:
1501:
1031:
1012:
769:
744:
712:
626:
570:
556:
442:
360:
282:
274:
266:
46:
1235:
2334:
2268:
2248:
1942:
1901:
1893:
1853:
1725:
356:
231:
91:
56:
1699:"Scaling of limb proportions in monitor lizards (Squamata: Varanidae)"
3174:
3143:
2976:
2920:
2392:
2223:
2163:
2138:
2118:
1993:
1835:
1319:
1053:
1016:
689:
368:
364:
181:
133:
96:
40:
3161:
3038:
1757:
1717:
1165:
Dong L, Wang YQ, Zhao Q, Vasilyan D, Wang Y, Evans SE (March 2022).
996:
3148:
3061:
2674:
2364:
2358:
2352:
2258:
2088:
2060:
2003:
1998:
1926:
1819:
1807:
1788:
1060:
1027:
1008:
221:
201:
86:
81:
66:
61:
51:
33:
1311:
440:(1988) as the clade containing the most recent common ancestor of
3018:
2080:
1970:
1813:
931:
673:
520:
376:
329:
211:
127:
101:
76:
2930:
2862:
2346:
2238:
1801:
1023:
648:
640:
409:
349:
322:
255:
191:
156:
151:
141:
3247:
3135:
2303:
2283:
1911:
1845:
1487:
1059:
Varanids possess unidirectional pulmonary airflow, including
527:
372:
1612:
10.1890/0012-9658(2002)083[1870:LHRREO]2.0.CO;2
1686:
1438:
1108:
1624:
380:
and a number of extinct genera more closely related to
1580:
1527:
420:. The oldest members of the family are known from the
2875:
1164:
654:
636:
622:
608:
594:
580:
566:
552:
538:
516:
1692:
1618:
1300:Bulletin of the American Museum of Natural History
1264:
1213:
1490:Bulletin of the Peabody Museum of Natural History
3280:
1586:
2319:
1773:
1625:Clemente CJ, Withers PC, Thompson GG (2009).
1392:"Molecular evidence and marine snake origins"
505:(2012) classify it as a member of Varanidae.
1666:. Chicago: Field Museum of Natural History.
1528:Conrad JL, Balcarcel AM, Mehling CM (2012).
1104:
1102:
2326:
2312:
1780:
1766:
1269:. Stanford University Press. p. 166.
485:; this definition explicitly excludes the
140:
112:
1646:
1635:Biological Journal of the Linnean Society
1563:
1553:
1523:
1521:
1519:
1464:
1415:
1374:
1291:
1190:
1134:
1099:
1258:
995:
530:included in Varanidae according to Dong
3281:
1661:
1655:
1516:
1340:
1297:
1207:
3043:
3042:
2860:
2377:
2307:
1787:
1761:
1334:
477:and all taxa more closely related to
3253:592C4D59-0A61-FFF1-2BC0-3E8E5D32B4E3
2647:Northern Sierra Madre forest monitor
2521:Black-spotted ridge-tailed monitor (
1353:(2686). The Royal Society: 720β722.
1160:
1158:
1156:
1154:
155:, an extinct varanid known from the
1389:
13:
3299:Extant Campanian first appearances
2378:
14:
3310:
1746:
1151:
1648:10.1111/j.1095-8312.2009.01207.x
168:
44:
2861:
1732:
547:Norell, Gao, & Conrad, 2008
2333:
1481:
1432:
1383:
1:
1093:
400:(the largest living lizard),
371:lizards, includes the living
3294:Taxa named by Blasius Merrem
2747:Short-tailed pygmy monitor (
1555:10.1371/journal.pone.0041767
683:
489:from the Varanidae. Whether
7:
1066:
856:Proplatynotia longirostrata
633:(Mongolia, Late Cretaceous)
619:(Mongolia, Late Cretaceous)
605:(Mongolia, Late Cretaceous)
591:(Mongolia, Late Cretaceous)
577:(Mongolia, Late Cretaceous)
563:(Mongolia, Late Cretaceous)
549:(Mongolia, Late Cretaceous)
470:) in the family Varanidae.
