173:
858:
338:
960:
810:
693:
1079:
1009:
971:
847:
1144:
1124:
1068:
927:
748:
715:
668:
549:
414:
328:
287:
1134:
1094:
1057:
938:
913:
872:
788:
759:
737:
682:
640:
625:
505:
490:
449:
395:
376:
348:
949:
894:
799:
611:
575:
538:
516:
479:
464:
836:
825:
704:
564:
527:
1042:
998:
883:
726:
654:
597:
439:
1031:
774:
586:
1345:
1020:
425:
1109:
273:
1371:
310:
362:
1319:
1358:
1363:
219:
Sandokanidae range in body size from about 2–11 mm. Their legs are relatively short and stout. Most species are
1376:
1237:
17:
1404:
172:
1285:
1280:
1248:
1350:
1306:
8:
1399:
1196:
40:
211:
The name of the type genus is based on the name of a fictional
Bornean prince-pirate.
1293:
1176:
384:
1200:
1188:
1298:
1271:
1050:
236:
123:
1393:
1087:
195:(the name was replaced because of the secondary homonymy of the type genus
159:
1332:
782:
299:
260:
192:
111:
1324:
457:
240:
188:
184:
99:
63:
1337:
1177:"Replacement Names for Oncopus and Oncopodidae (Arachnida, Opiliones)"
1192:
866:
767:
87:
75:
1242:
1265:
1221:(2007): Oncopodidae. Kury & Pérez G., 2002. In: Pinto-da-Rocha
676:
557:
433:
248:
921:
902:
818:
403:
318:
1311:
662:
633:
619:
407:
244:
51:
906:
605:
498:
472:
388:
295:
220:
648:
223:
colored with some dark brown patterns. A few undescribed
259:
Sandokanidae are probably the sister group to all other
1175:Özdikmen, Hüseyin; Kury, Adriano Brilhante (2007).
203:; this made the change of family name mandatory)
1391:
444:Schwendinger, in Martens & Schwendinger 1998
1214:
1212:
1210:
176:Drawing of female Oncopus truncatus Thorrell
1174:
1207:
171:
14:
1392:
1247:
1246:
1170:
1168:
1166:
24:
1084:(Schwendinger & Martens, 2004)
1062:(Schwendinger & Martens, 2004)
1047:(Schwendinger & Martens, 2004)
1014:(Schwendinger & Martens, 2004)
25:
1416:
1163:
1149:Martens & Schwendinger, 1998
1139:Martens & Schwendinger, 1998
1129:Martens & Schwendinger, 1998
1114:Martens & Schwendinger, 1998
976:Schwendinger & Martens, 2002
965:Schwendinger & Martens, 2002
918:Schwendinger & Martens, 2002
899:Schwendinger & Martens, 2002
888:Schwendinger & Martens, 2002
863:Schwendinger & Martens, 2002
841:Schwendinger & Martens, 2002
804:Schwendinger & Martens, 2002
793:Schwendinger & Martens, 2002
764:Schwendinger & Martens, 2002
753:Schwendinger & Martens, 2002
742:Martens & Schwendinger, 1998
731:Schwendinger & Martens, 2002
720:Martens & Schwendinger, 1998
709:Schwendinger & Martens, 2002
698:Schwendinger & Martens, 2002
687:Martens & Schwendinger, 1998
645:Schwendinger & Martens, 2002
591:Schwendinger & Martens, 2002
580:Martens & Schwendinger, 1998
569:Schwendinger & Martens, 2002
543:Schwendinger & Martens, 2002
521:Martens & Schwendinger, 1998
510:Schwendinger & Martens, 2002
484:Martens & Schwendinger, 1998
469:Schwendinger & Martens, 2002
454:Schwendinger & Martens, 2002
430:Martens & Schwendinger, 1998
353:Martens & Schwendinger, 1998
333:Martens & Schwendinger, 1998
315:Martens & Schwendinger, 1998
292:Martens & Schwendinger, 1998
278:Martens & Schwendinger, 1998
254:
239:from Indonesia, almost reaching
230:
214:
13:
1:
1156:
7:
10:
1421:
266:
235:This family is known from
191:, formerly referred to as
166:5 genera, > 60 species
1255:
989:Özdikmen & Kury, 2007
165:
158:
151:
146:
141:Özdikmen & Kury, 2007
41:Scientific classification
39:
34:
206:
1181:Journal of Arachnology
859:Gnomulus rostratoideus
339:Caenoncopus cuspidatus
177:
961:Gnomulus tuberculatus
811:Gnomulus palawanensis
694:Gnomulus latoperculum
175:
1236:'s Biology Catalog:
1080:Sandokan tiomanensis
1073:(Schwendinger, 1992)
1010:Sandokan expatriatus
972:Gnomulus tumidifrons
943:(Schwendinger, 1992)
848:Gnomulus pulvillatus
673:(Schwendinger, 1992)
495:(Schwendinger, 1992)
343:(Schwendinger, 1992)
247:, up north into the
227:species are orange.
