1130:
1110:
1083:
967:
1022:
399:
56:
31:
680:
1445:
494:
2132:
823:
297:) and having at least 207 species. In addition, some authorities expand the order to include the suborder Nandoidei, which includes three families - the
2157:
2112:
895:
Below shows the phylogenetic relationships among the
Anabantiform families after Collins et al. (2015), here including the Nandoidei as Anabantiforms:
2182:
855:
2142:
2127:
563:
2147:
2299:
819:
815:
125:
1211:
912:
1594:
1531:"The subterranean Aenigmachanna gollum , a new genus and species of snakehead (Teleostei: Channidae) from Kerala, South India"
1232:
1308:
1341:
309:- that appear to be closely related to the Anabantiformes. The order, and these three related families (classified as
1442:
1399:
752:
698:
2304:
1298:
1413:"A Fossil Gourami (teleostei, Anabantoidei) from Probable Eocene Deposits of the Ombilin Basin, Sumatra, Indonesia"
602:
508:
1242:
1587:
1129:
55:
2278:
2273:
1109:
1082:
2264:
873:
655:
452:
392:
361:
1489:
1467:
966:
2353:
1580:
1263:
2330:
431:
There are two suborders and five families currently recognized within the order
Anabantiformes:
2226:
522:
1206:
2325:
2317:
1530:
412:
403:
348:. This clade is sometimes referred to as the Carangaria but is left unnamed and unranked in
2286:
1572:
1228:
694:
186:
147:
8:
1865:
1515:. Illinois biological monographs. Vol. 30. Urbana,University of Illinois Press,1963.
641:
317:
2082:
2007:
1961:
1812:
550:
168:
141:
50:
38:
1411:
Murray, A. M.; Zaim, Y.; Rizal, Y.; Aswan, Y.; Gunnell, G. F.; Ciochon, R. L. (2015).
1205:
R. Betancur-Rodriguez, E. Wiley, N. Bailly, A. Acero, M. Miya, G. Lecointre, G. Ortí:
2312:
2177:
1784:
1776:
1558:
1550:
1487:
1424:
1395:
1304:
1237:
1021:
841:
663:
466:
345:
419:. In addition to being aquarium fish, some of the larger anabantiforms (such as the
2187:
2162:
2047:
2012:
1817:
1761:
1717:
1688:
1542:
1356:
1037:
1017:
1004:
796:
760:
380:
372:
341:
333:
278:
274:
204:
398:
2197:
2137:
2097:
2037:
1994:
1873:
1449:
1215:
1105:
827:
612:
577:
530:
384:
379:
are united by the presence of the labyrinth organ, which is a highly folded supra
290:
259:
2249:
2077:
2022:
1969:
1941:
1931:
1922:
1855:
1847:
1789:
1604:
1546:
1513:
The comparative osteology and phylogeny of the
Anabantoidei (Teleostei, Pisces)
1366:
726:
675:
617:
536:
518:
488:
480:
311:
266:
182:
97:
1529:
Raghavan, Rajeev; Dahanukar, Neelesh; Anoop, V. K.; Britz, Ralf (2019-05-09).
2347:
2102:
2092:
2062:
2042:
2017:
1832:
1827:
1822:
1635:
1554:
1428:
1342:"Phylogenetic systematics of leaffishes (Teleostei: Polycentridae, Nandidae)"
1125:
859:
809:
635:
625:
546:
420:
337:
294:
282:
2167:
2122:
2107:
2087:
2072:
2002:
1946:
1936:
1911:
1906:
1807:
1641:
1562:
1058:
688:
436:
388:
376:
368:
134:
2258:
2192:
2152:
2117:
2032:
2027:
1979:
1888:
1751:
1743:
1699:
1680:
1670:
1314:
1078:
925:
706:
596:
587:
448:
329:
306:
286:
110:
1412:
2172:
1984:
1883:
1878:
1799:
1756:
1722:
1629:
1488:
Richard van der Laan; William N. Eschmeyer & Ronald Fricke (2014).
