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Gastropod shell

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ones, and generally are absent in the young shell, except in some highly accelerated types. Tertiary spirals are intercalated between the preceding groups in more specialized species. Ribs are regular transverse foldings of the shell, which generally extend from the suture to suture. They are usually spaced uniformly and crossed by the spirals. In specialized types, when a shoulder angle is formed, they become concentrated as nodes upon this angle, disappearing from the shoulder above and the body below. Spines may replace the nodes in later stages. They form as notches in the margin of the shell and are subsequently abandoned, often remaining open in front. Irregular spines may also arise on various parts of the surface of the shell (see
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Tanke, D.H. and Brett-Surman, M.K. 2001. Evidence of Hatchling and Nestling-Size Hadrosaurs (Reptilia:Ornithischia) from Dinosaur Provincial Park (Dinosaur Park Formation: Campanian), Alberta, Canada. pp. 206–218. In: Mesozoic Vertebrate Life – New Research Inspired by the Paleontology of Philip J.
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of the gastropod shell consists of longitudinal ridges, and/or transverse ridges. Primary spirals may appear in regular succession on either side of the first primary, which generally becomes the shoulder angle if angulation occurs. Secondary spirals may appear by intercalation between the primary
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of the shell gradually increase in size. Normally the whorls are circular or elliptical in section. The spire can be high or low, broad or slender, according to the way the coils of the shell are arranged, and the apical angle of the shell varies accordingly. The whorls sometimes rest loosely upon
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living in isolated colonies on limestone outcrops. Several sets of species differ only in the direction of coiling, but the evidence is inconclusive as to whether left- and right-handed shells live together. Soos (1928, pp. 372–385) summarized previous discussions of the problem and concluded
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In a few cases, both left- and right-handed coiling are found in the same population. Sinistral mutants of normally dextral species and dextral mutants of normally sinistral species are rare but well documented occurrences among land snails in general. Populations or species with normally mixed
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The apertural end of the gastropod shell is the anterior end, nearest to the head of the animal; the apex of the spire is often the posterior end or at least is the dorsal side. Most authors figure the shells with the apex of the spire uppermost. In life, when the soft parts of these snail are
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In this context, the height (or the length) of a shell is its maximum measurement along the central axis. The width (or breadth, or diameter) is the maximum measurement of the shell at right angles to the central axis. Both terms are only related to the description of the shell and not to the
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If a coiled gastropod shell is held with the spire pointing upwards and the aperture more or less facing the observer, a dextral shell will have the aperture on the right-hand side, and a sinistral shell will have the aperture on the left-hand side. This chirality of gastropods is sometimes
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zone, are usually inhabited by snails whose shells have a wide aperture, a relatively low surface area, and a high growth rate per revolution. High-spired and highly sculptured forms become more common in quiet water environments. The shell of burrowing forms, such as the
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or peristome of the shell may be simple or variously modified. An outer and an inner (columellar) lip are generally recognized. These may be continuous with each other, or may be divided by an anterior notch. This, in some types
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overlooked when photographs of coiled gastropods are "flipped" by a non-expert prior to being used in a publication. This image "flipping" results in a normal dextral gastropod appearing to be a rare or abnormal sinistral one.
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The rate of translation of the generating curve along the axis of coiling, controlling how high-spired the resulting shell becomes. This may range from zero, a flat planispiral shell, to nearly the diameter of the
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Irregularities or "sculpturing" such as ribs, spines, knobs, and varices made by the snail regularly changing the shape of the generating curve during the course of growth, for instance in the many species of
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In large enough quantities, gastropod shells can have enough of an impact on environmental conditions to affect the ability of organic remains in the local environment to fossilize. For example, in the
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Schematic representation of the apical, apertural and basal view of a shell, showing 14 different commonly used measurements. Dotted lines represent the orientation axes (except of lower image).
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that the right- and left-handed populations were distinct species. Others have stated that these populations were not distinct, and the question is far from settled. The Peruvian clausiliid,
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is an imaginary axis along the length of a shell, around which, in a coiled shell, the whorls spiral. The central axis passes through the columella, the central pillar of the shell.
1313:). They also can overlap the earlier whorls such that the earlier whorls may be largely or wholly covered by the later ones. When an angulation occurs, the space between it and the 1044:. This left-handedness seems to be more common in freshwater and land pulmonates. But still the dextral living species in gastropods seem to account for 99% of the total number. 1021:) show a mixture of dextral and sinistral individuals. There occur also aberrantly sinistral forms of dextral species and some of these are highly sought by shell collectors. 2021: 1055:
is involved. A more recent study (2013) correlates the asymmetric coiling of the shell by the left-right asymmetric expression of the decapentaplegic gene in the mantle.
2469: 2618: 1568:: in shells where the whorls move apart as they grow, on the underside of the shell there is a deep depression reaching up towards the spire; this is the umbilicus 139:, which protects from predators, mechanical damage, and dehydration, but also serves for muscle attachment and calcium storage. Some gastropods appear shell-less ( 2769:
Wilson A. B., Glaubrecht M. & Meyer A. (March 2004) "Ancient lakes as evolutionary reservoirs: evidence from the thalassoid gastropods of Lake Tanganyika".
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sites, both of which are predominated by the preserved shells of invertebrate life, including gastropods. It was the slow dissolution of these shells releasing
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Some shell shapes are found more often in certain environments, though there are many exceptions. Wave-washed high-energy environments, such as the rocky
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above it constitutes the area known as the "shoulder" of the shell. The shoulder angle may be smooth or keeled, and may sometimes have nodes or spines.
2721: 2405:"How to Know the eastern land snails; pictured-key for determining the land snails of the United States occurring east of the Rocky Mountain Divide" 2812:
Pfenninger M., Hrabáková M., Steinke D. & Dèpraz A. (4 November 2005) "Why do snails have hairs? A Bayesian inference of character evolution".
3091:– Information on some gastropods of the tropical Western Atlantic, specifically the Caribbean Sea, with relevance to the fisheries in that region 624: 2157: 3094: 3042:
van Batenburg1 F. H. D. & Gittenberger E. (1996). "Ease of fixation of a change in coiling: computer experiments on chirality in snails".
