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Tapinocephalus Assemblage Zone

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Sidor, Christian A.; Angielczyk, Kenneth D.; Smith, Roger M. H.; Goulding, Adam K.; Nesbitt, Sterling J.; Peecook, Brandon R.; Steyer, J. Sébastien; Tolan, Stephen (2014-06-07). "Tapinocephalids (Therapsida, Dinocephalia) from the Permian Madumabisa Mudstone Formation (Lower Karoo, Mid-Zambezi Basin)
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Angielczyk, Kenneth D.; Rubidge, Bruce S.; Day, Michael O.; Lin, Florence (2016-02-06). "A reevaluation ofBrachyprosopus broomiandChelydontops altidentalis, dicynodonts (Therapsida, Anomodontia) from the middle PermianTapinocephalusAssemblage Zone of the Karoo Basin, South Africa".
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Rubidge, Bruce S.; Day, Michael O.; Barbolini, Natasha; Hancox, P. John; Choiniere, Jonah N.; Bamford, Marion K.; Viglietti, Pia A.; McPhee, Blair W.; Jirah, Sifelani (2016), "Advances in Nonmarine Karoo Biostratigraphy: Significance for Understanding Basin Development",
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Boos, A.D.S.; Kammerer, C.F.; Schultz, C.L.; Paes Neto, V.D. (2015-11-01). "A tapinocephalid dinocephalian (Synapsida, Therapsida) from the Rio do Rasto Formation (Paraná Basin, Brazil): Taxonomic, ontogenetic and biostratigraphic considerations".
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Day, Michael Oliver; Rubidge, Bruce Sidney (2014-12-01). "A brief lithostratigraphic review of the Abrahamskraal and Koonap formations of the Beaufort Group, South Africa: Towards a basin-wide stratigraphic scheme for the Middle Permian Karoo".
1942:
Kammerer, Christian F. (2013-09-21), "A Redescription of Eriphostoma microdon Broom, 1911 (Therapsida, Gorgonopsia) from the Tapinocephalus Assemblage Zone of South Africa and a Review of Middle Permian Gorgonopsians",
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Reisz, Robert R.; Dilkes, David W.; Berman, David S (1998-09-15). "Anatomy and relationships ofElliotsmithia longicepsBroom, a small synapsid (Eupelycosauria: Varanopseidae) from the late Permian of South Africa".
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ANGIELCZYK, KENNETH D.; RUBIDGE, BRUCE S. (September 2010). "A new pylaecephalid dicynodont (Therapsida, Anomodontia) from the Tapinocephalus Assemblage Zone, Karoo Basin, Middle Permian of South Africa".
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Kammerer, C.F., Smith, R.M., Day, M.O. and Rubidge, B.S., 2015. New information on the morphology and stratigraphic range of the mid‐Permian gorgonopsian E riphostoma microdon Broom, 1911.
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Canoville, A. and Chinsamy, A., 2017. Bone Microstructure of Pareiasaurs (Parareptilia) from the Karoo Basin, South Africa: Implications for Growth Strategies and Lifestyle Habits.
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Jirah, Sifelani; Rubidge, Bruce S. (2014-12-01). "Refined stratigraphy of the Middle Permian Abrahamskraal Formation (Beaufort Group) in the southern Karoo Basin".
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Rubidge, Bruce S.; Sidor, Christian A. (2002-07-08). "On the cranial morphology of the basal therapsids Burnetia and Proburnetia (Therapsida: Burnetiidae)".
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Boonstra, L.D. and Broom, R., 1936. Some features of the cranial morphology of the tapinocephalid deinocephalians. Bulletin of the AMNH; v. 72, article 2.
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Hancox, P.J; Rubidge, B.S (2001-01-01). "Breakthroughs in the biodiversity, biogeography, biostratigraphy, and basin analysis of the Beaufort group".
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Middle Permian continental biodiversity changes as reflected in the Beaufort Group of South Africa: a bio- and lithostratigraphic review of the
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Damiani, Ross J. (2004-01-01). "Temnospondyls from the Beaufort Group (Karoo Basin) of South Africa and Their Biostratigraphy".
1290:"Researches on the Structure, Organization, and Classification of the Fossil Reptilia. Part IX., Section 4. On the Gomphodontia" 820:
are not especially common, however are most commonly discovered as articulated single specimens within or associated with the
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defined the lower, middle, and upper sections of this biozone. In the 1970s, Keyser and Smith proposed the renaming of the
563:. Outcrops of this biozone are known from the south-western and central margins of the Abrahamskraal Formation where it 1992:
Cisneros, Juan Carlos; Abdala, Fernando; Jashashvili, Tea; de Oliveira Bueno, Ana; Dentzien-Dias, Paula (July 2015).
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Boonstra, Lieuwe D. (1938-03-01). "A Report on some Karroo Reptiles from the Luangwa Valley, Northern Rhodesia".
