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Slavic migrations to the Balkans

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event seen in our six Eastern Europe populations between northern European and southern European ancestral sources may correspond to the expansion of Slavic language speaking groups (commonly referred to as the Slavic expansion) across this region at a similar time, perhaps related to displacement caused by the Eurasian steppe invaders (38; 58). Under this scenario, the northerly source in the second event might represent DNA from Slavic-speaking migrants (sampled Slavic-speaking groups are excluded from being donors in the EastEurope I analysis). To test consistency with this, we repainted these populations adding the Polish as a single Slavic-speaking donor group ("East Europe II" analysis; see Note S7.6) and, in doing so, they largely replaced the original North European component (Figure S21), although we note that two nearby populations, Belarus and Lithuania, are equally often inferred as sources in our original analysis (Table S12). Outside these six populations, an admixture event at the same time (910CE, 95% CI:720-1140CE) is seen in the southerly neighboring Greeks, between sources represented by multiple neighboring Mediterranean peoples (63%) and the Polish (37%), suggesting a strong and early impact of the Slavic expansions in Greece, a subject of recent debate (37). These shared signals we find across East European groups could explain a recent observation of an excess of IBD sharing among similar groups, including Greece, that was dated to a wide range between 1,000 and 2,000 years ago (37)
397:, "impeded coordinated military resistance against the enemy", which put them in a situation of being under foreign political leadership. When the Slavs and later the Avars entered the southeast of Europe they lacked advanced siege-warfare tactics, but around 587 they acquired this knowledge from contact with Byzantine culture, and because of this no urban settlement or fort could oppose them any more. With the destruction of Roman fortifications came a loss of Byzantine military and administrative power in Roman provinces. The native population was often decimated, and smaller or larger groups of Slavs settled in the devastated lands. Settlement among the natives, often replacing them, happened in the autumn, when winter supplies were secured for the people and animals. After mixing with the natives who survived in smaller communities, depending on the region, the Slavic tribes mostly had names of 887:, based on 146 samples, confirmed that the spread of Slavic language and identity was because of large movements of people of both males and females with specific Eastern European ancestry and that "more than half of the ancestry of most peoples in the Balkans today comes from the Slavic migrations, with around a third Slavic ancestry even in countries like Greece where no Slavic languages are spoken today". The big data set also showed that the Y-DNA haplogroups I2a-L621 and R1a-Z282 are absent in the antiquity and appear only since the Early Middle Ages "always associated with Eastern European related ancestry in the autosomal genome, which supports that these lineages were introduced in the Balkans by Eastern European migrants during the Early Medieval period". 2374:
around 300–500 CE... The SNP-based age of the Eastern European CTS10228 branch is 2200 ± 300 years old. The carriers of the most ancient subgroup live in Southeast Poland, and it is likely that the rapid demographic expansion which brought the marker to other regions in Europe began there. The largest demographic explosion occurred in the Balkans, where the subgroup is dominant in 50.5% of Croatians, 30.1% of Serbs, 31.4% of Montenegrins, and in about 20% of Albanians and Greeks. As a result, this subgroup is often called Dinaric. It is interesting that while it is dominant among modern Balkan peoples, this subgroup has not been present yet during the Roman period, as it is almost absent in Italy as well (see Online Resource 5; ESM_5).
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sikeresen integrálódott egy olyan társadalomba, amely hamarosan erőteljes demográfiai expanzióba kezdett. Ez is mutatja, hogy nem feltétlenül népek, mintsem családok sikerével, nemzetségek elterjedésével is számolnunk kell, és ezt a jelenlegi etnikai identitással összefüggésbe hozni lehetetlen. A csoport elterjedése alapján valószínűsíthető, hogy a szláv népek migrációjában vett részt, így válva az R1a-t követően a második legdominánsabb csoporttá a mai Kelet-Európában. Nyugat-Európából viszont teljes mértékben hiányzik, kivéve a kora középkorban szláv nyelvet beszélő keletnémet területeket.
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present-day Ukraine... The calculated age of this specific haplogroup together with the variation peak detected in the suggested Slavic homeland could represent a signal of Slavic migration arising from medieval Slavic expansions. However, the strong genetic barrier around the area of Bosnia and Herzegovina, associated with the high frequency of the I2a1b-M423 haplogroup, could also be a consequence of a Paleolithic genetic signal of a Balkan refuge area, followed by mixing with a medieval Slavic signal from modern-day Ukraine.
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Slavic languages". According to Kushniarevich et al. 2015, the Hellenthal et al. 2014 IBD analysis, also found "multi-directional admixture events among East Europeans (both Slavic and non-Slavic), dated to around 1,000–1,600 YBP" which coincides with "the proposed time-frame for the Slavic expansion". The Slavic influence is "dated to 500-900 CE or a bit later with over 40-50% among
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Based on SNP analysis, the CTS10228 group is 2200 ± 300 years old. The group's demographic expansion may have begun in Southeast Poland around that time, as carriers of the oldest subgroup are found there today. The group cannot solely be tied to the Slavs, because the proto-Slavic period was later,
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R1a-M458 exceeds 20% in the Czech Republic, Slovakia, Poland, and Western Belarus. The lineage averages 11–15% across Russia and Ukraine and occurs at 7% or less elsewhere (Figure 2d). Unlike hg R1a-M458, the R1a-M558 clade is also common in the Volga-Uralic populations. R1a-M558 occurs at 10–33% in
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Syriac chroniclers (along with their Arab, Byzantine, Latin, Armenian, and Georgian counterparts) did not use ethnonyms as specifically as modern scholars do. As K. Czeglédy notes, "some sources... use the ethnonyms of the various steppe peoples, in particular those of the Scythians, Huns and Turkic
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Admixture analysis of autosomal SNPs of the Southeast Europe in a global context on the resolution level of 7 assumed ancestral populations: the African (brown), South/West European (light blue), Asian (yellow), Middle Eastern (orange), South Asian (green), North/East European (dark blue) and beige
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S7.6 "East Europe": The difference between the 'East Europe I' and 'East Europe II' analyses is that the latter analysis included the Polish as a potential donor population. The Polish were included in this analysis to reflect a Slavic language speaking source group." "We speculate that the second
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analysis, the South Slavs show a genetic uniformity, with a modeled ancestral genetic component in the study peaking in Baltic speakers, being high in East Slavs (80-95%) as well as Western and North-Western Europeans (Germans, Orcadians, Swedes), and between 55-70% among South Slavs. According to
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and Slavic expansion, which was a "relatively small population that expanded over a large geographic area", particularly "the expansion of the Slavic populations into regions of low population density beginning in the sixth century" and that it is "highly coincident with the modern distribution of
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Az I2-CTS10228 (köznevén „dinári-kárpáti") alcsoport legkorábbi közös őse 2200 évvel ezelőttre tehető, így esetében nem arról van szó, hogy a mezolit népesség Kelet-Európában ilyen mértékben fennmaradt volna, hanem arról, hogy egy, a mezolit csoportoktól származó szűk család az európai vaskorban
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However, a study by Battaglia et al. (2009) showed a variance peak for I2a1 in the Ukraine and, based on the observed pattern of variation, it could be suggested that at least part of the I2a1 haplogroup could have arrived in the Balkans and Slovenia with the Slavic migrations from a homeland in
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in Southeastern Europe. Pre-Slavic Southeast European populations have the most of the Anatolian Neolithic component of ancestry, whereas present-day Slavs outside the Southeast Europe have the least, "with present-day people from Southeastern Europe intermediate between the two extremes". Among
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parts of Russia, exceeds 26% in Poland and Western Belarus, and varies between 10 and 23% in the Ukraine, whereas it drops 10-fold lower in Western Europe. In general, both R1a-M458 and R1a-M558 occur at low but informative frequencies in Balkan populations with known Slavonic heritage.
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Hg I2a1a2b-L621 was present in 5 Conqueror samples, and a 6th sample form Magyarhomorog (MH/9) most likely also belongs here, as MH/9 is a likely kin of MH/16 (see below). This Hg of European origin is most prominent in the Balkans and Eastern Europe, especially among Slavic speaking
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Ulf Büntgen; Vladimir S. Myglan; Fredrik Charpentier Ljungqvist; Michael McCormick; Nicola Di Cosmo; Michael Sigl; Johann Jungclaus; Sebastian Wagner; Paul J. Krusic; Jan Esper; Jed O. Kaplan; Michiel A. C. de Vaan; Jürg Luterbacher; Lukas Wacker; Willy Tegel (2016).