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When a row of spines is formed at the edge or outer lip of the shell during a resting period, this feature sometimes remains behind as a
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coiling are much rarer, and, so far as is known, are confined, with one exception, to a few genera of arboreal tropical snails. Besides
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Keisuke Shimizu, Minoru Iijima, Davin HE SetiamargaIsao Sarashina, Tetsuhiro Kudoh, Takahiro Asami, Edi Gittenberger, Kazuyoshi Endo;
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shells, when the animal is alive or the shell is freshly empty, have an uppermost shell layer of horny, smooth, or hairy epidermis or
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An upper or posterior notch is present in certain taxa, and this may result in the formation of a ridge or shelf next to the suture (
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Some of these factors can be modelled mathematically and programs exist to generate extremely realistic images. Early work by
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Britton J. C (1995) "The relationship between position on shore and shell ornamentation in 2 size-dependent morphotypes of
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The shape of the generating curve, roughly equivalent to the shape of the aperture. It may be round, for instance in the
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The overall shape of the shell varies. For example, three groups can be distinguished based on the height – width ratio:
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Coeloconoid a slightly concave conical shell in which the incremental angle increases steadily during growth (see:
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Turreted, turriculate, babylonic; an elongated shell with the whorls angulated or shouldered on their upper part.
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of the shell, the height of the aperture and the width of the aperture. The number of whorls is also often used.
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on the analog computer also revealed many possible combinations that were never adopted by any actual gastropod.
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Gastropod shell morphology is usually quite constant among individuals of a species. Controlling variables are:
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vegetation would have caused the ancient waters to become acidic. Eggshell fragments are present in only two
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The rate of growth per revolution around the coiling axis. High rates give wide-mouthed forms such as the
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Shells of two different species of sea snail: on the left is the normally sinistral (left-handed) shell of
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Ontologic growth changes as the animal reaches adulthood. Good examples are the flaring lip of the adult
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Wandelt J. & Nagy L. M. (24 August 2004) "Left-Right Asymmetry: More Than One Way to Coil a Shell".
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In photographs or illustrations, a gastropod shell can be shown oriented in a number of standard ways:
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Currie. Edited by D.H. Tanke and K. Carpenter. Indiana University Press: Bloomington. xviii + 577 pp.
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Collin R. & Cipriani R. (22 December 2003) "Dollo's law and the re-evolution of shell coiling".
1523:: the nuclear or embryonic whorl; the larval shell, often remains in position even on an adult shell 2672:
Hershler R. & Ponder W. F.(1998). "A Review of Morphological Characters of Hydrobioid Snails".
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Biconic: shaped like two conical shapes that are touching their bases, and tapering at both ends:
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Goodfriend G. A. (1986) "Variation in land-snail shell form and size and its causes – a Review".
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This article incorporates public domain text from references, and CC-BY-2.0 text from reference.
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high enough to prevent the eggshell fragments from dissolving before they could be fossilized.
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Gibbous. Whorls swelled beyond the normal contour of increase (usually on the aperture side ).
778: 603: 2728:". The Marine Flora and Fauna of Dampier, Western Australia. Western Australian Museum, Perth. 1973: 1961: 3338: 3217: 3128:
Antonio Ruiz Ruiz, Ángel Cárcaba Pozo, Ana I. Porras Crevillen & José R. Arrébola Burgos
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retracted, in some groups the aperture of the shell is closed by using a horny or calcareous
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Most gastropod shells are spirally coiled. The majority (over 90%) of gastropod species have
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Pilsbry (1949, pp. 216–217), also has been described as being an amphidromine species.
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globose or conical shell – the height and the width of the shell are approximately the same
373: 147:). Some snails also possess an operculum that seals the opening of the shell, known as the 1925: 8: 3556: 3485: 3450: 3297: 3262: 2418: 2377: 2218: 1872: 1647: 1565: 1526: 1505:: a thin layer of organic "skin" which forms the outer layer of the shell of many species 1484: 1399: 1321: 1314: 1225: 1110: 1040:
Sinistrality arose independently 19 times among marine gastropods since the start of the
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Schilthuizen M. & Davison A. (2005). "The convoluted evolution of snail chirality".
3475: 3197: 3044: 1985: 1880: 1722: 1552: 1456: 1352: 1011:(right-handed) shells, but a small minority of species and genera are virtually always 828: 818: 669: 635: 552: 545: 456: 305: 233: 148: 115: 93: 83: 35: 2054: 999:, cement the shell to, and grow along, solid surfaces such as rocks, or other shells. 3510: 3414: 3369: 3252: 3135: 2855: 2426: 2285: 2193: 1913: 1888: 1829: 1813: 1663: 1514: 1309: 1282: 833: 743: 412: 346: 271: 190: 78: 53: 47: 41: 2470:"The Geography of Evolutionary Opportunity: Hypothesis and Two Cases in Gastropods". 109: 3500: 3424: 3419: 3384: 3292: 3212: 3115: 3072: 3052: 2982: 2848: 2820: 2777: 2753: 2682: 2534: 2476: 2434: 2253: 2046: 1705: 1145: 1081: 793: 783: 763: 380: 300: 186: 172: 31: 1562:
Teleoconch : the entire shell without the protoconch; the postnuclear whorls.
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include mechanical stability, defense against predators and climatic selection.
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Convolute: aperture as long as the shell, nearly or quite concealing the spire.
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The genetics of reverse coiling in a rare dextral mutant of another clausiliid,
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The shells of some gastropods have been reduced or partly reduced during their
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The following are the principal modifications of form in the gastropod shell.
1149:(Montagu), has been studied by Degner (1952). The mechanism is the same as in 3591: 3571: 2925: 2916: 2269: 2149: 1754: 1376: 1151: 823: 768: 608: 571: 434: 392: 363: 358: 176: 152: 66: 2334: 2077: 3237: 2859: 2852: 2824: 2781: 2538: 2480: 2430: 2372:
Thin section in plane-polarized light of microscopic gastropod shell, from
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are sometimes found, and these occasionally may be strong and tooth-like (
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Some taxa lost the coiling of their shell during evolution. According to
1671: 1435: 1371: 1100: 1066: 983: 921: 788: 773: 716: 630: 397: 329: 136: 3056: 1024: 279: 159:. The biological study of gastropods, and other molluscs in general, is 3515: 3490: 3409: 3399: 3328: 3267: 3257: 3207: 3180: 2997: 2956: 2940: 2757: 2584: 2236: 2185: 1839: 1821: 1771: 1730: 1520: 1478: 1345: 1327: 1295: 1130: 1115: 996: 978: 971: 932: 916: 368: 198: 160: 156: 128: 98: 2771:
Proceedings of the Royal Society London Series B – Biological Sciences
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in the mantle of gastropods correlates with asymmetric shell coiling."
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The largest height of any shell is found in the marine snail species
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or have an irregular shape with a siphonal canal extension, as in the
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Asami, T. (1993) "Genetic variation of coiling chirality in snails".
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Cyrtoconoid: approaching a conical shape but with convex sides (see:
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The most frequently used measurements of a gastropod shell are: the
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The terminology used to describe the shells of gastropods includes:
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On the inside of the outer lip, various ridges or plications called
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Falkner G., Obrdlík P., Castella E. & Speight M. C. D. (2001).