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Day, Michael O.; Güven, Saniye; Abdala, Fernando; Jirah, Sifelani; Rubidge, Bruce; Almond, John (April 2015).
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Keyser, A.W., 1979. A review of the biostratigraphy of the Beaufort Group in the Karoo Basin of South Africa.
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Broom, R., 1912, December. On some new fossil reptiles from the Permian and Triassic beds of South Africa. In
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Lyson, Tyler R.; Bever, Gabe S.; Scheyer, Torsten M.; Hsiang, Allison Y.; Gauthier, Jacques A. (2013-06-17).
1744:"Morphology and systematic position of the dinocephalian Styracocephalus platyrhynchus (Amniota: Therapsida)" 1721:
Atayman, S., Rubidge, B.S. and Abdala, F., 2009. Taxonomic re-evaluation of tapinocephalid dinocephalians.
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Assemblage zone, and all other succeeding assemblage zone deposits, were deposited over millions of years.
568: 482: 96: 239: 2002:, two bizarre anomodonts (Synapsida, Therapsida) with dental occlusion from the Permian of Gondwana" 1183:
van der Walt, Merrill; Day, Michael; Rubidge, Bruce; Cooper, Antony; Netterberg, Inge (2010-12-31).
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layers. As the name for this biozone suggests, it is renowned by paleontologists for its diverse
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Assemblage Zone have been found in formations in different countries which correlate in age. As
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in 1856. However, it was not until 1892 that it was observed that the geological strata of the
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Vertebrate biozonation of the Beaufort Group with special reference to the western Karoo Basin
1459: 1135:"A new burnetiamorph (Therapsida: Biarmosuchia) from the Lower Beaufort Group of South Africa" 574:
in its south-western localities. In its northern and eastern localities it inter-fingers with
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Boonstra, L.D., 1965. The girdles and limbs of the Gorgonopsia of the Tapinocephalus Zone.
1617: 579: 513: 501: 493: 1818:"Youngest dinocephalian fossils extend the Tapinocephalus Zone, Karoo Basin, South Africa" 8: 1947:, Vertebrate Paleobiology and Paleoanthropology, Springer Netherlands, pp. 171–184, 1514:. Geological Survey, Department of Mineral And Energy Affairs, Republic of South Africa. 1501:
Boonstra, L.D., 1969. The fauna of the Tapinocephalus Zone (Beaufort beds of the Karoo).
2259: 2144: 2083: 2034: 1993: 1798: 1759: 1730: 1440: 1358: 1311: 1270: 1243:"On the Discovery of the Fossil Remains of Bidental and other Reptiles in South Africa" 747: 743: 512:. After embarking on further study of dinocephalian fossils and their biostratigraphy, 356: 215: 2175: 1650: 1428: 640:, which indicates a lack of deposition for long periods of time. In some deposits the 2278: 2263: 2251: 2215: 2179: 2148: 2136: 2087: 2075: 2039: 2021: 1956: 1924: 1888: 1880: 1839: 1790: 1743: 1686: 1654: 1559: 1555: 1471: 1444: 1432: 1362: 1350: 1274: 1262: 1223: 1219: 1184: 1113: 695: 564: 1802: 1185:"A new GIS-based biozone map of the Beaufort Group (Karoo Supergroup), South Africa" 2243: 2207: 2171: 2128: 2067: 2029: 2013: 1948: 1920: 1916: 1870: 1829: 1782: 1755: 1678: 1646: 1551: 1424: 1342: 1301: 1258: 1254: 1215: 1146: 874: 870: 844: 824: 728: 613: 309: 295: 219: 2071: 1834: 1817: 1595: 2247: 2211: 2132: 1952: 1682: 888: 751: 732: 1089: 539:
which was accepted over Keyser and Smith’s proposal. However, the zoning of the
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are fine-grained and mainly tabular, indicating that deposition of these
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Broom, R., 1906. V.—On the Permian and Triassic Faunas of South Africa.
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species which have been discovered from this biozone are the advanced
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The braincase, basicranial axis and median septum in the Dinocephalia
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plains flowing northwards from a southerly source area in the rising
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pulses from the growing Gondwanides mountain chain and associated
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Assemblage Zone consist mainly of maroon to greyish red or purple
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where outcrops start to only be found in the south-east, north of
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first appeared in this biozone and remained ubiquitous until the
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fossils are frequently found disarticulated with much of their
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deposits. Alternating beds of light grey to dark greenish grey
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fossil species where almost all members of this family – the
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Assemblage Zone were first included with those of the lower
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Watson (1914), Keyser & Smith (1977-78), Kitching (1984)
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10.1671/0272-4634(2002)022[0257:otcmot]2.0.co;2
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Philosophical Transactions of the Royal Society of London B
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Origin and Evolution of the Cape Mountains and Karoo Basin
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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Biostratigraphy of the Beaufort Group (Karoo Supergroup).
