31:
51:
1363:
260:
ancestors that dispersed to India during the middle Eocene, before India collided with Asia. This is thought to have occurred due to India drifting close enough to
Southeast Asia to allow for biotic exchange between both regions. As the Gecarcinucidae are a freshwater group that could not disperse
30:
265:
may have formed in the Eocene between India and
Southeast Asia, allowing for the dispersal of freshwater organisms to India while it was still isolated. Following the
256:
origins themselves (unlike other lineages that are thought to have originated in
Insular India). Divergence estimates indicate that the Gecarcinucidae originate from
266:
608:
868:
738:
678:
638:
568:
418:
348:
308:
558:
508:
358:
858:
838:
828:
708:
648:
528:
408:
1260:
908:
898:
778:
698:
668:
478:
918:
438:
368:
1286:
878:
578:
538:
748:
448:
768:
798:
598:
458:
298:
888:
728:
718:
398:
338:
318:
1234:
996:
1273:
946:
788:
1428:
961:
1278:
942:
1404:
1335:
1291:
1053:"When Indian crabs were not yet Asian - biogeographic evidence for Eocene proximity of India and Southeast Asia"
50:
1182:
1159:
1004:
969:
1187:
1026:
1433:
1340:
1150:
1423:
1397:
1317:
1112:
237:, within the Gecarcinucidae. "Family" Parathelphusidae is now considered as a junior synonym.
962:"Systema Brachyurorum: Part I. An annotated checklist of extant Brachyuran crabs of the world"
1327:
1312:
1265:
1304:
1208:
1051:
Klaus, Sebastian; Schubart, Christoph D.; Streit, Bruno; Pfenninger, Markus (2010-09-17).
8:
1438:
241:
1378:
1009:
1390:
1087:
1052:
269:, the Gecarcinucidae dispersed back into mainland Asia, and from there to Australasia.
45:
1164:
1299:
1195:
1092:
1074:
808:
1200:
1082:
1064:
978:
818:
618:
234:
198:
378:
328:
202:
167:
152:
1374:
628:
488:
388:
285:
257:
210:
994:
1417:
1135:
1078:
1069:
959:
428:
245:
142:
1096:
658:
548:
273:
92:
1174:
1247:
1144:
688:
518:
262:
132:
112:
995:
Sammy De Grave; N. Dean
Pentcheff; Shane T. Ahyong; et al. (2009).
1239:
498:
277:
214:
206:
1252:
1221:
997:"A classification of living and fossil genera of decapod crustaceans"
848:
281:
249:
218:
82:
62:
1106:
1226:
1129:
468:
253:
102:
758:
588:
1213:
72:
1050:
176:
1362:
1370:
122:
960:
Peter K. L. Ng; Danièle Guinot; Peter J. F. Davie (2008).
240:
261:via marine habitats, this indicates that temporary
1415:
233:is now demoted to the rank of subfamily, as the
1398:
1405:
1391:
29:
1086:
1068:
272:The Gecarcinucidae are thought to be the
947:Integrated Taxonomic Information System
1416:
1111:
1110:
1357:
1046:
1044:
988:
13:
1008:. Suppl. 21: 1–109. Archived from
323:Pati, Bajantri & Hegde, 2023
14:
1450:
1041:
953:
1361:
1029:. Barcode of Life Systems (BOLD)
280:(the superfamily comprising the
49:
252:, despite not being of ancient
217:, with a single genus found in
1019:
935:
853:O. K. S. Chia & Ng, 2006
633:O. K. S. Chia & Ng, 2006
1:
1429:Taxa named by Mary J. Rathbun
928:
1377:. You can help Knowledge by
205:. They are found throughout
7:
1005:Raffles Bulletin of Zoology
970:Raffles Bulletin of Zoology
553:Kumar, Raj & Ng, 2017
224:
184:
10:
1455:
1356:
873:Ng & Naiyanetr, 1993
403:S. H. Tan & Ng, 1998
1119:
1027:"Gecarcinucidae {family}"
473:Pati & Sharma, 2014
291:
182:
175:
46:Scientific classification
44:
37:
28:
23:
1070:10.1186/1471-2148-10-287
1057:BMC Evolutionary Biology
673:H. Milne Edwards, 1853
977:: 1–286. Archived from
913:Bahir & Yeo, 2007
793:Bahir & Yeo, 2007
723:Bahir & Yeo, 2007
563:Bahir & Yeo, 2007
363:Bahir & Yeo, 2007
923:Bahir & Yeo, 2007
1313:Paleobiology Database
443:Milne Edwards, 1844
803:Yeo & Ng, 2012
683:Ng & Tay, 2001
593:Ng & Tay, 2001
513:Ng & Lim, 1986
503:Yeo & Ng, 2010
393:Ng & Tay, 2001
267:India-Asia collision
242:Indian subcontinent
609:Maydelliathelphusa
1386:
1385:
1351:
1350:
1300:Open Tree of Life
1113:Taxon identifiers
924:
914:
904:
894:
884:
874:
864:
854:
844:
834:
824:
814:
809:Somanniathelphusa
804:
794:
784:
774:
764:
763:Naiyanetr, 1994
754:
744:
734:
724:
714:
704:
694:
684:
674:
664:
654:
644:
634:
624:
623:Naiyanetr, 1985
614:
604:
594:
584:
574:
564:
554:
544:
534:
524:
514:
504:
494:
484:
474:
464:
454:
444:
434:
433:Naiyanetr, 1994
424:
414:
404:
394:
384:
374:
364:
354:
344:
334:
324:
314:
304:
191:
190:
171:
1446:
1434:Decapod families
1407:
1400:
1393:
1365:
1358:
1344:
1343:
1331:
1330:
1321:
1320:
1308:
1307:
1295:
1294:
1282:
1281:
1269:
1268:
1256:
1255:
1243:
1242:
1230:
1229:
1217:
1216:
1204:
1203:
1191:
1190:
1178:
1177:
1168:
1167:
1155:
1154:
1153:
1140:
1139:
1138:
1108:
1107:
1101:
1100:
1090:
1072:
1048:
1039:
1038:
1036:
1034:
1023:
1017:
1016:
1014:
1001:
992:
986:
985:
983:
966:
957:
951:
950:
943:"Gecarcinucidae"
939:
922:
912:
902:
892:
882:
872:
869:Thaksinthelphusa
862:
852:
842:
832:
822:
819:Spiralothelphusa
812:
802:
792:
782:
772:
762:
752:
742:
739:Salangathelphusa
732:
722:
712:
702:
692:
682:
672:
662:
652:
642:
639:Nautilothelphusa
632:
622:
619:Mekhongthelphusa
612:
602:
592:
582:
572:
569:Lepidopthelphusa
562:
552:
542:
532:
522:
512:
502:
492:
482:
472:
462:
452:
442:
432:
422:
419:Cylindrotelphusa
412:
402:
392:
382:
372:
362:
352:
349:Balssiathelphusa
342:
332:
322:
312:
309:Arachnothelphusa
302:
246:island continent
235:Parathelphusinae
231:Parathelphusidae
203:freshwater crabs
166:
54:
53:
39:Thelphusa indica
33:
21:
20:
1454:
1453:
1449:
1448:
1447:
1445:
1444:
1443:
1414:
1413:
1412:
1411:
1354:
1352:
1347:
1339:
1334:
1326:
1324:
1316:
1311:
1303:
1298:
1290:
1285:
1277:
1272:
1264:
1259:
1251:
1246:
1238:
1233:
1225:
1220:
1212:
1207:
1199:
1194:
1186:
1181:
1173:
1171:
1163:
1158:
1149:
1148:
1143:
1134:
1133:
1128:
1115:
1105:
1104:
1049:
1042:
1032:
1030:
1025:
1024:
1020:
1012:
999:
993:
989:
981:
964:
958:
954:
941:
940:
936:
931:
509:Heterothelphusa
379:Ceylonthelphusa
329:Austrothelphusa
294:
258:Southeast Asian
244:when it was an
227:
165:
153:Gecarcinucoidea
48:
24:Gecarcinucidae
17:
16:Family of crabs
12:
11:
5:
1452:
1442:
1441:
1436:
1431:
1426:
1424:Gecarcinucidae
1410:
1409:
1402:
1395:
1387:
1384:
1383:
1366:
1349:
1348:
1346:
1345:
1332:
1322:
1309:
1296:
1283:
1270:
1257:
1244:
1231:
1218:
1205:
1192:
1179:
1169:
1165:Gecarcinucidae
1156:
1151:Gecarcinucidae
1141:
1125:
1123:
1121:Gecarcinucidae
1117:
1116:
1103:
1102:
1040:
1018:
1015:on 2011-06-06.
987:
984:on 2011-06-06.
