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Tenontosaurus

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stretches from southwest Arkansas through southeastern Oklahoma and into northeastern Texas. This geological formation has not been dated radiometrically. Scientists have used biostratigraphic data and the fact that it shares several of the same genera as the Trinity Group of Texas, to surmise that
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must therefore have been a pack hunter, this view has been challenged based on both a supposed lack of evidence for coordinated hunting (rather than mobbing behavior as in most modern birds and reptiles, though crocodilians have been documented to hunt cooperatively on occasion) as well as evidence
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resembles quadrupedal ornithischians in having a tibia that is shorter than the femur and an anterolateral process on the ulna, but it resembles bipedal ornithischians in having a relatively narrow pelvis and a pendant fourth trochanter. The manus (the hand/front foot) shows a mixture of traits
527:(AMNH) expedition in 1903. Subsequent digs in the same area during the 1930s unearthed 18 more specimens, and four specimens were found during the 1940s. Despite the large number of fossil specimens, the animal was not named or scientifically described during this time, though 1278:, especially juveniles, to eating low-growing ferns and shrubs. Its powerful, U-shaped beak and the angled cutting surfaces of its teeth, however, meant it was not limited to which part of the plant it consumed. Leaves, wood, and even fruit may have formed part of its diet. 1790:
remains are common in two distinct rock units: the more ancient Little Sheep Mudstone Member (Cloverly Formation unit V) and the more recent Himes Member (units VI and VII). The oldest part of the formation, the Pryor Conglomerate, contains no
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would have been 6 metres (20 ft) and weighed 600 kilograms (1,300 lb), but Campione and Evans in 2020 estimated that this species would have weighed up to 971–1,019 kilograms (2,141–2,247 lb). Paul also estimated that
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Weishampel, David B.; Barrett, Paul M.; Coria, Rodolfo A.; Le Loeuff, Jean; Xu Xing; Zhao Xijin; Sahni, Ashok; Gomani, Elizabeth, M.P.; and Noto, Christopher R. (2004). "Dinosaur Distribution", in The Dinosauria (2nd), p.
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Brinkman, Daniel L.; Cifelli, Richard L.; Czaplewski, Nicholas J. (1998). "First Occurrence of Deinonychus antirrhopus (Dinosauria: Theropoda) from the Antlers Formation (Lower Cretaceous: Aptain-Albian) of Oklahoma".
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to semi-arid, dry, with seasonal periods of rainfall and occasional droughts. However, during a period of a few million years, the climate in the region shifted to one of increased rainfall, and the environment became
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reached a certain size, it passed out of range as a food source for the small theropods, though they may have scavenged larger individuals, or preyed on adults that were sick or injured. The fact that most
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was a facultative quadruped, capable of assuming either a bipedal or quadrupedal stance. It may have used a quadrupedal stance while feeding, but was probably incapable of rapid quadrupedal movement.
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Cifelli, Richard L.; Gardner, James D.; Nydam, Randall L.; Brinkman, Daniel L. (August 1997). "Additions to the vertebrate fauna of the Antlers Formation (Lower Cretaceous), southeastern Oklahoma".
1827:, though marked dry seasons persisted to create savannah-like environments as well. The change in rainfall levels is likely due to the advancing shoreline of the Skull Creek Seaway, a cycle of the 621:
would have been 7 meters (23 ft) long and weighed 1,000 kilograms (2,200 lb). Adult length estimates among other researchers range between 7 and 8 metres (23 and 26 ft).
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associated with bipeds and quadrupeds. It retains narrow claws, unlike the hoof-like unguals of quadrupedal ornithischians, has a short metacarpus, as in the bipedal
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very early in the evolution of dinosaurs, this suggests that dinosaurs in general produced medullary tissue and reached reproductive maturity before maximum size. A
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Forster, C. A. (1984). "The paleoecology of the ornithopod dinosaur Tenontosaurus tilletti from the Cloverly Formation, Big Horn Basin of Wyoming and Montana".
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Winkler, Dale A.; Murry, Phillip A.; Jacobs, Louis L. (1997). "A new species of Tenontosaurus (Dinosauria: Ornithopoda) from the Early Cretaceous of Texas".
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this formation was laid down during the Albian stage of the Early Cretaceous Period, approximately 110 mya. The area preserved in this formation was a large
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fossils, and they only appear in the uppermost, most recent part of the Little Sheep Mudstone Member. Catherine Forster, in a 1984 paper on the ecology of
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was a low browser, and an adult would have had a maximum browsing height of about 3 meters (10 ft) if it adopted a bipedal stance. This restricted
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of the AMNH gave it the informal name "Tenantosaurus", "sinew lizard", in reference to the extensive system of stiffening tendons in its back and tail.
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D'Emic, Michael D.; Melstrom, Keegan M.; Eddy, Drew R. (2012). "Paleobiology and geographic range of the large-bodied Cretaceous theropod dinosaur
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Ostrom, J. H. (1970). "Stratigraphy and paleontology of the Cloverly Formation (Lower Cretaceous) of the Bighorn Basin area, Wyoming and Montana".
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found it to be a quadruped, although the analysis only distinguished bipeds from quadrupeds without including a facultatively bipedal category.
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Kielan-Jaworowska, Zofia; Cifelli, Richard L (2001). "Primitive boreosphenidan mammal (?Deltatheroida) from the Early Cretaceous of Oklahoma".
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Nydam, Randall L.; Cifelli, Richard L. (2002-07-08). "Lizards from the Lower Cretaceous (Aptian–Albian) Antlers and Cloverly Formations".
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to have been a remarkably adaptable animal, which persisted for a long span of time in one area despite changes to its environment.
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Hunt, Tyler C.; Cifelli, Richard L.; Davies, Kyle L. (2021-07-16). "The hand of Tenontosaurus tilletti (Dinosauria, Ornithopoda)".
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Bishop, Peter J.; Bates, Karl T.; Allen, Vivian R.; Henderson, Donald M.; Randau, Marcela; Hutchinson, John R. (2020-10-16).
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was a medium-sized ornithopod, with both species weighing about 1,000 kilograms (2,200 lb). Paul in 2016 estimated that
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Maidment, Susannah; Barrett, Paul (2012-09-21). "Osteological correlates for quadrupedality in ornithischian dinosaurs".
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that drained into a shallow inland sea. Several million years later, this sea would expand to the north, becoming the
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period. The paleoenvironment of the Antlers Formation consisted of tropical or sub-tropical forests, floodplains,
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Butler, R.J.; Barrett, P.M. (2012). "26. Ornithopods". In Brett-Surman, M.K.; Holtz, T.R.; Farlow, J.O. (eds.).
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are half-grown individuals supports this view. It also lived in the same area as the large carnivorous dinosaur
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This dramatic shift in climate coincided with an apparent increase, rather than decrease, in the abundance of
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populations did not arrive in the Bighorn Basin area until the time of the late Little Sheep Mudstone Member.
5509: 5449: 5264: 2288:"An early-diverging iguanodontian (Dinosauria: Rhabdodontomorpha) from the Late Cretaceous of North America" 1977:
Forster, Catherine A. (1990). "The postcranial skeleton of the ornithopod dinosaur Tenontosaurus tilletti".
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Zanno, Lindsay E.; Gates, Terry A.; Avrahami, Haviv M.; Tucker, Ryan T.; Makovicky, Peter J. (2023-06-07).
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Dinets, Vladimir (2015). "Apparent coordination and collaboration in cooperatively hunting crocodilians".
423:. It had an unusually long, broad tail, which like its back was stiffened with a network of bony tendons. 142: 5454: 5444: 4894: 1504:
is by far the most common vertebrate, five times more abundant than the next most common, the ankylosaur
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Roach, B. T.; Brinkman, D. L. (2007). "A reevaluation of cooperative pack hunting and gregariousness in
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from the Quantou Formation (Cretaceous: Aptian–Cenomanian) of Jilin Province, north-eastern China".
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specimens have been reported, both from the Cloverly and other geological formations, including the
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varies between studies, with some studies finding it to be more closely related to dryomorphs than
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was the most common vertebrate in this region. The early mammals known from this region include
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in this region. The most common dinosaur in the paleoenvironment preserved in this formation is
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crocodile. After the major climate shift at the beginning of the Himes Member in the mid-
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is the only herbivorous dinosaur, and it shared its environment with the common predator
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have been collected from several geological formations throughout western North America.
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first appeared in Wyoming and Montana (the early Albian age), the regions climate was
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Maxwell, W. D.; Ostrom, J. H. (1995). "Taphonomy and paleobiological implications of
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age, several more dinosaurs entered the region, including the less common ornithopod
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Oreska, Matthew P. J.; Carrano, Matthew T.; Dzikiewicz, Katherine M. (March 2013).
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which, later in the Cretaceous period, would completely divide North America.
