665:
439:
48:
70:
652:
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2501:
382:, and that mouthbrooding has evolved on more than one occasion in the history of the genus. However, it was unable to establish a correlation with any of three habitat variables studied: whether a species was found in lowland or highland streams, whether it was found in
392:, and whether it was found in water with fast or slow currents. Mouthbrooding species tend to exhibit less sexual dimorphism, perhaps because they do not need to defend a territory as the bubble-nesters do.
551:
Bettas are commonly kept as pet fish, especially
Siamese fighting fish. Bettas are popular fish due to their color variety and ease to care for, although the males must be kept separated from other bettas.
538:
fish are hardy and eat almost any animal small enough to consume. This includes worms, larvae of mosquitoes or other insects, and smaller fish. Their natural environment is often resource-limited, so many
2091:
Rüber, Lukas; Britz, Ralf; Tan, Heok Hui; Ng, Peter K. L.; Zardoya, Rafael (April 2004). "Evolution of
Mouthbrooding and Life-History Correlates in the Fighting Fish Genus Betta".
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Tan, H.H.; Ng, P.K.L. (2005). "The fighting fishes (Teleostei: Osphronemidae: genus Betta) of
Singapore, Malaysia and Brunei".
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2388:
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2281:
605:
522:). The Thai phrase is not restricted to one specific strain. The term "fighting fish" is generalized to all members of the
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317:
species are small fishes, but they vary considerably in size, ranging from under 2.5 cm (1 in) total length in
518:
species complex (All have aggressive tendencies in the wild and all are extensively line-bred for aggression in eastern
2046:
336:. This accounts for their ability to thrive in low-oxygen water conditions that would kill most other fish, such as
2744:
2706:
2685:
2455:
Tan Heok Hui (2023). "A new species of black water fighting fish from
Singkep Island (Teleostei: Osphronemidae)".
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2338:
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is italicized and capitalized, when used as a common name it is usually neither italicized nor capitalized. The
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A phylogenetic study published in 2004 concluded tentatively that bubble-nesting was the ancestral condition in
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1807:
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367:. The mouthbrooding species are sometimes called "pseudo bettas", and are sometimes speculated to have
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2014:
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The vernacular name "plakat", often applied to the short-finned ornamental strains, derived from
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There are currently 76 recognized species in this genus. The currently described
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simply as "bettas". As of 2017, around 73 species are classified within the genus
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332:, which means they can breathe atmospheric air using a unique organ called the
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495:, and may be spelled with one 't'. The name of the genus is derived from the
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2238:"Top 25 Most Popular Freshwater Fish For Beginners - Everything Fishkeeping"
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2345:. International Betta Congress Species Maintenance Program. Archived from
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753:
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1958:
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is ubiquitous in the aquarium trade, other bettas are threatened. The
2646:
643:, as well as in captivity; the Red List classifies it as Vulnerable.
636:
621:
368:
81:
2523:
2546:
2512:
2104:
2023:
1915:
640:
519:
383:
302:
101:
2212:"Betta Fish Information; Plakats, Veiltails, Halfmoon, Crowntail"
617:
283:
131:
2190:"Betta splendens – Siamese Fighting Fish (Micracanthus marchei)"
2500:
389:
351:
The bettas exhibit two kinds of spawning behaviour: some build
345:
341:
91:
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416:
386:
272:
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469:
263:
257:
2370:. International Betta Congress Species Maintenance Program
2070:. International Betta Congress Species Maintenance Program
2209:
526:
species complex, including the
Siamese fighting fish.
449:, is often referred to simply as a "betta" in the U.S.
472:
466:
260:
254:
2412:
463:
251:
2099:(4). Society for the Study of Evolution: 799–813.
624:between 1970 and 1994. This likely refers to the
2726:
2090:
321:to 14 cm (5.5 in) in the Akar betta (
2086:
2084:
2473:
2454:
2432:Froese, Rainer; Pauly, Daniel, eds. (2023).