952:
933:
932:
930:
927:
926:
925:
915:
905:
895:
885:
875:
865:
859:Terrathelphusa
855:
845:
839:Sundathelphusa
835:
829:Stygothelphusa
825:
815:
805:
795:
785:
775:
765:
755:
745:
735:
725:
715:
713:Alcock, 1909
709:Phricotelphusa
705:
695:
685:
675:
665:
663:Muller, 1887
655:
649:Niasathelphusa
645:
635:
629:Migmathelphusa
625:
615:
605:
595:
585:
583:Alcock, 1909
575:
573:Colosi, 1920
565:
555:
545:
535:
529:Inglethelphusa
525:
515:
505:
495:
489:Gubernatoriana
485:
483:Alcock, 1909
475:
465:
455:
445:
435:
425:
423:Alcock, 1909
415:
409:Currothelphusa
405:
395:
389:Clinothelphusa
385:
375:
373:Alcock, 1909
365:
355:
345:
335:
325:
315:
305:
293:
290:
286:Potamonautidae
226:
223:
211:Southeast Asia
195:Gecarcinucidae
189:
188:
180:
179:
173:
172:
163:Gecarcinucidae
160:
156:
155:
150:
146:
145:
140:
136:
135:
130:
126:
125:
120:
116:
115:
110:
106:
105:
100:
96:
95:
90:
86:
85:
80:
76:
75:
70:
66:
65:
60:
56:
55:
42:
41:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
1451:
1440:
1437:
1435:
1432:
1430:
1427:
1425:
1422:
1421:
1419:
1408:
1403:
1401:
1396:
1394:
1389:
1388:
1382:
1380:
1376:
1373:article is a
1372:
1367:
1364:
1360:
1359:
1355:
1342:
1337:
1333:
1329:
1323:
1319:
1314:
1310:
1306:
1301:
1297:
1293:
1288:
1284:
1280:
1275:
1271:
1267:
1262:
1258:
1254:
1249:
1245:
1241:
1236:
1232:
1228:
1223:
1219:
1215:
1210:
1206:
1202:
1197:
1193:
1189:
1184:
1180:
1176:
1170:
1166:
1161:
1157:
1152:
1146:
1142:
1137:
1131:
1127:
1126:
1124:
1122:
1118:
1114:
1109:
1098:
1094:
1089:
1084:
1080:
1076:
1071:
1066:
1062:
1058:
1054:
1047:
1045:
1028:
1022:
1011:
1007:
1006:
998:
991:
980:
976:
972:
971:
963:
956:
948:
944:
938:
934:
921:
920:
916:
911:
910:
906:
901:
900:
899:Travancoriana
896:
891:
890:
886:
881:
880:
876:
871:
870:
866:
861:
860:
856:
851:
850:
846:
841:
840:
836:
831:
830:
826:
821:
820:
816:
811:
810:
806:
801:
800:
796:
791:
790:
786:
781:
780:
779:Siamthelphusa
776:
771:
770:
766:
761:
760:
756:
751:
750:
746:
741:
740:
736:
731:
730:
726:
721:
720:
716:
711:
710:
706:
701:
700:
699:Perithelphusa
696:
691:
690:
686:
681:
680:
676:
671:
670:
669:Parathelphusa
666:
661:
660:
656:
651:
650:
646:
643:Balss, 1933
641:
640:
636:
631:
630:
626:
621:
620:
616:
611:
610:
606:
601:
600:
596:
591:
590:
586:
581:
580:
576:
571:
570:
566:
561:
560:
556:
551:
550:
546:
541:
540:
536:
531:
530:
526:
521:
520:
516:
511:
510:
506:
501:
500:
496:
491:
490:
486:
481:
480:
479:Globitelphusa
476:
471:
470:
466:
461:
460:
456:
451:
450:
446:
441:
440:
436:
431:
430:
429:Esanthelphusa
426:
421:
420:
416:
411:
410:
406:
401:
400:
396:
391:
390:
386:
381:
380:
376:
371:
370:
366:
361:
360:
356:
351:
350:
346:
341:
340:
336:
331:
330:
326:
321:
320:
316:
311:
310:
306:
301:
300:
296:
295:
289:
287:
283:
279:
275:
270:
268:
264:
259:
255:
251:
247:
243:
238:
236:
232:
222:
220:
216:
212:
208:
204:
200:
196:
187:
186:
181:
178:
174:
169:
164:
161:
158:
157:
154:
151:
149:Superfamily:
148:
147:
144:
143:Heterotremata
141:
138:
137:
134:
131:
128:
127:
124:
121:
118:
117:
114:
111:
108:
107:
104:
101:
98:
97:
94:
91:
88:
87:
84:
81:
78:
77:
74:
71:
68:
67:
64:
61:
58:
57:
52:
47:
43:
40:
36:
32:
27:
22:
19:
1379:expanding it
1368:
1353:
1120:
1060:
1056:
1031:. Retrieved
1021:
1010:the original
1003:
990:
979:the original
974:
968:
955:
937:
917:
907:
903:Bott, 1969
897:
887:
883:Bott, 1969
877:
867:
857:
847:
843:Bott, 1969
837:
827:
823:Bott, 1968
817:
813:Bott, 1968
807:
797:
787:
783:Bott, 1968
777:
773:Bott, 1969
767:
757:
753:Bott, 1969
747:
743:Bott, 1968
737:
733:Bott, 1969
727:
717:
707:
697:
693:Bott, 1969
687:
677:
667:
659:Oziotelphusa
657:
647:
637:
627:
617:
613:Bott, 1969
607:
603:Bott, 1969
597:
587:
577:
567:
557:
547:
543:Bott, 1969
537:
533:Bott, 1970
527:
523:Bott, 1969
517:
507:
497:
493:Bott, 1970
487:
477:
467:
457:
453:Bott, 1974
447:
439:Gecarcinucus
437:
427:
417:
407:
397:
387:
383:Bott, 1969
377:
369:Barytelphusa
367:
357:
353:Bott, 1969
347:
337:
333:Bott, 1969
327:
317:
307:
303:Bott, 1969
297:
274:sister group
271:
263:land bridges
239:
230:
228:
194:
192:
183:
170:, 1904
162:
139:Subsection:
119:Infraorder:
93:Malacostraca
38:
18:
1248:iNaturalist
1145:Wikispecies
879:Thelphusula
703:Man, 1899
689:Perbrinckia
579:Liotelphusa
539:Irmengardia
519:Holthuisana
229:The family
133:Eubrachyura
113:Pleocyemata
1439:Crab stubs
1418:Categories
1063:(1): 287.
929:References
893:Ng, 1997
863:Ng, 1989
833:Ng, 1989
749:Sartoriana
653:Ng, 1991
499:Guinothusa
463:Ng, 1989
449:Geelvinkia
413:Ng, 1990
343:Ng, 1995
313:Ng, 1991
278:Potamoidea
215:New Guinea
207:South Asia
185:See genera
109:Suborder:
83:Arthropoda
1079:1471-2148
849:Syntripsa
769:Sendleria
282:Potamidae
254:Gondwanan
250:Paleogene
219:Australia
129:Section:
123:Brachyura
69:Kingdom:
63:Eukaryota
1172:BioLib:
1130:Wikidata
1097:20849594
799:Sodhiana
679:Pastilla
599:Mainitia
469:Ghatiana
459:Geithusa
299:Adeleana
225:Taxonomy
201:of true
159:Family:
103:Decapoda
79:Phylum:
73:Animalia
59:Domain:
1328:1456382
1240:3254308
1214:2606311
1136:Q994839
1088:2949875
889:Torhusa
759:Sayamia
729:Rouxana
719:Pilarta
589:Mahatha
559:Lamella
399:Coccusa
359:Baratha
339:Bakousa
319:Aradhya
276:to the
248:in the
168:Rathbun
99:Order:
89:Class:
1341:439035
1325:uBio:
1318:226027
1305:178271
1292:168676
1279:621514
1266:100424
1227:1PRTHF
1175:133123
1095:
1085:
1077:
1033:5 June
292:Genera
199:family
197:are a
177:Genera
1369:This
1336:WoRMS
1261:IRMNG
1253:85783
1201:7NHMY
1188:79124
1013:(PDF)
1000:(PDF)
982:(PDF)
965:(PDF)
909:Vanni
789:Snaha
1375:stub
1371:crab
1287:NCBI
1274:ITIS
1235:GBIF
1222:EPPO
1183:BOLD
1093:PMID
1075:ISSN
1035:2016
919:Vela
549:Kani
284:and
213:and
193:The
1209:EoL
1196:CoL
1160:AFD
1083:PMC
1065:doi
288:).
1420::
1338::
1315::
1302::
1289::
1276::
1263::
1250::
1237::
1224::
1211::
1198::
1185::
1162::
1147::
1132::
1091:.
1081:.
1073:.
1061:10
1059:.
1055:.
1043:^
1002:.
975:17
973:.
967:.
945:.
221:.
209:,
1406:e
1399:t
1392:v
1381:.
1099:.
1067::
1037:.
949:.
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