352:
80:
202:
415:
55:
216:
31:
382:
513:. The presence of this first known maternity colony in the country surprised the investigators: the bats roosted under the roof and in the various rooms instead of trees or abandoned mines: It might therefore be of interest to search for maternity colonies of the species in North Africa, not only in wooded areas and underground sites but also in abandoned opened buildings.
509:, the lesser horseshoe bat, though rarer than other bat species, is present in the humid climatic zone (i.e. the most northern part of the country). It had been mostly reported as isolated individuals or small groups of hibernating bats until 2011. Then, a maternity colony was found in the abandoned HĂ´tel des ChĂŞnes, South of
264:
The lesser horseshoe bat is one of the world's smallest bats, weighing only 5 to 9 grams (0.18 to 0.32 oz), with a wingspan of 192–254 millimetres (7.6–10.0 in) and a body length of 35–45 millimetres (1.4–1.8 in). It has strong feet that it uses to grasp rocks and branches, and can see
359:
The lesser horseshoe bat lives in warmer regions in foothills and highland, in particular wooded areas or areas of limestone, where it roosts in caves. In summer its range has been recorded up to 1160 m above sea level, and up to 2000 m in the winter, with the highest known nursery roost at 950 m.
332:
Females become sexually mature within their first year, with a study in
Czechoslovakia showing that about 15% of females give birth at one year of age. Animals have been seen to chase each other as a preliminary to mating, which is done with the male hanging himself behind and over the female.
497:. Roosts containing up to 50 individuals have been found in caves such as the Cova de Ca Na Rea (Ibiza), though most groups tend to be smaller. In the Balearics, the lesser horseshoe bat travels only short distances, usually around 2 km, and is also the most common bat species on Ibiza.
343:, but there is no direct mixing with other species. The move to maternity colonies occurs from April onwards, and the colony size can greatly vary from very few individuals to hundreds of them. Approximately two-thirds of females in a nursery roost give birth between mid-June and mid-July.
387:
383:
422:
A large breeding colony of lesser horseshoe bats populates the smallest SSSI (Site of
Special Scientific Interest) in the UK, a 7m barn in Gloucestershire, England. The species is threatened by a number of factors, including the disturbance or destruction of roosts, changes in
385:
324:
Lesser horseshoe bats mate in the autumn. Females give birth to one pup, normally between mid-June and the beginning of July. Pups weigh around 1.8 grams at birth, opening their eyes after around 10 days and becoming independent at six to seven weeks of age. The bats
284:
When hunting they are quick and agile, often flying within five metres of the ground while avoiding contact with bushes and shrubs. The lesser horseshoe bat eats small insects, most of which are gleaned from stones and branches. Their favorite types of prey include
372:
is widely distributed in the
Mediterranean area, occurring in North Africa and on most larger islands to Asia Minor and around the Black Sea. In Asia, it can be found in Kashmir, the Near East, Iran, Iraq and the Arabian Peninsula, as well as part of East Africa.
521:
The frequencies used by this bat species for echolocation lie between 93 and 111 kHz, have most energy at 110 kHz and have an average duration of 31.7 ms. Due to the frequency of their echolocation calls there are overlaps with those of the
280:
The base of its fur, which is soft and fluffy, is light grey in colour, with dorsal side fur smoky brown and the ventral side grey, with the exception of juvenile bats which are entirely dark grey. Ears and wing membranes are a light greyish brown.
386:
543:
553:
In order to highlight the importance of protecting this species at the
European scale, this species was selected as bat species of the Year 2018–2019 by the pan-European NGO BatLife Europe.
384:
2066:
865:"Variability in echolocation call design of 26 Swiss bat species: consequences, limits and options for automated field identification with a synergetic pattern recognition approach"
360:
The species is sedentary, with the average movement between summer and winter roosts between 5 and 10 kilometers, although the longest recorded distance is 153 kilometers.
713:
Calderoni, Gregorio; Krzemińska, Sandra; Górska, Monika; Marialuce, Paladini; Ren, Minrui; Apoznański, Grzegorz; Blum, Astrid; Bamberger, Axel; Kokurewicz, Tomasz (2024).
439:'s Hibernation Survey shows numbers in the UK are increasing significantly – by an average of 4.5% yearly between 1999 and 2012, a 77.2% total increase during that period.
368:
The species can be found in western
Ireland and south-west Britain, with some larger populations in parts of Germany and the Bavarian foothills of the Alps.
796:
McAney, K., O'Mahony, C., Kellecher, C., Taylor, A. and
Biggane, S. (2013). "The Lesser Horseshoe Bat in Ireland: surveys by The Vincent Wildlife Trust",
449:
The lesser horseshoe bat has been recorded in the counties Galway, South Mayo, Clare, Limerick, Cork and Kerry. The population is estimated at 13,000.