427:
10:
3315:
2629:Mitchell's water monitor (
2419:Rio Fuerte beaded lizard (
2410:Guatemalan beaded lizard (
991:
3051:
2947:
2911:
2874:
2870:
2856:
2822:
2473:
2449:
2391:
2387:
2373:
2341:
2209:
2171:
2162:
2079:
2026:
1979:
1951:
1938:
1925:
1892:
1844:
1828:
1796:
928:
910:
903:
885:
878:
851:
833:
826:
819:
783:
765:
758:
740:
733:
726:
708:
701:
508:
363:. The family, a group of
261:
254:
165:Scientific classification
163:
148:
139:
120:
111:
23:
2801:Yellow-spotted monitor (
2666:Peach-throated monitor (
2620:Merten's water monitor (
2584:Kimberely rock monitor (
2461:Earless monitor lizard (
1063:akin to those of birds.
838:Paravaranus angustifrons
647:(Europe, North America,
418:Chinese crocodile lizard
2720:Rennel Island monitor (
2428:Mexican beaded lizard (
2401:Chiapan beaded lizard (
692:from Dong et al. 2022.
617:Borsuk-Bialynicka, 1984
603:Borsuk-Bialynicka, 1984
589:Borsuk-Bialynicka, 1984
575:Borsuk-Bialynicka, 1984
2756:Spiny-tailed monitor (
2566:Emerald tree monitor (
2503:Black-headed monitor (
1706:Journal of Herpetology
1457:10.1098/rsbl.2012.0460
1408:10.1098/rsbl.2004.0282
1367:10.1098/rstb.1997.0005
1341:Warner R (June 1946).
1183:10.1098/rstb.2021.0041
1127:10.1098/rsbl.2010.0119
1019:
1005:Varanus olivaceus
788:Saniwides mongoliensis
487:earless monitor lizard
463:Lanthanotus borneensis
414:earless monitor lizard
386:earless monitor lizard
3235:Paleobiology Database
2729:Rosenberg's monitor (
2684:Pygmy mulga monitor (
2530:Blue-tailed monitor (
2485:Asian water monitor (
999:
2494:Black tree monitor (
1502:10.3374/014.053.0101
1390:Lee MS (June 2005).
745:Telmasaurus grangeri
561:Gao and Norell, 2000
2938:Varanus amnhophilis
2765:Turquoise monitor (
2738:Roughneck monitor (
2557:Dumeril's monitor (
2539:Crocodile monitor (
1546:2012PLoSO...741767C
1359:1997RSPTB.352...53L
1236:10.1038/nature04328
1228:2006Natur.439..584F
515:Genera marked with
2835:Cretaceous lizards
2823:Related categories
2783:Savannah monitor (
2611:Mangrove monitor (
1662:Greene HW (1986).
1177:(1847): 20210041.
1020:
915:Archaeovaranus lii
770:Aiolosaurus oriens
3276:
3275:
3222:Open Tree of Life
3045:Taxon identifiers
3036:
3035:
3032:
3031:
3028:
3027:
2852:
2851:
2848:
2847:
2785:V. exanthematicus
2711:Pilbara monitor (
2657:Peacock monitor (
2412:H. charlesbogerti
2301:
2300:
2297:
2296:
2022:
2021:
1953:Gymnophthalmoidea
1884:Sphaerodactylidae
1222:(7076): 584β588.
1037:Varanus olivaceus
1011:varanid from the
988:
987:
979:
978:
970:
969:
961:
960:
952:
951:
943:
942:
867:
866:
808:
807:
799:
798:
679:
668:
646:
632:
618:
604:
590:
576:
562:
548:
523:
501:, while Gauthier
402:crocodile monitor
338:
337:
250:
159:of North America
16:Family of lizards
3306:
3269:
3268:
3256:
3255:
3243:
3242:
3230:
3229:
3217:
3216:
3204:
3203:
3191:
3190:
3178:
3177:
3165:
3164:
3152:
3151:
3139:
3138:
3126:
3125:
3113:
3112:
3100:
3099:
3087:
3086:
3085:
3072:
3071:
3070:
3040:
3039:
2877:
2872:
2871:
2858:
2857:
2792:Yellow monitor (
2767:V. caerulivirens
2693:Quince monitor (
2575:Gray's monitor (
2548:Desert monitor (
2512:Bengal monitor (
2389:
2388:
2375:
2374:
2328:
2321:
2314:
2305:
2304:
2191:Leptotyphlopidae
2169:
2168:
1966:Gymnophthalmidae
1936:
1935:
1874:Phyllodactylidae
1859:Carphodactylidae
1782:
1775:
1768:
1759:
1758:
1741:
1736:
1730:
1729:
1703:
1690:
1684:
1683:
1659:
1653:
1652:
1650:
1622:
1616:
1615:
1606:(7): 1870β1885.