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The ecological community was similar in other regions, with dinosaurs like
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remains are only rarely found associated with other potential prey, like
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specimens have been found at over 50 sites, and 14 of those also contain
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Teeth and a number of skeletons belonging to the carnivorous theropod
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remains. According to one 1995 study, only six sites containing
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began an extensive, long-term dig in the Big Horn Basin area (
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10.1671/0272-4634(2002)022[0286:LFTLCA]2.0.CO;2
1437: 1247: 696: 590: 554:, a slight variation in spelling of Brown's informal name. 369: 357: 2662:"The Ontogenetic osteohistology of Tenontosaurus tilletti" 1440:
that are laying eggs, in reproduction. Additionally, like
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10.3374/0079-032X(2007)48[103:AROCPH]2.0.CO;2
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and dividing North America in two for nearly the entire
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Osteohistology of the diaphyseal femur of two juveniles
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may have been cannibalizing each other, as well as the
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in 2023 lent support for a close relationship between
381: 372: 363: 351: 345: 342: 2922: 2888: 354: 336: 1364:would have been capable of attacking a fully grown 360: 1890: 2572:Palaeogeography, Palaeoclimatology, Palaeoecology 2514:Bulletin of the Peabody Museum of Natural History 2185: 1863:Bulletin of the Peabody Museum of Natural History 5411: 1476:grew quickly early in life and during sub-adult 2607:Proceedings of the National Academy of Sciences 2545: 2543: 2147: 3044: 2816: 2143: 2141: 2139: 708:exhibits transitional characteristics between 667:. Subsequent phylogenetic analyses have found 4016: 2954: 2507: 2409: 2117: 2065: 1972: 1970: 1968: 1948: 1946: 1944: 1942: 1940: 1938: 1936: 1934: 1932: 1930: 1886: 1884: 1424:The presence of medullary bone tissue in the 2832: 2778: 2774: 2772: 2770: 2540: 2395:: CS1 maint: multiple names: authors list ( 1856: 1854: 1590:Mounted skeletons of an adult with juveniles 1510:. In the arid Little Sheep Mudstone Member, 1368:. While some scientists have suggested that 5425:Early Cretaceous dinosaurs of North America 2869: 2653: 2136: 2037:. Princeton University Press. p. 311. 1677:. Other vertebrates present at the time of 1306:have often been discovered associated with 394: 3151: 2961: 2947: 2712: 2600: 2458: 2346: 2222: 2181: 2179: 2026: 2024: 1965: 1927: 1881: 1602:as the most common large vertebrates. The 117: 2826: 2767: 2695: 2685: 2636: 2626: 2559: 2501: 2353:Richard J. Butler, Jin Liyong, Chen Jun, 2329: 2311: 2279: 2254: 2122:. Indiana University Press. p. 647. 2083: 1851: 1381:, in a feeding frenzy. It is likely that 2512:and other nonavian theropod dinosaurs". 2120:The Complete Dinosaur (Life of the Past) 1799:, used this as evidence to suggest that 1585: 1491: 1415: 1356:, however, were much smaller than adult 1340:fossils are found in close proximity to 1285: 659:was originally classified in the family 647: 496: 485: 5530:Multispecific non-avian dinosaur genera 2659: 2601:Lee, Andrew H.; Werning, Sarah (2008). 2176: 2021: 1976: 1952: 1520:as well as an indeterminate species of 686:. The precise phylogenetic position of 671:to be intermediate in position between 481: 5412: 2464: 2066:Campione, N. E.; Evans, D. C. (2020). 2034:The Princeton Field Guide to Dinosaurs 1860: 1436:used this tissue, today only found in 5306: 5305: 4910: 4003: 3138: 2980: 2942: 2779:Wedel, M. J.; Cifelli, R. L. (2005). 4911: 2594: 2030: 719:below follows an analysis by Butler 1500:Throughout the Cloverly Formation, 1464:diverged from the line that led to 13: 4004: 2835:Journal of Vertebrate Paleontology 2725:Journal of Vertebrate Paleontology 2420:Journal of Vertebrate Paleontology 2188:Journal of Vertebrate Paleontology 1979:Journal of Vertebrate Paleontology 1893:Journal of Vertebrate Paleontology 1643:with other dinosaurs, such as the 1547:, and an indeterminate species of 1246:ecosystem was likely dominated by 525:American Museum of Natural History 131:Perot Museum of Nature and Science 14: 5541: 2918: 643: 426:The genus is known from the late 316:Winkler, Murry & Jacobs, 1997 4893: 4887: 4882: 4876: 4869: 3986: 3979: 3972: 2925: 2467:Ethology Ecology & Evolution 2383:10.1111/j.1475-4983.2011.01046.x 1456:line of dinosaurs that includes 462:is known from only a handful of 332: 141: 49: 5250:List of informal dinosaur names 3139: 2968: 1487: 1432:of one specimen indicates that 1411: 1232: 5470:Aptian genus first appearances 5260:Timeline of hadrosaur research 2440:10.1080/02724634.1995.10011256 2111: 2059: 1999:10.1080/02724634.1990.10011815 1913:10.1080/02724634.1997.10010978 1360:, and it is unlikely a single 604: 1: 5430:Fossil taxa described in 1970 2897:Acta Palaeontologica Polonica 2208:10.1080/02724634.2021.1938591 2150:Acta Palaeontologica Polonica 1845: 712:and other rhabdodontomorphs. 5485:Ornithopods of North America 5465:Early Cretaceous ornithopods 2745:10.1080/02724634.2012.717567 2687:10.1371/journal.pone.0033539 2588:10.1016/j.palaeo.2012.03.003 2487:10.1080/03949370.2014.915432 2313:10.1371/journal.pone.0286042 1322:fossils contain no trace of 1293:with juveniles, in front of 1281: 546:. The expedition was led by 438:period sediments of western 7: 10: 5546: 2798:(2): 40–57. Archived from 2785:: Oklahoma's Native Giant" 2568:Acrocanthosaurus atokensis 2553:Oklahoma Geological Survey 1777: 1348:was the major predator of 16:Extinct genus of dinosaurs 5440:Taxa named by John Ostrom 5314: 5215: 5066: 5046: 4921: 4917: 4906: 4867: 4798: 4764: 4708: 4667: 4611: 4596: 4587: 4514: 4473: 4366: 4357: 4307: 4296: 4027: 4012: 3999: 3970: 3932: 3889: 3832: 3823: 3660: 3649: 3581: 3552: 3548: 3537: 3474: 3438: 3406: 3397: 3233: 3196: 3166: 3162: 3147: 3134: 3098: 3080: 3071: 3053: 3040: 2976: 2031:Paul, Gregory S. (2016). 