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to harvest, making holes in the crop, which can make individual items unsuitable to sell for aesthetic reasons, and can make the crop more vulnerable to rot and disease. Excessive buildup of slugs within some wastewater treatment plants with inadequate screening have been found to cause process issues resulting in increased energy and chemical use.
900:. In the banana slugs, the penis sometimes becomes trapped inside the body of the partner. Apophallation allows the slugs to separate themselves by one or both of the slugs chewing off the other's or its own penis. Once the penis has been discarded, banana slugs are still able to mate using only the female parts of the reproductive system. 1709: 1708: 1713: 1712: 1707: 1838:) are a commercially available biological control method that are effective against a wide range of common slug species. The nematodes are applied in water and actively seek out slugs in the soil and infect them, leading to the death of the slug. This control method is suitable for use in organic growing systems. 1714: 831:
provides some protection against predators, as it can make the slug hard to pick up and hold by a bird's beak, for example, or the mucus itself can be distasteful. Some slugs can also produce very sticky mucus which can incapacitate predators and can trap them within the secretion. Some species of slug, such as
1132:. Some slugs are selective towards certain parts or developmental stages of the fungi they eat, though this is very variable. Depending on the species and other factors, slugs eat only fungi at specific stages of development. In other cases, whole mushrooms can be eaten, without any selection or bias towards 882:, having both female and male reproductive organs. Once a slug has located a mate, they encircle each other and sperm is exchanged through their protruded genitalia. A few days later, the slugs lay approximately thirty eggs in a hole in the ground, or beneath the cover of an object such as a fallen log. 1788:
The great majority of slug species are harmless to humans and to their interests, but a small number of species are serious pests of agriculture and horticulture. They can destroy foliage faster than plants can grow, thus killing even fairly large plants. They also feed on fruits and vegetables prior
1738:. This, combined with the slippery mucus they produce, makes slugs more difficult for predators to grasp. The unpleasant taste of the mucus is also a deterrent. Slugs can also incapacitate predators through the production of a highly sticky and elastic mucus which can trap predators in the secretion. 830:
trail" a slug leaves behind has some secondary effects: other slugs coming across a slime trail can recognise the slime trail as produced by one of the same species, which is useful in finding a mate. Following a slime trail is also part of the hunting behaviour of some carnivorous slugs. Body mucus
823:. The thin mucus spreads from the foot's centre to its edges, whereas the thick mucus spreads from front to back. Slugs also produce thick mucus that coats the whole body of the animal. The mucus secreted by the foot contains fibres that help prevent the slug from slipping down vertical surfaces. 273:
families of land slugs form part of several quite different evolutionary lineages, which also include snails. Thus, the various families of slugs are not closely related, despite the superficial similarity in overall body form. The shell-less condition has arisen many times independently as an
1711: 306:, contains various clades of snails, semi-slugs (i.e. snails whose shells are too small for them to retract fully into), and slugs. The taxonomy of this group is in the process of being revised in light of DNA sequencing. It appears that pulmonates are paraphyletic and basal to the 206: 205: 211: 209: 204: 1971: 210: 708:, which is easier to see when open; also on the right side at the front are the genital opening and anus. Veronicellid slugs have a mantle covering the whole dorsal part of the body, they have no respiratory opening, and the anus opens posteriorly. 208: 3061:
Symondson, W. O. C.; Glen, D. M.; Erickson, M. L.; Liddell, J. E.; Langdon, C. J. (2000). "Do earthworms help to sustain the slug predator Pterostichus melanarius (Coleoptera: Carabidae) within crops? Investigations using monoclonal antibodies".
1832:, especially because domestic or wild animals may be exposed to the bait. The environmentally safer iron phosphate has been shown to be at least as effective as baits. Methiocarb baits are no longer widely used. Parasitic nematodes ( 3366: 746:: Most slugs retain a remnant of their shell, which is usually internalized. This organ generally serves as storage for calcium salts, often in conjunction with the digestive glands. An internal shell is present in the 2293:
Schilthuizen, M.; Thome, J. W. (2008). "Valiguna flava (Heynemann, 1885) from Indonesia and Malaysia: Redescription and Comparison with Valiguna siamensis (Martens, 1867)(Gastropoda: Soleolifera: Veronicellidae)".
1745:. In contrast to the general behavioral pattern, the Kerry slug retracts its head, lets go of the substrate, rolls up completely, and stays contracted in a ball-like shape. This is a unique feature among all the 1710: 808:(a 180° twisting of the internal organs) during development. Internally, slug anatomy clearly shows the effects of this rotation—but externally, the bodies of slugs appear more or less symmetrical, except the 792:. They must generate protective mucus to survive. Many species are most active just after a rain because of the moist ground or during nighttime. In drier conditions, they hide in damp places such as under 3526: 3373: 2941: 2396: 207: 3251: 1988: 1776:
conditions. Slugs are prone to attack during the summer, when the availability of resources is reduced. During winter, the aggressive responses are substituted by a
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When attacked, slugs can contract their body, making themselves harder and more compact and more still and round. By doing this, they become firmly attached to the
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Dayrat, B. (2009). "Review of the current knowledge of the systematics of Onchidiidae (Mollusca: Gastropoda: Pulmonata) with a checklist of nominal species".
1208:) includes slugs in its diet. Amphibians such as frogs and toads have long been regarded as important predators of slugs. Among them are species in the genus 252:
is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a small internal shell, particularly
3336:"Chapter 3: Dipteras as predators and parasitoids of terrestrial gastropods, with emphasis on Phoridae, Calliphoridae, Sarcophagidae, Muscidae and Fannidae" 2536: 3523: 1907: 526: 740:
that it travels on, which helps prevent damage to the foot tissues. Around the edge of the foot in some slugs is a structure called the 'foot fringe'.
2956: 3616: 1929: 1491:, are known to feed on slugs. Ants are a common predator of slugs; some ant species are deterred by the slug's mucus coating, while others such as 2423: 726:, have a prominent ridge running over their back along the middle of the tail (sometimes along the whole tail, sometimes only the posterior part). 2237:
Webb & Van Beneden, 1836, y consideraciones sobre la posicion sistematica de la familia Parmacellidae (Mollusca, Pulmonata, Stylommatophora)"
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Branson, B. A (1980). "The recent Gastropoda of Oklahoma, Part VIII. The slug families Limacidae, Arionidae, Veronicellidae, and Philomycidae".
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Rollo, C. D.; Wellington, W. G. (1979). "Intra- and inter-specific agonistic behavior among terrestrial slugs (Pulmonata: Stylommatophora)".
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Loest, R. A. (1979). "Ammonia Volatilization and Absorption by Terrestrial Gastropods_ a Comparison between Shelled and Shell-Less Species".
633: 436: 1156:, research investments have been made to discover and investigate potential predators in order to establish biological control strategies. 3870: 1842: 3858: 3153: 3248: 1841:
Other slug control methods are generally ineffective on a large scale, but can be somewhat useful in small gardens. These include
2973: 1184:, vertebrates can occasionally feed on, or be specialised predators of, slugs. Fish that feed on slugs include the brown trout ( 903:
In a temperate climate, slugs usually live one year outdoors. In greenhouses, many adult slugs may live for more than one year.
732:: The bottom side of a slug, which is flat, is called the 'foot'. Like almost all gastropods, a slug moves by rhythmic waves of 3524:'Evaluating Opportunities and Barriers to Improving the Energy Efficiency of Small Nebraska Wastewater Treatment Plants', pp.83 3594: 1880:
in the US to be more severely affected under low till because this increases organic matter, thus providing food and shelter.