2308:
2302:
2431:
2360:
2057:
2051:
2258:"2006 IUCN Red List of Threatened Species"
2081:
2005:
2003:
2001:
1999:
415:. A useful distinction is that, while the
46:
27:which is often called simply "betta", see
2406:
2331:
663:
650:
635:which continues to exist in the wild in
437:
2393:Integrated Taxonomic Information System
2250:
2009:
1996:
487:word bet. The name is often pronounced
19:This article is about the entire genus
2727:
375:to their fast-moving stream habitats.
2528:
2527:
2132:
2130:
2381:
2285:World Conservation Monitoring Centre
2282:United Nations Environment Programme
2270:
2169:"Common Names Summary - Betta picta"
606:United Nations Environment Programme
576:species as Vulnerable. In addition,
510:which means "fighting fish", is the
483:; the first part is the same as the
2319:IUCN Red List of Threatened Species
13:
2235:
2127:
308:
275:of small, active, often colorful,
14:
2771:
2493:
546:
2511:
2499:
2216:www.americanaquariumproducts.com
543:species are generalist feeders.
459:
289:(Osphronemidae). The best known
247:
68:
2480:The Raffles Bulletin of Zoology
2467:
2457:The Raffles Bulletin of Zoology
2448:
2425:
555:
2229:
2210:americanaquariumproducts.com.
2203:
2182:
2161:
2146:(4th ed.). Archived from
2031:
608:lists an unconfirmed species,
344:, drainage ditches, and large
1:
1990:
407:) are frequently sold in the
371:from the nest-builders in an
2760:Taxa named by Pieter Bleeker
2368:"Species Complex Management"
2143:American Heritage Dictionary
681:species can be grouped into
514:name for all members of the
7:
2415:Raffles Bulletin of Zoology
601:are Critically Endangered.
10:
2776:
646:
628:Singaporean population of
18:
2536:
1582:complex (fighting fish):
457:(or betta) is pronounced
193:
186:
173:
166:
65:Scientific classification
63:
54:
45:
38:
2278:"Extinctions since 1970"
2013:; Pauly, Daniel (eds.).
735:& K. K. P. Lim, 2004
2745:Fauna of Southeast Asia
1674:(Siamese fighting fish)
564:species are common and
529:
395:
2755:Freshwater fish genera
2474:Tan Heok Hui (2009). "
2287:. 2006. Archived from
674:
661:
450:
297:commonly known as the
2309:Kottelat, M. (1996).
2039:Caring for Betta Fish
2027:. March 2019 version.
667:
654:
503:("persistent fish").
447:Siamese fighting fish
441:
431:for example, is thus
401:Siamese fighting fish
301:and often kept as an
299:Siamese fighting fish
29:Siamese fighting fish
2508:at Wikimedia Commons
1626:Betta siamorientalis
1852:Betta chloropharynx
1601:Betta mahachaiensis
847:Betta albimarginata
582:is Endangered, and
572:classifies several
359:, while others are
57:Betta albimarginata
2041:Lulu Press, 2006.
2015:"Species in genus
1869:Betta hipposideros
675:
662:
451:
224:D. S. Jordan, 1919
23:. For the species
2722:
2721:
2694:Open Tree of Life
2530:Taxon identifiers
2504:Media related to
2058:Fernando, Yohan.