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fossils, along with other therapsid species found in the
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rocks remains as they were defined by Keyser and Smith.
214:, a fossiliferous and geologically important geological 2056: 1856: 1677:, Springer International Publishing, pp. 141–149, 1331:"V.—On the Permian and Triassic Faunas of South Africa" 324:. It is also the first biozone of the series where the 624:
climate, revealing the presence of paleocalcretes and
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frequently contain both symmetrical and asymmetrical
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https://core.ac.uk/download/pdf/39675877.pdf#page=88
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Assemblage Zone is characterized by the presence of
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species along with the appearance of other advanced
258:. Its northernmost exposures occur around the towns 1815: 1076:are good biostratigraphic markers for this period. 1742:Rubidge, Bruce; VAN DEN HEEVER, J.A (1997-01-01). 601:layers which exhibit blocky weathering at exposed 2117: 1905: 1016:, invertebrate trackways and burrows, vertebrate 2298: 754:that had previously begun to take course due to 1525:Geocongress, Geological Society of South Africa 1394:Proceedings of the Zoological Society of London 620:, both are indicative of a warm and seasonally 1084:Assemblage Zone, have been recovered from the 678:traveled downstream in a northerly direction. 304:. It is characterised by the presence of this 1636: 935:are confined to this biozone. One of the few 774:in the Karoo Basin where the deposits of the 1772: 1622:: CS1 maint: multiple names: authors list ( 1205: 1024:, and a variety of plant fossils, namely of 382:, which he named (from oldest to youngest): 1945:Early Evolutionary History of the Synapsida 1417:Quarterly Journal of the Geological Society 1247:Quarterly Journal of the Geological Society 954:are likewise found, including a variety of 1540: 903:Despite the rarity of fossils, the fossil 42: 2033: 1874: 1859:"Evolutionary Origin of the Turtle Shell" 1833: 1580:South African Committee of Stratigraphy. 1305: 1169:South African Committee of Stratigraphy. 1150: 939:species found in South African deposits, 2200:Journal of South American Earth Sciences 1941: 1457: 1414: 785: 758:of the Palaeo-pacific plate beneath the 702:are capped in the upper sections of the 666:which weather to light orange grey. The 2161: 911:Assemblage Zone are extremely diverse. 363:could be differentiated based on their 351:rocks that encompass the current eight 2299: 1593: 1510:Keyser, A.W. and Smith, R.M.H., 1978. 1287: 1133:Rubidge, B.S.; Kitching, J.W. (2003). 662:and greenish grey to light olive grey 488:under the name "Pareiasaurus beds" by 370:. The initial undertaking was done by 2276: 1460:"Early dichotomies in the Therapsids" 1328: 400:Zone of "highly specialized group of 1240: 719:The depositional environment of the 555:Assemblage Zone correlates with the 238:in its southernmost exposures, from 1594:Oliver, Day, Michael (2014-03-04). 1584:. Pretoria, Council for Geoscience. 1173:. Pretoria, Council for Geoscience. 1126: 1064:fossils are only known from Middle 13: 2236:Journal of Vertebrate Paleontology 2121:Journal of Vertebrate Paleontology 2102:Annals of the South African Museum 2060:Journal of Vertebrate Paleontology 1909:Journal of Vertebrate Paleontology 1775:Journal of Vertebrate Paleontology 1760:10.1111/j.1502-3931.1997.tb00457.x 1241:Bain, Andrew Geddes (1845-02-01). 