1152:. The predation of slugs has been the subject of studies for at least a century. Because some species of slugs are considered 938:
Most species of slugs are generalists, feeding on a broad spectrum of organic materials, including leaves from living plants,
3827: 3757: 3421: 2514: 2064:"Ten new complete mitochondrial genomes of pulmonates (Mollusca: Gastropoda) and their impact on phylogenetic relationships" 266:, which applies to gastropods that have a coiled shell large enough that they can fully retract their soft parts into it). 3579: 3547: 3347: 2755: 3838: 3776: 3038: 3642: 3260: 2348:"Adhesive defence mucus secretions in the red triangle slug (Triboniophorus graeffei) can incapacitate adult frogs" 2232: 2062:
White, T. R.; Conrad, M. M.; Tseng, R.; Balayan, S.; Golding, R.; de Frias Martins, A. M.; Dayrat, B. A. (2011).
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Avery, R. A. (1966). "Food and feeding habits of the Common lizard (Lacerta vivipara) in the west of England".
1293:), are exclusively slug eaters. Several lizards include slugs in their diet. This is the case in the slowworm ( 676:
on their head. The upper pair is light-sensing and has eyespots at the ends, while the lower pair provides the
3022: 3895: 2825:"The energetic consequences of dietary specialization in populations of the garter snake, Thamnophis elegans" 1834: 1610: 500: 479: 3793: 2040: 1955: 3900: 3200:
Teixeira, C. G.; Thiengo, S. C.; Thome, J. W.; Medeiros, A. B.; Camillo-Coura, L.; Agostini, A. A. (1993).
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and individuals from other species when competing for resources. This aggressiveness is also influenced by
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Slugs' bodies are made up mostly of water and, without a full-sized shell, their soft tissues are prone to
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from eating raw slugs. Live slugs that are accidentally eaten with improperly cleaned vegetables (such as
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underground during the winter in temperate climates, but in other species, the adults die in the autumn.
1616: 607: 583: 17: 3863: 3620: 3542: 3502:"SlugClear Ultra: Highly efficient protection against slugs and snails | Gardening tips and advice" 3890: 3570: 3286: 3282:
Monograph of the land and freshwater Mollusca of the British Isles. Testacellidae. Limacidae. Arionidae
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is documented, but varies greatly among slug species. Slugs often resort to aggression, attacking both
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As control measures, baits are commonly used in both agriculture and the garden. In recent years,
672:: Like other pulmonate land gastropods, the majority of land slugs have two pairs of 'feelers' or 1487: 1054: 3446: 1919: 1624:-causing nematode, have larval stages that can only live in molluscs, including slugs, such as 1553: 1041:. They also feed on carrots, peas, apples, and cabbage that are offered as a sole food source. 722: 1263:), is not a specialized predator of slugs but occasionally feeds on them. The redbelly snake ( 3867: 3384: 2347: 1523: 1378: 1259: 39: 3372:. Forest Service Department of Agriculture, Fisheries and Food. 12 July 2009. Archived from 3854: 3714: 3071: 2788: 2480: 2397:"Perverted cannibalistic hermaphrodites haunt the Pacific Northwest! " The Oyster's Garter" 2359: 2075: 1869: 1772:, because the availability of resources such as shelter and food may be compromised due to 1641:
of molluscs. To complete their development, many dipterans use slugs as hosts during their
1517: 1192: 733: 275: 1521:, is known to parasitise several dozen species of molluscs, including many slugs, such as 1164:
Slugs are preyed upon by virtually every major vertebrate group. With many examples among
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can often be seen swarming about their host's body, and live in its respiratory cavity.
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Leonard, J.L.; Pearse, J.S.; Harper, A.B. (2002). "Comparative reproductive biology of
2375: 2275: 2191: 2183: 2098: 2063: 1999: 1978: 1853: 1761: 1584: 1529: 1254: 1243: 1122: 1086: 1084:). Species of mushroom producing fungi used as food source by slugs include milk-caps ( 270: 120: 3280: 2809: 2442: 2324:. College of Agricultural, Consumer and Environmental Sciences, University of Illinois 2129: 3772: 3763: 3753: 3417: 3343: 3227: 3087: 3083: 3034: 2895: 2846: 2824: 2751: 2615: 2530: 2103: 1753:) a portion of their tail to help the slug escape from a predator. Some slug species 1565: 1481: 1394: 1346: 1313: 1153: 1014: 603: 408: 3664: 3222: 3201: 3099: 2890: 2865: 2500: 2379: 2317: 2195: 95: 3689: 3483: 3456: 3316: 3217: 3172: 3131: 3079: 2922: 2885: 2836: 2796: 2599: 2488: 2438: 2367: 2279: 2267: 2179: 2175: 2148: 2093: 2083: 1941: 1336: 1321: 696: 595: 414: 400: 113: 3176: 2977: 2801: 2772: 2492: 1891: 3530: 3255: 3060: 1806:), or improperly cooked slugs (for use in recipes requiring larger slugs such as 1604: 1589: 1541: 1426: 1356: 1295: 1289: 1283: 1271: 681: 487: 466: 325: 3120:(Schrank) – Acari-Trombidiformes and its relationship with pulmonate molluscs". 927: 919:. Most carnivorous slugs on occasion also eat dead specimens of their own kind. 3749: 1857: 1678: 1576:
Several species of nematodes are known to parasitise slugs. The nematode worms
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snakes are known predators of slugs. Coastal populations of the garter snake,
3879: 3460: 3320: 2899: 2874:(Semper, 1885) (Gastropoda; Veronicellidae): a trophic relationship revealed" 2864:
Maia, T; Dorigo, T. A.; Gomes, S. R.; Santos, S. B.; Rocha, C. F. D. (2012).
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Some species present different response behaviors when attacked, such as the
1651: 1646: 1626: 1598: 1578: 1559: 1547: 1449: 1399: 1326: 1110: 1066: 994: 932: 885: 879: 833: 591: 517: 445: 422: 307: 104: 80: 65: 3821: 3091: 2850: 2619: 2107: 1860:, and copper. Salt kills slugs by causing water to leave the body owing to 1847: 1819: 1664: 1440: 1186: 1149: 1049: 1038: 751: 508: 378: 86: 3411: 3231: 2863: 1120:
are also eaten by slugs. Slime molds used as food source by slugs include
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juvenile with eggs of another (larger) slug species, most likely Arionidae
2770: 2152: 1995: 1946: 1914: 1829: 1807: 1769: 1765: 1722: 1341: 1222: 1133: 1072: 1022: 975: 827: 819:: one is thin and watery, and the other thick and sticky. Both kinds are 809: 789: 777: 755: 705: 701: 553: 428: 363: 352: 343: 319: 303: 299: 295: 235: 192: 186: 181: 2187: 965: 804:, to help retain body moisture. Like all other gastropods, they undergo 2611: 1799: 1777: 1742: 1727: 1693: 1638: 1621: 1535: 1504: 1492: 1415: 1410: 1275:) feed mainly but not solely on slugs, while some species in the genus 1238: 1145: 998: 820: 700:. In stylommatophoran slugs, on the right-hand side of the mantle is a 642: 621: 562: 544: 496: 475: 457: 396: 387: 311: 167: 75: 70: 3303:
Robinson, W. H.; Foote, B. A. (1968). "Biology and Immature Stages of
2841: 2371: 3745: 2773:"Terrestrial and benthic foods in the diet of the shortjawed kokopu, 2462: 2130:"The physical properties of the pedal mucus of the terrestrial slug, 1962: 1754: 1746: 1683: 1673: 1660: 1656: 1634: 1602:. Species of widely known medical importance pertaining to the genus 1476: 1445: 1431: 1420: 1373: 1249: 1198: 1177: 1077: 1045: 982: 970: 959: 912: 896: 890: 850: 845: 801: 793: 747: 677: 369: 291: 279: 257: 242: 137: 109: 3487: 2603: 812:, which is on one side of the animal, normally the right-hand side. 736:
on the underside of its foot. It simultaneously secretes a layer of
2649: 2018: 1936: 1898: 1773: 1750: 1688: 1668: 1642: 1512: 1464: 1331: 1216: 1104: 1018: 951: 943: 253: 157: 31: 3342:. Wallingford, Oxfordshire, UK: CABI Publishing. pp. 88–124. 3543:"Health and Medicals News – Man's brain infected by eating slugs" 3416:(in German). Hamburg and Berlin: Verlag Paul Parey. p. 138. 2750:. Boundary Row, London, UK: Chapman & Hall. pp. 428 pp. 2672: 1877: 1861: 1803: 1435: 1383: 1365: 1351: 1165: 1116: 1034: 1026: 1002: 990: 947: 245: 3247:
Senanayake, S. N.; Pryor, D. S.; Walker, J.; Konecny, P. (2003).