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The geographic distributions of the major eastern European NRY haplogroups (R1a-Z282, I2a-P37) overlap with the area occupied by the present-day Slavs to a great extent, and it might be tempting to consider both haplogroups as Slavic-specic patrilineal
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Very soon after their arrival the typical Slavic archaeological culture was changed by the influence of native Byzantine cultures. According to the archaeological data the main movement of the Slavs was from the Middle and Lower Danube valley. The
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they lacked political organization, because of which the Byzantine Empire regained control there, and after 200 years the Slavs in the southern Balkans became assimilated by the Greek-speaking majority. In the territory of present-day
306:, with the intruders spending in 551 their time in Dalmatia "as if in their own land", but also occasioned peaceful permanent settlement on Byzantine territory, which began around the same time. Things changed with the arrival of the 854:
and their subclades R-M558, R-M458 and I-CTS10228 among South Slavs are in correlation with the spreading of Slavic languages during the medieval Slavic expansion from Eastern Europe, most probably from the territory of present-day
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The 2006 Y-DNA study results "suggest that the Slavic expansion started from the territory of present-day Ukraine, thus supporting the hypothesis that places the earliest known homeland of Slavs in the basin of the middle
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Like the name Scythian up to the early medieval period, the name Hun became a generic (usually pejorative) term in subsequent history for any steppe-warrior people, or even any enemy people, regardless of their actual
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According to archaeological data and historical sources, the Slavs mostly travelled along the river valleys, but while in the Southern Balkans, they travelled where they encountered greater resistance by the native
603:, alongside most of post-Roman Europe. Many Slavs soon began to accept the cultural customs of the highly civilized Roman provinces, and in order to expand their cultural and state influence on the South Slavs, the 745:. Those living near lakes, rivers, and seas also used various hooks and nets for fishing. They were known to be especially skilled in woodworking and shipbuilding, but also knew about metalworking and pottery. 2786:
Vedriš, Trpimir (2015). "Balkanske sklavinije i Bugarska – Hrvatska u međunarodnom kontekstu" [Balkan 'sklavinias' and Bulgaria – Croatia in the international context]. In Zrinka Nikolić Jakus (ed.).
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recorded that in 518 a large army of the Antae, "who dwell close to the Sclaveni", crossed the Danube River into Roman territory. Raids continued with ever-faster and stronger incursions during the time of
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Iosif Lazaridis; Songül Alpaslan-Roodenberg; Ayşe Acar; Ayşen Açıkkol; Anagnostis Agelarakis; Levon Aghikyan; Uğur Akyüz; Desislava Andreeva; Gojko Andrijašević; David Reich; et al. (26 August 2022).
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The ethnonym of the Huns, like those of Scythians and Türks, became a generic term for steppe-people (nomads) and invading enemies from the East, no matter their actual origin and identity.
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Fóthi, E.; Gonzalez, A.; Fehér, T.; et al. (2020), "Genetic analysis of male Hungarian Conquerors: European and Asian paternal lineages of the conquering Hungarian tribes",
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with some elements of the so-called Martinovka culture. V. V. Sedov considered that the majority of the Slavic population in Southeast Europe was descending from Antae, while
337:, Avars and Slavs made devastating intrusions along the Byzantine borders from Northern Italy to Southern Greece, and by the mid-7th century, the Slavs had settled in all the 251:
and of various Germanic tribes from the end of the 5th century CE. The first certain Slavic raids date to the early 6th century during the time of the Eastern Roman Emperor
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compared ancient, medieval and modern population samples and found that the medieval Slavic migrations "profoundly affected the region", resulting in the reduction of
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was pillaged almost every year by Huns, Sclaveni, and Antae, who did enormous damage to the native Roman population, making the region a "Scythian desert". As the
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present-day populations "Greeks and Albanians have more Anatolian Neolithic ancestry than their South Slavic neighbors". A 2023 archaeogenetic study published in
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Eventually the Slavs settled in the former Roman provinces of Pannonia and Dalmatia reached a substantial amount of autonomy or independence, establishing
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in which a council of nobles ruled the tribal community. This allowed Slav tribes to stay together regardless of environmental factors, but according to
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Koder, Johannes (2020). "On the Slavic Immigration in the Byzantine Balkans". In Johannes Preiser-Kapeller; Lucian Reinfandt; Yannis Stouraitis (eds.).
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since late 6th and 7th century, while Greece surely only since the 7th century (although military invasions could be argued since mid-6th century).
2899: 693:) near the border with Slovenia and Croatia are radiocarbon dated to the first-third of the 6th century. In Bulgaria and countries of former 1794:
Hellenthal, Garrett; Busby, George B.J.; Band, Gavin; Wilson, James F.; Capelli, Cristian; Falush, Daniel; Myers, Simon (14 February 2014).
4144: 4139: 3816: 3804: 3775: 3729: 3634: 3399: 3387: 3079: 2872: 3610: 3569: 3416: 3282: 3196: 3164: 3038: 2972: 1270:(1981). "The Evolution of Slavic Society and the Slavic Invasions in Greece. The First Major Slavic Attack on Thessaloniki, A. D. 597". 835:
2017 admixture study of Peloponnesian Greek population, "the Slavic ancestry of Peloponnesean subpopulations ranges from 0.2 to 14.4%".
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Migration Histories of the Medieval Afroeurasian Transition Zone: Aspects of Mobility Between Africa, Asia and Europe, 300-1500 C.E.
3717: 3507: 3792: 2241: 1668:"Standing at the Gateway to Europe - The Genetic Structure of Western Balkan Populations Based on Autosomal and Haploid Markers" 109:, Byzantium was not able to wage war on two fronts and regain its lost territories, so it reconciled with the establishment of 826:
and that there's an "even patterns of IBD sharing among East-West Slavs–'inter-Slavic' populations (Hungarians, Romanians and
3995: 2882: 2850: 2804: 2768: 2333: 1522: 1503: 1481: 1450: 1419: 1202: 1183: 988: 906: 896: 417: 1922:"Genetic Heritage of the Balto-Slavic Speaking Populations: A Synthesis of Autosomal, Mitochondrial and Y-Chromosomal Data" 685:
in Moldova are dated to the 5th century, in Romania since 6th century, from there to Transylvania in mid-6th century (with
3918: 3809: 2426: 2181: 2038: 405: 2825: 2725: 2704: 2680: 2624: 2603: 2582: 2233:Генофонд українців за різними системами генетичних маркерів: походження і місце на європейському генетичному просторі 1122: 844: 2085:
Rebała, K; Mikulich, AI; Tsybovsky, IS; Siváková, D; Dzupinková, Z; Szczerkowska-Dobosz, A; Szczerkowska, Z (2007).
1543:[First contacts of the Danubian Slavs and Avars: the evidences from the Byzantine writers and archaeology]. 4159: 3043: 2257:"Y-chromosome haplogroups from Hun, Avar and conquering Hungarian period nomadic people of the Carpathian Basin" 2238:
The gene pool of Ukrainians revealed by different systems of genetic markers: the origin and statement in Europe
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The settlement was facilitated by the substantial decrease of the Southeastern European population during the
4154: 2034:"Genetics of the peloponnesean populations and the theory of extinction of the medieval peloponnesean Greeks" 663: 612: 381: 357: 78: 1016: 193:(from 541–549 until the mid-8th century) decimated the native population, resulting in the weakening of the 4149: 2486:"Ancient DNA analysis reveals how the rise and fall of the Roman Empire shifted populations in the Balkans" 769:
survey "of recent genealogical ancestry over the past 3,000 years at a continental scale", the speakers of
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Underhill, Peter A. (2015), "The phylogenetic and geographic structure of Y-chromosome haplogroup R1a",
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assimilation and additional Slavic waves since mid-7th century). In Southwestern Hungary (southwest of
1784:"Companion website for "A genetic atlas of human admixture history", Hellenthal et al, Science (2014)" 2690: 492: 443: 299: 2891: 2128:"The paternal perspective of the Slovenian population and its relationship with other populations" 1854:
Hellenthal, G.; Busby, G. B.; Band, G.; Wilson, J. F.; Capelli, C.; Falush, D.; Myers, S. (2014).