1296:
849:
787:
771:
746:
736:
391:
expansion of the epibranchial bone of the first gill arch and used for
263:
1394:
Pinter, H. (1986). Labyrinth Fish. Barron's
Educational Series, Inc.,
1361:
1974:
1951:
1651:
837:
474:
411:
Many species are popular as aquarium fish - the most notable are the
322:
67:
30:
2220:
2243:
2067:
2057:
1898:
1709:
1623:
962:
571:
558:
502:
440:
298:
270:
87:
2291:
1837:
1733:
947:
649:
416:
302:
1659:
1617:
867:
460:
357:
77:
243:
716:
367:
These fish are characterized by the presence of teeth on the
325:
1602:
237:
1528:
1349:
Journal of
Zoological Systematics and Evolutionary Research
353:
213:
1766:
423:) are also harvested for food in their native countries.
231:
222:
207:
1443:
Diseases of Giant
Gourami, Osphronemus goramy (Lacepede)
1454:
The
Aquatic Animal Health Research Institute Newsletter
1227:
1208:
1201:
1199:
246:
234:
225:
210:
1410:
1297:
J. S. Nelson; T. C. Grande; M. V. H. Wilson (2016).
228:
219:
1196:
240:
216:
1231:; Fricke, Ron & van der Laan, Richard (eds.).
1339:
1335:
1333:
1331:
2345:
407:is the only surviving described anabantid fossil
1465:
1328:
1588:
1510:
1340:Collins, R.A.; Britz, R.; Rüber, L. (2015).
1595:
1581:
885:
29:
1360:
1292:
1290:
1288:
1286:
1284:
1390:
1388:
1386:
397:
2346:
1524:
1522:
1281:
332:, the other orders in the clade being
2225:
2224:
1576:
1490:"Family-group names of Recent fishes"
1481:
1383:
1504:
1303:(5th ed.). Wiley. p. 390.
1519:
1221:
13:
1417:Journal of Vertebrate Paleontology
360:, with some species introduced in
352:. This group of fish are found in
14:
2365:
1128:
1108:
1081:
1020:
965:
203:
54:
328:which is a sister clade to the
1459:
1435:
1404:
1256:
1243:California Academy of Sciences
1:
1466:Froese, R.; D. Pauly (eds.).
1441:Chanphong, Jitkasem. (1995).
1189:
668:(Prashad & Mukerji, 1929)
890:
7:
1468:"Trichogaster trichopterus"
426:
10:
2370:
1547:10.11646/zootaxa.4603.2.10
2233:
1993:
1960:
1920:
1897:
1864:
1846:
1798:
1775:
1742:
1731:
1708:
1697:
1679:
1668:
1650:
1612:
1123:
1103:
1096:
1076:
1069:
1062:
1035:
1015:
1008:
998:
960:
945:
938:
923:
916:
906:
254:, collectively known as
174:
167:
160:
155:
140:
133:
51:Scientific classification
49:
37:
28:
23:
1448:January 6, 2007, at the
362:United States of America
886:Alternative systematics
415:and several species of
156:Suborders and Families
1511:Karel F. Liem (1963).
591:(van der Hoeven, 1832)
408:
315:by the 5th edition of
2326:Paleobiology Database
1229:Eschmeyer, William N.