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McDowall, R. M.; Main, M. R.; West, D. W.; Lyon, G. L. (1996).
1593: 1460: 1277: 1253:, while inland populations have a generalized diet. One of its 1234: 1173: 1010: 1006: 939: 870: 863: 858: 147: 3437:
Pekarinen, E. (1994). "Autotomy in arionid and limacid slug".
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Slug contracts itself and retracts its tentacles when attacked
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USDA Farmer's Bulletin No. 1895. Revised 1959. Hosted by the
3249:"First report of human angiostrongyliasis acquired in Sydney" 2346:
Gould, John; Valdez, Jose W.; Upton, Rose (8 February 2019).
2013: 1873: 1508: 1404: 1388: 1230: 1030: 986: 955: 916: 816: 737: 657: 262: 220: 56: 35: 3412:
Kerney, M. P; Cameron, A. D. & Jungbluth, J. H. (1983).
3199: 2292: 1810:), can act as a vector for a parasitic infection in humans. 1749:, and among most other slugs. Some slugs can self-amputate ( 714:: The part of a slug behind the mantle is called the 'tail'. 694:: On top of the slug, behind the head, is the saddle-shaped 2588:"Feeding activities of slugs on Myxomycetes and macrofungi" 1369: 1210: 1181: 1169: 837:, secrete slime cords to suspend a pair during copulation. 2517:. 4 April 2005. Archived from the original on 4 April 2005 2421: 2241:
Boletín de la Sociedad de Historia Natural de les Baleares
68:
land slugs, including (from the top) the larger drawings:
1828:
baits have emerged and are preferred over the more toxic
1456: 1361: 3033:. Sofia, Moscow: PENSOFT Publishers. pp. 221–227. 2061: 1247:, have a specialised diet consisting of slugs, such as 888:
has been reported only in some species of banana slug (
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The external anatomy of a slug includes the following:
3595:"Less toxic iron phosphate slug bait proves effective" 3473: 3031:
Natural History and Applied Ecology of Carabid Beetles
2230: 310:, which are a terminal branch of the tree. The family 2781:
New Zealand Journal of Marine and Freshwater Research
2399:. Theoystersgarter.com. 24 March 2008. Archived from 3302: 2391: 2389: 1872:
worsens slug infestations. Hammond et al. 1999 find
3202:"On the diversity of mollusc intermediate hosts of 3151: 2823:Britt, E. J.; Hicks, J. W.; Bennett, A. F. (2006). 2127: 2585: 3617:"~ Slug Traps ~ Death by Beer Offers and Reviews" 3361: 3359: 2386: 1864:but this is not used for agricultural control as 3877: 2581: 2579: 2577: 2575: 2573: 2535:: CS1 maint: bot: original URL status unknown ( 2424:"Penis-biting slugs: wild claims and confusions" 2345: 1495:will roll the slug in dirt to absorb its mucus. 3333: 3206:Morera & Cespedes, 1971 in southern Brazil" 2748:Terrestrial Slugs: Biology, ecology and control 2699: 3356: 3309:Annals of the Entomological Society of America 3021:Brandmayr, P.; Paill, W.; et al. (2000). 3020: 2214:Proceedings of the Oklahoma Academy of Science 3791: 3729: 3597:. Extension.oregonstate.edu. 25 February 2008 3436: 3327: 3054: 2570: 2211: 2207: 2205: 2123: 2121: 2119: 2117: 3296: 2870:(Ihering, 1911) (Serpentes; Dipsadidae) and 2741: 2739: 2737: 2735: 2733: 2731: 2729: 2727: 2550: 2548: 2546: 2035: 2033: 1697:, are facultative parasitoids of Arionidae. 1498: 302:– solely comprise slugs. A third group, the 3568: 3195: 3193: 3147: 3145: 2473:Invertebrate Reproduction & Development 2311: 2309: 1792:In a few rare cases, humans have developed 1229:Reptiles that feed on slugs include mainly 1076:), which respectively feed on slime molds ( 3243: 3241: 3111: 3109: 3023:"Slugs as prey for larvae and imagines of 2712:. University of Michigan Museum of Zoology 2257: 2202: 2114: 94: 55: 3467: 3450: 3414:Die Landschnecken Nord- und Mitteleuropas 3221: 3014: 2906: 2889: 2840: 2816: 2800: 2724: 2543: 2515:"Slugs and Their Management, HYG-2010-95" 2097: 2087: 2030: 637:Jesús Ortea and Juan José Bacallado, 2016 3855:Land Slugs and Snails and Their Control. 3735: 3190: 3142: 2764: 2306: 2159: 2055: 1721: 1704: 964: 926: 869: 857: 844: 800:, rocks and manmade structures, such as 770: 656: 260:(this is in contrast to the common name 201: 27:Shell-less terrestrial gastropod mollusc 3340:Natural Enemies of Terrestrial Molluscs 3285:. Taylor Brothers, Leeds. Introduction 3238: 3106: 2695: 2693: 2322:Home, Yard & Garden Pest Newsletter 1982:, the world's largest terrestrial slug. 14: 3878: 3619:. Gardening-guru.co.uk. Archived from 3405: 3270: 1372:. Studies on slug predation also cite 1202:slug. Similarly, the shortjaw kokopu ( 915:by eating decaying plant material and 3152:Barker, G. M.; Ramsay, G. W. (1978). 3115: 2939: 2912: 2745: 2422:Reise, H; Hutchinson, J.M.C. (2002). 2341: 2339: 2165: 2128:Denny, M. W.; Gosline, J. M. (1980). 1730:, which is found only in this species 720:: Some species of slugs, for example 3665:"Using Diatomaceous Earth for Slugs" 3643:"How to Get Rid of Slugs and Snails" 3573:Man's brain infected by eating slugs 3367:"Forestry and Kerry Slug Guidelines" 3158:(Acari: Ereynetidae) in New Zealand" 2690: 2586:Keller, H. W.; Snell, K. L. (2002). 981:Slugs can feed on a wide variety of 911:Slugs play an important role in the 3859:UNT Government Documents Department 3835:"The Secret Lives of Jumping Slugs" 3580:Australian Broadcasting Corporation 3548:Australian Broadcasting Corporation 3307:, a Phorid Predator of Slug Eggs". 