1631:in what is now Oklahoma, 1133: 1118: 1111: 1082: 1065: 1058: 1046: 1031: 1024: 1007: 1000: 983: 976: 947: 930: 923: 916: 890: 873: 866: 859: 833: 816: 809: 792: 775: 758: 741: 734: 289: 282: 138:Scientific classification 136: 125: 116: 23: 5520:Paleontology in Maryland 5490:Paleontology in Oklahoma 5475:Albian genus extinctions 1786:of Montana and Wyoming, 587:Cedar Mountain Formation 468:Twin Mountains Formation 2628:10.1073/pnas.0708903105 2510:Deinonychus antirrhopus 1829:Western Interior Seaway 1693:Atokasaurus metarsiodon 1669:, which was likely the 1613:Western Interior Seaway 1237: 2878:Oklahoma Geology Notes 2792:Oklahoma Geology Notes 2359:Changchunsaurus parvus 2107:Supporting Information 1591: 1497: 1421: 1308:Tenontosaurus tilletti 1297: 704:and rhabdodontids, as 653: 652:Front part of skeleton 557:Since 1970, many more 505: 502:Tenontosaurus tilletti 494: 444:Tenontosaurus tilletti 5500:Paleontology in Idaho 5392:Paleobiology Database 5230:Basturs Poble bonebed 2162:10.4202/app.2012.0065 1589: 1495: 1419: 1289: 679:-like taxa, as a non- 651: 500: 489: 5510:Paleontology in Utah 5450:Ornithischian genera 5252:(contains hadrosaur 4650:Probrachylophosaurus 2660:Werning, S. (2012). 482:History of discovery 5525:Cretaceous Maryland 5495:Cretaceous Oklahoma 2737:2013JVPal..33..264O 2678:2012PLoSO...733539W 2619:2008PNAS..105..582L 2580:2012PPP...333...13D 2479:2015EtEcE..27..244D 2432:1995JVPal..15..707M 2375:2011Palgy..54..667B 2304:2023PLoSO..1886042Z 2256:10.1017/pab.2020.47 2247:2020Pbio...46..550B 2200:2021JVPal..41E8591H 1991:1990JVPal..10..273F 1905:1997JVPal..17..330W 1765:Atokatherium boreni 1652:(Pleurocoelus) and 1398:remains found with 466:collected from the 452:Tenontosaurus dossi 5455:Cretaceous Montana 5445:Fossils of Montana 5002:Ornithopodichnites 4613:Brachylophosaurini 3198:Hypsilophodontidae 2574:. 333–334: 13–23. 2451:2007-09-27 at the 2072:Biological Reviews 1784:Cloverly Formation 1744:, and the turtles 1738:, the crocodilian 1697:Ptilotodon wilsoni 1592: 1498: 1472:study showed that 1422: 1298: 1242:Plant life in the 665:Hypsilophodontidae 654: 540:Cloverly Formation 534:During the 1960s, 506: 495: 129:on exhibit in the 5407: 5406: 5379:Open Tree of Life 5308:Taxon identifiers 5299: 5298: 5295: 5294: 5291: 5290: 5009:Shiraminesauropus 4902: 4901: 4865: 4864: 4861: 4860: 4857: 4856: 4853: 4852: 4748:Naashoibitosaurus 4669:Austrokritosauria 4629:Brachylophosaurus 4583: 4582: 3995: 3994: 3968: 3967: 3964: 3963: 3960: 3959: 3956: 3955: 3952: 3951: 3645: 3644: 3533: 3532: 3529: 3528: 3513:Transylvanosaurus 3399:Rhabdodontomorpha 3345:Notohypsilophodon 3316:Macrogryphosaurus 3130: 3129: 3126: 3125: 3122: 3121: 3118: 3117: 2085:10.1111/brv.12638 2044:978-1-78684-190-2 1629:Antlers Formation 1604:Antlers Formation 1577:, and species of 1385:favored juvenile 1336:. In all, 20% of 1229: 1228: 1220: 1219: 1211: 1210: 1202: 1201: 1193: 1192: 1184: 1183: 1175: 1174: 1166: 1165: 1157: 1156: 1097: 1096: 962: 961: 905: 904: 848: 847: 595:Arundel Formation 563:Antlers Formation 542:) of Montana and 323: 322: 317: 306: 278: 5537: 5505:Cretaceous Idaho 5480:Cretaceous Texas 5460:Fossils of Texas 5400: 5399: 5387: 5386: 5374: 5373: 5361: 5360: 5348: 5347: 5335: 5334: 5333: 5303: 5302: 5276:mummy S.M. R4036 5150:Microhadrosaurus 5143:Mandschurosaurus 4953:Camptosaurichnus 4946:Bonaparteichnium 4919: 4918: 4912:Related articles 4908: 4907: 4897: 4891: 4886: 4881: 4880: 4873: 4594: 4593: 4475:Parasaurolophini 4386:Angulomastacator 4364: 4363: 4305: 4304: 4224:Shuangmiaosaurus 4125:Huehuecanauhtlus 4111:Gongpoquansaurus 4025: 4024: 4014: 4013: 4001: 4000: 3990: 3984: 3983: 3976: 3830: 3829: 3825:Hadrosauriformes 3793:Ratchasimasaurus 3658: 3657: 3550: 3549: 3546: 3545: 3428:Muttaburrasaurus 3404: 3403: 3337:Muttaburrasaurus 3251:Atlascopcosaurus 3186:Thescelosauridae 3164: 3163: 3160: 3159: 3149: 3148: 3136: 3135: 3078: 3077: 3051: 3050: 3042: 3041: 3035: 3034: 2978: 2977: 2963: 2956: 2949: 2940: 2939: 2935: 2933:Dinosaurs portal 2930: 2929: 2928: 2913: 2912: 2892: 2886: 2885: 2873: 2867: 2866: 2830: 2824: 2820: 2814: 2813: 2811: 2810: 2804: 2789: 2776: 2765: 2764: 2716: 2710: 2709: 2699: 2689: 2657: 2651: 2650: 2640: 2630: 2598: 2592: 2591: 2563: 2557: 2556: 2547: 2538: 2537: 2505: 2499: 2498: 2462: 2456: 2443: 2407: 2401: 2400: 2394: 2386: 2355:Pascal Godefroit 2350: 2344: 2343: 2333: 2315: 2283: 2277: 2276: 2258: 2226: 2220: 2219: 2183: 2174: 2173: 2145: 2134: 2133: 2115: 2109: 2105: 2087: 2078:(6): 1759–1797. 2063: 2057: 2056: 2028: 2019: 2018: 1974: 1963: 1962: 1950: 1925: 1924: 1888: 1879: 1878: 1858: 1665:Acrocanthosaurus 1641:paleoenvironment 1405:Acrocanthosaurus 1389:, and that when 1114: 1113: 1061: 1060: 1049: 1048: 1027: 1026: 1003: 1002: 979: 978: 926: 925: 919: 918: 869: 868: 862: 861: 812: 811: 737: 736: 727: 726: 571:Paluxy Formation 436:Early Cretaceous 408: 405: 402: 399: 396: 389: 385: 379: 378: 375: 374: 371: 368: 365: 362: 359: 356: 353: 350: 347: 344: 341: 338: 315: 300: 270: 263: 250: 234: 218: 202: 146: 145: 121: 111: 48: 31:Early Cretaceous 29:Temporal range: 21: 20: 5545: 5544: 5540: 5539: 5538: 5536: 5535: 5534: 5515:Cretaceous Utah 5410: 5409: 5408: 5403: 5395: 5390: 5382: 5377: 5369: 5364: 5356: 5351: 5343: 5338: 5329: 5328: 5323: 5310: 5300: 5287: 5268:mummy AMNH 5060 5235:Dakota (fossil) 5211: 5062: 5042: 5030:Tetrapodosaurus 4981:Hadrosaurichnus 4913: 4898: 4875: 4849: 4844:Shantungosaurus 4794: 4775:Augustynolophus 4760: 4704: 4678:Bonapartesaurus 4663: 4607: 4579: 4510: 4498:Parasaurolophus 4469: 4353: 4301: 4292: 4217:Qianjiangsaurus 4203:Probactrosaurus 4182:Penelopognathus 4021: 4008: 3991: 3978: 3948: 3928: 3885: 3819: 3800:Riabininohadros 3673:Bayannurosaurus 3654: 3641: 3577: 3542: 3525: 3470: 3440:Tenontosauridae 3434: 3393: 3229: 3208:Gideonmantellia 3192: 3156: 3143: 3114: 3094: 3067: 3061:Avemetatarsalia 3046:Avemetatarsalia 3036: 2983: 2982: 2972: 2967: 2931: 2926: 2924: 2921: 2916: 2893: 2889: 2874: 2870: 2831: 2827: 2821: 2817: 2808: 2806: 2802: 2787: 2777: 2768: 2717: 2713: 2658: 2654: 2599: 2595: 2564: 2560: 2548: 2541: 2506: 2502: 2463: 2459: 2453:Wayback Machine 2418:associations". 