31:
75:
50:
306:
627:
197:
415:. The bird's droppings were such an important source of fertilizer to the peoples of the Andes that it was protected by Inca rulers, who supposedly made disturbing the cormorants in any way punishable by death. The common name is an adaptation of the South American Spanish
329:. It has roseate feet. The head, neck and back are black as are the outer parts of the thighs. The throat patch, breast and belly are white. In breeding plumage it has a few white feathers on the sides of the head and neck.
381:
and degradation and over-fishing have resulted in a steady decline of the population of about 30% from an estimated figure of three million birds in 1984. This species is listed as
1589:
1629:
1509:
631:
1483:
1496:
393:
1624:
1501:
659:
1448:
340:
on flat surfaces on offshore islands or remote headlands. There are up to three nests per square meter in high-density
1553:
591:
1599:
1514:
412:
1619:
1462:
1457:
464:
186:
74:
1614:
1102:
486:
Cullen, David J. (1988). "Mineralogy of nitrogenous guano on the Bounty
Islands, SW Pacific Ocean".
325:
to the end of the tail. Its bill is grayish with some red at the base. The face is red with a green
1237:
1132:
652:
1604:
745:
1609:
1410:
1112:
1318:
1199:
1173:
1107:
755:
750:
169:
1548:
1287:
1247:
1217:
637:
495:
1566:
1540:
8:
1594:
1242:
1209:
1008:
912:
645:
441:
360:
39:
499:
1385:
1137:
765:
507:
352:
209:
69:
1535:
1470:
1360:
1204:
1158:
1082:
882:
839:
760:
587:
299:
1475:
1163:
1117:
1035:
939:
859:
792:
690:
503:
459:
368:
252:
1522:
1433:
1168:
1127:
998:
902:
668:
382:
54:
30:
1380:
1375:
1370:
1178:
1122:
1087:
1003:
844:
146:
1583:
1354:
1344:
1339:
1267:
1073:
450:
404:
256:
59:
1334:
1324:
1278:
1097:
849:
521:
378:
282:.) After breeding it spreads south to southern parts of Chile and north to
1527:
1442:
1365:
1349:
1228:
907:
864:
854:
736:
686:
341:
196:
1561:
1488:
1329:
1298:
1149:
672:
398:
279:
126:
685:
295:
1307:
1092:
1031:
581:
333:
332:
Breeding occurs year round with a peak in
November and December. The
314:
272:
268:
249:
136:
86:
1404:
1427:
345:
326:
305:
291:
275:
106:
1258:
1059:
993:
970:
816:
788:
722:
283:
934:
984:
935:
893:
626:
287:
96:
294:– probably a result of mass dispersal due to food shortage in
1190:
1048:
959:
943:
830:
805:
711:
694:
372:
337:
264:
667:
433:
386:
322:
260:
116:
1030:
546:
787:
564:
321:, but larger, measuring 78 cm from the tip of the
601:
439:
411:
The scientific name commemorates the French explorer
313:
The guanay cormorant is similar in coloration to the
1214:
348:
of approximately 63 × 40 mm in size.
465:10.2305/IUCN.UK.2018-2.RLTS.T22696810A133553624.en
1581:
582:de la Peña, Martín R.; Rumboll, Maurice (1998).
371:. The guanay cormorant is the main producer of
584:Birds of Southern South America and Antarctica
551:Species factsheet: Phalacrocorax bougainvillii
653:
286:, and has also been recorded as far north as
302:include shallow seawater and rocky shores.
660:
646:
392:Some taxonomic authorities, including the
195:
48:
29:
463:
344:. The guanay cormorant lays two or three
204:Distribution map of the guanay cormorant
304:
1582:
485:
1590:IUCN Red List near threatened species
1409:
1408:
1029:
933:
786:
684:
641:
419:, a plural of the English equivalent
1630:Western South American coastal birds
1357:(imperial cormorant, blue-eyed shag)
606:Global Register of Migratory Species
569:Red List of Threatened Species, IUCN
586:. London: HarperCollinsPublishers.