2641:"Worm-eating slug found in garden ( 2633: 1320:Birds that prey upon slugs include 1287:) and the common slug eater snake ( 862:Pair of mating slugs, suspended by 24: 3785: 3210:Memórias do Instituto Oswaldo Cruz 3116:Baker, R. A. (1978). "The food of 2927:10.1111/j.1469-7998.1966.tb03886.x 2336: 1783: 1726:The alarm response posture of the 1608:are also parasites of slugs. Both 1044:Slugs from different families are 278:, and thus the category "slug" is 25: 3912: 3809: 3690:"Guide to Using Copper Slug Tape" 2471:(Gastropoda; Stylommiatophora)". 2315: 2231:Alonso, M. R.; Ibañe, M. (1981). 1257:, the Northwestern garter snake ( 1144:Slugs are preyed upon by various 922: 3830:Ohio State University Extension. 3815: 3261:The Medical Journal of Australia 3084:10.1046/j.1365-294x.2000.01006.x 2808: 1987: 1970: 1954: 1928: 1906: 1890: 1507:by several organisms, including 1470: 974:sp. feeding on a small fruit in 680:. Both pairs are retractable in 124: 3707: 3682: 3657: 3635: 3609: 3587: 3569:Anna Salleh (20 October 2003). 3562: 3535: 3516: 3494: 3430: 3223:10.1590/S0074-02761993000300020 2992: 2976:. Cardiff.ac.uk. Archived from 2966: 2933: 2891:10.5007/2175-7925.2012v25n1p211 2857: 2829:Journal of Experimental Biology 2777:Clarke (Teleostei: Galaxiidae)" 2665: 2507: 2456: 2431:Trends in Ecology and Evolution 2415: 2141:Journal of Experimental Biology 1671:, specially those in the genus 1455:Mammals that eat slugs include 1190:), which occasionally feeds on 840: 3792:Burton, D. W. (January 1982). 3000:"Ecological Benefits of Slugs" 2286: 2251: 2224: 2180:10.1086/physzool.52.4.30155937 1645:. Some species of blow-flies ( 1159: 661:The external anatomy of a slug 652: 238:for any apparently shell-less 13: 1: 3204:Angiostrongylus costaricensis 3177:10.1080/00779962.1978.9722316 2802:10.1080/00288330.1996.9516713 2493:10.1080/07924259.2002.9652738 2443:10.1016/S0169-5347(02)02453-9 2024: 1835:Phasmarhabditis hermaphrodita 1813: 1611:Angiostrongylus costaricensis 766: 3439:Journal of Molluscan Studies 3279:(1902). Part 8, pages 1–52. 1691:, mainly those in the genus 1687:. House flies in the family 1681:, including many species of 1139: 1090:spp.), the oyster mushroom ( 989:, including flowers such as 7: 3828:Slugs and Their Management. 3738:Handbook of vegetable pests 3476:Canadian Journal of Zoology 2007: 1795:Angiostrongylus cantonensis 1700: 1617:Angiostrongylus cantonensis 815:Slugs produce two types of 285: 10: 3917: 3123:Journal of Natural History 2872:Potamojanuarius lamellatus 1883: 1817: 1760:Intra- and inter-specific 906: 684:slugs, but contractile in 29: 3338:. In Barker, G. M (ed.). 3136:10.1080/00222937000770481 2955:(3): 8–11. Archived from 2700:Sandy; Misner, L; Balog. 2272:10.11646/zootaxa.2068.1.1 2235:Parmacella valenciannesii 1868:is detrimental to crops. 1655:are known parasitoids of 1588:respectively live in the 1499:Parasites and parasitoids 1452:birds as slug predators. 1311:) and the common lizard ( 1308:Cyclodomorphus casuarinae 1266:Storeria occipitomaculata 750:and Parmacellidae. Adult 461:Gude & Woodward, 1921 191:Multiple lineages within 178: 173: 121:Scientific classification 119: 102: 93: 63: 54: 49: 3334:Coupland, J. B. (2004). 3165:New Zealand Entomologist 3027:(Coleoptera: Carabidae)" 2089:10.1186/1471-2148-11-295 2068:BMC Evolutionary Biology 1129:Symphytocarpus flaccidus 1102:). Other genera such as 1048:. It is the case in the 954:and eat other slugs and 3851:The Nature Conservancy. 3736:Capinera, John (2020). 1488:Pterostichus melanarius 1299:), the bobtail lizard ( 1269:) and the brown snake ( 1073:Ariolimax californianus 1055:Philomycus carolinianus 512:H. & A. Adams, 1855 314:are also polyphyletic. 3645:. asthegardenturns.com 3461:10.1093/mollus/60.1.19 3392:Cite journal requires 3321:10.1093/aesa/61.6.1587 2868:Sibynomorphus neuwiedi 2465:Ariolimax californicus 1920:Geotrupes stercorarius 1731: 1719: 1667:. Flies in the family 1554:Tandonia budapestensis 1511:and a wide variety of 1475:Beetles in the family 1305:), the she-oak skink ( 1061:Phylomicus flexuolaris 978: 935: 875: 874:A pair of mating slugs 867: 866:thread and pad on wall 855: 785: 723:Tandonia budapestensis 662: 224: 38:. For other uses, see 30:For marine slugs, see 3868:University of Florida 2940:Hewer, A. M. (1948). 2168:Physiological Zoology 2132:Ariolimax columbianus 1923:) moving a dead slug. 1856:, crushed eggshells, 1725: 1717: 1677:, are parasitoids of 1524:Deroceras reticulatum 1379:Deroceras reticulatum 1260:Thamnophis ordinoides 1096:) and the penny bun ( 1037:, and fruits such as 968: 930: 894:) and one species of 873: 861: 848: 774: 660: 290:Of the six orders of 214: 40:Slug (disambiguation) 3896:Mollusc common names 3824:at Wikimedia Commons 3794:"How to be sluggish" 3717:. Newton.dep.anl.gov 3522:Thompson, M. (2018) 3254:30 July 2022 at the 3156:Riccardoella limacum 3118:Riccardoella limacum 2949:Tazmanian Naturalist 2710:Animal Diversity Web 2673:"What Do Slugs Eat?" 2556:"What Do Slugs Eat?" 2153:10.1242/jeb.88.1.375 2041:"How to be sluggish" 1870:Conservation tillage 1518:Riccardoella limacum 1193:Arion circumscriptus 734:muscular contraction 521:Albers-Martens, 1860 276:convergent evolution 215:A slug on a wall in 3901:Polyphyletic groups 3715:"Slugs and Osmosis" 3623:on 18 November 2012 3529:6 June 2022 at the 3504:. LoveTheGarden.com 3076:2000MolEc...9.1279S 2942:"Tazmanian lizards" 2793:1996NZJMF..30..257M 2775:Galaxias postvectis 2485:2002InvRD..41...83L 2364:2019Ethol.125..587G 2080:2011BMCEE..11..295W 1994:An illustration by 1945:) being eaten by a 1205:Galaxias postvectis 1093:Pleurotus ostreatus 783:Fremont, California 64:Various species of 3305:Megaselia aequalis 2915:Journal of Zoology 2746:South, A. (1992). 