2408: 2404: 2388: 2387: 2351: 2347: 2298:(6): –0286042. 2284: 2280: 2227: 2223: 2184: 2177: 2146: 2137: 2130: 2116: 2112: 2064: 2060: 2045: 2029: 2022: 1975: 1966: 1951: 1928: 1889: 1882: 1859: 1852: 1848: 1780: 1691:, the reptiles 1617:Late Cretaceous 1549:titanosauriform 1496:Complete fossil 1490: 1414: 1284: 1240: 1235: 1230: 1221: 1212: 1203: 1194: 1185: 1176: 1167: 1158: 1098: 963: 906: 849: 820:Changchunsaurus 675:-like taxa and 646: 607: 579:Wayan Formation 536:Yale University 517:Big Horn County 490:Restoration of 484: 406: 403: 400: 397: 387: 383: 335: 331: 269: 261: 252:Tenontosauridae 248: 232: 216: 200: 140: 112: 110: 109: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 43: 42: 27: 17: 12: 11: 5: 5543: 5533: 5532: 5527: 5522: 5517: 5512: 5507: 5502: 5497: 5492: 5487: 5482: 5477: 5472: 5467: 5462: 5457: 5452: 5447: 5442: 5437: 5432: 5427: 5422: 5420:Cloverly fauna 5405: 5404: 5402: 5401: 5388: 5375: 5362: 5349: 5336: 5320: 5318: 5312: 5311: 5297: 5296: 5293: 5292: 5289: 5288: 5286: 5285: 5278: 5270: 5262: 5257: 5247: 5245:Hypsilophodont 5242: 5240:Hadrosaur diet 5237: 5232: 5227: 5219: 5217: 5216:Other articles 5213: 5212: 5210: 5209: 5202: 5195: 5188: 5181: 5178:Stephanosaurus 5174: 5167: 5160: 5153: 5146: 5139: 5132: 5125: 5118: 5115:Fulgurotherium 5111: 5104: 5097: 5089: 5082: 5074: 5072: 5064: 5063: 5061: 5060: 5052: 5050: 5044: 5043: 5041: 5040: 5033: 5026: 5019: 5012: 5005: 4998: 4991: 4988:Iguanodontipus 4984: 4977: 4970: 4963: 4956: 4949: 4942: 4935: 4927: 4925: 4915: 4914: 4904: 4903: 4900: 4899: 4868: 4866: 4863: 4862: 4859: 4858: 4855: 4854: 4851: 4850: 4848: 4847: 4840: 4837:Laiyangosaurus 4833: 4826: 4819: 4812: 4804: 4802: 4800:Edmontosaurini 4796: 4795: 4793: 4792: 4785: 4782:Prosaurolophus 4778: 4770: 4768: 4762: 4761: 4759: 4758: 4751: 4744: 4737: 4730: 4727:Coahuilasaurus 4723: 4714: 4712: 4706: 4705: 4703: 4702: 4695: 4688: 4681: 4673: 4671: 4665: 4664: 4662: 4661: 4653: 4646: 4639: 4632: 4625: 4617: 4615: 4609: 4608: 4606: 4605: 4597: 4591: 4585: 4584: 4581: 4580: 4578: 4577: 4570: 4563: 4556: 4549: 4542: 4535: 4528: 4520: 4518: 4512: 4511: 4509: 4508: 4501: 4494: 4487: 4479: 4477: 4471: 4470: 4468: 4467: 4460: 4453: 4446: 4439: 4431: 4424: 4417: 4410: 4403: 4396: 4389: 4382: 4375: 4367: 4361: 4359:Lambeosaurinae 4355: 4354: 4352: 4351: 4344: 4337: 4330: 4323: 4316: 4308: 4302: 4297: 4294: 4293: 4291: 4290: 4283: 4276: 4269: 4262: 4255: 4248: 4241: 4234: 4227: 4220: 4213: 4206: 4199: 4192: 4185: 4178: 4175:Nanyangosaurus 4171: 4168:Nanningosaurus 4164: 4157: 4150: 4142: 4135: 4128: 4121: 4114: 4107: 4100: 4097:Gilmoreosaurus 4093: 4086: 4079: 4072: 4065: 4058: 4051: 4043: 4036: 4028: 4022: 4019:Hadrosauroidea 4017: 4010: 4009: 4006:Hadrosauroidea 3997: 3996: 3993: 3992: 3971: 3969: 3966: 3965: 3962: 3961: 3958: 3957: 3954: 3953: 3950: 3949: 3947: 3946: 3945: 3944: 3938: 3936: 3934:Hadrosauroidea 3930: 3929: 3927: 3926: 3922:Mantellisaurus 3918: 3911: 3904: 3895: 3893: 3891:Iguanodontidae 3887: 3886: 3884: 3883: 3876: 3869: 3866:Mantellisaurus 3862: 3855: 3848: 3841: 3833: 3827: 3821: 3820: 3818: 3817: 3810: 3803: 3796: 3789: 3782: 3775: 3768: 3761: 3754: 3747: 3740: 3733: 3730:Iguanacolossus 3726: 3719: 3712: 3705: 3697: 3690: 3683: 3676: 3669: 3661: 3655: 3652:Ankylopollexia 3650: 3647: 3646: 3643: 3642: 3640: 3639: 3632: 3624: 3616: 3613:Eousdryosaurus 3609: 3602: 3599:Dysalotosaurus 3595: 3587: 3585: 3579: 3578: 3576: 3575: 3568: 3561: 3553: 3543: 3538: 3535: 3534: 3531: 3530: 3527: 3526: 3524: 3523: 3516: 3509: 3502: 3495: 3488: 3480: 3478: 3476:Rhabdodontidae 3472: 3471: 3469: 3468: 3461: 3454: 3445: 3443: 3436: 3435: 3433: 3432: 3424: 3416: 3407: 3401: 3395: 3394: 3392: 3391: 3387:Weewarrasaurus 3383: 3376: 3369: 3362: 3355: 3348: 3341: 3333: 3326: 3319: 3312: 3309:Leaellynasaura 3305: 3297: 3290: 3287:Gasparinisaura 3283: 3280:Galleonosaurus 3276: 3268: 3261: 3254: 3247: 3239: 3237: 3231: 3230: 3228: 3227: 3219: 3212: 3203: 3201: 3194: 3193: 3191: 3190: 3182: 3175: 3167: 3157: 3152: 3145: 3144: 3132: 3131: 3128: 3127: 3124: 3123: 3120: 3119: 3116: 3115: 3113: 3112: 3111: 3110: 3104: 3102: 3096: 3095: 3093: 3092: 3091: 3090: 3081: 3075: 3069: 3068: 3066: 3065: 3064: 3063: 3054: 3048: 3038: 3037: 3033: 3032: 3023: 3021:Neornithischia 3014: 3005: 2996: 2990: 2974: 2973: 2966: 2965: 2958: 2951: 2943: 2937: 2936: 2920: 2919:External links 2917: 2915: 2914: 2903:(3): 377–391. 2887: 2868: 2841:(2): 286–298. 2825: 2815: 2766: 2731:(2): 264–292. 2711: 2652: 2613:(2): 582–587. 2593: 2558: 2539: 2520:(1): 103–138. 2500: 2473:(2): 244–250. 2457: 2426:(4): 707–712. 2402: 2369:(3): 667–683. 2345: 2278: 2241:(4): 550–568. 2221: 2175: 2135: 2129:978-0253357014 2128: 2110: 2058: 2043: 2020: 1985:(3): 273–294. 1964: 1926: 1899:(2): 330–348. 