682:cracks which are infilled by fine 109:up to 6,561.68 feet (2,000 m) 14: 2328: 1639:Journal of African Earth Sciences 1544:Journal of African Earth Sciences 1429:10.1144/GSL.JGS.1938.094.01-04.14 1208:Journal of African Earth Sciences 412:named were subdivided further by 1822:South African Journal of Science 1556:10.1016/j.jafrearsci.2014.07.001 1464:South African Journal of Science 1220:10.1016/j.jafrearsci.2014.06.014 770:created accommodation space for 727:material being deposited in the 500:Assemblage Zone. As a young man 496:redefined the boundaries of the 254:, reaching up to areas south of 141: 2277:Sidor, Christian (2015-04-09). 2270: 2226: 2190: 2155: 2111: 2094: 2050: 1985: 1968: 1935: 1899: 1850: 1809: 1766: 1735: 1715: 1698: 1665: 1630: 1587: 1570: 1534: 1517: 1504: 1495: 1482: 1451: 1408: 1399: 781: 636:are also commonly found in the 206:which correlates to the middle 2307:South African assemblage zones 1921:10.1080/02724634.1998.10011087 1386: 1369: 1322: 1281: 1259:10.1144/GSL.JGS.1845.001.01.72 1234: 1199: 1176: 1159: 1054:species that are found in the 1045: 907:that have been found from the 723:Assemblage Zone was formed by 651:which have been attributed to 628:precipitation respectively in 268:Tapinocephalus Assemblage Zone 1: 2176:10.1016/S1342-937X(05)70315-4 2072:10.1080/02724634.2016.1078342 1651:10.1016/S0899-5362(01)00081-1 1458:Boonstra, L.D. (1963-05-01). 1119: 889:Styracocephalus platyrhynchus 2248:10.1080/02724634.2013.826669 2212:10.1016/j.jsames.2015.09.003 2133:10.1080/02724634.2010.501447 1953:10.1007/978-94-007-6841-3_11 1683:10.1007/978-3-319-40859-0_14 1576:Rubidge, B. S. (ed.) 1995b. 1165:Rubidge, B. S. (ed.) 1995b. 926:and the perplexing putative 865:respectively. Other notable 546: 7: 1835:10.17159/sajs.2015/20140309 1107: 586:is considered to be Middle 559:, Adelaide Subgroup of the 424:(from oldest to youngest): 210:, Adelaide Subgroup of the 10: 2333: 2006:Royal Society Open Science 1329:Broom, R. (January 1906). 1012:, invertebrate fossils of 862:Tapinocephalus atherstonei 803:Tapinocephalus atherstonei 338: 288:Tapinocephalus atherstonei 175:Tapinocephalus atherstonei 49:Tapinocephalus atherstonei 1876:10.1016/j.cub.2013.05.003 1582:Biostratigraphic Series 1 1347:10.1017/S001675680012271X 1171:Biostratigraphic Series 1 980:Permian-Triassic boundary 524:Assemblage Zone. In 1984 477:The rocks of the current 181: 169: 164: 154: 136: 118: 113: 105: 92: 79: 65: 55: 41: 26: 18: 1005:Rhinesuchoides tenuiceps 408:These proposed biozones 2283:Palaeontologia Africana 2000:Anomocephalus africanus 1996:Tiarajudens eccentricus 1976:Papers in Palaeontology 1723:Palaeontologia africana 1189:Palaeontologia Africana 1152:10.1111/1475-4983.00294 1042:, have been recovered. 1000:temnospondyl amphibians 942:Elliotsmithia longiceps 557:Abrahamskraal Formation 420:proposed the following 416:between 1906 and 1909. 208:Abrahamskraal Formation 70:Abrahamskraal Formation 1488:Boonstra, L.D., 1968. 1307:10.1098/rstb.1895.0001 1288:Seeley, H. G. (1895). 1086:Rio do Rasto Formation 1008:, remains of the fish 998:. Finally, fossils of 986:species found include 932:Eunotosaurus africanus 850:Anteosaurus magnificus 816:fossils found in this 793: 738:) by vast, low-energy 590:(Guadalupian) in age. 335:species first appear. 2234:of southern Zambia". 