2000:bicolored antpitta 1979:Limax cinereoniger 1854:diatomaceous earth 1762:agonistic behavior 1732: 1720: 1585:Angiostoma limacis 1530:Arianta arbustorum 1244:Thamnophis elegans 1154:agricultural pests 1123:Stemonitis axifera 979: 936: 876: 868: 856: 786: 663: 225: 34:. For snails, see 3891:Gastropod anatomy 3820:Media related to 3759:978-0-12-814488-6 3423:978-3-490-17918-0 3064:Molecular Ecology 3025:Carabus violaceus 2962:on 25 April 2013. 2842:10.1242/jeb.02366 2835:(16): 3164–3169. 2704:Arion lusitanicus 2469:A. dolichophallus 2372:10.1111/eth.12875 1715: 1566:Tandonia sowerbyi 1515:. The slug mite, 1482:Carabus violaceus 1347:Corvus frugilegus 1322:common blackbirds 1080:) and mushrooms ( 1027:tuberous begonias 950:. Some slugs are 638: 629: 611: 604:Strophocheiloidea 599: 587: 575: 558: 549: 540: 531: 530:Moellendorf, 1900 522: 513: 504: 492: 491:Bourguignat, 1877 483: 471: 462: 453: 441: 409:Athoracophoroidea 404: 392: 383: 374: 357: 348: 339: 330: 212: 200: 199: 16:(Redirected from 3908: 3864:Slugs of Florida 3850: 3848: 3846: 3841:on 13 April 2010 3837:. Archived from 3819: 3805: 3780: 3771: 3733: 3727: 3726: 3724: 3722: 3711: 3705: 3704: 3702: 3700: 3686: 3680: 3679: 3677: 3675: 3661: 3655: 3654: 3652: 3650: 3639: 3633: 3632: 3630: 3628: 3613: 3607: 3606: 3604: 3602: 3591: 3585: 3584: 3566: 3560: 3559: 3557: 3555: 3539: 3533: 3520: 3514: 3513: 3511: 3509: 3498: 3492: 3491: 3471: 3465: 3464: 3454: 3434: 3428: 3427: 3409: 3403: 3401: 3395: 3390: 3388: 3380: 3379:on 19 July 2013. 3378: 3371: 3363: 3354: 3353: 3331: 3325: 3324: 3315:(6): 1587–1594. 3300: 3294: 3274: 3268: 3245: 3236: 3235: 3225: 3197: 3188: 3187: 3185: 3179:. Archived from 3162: 3149: 3140: 3139: 3113: 3104: 3103: 3070:(9): 1279–1292. 3058: 3052: 3051: 3049: 3047: 3018: 3012: 3011: 3009: 3007: 2996: 2990: 2989: 2987: 2985: 2970: 2964: 2963: 2961: 2946: 2937: 2931: 2930: 2910: 2904: 2903: 2893: 2861: 2855: 2854: 2844: 2820: 2814: 2813: 2812: 2806: 2804: 2768: 2762: 2761: 2743: 2722: 2721: 2719: 2717: 2697: 2688: 2687: 2685: 2683: 2669: 2663: 2662: 2660: 2658: 2637: 2631: 2630: 2628: 2626: 2583: 2568: 2567: 2565: 2563: 2558:. animals.mom.me 2552: 2541: 2540: 2534: 2526: 2524: 2522: 2511: 2505: 2504: 2460: 2454: 2453: 2451: 2449: 2428: 2419: 2413: 2412: 2410: 2408: 2403:on 13 April 2008 2393: 2384: 2383: 2343: 2334: 2333: 2331: 2329: 2313: 2304: 2303: 2290: 2284: 2283: 2255: 2249: 2248: 2228: 2222: 2221: 2209: 2200: 2199: 2163: 2157: 2156: 2138: 2125: 2112: 2111: 2101: 2091: 2059: 2053: 2052: 2037: 2002:catching a slug. 1991: 1974: 1958: 1942:Amanita muscaria 1932: 1910: 1894: 1851: 1716: 1633:Insects such as 1337:Sturnus vulgaris 1314:Zootoca vivipara 931:Large slug near 682:stylommatophoran 636: 624: 606: 594: 582: 573: 556: 547: 538: 529: 527:Plectopylidoidea 520: 511: 499: 490: 478: 469: 460: 448: 439: 415:Athoracophoridae 403:in Turnton, 1840 399: 390: 381: 372: 355: 346: 337: 328: 213: 174:Groups included 129: 128: 98: 59: 47: 46: 21: 3916: 3915: 3911: 3910: 3909: 3907: 3906: 3905: 3886:Stylommatophora 3876: 3875: 3844: 3842: 3833: 3812: 3788: 3786:Further reading 3783: 3760: 3734: 3730: 3720: 3718: 3713: 3712: 3708: 3698: 3696: 3688: 3687: 3683: 3673: 3671: 3663: 3662: 3658: 3648: 3646: 3641: 3640: 3636: 3626: 3624: 3615: 3614: 3610: 3600: 3598: 3593: 3592: 3588: 3567: 3563: 3553: 3551: 3541: 3540: 3536: 3531:Wayback Machine 3521: 3517: 3507: 3505: 3500: 3499: 3495: 3488:10.1139/z79-104 3472: 3468: 3452:10.1.1.873.8562 3435: 3431: 3424: 3410: 3406: 3393: 3391: 3382: 3381: 3376: 3369: 3365: 3364: 3357: 3350: 3332: 3328: 3301: 3297: 3275: 3271: 3256:Wayback Machine 3246: 3239: 3198: 3191: 3186:on 5 July 2008. 3183: 3160: 3154:"The slug mite 3150: 3143: 3114: 3107: 3059: 3055: 3045: 3043: 3041: 3019: 3015: 3005: 3003: 2998: 2997: 2993: 2983: 2981: 2980:on 2 April 2013 2972: 2971: 2967: 2959: 2944: 2938: 2934: 2911: 2907: 2862: 2858: 2821: 2817: 2807: 2769: 2765: 2758: 2744: 2725: 2715: 2713: 2698: 2691: 2681: 2679: 2671: 2670: 2666: 2656: 2654: 2639: 2638: 2634: 2624: 2622: 2604:10.2307/3761690 2584: 2571: 2561: 2559: 2554: 2553: 2544: 2528: 2527: 2520: 2518: 2513: 2512: 2508: 2461: 2457: 2447: 2445: 2426: 2420: 2416: 2406: 2404: 2395: 2394: 2387: 2344: 2337: 2327: 2325: 2314: 2307: 2291: 2287: 2256: 2252: 2229: 2225: 2210: 2203: 2164: 2160: 2136: 2126: 2115: 2060: 2056: 2039: 2038: 2031: 2027: 2010: 2003: 1992: 1983: 1975: 1966: 1959: 1950: 1933: 1924: 1911: 1902: 1895: 1886: 1845: 1822: 1816: 1786: 1784:Human relevance 1705: 1703: 1649:) in the genus 1605:Angiostrongylus 1590:salivary glands 1542:Arion hortensis 1501: 1473: 1427:Arion hortensis 1357:Corvus monedula 1296:Anguis fragilis 1290:Duberria lutrix 1284:Dipsas neuwiedi 1272:Storeria dekayi 1162: 1142: 925: 909: 843: 769: 744:Vestigial shell 655: 488:Helixarionoidea 467:Gastrodontoidea 326:Achatinelloidea 288: 202: 123: 43: 28: 23: 22: 15: 12: 11: 5: 3914: 3904: 3903: 3898: 3893: 3888: 3874: 3873: 3861: 3852: 3831: 3825: 3811: 3810:External links 3808: 3807: 3806: 3787: 3784: 3782: 3781: 3758: 3750:Academic Press 3740:(2 ed.). 3728: 3706: 3681: 3656: 3634: 3608: 3586: 3561: 3534: 3515: 3493: 3482:(4): 846–855. 3466: 3429: 3422: 3404: 3394:|journal= 3355: 3349:978-0851993195 3348: 3326: 3295: 3269: 3237: 3189: 3171:(4): 441–443. 3141: 3130:(4): 521–530. 3105: 3053: 3039: 3013: 2991: 2974:"Slug Control" 2965: 2932: 2921:(2): 115–121. 2905: 2884:(1): 211–213. 2856: 2815: 2787:(2): 257–269. 2763: 2757:978-0412368103 2756: 2723: 2689: 2664: 2653:. 10 July 2008 2632: 2598:(5): 757–760. 2569: 2542: 2506: 2479:(1–3): 83–93. 2455: 2414: 2385: 2358:(8): 587–591. 