1880: 1849: 1847: 1844: 1779: 1776: 1540:oviraptorosaur 1489: 1486: 1482:iguanodontians 1413: 1410: 1283: 1280: 1239: 1236: 1234: 1231: 1227: 1226: 1223: 1222: 1218: 1217: 1214: 1213: 1209: 1208: 1205: 1204: 1200: 1199: 1196: 1195: 1191: 1190: 1187: 1186: 1182: 1181: 1178: 1177: 1173: 1172: 1169: 1168: 1164: 1163: 1160: 1159: 1155: 1154: 1151: 1150: 1148:, and others) 1136:Ankylopollexia 1132: 1129: 1128: 1125: 1124: 1117: 1112: 1110: 1104: 1103: 1100: 1099: 1095: 1094: 1091: 1090: 1081: 1078: 1077: 1074: 1073: 1064: 1059: 1057: 1047: 1045: 1042: 1041: 1038: 1037: 1034:Rhabdodontidae 1030: 1025: 1023: 1020: 1019: 1016: 1015: 1006: 1001: 999: 996: 995: 992: 991: 982: 977: 975: 969: 968: 965: 964: 960: 959: 956: 955: 951:Thescelosaurus 946: 943: 942: 939: 938: 929: 924: 922: 917: 915: 912: 911: 908: 907: 903: 902: 899: 898: 889: 886: 885: 882: 881: 877:Gasparinisaura 872: 867: 865: 860: 858: 855: 854: 851: 850: 846: 845: 842: 841: 832: 829: 828: 825: 824: 815: 810: 808: 805: 804: 801: 800: 791: 788: 787: 784: 783: 774: 771: 770: 767: 766: 757: 754: 753: 750: 749: 740: 735: 733: 725: 661:Iguanodontidae 645: 644:Classification 642: 606: 603: 483: 480: 446:(described by 321: 320: 319: 318: 307: 301:Ostrom, 1970 ( 287: 286: 280: 279: 259: 255: 254: 246: 239: 238: 230: 223: 222: 220:Neornithischia 214: 207: 206: 198: 191: 190: 185: 178: 177: 172: 168: 167: 162: 158: 157: 152: 148: 147: 134: 133: 123: 122: 114: 113: 105: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 28: 15: 9: 6: 4: 3: 2: 5542: 5531: 5528: 5526: 5523: 5521: 5518: 5516: 5513: 5511: 5508: 5506: 5503: 5501: 5498: 5496: 5493: 5491: 5488: 5486: 5483: 5481: 5478: 5476: 5473: 5471: 5468: 5466: 5463: 5461: 5458: 5456: 5453: 5451: 5448: 5446: 5443: 5441: 5438: 5436: 5433: 5431: 5428: 5426: 5423: 5421: 5418: 5417: 5415: 5398: 5393: 5389: 5385: 5380: 5376: 5372: 5367: 5363: 5359: 5354: 5350: 5346: 5341: 5337: 5332: 5326: 5322: 5321: 5319: 5317: 5316:Tenontosaurus 5313: 5309: 5304: 5284: 5283: 5279: 5277: 5275: 5274:Edmontosaurus 5271: 5269: 5267: 5266:Edmontosaurus 5263: 5261: 5258: 5255: 5251: 5248: 5246: 5243: 5241: 5238: 5236: 5233: 5231: 5228: 5226: 5225: 5221: 5220: 5218: 5214: 5208: 5207: 5203: 5201: 5200: 5196: 5194: 5193: 5189: 5187: 5186: 5185:Syngonosaurus 5182: 5180: 5179: 5175: 5173: 5172: 5168: 5166: 5165: 5161: 5159: 5158: 5154: 5152: 5151: 5147: 5145: 5144: 5140: 5138: 5137: 5133: 5131: 5130: 5129:Koutalisaurus 5126: 5124: 5123: 5119: 5117: 5116: 5112: 5110: 5109: 5108:Eucercosaurus 5105: 5103: 5102: 5098: 5095: 5094: 5090: 5088: 5087: 5083: 5081: 5080: 5079:Arstanosaurus 5076: 5075: 5073: 5071: 5070: 5065: 5059: 5058: 5054: 5053: 5051: 5049: 5045: 5039: 5038: 5034: 5032: 5031: 5027: 5025: 5024: 5020: 5018: 5017: 5013: 5011: 5010: 5006: 5004: 5003: 4999: 4997: 4996: 4995:Orcauichnites 4992: 4990: 4989: 4985: 4983: 4982: 4978: 4976: 4975: 4971: 4969: 4968: 4964: 4962: 4961: 4957: 4955: 4954: 4950: 4948: 4947: 4943: 4941: 4940: 4936: 4934: 4933: 4932:Amblydactylus 4929: 4928: 4926: 4924: 4920: 4916: 4909: 4905: 4896: 4892: 4890: 4885: 4879: 4872: 4846: 4845: 4841: 4839: 4838: 4834: 4832: 4831: 4830:Kundurosaurus 4827: 4825: 4824: 4823:Kerberosaurus 4820: 4818: 4817: 4813: 4811: 4810: 4809:Edmontosaurus 4806: 4805: 4803: 4801: 4797: 4791: 4790: 4786: 4784: 4783: 4779: 4777: 4776: 4772: 4771: 4769: 4767: 4763: 4757: 4756: 4752: 4750: 4749: 4745: 4743: 4742: 4738: 4736: 4735: 4731: 4729: 4728: 4724: 4721: 4720: 4719:Anasazisaurus 4716: 4715: 4713: 4711: 4707: 4701: 4700: 4699:Secernosaurus 4696: 4694: 4693: 4692:Kelumapusaura 4689: 4687: 4686: 4682: 4680: 4679: 4675: 4674: 4672: 4670: 4666: 4659: 4658: 4654: 4652: 4651: 4647: 4645: 4644: 4640: 4638: 4637: 4633: 4631: 4630: 4626: 4624: 4623: 4619: 4618: 4616: 4614: 4610: 4604: 4603: 4599: 4598: 4595: 4592: 4590: 4589:Saurolophinae 4586: 4576: 4575: 4571: 4569: 4568: 4564: 4562: 4561: 4557: 4555: 4554: 4550: 4548: 4547: 4543: 4541: 4540: 4539:Hypacrosaurus 4536: 4534: 4533: 4532:Corythosaurus 4529: 4527: 4526: 4522: 4521: 4519: 4517: 4516:Lambeosaurini 4513: 4507: 4506: 4502: 4500: 4499: 4495: 4493: 4492: 4491:Charonosaurus 4488: 4486: 4485: 4481: 4480: 4478: 4476: 4472: 4466: 4465: 4464:Tsintaosaurus 4461: 4459: 4458: 4457:Pararhabdodon 4454: 4452: 4451: 4450:Nipponosaurus 4447: 4445: 4444: 4440: 4437: 4436: 4432: 4430: 4429: 4425: 4423: 4422: 4421:Jaxartosaurus 4418: 4416: 4415: 4411: 4409: 4408: 4404: 4402: 4401: 4397: 4395: 4394: 4390: 4388: 4387: 4383: 4381: 4380: 4376: 4374: 4373: 4372:Adynomosaurus 4369: 4368: 4365: 4362: 4360: 4356: 4350: 4349: 4345: 4343: 4342: 4338: 4336: 4335: 4334:Lapampasaurus 4331: 4329: 4328: 4324: 4322: 4321: 4317: 4315: 4314: 4310: 4309: 4306: 4303: 4300: 4299:Hadrosauridae 4295: 4289: 4288: 4284: 4282: 4281: 4280:Zhanghenglong 4277: 4275: 4274: 4270: 4268: 4267: 4263: 4261: 4260: 4256: 4254: 4253: 4252:Telmatosaurus 4249: 4247: 4246: 4242: 4240: 4239: 4235: 4233: 4232: 4228: 4226: 4225: 4221: 4219: 4218: 4214: 4212: 4211: 4207: 4205: 4204: 4200: 4198: 4197: 4196:Portellsaurus 4193: 4191: 4190: 4186: 4184: 4183: 4179: 4177: 4176: 4172: 4170: 4169: 4165: 4163: 4162: 4158: 4156: 4155: 4151: 4148: 4147: 4143: 4141: 4140: 4136: 4134: 4133: 4129: 4127: 4126: 4122: 4120: 4119: 4115: 4113: 4112: 4108: 4106: 4105: 4101: 4099: 4098: 4094: 4092: 4091: 4087: 4085: 4084: 4080: 4078: 4077: 4073: 4071: 4070: 4066: 4064: 4063: 4059: 4057: 4056: 4052: 4049: 4048: 4044: 4042: 4041: 4037: 4035: 4034: 4030: 4029: 4026: 4023: 4020: 4015: 4011: 4007: 4002: 3998: 3989: 3985: 3982: 3975: 3942: 3941: 3940: 3939: 3937: 3935: 3931: 3924: 3923: 3919: 3917: 3916: 3912: 3910: 3909: 3905: 3902: 3901: 3897: 3896: 3894: 3892: 3888: 3882: 3881: 3877: 3875: 3874: 3870: 3868: 3867: 3863: 3861: 3860: 3859:Jinzhousaurus 3856: 3854: 3853: 3852:Hypselospinus 3849: 3847: 3846: 3842: 3840: 3839: 3835: 3834: 3831: 3828: 3826: 3822: 3816: 3815: 3811: 3809: 3808: 3807:Theiophytalia 3804: 3802: 3801: 3797: 3795: 3794: 3790: 3788: 3787: 3783: 3781: 3780: 3776: 3774: 3773: 3769: 3767: 3766: 3762: 3760: 3759: 3755: 3753: 3752: 3748: 3746: 3745: 3741: 3739: 3738: 3737:Lanzhousaurus 3734: 3732: 3731: 3727: 3725: 3724: 3720: 3718: 3717: 3713: 3711: 3710: 3706: 3703: 3702: 3698: 3696: 3695: 3691: 3689: 3688: 3684: 3682: 3681: 3677: 3675: 3674: 3670: 3668: 3667: 3663: 3662: 3659: 3656: 3653: 3648: 3638: 3637: 3633: 3630: 3629: 3625: 3622: 3621: 3617: 3615: 3614: 3610: 3608: 3607: 3606:Elrhazosaurus 3603: 3601: 3600: 3596: 3594: 3593: 3589: 3588: 3586: 3584: 3580: 3574: 3573: 3569: 3567: 3566: 3562: 3560: 3559: 3555: 3554: 3551: 3547: 3544: 3541: 3536: 3522: 3521: 3517: 3515: 3514: 3510: 3508: 3507: 3503: 3501: 3500: 3496: 3494: 3493: 3489: 3487: 3486: 3482: 3481: 3479: 3477: 3473: 3467: 3466: 3465:Tenontosaurus 3462: 3460: 3459: 3455: 3452: 3451: 3450:Convolosaurus 3447: 3446: 3444: 