1597:
1584:
1578:
1577:
1567:
1557:
1525:
1514:
1513:
1485:
1479:
1478:
1468:
1436:
1430:
1429:
1419:
1387:
1381:
1380:
1378:
1338:
1332:
1331:
1295:
1289:
1288:
1262:
1256:
1255:
1211:
1205:
1204:
1194:
1162:
1149:
1148:
1138:
1106:
1049:Varanus bitatawa
1043:Varanus mabitang
906:
905:
881:
880:
829:
828:
822:
821:
761:
760:
736:
735:
729:
728:
704:
703:
696:
695:
677:
662:
656:
644:
638:
630:
624:
616:
610:
602:
596:
588:
582:
574:
568:
560:
554:
546:
540:
518:
514:
406:savannah monitor
245:
173:
172:
144:
116:
106:
43:
27:Temporal range:
21:
20:
3314:
3313:
3309:
3308:
3307:
3305:
3304:
3303:
3289:Lizard families
3279:
3278:
3277:
3272:
3264:
3259:
3251:
3246:
3238:
3233:
3225:
3220:
3212:
3207:
3199:
3194:
3186:
3181:
3173:
3168:
3160:
3155:
3147:
3142:
3134:
3129:
3121:
3116:
3108:
3103:
3095:
3090:
3081:
3080:
3075:
3066:
3065:
3060:
3047:
3037:
3024:
2943:
2923:Varanus priscus
2907:
2902:Paranecrosaurus
2879:Palaeovaranidae
2866:
2844:
2830:Monitor lizards
2818:
2810:Yemen monitor (
2593:Komodo dragon (
2469:
2445:
2383:
2369:
2337:
2332:
2302:
2293:
2254:Cylindrophiidae
2234:Atractaspididae
2205:
2201:Xenotyphlopidae
2158:
2144:Phrynosomatidae
2075:
2018:
1975:
1947:
1921:
1888:
1864:Diplodactylidae
1840:
1824:
1792:
1786:
1749:
1744:
1737:
1733:
1718:10.2307/1565513
1701:
1691:
1687:
1660:
1656:
1623:
1619:
1595:
1585:
1581:
1526:
1517:
1486:
1482:
1445:Biology Letters
1437:
1433:
1396:Biology Letters
1388:
1384:
1339:
1335:
1296:
1292:
1277:
1263:
1259:
1212:
1208:
1163:
1152:
1115:Biology Letters
1107:
1100:
1096:
1073:Aigialosauridae
1069:
1030:species in the
1015:that primarily
994:
989:
980:
971:
962:
953:
944:
890:Saniwa ensidens
868:
809:
800:
686:
511:
430:
422:Late Cretaceous
244:
167:
107:
105:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
38:
37:
30:Late Cretaceous
28:
25:
17:
12:
11:
5:
3312:
3302:
3301:
3296:
3291:
3274:
3273:
3271:
3270:
3257:
3244:
3231:
3218:
3205:
3192:
3179:
3166:
3153:
3140:
3127:
3114:
3101:
3088:
3073:
3057:
3055:
3049:
3048:
3034:
3033:
3030:
3029:
3026:
3025:
3023:
3022:
3015:
3008:
3001:
2994:
2987:
2980:
2973:
2966:
2959:
2951:
2949:
2945:
2944:
2942:
2941:
2934:
2927:
2917:
2915:
2909:
2908:
2906:
2905:
2898:
2891:
2883:
2881:
2868:
2867:
2854:
2853:
2850:
2849:
2846:
2845:
2843:
2842:
2840:Helodermatidae
2837:
2832:
2826:
2824:
2820:
2819:
2817:
2816:
2807:
2798:
2789:
2780:
2771:
2762:
2753:
2749:V. brevicauda
2744:
2735:
2731:V. rosenbergi
2726:
2722:V. juxtindicus
2717:
2708:
2704:V. albigularis
2702:Rock monitor (
2699:
2690:
2681:
2672:
2663:
2659:V. auffenbergi
2654:
2644:
2638:Nile monitor (
2635:
2626:
2617:
2608:
2602:Lace monitor (
2599:
2595:V. komodoensis
2590:
2581:
2572:
2563:
2554:
2545:
2536:
2527:
2518:
2514:V. bengalensis
2509:
2500:
2491:
2481:
2479:
2471:
2470:
2468:
2467:
2457:
2455:
2447:
2446:
2444:
2443:
2437:Gila monster (
2434:
2425:
2421:H. exasperatum
2416:
2407:
2397:
2395:
2393:Helodermatidae
2385:
2384:
2371:
2370:
2368:
2367:
2361:
2355:
2349:
2342:
2339:
2338:
2331:
2330:
2323:
2316:
2308:
2299:
2298:
2295:
2294:
2292:
2291:
2286:
2281:
2276:
2274:Tropidophiidae
2271:
2266:
2261:
2256:
2251:
2246:
2241:
2236:
2231:
2226:
2221:
2215:
2213:
2211:Alethinophidia
2207:
2206:
2204:
2203:
2198:
2193:
2188:
2183:
2181:Anomalepididae
2177:
2175:
2166:
2160:
2159:
2157:
2156:
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2141:
2136:
2131:
2126:
2124:Leiocephalidae
2121:
2116:
2111:
2106:
2101:
2099:Corytophanidae
2096:
2094:Chamaeleonidae
2091:
2085:
2083:
2077:
2076:
2074:
2073:
2071:Diploglossidae
2068:
2063:
2058:
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2048:
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2032:
2030:
2024:
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2020:
2019:
2017:
2016:
2011:
2006:
2001:
1996:
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1989:Amphisbaenidae
1985:
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1961:Alopoglossidae
1957:
1955:
1949:
1948:
1946:
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1907:Gerrhosauridae
1904:
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1747:External links
1745:
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1712:(2): 219β230.
1685:
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1641:(3): 664β676.
1617:
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1451:(5): 853β855.
1431:
1402:(2): 227β230.
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1121:(5): 654β658.
1097:
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457:Varanus varius
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315:Archaeovaranus
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2918:
2916:
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2888:Palaeovaranus
2885:
2884:
2882:
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2873:
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2841:
2838:
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2812:V. yemenensis
2808:
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2799:
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2794:V. flavescens
2790:
2788:
2786:
2781:
2779:
2777:
2774:Sand goanna (
2772:
2770:
2768:
2763:
2761:
2759:
2758:V. acanthurus
2754:
2752:
2750:
2745:
2743:
2741:
2740:V. rudicollis
2736:
2734:
2732:
2727:
2725:
2723:
2718:
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2573:
2571:
2569:
2564:
2562:
2560:
2555:
2553:
2551:
2546:
2544:
2542:
2541:V. salvadorii
2537:
2535:
2533:
2528:
2526:
2524:
2519:
2517:
2515:
2510:
2508:
2506:
2501:
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2497:
2492:
2490:
2488:
2483:
2482:
2480:
2478:
2477:
2472:
2466:
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2463:L. borneensis
2459:
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2257:
2255:
2252:
2250:
2247:
2245:
2242:
2240:
2237:
2235:
2232:
2230:
2229:Anomochilidae
2227:
2225:
2222:
2220:
2219:Acrochordidae
2217:
2216:
2214:
2212:
2208:
2202:
2199:
2197:
2194:
2192:
2189:
2187:
2186:Gerrhopilidae
2184:
2182:
2179:
2178:
2176:
2174:
2173:Scolecophidia
2170:
2167:
2165:
2161:
2155:
2152:
2150:
2149:Polychrotidae
2147:
2145:
2142:
2140:
2137:
2135:
2132:
2130:
2127:
2125:
2122:
2120:
2117:
2115:
2114:Hoplocercidae
2112:
2110:
2107:
2105:
2104:Crotaphytidae
2102:
2100:
2097:
2095:
2092:
2090:
2087:
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2072:
2069:
2067:
2064:
2062:
2059:
2057:
2054:
2052:
2049:
2047:
2044:
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2041:Lanthanotidae
2039:
2037:
2036:Shinisauridae
2034:
2033:
2031:
2029:
2025:
2015:
2014:Trogonophidae
2012:
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1867:
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1540:(8): e41767.