2335: 2305: 2285: 2250: 2223: 2201: 2174:(4): 461–469. 2158: 2147:(1): 375–393. 2113: 2054: 2028: 2026: 2023: 2022: 2021: 2016: 2009: 2006: 2005: 2004: 1993: 1986: 1984: 1976: 1969: 1967: 1960: 1953: 1951: 1934: 1927: 1925: 1912: 1905: 1903: 1896: 1889: 1885: 1882: 1858:coffee grounds 1826:iron phosphate 1818:Main article: 1815: 1812: 1785: 1782: 1702: 1699: 1679:Agriolimacidae 1500: 1497: 1472: 1469: 1302:Tiliqua rugosa 1161: 1158: 1141: 1138: 1099:Boletus edulis 1082:basidiomycetes 995:chrysanthemums 924: 923:Feeding habits 921: 908: 905: 880:hermaphrodites 842: 839: 768: 765: 764: 763: 760:Veronicellidae 741: 727: 715: 709: 689: 678:sense of smell 654: 651: 650: 649: 648: 647: 646: 645: 630: 618: 616:Parmacelloidea 612: 600: 588: 580:Staffordioidea 576: 567: 566: 565: 550: 541: 532: 523: 514: 505: 493: 484: 472: 463: 454: 442: 433: 432: 431: 419: 418: 417: 405: 393: 384: 382:Swainson, 1840 375: 362:Subinfraorder 360: 359: 358: 349: 340: 335:Cochlicopoidea 331: 318:Subinfraorder 308:opisthobranchs 287: 284: 198: 197: 196: 195: 189: 184: 176: 175: 171: 170: 165: 161: 160: 155: 151: 150: 145: 141: 140: 135: 131: 130: 117: 116: 100: 99: 91: 90: 61: 60: 52: 51: 26: 9: 6: 4: 3: 2: 3913: 3902: 3899: 3897: 3894: 3892: 3889: 3887: 3884: 3883: 3881: 3872: 3869: 3865: 3862: 3860: 3856: 3853: 3840: 3836: 3832: 3829: 3826: 3823: 3818: 3814: 3813: 3803: 3799: 3795: 3790: 3789: 3778: 3777:9780128144893 3774: 3769: 3765: 3761: 3755: 3751: 3747: 3743: 3739: 3732: 3716: 3710: 3695: 3691: 3685: 3670: 3666: 3660: 3644: 3638: 3622: 3618: 3612: 3596: 3590: 3582: 3581: 3576: 3574: 3565: 3550: 3549: 3544: 3538: 3532: 3528: 3525: 3519: 3503: 3497: 3489: 3485: 3481: 3477: 3470: 3462: 3458: 3453: 3448: 3444: 3440: 3433: 3425: 3419: 3415: 3408: 3399: 3386: 3375: 3368: 3362: 3360: 3351: 3345: 3341: 3337: 3330: 3322: 3318: 3314: 3310: 3306: 3299: 3292: 3288: 3284: 3283: 3278: 3273: 3267:(8): 430–431. 3266: 3263: 3262: 3257: 3253: 3250: 3244: 3242: 3233: 3229: 3224: 3219: 3215: 3211: 3207: 3205: 3196: 3194: 3182: 3178: 3174: 3170: 3166: 3159: 3157: 3148: 3146: 3137: 3133: 3129: 3125: 3124: 3119: 3112: 3110: 3101: 3097: 3093: 3089: 3085: 3081: 3077: 3073: 3069: 3065: 3057: 3042: 3040:9789546421005 3036: 3032: 3028: 3026: 3017: 3002:. thenest.com 3001: 2995: 2979: 2975: 2969: 2958: 2954: 2950: 2943: 2936: 2928: 2924: 2920: 2916: 2909: 2901: 2897: 2892: 2887: 2883: 2879: 2875: 2873: 2869: 2860: 2852: 2848: 2843: 2838: 2834: 2830: 2826: 2819: 2811: 2803: 2798: 2794: 2790: 2786: 2782: 2778: 2776: 2767: 2759: 2753: 2749: 2742: 2740: 2738: 2736: 2734: 2732: 2730: 2728: 2711: 2707: 2705: 2696: 2694: 2678: 2674: 2668: 2652: 2651: 2646: 2644: 2636: 2621: 2617: 2613: 2609: 2605: 2601: 2597: 2593: 2589: 2582: 2580: 2578: 2576: 2574: 2557: 2551: 2549: 2547: 2538: 2532: 2516: 2510: 2502: 2498: 2494: 2490: 2486: 2482: 2478: 2474: 2470: 2466: 2459: 2444: 2440: 2436: 2432: 2425: 2418: 2402: 2398: 2392: 2390: 2381: 2377: 2373: 2369: 2365: 2361: 2357: 2353: 2349: 2342: 2340: 2323: 2319: 2312: 2310: 2302:(3): 163–170. 2301: 2297: 2289: 2281: 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1599:Limax maximus 1595: 1591: 1587: 1586: 1581: 1580: 1579:Agfa flexilis 1574: 1572: 1568: 1567: 1562: 1561: 1560:Milax gagates 1556: 1555: 1550: 1549: 1548:Limax maximus 1544: 1543: 1538: 1537: 1532: 1531: 1526: 1525: 1520: 1519: 1514: 1510: 1506: 1496: 1494: 1490: 1489: 1484: 1483: 1478: 1471:Invertebrates 1468: 1466: 1462: 1458: 1453: 1451: 1450:charadriiform 1447: 1443: 1442: 1437: 1433: 1429: 1428: 1423: 1422: 1417: 1413: 1412: 1407: 1406: 1401: 1397: 1396: 1391: 1390: 1385: 1381: 1380: 1375: 1371: 1367: 1363: 1359: 1358: 1353: 1349: 1348: 1343: 1339: 1338: 1333: 1329: 1328: 1327:Turdus merula 1323: 1318: 1316: 1315: 1310: 1309: 1304: 1303: 1298: 1297: 1292: 1291: 1286: 1285: 1280: 1279: 1274: 1273: 1268: 1267: 1262: 1261: 1256: 1252: 1251: 1246: 1245: 1240: 1236: 1232: 1227: 1225: 1224: 1219: 1218: 1213: 1212: 1207: 1206: 1201: 1200: 1195: 1194: 1189: 1188: 1183: 1179: 1175: 1171: 1167: 1157: 1155: 1151: 1150:invertebrates 1147: 1137: 1135: 1131: 1130: 1125: 1124: 1119: 1118: 1113: 1112: 1111:Pleurocybella 1107: 1106: 1101: 1100: 1095: 1094: 1089: 1088: 1083: 1079: 1075: 1074: 1069: 1068: 1067:Ariolimacidae 1063: 1062: 1057: 1056: 1051: 1047: 1042: 1040: 1036: 1032: 1028: 1024: 1020: 1016: 1012: 1008: 1004: 1000: 996: 992: 988: 984: 977: 973: 972: 967: 963: 961: 957: 953: 949: 945: 941: 934: 933:Manali, India 929: 920: 918: 914: 904: 901: 899: 898: 893: 892: 887: 886:Apophallation 883: 881: 872: 865: 860: 853: 852: 847: 838: 836: 835: 834:Limax maximus 829: 824: 822: 818: 813: 811: 807: 803: 799: 795: 791: 784: 780: 779: 773: 761: 757: 753: 749: 745: 742: 739: 735: 731: 728: 725: 724: 719: 716: 713: 710: 707: 704:opening, the 703: 699: 698: 693: 690: 687: 683: 679: 675: 671: 668: 667: 666: 659: 644: 640: 639: 635: 631: 627: 623: 619: 617: 613: 609: 605: 601: 597: 593: 592:Streptaxoidea 589: 585: 581: 577: 574:Pilsbry, 1895 572: 568: 564: 560: 559: 557:Pilsbry, 1893 555: 551: 546: 542: 539:Pilsbry, 1894 537: 533: 528: 524: 519: 518:Orthalicoidea 515: 510: 506: 502: 498: 494: 489: 485: 481: 477: 473: 468: 464: 459: 455: 451: 447: 446:Clausilioidea 443: 440:Pilsbry, 1895 438: 434: 430: 426: 425: 424: 423:Orthalicoidea 420: 416: 412: 411: 410: 406: 402: 398: 394: 389: 385: 380: 376: 373:Pilsbry, 1895 371: 367: 366: 365: 361: 354: 350: 347:Pilsbry, 1900 345: 341: 338:Pilsbry, 1900 336: 332: 327: 323: 322: 321: 317: 316: 315: 313: 309: 305: 301: 297: 293: 283: 281: 277: 272: 267: 265: 264: 259: 255: 251: 247: 244: 241: 237: 233: 229: 222: 218: 194: 190: 188: 185: 183: 180: 179: 177: 172: 169: 166: 163: 162: 159: 156: 153: 152: 149: 146: 143: 142: 139: 136: 133: 132: 127: 122: 118: 115: 111: 107: 106: 101: 97: 92: 89: 88: 83: 82: 81:Limax maximus 77: 73: 72: 67: 62: 58: 53: 48: 45: 41: 37: 33: 19: 3843:. Retrieved 3839:the original 3801: 3797: 3737: 3731: 3719:. Retrieved 3709: 3697:. Retrieved 3694:Slug Cuisine 3693: 3684: 3672:. Retrieved 3669:Slug Cuisine 3668: 3659: 3647:. Retrieved 3637: 3625:. Retrieved 3621:the original 3611: 3599:. Retrieved 3589: 3578: 3572: 3564: 3552:. Retrieved 3546: 3537: 3518: 3506:. Retrieved 3496: 3479: 3475: 3469: 3445:(1): 19–23. 