3441: 3437: 3430: 3429: 3425: 3422: 3421: 3417: 3414: 3413: 3412:Ampelognathus 3409: 3408: 3405: 3402: 3400: 3396: 3389: 3388: 3384: 3382: 3381: 3377: 3375: 3374: 3370: 3368: 3367: 3363: 3361: 3360: 3356: 3354: 3353: 3349: 3347: 3346: 3342: 3339: 3338: 3334: 3332: 3331: 3327: 3325: 3324: 3323:Mahuidacursor 3320: 3318: 3317: 3313: 3311: 3310: 3306: 3303: 3302: 3298: 3296: 3295: 3291: 3289: 3288: 3284: 3282: 3281: 3277: 3274: 3273: 3269: 3267: 3266: 3262: 3260: 3259: 3255: 3253: 3252: 3248: 3246: 3245: 3241: 3240: 3238: 3236: 3232: 3225: 3224: 3220: 3218: 3217: 3216:Hypsilophodon 3213: 3210: 3209: 3205: 3204: 3202: 3199: 3195: 3188: 3187: 3183: 3181: 3180: 3179:Callovosaurus 3176: 3174: 3173: 3172:Burianosaurus 3169: 3168: 3165: 3161: 3158: 3155: 3150: 3146: 3142: 3137: 3133: 3108: 3107: 3106: 3105: 3103: 3101: 3097: 3089: 3085: 3084: 3083: 3082: 3079: 3076: 3074: 3070: 3062: 3058: 3057: 3056: 3055: 3052: 3049: 3047: 3043: 3039: 3031: 3027: 3024: 3022: 3018: 3015: 3013: 3009: 3006: 3004: 3000: 2997: 2995: 2991: 2989: 2985: 2984: 2979: 2975: 2971: 2964: 2959: 2957: 2952: 2950: 2945: 2944: 2941: 2934: 2923: 2910: 2906: 2902: 2898: 2891: 2884:(4): 124–131. 2883: 2879: 2872: 2864: 2860: 2856: 2852: 2848: 2844: 2840: 2836: 2829: 2819: 2805:on 2008-07-05 2801: 2797: 2793: 2786: 2784: 2783:Sauroposeidon 2775: 2773: 2771: 2762: 2758: 2754: 2750: 2746: 2742: 2738: 2734: 2730: 2726: 2722: 2715: 2707: 2703: 2698: 2693: 2688: 2683: 2679: 2675: 2672:(3): e33539. 2671: 2667: 2663: 2656: 2648: 2644: 2639: 2634: 2629: 2624: 2620: 2616: 2612: 2608: 2604: 2597: 2589: 2585: 2581: 2577: 2573: 2569: 2562: 2554: 2546: 2544: 2535: 2531: 2527: 2523: 2519: 2515: 2511: 2504: 2496: 2492: 2488: 2484: 2480: 2476: 2472: 2468: 2461: 2454: 2450: 2447: 2441: 2437: 2433: 2429: 2425: 2421: 2417: 2413: 2412:Tenontosaurus 2406: 2398: 2392: 2384: 2380: 2376: 2372: 2368: 2364: 2363:Palaeontology 2360: 2356: 2349: 2341: 2337: 2332: 2327: 2323: 2319: 2314: 2309: 2305: 2301: 2297: 2293: 2289: 2282: 2274: 2270: 2266: 2262: 2257: 2252: 2248: 2244: 2240: 2236: 2232: 2225: 2217: 2213: 2209: 2205: 2201: 2197: 2193: 2189: 2182: 2180: 2171: 2167: 2163: 2159: 2155: 2151: 2144: 2142: 2140: 2131: 2125: 2121: 2114: 2108: 2103: 2099: 2095: 2091: 2086: 2081: 2077: 2073: 2069: 2062: 2054: 2050: 2046: 2040: 2036: 2035: 2027: 2025: 2016: 2012: 2008: 2004: 2000: 1996: 1992: 1988: 1984: 1980: 1973: 1971: 1969: 1960: 1956: 1949: 1947: 1945: 1943: 1941: 1939: 1937: 1935: 1933: 1931: 1922: 1918: 1914: 1910: 1906: 1902: 1898: 1894: 1887: 1885: 1876: 1872: 1868: 1864: 1857: 1855: 1850: 1843: 1841: 1840:Tenontosaurus 1838:. This shows 1837: 1836:Tenontosaurus 1832: 1830: 1826: 1822: 1818: 1813: 1809: 1808:Tenontosaurus 1804: 1802: 1801:Tenontosaurus 1798: 1797:Tenontosaurus 1794: 1793:Tenontosaurus 1789: 1788:Tenontosaurus 1785: 1775: 1773: 1771: 1770:Paracimexomys 1766: 1762: 1761: 1757: 1753: 1749: 1748: 1743: 1742: 1737: 1735: 1730: 1726: 1724: 1719: 1717: 1712: 1711:cartilaginous 1708: 1707: 1702: 1698: 1694: 1690: 1688: 1684: 1681:included the 1680: 1679:Tenontosaurus 1676: 1675:Tenontosaurus 1672: 1671:apex predator 1668: 1666: 1662: 1658: 1656: 1655:Sauroposeidon 1651: 1650: 1646: 1642: 1639:shared their 1638: 1634: 1633:Tenontosaurus 1630: 1626: 1622: 1618: 1614: 1610: 1605: 1601: 1597: 1596:Tenontosaurus 1588: 1584: 1583: 1580: 1576: 1575: 1570: 1566: 1565: 1561: 1557: 1553: 1550: 1546: 1545: 1541: 1537: 1536: 1535:Zephyrosaurus 1531: 1527: 1524:theropod and 1523: 1519: 1518: 1513: 1512:Tenontosaurus 1509: 1508: 1503: 1502:Tenontosaurus 1494: 1485: 1483: 1479: 1475: 1471: 1467: 1466:Tenontosaurus 1463: 1462:Tyrannosaurus 1459: 1455: 1451: 1450: 1445: 1444: 1443:Tyrannosaurus 1439: 1435: 1434:Tenontosaurus 1431: 1427: 1418: 1409: 1407: 1406: 1401: 1397: 1396:Tenontosaurus 1392: 1391:Tenontosaurus 1388: 1387:Tenontosaurus 1384: 1380: 1379:Tenontosaurus 1376: 1371: 1367: 1366:Tenontosaurus 1363: 1359: 1358:Tenontosaurus 1355: 1351: 1350:Tenontosaurus 1347: 1343: 1339: 1338:Tenontosaurus 1335: 1334: 1329: 1325: 1324:Tenontosaurus 1321: 1317: 1313: 1312:Tenontosaurus 1309: 1305: 1304: 1296: 1292: 1288: 1279: 1277: 1276:Tenontosaurus 1273: 1272:Tenontosaurus 1269: 1265: 1261: 1257: 1253: 1249: 1245: 1244:Tenontosaurus 1225: 1224: 1216: 1215: 1207: 1206: 1198: 1197: 1189: 1188: 1180: 1179: 1171: 1170: 1162: 1161: 1153: 1152: 1149: 1147: 1143: 1142: 1137: 1131: 1130: 1127: 1126: 1123: 1122: 1116: 1115: 1109: 1106: 1105: 1102: 1101: 1093: 1092: 1089: 1088: 1087: 1080: 1079: 1076: 1075: 1072: 1071: 1070: 1063: 1062: 1056: 1055: 1054:Tenontosaurus 1051: 1050: 1044: 1043: 1040: 1039: 1036: 1035: 1029: 1028: 1022: 1021: 1018: 1017: 1014: 1013: 1012: 1005: 1004: 998: 997: 994: 993: 990: 989: 988: 981: 980: 974: 971: 970: 967: 966: 958: 957: 954: 953: 952: 945: 944: 941: 940: 937: 936: 935: 928: 927: 921: 920: 914: 913: 910: 909: 901: 900: 897: 896: 895: 888: 887: 884: 883: 880: 879: 878: 871: 870: 864: 863: 857: 856: 853: 852: 844: 843: 840: 839: 838: 831: 830: 827: 826: 823: 822: 821: 814: 813: 807: 806: 803: 802: 799: 798: 797: 790: 789: 786: 785: 782: 781: 780: 779:Zephyrosaurus 773: 772: 769: 768: 765: 764: 763: 762:Hypsilophodon 756: 755: 752: 751: 748: 747: 746: 739: 738: 732: 729: 728: 724: 722: 718: 713: 711: 710:Tenontosaurus 707: 703: 702:Tenontosaurus 699: 698: 693: 692:rhabdodontids 689: 688:Tenontosaurus 685: 682: 678: 674: 673:Hypsilophodon 670: 669:Tenontosaurus 666: 662: 658: 657:Tenontosaurus 650: 641: 639: 638:Tenontosaurus 635: 634:Hypsilophodon 630: 629:Tenontosaurus 626: 625:Tenontosaurus 622: 620: 615: 611: 610:Tenontosaurus 602: 600: 596: 592: 588: 584: 580: 576: 572: 568: 564: 560: 559:Tenontosaurus 555: 553: 552:Tenontosaurus 549: 545: 541: 537: 532: 530: 526: 522: 518: 515:was found in 514: 511: 510:Tenontosaurus 503: 499: 493: 488: 479: 477: 473: 472:Parker County 469: 465: 461: 457: 453: 450:in 1970) and 449: 445: 441: 440:North America 437: 433: 429: 424: 422: 419: 416: 415:iguanodontian 412: 392: 391: 377: 329: 328: 327:Tenontosaurus 