1984:
1971:
1954:
1942:Betta spilotogena
1937:
1924:
1911:
1898:
1881:
1864:
1847:
1825:
1816:Betta unimaculata
1811:
1794:
1777:
1763:
1750:
1733:
1716:
1691:
1673:
1659:
1646:
1621:
1595:
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1505:
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1461:
1444:
1426:
1413:
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1383:
1366:
1349:
1323:
1310:
1296:
1282:
1257:
1239:
1226:
1209:
1196:
1188:Betta dennisyongi
1175:
1154:
1137:
1116:
1099:
1086:
1069:
1056:
1039:
1022:
1005:
992:
975:
950:
932:
911:
897:
889:Betta anabatoides
876:
859:
834:
817:
800:
779:
762:
749:
736:
723:
705:
683:species complexes
616:as having become
280:ray-finned fishes
239:
238:
233:
225:
217:
209:
201:
200:van Hasselt, 1859
176:Betta trifasciata
162:
2767:
2750:Fish of Thailand
2715:
2714:
2702:
2701:
2689:
2688:
2676:
2675:
2663:
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2650:
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2637:
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2598:
2597:
2585:
2584:
2572:
2571:
2570:
2557:
2556:
2555:
2525:
2524:
2516:Data related to
2515:
2503:
2488:
2487:
2476:Betta pardalotos
2471:
2465:
2464:
2452:
2446:
2445:
2436:Betta breviobesa
2429:
2423:
2422:
2410:
2404:
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2401:
2399:
2385:
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2376:
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2224:
2222:
2207:
2201:
2200:
2198:
2196:
2186:
2180:
2179:
2177:
2176:
2165:
2159:
2158:
2156:
2155:
2134:
2125:
2124:
2088:
2079:
2078:
2076:
2075:
2055:
2049:
2035:
2029:
2028:
2007:
1979:
1962:
1945:
1932:
1919:
1906:
1903:Betta pardalotos
1889:
1872:
1855:
1842:
1819:
1802:
1785:
1772:
1758:
1755:Betta macrostoma
1741:
1724:
1707:
1682:
1668:
1654:
1651:Betta smaragdina
1629:
1618:Sriwattanarothai
1604:
1596:(crescent betta)
1590:
1565:
1548:
1531:
1514:
1499:
1486:
1469:
1452:
1435:
1421:
1404:
1391:
1374:
1357:
1354:Betta breviobesa
1340:
1318:
1305:
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1273:
1248:
1234:
1217:
1204:
1191:
1170:
1145:
1132:
1107:
1094:
1077:
1064:
1061:Betta persephone
1047:
1044:Betta miniopinna
1030:
1013:
1000:
983:
966:
941:
927:
906:
892:
867:
864:Betta channoides
850:
840:B. albimarginata
825:
808:
787:
770:
757:
744:
731:
714:
700:
658:Betta smaragdina
493:American English
490:
482:
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471:
468:
465:
270:
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265:
262:
259:
256:
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231:
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215:
213:Parophiocephalus
207:
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157:
73:
72:
50:
36:
35:
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2236:Thomas, David.
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2089:
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2052:
2036:
2032:
2008:
1997:
1993:
1782:Betta pallifina
1764:(spotfin betta)
1721:Betta gladiator
1704:Betta compuncta
1665:Betta splendens
1545:Betta schalleri
1297:(spotted betta)
1129:Betta dimidiata
980:Betta burdigala
963:Betta brownorum
697:Betta akarensis
649:
558:
549:
532:
462:
458:
443:Betta splendens
398:
311:
309:Characteristics
250:
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182:
179:
156:
67:
32:
25:Betta splendens
17:
12:
11:
5:
2773:
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2757:
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2747:
2742:
2740:Macropodusinae
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2703:
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2540:
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2522:
2521:
2520:at Wikispecies
2509:
2495:
2494:External links
2492:
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2424:
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2181:
2171:. Fishbase.org
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2126:
2105:10.1554/03-364
2080:
2050:
2030:
2011:Froese, Rainer
1994:
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1989:
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1972:
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1938:
1925:
1912:
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1848:
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1826:(Howong betta)
1812:
1804:M. C. W. Weber
1795:
1778:
1769:Betta ocellata
1765:
1751:
1734:
1717:
1697:B. unimaculata
1694:
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1692:
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1587:Betta imbellis
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1562:Betta stigmosa
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1506:(Penang betta)
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1324:(Borneo betta)
1315:Betta taeniata
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1210:
1201:Betta foerschi
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1142:Betta krataios
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1074:Betta rutilans
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882:B. anabatoides
879:
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860:
837:
836:
835:
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801:
797:Casey K. C. Ng
789:N. S. S. Kamal
780:
767:Betta ibanorum
763:
750:
737:
728:Betta aurigans
724:
707:
648:
645:
598:B. spilotogena
557:
554:
548:
547:Bettas as pets
545:
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397:
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340:, slow-moving
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142:Macropodusinae
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122:Anabantiformes
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112:Actinopterygii
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2539:
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2526:
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2514:
2510:
2507:
2502:
2498:
2497:
2486:(2): 501–504.