440:
715:"First characterisation and conservation needs of a large maternity colony of lesser horseshoe bats (Rhinolophus hipposideros) in Aljezur, Portugal"
849:
673:
2170:
956:
2248:
2209:
2421:
2022:
2367:
2144:
2380:
2183:
996:
812:"Acoustic identification of twelve species of echolocating bat by discriminant function analysis and artificial neural networks"
2227:
965:
940:
687:
Bontadina, Fabio; Arlettaz, Raphaël; Fankhauser, Toni; Lutz, Miriam; Mühlethaler, Erich; Theiler, Alex; Zingg, Peter (2000).
2385:
2188:
748:"Importance of night roosts for bat conservation: roosting behaviour of the lesser horseshoe bat Rhinolophus hipposideros"
550:
convention. Several national legislation are also protecting this species and its habitats in many countries and regions.
2461:
2446:
2354:
2105:
1995:
628:
2123:
2393:
2300:
2253:
2441:
2136:
1433:
2436:
1919:
1375:
689:"The lesser horseshoe bat Rhinolophus hipposideros in Switzerland: Present status and research recommendations"
2235:
1844:
1774:
1597:
1345:
1255:
1355:
646:
1814:
1205:
407:
The UK distribution of the lesser horseshoe bat can be found on the
National Biodiversity Network website
191:
2222:
2040:
1824:
1543:
1416:
1315:
1215:
1118:
989:
523:
2045:
79:
1908:
1854:
1834:
1724:
1657:
1647:
1493:
1275:
1135:
1088:
436:
2398:
2305:
1925:
1888:
1882:
1864:
1764:
1667:
1637:
1581:
1570:
1453:
1325:
1285:
1265:
527:
490:
339:(also known as nursery colonies or nursery roosts) are often shared with other species, such as the
1704:
1617:
1483:
1463:
1175:
1165:
1129:
594:
2456:
2426:
2009:
1734:
1678:
1523:
1503:
1295:
1078:
1068:
400:
340:
2292:
2451:
2175:
1957:
1804:
1607:
1427:
1305:
1195:
982:
648:
649:"Bat fauna of Tunisia: Review of records and new records, morphometrics and echolocation data"
2431:
2287:
1898:
1560:
1554:
1533:
1443:
1365:
1335:
1235:
1225:
1155:
1145:
1108:
1098:
902:
864:
843:
667:
245:
174:
2279:
2201:
2079:
2004:
1794:
1744:
1714:
1627:
1587:
1513:
1473:
1410:
1393:
1245:
2071:
316:
showed no obvious morphological differences between North
African and European specimens.
8:
1754:
1684:
974:
408:
270:
44:
329:
during the winter months in dark caves, mines, old buildings, and sometimes in cellars.
1784:
1694:
1185:
884:
539:
74:
2341:
2274:
2053:
1935:
1875:
1386:
936:
831:
647:
Sebastien J. Puechmaille, Wassim M. Hizem, Benjamin
Allegrini, Awatef Abiadh (2012).
624:
571:
432:
888:
811:
776:
2346:
2332:
2058:
876:
823:
759:
726:
589:
470:
336:
266:
962:
731:
714:
2261:
969:
2240:
2214:
351:
2131:
2118:
1042:
396:
265:
well in spite of its small eyes. Like most bats, lesser horseshoe bats live in
688:
2415:
2196:
1980:
1059:
1006:
931:
Schober, Wilfried; Eckard
Grimmberger (1989). Dr. Robert E. Stebbings (ed.).
580:
249:
151:
141:
64:
59:
2359:
827:
510:
201:
2110:
835:
424:
880:
787:
Stuart Bell and Donald McGillivray, Environmental Law, 7th ed 2008, p. 690
414:
309:
is 3 years, although some individuals have been known to live far longer.
2157:
1989:
428:
326:
238:
2032:
2372:
2266:
2149:
494:
392:
274:
215:
2162:
2092:
763:
294:
253:
91:
1951:
431:, which reduce prey availability) and the loss of suitable foraging
411:, but they are mostly found in Wales and the Southwest of England.
2326:
2097:
1974:
1036:
1024:
747:
547:
478:
290:
111:
712:
252:, the species gets its name from its distinctive horseshoe-shaped
506:
482:
313:
30:
2084:
2017:
1030:
1018:
686:
458:
302:
234:
121:
101:
863:
Obrist, Martin K.; Boesch, Ruedi; FlĂĽckiger, Peter F. (2004).
486:
298:
1004:
538:
This species is protected in the European Union under the
1048:
930:
809:
286:
241:
131:
777:
Rhinolophus hipposideros – Science for Nature Foundation
862:
623:. Palma de Mallorca: Editorial Moll. pp. 90–91.