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1128:
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1120:
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1088:Necrosauridae
1086:
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1070:
1064:
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1009:tree-dwelling
1006:
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643:
642:
635:
631:Gilmore, 1943
629:
628:
621:
615:
614:
613:Proplatynotia
607:
601:
600:
593:
587:
586:
579:
573:
572:
565:
559:
558:
551:
545:
544:
537:
536:
535:
533:
529:
522:
513:
512:
506:
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499:Lanthanotidae
496:
492:
491:L. borneensis
488:
484:
480:
476:
471:
469:
468:L. borneensis
465:
464:
459:
458:
453:
449:
445:
444:
439:
435:
425:
424:of Mongolia.
423:
419:
415:
411:
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403:
399:
398:Komodo dragon
396:includes the
395:
391:
387:
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355:
351:
347:
343:
332:
331:
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325:
324:
319:
317:
316:
311:
309:
308:
307:Proplatynotia
303:
301:
300:
295:
293:
292:
287:
285:
284:
279:
277:
276:
271:
269:
268:
263:
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253:
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243:
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230:
228:Superfamily:
227:
226:
223:
220:
217:
216:
213:
210:
207:
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158:
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147:
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88:
83:
78:
73:
68:
63:
58:
53:
48:
42:
35:
31:
22:
19:
3052:
3017:
3010:
3005:Palaeosaniwa
3003:
2996:
2989:
2984:Morohasaurus
2982:
2975:
2968:
2961:
2956:Arcanosaurus
2954:
2936:
2929:
2922:
2912:
2900:
2893:
2886:
2811:
2802:
2793:
2784:
2775:
2766:
2757:
2748:
2739:
2730:
2721:
2712:
2703:
2694:
2685:
2677:V. giganteus
2676:
2668:V. jobiensis
2667:
2658:
2650:
2640:V. niloticus
2639:
2631:V. mitchelli
2630:
2621:
2612:
2603:
2594:
2585:
2577:V. olivaceus
2576:
2567:
2559:V. dumerilii
2558:
2549:
2540:
2531:
2522:
2513:
2504:
2495:
2486:
2474:
2462:
2450:
2439:H. suspectum
2438:
2429:
2420:
2411:
2402:
2289:Xenopeltidae
2154:Tropiduridae
2129:Leiosauridae
2056:Xenosauridae
2045:
1981:Amphisbaenia
1734:
1709:
1705:
1695:Garland Jr T
1688:
1663:
1657:
1638:
1634:
1628:
1620:
1603:
1599:
1589:Garland Jr T
1582:
1537:
1533:
1496:(1): 3β308.
1493:
1489:
1483:
1448:
1444:
1434:
1399:
1395:
1385:
1350:
1346:
1336:
1303:
1299:
1293:
1266:
1260:
1219:
1215:
1209:
1174:
1170:
1118:
1114:
1083:Mosasauridae
1058:
1047:
1041:
1035:
1021:
1004:
930:
929:
913:
911:
888:
886:
854:
852:
836:
834:
786:
784:
768:
766:
743:
741:
711:
709:
687:
672:
664:
657:
639:
625:
611:
597:
583:
569:
555:
541:
531:
526:
502:
494:
490:
482:
478:
474:
472:
467:
461:
455:
451:
447:
441:
431:
393:
389:
384:than to the
381:
375:
341:
339:
328:
321:
313:
305:
297:
289:
281:
273:
265:
241:
150:
132:) fighting,
126:
18:
3170:iNaturalist
3077:Wikispecies
2998:Telmasaurus
2970:Cherminotus
2963:Aiolosaurus
2921:Megalania (
2803:V. panoptes
2686:V. gilleni
2651:V. bitatawa
2622:V. mertensi
2586:V. glauerti
2568:V. prasinus
2532:V. doreanus
2496:V. beccarii
2487:V. salvator
2452:Lanthanotus
2430:H. horridum
2403:H. alvarezi
2279:Uropeltidae
2264:Loxocemidae
2244:Bolyeriidae
2196:Typhlopidae
2134:Liolaemidae
2109:Dactyloidae
2066:Anniellidae
2028:Anguimorpha
2009:Rhineuridae
1931:Lacertoidea
1917:Xantusiidae
1879:Pygopodidae
1032:Philippines
1013:Philippines
713:Ovoo gurvel