2485:
2481:
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2443:
2439:
2437:
2428:
2420:
2416:
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2349:on 2013-04-14
2348:
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2291:on 2007-09-30
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2273:
2259:
2253:
2239:
2232:
2217:
2213:
2206:
2191:
2185:
2170:
2164:
2150:on 2007-10-30
2149:
2145:
2144:
2139:
2133:
2131:
2122:
2118:
2114:
2110:
2106:
2102:
2098:
2094:
2087:
2085:
2069:
2067:
2063:
2062:Betta edithae
2054:
2048:
2047:1-4116-9365-5
2044:
2040:
2037:Marcus Song,
2034:
2026:
2025:
2020:
2018:
2012:
2006:
2004:
2002:
2000:
1995:
1982:
1978:
1977:
1973:
1969:
1965:
1961:
1960:
1956:
1952:
1948:
1944:
1943:
1939:
1935:
1931:
1930:
1926:
1922:
1918:
1917:
1913:
1909:
1905:
1904:
1900:
1896:
1892:
1888:
1887:
1883:
1879:
1875:
1871:
1870:
1866:
1862:
1858:
1854:
1853:
1849:
1845:
1841:
1840:
1836:
1835:
1833:
1830:
1823:
1818:
1817:
1813:
1809:
1805:
1801:
1800:
1796:
1792:
1788:
1784:
1783:
1779:
1775:
1771:
1770:
1766:
1761:
1757:
1756:
1752:
1748:
1744:
1740:
1739:
1735:
1731:
1727:
1723:
1722:
1718:
1714:
1710:
1706:
1705:
1701:
1700:
1698:
1695:
1689:
1685:
1681:
1680:
1679:Betta stiktos
1676:
1671:
1667:
1666:
1662:
1657:
1653:
1652:
1648:
1644:
1640:
1636:
1632:
1628:
1627:
1623:
1619:
1615:
1611:
1607:
1603:
1602:
1598:
1593:
1589:
1588:
1584:
1583:
1581:
1578:
1572:
1568:
1564:
1563:
1559:
1555:
1551:
1547:
1546:
1542:
1538:
1534:
1530:
1529:
1525:
1521:
1517:
1513:
1512:
1511:Betta pulchra
1508:
1503:
1498:
1497:
1493:
1489:
1485:
1484:
1480:
1476:
1472:
1468:
1467:
1466:Betta pallida
1463:
1459:
1455:
1451:
1450:
1446:
1442:
1438:
1434:
1433:
1432:Betta kuehnei
1429:
1427:(dusky betta)
1424:
1420:
1419:
1415:
1411:
1407:
1403:
1402:
1398:
1394:
1390:
1389:
1385:
1381:
1377:
1373:
1372:
1371:Betta cracens
1368:
1364:
1360:
1356:
1355:
1351:
1347:
1343:
1339:
1338:
1337:Betta apollon
1334:
1333:
1331:
1328:
1321:
1317:
1316:
1312:
1308:
1304:
1303:
1302:Betta simplex
1299:
1294:
1289:
1288:
1284:
1280:
1276:
1272:
1271:
1267:
1266:
1264:
1261:
1255:
1251:
1247:
1246:
1242:
1237:
1233:
1232:
1228:
1224:
1220:
1216:
1215:
1211:
1207:
1203:
1202:
1198:
1194:
1190:
1189:
1185:
1184:
1182:
1179:
1173:
1169:
1168:
1167:Betta edithae
1164:
1163:
1161:
1158:
1152:
1148:
1144:
1143:
1139:
1135:
1134:T. R. Roberts
1131:
1130:
1126:
1125:
1123:
1120:
1114:
1110:
1106:
1105:
1101:
1097:
1093:
1092:
1091:Betta tussyae
1088:
1084:
1080:
1076:
1075:
1071:
1067:
1063:
1062:
1058:
1054:
1050:
1046:
1045:
1041:
1037:
1033:
1029:
1028:
1024:
1020:
1016:
1012:
1011:
1007:
1003:
999:
998:
997:Betta coccina
994:
990:
986:
982:
981:
977:
973:
969:
965:
964:
960:
959:
957:
954:
948:
944:
940:
939:
938:Betta simorum
935:
930:
926:
925:
924:Betta bellica
921:
920:
918:
915:
909:
905:
904:
900:
898:(giant betta)
895:
891:
890:
886:
885:
883:
880:
874:
870:
866:
865:
861:
857:
853:
849:
848:
844:
843:
841:
838:
832:
828:
824:
823:
822:Betta pinguis
819:
815:
811:
807:
806:
805:Betta obscura
802:
798:
794:
790:
786:
785:
784:Betta nuluhon
781:
777:
773:
769:
768:
764:
760:
756:
755:
751:
747:
743:
742:
741:Betta balunga
738:
734:
730:
729:
725:
721:
717:
713:
712:
708:
703:
699:
698:
694:
693:
691:
688:
687:
686:
684:
680:
673:
672:
671:Betta tussyae
666:
660:
659:
653:
644:
642:
638:
634:
632:
627:
623:
619:
615:
611:
607:
602:
600:
599:
594:
592:
591:B. persephone
588:
586:
585:B. miniopinna
581:
580:
575:
571:
570:IUCN Red List
567:
563:
553:
544:
542:
537:
527:
525:
521:
517:
513:
509:
504:
502:
498:
494:
486:
480:
456:
448:
444:
440:
436:
434:
430:
426:
422:
418:
414:
410:
409:United States
406:
402:
393:
391:
388:
385:
381:
376:
374:
370:
366:
362:
361:mouthbrooders
358:
354:
349:
347:
343:
339:
335:
331:
326:
324:
320:
316:
306:
304:
300:
296:
295:B. splendens,
292:
288:
285:
281:
278:
274:
268:
245:
244:
232:Richter, 1981
230:
227:
222:
219:
214:
211:
208:Sauvage, 1879
206:
203:
198:
195:
194:
192:
189:
185:
181:Bleeker, 1850
178:
177:
172:
169:
165:
160:
155:
154:
150:
147:
146:
143:
140:
137:
136:
133:
132:Osphronemidae
130:
127:
126:
123:
120:
117:
116:
113:
110:
107:
106:
103:
100:
97:
96:
93:
90:
87:
86:
83:
80:
77:
76:
71:
66:
62:
59:
58:
53:
49:
44:
41:
37:
34:
30:
26:
22:
16:Genus of fish
2537:
2483:
2479:
2475:
2469:
2460:
2456:
2450:
2441:
2435:
2427:
2418:
2414:
2408:
2396:. Retrieved
2383:
2372:. Retrieved
2362:
2351:. Retrieved
2347:the original
2340:
2333:
2322:. Retrieved
2312:
2304:
2293:. Retrieved
2289:the original
2272:
2261:. Retrieved
2252:
2241:. Retrieved
2231:
2219:. Retrieved
2215:
2205:
2193:. Retrieved
2184:
2173:. Retrieved
2163:
2152:. Retrieved
2148:the original
2141:
2096:
2092:
2072:. Retrieved
2065:
2061:
2053:
2038:
2033:
2022:
2016:
1981:Krummenacher
1976:Betta waseri