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Caracoles Terrestres de Andalúcia. Guía y manual de identificación
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Among proposed roles invoked for the variability of shells during
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Pear-shaped: a combination of two shapes: ovate-conic and conic.
1805: 1763: 1746: 1713: 1699: 1691: 1394: 1358: 988: 911: 694: 619: 540: 429: 315: 132: 1499:: a ridge on the inner lip of the aperture in certain gastropods 208: 3389: 3374: 3348: 2161: 1780: 1404: 1291: 1155:(Müller), with the direction of coiling determined by a simple 1090: 723: 51:, on the right is the normally dextral (right-handed) shell of 3394: 3343: 3232: 2961: 2719:(Linnaeus, 1758) (Turbinellidae) the largest living gastropod 2320: 1680: 1340: 1290:
The shell begins with the larval shell, the (usually) minute
1077: 1052: 963: 959: 951: 937: 439: 189:. The calcareous central layer, tracum, is typically made of 180: 30:"Snail shell" redirects here. For the cave in Tennessee, see 2160:) a coiled shell from the previous condition of an uncoiled 1487:: the "little column" at the axis of revolution of the shell 242:
Central image: Lateral view showing the profile of the shell
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Wells F. E., Walker D. I. & Jones D. S. (eds.) (2003) "
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The gastropod shell has three major layers secreted by the
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WM. C. Brown Company Publishers, Dubuque, Iowa. 214 pages.
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Morphology of typical spirally coiled shell. The shell of
1267:: the shell is shown looking down directly onto the apex. 244:
Lower image: Basal view showing umbilicus in the centre.
1580:: each one of the complete rotations of the shell spiral 2865: 2197: 1047:
The chirality in gastropods appears in early cleavage (
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A possible exception may concern some of the European
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depressed – the width is much bigger than the height
1543:: an extension of the aperture in certain gastropods 1535:: raised lines or ridges on the surface of the shell 1379:), or by periodic renewal as a succession of holes ( 2715:Food of giants – field observations on the diet of 1162: 1015:(left-handed), and a very few species (for example 2808: 2806: 151:, which provides further protection. The study of 3101:"Effect of Ribonuclease on Shell Regeneration in 1803:Trochiform, pyramidal, conical with a flat base. 1593:oblong – the height is much bigger than the width 1108:), as well as several species of Pacific islands 240:Upper image: Dorsal view, showing whorls and apex 3589: 3099:Nair K. K. & Muthe P. T. (18 November 1961) 1529:: ornamentation on the outer surface of a shell 254:apical view, an apertural view and a basal view 163:. Shell morphology terms vary by species group. 2803: 2647:. München: Friedrich-Held-Gesellschaft, 267 pp. 2571: 2569: 2567: 2102:orientation of the shell on the living animal. 197:) precipitated into an organic matrix known as 2630:. A. G. Seiler & Company, New York. pages 2565: 2563: 2561: 2559: 2557: 2555: 2553: 2551: 2549: 2547: 2033:has the largest shell of any living gastropod. 2003:elongate-conic or turriform or cockscrew shape 3165: 3017: 1549:: the part of the shell above the body whorl. 914:, low rates give highly coiled forms such as 887: 2399: 2397: 1058: 2544: 3172: 3158: 3024: 3010: 2167: 1895:Detailed distinction of the shape can be: 1686:Contabulate, short, with shouldered whorls 894: 880: 64:The shell of a large land snail (probably 2834: 2832: 2394: 2152:that changed their developmental timing ( 1362:, etc.) it is drawn out into an anterior 34:. For the They Might Be Giants song, see 2610: 2608: 2452:"Amphidromus perversus (Linnaeus, 1758)" 2076: 1811:Turbinated: conical, with rounded base. 1434: 1275: 1023: 108: 59: 40: 27:Part of the body of a gastropod or snail 2637: 2196:into the water that raised the water's 2121: 1443:form of the shell of the garden snail, 14: 3590: 3095:Radiocarbon Dating of Gastropod Shells 2829: 1475:: the smallest few whorls of the shell 1229:also shows an apical view of its shell 1223:This dorsal view of the living animal 3179: 3153: 2707: 2650: 2605: 2524:"Left-right asymmetric expression of 1584: 1430: 2675:Smithsonian Contributions to Zoology 2661:Structural and systematic conchology 2645:Shelled Gastropoda of Western Europe 2137:. This reduction can be seen in all 237:, a land snail, has dextral coiling. 2473:Integrative and Comparative Biology 2222:, many-whorled shell of tower snail 2203: 1955:depressed trochiform or valvatiform 1827:Scalariform, whorls not impinging. 