1706:The Anatomical Record 995:Pristerodon brachyops 951:Hipposaurus boonstrai 789: 528:proposed to name the 2312:Permian South Africa 1010:Namaichthys digitata 915:species such as the 744:Gondwanide mountains 694:was in a low-energy 686:are also found. The 674:which indicate that 644:layers contain thin 494:Lieuwe Dirk Boonstra 2018:10.1098/rsos.150090 1377:Geological Magazine 1335:Geological Magazine 1098:Madumabisa Mudstone 750:were the result of 374:who subdivided the 372:Harry Govier Seeley 355:were discovered by 347:to be found in the 202:assemblage zone or 28:Stratigraphic range 1712:(6), pp.1039-1066. 989:Robertia broomiana 975:Diictodon feliceps 886:, and the unusual 856:Jonkeria boonstrai 794: 504:collaborated with 357:Andrew Geddes Bain 331:Diictodon feliceps 2164:Gondwana Research 2108:(13), pp.237-249. 1869:(12): 1113–1119. 1729:, p.88Á90. Link: 1114:List of synapsids 809:Bradysaurus baini 791:Bradysaurus baini 698:environment. The 593:The rocks of the 321:Bradysaurus baini 189: 188: 2324: 2291: 2290: 2274: 2268: 2267: 2230: 2224: 2223: 2194: 2188: 2187: 2159: 2153: 2152: 2127:(5): 1396–1409. 2115: 2109: 2098: 2092: 2091: 2054: 2048: 2047: 2037: 1989: 1983: 1982:(2), pp.201-221. 1972: 1966: 1965: 1939: 1933: 1932: 1903: 1897: 1896: 1878: 1854: 1848: 1847: 1837: 1813: 1807: 1806: 1770: 1764: 1763: 1739: 1733: 1719: 1713: 1702: 1696: 1695: 1669: 1663: 1662: 1645:(3–4): 563–577. 1634: 1628: 1627: 1621: 1613: 1609:assemblage zones 1591: 1585: 1574: 1568: 1567: 1538: 1532: 1521: 1515: 1508: 1502: 1499: 1493: 1486: 1480: 1479: 1455: 1449: 1448: 1423:(1–4): 371–384. 1412: 1406: 1403: 1397: 1390: 1384: 1373: 1367: 1366: 1326: 1320: 1319: 1309: 1285: 1279: 1278: 1238: 1232: 1231: 1203: 1197: 1196: 1180: 1174: 1163: 1157: 1156: 1154: 1130: 892:. Unfortunately 875:Struthiocephalus 845:Tapinocephalidae 315:, and the large 281:The name of the 220:Karoo Supergroup 147: 145: 144: 46: 16: 15: 2332: 2331: 2327: 2326: 2325: 2323: 2322: 2321: 2297: 2296: 2295: 2294: 2275: 2271: 2231: 2227: 2195: 2191: 2160: 2156: 2116: 2112: 2099: 2095: 2066:(2): e1078342. 2055: 2051: 1990: 1986: 1973: 1969: 1963: 1940: 1936: 1904: 1900: 1863:Current Biology 1855: 1851: 1814: 1810: 1771: 1767: 1740: 1736: 1720: 1716: 1703: 1699: 1693: 1670: 1666: 1635: 1631: 1615: 1614: 1607:Pristerognathus 1592: 1588: 1575: 1571: 1539: 1535: 1522: 1518: 1509: 1505: 1500: 1496: 1487: 1483: 1456: 1452: 1413: 1409: 1404: 1400: 1391: 1387: 1374: 1370: 1327: 1323: 1286: 1282: 1239: 1235: 1204: 1200: 1181: 1177: 1164: 1160: 1131: 1127: 1122: 1110: 1057:Tapinochephalus 1048: 871:tapinocephalids 784: 672:ripple surfaces 572:Assemblage Zone 549: 508:on research of 486:Assemblage Zone 341: 226:. The thickest 196:Assemblage Zone 142: 140: 100:Assemblage Zone 87:Assemblage Zone 85:Pristerognathus 51: 22:Assemblage Zone 12: 11: 5: 2330: 2320: 2319: 2314: 2309: 2293: 2292: 2269: 2242:(4): 980–986. 2225: 2189: 2170:(1): 165–173. 2154: 2110: 2093: 2049: 1984: 1967: 1961: 1934: 1915:(3): 602–611. 1898: 1849: 1808: 1781:(2): 257–267. 1765: 1734: 1714: 1697: 1691: 1664: 1629: 1603:Tapinocephalus 1586: 1569: 1533: 1516: 1503: 1494: 1481: 1450: 1407: 1398: 1385: 1383:(1), pp.29-30. 1368: 1321: 1280: 1253:(1): 317–318. 1233: 1198: 1175: 1158: 1145:(1): 199–210. 