37:
1204:; Crawford, Daniel J.; Ruiz, Eduardo A. (2018). "Chapter 13: Patterns of Phylogeny". In Stuessy, Tod F.; Crawford, Daniel J.; LĂłpez-SepĂşlvada, Patricio; Baeza, Carlos M.; Ruiz, Eduardo A. (eds.).
269:. It has leathery, thrice-divided leaves with dark brown axes; the leaves are coated with white powder below. First described as a species in 1853, it was transferred to the new genus
316:
The fronds arise in clumps from the rhizome. From base to tip of leaf, they are 5 to 15 centimeters (2.0 to 5.9 in) long. Of this length, about half is made up by the
335:(cut into pinnae, pinnules, and pinnulets), typically bearing about five pairs of pinnae. The texture of the blade is leathery; the underside is densely covered in white
1150:; Huiet, Layne; Yatskievych, George; Pryer, Kathleen M. (2011). "Species Relationships and Farina Evolution in the Cheilanthoid Fern Genus Argyrochosma (Pteridaceae)".
426:, attributing that name to FĂ©e. While he did not explain his choice of epithet, it presumably reflects the Chilean origin of the specimens he examined. In 1858,
366:
lie along the veins, in the one-third to one-fourth of the vein closest to the leaf edge. The leaf edges are curved under, but not otherwise modified into false
1543:
1147:
573:
159:
1201:
1435:
359:(blunt) at the tip, and are generally attached directly to the costule (pinnule axis), without stalks. They are not jointed at the base.
1206:
Plants of
Oceanic Islands: Evolution, Biogeography and Conservation of the Flora of the Juan Fernández (Robinson Crusoe) Archipelago
1396:
374:
contains 64 spores. The spores are covered with a network of raised crests, although portions of the surface lack elevated crests.
994:
717:
based on morphology extremely difficult, and scientists have frequently disagreed on the placement of species such as this.
1325:
1018:
383:
320:(the stalk of the leaf, below the blade), which is round, a dark chestnut brown in color, and lacks scales and hairs.
151:
1533:
1440:
340:
309:
scales 3 millimeters (0.1 in) long that terminate in a hair attached by a joint, of a bright brown color, with
1374:
348:
1422:
356:
352:
310:
306:
473:
which, however, had a broader blade, more heavily dissected and with more leaf tissue, than the leaves of true
328:
324:
1479:
1004:
441:
245:
36:
680:
It is found on rocks in dry, exposed situations, at altitudes from 20 to 40 meters (70 to 100 ft).
1474:
974:
595:
215:
204:
1319:
666:
262:
674:
651:
551:
608:, as part of a program to consolidate the cheilanthoid ferns into that genus. He also transferred
1538:
1103:
688:
The horticulturist George
Schneider considered the species suitable for greenhouse cultivation.
1336:
670:
639:
490:
1528:
1237:
1187:
498:
193:
134:
1453:
1213:
523:
427:
226:
987:
Plant
Gateway's the Global Flora: A practical flora to vascular plant species of the world
336:
275:(the "false cloak ferns") in 1987, recognizing their distinctness from the "cloak ferns" (
8:
1466:
1066:
978:
518:
391:
351:(diamond-shaped) terminal pinnule; the other pinnules are cut into pinnulets, which are
1500:
1300:
1267:
1259:
1233:
1175:
1167:
1123:
1090:
1052:
624:
536:
464:
189:
167:
31:
1448:
1383:
1271:
990:
514:
1241:
1217:
1191:
1179:
1136:
1127:
1102:
Morbelli, Marta A.; Ponce, M. MĂłnica; MacLuf, C. Cecilia; Piñeiro, MarĂa R. (2001).