413:Siamese fighting fish
404:Ombilinichthys yamini
401:
1866:Protacanthopterygii
1856:Lepidogalaxiiformes
1300:Fishes of the World
350:Fishes of the World
318:Fishes of the World
2083:Cyprinodontiformes
2008:Trachichthyiformes
1962:Paracanthopterygii
1813:Alepocephaliformes
1607:orders by subclass
1214:2017-07-11 at the
776:(Canestrini, 1860)
409:
387:. It is formed by
142:Anabas testudineus
43:Anabas testudineus
2341:
2340:
2313:Open Tree of Life
2227:Taxon identifiers
2218:
2217:
2214:
2213:
2210:
2209:
2206:
2205:
2178:Tetraodontiformes
2133:Chaetodontiformes
2068:Pleuronectiformes
2023:Batrachoidiformes
1785:Osteoglossiformes
1777:Osteoglossomorpha
1362:10.1111/jzs.12103
1310:978-1-118-34233-6
1238:Catalog of Fishes
1186:
1185:
1177:
1176:
1168:
1167:
1159:
1158:
1150:
1149:
1141:
1140:
1048:
1047:
987:
986:
978:
977:
877:
863:
845:
831:
805:
792:
777:
766:
756:
742:
732:
722:
712:
702:
684:
669:
664:Parasphaerichthys
659:
645:
631:
621:
606:
592:
581:
567:
554:
540:
526:
512:
498:
484:
470:
456:
444:
346:Pleuronectiformes
321:), are part of a
262:of air-breathing
196:
195:
190:
129:
2361:
2334:
2333:
2321:
2320:
2308:
2307:
2295:
2294:
2282:
2281:
2269:
2268:
2267:
2254:
2253:
2252:
2222:
2221:
2188:Centrarchiformes
2163:Priacanthiformes
2158:Scatophagiformes
2113:Uranoscopiformes
2058:Istiophoriformes
2048:Synbranchiformes
2013:Holocentriformes
1980:Stylephoriformes
1932:Ateleopodiformes
1818:Gonorynchiformes
1762:Notacanthiformes
1740:
1739:
1736:
1718:Lepisosteiformes
1706:
1705:
1702:
1689:Acipenseriformes
1677:
1676:
1673:
1597:
1590:
1583:
1574:
1573:
1567:
1566:
1526:
1517:
1516:
1508:
1502:
1501:
1485:
1479:
1478:
1476:
1475:
1463:
1457:
1439:
1433:
1432:
1408:
1402:
1392:
1381:
1380:
1378:
1377:
1371:
1365:. Archived from
1364:
1346:
1337:
1326:
1325:
1323:
1322:
1313:. Archived from
1294:
1279:
1278:
1276:
1274:
1260:
1254:
1253:
1251:
1249:
1225:
1219:
1203:
1132:
1112:
1099:
1098:
1085:
1072:
1071:
1065:
1064:
1038:Aenigmachannidae
1024:
1011:
1010:
1001:
1000:
969:
941:
940:
919:
918:
909:
908:
899:
898:
871:
853:
840:
813:
799:
797:Aenigmachannidae
790:
775:
764:
761:Pseudosphromenus
750:
741:(Lacepède, 1801)
740:
730:
720:
711:(Lacepède, 1801)
710:
692:
678:
667:
653:
639:
629:
615:
600:
590:
575:
561:
549:
534:
521:
506:
492:
478:
464:
451:
439:
342:Istiophoriformes
334:Synbranchiformes
283:Aenigmachannidae
253:
252:
249:
248:
245:
242:
239:
236:
233:
230:
227:
224:
221:
218:
215:
212:
209:
180:
124:
59:
58:
33:
21:
20:
2369:
2368:
2364:
2363:
2362:
2360:
2359:
2358:
2344:
2343:
2342:
2337:
2329:
2324:
2316:
2311:
2303:
2298:
2290:
2285:
2277:
2272:
2263:
2262:
2257:
2248:
2247:
2242:
2229:
2219:
2202:
2198:Scorpaeniformes
2138:Acanthuriformes
2098:Gobiesociformes
2038:Syngnathiformes
1995:Acanthopterygii
1989:
1956:
1952:Polymixiiformes
1916:
1893:
1874:Argentiniformes
1860:
1842:
1794:
1771:
1732:
1727:
1698:
1693:
1669:
1664:
1660:Polypteriformes
1646:
1608:
1601:
1571:
1570:
1527:
1520:
1509:
1505:
1486:
1482:
1473:
1471:
1464:
1460:
1450:Wayback Machine
1440:
1436:
1409:
1405:
1393:
1384:
1375:
1373:
1369:
1344:
1338:
1329:
1320:
1318:
1311:
1295:
1282:
1272:
1270:
1262:
1261:
1257:
1247:
1245:
1226:
1222:
1216:Wayback Machine
1204:
1197:
1192:
1187:
1178:
1169:
1160:
1151:
1142:
1049:
988:
979:
926:Pristolepididae
893:
888:
731:(Bleeker, 1877)
721:(Bleeker, 1850)
630:(Bleeker, 1851)
613:Luciocephalinae
429:
385:breathing organ
307:Pristolepididae
267:ray-finned fish
206:
202:
151:
144:
123:
53:
24:Anabantiformes
17:
16:Order of fishes
12:
11:
5:
2367:
2357:
2356:
2354:Anabantiformes
2339:
2338:
2336:
2335:
2322:
2309:
2296:
2283:
2270:
2265:Anabantiformes
2255:
2239:
2237:
2235:Anabantiformes
2231:
2230:
2216:
2215:
2212:
2211:
2208:
2207:
2204:
2203:
2201:
2200:
2195:
2190:
2185:
2183:Pempheriformes
2180:
2175:
2170:
2165:
2160:
2155:
2150:
2145:
2140:
2135:
2130:
2125:
2120:
2115:
2110:
2105:
2100:
2095:
2090:
2085:
2080:
2078:Atheriniformes
2075:
2070:
2065:
2060:
2055:
2053:Anabantiformes
2050:
2045:
2040:
2035:
2030:
2025:
2020:
2015:
2010:
2005:
1999:
1997:
1991:
1990:
1988:
1987:
1982:
1977:
1972:
1970:Percopsiformes
1966:
1964:
1958:
1957:
1955:
1954:
1949:
1944:
1942:Myctophiformes
1939:
1934:
1928:
1926:
1923:incertae sedis
1918:
1917:
1915:
1914:
1909:
1903:
1901:
1895:
1894:
1892:
1891:
1886:
1881:
1876:
1870:
1868:
1862:
1861:
1859:
1858:
1852:
1850:
1844:
1843:
1841:
1840:
1835:
1830:
1825:
1820:
1815:
1810:
1804:
1802:
1796:
1795:
1793:
1792:
1790:Hiodontiformes
1787:
1781:
1779:
1773:
1772:
1770:
1769:
1767:Anguilliformes
1764:
1759:
1754:
1748:
1746:
1737:
1729:
1728:
1726:
1725:
1720:
1714:
1712:
1703:
1695:
1694:
1692:
1691:
1685:
1683:
1674:
1666:
1665:
1663:
1662:
1656:
1654:
1648:
1647:
1645:
1644:
1638:
1632:
1626:
1620:
1613:
1610:
1609:
1605:Actinopterygii
1600:
1599:
1592:
1585:
1577:
1569:
1568:
1541:(2): 377–388.
1518:
1503:
1480:
1458:
1434:
1403:
1382:
1355:(4): 259–272.
1327:
1309:
1280:
1255:
1220:
1194:
1193:
1191:
1188:
1184:
1183:
1180:
1179:
1175:
1174:
1171:
1170:
1166:
1165:
1162:
1161:
1157:
1156:
1153:
1152:
1148:
1147:
1144:
1143:
1139:
1138:
1135:
1134:
1122:
1119:
1118:
1115:
1114:
1106:Helostomatidae
1102:
1097:
1095:
1092:
1091:
1088:
1087:
1075:
1070:
1068:
1063:
1061:
1055:
1054:
1051:
1050:
1046:
1045:
1042:
1041:
1034:
1031:
1030:
1027:
1026:
1014:
1009:
1007:
999:
997:
994:
993:
990:
989:
985:
984:
981:
980:
976:
975:
972:
971:
959:
956:
955:
952:
951:
944:
939:
937:
934:
933:
930:
929:
922:
917:
915:
907:
905:
903:Anabantiformes
897:
892:
889:
887:
884:
883:
882:
881:
880:
879:
878:
864:
834:
833:
832:
784:
783:
782:
781:
780:
779:
778:
768:
765:(Bleeker, 1879
757:
743:
733:
727:Parosphromenus
723:
713:
703:
676:Macropodusinae
672:
671:
670:
660:
646:
632:
609:
608:
607:
584:
583:
582:
551:van der Hoeven
543:
542:
541:
519:Helostomatidae
515:
514:
513:
499:
489:Microctenopoma
485:
471:
428:
425:
312:incertae sedis
291:Helostomatidae