3442: 3438: 3432: 3413: 3407: 3385:cite journal 3374:the original 3339: 3329: 3312: 3308: 3304: 3298: 3281: 3277:Taylor J. W. 3272: 3264: 3259: 3216:(3): 487–9. 3213: 3209: 3203: 3181:the original 3168: 3164: 3155: 3127: 3121: 3117: 3067: 3063: 3056: 3044:. Retrieved 3030: 3024: 3016: 3004:. Retrieved 2994: 2982:. Retrieved 2978:the original 2968: 2957:the original 2952: 2948: 2935: 2918: 2914: 2908: 2881: 2877: 2871: 2867: 2859: 2832: 2828: 2818: 2784: 2780: 2774: 2766: 2747: 2714:. Retrieved 2709: 2703: 2680:. Retrieved 2677:Slug Cuisine 2676: 2667: 2655:. Retrieved 2648: 2642: 2635: 2623:. Retrieved 2595: 2591: 2560:. 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Retrieved 2321: 2299: 2295: 2288: 2263: 2259: 2253: 2244: 2240: 2234: 2233:"Estudio de 2226: 2217: 2213: 2171: 2167: 2161: 2144: 2140: 2131: 2071: 2067: 2057: 2048: 2044: 1977: 1940: 1918: 1897: 1840: 1833: 1823: 1820:Molluscicide 1808:banana slugs 1793: 1791: 1787: 1766:conspecifics 1759: 1740: 1733: 1692: 1682: 1672: 1665:Philomycidae 1650: 1632: 1625: 1615: 1609: 1603: 1597: 1583: 1577: 1575: 1570: 1564: 1558: 1552: 1546: 1540: 1534: 1528: 1522: 1516: 1502: 1486: 1480: 1474: 1454: 1441:Limax flavus 1439: 1425: 1419: 1409: 1403: 1393: 1387: 1386:(feeding on 1377: 1376:(feeding on 1355: 1345: 1335: 1325: 1319: 1312: 1306: 1300: 1294: 1288: 1282: 1276: 1270: 1264: 1258: 1248: 1242: 1228: 1221: 1215: 1209: 1203: 1197: 1191: 1187:Salmo trutta 1185: 1163: 1143: 1127: 1121: 1115: 1109: 1103: 1097: 1091: 1085: 1071: 1065: 1059: 1053: 1050:Philomycidae 1043: 1039:strawberries 980: 969: 937: 910: 902: 895: 889: 884: 877: 849: 841:Reproduction 832: 825: 814: 787: 776: 762:lack shells. 752:Philomycidae 743: 729: 721: 717: 711: 695: 691: 686:veronicellid 673: 669: 664: 632:Superfamily 620:Superfamily 614:Superfamily 602:Superfamily 590:Superfamily 578:Superfamily 571:Sagdidoidera 569:Superfamily 552:Superfamily 543:Superfamily 536:Polygyroidea 534:Superfamily 525:Superfamily 516:Superfamily 509:Oleacinoidea 507:Superfamily 495:Superfamily 486:Superfamily 474:Superfamily 465:Superfamily 456:Superfamily 444:Superfamily 435:Superfamily 421:Superfamily 407:Superfamily 395:Superfamily 386:Superfamily 379:Achatinoidea 377:Superfamily 368:Superfamily 356:Turton, 1831 351:Superfamily 342:Superfamily 333:Superfamily 329:Gulick, 1873 324:Superfamily 294:, two – the 289: 280:polyphyletic 268: 261: 249: 231: 227: 226: 103: 87:Limax flavus 85: 79: 69: 44: 3804:(2): 48–63. 3748:, CA, USA: 3627:16 November 3508:19 November 2716:14 December 2328:14 December 1996:Joseph Smit 1947:banana slug 1915:dung beetle 1899:Arion rufus 1846: [ 1830:metaldehyde 1770:seasonality 1639:parasitoids 1505:parasitised 1493:driver ants 1223:Ceratophrys 1160:Vertebrates 1146:vertebrates 1134:ontogenetic 1078:myxomycetes 976:Mexico City 946:, and even 821:hygroscopic 810:pneumostome 790:desiccation 778:Ambigolimax 756:Onchidiidae 706:pneumostome 702:respiratory 653:Description 634:Quijotoidea 554:Rhytidoidea 548:Morse, 1864 470:Tryon, 1866 437:Camaenoidea 429:Bulimulinae 391:Baker, 1960 364:Sigmurethra 353:Pupilloidea 344:Partuloidea 320:Orthurethra 304:Sigmurethra 300:Soleolifera 296:Onchidiacea 274:example of 248:. The word 240:terrestrial 236:common name 193:Sigmurethra 187:Soleolifera 182:Onchidiacea 18:Keel (slug) 3880:Categories 3845:9 February 3768:1152284558 2437:(4): 163. 2316:Nixon, P. 2247:: 103–124. 2074:(1): 295. 2051:(2): 48–63 2025:References 1963:vegetables 1874:maize/corn 1843:beer traps 1814:Prevention 1800:meningitis 1780:behavior. 1778:gregarious 1743:Kerry slug 1728:Kerry slug 1694:Sarcophaga 1637:are known 1622:meningitis 1571:R. limacum 1536:Arion ater 1503:Slugs are 1479:, such as 1446:rock doves 1432:game birds 1416:red grouse 1411:Arion ater 1374:fieldfares 1178:amphibians 1046:fungivores 1031:hollyhocks 983:vegetables 960:earthworms 878:Slugs are 775:An active 767:Physiology 643:Quijotidae 622:Zonitoidea 563:Rhytididae 545:Punctoidea 501:Rafinesque 497:Limacoidea 480:Rafinesque 476:Helicoidea 458:Dyakioidea 427:Subfamily 397:Arionoidea 388:Aillyoidea 312:Ellobiidae 258:semi-slugs 168:Gastropoda 108:sp., from 76:Kerry slug 71:Arion ater 3746:San Diego 3447:CiteSeerX 3046:15 August 3006:15 August 2984:15 August 2900:2175-7925 2657:15 August 2625:15 August 2592:Mycologia 2562:15 August 2448:15 August 2407:15 August 1798:-induced 1755:hibernate 1747:Arionidae 1736:substrate 1684:Deroceras 1674:Megaselia 1661:Limacidae 1657:Arionidae 1635:dipterans 1513:nematodes 1477:Carabidae 1465:hedgehogs 1421:Deroceras 1332:starlings 1255:congeners 1250:Ariolimax 1140:Predators 1087:Lactarius 1035:marigolds 1023:primroses 1015:narcissus 971:Lehmannia 952:predators 944:mushrooms 913:ecosystem 897:Deroceras 891:Ariolimax 851:Deroceras 796:, fallen 794:tree bark 748:Limacidae 674:tentacles 670:Tentacles 596:J.E. Gray 401:J.E. Gray 370:Acavoidea 292:Pulmonata 271:taxonomic 254:sea slugs 243:gastropod 232:land slug 144:Kingdom: 138:Eukaryota 110:Vancouver 3649:8 August 3554:15 March 3527:Archived 3289:, pages 3287:page XV. 3252:Archived 3100:23921693 3092:10972768 2878:Biotemas 2851:16888064 2650:BBC News 2620:21156549 2531:cite web 2501:83829239 2380:92677691 2352:Ethology 2266:: 1–26. 