1124: 1123: 1121: 1118: 1117: 1116: 1109: 1106: 1082:Tapinocephalus 1074:dinocephalians 1047: 1044: 967:therocephalian 909:Tapinocephalus 841:Titanosuchidae 798:Tapinocephalus 783: 780: 776:Tapinocephalus 752:crustal uplift 736:foreland basin 721:Tapinocephalus 595:Tapinocephalus 561:Beaufort Group 553:Tapinocephalus 548: 545: 535:Tapinocephalus 526:James Kitching 510:dinocephalians 498:Tapinocephalus 492:. Years later 479:Tapinocephalus 475: 474: 466: 458: 450: 442: 434: 406: 405: 398: 391: 388:Pareiasaurians 376:Beaufort Group 361:Beaufort Group 349:Beaufort Group 340: 337: 313:dinocephalians 310:tapinocephalid 296:tapinocephalid 276:Beaufort Group 270:is the second 212:Beaufort Group 194:Tapinocephalus 187: 186: 183: 179: 178: 171: 167: 166: 162: 161: 156: 152: 151: 138: 134: 133: 120: 116: 115: 111: 110: 107: 103: 102: 94: 90: 89: 81: 77: 76: 74:Beaufort Group 67: 63: 62: 57: 53: 52: 47: 39: 38: 32:Middle Permian 24: 23: 20:Tapinocephalus 9: 6: 4: 3: 2: 2329: 2318: 2315: 2313: 2310: 2308: 2305: 2304: 2302: 2288: 2284: 2280: 2273: 2265: 2261: 2257: 2253: 2249: 2245: 2241: 2237: 2229: 2221: 2217: 2213: 2209: 2205: 2201: 2193: 2185: 2181: 2177: 2173: 2169: 2165: 2158: 2150: 2146: 2142: 2138: 2134: 2130: 2126: 2122: 2114: 2107: 2103: 2097: 2089: 2085: 2081: 2077: 2073: 2069: 2065: 2061: 2053: 2045: 2041: 2036: 2031: 2027: 2023: 2019: 2015: 2012:(7): 150090. 2011: 2007: 2003: 2001: 1997: 1988: 1981: 1977: 1971: 1964: 1962:9789400768406 1958: 1954: 1950: 1946: 1938: 1930: 1926: 1922: 1918: 1914: 1910: 1902: 1894: 1890: 1886: 1882: 1877: 1872: 1868: 1864: 1860: 1853: 1845: 1841: 1836: 1831: 1827: 1823: 1819: 1812: 1804: 1800: 1796: 1792: 1788: 1784: 1780: 1776: 1769: 1761: 1757: 1753: 1749: 1745: 1738: 1732: 1728: 1724: 1718: 1711: 1707: 1701: 1694: 1692:9783319408583 1688: 1684: 1680: 1676: 1668: 1660: 1656: 1652: 1648: 1644: 1640: 1633: 1625: 1619: 1611: 1610: 1606: 1602: 1598: 1590: 1583: 1579: 1573: 1565: 1561: 1557: 1553: 1549: 1545: 1537: 1530: 1526: 1520: 1513: 1507: 1498: 1491: 1485: 1477: 1473: 1469: 1465: 1461: 1454: 1446: 1442: 1438: 1434: 1430: 1426: 1422: 1418: 1411: 1402: 1395: 1389: 1382: 1378: 1372: 1364: 1360: 1356: 1352: 1348: 1344: 1340: 1336: 1332: 1325: 1317: 1313: 1308: 1303: 1299: 1295: 1291: 1284: 1276: 1272: 1268: 1264: 1260: 1256: 1252: 1248: 1244: 1237: 1229: 1225: 1221: 1217: 1213: 1209: 1202: 1194: 1190: 1186: 1179: 1172: 1168: 1162: 1153: 1148: 1144: 1140: 1139:Palaeontology 1136: 1129: 1125: 1115: 1112: 1111: 1105: 1103: 1099: 1096:and from the 1095: 1091: 1087: 1083: 1079: 1078:Dinocephalian 1075: 1071: 1067: 1063: 1062:dinocephalian 1059: 1058: 1053: 1052:dinocephalian 1043: 1041: 1040: 1035: 1034: 1029: 1028: 1023: 1019: 1015: 1011: 1007: 1006: 1001: 997: 996: 991: 990: 985: 981: 977: 976: 972: 969:species. The 968: 964: 960: 957: 953: 952: 948: 947:biarmosuchian 944: 943: 938: 934: 933: 929: 925: 923: 918: 914: 910: 906: 901: 899: 895: 894:dinocephalian 891: 890: 885: 883: 878: 876: 872: 868: 867:dinocephalian 864: 863: 858: 857: 852: 851: 846: 842: 838: 837:Anteosauridae 834: 833:Dinocephalian 830: 826: 823: 819: 815: 811: 810: 805: 804: 799: 792: 788: 779: 777: 773: 772:sedimentation 769: 765: 761: 757: 753: 749: 745: 741: 737: 734: 730: 726: 722: 717: 715: 714:conglomerates 712: 709: 705: 701: 697: 693: 689: 685: 681: 677: 676:paleocurrents 673: 669: 665: 661: 657: 654: 650: 647: 643: 639: 635: 631: 627: 623: 619: 615: 612: 608: 604: 600: 596: 591: 589: 585: 581: 577: 573: 571: 567:overlies the 566: 562: 558: 554: 544: 542: 538: 536: 531: 527: 523: 522:Dinocephalian 519: 515: 511: 507: 503: 499: 495: 491: 487: 485: 480: 472: 471: 467: 464: 463: 459: 456: 455: 451: 448: 447: 446:Kistecephalus 443: 440: 439: 435: 432: 431: 427: 426: 425: 423: 419: 415: 411: 403: 399: 396: 392: 389: 385: 384: 383: 381: 377: 373: 369: 366: 362: 358: 354: 350: 346: 336: 334: 