1070:
1056:
1042:
1022:
1505:
1292:
1249:
1159:
1115:
1082:
1008:
567:
469:
1388:
566:
and
Weatherby himself had suggested in the 1940s that a group of ferns related to
1359:
446:
1280:
1104:"Palynology of South American Argyrochosma and Notholaena (Pteridaceae) species"
414:. RĂ©my published his flora the following year, in the sixth botanical volume of
1461:
563:
317:
61:
1119:
1522:
1163:
982:
456:
282:
401:
155:
1492:
927:
714:
586:
271:
111:
91:
493:. He attributed its introduction to British cultivation to John Riley, of
1409:
1368:
1038:
576:, who was carrying out phylogenetic studies of these genera. He elevated
482:
396:
101:
1263:
1171:
572:
might represent a distinct genus. This was finally addressed in 1987 by
1401:
1304:
1094:
643:
630:
600:
494:
432:
415:
371:
332:
298:
277:
1414:
347:(leaf axis) by short stalks. Each pinna is tipped with a three-lobed,
1254:
759:
710:
540:
478:
1330:
1296:
1086:
339:(powder), which is absent from the upper side. The pinnae are ovate-
1487:
1353:
1027:
Mémoires de la Société du muséum d'histoire naturelle de
Strasbourg
406:
367:
1427:
939:
915:
903:
411:
404:, who was preparing a flora of Chile, considered it to belong to
302:
1138:
The book of choice ferns for the garden, conservatory and stove
467:
described a fern being cultivated in
Britain under the name of
344:
749:
747:
745:
743:
741:
739:
737:
735:
733:
363:
266:
48:
1222:
1218:"Enumeratio plantarum vascularum cryptogamicarum Chilensium"
1145:
933:
562:
into sections based on the data available at the time, both
1246:
Contributions from the Gray
Herbarium of Harvard University
855:
730:
539:
designated a specimen of Riley's cultivation at Kew as the
327:, broadest at the lowest or second pair of pinnae, with an
81:
71:
1208:. Cambridge: Cambridge University Press. pp. 209–275.
1101:
765:
807:
951:
477:. He was unable to find a corresponding specimen in the
619:
A phylogenetic analysis including a single specimen of
1047:. Vol. Botanica v. 6. Paris: E. Thunot & Co.
891:
879:
867:
819:
485:, and described it as a new species, which he called
543:, and concluded that it was in fact synonymous with
1200:
1013:. Vol. Fascicles 1-12. Copenhagen: H. Hagerup.
973:
945:
921:
909:
831:
795:
558:, considered it impossible to reasonably subdivide
1242:"A revision of the American species of Notholaena"
843:
1281:"Argyrochosma, a new genus of cheilanthoid ferns"
783:
771:
1520:
590:, and transferred this species to that genus as
382:The first scientist to mention the species was
1232:
861:
753:
430:transferred it into a broadly-construed genus
531:, as named by Eaton, was simply a synonym of
1061:. Vol. 1. London: Groombridge and Sons.
623:found it nested within a clade representing
1544:Endemic flora of the Juan Fernández Islands
1003:
813:
713:has made the delimitation of genera in the
657:
634:. It probably evolved by the dispersal of
1253:
1196:(1st ed.). London: Robert Hardwicke.
1134:
957:
16:Species of fern in the family Pteridaceae
521:, when devising the replacement name of
1278:
897:
885:
873:
691:
1521:
1141:. Vol. 2. London: L. Upcott Gill.
527:for Eaton's fern in 1915, opined that
1335:
1334:
1212:
1186:
1065:
849:
837:
801:
305:is stout and upright. It bears thin,
1051:
825:
1044:Historia fisica y politica de Chile
1037:
1017:
789:
777:
420:Historia fisica y politica de Chile
13:
14:
1555:
1313:
946:Stuessy, Crawford & Ruiz 2018
922:Christenhusz, Fay & Byng 2018
910:Christenhusz, Fay & Byng 2018
1480:urn:lsid:ipni.org:names:272600-2
35:
1023:"Genera filicum. Polypodiacées"
422:, and described the species as
386:in 1852, who referred to it as
384:Antoine Laurent Apollinaire FĂ©e
966:
703:
683:
650:and its spread to the younger
444:, by contrast, assigned it to
289:
1:
724:
1279:Windham, Michael D. (1987).