2220:: 29–35. 2196:87142440 2188:30155937 2108:21985526 2019:Sea slug 2008:See also 1937:mushroom 1774:climatic 1751:autotomy 1701:Behavior 1689:Muscidae 1669:Phoridae 1643:ontogeny 1436:wrynecks 1400:thrushes 1384:redwings 1366:vultures 1352:jackdaws 1239:colubrid 1217:Rhinella 1166:reptiles 1136:stages. 1105:Agaricus 1019:gentians 991:petunias 802:planters 781:slug in 286:Taxonomy 269:Various 217:Kanagawa 158:Mollusca 154:Phylum: 148:Animalia 134:Domain: 32:Sea slug 3798:Tuatara 3721:2 March 3699:25 July 3674:25 July 3601:2 March 3232:8107609 3072:Bibcode 2789:Bibcode 2682:25 July 2612:3761690 2521:12 July 2481:Bibcode 2360:Bibcode 2318:"Slugs" 2296:Veliger 2280:4821033 2260:Zootaxa 2099:3198971 2076:Bibcode 2045:Tuatara 1884:Gallery 1878:soybean 1862:osmosis 1804:lettuce 1652:Melinda 1461:badgers 1237:. Some 1235:lizards 1199:arionid 1174:mammals 1117:Russula 1052:(e. g. 1011:dahlias 1003:lobelia 999:daisies 948:carrion 940:lichens 907:Ecology 806:torsion 641:Family 561:Family 413:Family 246:mollusc 234:, is a 164:Class: 66:British 3775:  3766:  3756:  3742:London 3449:  3420:  3346:  3230:  3098:  3090:  3037:  2898:  2849:  2754:  2618:  2610:  2499:  2378:  2278:  2194:  2186:  2106:  2096:  1594:rectum 1563:, and 1281:(e.g. 1278:Dipsas 1231:snakes 1064:) and 1007:lilies 956:snails 864:mucous 697:mantle 692:Mantle 688:slugs. 628:, 1864 610:, 1926 608:Thiele 598:, 1806 586:, 1931 584:Thiele 503:, 1815 482:, 1815 452:, 1864 3402:9 pp. 3377:(PDF) 3370:(PDF) 3184:(PDF) 3161:(PDF) 3096:S2CID 2960:(PDF) 2945:(PDF) 2643:video 2608:JSTOR 2497:S2CID 2427:(PDF) 2376:S2CID 2276:S2CID 2192:S2CID 2184:JSTOR 2137:(PDF) 2014:Snail 1998:of a 1850:] 1509:acari 1457:foxes 1405:Limax 1395:Arion 1389:Limax 1370:ducks 1342:rooks 1196:, an 1170:birds 987:herbs 958:, or 917:fungi 828:slime 826:The " 817:mucus 738:mucus 626:Mörch 450:Mörch 263:snail 230:, or 221:Japan 105:Arion 50:Slug 36:Snail 3871:IFAS 3847:2010 3822:Slug 3773:ISBN 3764:OCLC 3754:ISBN 3723:2014 3701:2024 3676:2024 3651:2016 3629:2012 3603:2014 3556:2006 3510:2012 3418:ISBN 3398:help 3344:ISBN 3293:–52. 3228:PMID 3088:PMID 3048:2016 3035:ISBN 3008:2016 2986:2016 2896:ISSN 2847:PMID 2752:ISBN 2718:2012 2684:2024 2659:2016 2627:2016 2616:PMID 2564:2016 2537:link 2523:2023 2467:and 2450:2016 2409:2016 2330:2012 2264:2068 2104:PMID 1876:and 1663:and 1620:, a 1614:and 1592:and 1582:and 1485:and 1463:and 1448:and 1438:(on 1424:and 1418:(on 1408:and 1402:(on 1392:and 1368:and 1362:owls 1233:and 1220:and 1211:Bufo 1182:fish 1180:and 1148:and 1126:and 1114:and 1058:and 985:and 798:logs 758:and 730:Foot 718:Keel 712:Tail 298:and 256:and 250:slug 228:Slug 84:and 3484:doi 3457:doi 3317:doi 3265:179 3218:doi 3173:doi 3132:doi 3080:doi 2923:doi 2919:149 2886:doi 2837:doi 2833:209 2797:doi 2600:doi 2489:doi 2439:doi 2368:doi 2356:125 2268:doi 2176:doi 2149:doi 2094:PMC 2084:doi 1596:of 1444:), 1430:), 1414:), 1398:), 1382:), 1360:), 1350:), 1340:), 1330:), 1317:). 3882:: 3866:. 3802:25 3800:. 3796:. 3762:. 3752:. 3744:, 3692:. 3667:. 3577:. 3545:. 3480:57 3478:. 3455:. 3443:60 3441:. 3389:: 3387:}} 3383:{{ 3358:^ 3313:61 3311:. 3291:34 3258:. 3240:^ 3226:. 3214:88 3212:. 3208:. 3192:^ 3167:. 3163:. 3144:^ 3126:. 3108:^ 3094:. 3086:. 3078:. 3066:. 3029:. 2951:. 2947:. 2917:. 2894:. 2882:25 2880:. 2876:. 2845:. 2831:. 2827:. 2795:. 2785:30 2783:. 2779:. 2726:^ 2708:. 2692:^ 2675:. 2647:. 2645:)" 2614:. 2606:. 2596:94 2594:. 2590:. 2572:^ 2545:^ 2533:}} 2529:{{ 2495:. 2487:. 2477:41 2475:. 2435:17 2433:. 2429:. 2388:^ 2374:. 2366:. 2354:. 2350:. 2338:^ 2320:. 2308:^ 2300:50 2298:. 2274:. 2262:. 2245:25 2243:. 2239:. 2218:60 2216:. 2204:^ 2190:. 2182:. 2172:52 2170:. 2145:88 2143:. 2139:. 2116:^ 2102:. 2092:. 2082:. 2072:11 2070:. 2066:. 2049:25 2047:, 2043:, 2032:^ 1935:A 1913:A 1852:, 1848:de 1659:, 1630:. 1569:. 1557:, 1551:, 1545:, 1539:, 1533:, 1527:, 1467:. 1459:, 1434:, 1364:, 1226:. 1214:, 1176:, 1172:, 1168:, 1108:, 1033:, 1029:, 1025:, 1021:, 1017:, 1013:, 1009:, 1005:, 1001:, 997:, 993:, 962:. 942:, 754:, 282:. 219:, 114:BC 112:, 78:, 74:, 3849:. 3779:. 3770:. 3725:. 3703:. 3678:. 3653:. 3631:. 3605:. 3583:. 3575:" 3571:" 3558:. 3512:. 3490:. 3486:: 3463:. 3459:: 3426:. 3400:) 3396:( 3352:. 3323:. 3319:: 3234:. 3220:: 3175:: 3169:6 3138:. 3134:: 3128:4 3102:. 3082:: 3074:: 3068:9 3050:. 3010:. 2988:. 2953:1 2929:. 2925:: 2902:. 2888:: 2866:" 2853:. 2839:: 2805:. 2799:: 2791:: 2760:. 2720:. 2706:" 2702:" 2686:. 2661:. 2629:. 2602:: 2566:. 2539:) 2525:. 2503:. 2491:: 2483:: 2452:. 2441:: 2411:. 2382:. 2370:: 2362:: 2332:. 2282:. 2270:: 2198:. 2178:: 2155:. 2151:: 2134:" 2110:. 2086:: 2078:: 1965:. 1949:. 1939:( 1917:( 1354:( 1344:( 1334:( 1324:( 1070:( 223:. 42:. 20:)

Index

Keel (slug)
Sea slug
Snail
Slug (disambiguation)
Various species of British land slugs, including (from the top) the larger drawings: Arion ater, Kerry slug, Limax maximus and Limax flavus
British
Arion ater
Kerry slug
Limax maximus
Limax flavus
Arion sp., from Vancouver, BC
Arion
Vancouver
BC
Scientific classification
Edit this classification
Eukaryota
Animalia
Mollusca
Gastropoda
Onchidiacea
Soleolifera
Sigmurethra
Kanagawa
Japan
common name
terrestrial
gastropod
mollusc
sea slugs

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