995:A few gastropods, for instance the 24: 2887: 2841:Proceedings of the Royal Society B 212: 25: 3614: 3521:Reproductive system of gastropods 3082: 2380:, Bahamas. Scale bar 500 μm. 2111:, which can be up to 91 cm. 962:and the inward-coiled lip of the 3526:Respiratory system of gastropods 3441:Circulatory system of gastropods 2907: 2575: 2365: 2349: 2333: 2313: 2296: 2277: 2261: 2244: 2227: 2210: 2065: 2053: 2038: 2020: 2008: 1996: 1984: 1972: 1960: 1948: 1936: 1924: 1912: 1900: 1639: 1622: 1605: 1216: 1201: 1186: 1169: 1163:Standard ways of viewing a shell 861: 860: 278: 222: 2787: 2763: 2731: 2694: 2666: 2376:lagoonal sediment of Rice Bay, 1744:Elongated, subulate, elevated. 1366:, of greater or lesser length. 625:microbial calcite precipitation 166: 3466:Excretory system of gastropods 3456:Digestive system of gastropods 2516: 2499: 2486: 2462: 2444: 2410: 2251:Shell of marine cowry snail – 2027:spindle shape – the sea snail 1271: 13: 1: 2385: 2085: 1862:Boat-shaped, slipper-shaped. 1837:Irregularly spiral, evolute. 1493:: the "trapdoor" of the shell 1193:Abapertural view of shell of 585:marine biogenic calcification 204: 3531:Sensory organs of gastropods 3496:Nervous system of gastropods 2623:North American Index Fossils 2505:Myers P. Z. (13 April 2009) 1632:Sphincterochila candidissima 1469:: the margin of the aperture 1002: 210: 7: 3562:Circumesophageal nerve ring 1208:Umbilical view of shell of 1176:Apertural view of shell of 814:Biomineralising polychaetes 580:amorphous calcium carbonate 266:Part of a series related to 10: 3619: 1870:Conical or limpet-shaped. 1761:Fusiform, spindle-shaped. 1459:: the opening of the shell 846:Burgess Shale preservation 229: 170: 29: 3539: 3433: 3357: 3321: 3188: 3077:10.1016/j.cub.2004.08.010 2439:10.1007/s00114-005-0045-2 1517:: folds on the columella. 1439:A drawing of an abnormal 1059:Mixed coiling populations 808:Cupriavidus metallidurans 127:is part of the body of a 113:Four views of a shell of 3089:Gastropods by J. H. Leal 2814:BMC Evolutionary Biology 2627:Invertebrates. Volume I. 1320:The most simple form of 1028:The left-handed turrid ( 529:Teeth, scales, tusks etc 2657:George Washington Tryon 2593:: a synoptic catalogue" 2468:Geerat Vermeij (2002). 2403:John. B. Burch (1962). 2178:Dinosaur Park Formation 1720:Depressed, lenticular. 1697:Cylindrical, pupiform. 1661:Bulloid: bubble-shaped 1555:: The junction between 590:calcareous nannofossils 386:Choanoflagellate lorica 2853:10.1098/rspb.2003.2517 2825:10.1186/1471-2148-5-59 2782:10.1098/rspb.2003.2624 2589:"The land snail genus 2539:10.1186/2041-9139-4-15 2513:, accessed 3 May 2009. 2323:image of the shell of 2082: 1678:Cone-shaped, obconic. 1449: 1287: 1280:Spiral tower shell of 1129:. They are obligatory 1033: 779:Magnetotactic bacteria 604:oolitic aragonite sand 462:scaly-foot snail shell 248:Photo of the shell of 217: 120: 106: 57: 2743:Melarhaphe neritoides 2080: 1615:Bulgarica denticulata 1438: 1279: 1027: 1018:Amphidromus perversus 931:, elongate as in the 216: 112: 63: 44: 2949:About mollusc shells 2507:"Snails have nodal!" 2481:10.1093/icb/42.5.935 2122:Evolutionary changes 1795:Short, bucciniform. 1789:Millerelix peregrina 1294:whorls known as the 1137:Nenia callistoglypta 131:or snail, a kind of 3486:Hypobranchial gland 3434:Internal soft parts 3358:External soft parts 3057:10.1038/hdy.1996.41 2895:Part of a series on 2847:(1533): 2551–2555. 2419:Naturwissenschaften 2378:San Salvador Island 2219:Turritella communis 2156:) and gained back ( 1648:Escargot de Quimper 1646:depressed shell of 1307:one another (as in 1226:Calliostoma bairdii 1085:(Linnaeus) (family 506:Vertebrate skeleton 296:Mineralized tissues 2795:Systematic Zoology 2758:10.1007/BF00014480 2724:2009-10-03 at the 2094:of the shell, the 2083: 1881:Fasciolaria tulipa 1723:Ethalia carneolata 1658:Regularly spiral: 1585:Shape of the shell 1559:of most gastropods 1450: 1431:Parts of the shell 1344:) and many of the 1288: 1098:and many Hawaiian 1034: 670:diatomaceous earth 636:Great Calcite Belt 553:Scale microfossils 546:otolithic membrane 457:small shelly fauna 430:echinoderm stereom 306:Biocrystallization 250:Zonitoides nitidus 234:Zonitoides nitidus 218: 135:. The shell is an 121: 116:Arianta arbustorum 107: 58: 36:John Henry (album) 3598:Gastropod anatomy 3585: 3584: 3511:Oesophageal pouch 3370:Caudal mucous pit 3071:(16): R654–R656. 3034: 3033: 2988:Crustacean shells 2983:Brachiopod shells 2739:Littorina striata 2663:, 1882, pp. 43–44 2597:Fieldiana Zoology 2511:The Panda's Thumb 2286:Epitonium scalare 2194:calcium carbonate 1830:Epitonium scalare 1629:globose shell of 1310:Epitonium scalare 1283:Epitonium scalare 1125:of the subfamily 1030:Antiplanes vinosa 904: 903: 834:permineralization 819:Mineral nutrients 744:Mineral evolution 413:foraminifera test 272:Biomineralization 258: 257: 191:calcium carbonate 54:Neptunea despecta 48:Neptunea angulata 16:(Redirected from 3610: 3501:Nidamental gland 3425:Suprapedal gland 3322:Other hard parts 3174: 3167: 3160: 3151: 3150: 3127: 3120:10.1038/192674b0 3026: 3019: 3012: 2998:Sea urchin tests 2911: 2892: 2891: 2882: 2878: 2863: 2836: 2827: 2810: 2801: 2791: 2785: 2767: 2761: 2735: 2729: 2711: 2705: 2698: 2692: 2690: 2670: 2664: 2654: 2648: 2641: 2635: 2612: 2603: 2579: 2573: 2542: 2520: 2514: 2503: 2497: 2490: 2484: 2475:42(5): 9359–40. 