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in the summer of 626, they remained in the wider Southeast Europe area after they had settled the
2485: 2132: 1333: 1108: 770: 322: 140: 2423: 877: 682: 667: 641: 116: 1020:"Cooling and societal change during the Late Antique Little Ice Age from 536 to around 660 AD" 851: 34:. The rapid demographic spread of the Slavs was followed by a population exchange, mixing and 4103: 2789:
Nova zraka u europskom svjetlu: Hrvatske zemlje u ranome srednjem vijeku (oko 550 − oko 1150)
365: 3797: 843:". According to genetic studies until 2020, the distribution, variance and frequency of the 2672: 2517: 2274: 2087:"Y-STR variation among Slavs: Evidence for the Slavic homeland in the middle Dnieper basin" 1935: 1867: 1809: 1681: 1031: 645: 545: 182: 70: 1541:"Первые контакты дунайских славян и авар: свидетельства византийских авторов и археологии" 8: 3888: 3241: 2717:
Byzantium and the Avars, 6th-9th Century AD: Political, Diplomatic and Cultural Relations
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Empires of the Silk Road: A History of Central Eurasia from the Bronze Age to the Present
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The Early Medieval Balkans: A Critical Survey from the Sixth to the Late Twelfth Century
2546: 2454: 2427:"A genetic probe into the ancient and medieval history of Southern Europe and West Asia" 2278: 1939: 1871: 1813: 1685: 1035: 3935: 3586: 2774: 2467: 2295: 2261: 2256: 2203: 2157: 2062: 2033: 2009: 1982: 1958: 1921: 1888: 1855: 1838: 1795: 1760: 1731: 1704: 1667: 1287: 2595:
The Making of the Slavs: History and Archaeology of the Lower Danube Region, c.500–700
2399: 3753: 3682: 3534: 3324: 2878: 2861: 2846: 2821: 2800: 2778: 2764: 2721: 2700: 2676: 2668: 2650: 2620: 2599: 2578: 2471: 2459: 2387: 2329: 2300: 2231: 2208: 2149: 2127: 2108: 2067: 2014: 1963: 1893: 1843: 1825: 1800: 1765: 1709: 1118: 984: 872: 831: 633: 530: 394: 346: 263: 231: 31: 2161: 4005: 3181: 3028: 2874:Јужни Словени под византијском влашћу (South Slavs under Byzantine rule 600-1025AD) 2756: 2541: 2531: 2449: 2441: 2395: 2362: 2290: 2282: 2198: 2190: 2141: 2098: 2057: 2047: 2004: 1994: 1953: 1943: 1883: 1875: 1833: 1817: 1755: 1745: 1699: 1689: 1556: 1279: 1039: 774: 726: 702: 671: 629: 588: 373: 361: 128: 4010: 3780: 3768: 3274: 2836: 2815: 2796: 2750: 2734: 2715: 2694: 2658: 2654: 2646: 2638: 2634: 2614: 2593: 2572: 2323: 2270: 2145: 1948: 1750: 1694: 1536: 1112: 978: 675: 596: 432: 334: 307: 210: 194: 62: 23: 3360: 404:
Slavs established dense settlements in Southeast Europe, more precisely in the
3746: 3392: 3334: 3105: 2662: 2536: 2522: 2367: 2286: 2240:] (PhD) (in Ukrainian). National Research Center for Radiation Medicine of 1983:"Genetic Landscape of Slovenians: Past Admixture and Natural Selection Pattern" 1267: 883: 867: 847: 790: 515: 387:
According to Procopius, Slavic social and political organization was a kind of
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began the reconstruction of fortresses, cities, and Christianity. However, the
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gave more emphasis on the Sclaveni as immigration started in Western Ukraine.
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lacked garrisons, in 545 Justinian I made an alliance with the Antae to stop
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Pamjav, Horolma; Fehér, Tibor; Németh, Endre; Koppány Csáji, László (2019).
1879: 1821: 1310: 1019: 500: 3999: 3943: 3851: 3690: 3667: 3639: 3574: 3512: 3472: 3433: 3421: 3411: 3404: 3299: 3287: 3213: 3110: 2989: 2942: 2518:"A genetic history of the Balkans from Roman frontier to Slavic migrations" 2490: 2463: 2304: 2212: 2153: 2112: 2071: 2018: 1967: 1897: 1847: 1769: 1736: 1713: 1272:
Hesperia: The Journal of the American School of Classical Studies at Athens
710: 690: 549: 477: 447: 421: 148: 86: 45: 3627: 247:. Small groups of Slavs had probably participated in the campaigns of the 3968: 3903: 3872: 3821: 3705: 3460: 3377: 3264: 3225: 3033: 2194: 2052: 1856:"Supplementary Material for "A genetic atlas of human admixture history"" 1334:"On the Baptism of the Serbs and Croats in the Time of Basil I (867–886)" 811: 722: 706: 569: 538: 463: 342: 272: 206: 186: 131:, the population of the Southeast Europe was composed of Ancient Greeks, 74: 4065: 504: 4030: 3978: 3963: 3564: 3557: 3372: 3367: 3304: 3249: 3186: 3133: 3016: 2930: 983:. Cambridge University Press. pp. 15, 24, 116, 118, 125, 286–287. 823: 819: 807: 799: 694: 620: 565: 561: 459: 389: 303: 295: 160: 111: 105:. Exhausted by several factors and reduced to the coastal parts of the 98: 2078: 1291: 653:
the Albanian-speaking majority assimilated the local Slavic settlers.
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Wright, David Curtis (1997). "The Hsiung-Nu-Hun Equation Revisited".
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and by the Byzantine Empire. In most parts of the former dioceses of
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The Slavs who settled in Southeast Europe comprised two groups: the
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in the mid-6th century and first decades of the 7th century in the
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Approximate location of South Slavic tribes, per V. V. Sedov 1995.
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The War of the Three Gods: Romans, Persians and the Rise of Islam
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influence and created an alliance with them against the Avar and
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forces, along the mountain ranges. They mostly were engaged in
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After the settlement of the Slavs, Southeast Europe turned to
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area. This caused more Sclaveni intrusions from the region of
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from 536 to around 660 CE and the series of wars between the
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Croatian lands in the Early Middle Ages (o. 550. – o. 1150.)
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Sloveni u ranom srednjem veku (Slavs in Early Middle Ages)
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The Roman Eastern Frontier and the Persian Wars AD 363-628