256:labyrinth fish
200:Anabantiformes
194:
193:
192:
191:
172:
171:
165:
164:
158:
157:
153:
152:
145:
138:
137:
131:
130:
121:Anabantiformes
118:
114:
113:
108:
101:
100:
98:Actinopterygii
95:
91:
90:
85:
81:
80:
75:
71:
70:
65:
61:
60:
47:
46:
39:Climbing perch
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
2366:
2355:
2352:
2351:
2349:
2332:
2327:
2323:
2319:
2314:
2310:
2306:
2301:
2297:
2293:
2288:
2284:
2280:
2275:
2271:
2266:
2260:
2256:
2251:
2245:
2241:
2240:
2238:
2236:
2232:
2228:
2223:
2199:
2196:
2194:
2191:
2189:
2186:
2184:
2181:
2179:
2176:
2174:
2171:
2169:
2166:
2164:
2161:
2159:
2156:
2154:
2151:
2149:
2146:
2144:
2143:Lutjaniformes
2141:
2139:
2136:
2134:
2131:
2129:
2128:Ephippiformes
2126:
2124:
2121:
2119:
2116:
2114:
2111:
2109:
2106:
2104:
2103:Blenniiformes
2101:
2099:
2096:
2094:
2091:
2089:
2086:
2084:
2081:
2079:
2076:
2074:
2071:
2069:
2066:
2064:
2063:Carangiformes
2061:
2059:
2056:
2054:
2051:
2049:
2046:
2044:
2043:Scombriformes
2041:
2039:
2036:
2034:
2031:
2029:
2026:
2024:
2021:
2019:
2018:Ophidiiformes
2016:
2014:
2011:
2009:
2006:
2004:
2001:
2000:
1998:
1996:
1992:
1986:
1983:
1981:
1978:
1976:
1973:
1971:
1968:
1967:
1965:
1963:
1959:
1953:
1950:
1948:
1945:
1943:
1940:
1938:
1935:
1933:
1930:
1929:
1927:
1925:
1924:
1919:
1913:
1910:
1908:
1905:
1904:
1902:
1900:
1896:
1890:
1887:
1885:
1884:Salmoniformes
1882:
1880:
1879:Galaxiiformes
1877:
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1400:0-8120-5635-3
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27:
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2173:Lophiiformes
2168:Caproiformes
2148:Lobotiformes
2123:Moroniformes
2108:Gerreiformes
2093:Mugiliformes
2088:Beloniformes
2073:Cichliformes
2052:
2003:Beryciformes
1947:Lampriformes
1937:Aulopiformes
1921:
1912:Stomiiformes
1907:Osmeriformes
1838:Siluriformes
1808:Clupeiformes
1757:Albuliformes
1642:Osteichthyes
1634:Infraphylum
1538:
1534:
1512:
1506:
1497:
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1483:
1472:. Retrieved
1461:
1453:
1437:
1420:
1416:
1406:
1374:. Retrieved
1367:the original
1352:
1348:
1319:. Retrieved
1315:the original
1299:
1271:. Retrieved
1267:
1258:
1246:. Retrieved
1236:
1223:
1207:
1124:
1104:
1077:
1059:Anabantoidei
1036:
1016:
961:
946:
924:
902:
894:
866:
848:
808:
801:
770:
759:
753:Deraniyagala
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689:Trichogaster
687:
662:
648:
634:
624:
595:
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529:
501:
487:
473:
459:
437:Anabantoidei
430:
410:
402:
389:vascularized
369:parasphenoid
366:
349:
323:monophyletic
316:
310:
255:
199:
197:
187:Valenciennes
178:Labyrinthici
177:
161:
135:Type species
120:
104:
42:
18:
2259:Wikispecies
2193:Perciformes
2153:Spariformes
2118:Labriformes
2033:Gobiiformes
2028:Kurtiformes
1889:Esociformes
1752:Elopiformes
1744:Elopomorpha
1700:Neopterygii
1681:Chondrostei
1671:Actinopteri
1640:Superclass
1423:(2): 1–11.