688:Below is a
645:Leidy, 1870
627:Telmasaurus
599:Paravaranus
571:Cherminotus
557:Aiolosaurus
483:Lanthanotus
443:Lanthanotus
390:Lanthanotus
369:frugivorous
365:carnivorous
361:Anguimorpha
354:superfamily
299:Paravaranus
283:Aiolosaurus
275:Telmasaurus
130:komodoensis
3283:Categories
2776:V. gouldii
2695:V. melinus
2675:Perentie (
2613:V. indicus
2550:V. griseus
2523:V. baritji
2505:V. tristis
2335:Varanoidea
2269:Pythonidae
2249:Colubridae
1943:Lacertidae
1902:Cordylidae
1894:Scincoidea
1854:Gekkonidae
1094:References
699:Varanidae
678:Shaw, 1790
434:de Queiroz
359:and order
357:Varanoidea
232:Varanoidea
149:Fossil of
39:80β0
3097:Varanidae
3083:Varanidae
3053:Varanidae
3012:Saniwides
2977:Gobiderma
2913:Varanidae
2604:V. varius
2345:Kingdom:
2284:Viperidae
2224:Aniliidae
2164:Serpentes
2139:Opluridae
2119:Iguanidae
2046:Varanidae
1994:Bipedidae
1912:Scincidae
1836:Dibamidae
1800:Kingdom:
1753:Varanidae
1587:Perry G,
1320:2246/5915
1306:: 1β182.
1054:allometry
690:cladogram
684:Phylogeny
585:Saniwides
342:Varanidae
291:Saniwides
242:Varanidae
188:Kingdom:
182:Eukaryota
134:Indonesia
3068:Q3082684
3062:Wikidata
2895:Eosaniwa
2713:V. bushi
2365:Squamata
2359:Reptilia
2353:Chordata
2351:Phylum:
2347:Animalia
2259:Elapidae
2089:Agamidae
2061:Anguidae
2004:Cadeidae
1999:Blanidae
1927:Laterata
1820:Squamata
1814:Reptilia
1808:Chordata
1806:Phylum:
1802:Animalia
1791:families
1789:Squamata
1697:(1996).
1680:7155983M
1672:14915452
1591:(2002).
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1534:PLOS ONE
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1201:35125002
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1067:See also
1061:air-sacs
1028:arboreal
667:, 2022 (
481:than to
438:Gauthier
428:Taxonomy
238:Family:
222:Squamata
212:Reptilia
202:Chordata
198:Phylum:
192:Animalia
178:Domain:
34:Holocene
24:Varanids
3019:Estesia
2476:Varanus
2382:species
2363:Order:
2357:Class:
2081:Iguania
1971:Teiidae
1846:Gekkota
1829:Dibamia
1818:Order:
1812:Class:
1726:1565513
1629:Varanus
1600:Ecology
1565:3416840
1542:Bibcode
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1417:1626205
1376:1691912
1355:Bibcode
1347:Science
1252:4386245
1224:Bibcode
1192:8819366
1136:2936141
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992:Biology
932:Varanus
674:Varanus
534:, 2022
521:extinct
479:Varanus
475:Varanus
448:Varanus
410:goannas
394:Varanus
382:Varanus
377:Varanus
352:in the
350:lizards
330:Varanus
218:Order:
208:Class:
128:Varanus
3266:395082
3227:713776
3201:174114
3188:103447
3149:1VARAF
2948:Others
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649:Eocene
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528:Genera
509:Genera
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495:et al.
452:et al.
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346:family
344:are a
323:Saniwa
256:Genera
249:, 1820
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157:Eocene
152:Saniwa
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3248:Plazi
3240:38035
3183:IRMNG
3175:39392
3123:6273W
1722:JSTOR
1702:(PDF)
1596:(PDF)
1506:S2CID
1324:S2CID
1248:S2CID
663:Dong
373:genus
121:Male
3214:8555
3209:NCBI
3196:ITIS
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3157:GBIF
3144:EPPO
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3110:1099
3105:BOLD
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2865:taxa
1668:OCLC
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1471:PMID
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1271:ISBN
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