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share a very high number of common ancestors dated to the
4115:= generally considered synonym for early medieval Slovaks 1853: 1793: 1359: 1298: 1173: 1171: 360:(582–602) did not manage to stop the successful siege of 4109:= some of the Silesian tribes are Germanic, for example 1732:"The Geography of Recent Genetic Ancestry across Europe" 1644: 1605: 1593: 1488: 1347: 1209: 980:
Plague and the End of Antiquity: The Pandemic of 541-750
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subjects. There may have also been small communities of
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Encyclopedia of Slavic Languages and Linguistics Online
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Encyclopedia of Slavic Languages and Linguistics Online
1622: 1620: 1438: 1077: 958: 721:. They grew various fruits and vegetables, and learned 681:
For now archaeologically the earliest Slavic sites and
372:(617), and the destruction of various cities including 2859: 2025: 1581: 1510: 1371: 1238: 1236: 1168: 1067: 1065: 2817:
The Avars: A Steppe Empire in Central Europe, 567–822
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Medieval Syriac Historians' Perceptions of the Turks
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for military and agricultural purposes, and raising
325:(568-626). With the Byzantines preoccupied with the 1233: 1190: 1062: 2713: 2516:Olalde, Iñigo; Carrión, Pablo (December 7, 2023). 2352: 1089: 943: 73:. The backbone of the Avar Khaganate consisted of 1908: 16:Overview of Slavic migrations to Southeast Europe 4126: 2923:Early Slavic ethnic groups (7th–12th centuries) 2613:Dodgeon, Michael H.; Lieu, Samuel N. C. (1991). 2315: 2254: 1919: 1661: 1659: 976: 741:. Those living in hilly terrain mostly lived as 2570: 364:(580 to 582), though his generals triumphed at 2863:On the Origins of Proto-Croats and Proto-Serbs 2664:De Administrando Imperio: Volume II Commentary 2386:Kushniarevich, Alena; Kassian, Alexei (2020), 282:), with Procopius recording that the whole of 2907: 2591: 2515: 2511: 2509: 2417: 1656: 2813: 2229: 1980: 1796:"A Genetic Atlas of Human Admixture History" 1725: 1723: 1665: 583:Some Slavs in Thrace were also relocated to 2699:. Ann Arbor: University of Michigan Press. 2379: 2355:Archaeological and Anthropological Sciences 2032:G. Stamatoyannopoulos; et al. (2017). 1521:sfn error: no target: CITEREFKazanski2020 ( 1502:sfn error: no target: CITEREFKazanski2020 ( 1480:sfn error: no target: CITEREFKazanski2020 ( 1449:sfn error: no target: CITEREFKazanski2020 ( 1418:sfn error: no target: CITEREFKazanski2020 ( 1182:sfn error: no target: CITEREFKazanski2020 ( 2914: 2900: 2506: 2328:(in Hungarian). Napkút Kiadó. p. 58. 2248: 1974: 1788:A genetic atlas of human admixture history 1151:tribes, in the generic sense of 'nomads'". 1117:. Princeton University Press. p. 99. 2545: 2535: 2453: 2366: 2294: 2202: 2178: 2102: 2061: 2051: 2008: 1998: 1957: 1947: 1887: 1837: 1759: 1749: 1720: 1703: 1693: 1560: 1341:Studia Slavica et Balcanica Petropolitana 970: 609:Ecumenical Patriarchate of Constantinople 2870: 2125: 1729: 1535: 1516: 1497: 1475: 1444: 1413: 1328: 1201:sfn error: no target: CITEREFCurta2001 ( 1177: 1107: 810:". The 2015 IBD analysis found that the 752: 230: 44: 2633: 2255:Neparáczki, Endre; et al. (2019). 2242:National Academy of Sciences of Ukraine 2223: 1266: 1148:. University of Cambridge. p. 19. 1141: 1056: 298:intrusions from Antae territory in the 4127: 2785: 1920:A. Kushniarevich; et al. (2015). 1365: 1319:, p. 212–218, 382, 413, 444, 458. 793:. It is concluded to be caused by the 666:was composed of a mixture of Sclaveni 49:Slavic migrations to Southeast Europe. 2895: 2835:Sedov, Valentin Vasilyevich (2013) . 2834: 2748: 2346: 1650: 1638: 1626: 1611: 1599: 1587: 1575: 1463: 1432: 1401: 1389: 1377: 1353: 1316: 1304: 1254: 1242: 1227: 1215: 1196: 1162: 1095: 1083: 1071: 964: 952: 907:Outline of Slavic history and culture 518:and Moesia Secunda were inhabited by 418:Slavic settlement of the Eastern Alps 167:. After the destructive campaigns of 2739:Archaeology of the Slavic Migrations 2689: 1545:Antichnaya Drevnost's I Srednie Veka 181:, which resulted in the fall of the 4145:Wars involving the Byzantine Empire 4140:7th century in the Byzantine Empire 2845:]. Novi Sad: Akademska knjiga. 777:approximately 1,500 years ago with 13: 2182:European Journal of Human Genetics 2039:European Journal of Human Genetics 1981:P. M. Delser; et al. (2018). 1666:L. Kovačević; et al. (2014). 406:Praetorian prefecture of Illyricum 262:), coinciding with the end of the 14: 4171: 2400:10.1163/2589-6229_ESLO_COM_032367 1010: 897:Extreme weather events of 535–536 725:. They were actively engaged in 2860:Sotirović, Vladimir B. (2018). 2478: 2388:"Genetics and Slavic languages" 2172: 2119: 1776: 1529: 1322: 1260: 1135: 1101: 924: 351: 316: 277: 257: 2838:Славяне в раннем Средневековье 2390:, in Marc L. Greenberg (ed.), 2126:A. Zupan; et al. (2013). 1730:P. Ralph; et al. (2013). 997: 977:Lester K. Little, ed. (2007). 656: 412:In the late Roman province of 205:, pushed the migration of the 1: 2795:] (in Croatian). Zagreb: 937: 382:Siege of Constantinople (626) 122: 2820:. Cornell University Press. 2146:10.3109/03014460.2013.813584 1949:10.1371/journal.pone.0135820 1751:10.1371/journal.pbio.1001555 1695:10.1371/journal.pone.0105090 878:Anatolian Neolithic ancestry 587:, and later became known as 7: 2714:Kardaras, Georgios (2018). 890: 748: 640:fell under the rule of the 209:, who were also led by the 59:Late Antique Little Ice Age 10: 4176: 2564: 2537:10.1016/j.cell.2023.10.018 2368:10.1007/s12520-019-00996-0 2287:10.1038/s41598-019-53105-5 912:Pre-modern human migration 220: 216: 4092: 3991: 3987: 3929: 3880: 3871: 3837: 3681: 3543: 3483: 3342: 3333: 3273: 3240: 3150: 3141: 3132: 3065: 3058: 3007: 2998: 2938: 2929: 2761:10.1163/9789004425613_004 2104:10.1007/s10038-007-0125-6 2091:Journal of Human Genetics 1562:10.15826/adsv.2023.51.004 1005:Eurasian Studies Yearbook 713:, which they introduced, 664:Ipotesti–Candesti culture 368:(599). Subsequently the 57:. Another reason was the 2871:Živković, Tibor (2007). 2755:Brill. pp. 81–100. 2598:. Cembridge University. 2571:Crawford, Peter (2013). 2244:. pp. 219–226, 302. 