619:
Antoni Alcover, J., ed. (1988). "MamĂfers Actuals".
618:
418:A torpid lesser horseshoe bat, Gloucestershire, UK
2413:
391:Conservation of the Lesser horseshoe bat in the
595:10.2305/IUCN.UK.2016-2.RLTS.T19518A21972794.en
990:
935:(1st ed.). UK: Hamlyn Publishing Group.
848:: CS1 maint: multiple names: authors list (
672:: CS1 maint: multiple names: authors list (
277:from specialized round pads in their mouth.
745:
355:Lesser horseshoe bat in cave during winter.
997:
983:
200:
53:
29:
730:
593:
546:and is specifically targeted by the UNEP-
563:
477:lives on all three of the main islands:
427:practices (such as the increased use of
413:
380:
350:
214:
259:
2414:
614:
612:
569:
1956:
1955:
978:
933:A Guide to Bats of Britain and Europe
746:Knight, Tessa; Jones, Gareth (2009).
542:. This species is also listed in the
2137:faa20a96-599d-487e-bde8-5a2201ae86e3
642:
640:
457:The lesser horseshoe bat is rare in
244:, related to its larger cousin, the
2422:IUCN Red List least concern species
609:
581:IUCN Red List of Threatened Species
464:
13:
963:Woodland Management For Bats Guide
924:
810:Parsons, S. and Jones, G. (2000).
14:
2473:
950:
637:
489:, as well as the smaller islands
376:
78:
1434:Philippine forest horseshoe bat
895:
856:
803:
719:The European Zoological Journal
533:
516:
363:
790:
781:
770:
739:
706:
680:
312:Detailed observations done in
219:Lesser horseshoe bat in flight
1:
1845:Little Nepalese horseshoe bat
1598:Formosan lesser horseshoe bat
1346:Formosan woolly horseshoe bat
1256:Little Japanese horseshoe bat
732:10.1080/24750263.2024.2312930
556:
1815:Chinese rufous horseshoe bat
903:"BatLife Europe Bat of Year"
7:
1755:Lesser woolly horseshoe bat
1376:Hildebrandt's horseshoe bat
1316:Mediterranean horseshoe bat
1119:Lesser woolly horseshoe bat
752:Endangered Species Research
621:els MamĂfers de les Balears
524:Mediterranean horseshoe bat
209:Lesser horseshoe bat range
10:
2478:
2462:Habitats Directive species
1909:Yellow-faced horseshoe bat
1855:Small rufous horseshoe bat
1835:Lesser brown horseshoe bat
1775:Shortridge's horseshoe bat
1725:Large rufous horseshoe bat
1089:Intermediate horseshoe bat
500:
444:
437:the Bat Conservation Trust
346:
305:. The average lifespan of
2447:Mammals described in 1800
2316:
1964:
1668:Large-eared horseshoe bat
1326:Broad-eared horseshoe bat
1206:Chiewkwee's horseshoe bat
1057:
1013:
1005:Extant species of family
452:
319:
208:
199:
180:
173:
75:Scientific classification
73:
51:
42:
37:
28:
23:
2318:Vespertilio hipposideros
2072:rhinolophus-hipposideros
2023:rhinolophus-hipposideros
2010:Rhinolophus_hipposideros
1996:Rhinolophus hipposideros
1966:Rhinolophus hipposideros
1825:Smithers's horseshoe bat
1705:Peninsular horseshoe bat
1618:Mozambican horseshoe bat
1544:Marshall's horseshoe bat
1484:Mount Mabu horseshoe bat
1417:Imaizumi's horseshoe bat
1216:Geoffroy's horseshoe bat
574:Rhinolophus hipposideros
475:Rhinolophus hipposideros
370:Rhinolophus hipposideros
307:Rhinolophus hipposideros
230:Rhinolophus hipposideros
184:Rhinolophus hipposideros
1735:Ruwenzori horseshoe bat
1658:Pearson's horseshoe bat
1648:Bourret's horseshoe bat
1524:Maendeleo horseshoe bat
1504:Big-eared horseshoe bat
1494:Maclaud's horseshoe bat
1356:RĂĽppell's horseshoe bat
1276:Darling's horseshoe bat
1136:Blasius's horseshoe bat
1069:Acuminate horseshoe bat
828:10.1242/jeb.203.17.2641
341:greater mouse-eared bat
269:and hunt their prey by
2442:Mammals of Afghanistan
1926:Dobson's horseshoe bat
1889:Thomas's horseshoe bat
1865:Swinny's horseshoe bat
1805:Bushveld horseshoe bat
1765:Shamel's horseshoe bat
1638:Osgood's horseshoe bat
1608:Timorese horseshoe bat
1571:Mehely's horseshoe bat
1454:Lander's horseshoe bat
1306:Eloquent horseshoe bat
1286:Decken's horseshoe bat
1266:Creagh's horseshoe bat
1196:Sulawesi horseshoe bat
528:Mehely's horseshoe bat
419:
404:
356:
220:
2437:Mammals of Azerbaijan
2288:Paleobiology Database
1920:R. xinanzhongguoensis
1899:Trefoil horseshoe bat
1561:Smaller horseshoe bat
1534:Malayan horseshoe bat
1464:Blyth's horseshoe bat
1444:Insular horseshoe bat
1366:Guinean horseshoe bat
1336:Greater horseshoe bat
1236:Andaman horseshoe bat
1226:Croslet horseshoe bat
1176:Cohen's horseshoe bat
1166:Canut's horseshoe bat
1156:Bornean horseshoe bat
1146:Bokhara horseshoe bat
1109:Arcuate horseshoe bat
1099:Halcyon horseshoe bat
881:10.1515/mamm.2004.030
588:: e.T19518A21972794.