332: 327: 323: 322: 318: 314: 311: 307: 306:dinocephalian 303: 300: 299:dinocephalian 297: 294: 290: 289: 284: 279: 277: 273: 269: 265: 264:Victoria West 261: 257: 256:Graaff-Reinet 253: 249: 245: 244:Beaufort West 241: 237: 233: 229: 225: 221: 217: 213: 209: 205: 201: 197: 195: 184: 180: 177: 176: 172: 168: 163: 160: 157: 153: 150: 139: 135: 132: 128: 124: 121: 117: 112: 108: 104: 101: 99: 95: 91: 88: 86: 82: 78: 75: 71: 68: 64: 61: 58: 54: 50: 45: 40: 37: 33: 29: 25: 21: 17: 2286: 2282: 2272: 2239: 2235: 2228: 2203: 2199: 2192: 2167: 2163: 2157: 2124: 2120: 2113: 2105: 2101: 2096: 2063: 2059: 2052: 2009: 2005: 1999: 1995: 1987: 1979: 1975: 1970: 1944: 1937: 1912: 1908: 1901: 1866: 1862: 1852: 1828:(3–4): 1–5. 1825: 1821: 1811: 1778: 1774: 1768: 1751: 1747: 1737: 1726: 1722: 1717: 1709: 1705: 1700: 1674: 1667: 1642: 1638: 1632: 1608: 1604: 1600: 1596: 1589: 1581: 1577: 1572: 1547: 1543: 1536: 1528: 1524: 1519: 1511: 1506: 1497: 1489: 1484: 1467: 1463: 1453: 1420: 1416: 1410: 1401: 1393: 1388: 1380: 1376: 1371: 1341:(1): 29–30. 1338: 1334: 1324: 1297: 1293: 1283: 1250: 1246: 1236: 1211: 1207: 1201: 1192: 1188: 1178: 1170: 1166: 1161: 1142: 1138: 1128: 1090:Paraná Basin 1081: 1072:) deposits, 1055: 1049: 1037: 1031: 1027:Glossopteris 1025: 1009: 1003: 993: 987: 973: 949: 940: 930: 928:pantestudine 920: 908: 902: 887: 880: 873: 860: 854: 848: 807: 801: 797: 795: 790: 782:Paleontology 775: 720: 718: 594: 592: 569: 552: 550: 533: 497: 483: 478: 476: 468: 460: 454:Lystrosaurus 452: 444: 436: 430:Pareiasaurus 428: 414:Robert Broom 407: 342: 329: 319: 286: 280: 267: 224:South Africa 193: 192: 190: 173: 165:Type section 149:South Africa 131:Eastern Cape 97: 84: 48: 36:Late Permian 19: 2317:Guadalupian 2206:: 375–384. 1754:: 157–168. 1618:cite thesis 1599:Eodicynodon 1550:: 227–242. 1531:, pp.13-31. 1214:: 121–135. 1070:Guadalupian 1046:Correlation 1039:Schizoneura 959:gorgonopsid 922:Bradysaurus 913:Parareptile 748:Gondwanides 729:Karoo Basin 725:sedimentary 680:Desiccation 630:playa lakes 570:Eodicynodon 565:conformably 484:Eodicynodon 470:Cynognathus 462:Procolophon 438:Endothiodon 402:theriodonts 395:Dicynodonts 378:into three 293:herbivorous 242:through to 159:Karoo Basin 98:Eodicynodon 2301:Categories 1120:References 1022:therapsids 1018:footprints 984:dicynodont 971:dicynodont 945:, and the 937:pelycosaur 917:pareiasaur 898:postcrania 822:calcareous 814:Vertebrate 768:subduction 756:subduction 700:sandstones 692:sandstones 688:sandstones 668:siltstones 664:sandstones 653:silicified 618:limestones 616:and sheet 611:calcareous 576:Ecca Group 343:The first 326:dicynodont 317:pareiasaur 291:, a large 285:refers to 260:Fraserburg 252:Willowmore 240:Sutherland 236:Leeu-Gamka 232:Merweville 2264:128431441 2256:0272-4634 2220:0895-9811 2184:1342-937X 2149:129846697 2141:0272-4634 2088:130520407 2080:0272-4634 2026:2054-5703 1929:0272-4634 1885:0960-9822 1844:0038-2353 1795:0272-4634 1659:1464-343X 1612:(Thesis). 1564:1464-343X 1476:0038-2353 1445:129931146 1437:0370-291X 1363:129265956 1355:1469-5081 1275:128602890 1267:0370-291X 1228:1464-343X 1088:from the 1033:Dadoxylon 963:anomodont 900:missing. 760:Gondwanan 733:retro-arc 684:sandstone 660:siltstone 638:mudstones 634:Paleosols 626:carbonate 607:mudstones 547:Lithology 393:Zone of " 386:Zone of " 302:therapsid 248:Oudshoorn 170:Named for 106:Thickness 80:Underlies 2044:26587266 1893:23727095 1803:86207308 1300:: 1–57. 