1193:Ferns, British & Foreign
1071:"Note on American Ferns: IX"
550:While Tryon, when finishing
7:
975:Christenhusz, Maarten J. M.
598:transferred the species to
513:" was mistakenly reused by
377:
10:
1560:
1135:Schneider, George (1892).
862:Tryon & Weatherby 1956
754:Tryon & Weatherby 1956
596:Maarten J. M. Christenhusz
343:to ovate, attached to the
1343:
1120:10.1080/00173130152987517
1058:Ferns: British and Exotic
261:is a fern endemic to the
173:
166:
140:
133:
32:Scientific classification
30:
23:
1534:Plants described in 1853
1164:10.1600/036364411X583547
696:
675:Alejandro Selkirk Island
658:Distribution and habitat
652:Alejandro Selkirk Island
554:'s revision of American
552:Charles Alfred Weatherby
331:(pointed) tip. They are
1326:Endemic Plants of Chile
1322:at JSTOR Plant Science.
410:, in the sense used by
1375:Argyrochosma chilensis
1345:Argyrochosma chilensis
1320:Isotype of the species
671:Robinson Crusoe Island
667:Juan Fernández Islands
663:Argyrochosma chilensis
640:Robinson Crusoe Island
491:William Jackson Hooker
295:Argyrochosma chilensis
263:Juan Fernández Islands
258:Argyrochosma chilensis
144:Argyrochosma chilensis
25:Argyrochosma chilensis
1285:American Fern Journal
1075:American Fern Journal
313:(toothless) margins.
766:Morbelli et al. 2001
692:Notes and references
584:to become the genus
524:Notholaena standleyi
428:Johann Wilhelm Sturm
424:Cincinalis chilensis
394:(making that name a
323:The leaf blades are
222:Notholaena chilensis
200:Hemionitis chilensis
177:Cincinalis chilensis
1148:Windham, Michael D.
519:William Ralph Maxon
392:species description
390:without offering a
828:, pp. 34, 59.
665:is endemic to the
574:Michael D. Windham
537:Rolla M. Tryon Jr.
501:transferred it to
487:Notholaena hookeri
465:Edward Joseph Lowe
400:), but noted that
233:Notholaena hookeri
211:Hemionitis hookeri
185:Cincinalis hookeri
1516:
1515:
1449:Open Tree of Life
1337:Taxon identifiers
1238:Weatherby, Una F.
1152:Systematic Botany
1067:Maxon, William R.
1005:Christensen, Carl
996:978-0-9929993-9-1
934:Sigel et al. 2011
648:A. chilensis
621:A. chilensis
606:H. chilensis
592:A. chilensis
545:N. chilensis
515:Daniel Cady Eaton
452:P. chilensis
438:N. chilensis
388:Pellaea chilensis
265:off the coast of
254:
253:
248:
241:Pellaea chilensis
237:
229:
218:
207:
196:
181:
126:A. chilensis
1551:
1509:
1508:
1496:
1495:
1483:
1482:
1470:
1469:
1457:
1456:
1444:
1443:
1431:
1430:
1418:
1417:
1405:
1404:
1392:
1391:
1379:
1378:
1377:
1364:
1363:
1362:
1332:
1331:
1308:
1275:
1257:
1255:10.5962/p.336378
1229:
1209:
1197:
1183:
1146:Sigel, Erin M.;
1142:
1131:
1098:
1062:
1048:
1034:
1014:
1000:
961:
955:
949:
943:
937:
931:
925:
919:
913:
907:
901:
895:
889:
883:
877:
871:
865:
859:
853:
847:
841:
835:
829:
823:
817:
814:Christensen 1906
811:
805:
799:
793:
787:
781:
775:
769:
763:
757:
751:
718:
707:
470:Notholaena nivea
442:Carl Christensen
244:(FĂ©e ex J.RĂ©my)
243:
235:
225:(FĂ©e ex J.RĂ©my)
224:
213:
203:(FĂ©e ex J.RĂ©my)
202:
187:
180:FĂ©e ex J.RĂ©my
179:
146:
40:
39:
21:
20:
1559:
1558:
1554:
1553:
1552:
1550:
1549:
1548:
1519:
1518:
1517:
1512:
1504:
1499:
1491:
1486:
1478:
1473:
1465:
1460:
1452:
1447:
1439:
1434:
1426:
1421:
1413:
1408:
1400:
1395:
1387:
1382:
1373:
1372:
1367:
1358:
1357:
1352:
1339:
1316:
1311:
1297:10.2307/1547438
1234:Tryon, Rolla M.