417:
390:
354:
246:greater horseshoe bat
233:) is a type of small
218:
24:Lesser horseshoe bat
2132:Fauna Europaea (new)
1795:Forest horseshoe bat
1745:Sakeji horseshoe bat
1715:Rufous horseshoe bat
1628:Neriad horseshoe bat
1588:Mitred horseshoe bat
1514:Madura horseshoe bat
1474:Woolly horseshoe bat
1400:Lesser horseshoe bat
1296:Dent's horseshoe bat
1246:Convex horseshoe bat
1079:Adam's horseshoe bat
822:(Pt 17): 2641–2656.
260:Physical description
225:lesser horseshoe bat
166:R. hipposideros
1936:Ziama horseshoe bat
1685:Least horseshoe bat
959:Photographs, Video.
907:batlife-europe.info
570:Taylor, P. (2016).
399:; a short video by
45:Conservation status
1785:Thai horseshoe bat
1695:King horseshoe bat
1186:Cape horseshoe bat
968:2013-03-19 at the
540:Habitats Directive
420:
405:
357:
337:Maternity colonies
237:and North African
221:
2409:
2408:
2275:Open Tree of Life
1958:Taxon identifiers
1949:
1948:
1672:R. philippinensis
1652:R. paradoxolophus
942:978-0-600-56424-9
388:
213:
212:
68:
2469:
2402:
2401:
2389:
2388:
2376:
2375:
2363:
2362:
2350:
2349:
2337:
2336:
2335:
2309:
2308:
2296:
2295:
2283:
2282:
2270:
2269:
2257:
2256:
2244:
2243:
2241:NHMSYS0000080177
2231:
2230:
2218:
2217:
2205:
2204:
2192:
2191:
2179:
2178:
2166:
2165:
2153:
2152:
2140:
2139:
2127:
2126:
2114:
2113:
2101:
2100:
2088:
2087:
2075:
2074:
2062:
2061:
2049:
2048:
2036:
2035:
2026:
2025:
2013:
2012:
2000:
1999:
1998:
1985:
1984:
1983:
1953:
1952:
1883:R. thailandensis
1582:R. microglobosus
1340:R. ferrumequinum
999:
992:
985:
976:
975:
946:
918:
917:
915:
913:
899:
893:
892:
860:
854:
853:
847:
839:
807:
801:
794:
788:
785:
779:
774:
768:
767:
764:10.3354/esr00194
743:
737:
736:
734:
710:
704:
703:
693:
684:
678:
677:
671:
663:
653:
644:
635:
634:
616:
607:
606:
604:
602:
597:
567:
544:Berne Convention
471:Balearic Islands
465:Balearic Islands
389:
204:
186:
83:
82:
62:
57:
56:
33:
21:
20:
2477:
2476:
2472:
2471:
2470:
2468:
2467:
2466:
2412:
2411:
2410:
2405:
2397:
2392:
2384:
2379:
2371:
2366:
2358:
2353:
2345:
2340:
2331:
2330:
2325:
2312:
2304:
2299:
2291:
2286:
2278:
2273:
2265:
2262:Observation.org
2260:
2252:
2247:
2239:
2234:
2226:
2221:
2213:
2208:
2200:
2195:
2187:
2182:
2174:
2169:
2161:
2156:
2148:
2143:
2135:
2130:
2122:
2117:
2109:
2104:
2096:
2091:
2083:
2078:
2070:
2065:
2057:
2052:
2044:
2039:
2031:
2029:
2021:
2016:
2008:
2003:
1994:
1993:
1988:
1979:
1978:
1973:
1960:
1950:
1945:
1404:R. hipposideros
1380:R. hildebrandti
1230:R. coelophyllus
1130:R. belligerator
1053:
1009:
1003:
970:Wayback Machine
953:
943:
927:
925:Further reading
922:
921:
911:
909:
901:
900:
896:
861:
857:
841:
840:
808:
804:
795:
791:
786:
782:
775:
771:
744:
740:
711:
707:
691:
685:
681:
665:
664:
651:
645:
638:
631:
617:
610:
600:
598:
568:
564:
559:
536:
519:
503:
467:
455:
447:
381:
379:
366:
349:
322:
262:
195:
188:
182:
169:
77:
69:
58:
54:
47:
17:
12:
11:
5:
2475:
2465:
2464:
2459:
2457:Bats of Africa