2466: 2460: 2459: 2448: 2442: 2414: 2408: 2401: 2369: 2353: 2337: 2317: 2300: 2281: 2265: 2254:Cypraea nebrites 2248: 2231: 2214: 2204:Variety of forms 2069: 2057: 2047:Venus Comb Murex 2042: 2024: 2012: 2000: 1988: 1976: 1964: 1952: 1940: 1928: 1916: 1904: 1643: 1626: 1612:oblong shell of 1609: 1245:abapertural view 1220: 1205: 1190: 1173: 1146:Alinda biplicata 1082:Liguus virgineus 896: 889: 882: 869: 864: 863: 784:Magnetoreception 764:Ballast minerals 359:Cephalopod shell 354:Brachiopod shell 301:Remineralisation 282: 263: 262: 226: 209: 173:mantle (mollusc) 79:columellar plait 32:Snail Shell Cave 21: 3618: 3617: 3613: 3612: 3611: 3609: 3608: 3607: 3588: 3587: 3586: 3581: 3535: 3476:Hancock's organ 3429: 3353: 3317: 3288:Stromboid notch 3184: 3178: 3125: 3085: 3065:Current Biology 3037:About chirality 3030: 2993:Horseshoe crabs 2975:Other seashells 2890: 2888:Further reading 2885: 2879: 2866: 2837: 2830: 2811: 2804: 2792: 2788: 2768: 2764: 2736: 2732: 2726:Wayback Machine 2712: 2708: 2699: 2695: 2686: 2671: 2667: 2655: 2651: 2642: 2638: 2613: 2606: 2574: 2545: 2521: 2517: 2504: 2500: 2491: 2487: 2467: 2463: 2450: 2449: 2445: 2425:(11): 504–515. 2415: 2411: 2402: 2395: 2388: 2381: 2370: 2361: 2354: 2345: 2338: 2329: 2318: 2309: 2303:X-ray image of 2301: 2290: 2282: 2273: 2266: 2257: 2249: 2240: 2234:X-ray image of 2232: 2223: 2215: 2206: 2173: 2124: 2088: 2073: 2072:irregular shape 2070: 2061: 2058: 2049: 2043: 2034: 2025: 2016: 2013: 2004: 2001: 1992: 1989: 1980: 1977: 1968: 1965: 1956: 1953: 1944: 1941: 1932: 1929: 1920: 1917: 1908: 1905: 1854:Shield-shaped. 1650: 1644: 1635: 1627: 1618: 1610: 1587: 1497:Parietal callus 1433: 1302:, the coils or 1274: 1230: 1221: 1212: 1210:Valvata sincera 1206: 1197: 1195:Valvata sincera 1191: 1182: 1179:Valvata sincera 1174: 1165: 1073:Liguus vittatus 1061: 1051:) and the gene 1049:spiral cleavage 1005: 920:or some of the 900: 859: 852: 851: 850: 738: 730: 729: 728: 684: 676: 675: 674: 660:biogenic silica 654: 644: 643: 642: 627: 615: 594: 574: 564: 563: 562: 530: 522: 521: 520: 500: 485: 484: 483: 450:gastropod shell 418:testate amoebae 408:diatom frustule 333: 322: 321: 320: 290: 260: 253: 243: 241: 238: 207: 196: 183: 171:Main articles: 169: 125:gastropod shell 101: 96: 91: 86: 81: 76: 71: 39: 28: 23: 22: 15: 12: 11: 5: 3616: 3606: 3605: 3603:Mollusc shells 3600: 3583: 3582: 3580: 3579: 3574: 3569: 3564: 3559: 3554: 3549: 3543: 3541: 3537: 3536: 3534: 3533: 3528: 3523: 3518: 3513: 3508: 3503: 3498: 3493: 3488: 3483: 3481:Hepatopancreas 3478: 3473: 3471:Gastric shield 3468: 3463: 3458: 3453: 3448: 3443: 3437: 3435: 3431: 3430: 3428: 3427: 3422: 3417: 3415:Semper's organ 3412: 3407: 3402: 3397: 3392: 3387: 3382: 3377: 3372: 3367: 3365:Caryophyllidia 3361: 3359: 3355: 3354: 3352: 3351: 3346: 3341: 3336: 3331: 3325: 3323: 3319: 3318: 3316: 3315: 3310: 3305: 3300: 3295: 3290: 3285: 3280: 3278:Siphonal notch 3275: 3273:Siphonal canal 3270: 3265: 3260: 3255: 3250: 3245: 3240: 3235: 3230: 3225: 3220: 3215: 3210: 3205: 3200: 3194: 3192: 3186: 3185: 3177: 3176: 3169: 3162: 3154: 3148: 3147: 3123: 3097: 3092: 3084: 3083:External links 3081: 3080: 3079: 3060: 3039: 3038: 3032: 3031: 3029: 3028: 3021: 3014: 3006: 3003: 3002: 3001: 3000: 2995: 2990: 2985: 2977: 2976: 2972: 2971: 2970: 2969: 2964: 2959: 2951: 2950: 2946: 2945: 2944: 2943: 2938: 2933: 2928: 2920: 2919: 2917:Mollusc shells 2913: 2912: 2904: 2903: 2897: 2896: 2889: 2886: 2884: 2883: 2864: 2828: 2802: 2786: 2762: 2730: 2717:Syrinx aruanus 2706: 2693: 2665: 2649: 2636: 2604: 2543: 2515: 2498: 2485: 2461: 2443: 2409: 2392: 2387: 2384: 2383: 2382: 2371: 2364: 2362: 2355: 2348: 2346: 2339: 2332: 2330: 2319: 2312: 2310: 2302: 2295: 2292: 2291: 2283: 2276: 2274: 2267: 2260: 2258: 2250: 2243: 2241: 2233: 2226: 2224: 2216: 2209: 2205: 2202: 2172: 2166: 2123: 2120: 2108:Syrinx aruanus 2087: 2084: 2075: 2074: 2071: 2064: 2062: 2059: 2052: 2050: 2044: 2037: 2035: 2030:Syrinx aruanus 2026: 2019: 2017: 2014: 2007: 2005: 2002: 1995: 1993: 1990: 1983: 1981: 1978: 1971: 1969: 1966: 1959: 1957: 1954: 1947: 1945: 1942: 1935: 1933: 1930: 1923: 1921: 1918: 1911: 1909: 1906: 1899: 1893: 1892: 1884: 1876: 1868: 1860: 1852: 1849: 1835: 1834: 1833: 1825: 1817: 1809: 1801: 1793: 1786:Many-whorled. 