2000:10.3389/fgene.2018.00551 1109:Beckwith, Christopher I. 917: 814:have lower proximity to 789:cluster among others in 548:were numerous tribes of 345:. The Byzantine Emperor 185:, Eastern Roman Emperor 2877:. Čigoja. p. 504. 2446:10.1126/science.abq0755 2133:Annals of Human Biology 1880:10.1126/science.1243518 1822:10.1126/science.1243518 1435:, p. 219, 221–222. 1404:, p. 218, 220–221. 1230:, p. 209–212, 215. 771:Serbo-Croatian language 323:almost century-long war 85:provinces south of the 79:siege of Constantinople 4160:History of the Balkans 2592:Curta, Florin (2001). 2325:Genetika és őstörténet 1142:Dickens, Mark (2004). 762:According to the 2013 759: 642:First Bulgarian Empire 380:, culminated with the 236: 175:, who were previously 129:great migration period 50: 4135:7th-century conflicts 2866:. Vilnius University. 2814:Pohl, Walter (2018). 2530:(25): P5472-5485.E9. 2230:O.M. Utevska (2017). 1987:Frontiers in Genetics 1553:Ural University Press 756: 611:began the process of 370:siege of Thessalonica 234: 48: 4155:South Slavic history 2799:. pp. 581–608. 2691:Fine, John V. A. Jr. 2673:University of London 2619:. Psychology Press. 2195:10.1038/ejhg.2014.50 2053:10.1038/ejhg.2017.18 646:diocese of Macedonia 546:Diocese of Macedonia 183:Western Roman Empire 71:Eastern Roman Empire 4150:Invasions of Greece 3889:Seven Slavic tribes 2279:2019NatSR...916569N 1940:2015PLoSO..1035820K 1872:2014Sci...343..747H 1814:2014Sci...343..747H 1686:2014PLoSO...9j5090K 1307:, p. 212, 215. 1036:2016NatGe...9..231B 870:study published in 861:Southeastern Poland 758:Caucasus component. 624:influenced both by 520:Seven Slavic tribes 446:was settled by the 203:population pressure 191:Plague of Justinian 77:. After the failed 55:Plague of Justinian 28:Southeastern Europe 3894:Southern Severians 2494:. December 7, 2023 2262:Scientific Reports 1653:, p. 227–228. 1614:, p. 225–226. 1602:, p. 225–227. 1478:, p. 7–8, 10. 1368:, p. 581–603. 1356:, p. 229–232. 1218:, p. 208–209. 1086:, p. 207–208. 967:, p. 206–207. 760: 495:were inhabited by 237: 51: 4122: 4121: 4088: 4087: 4084: 4083: 4015:Pannonian Dulebes 3867: 3866: 3677: 3676: 3236: 3235: 3128: 3127: 3124: 3123: 3054: 3053: 2943:Carpathian Croats 2884:978-86-7558-501-5 2852:978-86-6263-026-1 2806:978-953-150-942-8 2770:978-90-04-42561-3 2669:The Athlone Press 2651:Dimitri Obolensky 2577:. Pen and Sword. 2440:(6609): 940–951. 2335:978-963-263-855-3 1866:(6172): 747–751. 1808:(6172): 747–751. 1024:Nature Geoscience 990:978-0-521-84639-4 845:Y-DNA haplogroups 531:Diocese of Thrace 514:The provinces of 487:The provinces of 437:Pannonian Dulebes 117:Bulgar Khaganates 93:rivers, from the 32:Early Middle Ages 4167: 4006:Asia Minor Slavs 3989: 3988: 3922: 3881:Bulgarian tribes 3878: 3877: 3855: 3825: 3813: 3801: 3784: 3772: 3750: 3738: 3726: 3713:Bohemian Dulebes 3709: 3655: 3643: 3631: 3619: 3607: 3590: 3578: 3561: 3516: 3476: 3464: 3437: 3425: 3408: 3396: 3364: 3340: 3339: 3291: 3229: 3217: 3205: 3173: 3148: 3147: 3139: 3138: 3088: 3063: 3062: 3047: 3005: 3004: 2981: 2936: 2935: 2916: 2909: 2902: 2893: 2892: 2888: 2867: 2856: 2831: 2810: 2782: 2735:Kazanski, Michel 2731: 2710: 2686: 2659:Jenkins, Romilly 2630: 2609: 2588: 2559: 2558: 2556: 2554: 2549: 2539: 2513: 2504: 2503: 2501: 2499: 2482: 2476: 2475: 2457: 2431: 2421: 2415: 2414: 2408: 2406: 2383: 2377: 2376: 2370: 2350: 2344: 2343: 2319: 2313: 2312: 2298: 2252: 2246: 2245: 2227: 2221: 2220: 2206: 2176: 2170: 2169: 2123: 2117: 2116: 2106: 2082: 2076: 2075: 2065: 2055: 2029: 2023: 2022: 2012: 2002: 1978: 1972: 1971: 1961: 1951: 1917: 1906: 1905: 1891: 1851: 1841: 1791: 1780: 1774: 1773: 1763: 1753: 1727: 1718: 1717: 1707: 1697: 1663: 1654: 1648: 1642: 1636: 1630: 1624: 1615: 1609: 1603: 1597: 1591: 1590:, p. 88–89. 1585: 1579: 1573: 1567: 1566: 1564: 1537:Kazanski, Michel 1533: 1527: 1526: 1514: 1508: 1507: 1495: 1486: 1485: 1473: 1467: 1461: 1455: 1454: 1442: 1436: 1430: 1424: 1423: 1411: 1405: 1399: 1393: 1387: 1381: 1380:, p. 91–95. 1375: 1369: 1363: 1357: 1351: 1345: 1344: 1338: 1326: 1320: 1314: 1308: 1302: 1296: 1295: 1264: 1258: 1252: 1246: 1240: 1231: 1225: 1219: 1213: 1207: 1206: 1194: 1188: 1187: 1175: 1166: 1160: 1154: 1153: 1139: 1133: 1132: 1105: 1099: 1093: 1087: 1081: 1075: 1069: 1060: 1054: 1048: 1047: 1044:10.1038/ngeo2652 1014: 1008: 1001: 995: 994: 974: 968: 962: 956: 950: 931: 928: 775:migration period 727:animal husbandry 672:Penkovka culture 613:Christianization 599:and entered the 589:Asia Minor Slavs 442:The province of 374:Justiniana Prima 358:Balkan campaigns 355: 353: 320: 318: 281: 279: 261: 259: 147:, as well as of 4175: 4174: 4170: 4169: 4168: 4166: 4165: 4164: 4125: 4124: 4123: 4118: 4080: 4011:Pannonian Slavs 3983: 3925: 3916: 3863: 3849: 3833: 3819: 3807: 3795: 3778: 3766: 3744: 3732: 3720: 3703: 3691:Bohemian Croats 3673: 3649: 3637: 3625: 3613: 3601: 3584: 3572: 3555: 3539: 3510: 3479: 3470: 3458: 3431: 3419: 3402: 3390: 3358: 3335:Polabian tribes 3329: 3300:Silesian Croats 3285: 3275:Silesian tribes 3269: 3232: 3223: 3211: 3199: 3167: 3120: 3082: 3059:Northern tribes 3050: 3041: 2994: 2975: 2925: 2920: 2885: 2853: 2828: 2807: 2797:Matica hrvatska 2771: 2728: 2707: 2683: 2655:Steven Runciman 2647:Gyula Moravcsik 2639:Romilly Jenkins 2635:Francis Dvornik 2627: 2606: 2585: 2567: 2562: 2552: 2550: 2514: 2507: 2497: 2495: 2484: 2483: 2479: 2429: 2422: 2418: 2404: 2402: 2384: 2380: 2351: 2347: 2336: 2320: 2316: 2271:Nature Research 2253: 2249: 2228: 2224: 2177: 2173: 2124: 2120: 2083: 2079: 2030: 2026: 1979: 1975: 1918: 1909: 1852: 1792: 1782: 1781: 1777: 1728: 1721: 1664: 1657: 1649: 1645: 1637: 1633: 1625: 1618: 1610: 1606: 1598: 1594: 1586: 1582: 1574: 1570: 1534: 1530: 1520: 1515: 1511: 1501: 1500:, p. 8–12. 