2454:
2449:
2444:
2439:
2434:
2429:
2427:Bats of Europe
2424:
2407:
2406:
2404:
2403:
2390:
2377:
2364:
2351:
2338:
2322:
2320:
2314:
2313:
2311:
2310:
2297:
2284:
2271:
2258:
2245:
2232:
2219:
2206:
2193:
2180:
2167:
2154:
2141:
2128:
2119:Fauna Europaea
2115:
2102:
2089:
2076:
2063:
2050:
2037:
2027:
2014:
2001:
1986:
1970:
1968:
1962:
1961:
1947:
1946:
1944:
1943:
1933:
1923:
1916:
1906:
1903:R. trifoliatus
1896:
1886:
1879:
1872:
1862:
1852:
1842:
1832:
1822:
1812:
1802:
1792:
1782:
1779:R. shortridgei
1772:
1762:
1752:
1749:R. sakejiensis
1742:
1732:
1722:
1712:
1702:
1692:
1682:
1679:R. proconsulis
1675:
1665:
1655:
1645:
1635:
1625:
1622:R. mossambicus
1615:
1605:
1595:
1585:
1578:
1568:
1565:R. megaphyllus
1558:
1551:
1541:
1531:
1521:
1511:
1501:
1491:
1481:
1471:
1461:
1451:
1441:
1431:
1424:
1414:
1407:
1397:
1390:
1383:
1373:
1363:
1353:
1343:
1333:
1323:
1313:
1303:
1293:
1283:
1273:
1263:
1253:
1243:
1233:
1223:
1213:
1210:R. chiewkweeae
1203:
1193:
1183:
1173:
1163:
1153:
1143:
1133:
1126:
1116:
1106:
1096:
1086:
1076:
1065:
1063:
1055:
1054:
1052:
1051:
1045:
1043:Laurasiatheria
1039:
1033:
1027:
1021:
1014:
1011:
1010:
1002:
1001:
994:
987:
979:
973:
972:
960:
952:
951:External links
949:
948:
947:
941:
926:
923:
920:
919:
894:
875:(4): 307–322.
855:
802:
789:
780:
769:
738:
725:(1): 235–244.
705:
679:
636:
629:
608:
561:
560:
558:
555:
535:
532:
518:
515:
502:
499:
466:
463:
454:
451:
446:
443:
397:Forest of Dean
378:
377:United Kingdom
375:
365:
362:
348:
345:
321:
318:
261:
258:
250:horseshoe bats
248:. As with all
211:
210:
206:
205:
197:
196:
189:
178:
177:
171:
170:
163:
161:
157:
156:
149:
145:
144:
139:
135:
134:
129:
125:
124:
119:
115:
114:
109:
105:
104:
99:
95:
94:
89:
85:
84:
71:
70:
52:
49:
48:
43:
40:
39:
35:
34:
26:
25:
16:Species of bat
15:
9:
6:
4:
3:
2:
2474:
2463:
2460:
2458:
2455:
2453:
2452:Rhinolophidae
2450:
2448:
2445:
2443:
2440:
2438:
2435:
2433:
2430:
2428:
2425:
2423:
2420:
2419:
2417:
2400:
2395:
2391:
2387:
2382:
2378:
2374:
2369:
2365:
2361:
2356:
2352:
2348:
2343:
2339:
2334:
2328:
2324:
2323:
2321:
2319:
2315:
2307:
2302:
2298:
2294:
2289:
2285:
2281:
2276:
2272:
2268:
2263:
2259:
2255:
2250:
2246:
2242:
2237:
2233:
2229:
2224:
2220:
2216:
2211:
2207:
2203:
2198:
2194:
2190:
2185:
2181:
2177:
2172:
2168:
2164:
2159:
2155:
2151:
2146:
2142:
2138:
2133:
2129:
2125:
2120:
2116:
2112:
2107:
2103:
2099:
2094:
2090:
2086:
2081:
2077:
2073:
2068:
2064:
2060:
2055:
2051:
2047:
2042:
2038:
2034:
2028:
2024:
2019:
2015:
2011:
2006:
2002:
1997:
1991:
1987:
1982:
1976:
1972:
1971:
1969:
1967:
1963:
1959:
1954:
1941:
1937:
1934:
1931:
1930:R. yunanensis
1927:
1924:
1922:
1921:
1917:
1914:
1910:
1907:
1904:
1900:
1897:
1894:
1890:
1887:
1885:
1884:
1880:
1878:
1877:
1873:
1870:
1866:
1863:
1860:
1856:
1853:
1850:
1846:
1843:
1840:
1836:
1833:
1830:
1826:
1823:
1820:
1816:
1813:
1810:
1806:
1803:
1800:
1799:R. silvestris
1796:
1793:
1790:
1786:
1783:
1780:
1776:
1773:
1770:
1766:
1763:
1760:
1756:
1753:
1750:
1746:
1743:
1740:
1739:R. ruwenzorii
1736:
1733:
1730:
1726:
1723:
1720:
1716:
1713:
1710:
1706:
1703:
1700:
1696:
1693:
1690:
1686:
1683:
1681:
1680:
1676:
1673:
1669:
1666:
1663:
1659:
1656:
1653:
1649:
1646:
1643:
1639:
1636:
1633:
1629:
1626:
1623:
1619:
1616:
1613:
1609:
1606:
1603:
1599:
1596:
1593:
1589:
1586:
1584:
1583:
1579:
1576:
1572:
1569:
1566:
1562:
1559:
1557:
1556:
1552:
1549:
1545:
1542:
1539:
1535:
1532:
1529:
1525:
1522:
1519:
1518:R. madurensis
1515:
1512:
1509:
1505:
1502:
1499:
1495:
1492:
1489:
1485:
1482:
1479:
1475:
1472:
1469:
1465:
1462:
1459:
1455:
1452:
1449:
1445:
1442:
1439:
1435:
1432:
1430:
1429:
1428:R. indorouxii
1425:
1422:
1418:
1415:
1413:
1412:
1408:
1405:
1401:
1398:
1396:
1395:
1391:
1389:
1388:
1384:
1381:
1377:
1374:
1371:
1370:R. guineensis
1367:
1364:
1361:
1357:
1354:
1351:
1347:
1344:
1341:
1337:
1334:
1331:
1327:
1324:
1321:
1317:
1314:
1311:
1307:
1304:
1301:
1297:
1294:
1291:
1287:
1284:
1281:
1277:
1274:
1271:
1267:
1264:
1261:
1257:
1254:
1251:
1247:
1244:
1241:
1237:
1234:
1231:
1227:
1224:
1221:
1217:
1214:
1211:
1207:
1204:
1201:
1200:R. celebensis
1197:
1194:
1191:
1187:
1184:
1181:
1177:
1174:
1171:
1167:
1164:
1161:
1160:R. borneensis
1157:
1154:
1151:
1150:R. bocharicus
1147:
1144:
1141:
1137:
1134:
1132:
1131:
1127:
1124:
1120:
1117:
1114:
1110:
1107:
1104:
1100:
1097:
1094:
1090:
1087:
1084:
1080:
1077:
1074:
1073:R. acuminatus
1070:
1067:
1066:
1064:
1062:
1061:
1056:
1050:
1046:
1044:
1040:
1038:
1034:
1032:
1028:
1026:
1022:
1020:
1016:
1015:
1012:
1008:
1007:Rhinolophidae
1000:
995:
993:
988:
986:
981:
980:
977:
971:
967:
964:
961:
958:
955:
954:
944:
938:
934:
929:
928:
908:
904:
898:
890:
886:
882:
878:
874:
870:
866:
859:
851:
845:
837:
833:
829:
825:
821:
817:
813:
806:
799:
793:
784:
778:
773:
765:
761:
757:
753:
749:
742:
733:
728:
724:
720:
716:
709:
701:
697:
696:Le Rhinolophe
690:
683:
675:
669:
661:
657:
650:
643:
641:
632:
630:84-273-0265-7
626:
622:
615:
613:
596:
591:
587:
583:
582:
577:
575:
566:
562:
554:
551:
549:
545:
541:
531:
529:
525:
514:
512:
508:
498:
496:
492:
488:
484:
480:
476:
472:
462:
460:
450:
442:
441:
438:
434:
430:
426:
416:
412:
410:
402:
398:
394:
374:
371:
361:
353:
344:
342:
338:
334:
330:
328:
317:
315:
310:
308:
304:
300:
296:
292:
288:
282:
278:
276:
272:
268:
257:
255:
251:
247:
243:
240:
239:insectivorous
236:
232:
231:
226:
217:
207:
203:
198:
193:
187:
185:
179:
176:
175:Binomial name
172:
168:
167:
162:
159:
158:
155:
154:
150:
147:
146:
143:
142:Rhinolophidae
140:
137:
136:
133:
130:
127:
126:
123:
120:
117:
116:
113:
110:
107:
106:
103:
100:
97:
96:
93:
90:
87:
86:
81:
76:
72:
66:
61:
60:Least Concern
50:
46:
41:
36:
32:
27:
22:
19:
2432:Bats of Asia
2317:
1965:
1939:
1929:
1918:
1912:
1902:
1892:
1881:
1874:
1868:
1858:
1849:R. subbadius
1848:
1838:
1829:R. smithersi
1828:
1818:
1809:R. simulator
1808:
1798:
1789:R. siamensis
1788:
1778:
1768:
1758:
1748:
1738:
1728:
1718:
1709:R. robinsoni
1708:
1698:
1688:
1677:
1671:
1662:R. pearsonii
1661:
1651:
1641:
1631:
1621:
1611:
1602:R. monoceros
1601:
1591:
1580:
1574:
1564:
1555:R. mcintyrei
1553:
1548:R. marshalli
1547:
1538:R. malayanus
1537:
1528:R. maendeleo
1527:
1517:
1507:
1497:
1488:R. mabuensis
1487:
1477:
1467:
1457:
1447:
1437:
1426:
1421:R. imaizumii
1420:
1409:
1403:
1399:
1392:
1385:
1379:
1369:
1360:R. fumigatus
1359:
1349:
1339:
1329:
1319:
1309:
1299:
1289:
1279:
1269:
1259:
1249:
1239:
1229:
1219:
1209:
1199:
1189:
1179:
1169:
1159:
1149:
1139:
1128:
1122:
1112:
1102:
1092:
1082:
1072:
1058:
1041:Superorder:
1035:Infraclass:
932:
912:September 3,
910:. Retrieved
906:
897:
872:
868:
858:
844:cite journal
819:
815:
805:
797:
792:
783:
772:
755:
751:
741:
722:
718:
708:
699:
695:
682:
668:cite journal
659:
655:
620:
599:. Retrieved
585:
579:
573:
565:
552:
537:
534:Conservation
520:
517:Echolocation
504:
474:
468:
456:
448:
429:insecticides
425:agricultural
421:
406:
369:
367:
364:Distribution
358:
335:
331:
323:
311:
306:
283:
279:
271:echolocation
263:
229:
228:
224:
222:
183:
181:
165:
164:
152:
18:
2158:iNaturalist
1990:Wikispecies
1859:R. subrufus
1689:R. pusillus
1612:R. montanus
1592:R. mitratus
1508:R. macrotis
1498:R. maclaudi
1448:R. keyensis
1411:R. huananus
1394:R. hillorum
1350:R. formosae
1330:R. euryotis
1310:R. eloquens
1290:R. deckenii
1280:R. darlingi
1260:R. cornutus
1250:R. convexus
1240:R. cognatus
1220:R. clivosus
1190:R. capensis
1123:R. beddomei
1113:R. arcuatus
1060:Rhinolophus
798:Ir. Nat. J.
656:Vespertilio
601:21 February
435:. However,
295:butterflies
273:, emitting
153:Rhinolophus
2416:Categories
2333:Q106447960
1893:R. thomasi
1869:R. swinnyi
1819:R. sinicus
1769:R. shameli
1759:R. sedulus
1642:R. osgoodi
1575:R. mehelyi
1468:R. lepidus
1458:R. landeri
1320:R. euryale
1270:R. creaghi
1180:R. cohenae
1103:R. alcyone
1093:R. affinis
1049:Chiroptera
816:J Exp Biol
758:(1–2): 2.
662:: 211–239.
557:References
511:Ain Draham
495:Formentera
393:Wye Valley
291:mosquitoes
275:ultrasound
132:Chiroptera
38:In France
1839:R. stheno
1719:R. rouxii
1632:R. nereis
1478:R. luctus
1170:R. canuti
1140:R. blasii
1017:Kingdom:
327:hibernate
192:Bechstein
160:Species:
98:Kingdom:
92:Eukaryota
2327:Wikidata
2228:13800555
2176:11041856
2030:BioLib:
1975:Wikidata
1940:R. ziama
1913:R. virgo
1876:R. tatar
1729:R. rufus
1438:R. inops
1387:R. hilli
1300:R. denti
1083:R. adami
1037:Eutheria
1031:Mammalia
1025:Chordata
1023:Phylum:
1019:Animalia
966:Archived
889:86180828
869:Mammalia
836:10934005
548:EUROBATS
479:Mallorca
433:habitats
395:and the
267:colonies
254:noseleaf
235:European
138:Family:
122:Mammalia
112:Chordata
108:Phylum:
102:Animalia
88:Domain:
65:IUCN 3.1
2399:1049410
2373:9085869
2306:1049409
2215:1004693
2150:2432614
1981:Q585476
1047:Order:
1029:Class:
800:1 – 38.