1784: 1776: 1767: 1759: 1750: 1742: 1734: 1726: 1718: 1709: 1695: 1687: 1684: 1676: 1667: 1652: 1651: 1645: 1638: 1636: 1628: 1621: 1619: 1611: 1604: 1601: 1600: 1597: 1594: 1586: 1583: 1582: 1581: 1575: 1569: 1563: 1560: 1550: 1544: 1541:Siphonal canal 1538: 1537: 1536: 1524: 1518: 1512: 1506: 1500: 1494: 1488: 1482: 1476: 1470: 1460: 1446:Cornu aspersum 1432: 1429: 1364:siphonal canal 1273: 1270: 1269: 1268: 1262: 1255:umbilical view 1252: 1242: 1239:apertural view 1232: 1231: 1222: 1215: 1213: 1207: 1200: 1198: 1192: 1185: 1183: 1175: 1168: 1164: 1161: 1106:Achatinellidae 1060: 1057: 1004: 1001: 968: 967: 956: 946: 942: 925: 902: 901: 899: 898: 891: 884: 876: 873: 872: 871: 870: 854: 853: 849: 848: 843: 842: 841: 836: 826: 821: 816: 811: 804: 799: 791: 786: 781: 776: 771: 766: 761: 760: 759: 757:immobilization 754: 752:mineralization 746: 740: 739: 736: 735: 732: 731: 727: 726: 721: 720: 719: 709: 708: 707: 702: 692: 686: 685: 682: 681: 678: 677: 673: 672: 667: 665:siliceous ooze 662: 656: 655: 652:Silicification 650: 649: 646: 645: 641: 640: 639: 638: 633: 628: 616: 614: 613: 612: 611: 606: 595: 593: 592: 587: 582: 576: 575: 570: 569: 566: 565: 561: 560: 555: 550: 549: 548: 538: 532: 531: 528: 527: 524: 523: 519: 518: 513: 508: 502: 501: 491: 490: 487: 486: 482: 481: 476: 474:Sponge spicule 471: 470: 469: 467:estuary shells 464: 459: 454: 453: 452: 447: 442: 432: 422: 421: 420: 415: 410: 405: 400: 390: 389: 388: 378: 377: 376: 371: 366: 356: 351: 350: 349: 344: 335: 334: 328: 327: 324: 323: 319: 318: 313: 308: 303: 298: 292: 291: 288: 287: 284: 283: 275: 274: 268: 267: 256: 255: 246: 228: 227: 219: 206: 203: 194: 168: 165: 153:mollusc shells 26: 9: 6: 4: 3: 2: 3615: 3604: 3601: 3599: 3596: 3595: 3593: 3578: 3575: 3573: 3572:Tyrian purple 3570: 3568: 3565: 3563: 3560: 3558: 3555: 3553: 3550: 3548: 3545: 3544: 3542: 3538: 3532: 3529: 3527: 3524: 3522: 3519: 3517: 3514: 3512: 3509: 3507: 3504: 3502: 3499: 3497: 3494: 3492: 3489: 3487: 3484: 3482: 3479: 3477: 3474: 3472: 3469: 3467: 3464: 3462: 3459: 3457: 3454: 3452: 3449: 3447: 3444: 3442: 3439: 3438: 3436: 3432: 3426: 3423: 3421: 3418: 3416: 3413: 3411: 3408: 3406: 3403: 3401: 3398: 3396: 3393: 3391: 3388: 3386: 3383: 3381: 3378: 3376: 3373: 3371: 3368: 3366: 3363: 3362: 3360: 3356: 3350: 3347: 3345: 3342: 3340: 3337: 3335: 3332: 3330: 3327: 3326: 3324: 3320: 3314: 3311: 3309: 3306: 3304: 3301: 3299: 3296: 3294: 3291: 3289: 3286: 3284: 3281: 3279: 3276: 3274: 3271: 3269: 3266: 3264: 3261: 3259: 3256: 3254: 3251: 3249: 3246: 3244: 3241: 3239: 3236: 3234: 3231: 3229: 3226: 3224: 3221: 3219: 3216: 3214: 3211: 3209: 3206: 3204: 3201: 3199: 3196: 3195: 3193: 3191: 3187: 3182: 3175: 3170: 3168: 3163: 3161: 3156: 3155: 3152: 3145: 3141: 3140:84-935194-2-1 3137: 3133: 3132: 3124: 3121: 3117: 3113: 3110: 3106: 3104: 3098: 3096: 3093: 3090: 3087: 3086: 3078: 3074: 3070: 3067: 3066: 3061: 3058: 3054: 3050: 3047: 3046: 3041: 3040: 3036: 3035: 3027: 3022: 3020: 3015: 3013: 3008: 3007: 3005: 3004: 2999: 2996: 2994: 2991: 2989: 2986: 2984: 2981: 2980: 2979: 2978: 2974: 2973: 2968: 2965: 2963: 2960: 2958: 2955: 2954: 2953: 2952: 2948: 2947: 2942: 2939: 2937: 2934: 2932: 2931:Chiton shells 2929: 2927: 2924: 2923: 2922: 2921: 2918: 2915: 2914: 2910: 2906: 2905: 2902: 2899: 2898: 2894: 2893: 2877: 2875: 2873: 2871: 2869: 2861: 2857: 2854: 2850: 2846: 2843: 2842: 2835: 2833: 2826: 2822: 2818: 2815: 2809: 2807: 2799: 2796: 2790: 2783: 2779: 2775: 2772: 2766: 2759: 2755: 2751: 2748: 2747:Hydrobiologia 2744: 2740: 2734: 2727: 2723: 2720: 2718: 2710: 2703: 2697: 2689: 2684: 2680: 2677: 2676: 2669: 2662: 2658: 2653: 2646: 2640: 2633: 2629: 2628: 2624: 2620: 2616: 2611: 2609: 2602:(4): 505–720. 2601: 2598: 2594: 2592: 2586: 2582: 2581:Laidlaw F. F. 2578: 2572: 2570: 2568: 2566: 2564: 2562: 2560: 2558: 2556: 2554: 2552: 2550: 2548: 2540: 2536: 2532: 2529: 2527: 2519: 2512: 2508: 2502: 2495: 2489: 2482: 2478: 2474: 2471: 2465: 2457: 2456:jaxshells.org 2453: 2447: 2440: 2436: 2432: 2428: 2424: 2421: 2420: 2413: 2406: 2400: 2398: 2393: 2391: 2379: 2375: 2368: 2363: 2360: 2359: 2352: 2347: 2344: 2343: 2336: 2331: 2328: 2327: 2322: 2316: 2311: 2308: 2307: 2299: 2294: 2293: 2288: 2287: 2280: 2275: 2272: 2271: 2270:Helix pomatia 2264: 2259: 2256: 2255: 2247: 2242: 2239: 2238: 2230: 2225: 2221: 2220: 2213: 2208: 2207: 2201: 2199: 2195: 2191: 2187: 2183: 2179: 2170: 2165: 2164:-like shell. 2163: 2159: 2155: 2151: 2150:Calyptraeidae 2147: 2142: 2140: 2136: 2131: 2129: 2119: 2117: 2112: 2110: 2109: 2103: 2099: 2097: 2093: 2079: 2068: 2063: 2056: 2051: 2048: 2045:club shape - 2041: 2036: 2032: 2031: 2023: 2018: 2011: 2006: 1999: 1994: 1987: 1982: 1975: 1970: 1963: 1958: 1951: 1946: 1939: 1934: 1927: 1922: 1915: 1910: 1903: 1898: 1897: 1896: 1891: 1890: 1885: 1883: 1882: 1877: 1875: 1874: 1869: 1867: 1866: 1861: 1859: 1858: 1853: 1850: 1848: 1847: 1842: 1841: 1836: 1832: 1831: 1826: 1824: 1823: 1818: 1816: 1815: 1810: 1808: 1807: 1802: 1800: 1799: 1794: 1791: 1790: 1785: 1783: 1782: 1777: 1774: 1773: 1768: 1766: 1765: 1760: 1757: 1756: 1755:Helix pomatia 1752:Few-whorled. 