314: 313: 308: 304: 299: 298: 294: 291: 290: 288: 285: 281: 277: 273: 268: 267: 266:Tenontosaurus 260: 257: 256: 253: 247: 244: 241: 240: 237: 231: 228: 225: 224: 221: 215: 212: 209: 208: 205: 199: 196: 193: 192: 189: 186: 183: 180: 179: 176: 173: 170: 169: 166: 163: 160: 159: 156: 153: 150: 149: 144: 139: 135: 132: 128: 124: 120: 115: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 47: 44:115–108  40: 36: 32: 26: 25:Tenontosaurus 22: 19: 5315: 5280: 5273: 5265: 5253: 5222: 5204: 5197: 5190: 5183: 5176: 5169: 5162: 5155: 5148: 5141: 5134: 5127: 5120: 5113: 5106: 5099: 5093:Claorhynchus 5091: 5084: 5077: 5069:Nomina dubia 5067: 5057:Guegoolithus 5055: 5035: 5028: 5023:Staurichnium 5021: 5016:Sousaichnium 5014: 5007: 5000: 4993: 4986: 4979: 4972: 4967:Dakotasaurus 4965: 4960:Caririchnium 4958: 4951: 4944: 4937: 4930: 4874: 4842: 4835: 4828: 4821: 4814: 4807: 4787: 4780: 4773: 4766:Saurolophini 4753: 4746: 4739: 4732: 4725: 4717: 4710:Kritosaurini 4697: 4690: 4685:Huallasaurus 4683: 4676: 4657:Wulagasaurus 4655: 4648: 4641: 4634: 4627: 4620: 4600: 4572: 4565: 4558: 4551: 4546:Lambeosaurus 4544: 4537: 4530: 4523: 4503: 4496: 4489: 4482: 4462: 4455: 4448: 4441: 4433: 4426: 4419: 4412: 4405: 4398: 4391: 4384: 4377: 4370: 4348:Yamatosaurus 4346: 4339: 4332: 4325: 4318: 4313:Aquilarhinus 4311: 4285: 4278: 4271: 4264: 4259:Tethyshadros 4257: 4250: 4243: 4236: 4229: 4222: 4215: 4208: 4201: 4194: 4189:Plesiohadros 4187: 4180: 4173: 4166: 4161:Lophorhothon 4159: 4152: 4144: 4137: 4130: 4123: 4116: 4109: 4102: 4095: 4088: 4081: 4074: 4067: 4060: 4053: 4047:Batyrosaurus 4045: 4040:Bactrosaurus 4038: 4031: 3977: 3920: 3913: 3906: 3898: 3880:Ouranosaurus 3878: 3871: 3864: 3857: 3850: 3845:Brighstoneus 3843: 3836: 3812: 3805: 3798: 3791: 3784: 3777: 3772:Osmakasaurus 3770: 3765:Oblitosaurus 3763: 3756: 3749: 3742: 3735: 3728: 3721: 3714: 3707: 3699: 3692: 3687:Camptosaurus 3685: 3678: 3671: 3664: 3634: 3628:Kangnasaurus 3626: 3618: 3611: 3604: 3597: 3590: 3583:Dryosauridae 3570: 3563: 3556: 3518: 3511: 3504: 3497: 3490: 3485:Matheronodon 3483: 3464: 3463: 3456: 3448: 3426: 3418: 3410: 3385: 3378: 3371: 3364: 3359:Sektensaurus 3357: 3352:Qantassaurus 3350: 3343: 3335: 3328: 3321: 3314: 3307: 3301:Kangnasaurus 3299: 3292: 3285: 3278: 3270: 3265:Diluvicursor 3263: 3256: 3249: 3242: 3223:Vectidromeus 3221: 3214: 3206: 3184: 3177: 3170: 3088:Ornithischia 3073:Ornithischia 3025: 3016: 3012:Ornithischia 3007: 2998: 2900: 2896: 2890: 2881: 2877: 2871: 2838: 2834: 2828: 2818: 2807:. Retrieved 2800:the original 2795: 2791: 2782: 2728: 2724: 2714: 2669: 2665: 2655: 2610: 2606: 2596: 2571: 2567: 2561: 2555:(164): 1–27. 2552: 2517: 2513: 2509: 2503: 2470: 2466: 2460: 2423: 2419: 2415: 2411: 2405: 2391:cite journal 2366: 2362: 2358: 2348: 2295: 2291: 2281: 2238: 2235:Paleobiology 2234: 2224: 2191: 2187: 2156:(1): 53–70. 2153: 2149: 2119: 2113: 2075: 2071: 2061: 2033: 1982: 1978: 1958: 1955:The Mosasaur 1954: 1896: 1892: 1866: 1862: 1839: 1835: 1833: 1807: 1806:At the time 1805: 1800: 1796: 1792: 1787: 1781: 1768: 1764: 1758: 1751: 1745: 1739: 1732: 1721: 1714: 1706:Bernissartia 1704: 1696: 1692: 1687:Albanerpeton 1685: 1678: 1674: 1663: 1653: 1647: 1636: 1632: 1621:river deltas 1599: 1595: 1593: 1581: 1572: 1562: 1556:ornithomimid 1544:Microvenator 1542: 1533: 1515: 1511: 1505: 1501: 1499: 1488:Paleoecology 1473: 1470:histological 1465: 1461: 1457: 1447: 1441: 1433: 1423: 1412:Reproduction 1403: 1399: 1395: 1390: 1386: 1382: 1378: 1374: 1369: 1365: 1361: 1357: 1353: 1349: 1345: 1341: 1337: 1331: 1327: 1323: 1319: 1315: 1311: 1307: 1301: 1299: 1294: 1290: 1275: 1271: 1243: 1241: 1233:Paleobiology 1146:hadrosaurids 1139: 1134: 1121:Dryosauridae 1119: 1085: 1084: 1083: 1068: 1067: 1066: 1053: 1052: 1032: 1009: 1008: 985: 984: 973:Iguanodontia 949: 948: 932: 931: 894:Parksosaurus 892: 891: 875: 874: 837:Jeholosaurus 835: 834: 818: 817: 794: 793: 777: 776: 760: 759: 743: 742: 720: 714: 709: 705: 701: 695: 687: 676: 672: 668: 656: 655: 637: 633: 628: 624: 623: 618: 613: 609: 608: 558: 556: 551: 533: 529:Barnum Brown 509: 507: 501: 491: 459: 455: 451: 443: 434:ages of the 425: 404:sinew lizard 326: 325: 324: 311: 310: 296: 295: 265: 264: 242: 226: 210: 204:Ornithischia 194: 181: 126: 24: 18: 5435:Iguanodonts 5282:Willinakaqe 5254:nomina nuda 5171:Siluosaurus 5136:Loncosaurus 4974:Dinehichnus 4816:Kamuysaurus 4789:Saurolophus 4741:Kritosaurus 4734:Gryposaurus 4567:Sahaliyania 4553:Magnapaulia 4525:Amurosaurus 4505:Tlatolophus 4484:Adelolophus 4428:Kazaklambia 4407:Blasisaurus 4400:Arenysaurus 4393:Aralosaurus 4327:Hadrosaurus 4320:Eotrachodon 4273:Yunganglong 4238:Sirindhorna 4210:Protohadros 4146:Koshisaurus 4139:Jintasaurus 3908:Comptonatus 3758:Napaisaurus 3744:Lurdusaurus 3716:Fukuisaurus 3694:Cedrorestes 3636:Valdosaurus 3499:Pareisactus 3330:Morrosaurus 3294:Isasicursor 3258:Chakisaurus 3154:Ornithopoda 3141:Ornithopoda 3100:Ornithopoda 2970:Ornithopoda 2416:Deinonychus 1817:subtropical 1760:Lepisosteus 1752:Naomichelys 1741:Goniopholis 1701:crurotarsan 1637:Deinonychus 1600:Deinonychus 1564:Gobiconodon 1526:goniopholid 1522:allosauroid 1517:Deinonychus 1474:T. tilletti 1400:Deinonychus 1383:Deinonychus 1375:Deinonychus 1370:Deinonychus 1362:Deinonychus 1354:Deinonychus 1346:Deinonychus 1342:Deinonychus 1328:Deinonychus 1320:Deinonychus 1316:Deinonychus 1303:Deinonychus 1295:Deinonychus 1291:T. tilletti 1260:gymnosperms 1138:(including 1069:T. tilletti 934:Bugenasaura 796:Yandusaurus 731:Ornithopoda 614:T. tilletti 605:Description 548:John Ostrom 492:T. tilletti 456:T. tilletti 448:John Ostrom 297:T. tilletti 236:Ornithopoda 5414:Categories 5224:Arkharavia 5199:Tichosteus 5192:Thespesius 5164:Pteropelyx 5157:Orthomerus 5037:Wintonopus 4622:Acristavus 4602:Barsboldia 4560:Olorotitan 4435:Latirhinus 4287:Zuoyunlong 4154:Levnesovia 4104:Gobihadros 4069:Datonglong 4062:Claosaurus 4033:Altirhinus 3943:see below↓ 3873:Morelladon 3751:Magnamanus 3723:Hippodraco 3592:Dryosaurus 3565:Hesperonyx 3540:Dryomorpha 3380:Trinisaura 3373:Tietasaura 3366:Talenkauen 3244:Anabisetia 3109:see below↓ 3003:Dinosauria 2809:2007-07-07 1961:: 151–163. 