1496: 1489: 1479: 1474: 1470: 1462: 1458: 1448: 1443: 1439: 1431: 1427: 1417: 1412: 1408: 1400: 1396: 1388: 1384: 1376: 1372: 1364: 1360: 1352: 1348: 1336: 1330:Živković, Tibor 1327: 1323: 1315: 1311: 1303: 1299: 1268:Vryonis, Speros 1265: 1261: 1253: 1249: 1241: 1234: 1226: 1222: 1214: 1210: 1200: 1195: 1191: 1181: 1176: 1169: 1161: 1157: 1140: 1136: 1125: 1106: 1102: 1094: 1090: 1082: 1078: 1070: 1063: 1055: 1051: 1015: 1011: 1002: 998: 991: 975: 971: 963: 959: 951: 944: 940: 935: 934: 929: 925: 920: 893: 751: 703:Byzantine Greek 676:Michel Kazanski 659: 644:, while in the 529:In part of the 480:was settled by 433:Pannonian Slavs 350: 335:Sasanian Empire 315: 308:Pannonian Avars 276: 264:Vitalian revolt 256: 229: 221:Main articles: 219: 211:Pannonian Avars 125: 63:Sasanian Empire 17: 12: 11: 5: 4173: 4163: 4162: 4157: 4152: 4147: 4142: 4137: 4120: 4119: 4117: 4116: 4113: 4107: 4100: 4093: 4090: 4089: 4086: 4085: 4082: 4081: 4079: 4078: 4073: 4068: 4063: 4058: 4053: 4048: 4043: 4038: 4033: 4028: 4023: 4018: 4008: 4003: 3992: 3985: 3984: 3982: 3981: 3976: 3971: 3966: 3961: 3956: 3951: 3946: 3940: 3938: 3927: 3926: 3924: 3923: 3911: 3906: 3901: 3896: 3891: 3884: 3882: 3875: 3869: 3868: 3865: 3864: 3862: 3861: 3856: 3843: 3841: 3835: 3834: 3832: 3831: 3826: 3814: 3802: 3790: 3785: 3773: 3761: 3756: 3751: 3739: 3727: 3715: 3710: 3698: 3693: 3687: 3685: 3679: 3678: 3675: 3674: 3672: 3671: 3661: 3656: 3644: 3632: 3620: 3608: 3596: 3591: 3579: 3567: 3562: 3549: 3547: 3541: 3540: 3538: 3537: 3532: 3527: 3522: 3517: 3505: 3500: 3495: 3489: 3487: 3481: 3480: 3478: 3477: 3465: 3453: 3448: 3443: 3438: 3426: 3414: 3409: 3397: 3385: 3380: 3375: 3370: 3365: 3352: 3350: 3337: 3331: 3330: 3328: 3327: 3322: 3317: 3312: 3307: 3302: 3297: 3292: 3279: 3277: 3271: 3270: 3268: 3267: 3262: 3257: 3252: 3246: 3244: 3238: 3237: 3234: 3233: 3231: 3230: 3218: 3206: 3194: 3189: 3184: 3179: 3174: 3162: 3157: 3151: 3145: 3136: 3130: 3129: 3126: 3125: 3122: 3121: 3119: 3118: 3113: 3108: 3103: 3102: 3101: 3095: 3089: 3077: 3066: 3060: 3056: 3055: 3052: 3051: 3049: 3048: 3036: 3031: 3026: 3021: 3020: 3019: 3008: 3002: 2996: 2995: 2993: 2992: 2987: 2982: 2970: 2965: 2960: 2955: 2950: 2945: 2939: 2933: 2927: 2926: 2919: 2918: 2911: 2904: 2896: 2890: 2889: 2883: 2868: 2857: 2851: 2832: 2826: 2811: 2805: 2783: 2769: 2746: 2732: 2726: 2711: 2705: 2687: 2681: 2631: 2625: 2610: 2604: 2589: 2583: 2566: 2563: 2561: 2560: 2505: 2477: 2416: 2378: 2345: 2334: 2314: 2247: 2222: 2189:(1): 124–131, 2171: 2140:(6): 515–526. 2118: 2077: 2046:(5): 637–645. 2024: 1973: 1907: 1775: 1744:(5): e105090. 1719: 1680:(8): e105090. 1655: 1643: 1641:, p. 227. 1631: 1629:, p. 226. 1616: 1604: 1592: 1580: 1578:, p. 216. 1568: 1528: 1509: 1487: 1468: 1466:, p. 221. 1456: 1447:, p. 7–9. 1437: 1425: 1406: 1394: 1392:, p. 217. 1382: 1370: 1358: 1346: 1321: 1309: 1297: 1284:10.2307/147879 1278:(4): 378–390. 1259: 1257:, p. 229. 1247: 1232: 1220: 1208: 1189: 1167: 1165:, p. 208. 1155: 1134: 1123: 1100: 1088: 1076: 1074:, p. 207. 1061: 1059:, p. 113. 1049: 1030:(3): 231–236. 1009: 996: 989: 969: 957: 941: 939: 936: 933: 932: 922: 921: 919: 916: 915: 914: 909: 904: 899: 892: 889: 868:archaeogenetic 791:Eastern Europe 750: 747: 709:, cultivating 668:Prague-Korchak 658: 655: 593: 592: 581: 544:In all of the 542: 527: 516:Dacia Ripensis 512: 485: 475: 440: 425: 395:Johannes Koder 354: 582–602 333:wars with the 319: 565–574 292:Danubian Limes 280: 527–565 260: 518–527 218: 215: 199:Danubian Limes 169:Attila the Hun 149:Roman Imperial 124: 121: 67:Avar Khaganate 36:language shift 15: 9: 6: 4: 3: 2: 4172: 4161: 4158: 4156: 4153: 4151: 4148: 4146: 4143: 4141: 4138: 4136: 4133: 4132: 4130: 4114: 4112: 4108: 4105: 4102:= supposedly 4101: 4098: 4095: 4094: 4091: 4077: 4076:Praedenecenti 4074: 4072: 4069: 4067: 4064: 4062: 4059: 4057: 4054: 4052: 4049: 4047: 4044: 4042: 4039: 4037: 4034: 4032: 4029: 4027: 4024: 4022: 4019: 4016: 4012: 4009: 4007: 4004: 4001: 3997: 3994: 3993: 3990: 3986: 3980: 3977: 3975: 3972: 3970: 3967: 3965: 3962: 3960: 3957: 3955: 3952: 3950: 3947: 3945: 3942: 3941: 3939: 3937: 3933: 3928: 3920: 3915: 3912: 3910: 3907: 3905: 3902: 3900: 3897: 3895: 3892: 3890: 3887:Union of the 3886: 3885: 3883: 3879: 3876: 3874: 3870: 3860: 3857: 3853: 3848: 3845: 3844: 3842: 3840: 3839:Slovak tribes 3836: 3830: 3827: 3823: 3818: 3815: 3811: 3806: 3803: 3799: 3794: 3791: 3789: 3786: 3782: 3777: 3774: 3770: 3765: 3762: 3760: 3757: 3755: 3752: 3748: 3743: 3740: 3736: 3731: 3728: 3724: 3719: 3716: 3714: 3711: 3707: 3702: 3699: 3697: 3694: 3692: 3689: 3688: 3686: 3684: 3680: 3669: 3665: 3662: 3660: 3657: 3653: 3648: 3645: 3641: 3636: 3633: 3629: 3624: 3621: 3617: 3612: 3609: 3605: 3600: 3597: 3595: 3592: 3588: 3583: 3580: 3576: 3571: 3568: 3566: 3563: 3559: 3554: 3551: 3550: 3548: 3546: 3542: 3536: 3533: 3531: 3528: 3526: 3523: 3521: 3518: 3514: 3509: 3506: 3504: 3501: 3499: 3496: 3494: 3491: 3490: 3488: 3486: 3482: 3474: 3469: 3466: 3462: 3457: 3454: 3452: 3449: 3447: 3444: 3442: 3439: 3435: 3430: 3427: 3423: 3418: 3415: 3413: 3410: 3406: 3401: 3398: 3394: 3389: 3386: 3384: 3381: 3379: 3376: 3374: 3371: 3369: 3366: 3362: 3357: 3354: 3353: 3351: 3349: 3345: 3341: 3338: 3336: 3332: 3326: 3323: 3321: 3318: 3316: 3313: 3311: 3308: 3306: 3303: 3301: 3298: 3296: 3293: 3289: 3284: 3281: 3280: 3278: 3276: 3272: 3266: 3263: 3261: 3258: 3256: 3253: 3251: 3248: 3247: 3245: 3243: 3239: 3227: 3222: 3219: 3215: 3210: 3207: 3203: 3198: 3195: 3193: 3190: 3188: 3185: 3183: 3180: 3178: 3175: 3171: 3166: 3163: 3161: 3158: 3156: 3153: 3152: 3149: 3146: 3144: 3143:Polish tribes 3140: 3137: 3135: 3131: 3117: 3114: 3112: 3109: 3107: 3104: 3099: 3096: 3093: 3090: 3086: 3081: 3078: 3076: 3073: 3072: 3071: 3068: 3067: 3064: 3061: 3057: 3045: 3040: 3037: 3035: 3032: 3030: 3027: 3025: 3022: 3018: 3015: 3014: 3013: 3010: 3009: 3006: 3003: 3001: 2997: 2991: 2988: 2986: 2983: 2979: 2974: 2971: 2969: 2966: 2964: 2961: 2959: 2956: 2954: 2951: 2949: 2946: 2944: 2941: 2940: 2937: 2934: 2932: 2928: 2924: 2917: 2912: 2910: 2905: 2903: 2898: 2897: 2894: 2886: 2880: 2876: 2875: 2869: 2865: 2864: 2858: 2854: 2848: 2844: 2840: 2839: 2833: 2829: 2827:9781501729409 2823: 2819: 2818: 2812: 2808: 2802: 2798: 2794: 2790: 2784: 2780: 2776: 2772: 2766: 2762: 2758: 2754: 2753: 2747: 2744: 2740: 2736: 2733: 2729: 2727:9789004248380 2723: 2719: 2718: 2712: 2708: 2706:0-472-08149-7 2702: 2698: 2697: 2692: 2688: 2684: 2682:9780884020219 2678: 2674: 2670: 2666: 2665: 2660: 2656: 2652: 2648: 2644: 2643:Bernard Lewis 2640: 2636: 2632: 2628: 2626:9780415465304 2622: 2618: 2617: 2611: 2607: 2605:9781139428880 2601: 2597: 2596: 2590: 2586: 2584:9781848846128 2580: 2576: 2575: 2569: 2568: 2548: 2543: 2538: 2533: 2529: 2525: 2524: 2519: 2512: 2510: 2493: 2492: 2487: 2481: 2473: 2469: 2465: 2461: 2456: 2451: 2447: 2443: 2439: 2435: 2428: 2420: 2413: 2401: 2397: 2393: 2389: 2382: 2375: 2369: 2364: 2360: 2356: 2349: 2342: 2337: 2331: 2327: 2326: 2318: 2311: 2306: 2302: 2297: 2292: 2288: 2284: 2280: 2276: 2272: 2268: 2264: 2263: 2258: 2251: 2243: 2239: 2235: 2234: 2226: 2219: 2214: 2210: 2205: 2200: 2196: 2192: 2188: 2184: 2183: 2175: 2168: 2163: 2159: 2155: 2151: 2147: 2143: 2139: 2135: 2134: 2129: 2122: 2114: 2110: 2105: 2100: 2097:(5): 406–14. 