1974:
1957:
1940:
1929:Betta renata
1927:
1914:
1901:
1884:
1867:
1850:
1839:Betta andrei
1837:
1831:
1814:
1799:Betta patoti
1797:
1780:
1767:
1753:
1736:
1719:
1702:
1696:
1677:
1663:
1660:(Blue betta)
1649:
1624:
1599:
1585:
1580:B. splendens
1579:
1560:
1543:
1526:
1509:
1496:Betta pugnax
1494:
1481:
1471:I. Schindler
1464:
1447:
1437:I. Schindler
1430:
1416:
1406:I. Schindler
1399:
1388:Betta enisae
1386:
1369:
1352:
1342:I. Schindler
1335:
1329:
1313:
1300:
1293:Valenciennes
1285:
1268:
1262:
1245:Betta strohi
1243:
1240:(Toba betta)
1229:
1214:Betta mandor
1212:
1199:
1186:
1180:
1165:
1159:
1140:
1127:
1122:B. dimidiata
1121:
1104:Betta uberis
1102:
1089:
1072:
1059:
1042:
1027:Betta livida
1025:
1015:I. Schindler
1010:Betta hendra
1008:
995:
978:
961:
955:
936:
933:(slim betta)
922:
916:
901:
887:
881:
862:
845:
839:
820:
803:
782:
765:
752:
739:
726:
711:Betta antoni
709:
706:(Akar betta)
695:
690:B. akarensis
689:
678:
676:
669:
656:
629:
613:
609:
603:
596:
589:
583:
577:
573:
566:B. splendens
565:
561:
559:
556:Conservation
550:
540:
535:
533:
524:B. splendens
523:
516:B. splendens
515:
507:
505:
500:
454:
452:
442:
433:Penang betta
428:
420:
412:
405:B. splendens
404:
399:
379:
377:
364:
357:B. splendens
356:
353:bubble nests
350:
338:rice paddies
327:
323:B. akarensis
322:
319:B. chanoides
318:
314:
312:
294:
290:
242:
241:
240:
228:
220:
212:
205:Micracanthus
204:
196:
174:
168:Type species
152:
151:
55:
39:
33:
24:
20:
2642:iNaturalist
2562:Wikispecies
2221:16 November
2195:16 November
2064:- a Pseudo
1964:P. K. L. Ng
1947:P. K. L. Ng
1886:Betta omega
1874:P. K. L. Ng
1861:P. K. L. Ng
1808:de Beaufort
1791:P. K. L. Ng
1774:de Beaufort
1747:P. K. L. Ng
1738:Betta ideii
1730:P. K. L. Ng
1713:P. K. L. Ng
1688:P. K. L. Ng
1571:P. K. L. Ng
1554:P. K. L. Ng
1537:P. K. L. Ng
1483:Betta prima
1458:P. K. L. Ng
1418:Betta fusca
1401:Betta ferox
1380:P. K. L. Ng
1287:Betta picta
1231:Betta rubra
1223:P. K. L. Ng
1181:B. foerschi
1151:P. K. L. Ng
1113:P. K. L. Ng
1079:K. E. Witte
1032:P. K. L. Ng
989:P. K. L. Ng
968:K. E. Witte
947:P. K. L. Ng
903:Betta midas
873:P. K. L. Ng
856:P. K. L. Ng
814:P. K. L. Ng
776:P. K. L. Ng
759:P. K. L. Ng
754:Betta chini
720:P. K. L. Ng
560:While many
425:common name
330:anabantoids
328:Bettas are
293:species is
271:is a large
229:Pseudobetta
216:Popta, 1905
138:Subfamily:
2729:Categories
2463:: 491–495.