1202:Stuessy, Tod F.
1087:10.2307/1544458
997:
989:. Vol. 4.
979:Fay, Michael F.
969:
964:
956:
952:
944:
940:
932:
928:
920:
916:
908:
904:
896:
892:
884:
880:
872:
868:
860:
856:
848:
844:
836:
832:
824:
820:
812:
808:
800:
796:
788:
784:
776:
772:
764:
760:
752:
731:
727:
722:
721:
708:
704:
699:
694:
686:
669:, particularly
660:
614:H. hookeri
610:N. hookeri
529:N. hookeri
511:N. hookeri
507:C. hookeri
380:
292:
162:
148:
142:
129:
34:
17:
12:
11:
5:
1557:
1547:
1546:
1541:
1539:Ferns of Chile
1536:
1531:
1514:
1513:
1511:
1510:
1506:wfo-0001115376
1497:
1484:
1471:
1458:
1445:
1432:
1419:
1406:
1393:
1380:
1365:
1349:
1347:
1341:
1340:
1329:
1328:
1323:
1315:
1314:External links
1312:
1310:
1309:
1276:
1248:(179): 1–106.
1230:
1210:
1198:
1184:
1158:(3): 554–564.
1143:
1132:
1114:(6): 280–291.
1099:
1063:
1049:
1035:
1015:
1001:
995:
983:Byng, James W.
970:
968:
965:
963:
962:
960:, p. 605.
958:Schneider 1892
950:
948:, p. 259.
938:
936:, p. 562.
926:
914:
902:
890:
878:
866:
854:
842:
840:, p. 178.
830:
818:
816:, p. 479.
806:
804:, p. 166.
794:
792:, p. 497.
782:
780:, p. 129.
770:
758:
728:
726:
723:
720:
719:
701:
700:
698:
695:
693:
690:
685:
682:
659:
656:
642:, followed by
564:Edwin Copeland
489:, in honor of
379:
376:
291:
288:
252:
251:
250:
249:
238:
230:
219:
208:
197:
182:
171:
170:
164:
163:
149:
138:
137:
131:
130:
123:
121:
117:
116:
109:
105:
104:
99:
95:
94:
89:
85:
84:
82:Polypodiopsida
79:
75:
74:
72:Polypodiophyta
69:
65:
64:
59:
52:
51:
46:
42:
41:
28:
27:
15:
9:
6:
4:
3:
2:
1556:
1545:
1542:
1540:
1537:
1535:
1532:
1530:
1527:
1526:
1524:
1507:
1502:
1498:
1494:
1489:
1485:
1481:
1476:
1472:
1468:
1463:
1459:
1455:
1450:
1446:
1442:
1437:
1433:
1429:
1424:
1420:
1416:
1411:
1407:
1403:
1398:
1394:
1390:
1385:
1381:
1376:
1370:
1366:
1361:
1355:
1351:
1350:
1348:
1346:
1342:
1338:
1333:
1327:
1324:
1321:
1318:
1317:
1306:
1302:
1298:
1294:
1290:
1286:
1282:
1277:
1273:
1269:
1265:
1261:
1256:
1251:
1247:
1243:
1239:
1235:
1231:
1227:
1223:
1219:
1215:
1211:
1207:
1203:
1199:
1195:
1194:
1189:
1185:
1181:
1177:
1173:
1169:
1165:
1161:
1157:
1153:
1149:
1144:
1140:
1139:
1133:
1129:
1125:
1121:
1117:
1113:
1109:
1105:
1100:
1096:
1092:
1088:
1084:
1080:
1076:
1072:
1068:
1064:
1060:
1059:
1054:
1050:
1046:
1045:
1040:
1036:
1032:
1028:
1024:
1020:
1019:FĂ©e, A. L. A.