451:IUCN Red List of Threatened Species
394:International Ornithologists' Union
13:
508:10.1111/j.1365-3091.1988.tb00995.x
396:, place this species in the genus
14:
1641:
619:
625:
73:
440:BirdLife International (2018).
402:. Others place it in the genus
514:
479:
222:Phalacrocorax bougainvilliorum
1:
426:
413:Louis Antoine de Bougainville
756:Christmas Island frigatebird
7:
1057:
968:
814:
720:
444:Leucocarbo bougainvilliorum
367:, which thrive in the cold
309:Immatures in Pucusana, Peru
245:Leucocarbo bougainvilliorum
218:Phalacrocorax bougainvillii
179:Leucocarbo bougainvilliorum
10:
1646:
1625:Taxa named by René Lesson
1417:
1296:
1276:
1256:
1226:
1188:
1147:
1071:
1056:
1047:
1043:
1025:
982:
967:
958:
954:
929:
891:
873:
828:
813:
804:
800:
782:
734:
719:
710:
706:
680:
458:: e.T22696810A133553624.
215:
208:
203:
194:
175:
168:
70:Scientific classification
68:
46:
37:
28:
23:
1567:Leucocarbo-bougainvillii
1449:Leucocarbo bougainvillii
1419:Leucocarbo bougainvillii
1238:Double-crested cormorant
1133:White-breasted cormorant
632:Leucocarbo bougainvillii
547:"BirdLife International"
226:Leucocarbo bougainvillii
161:L. bougainvilliorum
1600:Birds described in 1837
1181:(long-tailed cormorant)
746:Magnificent frigatebird
351:It feeds mainly on the
319:Leucocarbo magellanicus
1310:(Magellanic cormorant)
1140:(Temminck's cormorant)
1113:Little black cormorant
675:(Phalacrocoraciformes)
310:
1319:New Zealand king shag
1174:Little pied cormorant
1108:Black-faced cormorant
751:Ascension frigatebird
308:
248:) is a member of the
1288:Red-legged cormorant
1248:Flightless cormorant
1218:Spectacled cormorant
1038:: Phalacrocoracidae)
942:: Anhingidae ·
693:: Fregatidae ·
634:at Wikimedia Commons
278:coast appears to be
1243:Neotropic cormorant
1210:Red-faced cormorant
1009:Australasian darter
913:Australasian gannet
789:Gannets and boobies
500:1988Sedim..35..421C
361:Peruvian silverside
40:Conservation status
1620:Birds of Patagonia
1386:South Georgia shag
1321:(rough-faced shag)
1200:Brandt's cormorant
1138:Japanese cormorant
766:Lesser frigatebird
353:Peruvian anchoveta
311:
271:population on the
1577:
1576:
1536:Open Tree of Life
1411:Taxon identifiers
1402:
1401:
1398:
1397:
1394:
1393:
1361:Heard Island shag
1205:Pelagic cormorant
1159:Crowned cormorant
1083:Socotra cormorant
1067:
1066:
1021:
1020:
1017:
1016:
978:
977:
925:
924:
921:
920:
840:Blue-footed booby
824:
823:
778:
777:
774:
773:
761:Great frigatebird
730:
729:
630:Media related to
365:Odontesthes regia
357:Engraulis ringens
298:years. Its major
232:
231:
137:Phalacrocoracidae
63:
24:Guanay cormorant
1637:
1615:Birds of Ecuador
1570:
1569:
1557:
1556:
1544:
1543:
1531:
1530:
1518:
1517:
1505:
1504:
1492:
1491:
1479:
1478:
1466:
1465:
1453:
1452:
1451:
1438:
1437:
1436:
1406:
1405:
1314:Guanay cormorant
1216:
1164:Little cormorant
1118:Indian cormorant
1054:
1053:
1045:
1044:
1027:
1026:
965:
964:
956:
955:
931:
930:
860:Red-footed booby
811:
810:
802:
801:
784:
783:
717:
716:
708:
707:
682:
681:
662:
655:
648:
639:
638:
629:
615:
613:
612:
597:
578:
576:
575:
560:
558:
557:
538:
537:
535:
533:
518:
512:
511:
483:
477:
476:
474:
472:
467:
437:
369:Humboldt Current
236:guanay cormorant
199:
181:
78:
77:
57:
52:
51:
33:
21:
20:
1645:
1644:
1640:
1639:
1638:
1636:
1635:
1634:
1580:
1579:
1578:
1573:
1565:
1560:
1552:
1547:
1539:
1534:
1526:
1523:Observation.org
1521:
1513:
1508:
1500:
1495:
1487:
1482:
1474:
1469:
1461:
1456:
1447:
1446:
1441:
1432:
1431:
1426:
1413:
1403:
1390:
1292:
1272:
1252:
1222:
1184:
1169:Pygmy cormorant
1143:
1128:Great cormorant
1063:
1039:
1013:
999:Oriental darter
974:
950:
917:
903:Northern gannet
887:
869:
820:
796:
770:
726:
702:
676:
666:
622:
610:
608:
600:
594:
573:
571:
563:
555:
553:
545:
542:
541:
531:
529:
520:
519:
515:
484:
480:
470:
468:
438:
434:
429:
383:near threatened
224:
220:
190:
183:
177:
164:
72:
64:
55:Near Threatened
53:
49:
42:
17:
16:Species of bird
12:
11:
5:
1643:
1633:
1632:
1627:
1622:
1617:
1612:
1607:
1605:Birds of Chile
1602:
1597:
1592:
1575:
1574:
1572:
1571:
1558:
1545:
1532:
1519:
1506:
1493:
1480:
1467:
1454:
1439:
1423:
1421:
1415:
1414:
1400:
1399:
1396:
1395:
1392:
1391:
1389:
1388:
1383:
1381:Macquarie shag
1378:
1376:Kerguelen shag
1373:
1371:Antarctic shag
1368:
1363:
1358:
1352:
1347:
1342:
1337:
1332:
1327:
1322:
1316:
1311:
1304:
1302:
1294:
1293:
1291:
1290:
1284:
1282:
1274:
1273:
1271:
1270:
1264:
1262:
1254:
1253:
1251:
1250:
1245:
1240:
1234:
1232:
1224:
1223:
1221:
1220:
1212:
1207:
1202:
1196:
1194:
1186:
1185:
1183:
1182:
1179:Reed cormorant
1176:
1171:
1166:
1161:
1155:
1153:
1145:
1144:
1142:
1141:
1135:
1130:
1125:
1123:Cape cormorant
1120:
1115:
1110:
1105:
1103:Pied cormorant
1100:
1095:
1090:
1088:Bank cormorant
1085:
1079:
1077:
1069:
1068:
1065:
1064:
1051:
1041:
1040:
1023:
1022:
1019:
1018:
1015:
1014:
1012:
1011:
1006:
1004:African darter
1001:
996:
990:
988:
980:
979:
976:
975:
962:
952:
951:
927:
926:
923:
922:
919:
918:
916:
915:
910:
905:
899:
897:
889:
888:
886:
885:
883:Abbott's booby
879:
877:
871:
870:
868:
867:
862:
857:
852:
847:
845:Peruvian booby
842:
836:
834:
826:
825:
822:
821:
808:
798:
797:
780:
779:
776:
775:
772:
771:
769:
768:
763:
758:
753:
748:
742:
740:
732:
731:
728:
727:
714:
704:
703:
678:
677:
665:
664:
657:
650:
642:
636:
635:
621:
620:External links
618:
617:
616:
598:
592:
579:
561:
540:
539:
526:dictionary.com
513:
494:(3): 421–428.