507:Tunisia
501:Tunisia
491:Cabrera
483:Menorca
469:In the
445:Ireland
347:Habitat
314:Tunisia
303:spiders
194:, 1800)
148:Genus:
128:Order:
118:Class:
63: (
2386:946405
2293:376819
2280:610867
2189:631434
2124:305391
2098:RHLPHI
2085:328775
2018:ARKive
1699:R. rex
957:ARKive
939:
887:
834:
627:
459:Israel
453:Israel
320:Mating
301:, and
2394:WoRMS
2360:14558
2355:EUNIS
2347:7G2TQ
2301:WoRMS
2254:77218
2202:19518
2171:IRMNG
2163:40695
2106:EUNIS
2059:4S97S
2046:84662
885:S2CID
692:(PDF)
652:(PDF)
487:Ibiza
299:moths
287:flies
2381:ITIS
2368:GBIF
2249:NCBI
2197:IUCN
2184:ITIS
2145:GBIF
2111:1545
2093:EPPO
2041:BOLD
2033:2255
937:ISBN
914:2019
850:link
832:PMID
702:: 5.
674:link
625:ISBN
603:2022
586:2016
526:and
493:and
485:and
409:here
223:The
2342:CoL
2267:408
2236:NBN
2223:MSW
2210:MDD
2080:EoL
2067:CMS
2054:CoL
2005:ADW
877:doi
824:doi
820:203
760:doi
727:doi
590:doi
505:In
401:NRW
242:bat
2418::
2396::
2383::
2370::
2357::
2344::
2329::
2303::
2290::
2277::
2264::
2251::
2238::
2225::
2212::
2199::
2186::
2173::
2160::
2147::
2134::
2121::
2108::
2095::
2082::
2069::
2056::
2043::
2020::
2007::
1992::
1977::
905:.
883:.
873:68
871:.
867:.
846:}}
842:{{
830:.
818:.
814:.
754:.
750:.
723:91
721:.
717:.
700:14
698:.
694:.
670:}}
666:{{
660:16
658:.
654:.
639:^
611:^
584:.
578:.
530:.
481:,
473:,
461:.
297:,
293:,
289:,
256:.
1942:)
1938:(
1932:)
1928:(
1915:)
1911:(
1905:)
1901:(
1895:)
1891:(
1871:)
1867:(
1861:)
1857:(
1851:)
1847:(
1841:)
1837:(
1831:)
1827:(
1821:)
1817:(
1811:)
1807:(
1801:)
1797:(
1791:)
1787:(
1781:)
1777:(
1771:)
1767:(
1761:)
1757:(
1751:)
1747:(
1741:)
1737:(
1731:)
1727:(
1721:)
1717:(
1711:)
1707:(
1701:)
1697:(
1691:)
1687:(
1674:)
1670:(
1664:)
1660:(
1654:)
1650:(
1644:)
1640:(
1634:)
1630:(
1624:)
1620:(
1614:)
1610:(
1604:)
1600:(
1594:)
1590:(
1577:)
1573:(
1567:)
1563:(
1550:)
1546:(
1540:)
1536:(
1530:)
1526:(
1520:)
1516:(
1510:)
1506:(
1500:)
1496:(
1490:)
1486:(
1480:)
1476:(
1470:)
1466:(
1460:)
1456:(
1450:)
1446:(
1440:)
1436:(
1423:)
1419:(
1406:)
1402:(
1382:)
1378:(
1372:)
1368:(
1362:)
1358:(
1352:)
1348:(
1342:)
1338:(
1332:)
1328:(
1322:)
1318:(
1312:)
1308:(
1302:)
1298:(
1292:)
1288:(
1282:)
1278:(
1272:)
1268:(
1262:)
1258:(
1252:)
1248:(
1242:)
1238:(
1232:)
1228:(
1222:)
1218:(
1212:)
1208:(
1202:)
1198:(
1192:)
1188:(
1182:)
1178:(
1172:)
1168:(
1162:)
1158:(
1152:)
1148:(
1142:)
1138:(
1125:)
1121:(
1115:)
1111:(
1105:)
1101:(
1095:)
1091:(
1085:)
1081:(
1075:)
1071:(
998:e
991:t
984:v
945:.
916:.
891:.
879::
852:)
838:.
826::
766:.
762::
756:8
735:.
729::
676:)
633:.
605:.
592::
576:"
572:"
403:.
227:(
190:(
67:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.