1846:References 1729:ray-finned 1689:arthridion 1659:, and the 1609:floodplain 1507:Sauropelta 1458:Allosaurus 1449:Allosaurus 1426:thigh bone 1333:Sauropelta 1262:, such as 1252:tree ferns 1108:Dryomorpha 1011:Anabisetia 987:Talenkauen 745:Orodromeus 684:iguanodont 508:The first 418:ornithopod 188:Dinosauria 5206:Trachodon 5122:Glishades 5101:Diclonius 4939:Anomoepus 4923:Ichnotaxa 4636:Maiasaura 4574:Velafrons 4083:Equijubus 4055:Choyrodon 3915:Iguanodon 3779:Planicoxa 3709:Dakotadon 3680:Calvarius 3506:Rhabdodon 3492:Mochlodon 3235:Elasmaria 2986:Kingdom: 2909:0567-7920 2863:0272-4634 2855:1937-2809 2753:0272-4634 2322:1932-6203 2273:0094-8373 2265:1938-5331 2216:0272-4634 2170:0567-7920 2102:221404013 2053:985402380 2007:0272-4634 1875:0079-032X 1869:: 1–234. 1825:Louisiana 1734:Gyronchus 1683:amphibian 1667:atokensis 1661:carnosaur 1645:sauropods 1627:. In the 1625:Louisiana 1430:shin bone 1310:remains. 1282:Predators 1141:Iguanodon 717:cladogram 681:dryomorph 677:Iguanodon 464:specimens 161:Kingdom: 155:Eukaryota 5345:47460280 5325:Wikidata 5086:Cionodon 4755:Rhinorex 4643:Ornatops 4443:Minqaria 4414:Canardia 4341:Malefica 4266:Xuwulong 4231:Siamodon 4132:Jeyawati 4118:Gonkoken 4076:Eolambia 3900:Barilium 3701:Cumnoria 3666:Barilium 3572:Owenodon 3558:Draconyx 3520:Zalmoxes 3420:Fostoria 3272:Fostoria 3030:Cerapoda 2994:Chordata 2992:Phylum: 2988:Animalia 2761:83674985 2706:22470454 2666:PLOS ONE 2647:18195356 2534:84175628 2495:84672219 2449:Archived 2446:abstract 2340:37285376 2331:10246810 2292:PLOS ONE 2094:32869488 1821:tropical 1747:Glyptops 1723:Lissodus 1703:reptile 1657:proteles 1649:Astrodon 1579:lungfish 1574:Glyptops 1571:such as 1552:sauropod 1478:ontogeny 1454:theropod 1352:. Adult 1086:T. dossi 723:, 2011. 619:T. dossi 599:Maryland 567:Oklahoma 460:T. dossi 421:dinosaur 312:T. dossi 175:Chordata 171:Phylum: 165:Animalia 151:Domain: 127:T. dossi 5384:4947357 5371:1043766 5358:4823164 5331:Q131103 4379:Ajnabia 3814:Uteodon 2733:Bibcode 2697:3314665 2674:Bibcode 2638:2206579 2615:Bibcode 2576:Bibcode 2475:Bibcode 2428:Bibcode 2371:Bibcode 2300:Bibcode 2243:Bibcode 2196:Bibcode 2015:4523326 1987:Bibcode 1921:4523810 1901:Bibcode 1782:In the 1778:Climate 1736:dumblei 1716:Hybodus 1569:turtles 1270:trees. 1264:conifer 544:Wyoming 521:Montana 409:) is a 398:  284:Species 258:Genus: 5048:Ootaxa 4245:Tanius 3838:Bolong 2907:  2861:  2853:  2759:  2751:  2704:  2694:  2645:  2635:  2532:  2493:  2338:  2328:  2320:  2271:  2263:  2214:  2168:  2126:  2100:  2092:  2051:  2041:  2013:  2005:  1919:  1873:  1772:crossi 1727:, the 1725:anitae 1718:buderi 1709:, the 1699:, the 1560:mammal 1538:, the 1530:Albian 1326:, and 1268:ginkgo 1256:cycads 593:, and 523:by an 513:fossil 432:Albian 428:Aptian 272:Ostrom 39:Albian 35:Aptian 5397:38741 5366:IRMNG 4090:Fylax 3620:Iyuku 3026:Clade 3017:Clade 3008:Clade 2999:Clade 2851:eISSN 2803:(PDF) 2788:(PDF) 2757:S2CID 2530:S2CID 2491:S2CID 2261:eISSN 2194:(2). 2098:S2CID 2011:JSTOR 1917:JSTOR 1731:fish 1713:fish 1438:birds 1373:that 1248:ferns 721:et al 583:Idaho 575:Texas 476:Texas 411:genus 243:Clade 227:Clade 211:Clade 195:Clade 182:Clade 5353:GBIF 3786:Proa 3458:Iani 3086:see 3059:see 2905:ISSN 2859:ISSN 2823:264. 2749:ISSN 2702:PMID 2643:PMID 2397:link 2336:PMID 2318:ISSN 2269:ISSN 2212:ISSN 2166:ISSN 2124:ISBN 2090:PMID 2049:OCLC 2039:ISBN 2003:ISSN 1871:ISSN 1812:arid 1767:and 1750:and 1720:and 1695:and 1635:and 1598:and 1554:and 1460:and 1446:and 1428:and 1266:and 1250:and 1238:Diet 715:The 706:Iani 697:Iani 591:Utah 395:lit. 386:-tə- 303:type 276:1970 52:PreꞒ 5340:EoL 2843:doi 2741:doi 2692:PMC 2682:doi 2633:PMC 2623:doi 2611:105 2584:doi 2570:". 2522:doi 2483:doi 2436:doi 2379:doi 2326:PMC 2308:doi 2251:doi 2204:doi 2158:doi 2080:doi 1995:doi 1909:doi 1819:to 1756:gar 597:of 589:of 581:of 573:of 565:in 478:. 470:of 430:to 413:of 390:-əs 388:SOR 384:NON 382:ti- 367:ɔːr 41:), 37:to 5416:: 5394:: 5381:: 5368:: 5355:: 5342:: 5327:: 3028:: 3019:: 3010:: 3001:: 2901:46 2899:. 2882:57 2880:. 2857:. 2849:. 2839:22 2837:. 2796:65 2794:. 2790:. 2769:^ 2755:. 2747:. 2739:. 2729:33 2727:. 2723:. 2700:. 2690:. 2680:. 2668:. 2664:. 2641:. 2631:. 2621:. 2609:. 2605:. 2582:. 2542:^ 2528:. 2518:48 2516:. 2489:. 2481:. 2471:27 2469:. 2434:. 2424:15 2422:. 2393:}} 2389:{{ 2377:. 2367:54 2365:. 2334:. 2324:. 2316:. 2306:. 2296:18 2294:. 2290:. 2267:. 2259:. 2249:. 2239:46 2237:. 2233:. 2210:. 2202:. 2192:41 2190:. 2178:^ 2164:. 2154:59 2152:. 2138:^ 2096:. 2088:. 2076:95 2074:. 2070:. 2047:. 2023:^ 2009:. 2001:. 1993:. 1983:10 1981:. 1967:^ 1957:. 1929:^ 1915:. 1907:. 1897:17 1895:. 1883:^ 1867:35 1865:. 1853:^ 1774:. 1582:. 1567:, 1484:. 1408:. 1254:, 1144:, 601:. 585:, 577:, 569:, 519:, 474:, 393:; 274:, 245:: 229:: 213:: 197:: 184:: 102:Pg 46:Ma 5256:) 5096:? 4722:? 4660:? 4438:? 4149:? 4050:? 3925:? 3903:? 3704:? 3631:? 3623:? 3453:? 3442:? 3431:? 3423:? 3415:? 3390:? 3340:? 3304:? 3275:? 3226:? 3211:? 3200:? 3189:? 2962:e 2955:t 2948:v 2911:. 2865:. 2845:: 2812:. 2781:" 2763:. 2743:: 2735:: 2708:. 2684:: 2676:: 2670:7 2649:. 2625:: 2617:: 2590:. 2586:: 2578:: 2536:. 2524:: 2497:. 2485:: 2477:: 2455:) 2444:( 2442:. 2438:: 2430:: 2414:- 2399:) 2385:. 2381:: 2373:: 2342:. 2310:: 2302:: 2275:. 2253:: 2245:: 2218:. 2206:: 2198:: 2172:. 2160:: 2132:. 2104:. 2082:: 2055:. 2017:. 1997:: 1989:: 1959:2 1923:. 1911:: 1903:: 1877:. 407:' 401:' 376:/ 373:s 370:ə 364:s 361:ˈ 358:ə 355:t 352:n 349:ɒ 346:n 343:ˌ 340:ɪ 337:t 334:/ 330:( 309:† 305:) 293:† 262:† 249:† 233:† 217:† 201:† 107:N 97:K 92:J 87:T 82:P 77:C 72:D 67:S 62:O 57:Ꞓ 33:(

Index

Early Cretaceous
Aptian
Albian
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Perot Museum of Nature and Science
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Ornithischia
Neornithischia
Ornithopoda
Tenontosauridae
Tenontosaurus
Ostrom

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