1264:"Labyrinth"
1248:11 December
1079:Anabantidae
707:Trichopodus
597:Osphronemus
559:Belontiinae
449:Anabantidae
393:respiration
377:anabantoids
330:Ovalentaria
287:Anabantidae
111:Percomorpha
1985:Gadiformes
1800:Otocephala
1723:Amiiformes
1630:Vertebrata
1628:Subphylum
1474:2006-12-23
1470:. FishBase
1376:2017-12-27
1321:2019-12-08
1273:16 January
1190:References
1005:Channoidei
850:Parachanna
791:Berg, 1940
788:Channoidei
772:Trichopsis
747:Malpulutta
737:Macropodus
674:Subfamily
656:McClelland
642:Canestrini
611:Subfamily
586:Subfamily
557:Subfamily
383:accessory
373:snakeheads
264:freshwater
258:, are an
1975:Zeiformes
1734:Teleostei
1652:Cladistia
1555:1175-5334
1429:0272-4634
1018:Channidae
913:Nandoidei
891:Phylogeny
838:Channidae
824:Dahanukar
786:suborder
699:Schneider
531:Helostoma
509:Castelnau
475:Ctenopoma
453:Bonaparte
435:suborder
417:gouramies
381:branchial
279:Channidae
271:suborders
269:with two
162:see text
74:Kingdom:
68:Eukaryota
2348:Category
2292:39721936
2250:Q1744606
2244:Wikidata
1899:Stomiati
1710:Holostei
1624:Chordata
1618:Animalia
1616:Kingdom
1563:31717234
1446:Archived
1233:"Anabas"
1212:Archived
963:Nandidae
828:Raghavan
681:Hoedeman
603:Lacepède
572:Belontia
503:Sandelia
427:Taxonomy
395:in air.
375:and the
299:Nandidae
275:families
169:Synonyms
150:, 1792)
88:Chordata
84:Phylum:
78:Animalia
64:Domain:
2318:5554917
2305:1489900
1622:Phylum
1603:Extant
1535:Zootaxa
1494:Zootaxa
948:Badidae
876:, 1777)
874:Scopoli
862:, 1984)
856:Teugels
836:family
830:, 2019)
795:family
755:, 1937)
701:, 1801)
683:, 1948)
658:, 1845)
650:Ctenops
644:, 1860)
620:, 1852)
618:Bleeker
605:, 1801)
580:, 1923)
566:, 1962)
545:family
539:, 1829)
517:family
511:, 1861)
497:, 1995)
483:, 1844)
469:, 1816)
467:Cloquet
447:family
303:Badidae
273:, five
189:, 1828)
117:Order:
94:Class:
2331:266131
2279:747030
1561:
1553:
1427:
1398:
1307:
1218:(2016)
868:Channa
858:&
844:, 1934
842:Fowler
804:, 2020
802:et al.
800:Britz
697:&
553:, 1832
537:Cuvier
525:, 1872
495:Norris
481:Peters
461:Anabas
455:, 1831
443:, 1940
371:. The
358:Africa
293:, and
185:&
183:Cuvier
128:, 1995
1456:4(1).
1370:(PDF)
1345:(PDF)
860:Daget
820:Anoop
816:Britz
717:Betta
695:Bloch
578:Myers
326:clade
260:order
148:Bloch
126:Britz
105:Clade
2300:NCBI
2274:BOLD
1559:PMID
1551:ISSN
1539:4603
1498:3882
1425:ISSN
1396:ISBN
1305:ISBN
1275:2022
1250:2019
826:and
564:Liem
523:Gill
441:Berg
356:and
354:Asia
344:and
305:and
198:The
2287:EoL
1543:doi
1357:doi
1268:the
238:ɔːr
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2289::
2276::
2261::
2246::
1557:.
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1533:.
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1496:.
1492:.
1452:.
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1198:^
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217:ˈ
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