2374:2006-07-01
2353:2006-07-01
2341:Betta tomi
2324:2006-07-01
2313:Betta tomi
2295:2006-07-01
2263:2006-06-30
2243:2022-07-15
2175:2012-07-31
2154:2006-06-29
2074:2017-01-30
1991:References
1959:Betta tomi
1639:Ruenwongsa
1614:Ruenwongsa
1528:Betta raja
1475:J. Schmidt
1449:Betta lehi
1441:J. Schmidt
1410:J. Schmidt
1346:J. Schmidt
1270:Betta falx
1160:B. edithae
972:J. Schmidt
956:B. coccina
917:B. bellica
655:A pair of
626:extirpated
501:ikan betah
429:B. pugnax,
373:adaptation
363:, such as
355:, such as
277:freshwater
2121:198158291
2093:Evolution
1934:H. H. Tan
1921:H. H. Tan
1908:H. H. Tan
1891:H. H. Tan
1844:H. H. Tan
1834:complex:
1832:B. waseri
1787:H. H. Tan
1743:H. H. Tan
1726:H. H. Tan
1709:H. H. Tan
1699:complex:
1684:H. H. Tan
1643:Jeenthong
1631:Kowasupat
1606:Kowasupat
1567:H. H. Tan
1533:H. H. Tan
1520:S. H. Tan
1516:H. H. Tan
1454:H. H. Tan
1376:H. H. Tan
1359:H. H. Tan
1332:complex:
1330:B. pugnax
1275:H. H. Tan
1265:complex:
1219:H. H. Tan
1193:H. H. Tan
1183:complex:
1162:complex:
1147:H. H. Tan
1124:complex:
1109:H. H. Tan
1053:S. H. Tan
1049:H. H. Tan
958:complex:
943:H. H. Tan
919:complex:
908:H. H. Tan
884:complex:
842:complex:
827:H. H. Tan
810:H. H. Tan
793:H. H. Tan
772:H. H. Tan
733:H. H. Tan
716:H. H. Tan
692:complex:
637:Indonesia
622:Singapore
579:B. livida
453:The name
334:labyrinth
282:, in the
88:Kingdom:
82:Eukaryota
2553:Q4164940
2547:Wikidata
2442:FishBase
2421:: 43–99.
2024:FishBase
1968:Kottelat
1951:Kottelat
1916:Betta pi
1878:Kottelat
1857:Kottelat
1635:Panijpan
1610:Panijpan
1550:Kottelat
1488:Kottelat
1393:Kottelat
1363:Kottelat
1307:Kottelat
1279:Kottelat
1263:B. picta
1254:Kottelat
1250:Schaller
1096:Schaller
1083:Kottelat
1066:Schaller
1036:Kottelat
985:Kottelat
869:Kottelat
852:Kottelat
831:Kottelat
641:Malaysia
520:Thailand
489:/ˈbeɪtə/
365:B. picta
303:aquarium
197:Anostoma
188:Synonyms
128:Family:
102:Chordata
98:Phylum:
92:Animalia
78:Domain:
2699:1033359
2660:1268464
2634:2393980
2398:30 June
2389:"Betta"
2317:. 2006
2138:"Betta"
2113:3449277
1824:, 1905)
1656:Ladiges
1592:Ladiges
1504:, 1849)
1295:, 1846)
1236:Perugia
929:Sauvage
894:Bleeker
647:Species
631:B. tomi
618:extinct
508:pla kad
485:English
417:generic
390:forests
369:evolved
346:puddles
342:streams
284:gourami
221:Oshimia
159:Bleeker
148:Genus:
118:Order:
108:Class:
2712:268539
2686:158455
2673:172610
2119:
2111:
2066:Betta?