1108:See also 1014:molluscs 1002:such as 982:. Other 884:capensis 882:Moschops 829:mudstone 764:Orogenic 740:alluvial 708:mudstone 642:mudstone 609:contain 603:outcrops 599:mudstone 580:deposits 514:Boonstra 502:Boonstra 422:biozones 380:biozones 353:biozones 228:outcrops 200:tetrapod 182:Named by 123:Northern 114:Location 93:Overlies 2035:4632579 1748:Lethaia 1066:Permian 877:whaitsi 827:in the 825:nodules 818:biozone 762:Plate. 704:biozone 696:fluvial 614:nodules 588:Permian 584:biozone 582:. This 541:biozone 530:biozone 518:biozone 345:fossils 339:History 283:biozone 274:of the 272:biozone 218:of the 204:biozone 137:Country 127:Western 72:of the 66:Unit of 60:Biozone 2289:: 1–7. 2262:  2254:  2218:  2182:  2147:  2139:  2086:  2078:  2042:  2032:  2024:  1959:  1927:  1891:  1883:  1842:  1801:  1793:  1689:  1657:  1562:  1474:  1443:  1435:  1361:  1353:  1314:  1273:  1265:  1226:  1195:: 1–6. 1102:Zambia 1094:Brazil 1036:, and 965:, and 859:, and 843:, and 746:. The 649:lenses 605:. The 578:-aged 532:after 410:Seeley 365:fossil 266:. The 155:Extent 146:  129:& 119:Region 2260:S2CID 2145:S2CID 2084:S2CID 1799:S2CID 1470:(5). 1441:S2CID 1359:S2CID 1316:91793 1312:JSTOR 1271:S2CID 1050:Many 956:basal 711:clast 706:with 646:chert 506:Broom 490:Broom 418:Broom 216:Group 198:is a 2252:ISSN 2216:ISSN 2180:ISSN 2137:ISSN 2076:ISSN 2040:PMID 2022:ISSN 1998:and 1957:ISBN 1925:ISSN 1889:PMID 1881:ISSN 1840:ISSN 1791:ISSN 1687:ISBN 1655:ISSN 1624:link 1605:and 1560:ISSN 1472:ISSN 1433:ISSN 1351:ISSN 1263:ISSN 1224:ISSN 992:and 905:taxa 879:and 806:and 796:The 656:tuff 622:arid 551:The 473:beds 465:beds 457:beds 449:beds 441:beds 433:beds 368:taxa 262:and 250:and 234:and 191:The 56:Type 2244:doi 2208:doi 2172:doi 2129:doi 2068:doi 2030:PMC 2014:doi 1949:doi 1917:doi 1871:doi 1830:doi 1826:111 1783:doi 1756:doi 1710:300 1679:doi 1647:doi 1552:doi 1548:100 1425:doi 1343:doi 1302:doi 1298:186 1255:doi 1216:doi 1212:100 1147:doi 1100:in 1092:in 1020:of 731:(a 520:to 222:in 34:to 2303:: 2287:49 2285:. 2281:. 2258:. 2250:. 2240:34 2238:. 2214:. 2204:63 2202:. 2178:. 2166:. 2143:. 2135:. 2125:30 2123:. 2106:48 2104:, 2082:. 2074:. 2064:36 2062:. 2038:. 2028:. 2020:. 2008:. 2004:. 1978:, 1955:, 1923:. 1913:18 1911:. 1887:. 1879:. 1867:23 1865:. 1861:. 1838:. 1824:. 1820:. 1797:. 1789:. 1779:22 1777:. 1752:30 1750:. 1746:. 1727:44 1725:, 1708:, 1685:, 1653:. 1643:33 1641:. 1620:}} 1616:{{ 1601:, 1558:. 1546:. 1527:, 1468:59 1466:. 1462:. 1439:. 1431:. 1421:94 1419:. 1379:, 1357:. 1349:. 1337:. 1333:. 1310:. 1296:. 1292:. 1269:. 1261:. 1249:. 1245:. 1222:. 1210:. 1193:45 1191:. 1187:. 1143:46 1141:. 1137:. 1104:. 1030:, 961:, 919:, 853:, 839:, 812:. 716:. 632:. 328:, 278:. 125:, 30:: 2266:. 2246:: 2222:. 2210:: 2186:. 2174:: 2168:7 2151:. 2131:: 2090:. 2070:: 2046:. 2016:: 2010:2 1994:" 1980:1 1951:: 1931:. 1919:: 1895:. 1873:: 1846:. 1832:: 1805:. 1785:: 1762:. 1758:: 1681:: 1661:. 1649:: 1626:) 1566:. 1554:: 1529:2 1492:. 1478:. 1447:. 1427:: 1381:3 1365:. 1345:: 1339:3 1318:. 1304:: 1277:. 1257:: 1251:1 1230:. 1218:: 1155:. 1149:: 1068:( 924:, 537:, 404:" 397:" 390:" 333:,

Index

Stratigraphic range
Middle Permian
Late Permian

Biozone
Abrahamskraal Formation
Beaufort Group
Pristerognathus Assemblage Zone
Eodicynodon Assemblage Zone
Northern
Western
Eastern Cape
South Africa
Karoo Basin
Tapinocephalus atherstonei
tetrapod
biozone
Abrahamskraal Formation
Beaufort Group
Group
Karoo Supergroup
South Africa
outcrops
Merweville
Leeu-Gamka
Sutherland
Beaufort West
Oudshoorn
Willowmore
Graaff-Reinet

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