1751: 1749: 1748: 1743: 1741: 1740: 1735: 1733: 1732: 1727: 1725: 1724: 1719: 1716: 1715: 1710: 1708: 1707: 1702: 1701: 1696: 1694: 1693: 1688: 1685: 1683: 1682: 1677: 1674: 1673: 1668: 1666: 1665: 1660: 1659: 1657: 1656: 1655: 1649: 1642: 1637: 1634: 1633: 1625: 1620: 1617: 1616: 1608: 1603: 1602: 1598: 1595: 1592: 1591: 1590: 1579: 1576: 1573: 1570: 1567: 1564: 1561: 1558: 1554: 1551: 1548: 1545: 1542: 1539: 1534: 1531: 1530: 1528: 1525: 1522: 1519: 1516: 1513: 1510: 1507: 1504: 1501: 1498: 1495: 1492: 1489: 1486: 1483: 1480: 1477: 1474: 1471: 1468: 1464: 1461: 1458: 1455: 1454: 1453: 1448: 1447: 1442: 1437: 1428: 1426: 1420: 1418: 1414: 1409: 1407: 1406: 1401: 1397: 1396: 1391: 1386: 1384: 1383: 1378: 1377:Fissurellidae 1374: 1373: 1367: 1365: 1361: 1360: 1354: 1349: 1347: 1343: 1342: 1337: 1332: 1330: 1329: 1323: 1318: 1316: 1312: 1311: 1305: 1301: 1297: 1293: 1285: 1284: 1278: 1266: 1263: 1260: 1256: 1253: 1250: 1246: 1243: 1240: 1237: 1236: 1235: 1228: 1227: 1219: 1214: 1211: 1204: 1199: 1196: 1189: 1184: 1181: 1180: 1172: 1167: 1166: 1160: 1158: 1154: 1153: 1152:Radix peregra 1148: 1147: 1141: 1139: 1138: 1132: 1128: 1124: 1119: 1117: 1113: 1112: 1107: 1103: 1102: 1097: 1096: 1092: 1088: 1084: 1083: 1079: 1075: 1074: 1069: 1068: 1056: 1054: 1050: 1045: 1043: 1038: 1031: 1026: 1022: 1020: 1019: 1014: 1010: 1000: 998: 993: 991: 990: 985: 980: 975: 973: 965: 961: 957: 954: 953: 947: 943: 940: 939: 934: 930: 926: 923: 919: 918: 913: 909: 908: 907: 897: 892: 890: 885: 883: 878: 877: 875: 874: 868: 858: 857: 856: 855: 847: 844: 840: 837: 835: 832: 831: 830: 829:Fossilization 827: 825: 824:Microbial mat 822: 820: 817: 815: 812: 810: 809: 805: 803: 800: 798: 796: 792: 790: 787: 785: 782: 780: 777: 775: 772: 770: 769:Magnetofossil 767: 765: 762: 758: 755: 753: 750: 749: 747: 745: 742: 741: 734: 733: 725: 722: 718: 715: 714: 713: 710: 706: 703: 701: 698: 697: 696: 693: 691: 688: 687: 680: 679: 671: 668: 666: 663: 661: 658: 657: 653: 648: 647: 637: 634: 632: 629: 626: 623: 622: 621: 618: 617: 610: 609:aragonite sea 607: 605: 602: 601: 600: 597: 596: 591: 588: 586: 583: 581: 578: 577: 573: 572:Calcification 568: 567: 559: 556: 554: 551: 547: 544: 543: 542: 539: 537: 534: 533: 526: 525: 517: 514: 512: 509: 507: 504: 503: 498: 494: 493:Endoskeletons 489: 488: 480: 477: 475: 472: 468: 465: 463: 460: 458: 455: 451: 448: 446: 443: 441: 438: 437: 436: 435:mollusc shell 433: 431: 428: 427: 426: 423: 419: 416: 414: 411: 409: 406: 404: 401: 399: 396: 395: 394: 393:Protist shell 391: 387: 384: 383: 382: 379: 375: 372: 370: 367: 365: 364:cirrate shell 362: 361: 360: 357: 355: 352: 348: 345: 343: 340: 339: 337: 336: 331: 326: 325: 317: 314: 312: 309: 307: 304: 302: 299: 297: 294: 293: 286: 285: 281: 277: 276: 273: 270: 269: 265: 264: 261: 251: 247: 245: 236: 235: 230: 225: 220: 215: 211: 202: 200: 192: 188: 182: 178: 177:mollusc shell 174: 164: 162: 158: 154: 150: 146: 142: 138: 134: 130: 126: 118: 117: 111: 105: 100: 95: 90: 85: 80: 75: 69: 68: 67:Helix pomatia 62: 56: 55: 50: 49: 43: 37: 33: 19: 3238:Periostracum 3189: 3144:from website 3130: 3126:(in Spanish) 3111: 3108: 3102: 3068: 3063: 3048: 3043: 2936:Snail shells 2935: 2844: 2839: 2816: 2813: 2797: 2794: 2789: 2773: 2770: 2765: 2749: 2746: 2742: 2738: 2733: 2716: 2709: 2701: 2700:Dance P. S. 2696: 2678: 2673: 2668: 2660: 2652: 2644: 2639: 2626: 2622: 2619:Shimer H. W. 2615:Grabau A. W. 2599: 2596: 2590: 2533:2013, 4:15. 2530: 2525: 2518: 2510: 2501: 2496:, 8, 263–276 2493: 2488: 2472: 2464: 2455: 2446: 2422: 2417: 2412: 2389: 2358:Murex pecten 2356: 2340: 2324: 2304: 2289:spiral shell 2284: 2268: 2252: 2235: 2217: 2174: 2171:implications 2158:re-evolution 2154:heterochrony 2143: 2132: 2125: 2116:central axis 2115: 2113: 2106: 2104: 2100: 2095: 2091: 2089: 2028: 1894: 1887: 1879: 1871: 1863: 1855: 1844: 1838: 1828: 1820: 1812: 1804: 1796: 1787: 1779: 1770: 1762: 1753: 1745: 1737: 1736:Ear-shaped. 1729: 1721: 1712: 1704: 1698: 1690: 1679: 1670: 1662: 1653: 1630: 1613: 1588: 1503:Periostracum 1466: 1451: 1444: 1421: 1417:periostracum 1410: 1403: 1393: 1387: 1380: 1370: 1368: 1357: 1350: 1339: 1333: 1326: 1319: 1308: 1289: 1281: 1264: 1258: 1254: 1248: 1244: 1238: 1233: 1224: 1209: 1194: 1177: 1150: 1144: 1142: 1135: 1120: 1109: 1099: 1093: 1087:Orthalicidae 1080: 1076:(Swainson), 1071: 1070:, the Cuban 1065: 1062: 1046: 1039: 1035: 1029: 1016: 1012: 1008: 1006: 994: 987: 976: 969: 950: 936: 929:turban shell 915: 905: 839:petrifaction 806: 794: 789:Microfossils 536:Limpet teeth 516:Ossification 511:Bone mineral 449: 445:chiton shell 330:Exoskeletons 311:Biointerface 259: 249: 239: 232: 184: 167:Shell layers 155:is known as 124: 122: 114: 65: 52: 46: 3567:Trochophore 3552:Pseudofeces 3506:Odontophore 3461:Diverticula 3405:Pneumostome 3248:Planispiral 3134:. 