2045:
1983:, 1986
1970:, 1994
1966:&
1953:, 1994
1949:&
1936:, 1998
1923:, 1998
1910:, 2009
1897:, 2018
1893:&
1880:, 1994
1876:&
1863:, 1994
1859:&
1846:, 2023
1810:, 1922
1806:&
1793:, 2005
1789:&
1776:, 1933
1762:, 1910
1749:, 2006
1745:&
1732:, 2005
1728:&
1715:, 2006
1711:&
1690:, 2005
1686:&
1672:, 1910
1658:, 1972
1645:, 2012
1641:&
1620:, 2012
1616:&
1594:, 1975
1573:, 2005
1569:&
1556:, 1994
1552:&
1539:, 2005
1535:&
1522:, 1996
1518:&
1502:Cantor
1490:, 1994
1477:, 2004
1473:&
1460:, 2005
1456:&
1443:, 2008
1439:&
1425:, 1910
1412:, 2006
1408:&
1395:, 1995
1382:, 2005
1378:&
1365:, 1998
1361:&
1348:, 2006
1344:&
1322:, 1910
1309:, 1994
1281:, 1998
1277:&
1256:, 1989
1252:&
1238:, 1893
1225:, 2006
1221:&
1208:, 1979
1206:Vierke
1195:, 2013
1174:, 1984
1172:Vierke
1153:, 2006
1149:&
1136:, 1989
1115:, 2006
1111:&
1098:, 1985
1085:, 1991
1081:&
1068:, 1986
1055:, 1994
1051:&
1038:, 1992
1034:&
1021:, 2013
1017:&
1004:, 1979
1002:Vierke
991:, 1994
987:&
974:, 1992
970:&
949:, 1996
945:&
931:, 1884
910:, 2009
896:, 1851
875:, 1994
871:&
858:, 1994
854:&
833:, 1998
829:&
816:, 2005
812:&
799:, 2020
795:&
778:, 2004
774:&
761:, 1993
748:, 1940
722:, 2006
718:&
704:, 1910
445:, the
287:family
161:, 1850
2735:Betta
2707:WoRMS
2655:IRMNG
2647:86963
2608:47892
2595:Betta
2582:Betta
2568:Betta
2538:Betta
2518:Betta
2506:Betta
2117:S2CID
2109:JSTOR
2017:Betta
1895:Ahmad
1822:Popta
1760:Regan
1670:Regan
1423:Regan
1320:Regan
1019:Linke
746:Herre
702:Regan
679:Betta
668:Male
614:tomi,
610:Betta
574:Betta
562:Betta
541:Betta
536:Betta
534:Wild
499:term
497:Malay
455:Betta
421:Betta
419:name
413:Betta
387:swamp
380:Betta
315:Betta
305:pet.
291:Betta
273:genus
243:Betta
153:Betta
40:Betta
21:Betta
2681:NCBI
2668:ITIS
2629:GBIF
2621:39X6
2603:BOLD
2400:2006
2223:2018
2197:2018
2043:ISBN
639:and
612:cf.
604:The
595:and
530:Diet
512:Thai
396:Name
384:peat
313:All
2616:CoL
2590:AFD
2577:ADW
2101:doi
620:in
491:in
427:of
325:).
2731::
2709::
2696::
2683::
2670::
2657::
2644::
2631::
2618::
2605::
2592::
2579::
2564::
2549::
2484:57
2482:.
2461:71
2459:.
2440:.
2419:13
2417:.
2391:.
2280:.
2214:.
2140:.
2129:^
2115:.
2107:.
2097:53
2095:.
2083:^
2021:.
1998:^
1637:,
1633:,
1612:,
1608:,
791:,
685::
435:.
348:.
2444:.
2438:"
2434:"
2402:.
2377:.
2356:.
2343:"
2339:"
2327:.
2315:"
2311:"
2298:.
2266:.
2246:.
2225:.
2199:.
2178:.
2157:.
2123:.
2103::
2077:.
2068:"
2060:"
2019:"
1820:(
1500:(
1291:(
633:,
593:,
587:,
479:/
476:ə
473:t
470:ɛ
467:b
464:ˈ
461:/
403:(
267:/
264:ə
261:t
258:ɛ
255:b
252:ˈ
249:/
31:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