478:
431:
430:
428:
425:
230:
229:
213:
212:
206:
205:
201:
200:
192:
191:
184:
173:
172:
166:
165:
158:
156:
152:
151:
144:
140:
139:
134:
130:
129:
124:
120:
119:
114:
110:
109:
104:
100:
99:
94:
90:
89:
84:
80:
79:
66:
65:
47:
44:
43:
38:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
1642:
1631:
1628:
1626:
1623:
1621:
1618:
1616:
1613:
1611:
1610:Birds of Peru
1608:
1606:
1603:
1601:
1598:
1596:
1593:
1591:
1588:
1587:
1585:
1568:
1563:
1559:
1555:
1550:
1546:
1542:
1537:
1533:
1529:
1524:
1520:
1516:
1511:
1507:
1503:
1498:
1494:
1490:
1485:
1481:
1477:
1472:
1468:
1464:
1459:
1455:
1450:
1444:
1440:
1435:
1429:
1425:
1424:
1422:
1420:
1416:
1412:
1407:
1387:
1384:
1382:
1379:
1377:
1374:
1372:
1369:
1367:
1364:
1362:
1359:
1356:
1355:Imperial shag
1353:
1351:
1348:
1346:
1345:Campbell shag
1343:
1341:
1340:Auckland shag
1338:
1336:
1333:
1331:
1328:
1326:
1323:
1320:
1317:
1315:
1312:
1309:
1306:
1305:
1303:
1301:
1300:
1295:
1289:
1286:
1285:
1283:
1281:
1280:
1275:
1269:
1268:European shag
1266:
1265:
1263:
1261:
1260:
1255:
1249:
1246:
1244:
1241:
1239:
1236:
1235:
1233:
1231:
1230:
1225:
1219:
1213:
1211:
1208:
1206:
1203:
1201:
1198:
1197:
1195:
1193:
1192:
1187:
1180:
1177:
1175:
1172:
1170:
1167:
1165:
1162:
1160:
1157:
1156:
1154:
1152:
1151:
1146:
1139:
1136:
1134:
1131:
1129:
1126:
1124:
1121:
1119:
1116:
1114:
1111:
1109:
1106:
1104:
1101:
1099:
1096:
1094:
1091:
1089:
1086:
1084:
1081:
1080:
1078:
1076:
1075:
1074:Phalacrocorax
1070:
1062:
1061:
1055:
1052:
1050:
1046:
1042:
1037:
1033:
1028:
1024:
1010:
1007:
1005:
1002:
1000:
997:
995:
992:
991:
989:
987:
986:
981:
973:
972:
966:
963:
961:
957:
953:
948:
945:
941:
937:
932:
928:
914:
911:
909:
906:
904:
901:
900:
898:
896:
895:
890:
884:
881:
880:
878:
876:
872:
866:
863:
861:
858:
856:
853:
851:
848:
846:
843:
841:
838:
837:
835:
833:
832:
827:
819:
818:
812:
809:
807:
803:
799:
794:
790:
785:
781:
767:
764:
762:
759:
757:
754:
752:
749:
747:
744:
743:
741:
739:
738:
733:
725:
724:
718:
715:
713:
709:
705:
700:
696:
692:
688:
683:
679:
674:
670:
663:
658:
656:
651:
649:
644:
643:
640:
633:
628:
624:
623:
607:
603:
599:
595:
593:0-00-220077-5
589:
585:
580:
570:
566:
562:
552:
548:
544:
543:
527:
523:
517:
509:
505:
501:
497:
493:
489:
488:Sedimentology
482:
466:
461:
457:
453:
452:
447:
445:
436:
432:
424:
422:
418:
414:
409:
407:
406:
405:Phalacrocorax
401:
400:
395:
390:
388:
384:
380:
376:
374:
370:
366:
362:
358:
354:
349:
347:
343:
339:
335:
330:
328:
324:
320:
316:
307:
303:
301:
297:
293:
289:
285:
281:
277:
274:
270:
266:
263:and northern
262:
258:
255:found on the
254:
251:
247:
246:
241:
237:
228:
227:
223:
219:
214:
211:
207:
202:
198:
193:
188:
182:
180:
174:
171:
170:Binomial name
167:
163:
162:
157:
154:
153:
150:
149:
145:
142:
141:
138:
135:
132:
131:
128:
125:
122:
121:
118:
115:
112:
111:
108:
105:
102:
101:
98:
95:
92:
91:
88:
85:
82:
81:
76:
71:
67:
61:
56:
45:
41:
36:
32:
27:
22:
19:
1418:
1335:Chatham shag
1325:Foveaux shag
1313:
1297:
1279:Poikilocarbo
1277:
1257:
1227:
1189:
1148:
1098:Spotted shag
1072:
1058:
983:
969:
946:
892:
874:
850:Masked booby
829:
815:
735:
721:
698:
687:Frigatebirds
609:. Retrieved
605:
583:
572:. Retrieved
568:
554:. Retrieved
550:
530:. Retrieved
525:
516:
491:
487:
481:
469:. Retrieved
455:
449:
443:
435:
420:
416:
410:
403:
397:
391:
379:Habitat loss
377:
364:
356:
350:
336:is built of
331:
318:
312:
244:
243:
239:
235:
233:
225:
221:
217:
216:
178:
176:
160:
159:
147:
18:
1443:Wikispecies
1366:Crozet shag
1350:Bounty shag
1229:Nannopterum
908:Cape gannet
865:Brown booby
855:Nazca booby
471:12 November
269:Argentinian
240:guanay shag
1595:Leucocarbo
1584:Categories
1562:Xeno-canto
1330:Otago shag
1299:Leucocarbo
1150:Microcarbo
1032:Cormorants
795:: Sulidae)
673:Suliformes
611:2005-08-12
574:2005-08-12
565:"Red List"
556:2005-08-12
427:References
399:Leucocarbo
359:, and the
280:extirpated
273:Patagonian
187:Lesson, RP
148:Leucocarbo
127:Suliformes
1434:Q27074617
1308:Rock shag
1093:Pitt shag
315:rock shag
259:coast of
250:cormorant
155:Species:
93:Kingdom:
87:Eukaryota
1428:Wikidata
875:Papasula
522:"guanay"
342:colonies
327:eye-ring
300:habitats
292:Colombia
276:Atlantic
267:. (The
210:Synonyms
133:Family:
107:Chordata
103:Phylum:
97:Animalia
83:Domain:
60:IUCN 3.1
1489:6066498
1259:Gulosus
1060:Species
994:Anhinga
985:Anhinga
971:Species
947:Anhinga
936:Darters
817:Species
737:Fregata
723:Species
699:Fregata
602:"GROMS"
532:June 6,
496:Bibcode
296:El Niño
284:Ecuador
257:Pacific
189:, 1837)
143:Genus:
123:Order:
113:Class:
58: (
1515:146616
1502:824002
1463:803258
1036:family
940:family
793:family
691:family
590:
528:. 2023
417:guanae
288:Panama
253:family
1528:73428
1476:6PTKG
1191:Urile
1049:Genus
960:Genus
944:genus
894:Morus
806:Genus
712:Genus
695:genus
669:Order
421:guano
373:guano
338:guano
265:Chile
1554:9318
1541:9834
1510:NCBI
1497:ITIS
1484:GBIF
1458:BOLD
831:Sula
588:ISBN
534:2023
473:2021
456:2018
387:IUCN
346:eggs
334:nest
323:bill
290:and
261:Peru
234:The
117:Aves
1549:TSA
1471:CoL
504:doi
460:doi
423:.
385:by
238:or
1586::
1564::
1551::
1538::
1525::
1512::
1499::
1486::
1473::
1460::
1445::
1430::
697::
671::
604:.
567:.
549:.
524:.
502:.
492:35
490:.
454:.
448:.
408:.
389:.
375:.
363:,
355:,
317:,
1215:†
1034:(
949:)
938:(
791:(
701:)
689:(
661:e
654:t
647:v
614:.
596:.
577:.
559:.
536:.
510:.
506::
498::
475:.
462::
446:"
442:"
242:(
185:(
62:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