2096: 2092: 2088: 2081: 2073: 2069: 2064: 2059: 2054: 2049: 2045: 2041: 2040: 2035: 2028: 2020: 2016: 2011: 2006: 2001: 1996: 1992: 1988: 1984: 1977: 1969: 1965: 1960: 1955: 1950: 1945: 1941: 1937: 1933: 1929: 1928: 1923: 1916: 1914: 1912: 1904: 1899: 1895: 1890: 1885: 1881: 1877: 1873: 1869: 1865: 1861: 1857: 1849: 1845: 1840: 1835: 1831: 1827: 1823: 1819: 1815: 1811: 1807: 1803: 1802: 1797: 1789: 1785: 1779: 1771: 1767: 1762: 1757: 1752: 1747: 1743: 1739: 1738: 1733: 1726: 1724: 1715: 1711: 1706: 1701: 1696: 1691: 1687: 1683: 1679: 1675: 1674: 1669: 1662: 1660: 1652: 1647: 1640: 1635: 1628: 1623: 1621: 1613: 1608: 1601: 1596: 1589: 1584: 1577: 1572: 1563: 1558: 1554: 1550: 1546: 1542: 1538: 1532: 1524: 1519:, p. 13. 1518: 1517:Kazanski 2020 1513: 1505: 1499: 1498:Kazanski 2020 1494: 1492: 1483: 1477: 1476:Kazanski 2020 1472: 1465: 1460: 1452: 1446: 1445:Kazanski 2020 1441: 1434: 1429: 1421: 1415: 1414:Kazanski 2020 1410: 1403: 1398: 1391: 1386: 1379: 1374: 1367: 1362: 1355: 1350: 1342: 1335: 1331: 1325: 1318: 1313: 1306: 1301: 1293: 1289: 1285: 1281: 1277: 1273: 1269: 1263: 1256: 1251: 1245:, p. 83. 1244: 1239: 1237: 1229: 1224: 1217: 1212: 1204: 1199:, p. 86. 1198: 1193: 1185: 1180:, p. 12. 1179: 1178:Kazanski 2020 1174: 1172: 1164: 1159: 1152: 1147: 1146: 1138: 1131: 1126: 1124:9781400829941 1120: 1116: 1115: 1110: 1104: 1098:, p. 82. 1097: 1092: 1085: 1080: 1073: 1068: 1066: 1058: 1053: 1045: 1041: 1037: 1033: 1029: 1025: 1021: 1013: 1007:. 69: 77–112. 1006: 1000: 992: 986: 982: 981: 973: 966: 961: 955:, p. 84. 954: 949: 947: 942: 927: 923: 913: 910: 908: 905: 903: 900: 898: 895: 894: 888: 886: 885: 879: 875: 874: 869: 864: 862: 858: 853: 849: 846: 842: 836: 833: 829: 825: 821: 817: 813: 809: 805: 801: 796: 792: 788: 784: 780: 776: 772: 768: 765: 755: 746: 744: 740: 736: 732: 728: 724: 720: 716: 712: 708: 704: 698: 696: 692: 688: 684: 679: 677: 673: 669: 665: 654: 652: 647: 643: 639: 635: 631: 627: 623: 622: 616: 614: 610: 606: 602: 598: 590: 586: 582: 579: 575: 571: 567: 563: 559: 555: 551: 547: 543: 540: 536: 532: 528: 525: 521: 517: 513: 510: 509:Praedenecenti 506: 502: 498: 494: 490: 486: 483: 479: 476: 473: 469: 465: 461: 457: 453: 449: 445: 441: 438: 434: 430: 426: 423: 419: 415: 411: 410: 409: 407: 402: 400: 396: 392: 391: 385: 383: 379: 375: 371: 367: 363: 359: 348: 344: 340: 336: 332: 328: 324: 313: 309: 305: 301: 297: 293: 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Retrieved 2527: 2521: 2496:. Retrieved 2491:ScienceDaily 2489: 2480: 2437: 2433: 2419: 2410: 2403:, retrieved 2391: 2381: 2372: 2358: 2354: 2348: 2339: 2324: 2317: 2308: 2266: 2260: 2250: 2237: 2232: 2225: 2216: 2186: 2180: 2174: 2165: 2137: 2131: 2121: 2094: 2090: 2080: 2043: 2037: 2027: 1990: 1986: 1976: 1931: 1925: 1901: 1863: 1859: 1805: 1799: 1787: 1778: 1741: 1737:PLOS Biology 1735: 1677: 1671: 1646: 1634: 1607: 1595: 1583: 1571: 1548: 1544: 1531: 1512: 1471: 1459: 1440: 1428: 1416:, p. 8. 1409: 1397: 1385: 1373: 1361: 1349: 1340: 1324: 1312: 1300: 1275: 1271: 1262: 1250: 1223: 1211: 1192: 1158: 1149: 1144: 1137: 1128: 1113: 1103: 1091: 1079: 1057:Dvornik 1962 1052: 1027: 1023: 1012: 1004: 999: 979: 972: 960: 926: 882: 871: 865: 837: 761: 711:proso millet 699: 691:Lake Balaton 680: 660: 637: 619: 617: 605:Roman Church 594: 550:Drougoubitai 478:Praevalitana 448:White Croats 422:Carantanians 403: 388: 386: 300:Lower Danube 266:of 511–518. 238: 176: 126: 110: 97:towards the 69:against the 52: 38:to and from 18: 4104:Finno-Ugric 4071:Branichevci 4036:Zachlumians 3969:Belegezites 3917: [ 3904:Strymonites 3873:South Slavs 3850: [ 3820: [ 3808: [ 3796: [ 3779: [ 3767: [ 3745: [ 3733: [ 3721: [ 3704: [ 3668:White Serbs 3650: [ 3638: [ 3626: [ 3614: [ 3602: [ 3585: [ 3573: [ 3556: [ 3511: [ 3471: [ 3459: [ 3432: [ 3420: [ 3403: [ 3391: [ 3378:Tollensians 3359: [ 3286: [ 3265:Slovincians 3242:Pomeranians 3224: [ 3212: [ 3200: [ 3168: [ 3083: [ 3042: [ 3034:Dregoviches 2976: [ 2553:December 8, 2498:December 8, 2405:10 December 1934:(9): 6, 7. 1366:Vedriš 2015 1343:(1): 33–53. 812:South Slavs 723:viticulture 717:, but also 707:agriculture 657:Archaeology 570:Belegezites 539:Strymonites 468:Travunijans 464:Zachlumians 420:(including 343:Peloponnese 273:Justinian I 207:Early Slavs 187:Justinian I 127:Before the 4129:Categories 4041:Travunians 4031:Narentines 3979:Rhynchinoi 3964:Baiounitai 3817:Lupiglians 3805:Domazhlici 3776:Sedlichans 3730:Litomerici 3635:Neletiches 3565:Glomatians 3400:Neletyches 3388:Morzyczans 3373:Kessinians 3368:Circipania 3305:Dadosesani 3250:Kashubians 3187:Sieradzans 3134:West Slavs 3017:Volhynians 2931:East Slavs 2667:. London: 1651:Sedov 2013 1639:Sedov 2013 1627:Sedov 2013 1612:Sedov 2013 1600:Sedov 2013 1588:Koder 2020 1576:Sedov 2013 1464:Sedov 2013 1433:Sedov 2013 1402:Sedov 2013 1390:Sedov 2013 1378:Koder 2020 1354:Sedov 2013 1317:Sedov 2013 1305:Sedov 2013 1255:Sedov 2013 1243:Koder 2020 1228:Sedov 2013 1216:Sedov 2013 1197:Curta 2001 1163:Sedov 2013 1096:Koder 2020 1084:Sedov 2013 1072:Sedov 2013 965:Sedov 2013 953:Koder 2020 938:References 824:West Slavs 820:East Slavs 818:than with 808:Hungarians 800:Bulgarians 695:Yugoslavia 670:and Antae 638:Sklavinias 621:Sklavinias 566:Baiounitai 562:Rhynchinoi 501:Braničevci 460:Narentines 390:demokratia 366:Viminacium 304:Podunavlje 161:Langobards 145:Hellenized 123:Background 112:Sklavinias 101:up to the 4066:Timochans 4046:Kanalites 4026:Guduscani 3959:Sagudates 3936:Macedonia 3829:Znetalici 3754:Moravians 3611:Zhirmunts 3582:Lusatians 3570:Koledices 3545:Lusatians 3535:Polabians 3520:Smeldingi 3493:Bethenici 3485:Obotrites 3417:Redariers 3325:Silesians 3310:Golensizi 3283:Bezunzans 3260:Wolinians 3197:Kujawians 3192:Vistulans 3177:Masovians 3165:Lubuszans 3075:Polochans 3039:Narevyans 3024:Drevlians 2973:Don Slavs 2953:Severians 2948:Radimichs 2779:218997565 2737:(2020). 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Brill. 2693:(1991) . 2472:251844202 2394:, Brill, 2273:: 16569. 2269:(16569). 1830:0036-8075 1555:: 76–89. 1130:identity. 832:admixture 804:Romanians 764:autosomal 743:shepherds 683:artifacts 601:Dark Ages 558:Sagudates 524:Severians 482:Diocleans 472:Kanalites 452:Guduscani 431:were the 401:origin. 