1016:
1012:
1011:
1010:Index Filicum
1006:
1002:
998:
992:
988:
984:
980:
976:
972:
971:
959:
954:
947:
942:
935:
930:
924:, p. 15.
923:
918:
912:, p. 11.
911:
906:
900:, p. 40.
899:
894:
888:, p. 38.
887:
882:
876:, p. 37.
875:
870:
863:
858:
851:
846:
839:
834:
827:
822:
815:
810:
803:
798:
791:
786:
779:
774:
767:
762:
756:, p. 81.
755:
750:
748:
746:
744:
742:
740:
738:
736:
734:
729:
716:
715:cheilanthoids
712:
706:
702:
689:
681:
678:
676:
672:
668:
664:
655:
653:
649:
645:
641:
637:
636:A. nivea
633:
632:
628:
627:
626:A. nivea
622:
617:
615:
611:
607:
603:
602:
597:
593:
589:
588:
583:
579:
575:
571:
570:
569:N. nivea
565:
561:
557:
553:
548:
546:
542:
538:
534:
533:N. nivea
530:
526:
525:
520:
516:
512:
508:
504:
500:
496:
492:
488:
484:
480:
476:
475:N. nivea
472:
471:
466:
461:
459:
458:
457:Index Filicum
453:
449:
448:
443:
439:
435:
434:
429:
425:
421:
417:
413:
409:
408:
403:
399:
398:
393:
389:
385:
375:
373:
369:
365:
360:
358:
354:
350:
346:
342:
338:
334:
330:
326:
321:
319:
314:
312:
308:
304:
300:
296:
287:
285:
284:
283:sensu stricto
280:
279:
274:
273:
268:
264:
260:
259:
247:
242:
239:
234:
231:
228:
223:
220:
217:
212:
209:
206:
201:
198:
195:
191:
186:
183:
178:
175:
174:
172:
169:
165:
161:
157:
153:
147:
145:
139:
136:
135:Binomial name
132:
128:
127:
122:
119:
118:
115:
114:
110:
107:
106:
103:
100:
97:
96:
93:
90:
87:
86:
83:
80:
77:
76:
73:
70:
67:
66:
63:
62:Tracheophytes
60:
57:
54:
53:
50:
47:
44:
43:
38:
33:
29:
26:
22:
19:
1529:Argyrochosma
1467:tro-26609900
1344:
1291:(2): 37–41.
1288:
1284:
1245:
1225:
1221:
1214:Sturm, J. W.
1205:
1192:
1155:
1151:
1137:
1111:
1107:
1078:
1074:
1057:
1043:
1030:
1026:
1009:
986:
953:
941:
929:
917:
905:
898:Windham 1987
893:
886:Windham 1987
881:
874:Windham 1987
869:
864:, p. 6.
857:
852:, p. 1.
845:
833:
821:
809:
797:
785:
773:
761:
705:
687:
679:
662:
661:
647:
635:
629:
625:
620:
618:
613:
609:
605:
599:
591:
587:Argyrochosma
585:
582:Argyrochosma
581:
577:
568:
559:
555:
549:
544:
532:
528:
522:
510:
509:. The name "
506:
502:
486:
474:
468:
462:
455:
451:
445:
437:
431:
423:
419:
405:
395:
387:
381:
361:
322:
315:
294:
293:
281:
276:
272:Argyrochosma
270:
257:
256:
255:
240:
232:
221:
210:
199:
184:
176:
143:
141:
125:
124:
113:Argyrochosma
112:
92:Polypodiales
55:
24:
18:
1410:iNaturalist
1369:Wikispecies
1188:Smith, John
1053:Lowe, E. J.