303 pp., 3114:: 674–675. 3051:: 278–286. 2941:Tusk shells 2926:Clam shells 2776:: 529–536. 2752:: 129–142. 2591:Amphidromus 2326:Tonna galea 2190:microfossil 2184:from local 2146:Dollo's law 1979:ovate-conic 1943:planispiral 1728:Discoidal. 1672:Calliostoma 1372:Clavilithes 1272:Description 1265:apical view 1249:dorsal view 1159:recessive. 1131:calciphiles 1123:clausiliids 1101:Achatinella 1067:Amphidromus 922:Planorbidae 774:Magnetosome 717:phosphorite 683:Other forms 631:calcite sea 398:coccosphere 342:exoskeleton 137:exoskeleton 3592:Categories 3516:Osphradium 3491:Nephridium 3410:Rhinophore 3400:Parapodium 3329:Clausilium 3268:Selenizone 3258:Protoconch 3208:Body whorl 3103:Ariophanta 2957:Conchology 2800:: 204–223. 2688:10088/5530 2386:References 2237:Turritella 2186:coniferous 2169:Taphonomic 2086:Dimensions 1967:trochiform 1931:neritiform 1857:Umbraculum 1840:Siliquaria 1822:Turritella 1778:Globular. 1772:Streptaxis 1731:Elachorbis 1521:Protoconch 1479:Body whorl 1346:Ranellidae 1328:Platyceras 1296:protoconch 1259:basal view 1116:Partulidae 997:Vermetidae 979:intertidal 972:David Raup 933:cone shell 917:Turritella 369:cuttlebone 338:Arthropod 205:Morphology 199:conchiolin 161:malacology 157:conchology 99:body whorl 18:Teleoconch 3547:Epiphragm 3451:Ctenidium 3339:Operculum 3334:Love dart 3298:Umbilicus 3263:Sculpture 3243:Peristome 3218:Columella 3181:Gastropod 2901:Seashells 2135:evolution 2128:evolution 2060:egg shape 2015:top shape 1919:ear shape 1907:cap shape 1865:Crepidula 1566:Umbilicus 1527:Sculpture 1509:Peristome 1491:Operculum 1485:Columella 1467:peristome 1425:operculum 1400:umbilicus 1322:sculpture 1292:embryonic 1157:Mendelian 1127:Alopiinae 1095:Partulina 1013:sinistral 1003:Chirality 945:aperture. 795:engrailed 712:Phosphate 705:oil shale 599:Aragonite 403:coccolith 145:semi-slug 129:gastropod 89:columella 74:umbilicus 3380:Cnidosac 3198:Aperture 3045:Heredity 2860:14728776 2722:Archived 2681:: 1–55. 2632:page 582 2587:(1961). 2585:Solem A. 2431:16217668 2374:Holocene 2342:Charonia 1851:Tubular. 1846:Vermetus 1798:Buccinum 1739:Haliotis 1457:Aperture 1441:scalarid 1382:Haliotis 1353:aperture 1114:(family 1104:(family 1091:Hawaiian 1089:), some 1042:Cenozoic 867:Category 748:In soil 700:alginite 690:Bone bed 425:Seashell 332:(shells) 149:aperture 84:aperture 3577:Veliger 3557:Torsion 3183:anatomy 2621:(1909) 2531:EvoDevo 2306:Cypraea 2182:tannins 1873:Patella 1806:Trochus 1764:Fusinus 1747:Terebra 1714:Gibbula 1700:Lioplax 1692:Cypraea 1395:Nerinea 1359:Fusinus 1338:as in ( 1111:Partula 1078:Haitian 1009:dextral 989:Terebra 912:abalone 737:Related 695:Kerogen 620:Calcite 541:Otolith 374:gladius 347:cuticle 316:Biofilm 289:General 252:with an 133:mollusc 3420:Siphon 3390:Mentum 3385:Mantle 3375:Cerata 3349:Radula 3293:Suture 3213:Callus 3138:  3109:Nature 2858:  2819:: 59. 2702:Shells 2617:& 2583:& 2429:  2162:limpet 2092:height 1781:Natica 1557:whorls 1553:Suture 1413:Recent 1405:Natica 1315:suture 1304:whorls 865:  724:Pyrena 381:Lorica 187:mantle 179:, and 94:suture 3540:Other 3395:Notum 3344:Pearl 3313:Whorl 3308:Varix 3303:Valve 3283:Spire 3253:Plait 3233:Nacre 3190:Shell 2967:Valve 2962:Nacre 2634:–584. 2494:Forma 2321:X-ray 2139:slugs 2096:width 1991:conic 1889:Ficus 1814:Turbo 1681:Conus 1664:Bulla 1578:Whorl 1572:Varix 1547:Spire 1515:Plait 1411:Many 1390:lirae 1341:Murex 1336:varix 1300:spire 1053:NODAL 984:olive 964:cowry 960:conch 952:Murex 938:Murex 802:Druse 497:bones 440:nacre 193:(CaCO 181:nacre 141:slugs 3446:Crop 3228:Lira 3203:Apex 3136:ISBN 3105:sp." 2856:PMID 2427:PMID 2114:The 1706:Pupa 1533:Lira 1473:Apex 1351:The 1257:(or 1247:(or 986:and 797:gene 558:Tusk 479:Test 123:The 104:apex 102:7 – 97:6 – 92:5 – 87:4 – 82:3 – 77:2 – 72:1 – 3223:Lip 3142:. ( 3116:doi 3112:192 3073:doi 3053:doi 2849:doi 2845:270 2821:doi 2778:doi 2774:271 2754:doi 2750:309 2745:". 2683:hdl 2679:600 2535:doi 2526:dpp 2477:doi 2435:doi 1843:, 1463:Lip 1408:). 1331:). 3594:: 3107:. 3069:14 3049:76 2867:^ 2831:^ 2805:^ 2798:35 2659:, 2607:^ 2600:41 2595:. 2546:^ 2509:. 2454:. 2433:. 2423:92 2396:^ 2198:pH 1703:, 1465:: 175:, 3173:e 3166:t 3159:v 3146:) 3122:. 3118:: 3075:: 3059:. 3055:: 3025:e 3018:t 3011:v 2862:. 2851:: 2823:: 2817:5 2784:. 2780:: 2760:. 2756:: 2704:. 2691:. 2685:: 2541:. 2537:: 2483:. 2479:: 2458:. 2441:. 2437:: 1792:. 1775:. 1758:. 1717:) 1675:) 1356:( 1286:. 1032:) 966:. 955:. 941:. 924:. 895:e 888:t 881:v 499:) 495:( 221:| 195:3 38:. 20:)

Index

Teleoconch
Snail Shell Cave
John Henry (album)

Neptunea angulata
Neptunea despecta

Helix pomatia
umbilicus
columellar plait
aperture
columella
suture
body whorl
apex

Arianta arbustorum
gastropod
mollusc
exoskeleton
slugs
semi-slug
aperture
mollusc shells
conchology
malacology
mantle (mollusc)
mollusc shell
nacre
mantle

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