399:toponymic 356:) in his 312:Justin II 296:barbarian 284:Illyricum 268:Procopius 195:Pannonian 178:foederati 157:Bastarnae 141:Romanized 137:Thracians 133:Illyrians 103:Black Sea 83:Byzantine 24:migrating 4061:Moravens 3954:Melingoi 3949:Ezeritai 3914:Milcovci 3899:Smolyani 3847:Nitrians 3764:Pshovans 3759:Merehani 3647:Nizhices 3623:Zhitices 3553:Khutices 3441:Sprevane 3295:Bobrzans 3255:Prissani 3221:Thafnezi 3209:Wiercans 3160:Lendians 3106:Slovenes 3098:Smolensk 3070:Krivichs 2985:Zeriuani 2968:Vyatichi 2958:Tivertsi 2657:(1962). 2547:10752003 2464:36007020 2455:10019558 2412:lineages 2305:31719606 2213:24667786 2162:34621779 2154:23879710 2113:17364156 2072:28272534 2019:30510563 1968:26332464 1927:PLOS One 1898:24531965 1848:24531965 1770:23667324 1714:25148043 1673:PLOS One 1539:(2023). 1332:(2013). 1111:(2009). 891:See also 787:Bulgaria 749:Genetics 729:, using 597:paganism 585:Anatolia 578:Ezeritai 574:Melingoi 535:Smolyani 505:Timočani 497:Merehani 493:Dardania 444:Dalmatia 429:Pannonia 416:was the 253:Justin I 245:Sclaveni 243:and the 227:Sclaveni 171:and the 95:Adriatic 65:and the 4111:Silings 3974:Berziti 3909:Moratsi 3859:Slovaks 3788:Zlicans 3742:Luchans 3701:Dechans 3659:Nishans 3599:Suslowi 3594:Milceni 3530:Warnabi 3498:Drevani 3468:Zemcici 3456:Zamcici 3446:Hevelli 3429:Rechans 3356:Brizans 3320:Selpoli 3315:Opolans 3155:Goplans 3080:Pskov's 3029:Polans 3012:Buzhans 3000:Dulebes 2745:. BRILL 2661:(ed.). 2565:Sources 2434:Science 2310:groups. 2296:6851379 2275:Bibcode 2204:4266736 2063:5437898 2010:6252347 1993:: 551. 1959:4558026 1936:Bibcode 1889:4209567 1868:Bibcode 1860:Science 1839:4209567 1810:Bibcode 1801:Science 1761:3646727 1705:4141785 1682:Bibcode 1032:Bibcode 902:Bulgars 873:Science 866:A 2022 857:Ukraine 841:Dnieper 783:Romania 651:Albania 634:Thracia 626:Francia 554:Berziti 414:Noricum 362:Sirmium 347:Maurice 339:Balkans 331:602–628 327:572–591 217:History 107:Balkans 4106:tribes 4021:Croats 3932:Greece 3718:Lemuzi 3696:Czechs 3508:Reregs 3503:Linons 3451:Ukrani 3412:Rujani 3383:Doxani 3348:Lutici 3344:Veleti 3182:Polans 3116:Muroma 2963:Ulichs 2881:  2849:  2824:  2803:  2777:  2767:  2724:  2703:  2679:  2623:  2602:  2581:  2544:  2470:  2462:  2452:  2332:  2303:  2293:  2211:  2201:  2160:  2152:  2111:  2070:  2060:  2017:  2007:  1966:  1956:  1896:  1886:  1846:  1836:  1828:  1768:  1758:  1712:  1702:  1292:147879 1290:  1121:  987:  828:Gagauz 816:Greeks 806:, and 795:Hunnic 779:Poland 731:horses 687:Gepids 489:Moesia 470:, and 435:(with 378:Salona 288:Thrace 153:Heruli 99:Aegean 91:Danube 40:Slavic 22:began 4097:Notes 4056:Serbs 3921:] 3854:] 3824:] 3812:] 3800:] 3793:Hbans 3783:] 3771:] 3749:] 3737:] 3725:] 3708:] 3664:Sorbs 3654:] 3642:] 3630:] 3618:] 3606:] 3589:] 3577:] 3560:] 3525:Wagri 3515:] 3475:] 3463:] 3436:] 3424:] 3407:] 3395:] 3363:] 3290:] 3228:] 3216:] 3204:] 3172:] 3111:Merya 3087:] 3046:] 2980:] 2841:[ 2791:[ 2775:S2CID 2468:S2CID 2430:(PDF) 2361:(1), 2236:[ 2158:S2CID 1337:(PDF) 1288:JSTOR 918:Notes 739:goats 715:wheat 630:Dacia 533:were 456:Serbs 450:(and 241:Antae 223:Antae 173:Goths 165:Sciri 20:Slavs 3934:and 3346:and 3092:Tver 2879:ISBN 2847:ISBN 2822:ISBN 2801:ISBN 2765:ISBN 2722:ISBN 2701:ISBN 2677:ISBN 2621:ISBN 2600:ISBN 2579:ISBN 2555:2023 2523:Cell 2500:2023 2460:PMID 2407:2020 2330:ISBN 2301:PMID 2209:PMID 2150:PMID 2109:PMID 2068:PMID 2015:PMID 1964:PMID 1894:PMID 1844:PMID 1826:ISSN 1766:PMID 1710:PMID 1523:help 1504:help 1482:help 1451:help 1420:help 1203:help 1184:help 1119:ISBN 985:ISBN 884:Cell 859:and 850:and 822:and 781:and 737:and 735:oxen 719:flax 636:the 632:and 607:and 576:and 537:and 522:and 507:and 491:and 376:and 341:and 329:and 286:and 249:Huns 225:and 197:and 163:and 143:and 135:and 89:and 87:Sava 3930:in 2757:doi 2741:". 2542:PMC 2532:doi 2528:186 2450:PMC 2442:doi 2438:377 2396:doi 2363:doi 2291:PMC 2283:doi 2199:PMC 2191:doi 2142:doi 2099:doi 2058:PMC 2048:doi 2005:PMC 1995:doi 1954:PMC 1944:doi 1884:PMC 1876:doi 1864:343 1834:PMC 1818:doi 1806:343 1756:PMC 1746:doi 1700:PMC 1690:doi 1557:doi 1280:doi 1040:doi 863:. 848:R1a 767:IBD 615:. 454:), 427:In 408:: 26:to 4131:: 3919:ru 3852:pl 3822:pl 3810:ru 3798:uk 3781:pl 3769:cs 3747:cs 3735:pl 3723:cs 3706:pl 3652:pl 3640:pl 3628:pl 3616:pl 3604:pl 3587:pl 3575:pl 3558:pl 3513:pl 3473:pl 3461:pl 3434:pl 3422:pl 3405:pl 3393:pl 3361:pl 3288:de 3226:pl 3214:pl 3202:pl 3170:pl 3100:'s 3094:'s 3085:ru 3044:ru 2978:ru 2773:. 2763:. 2675:. 2671:, 2653:; 2649:; 2645:; 2641:; 2637:; 2540:. 2526:. 2520:. 2508:^ 2488:. 2466:. 2458:. 2448:. 2436:. 2432:. 2409:, 2371:, 2359:12 2357:, 2338:. 2307:. 2299:. 2289:. 2281:. 2265:. 2259:. 2215:, 2207:, 2197:, 2187:23 2185:, 2164:. 2156:. 2148:. 2138:40 2136:. 2130:. 2107:. 2095:52 2093:. 2089:. 2066:. 2056:. 2044:25 2042:. 2036:. 2013:. 2003:. 1989:. 1985:. 1962:. 1952:. 1942:. 1932:10 1930:. 1924:. 1910:^ 1900:. 1892:. 1882:. 1874:. 1862:. 1858:. 1842:. 1832:. 1824:. 1816:. 1804:. 1798:. 1786:. 1764:. 1754:. 1742:11 1740:. 1734:. 1722:^ 1708:. 1698:. 1688:. 1676:. 1670:. 1658:^ 1619:^ 1551:. 1549:23 1547:. 1490:^ 1339:. 1286:. 1276:50 1274:. 1235:^ 1170:^ 1127:. 1064:^ 1038:. 1026:. 1022:. 945:^ 852:I2 802:, 572:, 568:, 564:, 560:, 556:, 552:, 503:, 499:, 466:, 462:, 458:, 439:). 424:). 384:. 352:r. 317:r. 278:r. 258:r. 213:. 159:, 155:, 119:. 42:. 4017:) 4013:( 4002:) 3998:( 3670:) 3666:( 2915:e 2908:t 2901:v 2887:. 2855:. 2830:. 2809:. 2781:. 2759:: 2730:. 2709:. 2685:. 2629:. 2608:. 2587:. 2557:. 2534:: 2502:. 2474:. 2444:: 2398:: 2365:: 2285:: 2277:: 2267:9 2193:: 2144:: 2115:. 2101:: 2074:. 2050:: 2021:. 1997:: 1991:9 1970:. 1946:: 1938:: 1878:: 1870:: 1850:. 1820:: 1812:: 1790:. 1772:. 1748:: 1716:. 1692:: 1684:: 1678:9 1565:. 1559:: 1525:) 1506:) 1484:) 1453:) 1422:) 1294:. 1282:: 1205:) 1186:) 1046:. 1042:: 1034:: 1028:9 993:. 785:- 591:. 580:. 541:. 526:. 511:. 484:. 474:. 349:( 314:( 275:( 255:(

Index

Slavs
migrating
Southeastern Europe
Early Middle Ages
language shift
Slavic

Plague of Justinian
Late Antique Little Ice Age
Sasanian Empire
Avar Khaganate
Eastern Roman Empire
Slavic tribes
siege of Constantinople
Byzantine
Sava
Danube
Adriatic
Aegean
Black Sea
Balkans
Sklavinias
Bulgar Khaganates
great migration period
Illyrians
Thracians
Romanized
Hellenized
Roman Imperial
Heruli

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