1039:Gay, Claude
967:Works cited
709:Widespread
684:Cultivation
594:. In 2018,
497:. In 1866,
397:nomen nudum
297:is a small
290:Description
236:E.J.Lowe
214:(E.J.Lowe)
102:Pteridaceae
1523:Categories
1462:Plant List
1228:: 151–202.
1081:(1): 1–2.
850:Maxon 1915
838:Smith 1866
802:Sturm 1858
725:References
644:anagenesis
631:sensu lato
601:Hemionitis
578:Notholaena
560:Notholaena
556:Notholaena
503:Cincinalis
499:John Smith
495:Papplewick
433:Notholaena
416:Claude Gay
402:Jules RĂ©my
372:sporangium
341:lanceolate
333:tripinnate
301:fern. The
278:Notholaena
216:Christenh.
205:Christenh.
68:Division:
1360:Q15597061
1272:249085059
826:Lowe 1856
711:homoplasy
646:creating
541:lectotype
479:herbarium
463:In 1856,
460:of 1906.
299:epipetric
227:J.W.Sturm
120:Species:
45:Kingdom:
1493:26609900
1488:Tropicos
1428:272600-2
1354:Wikidata
1264:41764632
1240:(1956).
1216:(1858).
1190:(1866).
1180:16214744
1172:23028975
1128:85155458
1069:(1915).
1055:(1856).
1041:(1853).
1021:(1852).
1007:(1906).
985:(2018).
790:Gay 1853
778:FĂ©e 1852
407:Lindsaea
378:Taxonomy
349:rhomboid
190:E.J.Lowe
168:Synonyms
98:Family:
1441:1055470
1402:3750272
1305:1547438
1095:1544458
454:in his
447:Pellaea
412:Desvaux
370:. Each
368:indusia
303:rhizome
160:Windham
108:Genus:
88:Order:
78:Class:
49:Plantae
1454:420001
1415:963388
1303:
1270:
1262:
1178:
1170:
1126:
1093:
993:
580:sect.
357:obtuse
353:oblong
345:rachis
337:farina
311:entire
307:linear
246:C.Chr.
156:J.RĂ©my
1301:JSTOR
1268:S2CID
1260:JSTOR
1176:S2CID
1168:JSTOR
1124:S2CID
1108:Grana
1091:JSTOR
697:Notes
612:, as
329:acute
325:ovate
318:stipe
267:Chile
194:J.Sm.
56:Clade
1475:POWO
1436:NCBI
1423:IPNI
1397:GBIF
1389:GJMN
991:ISBN
673:and
364:sori
362:The
1501:WFO
1384:CoL
1293:doi
1250:doi
1160:doi
1116:doi
1083:doi
638:to
604:as
505:as
483:Kew
481:at
450:as
436:as
418:'s
286:).
154:ex
152:FĂ©e
1525::
1503::
1490::
1477::
1464::
1451::
1438::
1425::
1412::
1399::
1386::
1371::
1356::
1299:.
1289:77
1287:.
1283:.
1266:.
1258:.
1244:.
1236:;
1224:.
1220:.
1174:.
1166:.
1156:36
1154:.
1122:.
1112:40
1110:.
1106:.
1089:.
1077:.
1073:.
1029:.
1025:.
981:;
977:;
732:^
677:.
654:.
616:.
547:.
535:.
517:;
440:.
355:,
192:)
158:)
58::
1307:.
1295::
1274:.
1252::
1226:2
1182:.
1162::
1130:.
1118::
1097:.
1085::
1079:5
1033:.
1031:5
999:.
768:.
188:(
150:(
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