27:Family of harvestmen
1405:Harvestman families
1219:Schwendinger, Peter
1145:Palaeoncopus kerdil
1125:Palaeoncopus gunung
1069:Sandokan megachelis
928:Gnomulus sumatranus
749:Gnomulus marginatus
716:Gnomulus leyteensis
669:Gnomulus lannaianus
550:Gnomulus goodnighti
415:Gnomulus armillatus
329:Caenoncopus affinis
288:Biantoncopus fuscus
153:See text for list.
1135:Palaeoncopus katik
1095:Sandokan truncatus
1058:Sandokan malayanus
939:Gnomulus sundaicus
914:Gnomulus spiniceps
873:Gnomulus rostratus
789:Gnomulus monticola
779:N. Tsurusaki, 1990
760:Gnomulus matabesar
738:Gnomulus maculatus
733:— Borneo, Sumatra?
683:Gnomulus laruticus
641:Gnomulus javanicus
626:Gnomulus insularis
506:Gnomulus crassipes
491:Gnomulus conigerus
450:Gnomulus carinatus
396:Gnomulus annulipes
377:Gnomulus aborensis
349:Caenoncopus tenuis
178:
1387:
1386:
1249:Taxon identifiers
1150:
1140:
1130:
1115:
1100:
1085:
1074:
1063:
1048:
1037:
1026:
1015:
1004:
990:
977:
966:
955:
950:Gnomulus thorelli
944:
933:
919:
900:
895:Gnomulus sinensis
889:
878:
865:— West Malaysia,
864:
853:
842:
831:
816:
815:(S. Suzuki, 1982)
805:
800:Gnomulus obscurus
794:
780:
765:
754:
743:
732:
721:
710:
699:
688:
674:
660:
646:
631:
617:
612:Gnomulus imadatei
603:
592:
581:
576:Gnomulus hirsutus
570:
555:
544:
539:Gnomulus exsudans
533:
522:
517:Gnomulus crucifer
511:
496:
485:
480:Gnomulus coniceps
470:
465:Gnomulus claviger
455:
445:
431:
420:
401:
385:Arunachal Pradesh
382:
368:
354:
344:
334:
316:
293:
279:
170:
169:
16:(Redirected from
1412:
1380:
1379:
1367:
1366:
1354:
1353:
1341:
1340:
1328:
1327:
1315:
1314:
1302:
1301:
1289:
1288:
1276:
1275:
1274:
1244:
1243:
1235:
1226:
1220:
1216:
1205:
1204:
1193:10.1636/h05-46.1
1172:
1148:
1138:
1128:
1113:
1098:
1083:
1072:
1061:
1046:
1035:
1024:
1013:
1002:
988:
975:
964:
954:(Sørensen, 1932)
953:
942:
931:
917:
898:
887:
876:
862:
851:
840:
837:Gnomulus pilosus
829:
826:Gnomulus piliger
814:
803:
792:
778:
763:
752:
741:
730:
719:
708:
705:Gnomulus leofeae
697:
686:
672:
658:
644:
629:
615:
601:
590:
579:
568:
565:Gnomulus hamatus
553:
542:
531:
528:Gnomulus drescoi
520:
509:
494:
483:
468:
453:
443:
429:
418:
399:
380:
366:
352:
342:
332:
314:
291:
277:
187:in the suborder
32:
31:
21:
1420:
1419:
1415:
1414:
1413:
1411:
1410:
1409:
1390:
1389:
1388:
1383:
1375:
1370:
1362:
1357:
1349:
1344:
1336:
1331:
1323:
1318:
1310:
1305:
1297:
1292:
1284:
1279:
1270:
1269:
1264:
1251:
1233:
1230:
1229:
1218:
1217:
1208:
1173:
1164:
1159:
1064:— West Malaysia
1043:Sandokan lingga
999:Sandokan doriae
884:Gnomulus ryssie
854:— West Malaysia
843:— West Malaysia
795:— West Malaysia
727:Gnomulus lomani
689:— West Malaysia
655:Gnomulus laevis
602:(Martens, 1977)
598:Gnomulus hyatti
582:— West Malaysia
532:(Silhavy, 1962)
440:Gnomulus baharu
419:(Thorell, 1891)
269:
257:
233:
217:
209:
183:is a family of
142:
139:
138:
126:
114:
102:
90:
78:
66:
54:
28:
23:
22:
15:
12:
11:
5:
1418:
1408:
1407:
1402:
1385:
1384:
1382:
1381:
1368:
1355:
1342:
1329:
1316:
1303:
1290:
1277:
1261:
1259:
1253:
1252:
1241:
1240:
1228:
1227:
1206:
1187:(2): 407–408.
1161:
1160:
1158:
1155:
1154:
1153:
1152:
1151:
1141:
1131:
1118:
1117:
1105:
1104:
1103:
1102:
1091:
1076:
1065:
1054:
1051:Lingga Islands
1039:
1032:Sandokan hosei
1028:
1017:
1006:
992:
991:
982:
981:
980:
979:
968:
957:
946:
935:
924:
910:
891:
880:
869:
855:
852:(Pocock, 1903)
844:
833:
830:(Pocock, 1903)
822:
807:
796:
785:
775:Gnomulus minor
771:
756:
745:
734:
723:
712:
701:
690:
679:
665:
659:(Roewer, 1915)
651:
637:
630:(Roewer, 1927)
622:
616:(Suzuki, 1970)
608:
594:
587:Gnomulus hutan
583:
572:
561:
554:(Suzuki, 1977)
546:
535:
524:
513:
502:
487:
476:
461:
446:
436:
422:
411:
400:(Pocock, 1897)
392:
381:(Roewer, 1913)
370:
369:
358:
357:
356:
355:
345:
335:
322:
321:
306:
305:
304:
303:
281:
280:
268:
265:
256:
253:
237:Southeast Asia
232:
229:
216:
213:
208:
205:
199:, replaced by
168:
167:
163:
162:
156:
155:
149:
148:
144:
143:
140:
134:
132:
128:
127:
124:Phalangodoidea
122:
120:
116:
115:
110:
108:
104:
103:
98:
96:
92:
91:
86:
84:
80:
79:
74:
72:
68:
67:
62:
60:
56:
55:
50:
48:
44:
43:
37:
36:
26:
9:
6:
4:
3:
2:
1417:
1406:
1403:
1401:
1398:
1397:
1395:
1378:
1373:
1369:
1365:
1360:
1356:
1352:
1347:
1343:
1339:
1334:
1330:
1326:
1321:
1317:
1313:
1308:
1304:
1300:
1295:
1291:
1287:
1282:
1278:
1273:
1267:
1263:
1262:
1260:
1258:
1254:
1250:
1245:
1239:
1232:
1231:
1224:
1215:
1213:
1211:
1202:
1198:
1194:
1190:
1186:
1182:
1178:
1171:
1169:
1167:
1162:
1147:
1146:
1142:
1137:
1136:
1132:
1127:
1126:
1122:
1121:
1120:
1119:
1112:
1111:
1107:
1106:
1099:Thorell, 1891
1097:
1096:
1092:
1089:
1088:Tioman Island
1082:
1081:
1077:
1071:
1070:
1066:
1060:
1059:
1055:
1052:
1045:
1044:
1040:
1034:
1033:
1029:
1025:Thorell, 1890
1023:
1022:
1021:Sandokan feae
1018:
1012:
1011:
1007:
1003:Thorell, 1876
1001:
1000:
996:
995:
994:
993:
987:
984:
983:
974:
973:
969:
963:
962:
958:
952:
951:
947:
941:
940:
936:
932:Thorell, 1891
930:
929:
925:
923:
916:
915:
911:
908:
904:
897:
896:
892:
886:
885:
881:
877:Thorell, 1890
875:
874:
870:
868:
861:
860:
856:
850:
849:
845:
839:
838:
834:
828:
827:
823:
821:(Philippines)
820:
813:
812:
808:
802:
801:
797:
791:
790:
786:
784:
777:
776:
772:
769:
762:
761:
757:
751:
750:
746:
740:
739:
735:
729:
728:
724:
718:
717:
713:
707:
706:
702:
696:
695:
691:
685:
684:
680:
678:
671:
670:
666:
664:
657:
656:
652:
650:
643:
642:
638:
635:
628:
627:
623:
621:
614:
613:
609:
607:
600:
599:
595:
589:
588:
584:
578:
577:
573:
567:
566:
562:
560:(Philippines)
559:
552:
551:
547:
541:
540:
536:
530:
529:
525:
519:
518:
514:
508:
507:
503:
500:
493:
492:
488:
482:
481:
477:
475:(Philippines)
474:
467:
466:
462:
459:
452:
451:
447:
442:
441:
437:
435:
428:
427:
426:Gnomulus asli
423:
417:
416:
412:
409:
405:
398:
397:
393:
390:
386:
379:
378:
374:
373:
372:
371:
367:Thorell, 1890
365:
364:
360:
359:
351:
350:
346:
341:
340:
336:
331:
330:
326:
325:
324:
323:
320:
313:
312:
308:
307:
301:
297:
290:
289:
285:
284:
283:
282:
276:
275:
271:
270:
264:
262:
255:Relationships
252:
250:
246:
242:
238:
228:
226:
222:
212:
204:
202:
198:
194:
190:
186:
182:
174:
164:
161:
157:
154:
150:
145:
137:
133:
130:
129:
125:
121:
119:Superfamily:
118:
117:
113:
109:
106:
105:
101:
97:
94:
93:
89:
85:
82:
81:
77:
73:
70:
69:
65:
61:
58:
57:
53:
49:
46:
45:
42:
38:
35:Sandokanidae
33:
30:
19:
18:Sandokanoidea
1257:Sandokanidae
1256:
1222:
1184:
1180:
1143:
1133:
1123:
1110:Palaeoncopus
1108:
1093:
1078:
1067:
1056:
1041:
1036:Pocock, 1897
1030:
1019:
1008:
997:
985:
970:
959:
948:
937:
926:
912:
893:
882:
871:
857:
846:
835:
824:
809:
798:
787:
773:
758:
747:
736:
725:
714:
703:
692:
681:
667:
653:
639:
636:? (Malaysia)
624:
610:
596:
585:
574:
563:
548:
537:
526:
515:
504:
489:
478:
463:
448:
438:
424:
413:
394:
375:
361:
347:
337:
327:
309:
286:
274:Biantoncopus
272:
258:
234:
231:Distribution
224:
218:
210:
200:
196:
181:Sandokanidae
180:
179:
152:
136:Sandokanidae
135:
107:Infraorder:
29:
1333:iNaturalist
1238:Oncopodidae
1234:Joel Hallan
1225:2007: 211ff
1101:— Singapore
1053:(Indonesia)
1016:— Thailand?
978:— Halmahera
783:Philippines
770:(Indonesia)
460:(Indonesia)
387:(northeast
311:Caenoncopus
300:Philippines
261:Grassatores
215:Description
193:Oncopodidae
112:Grassatores
1400:Harvestmen
1394:Categories
1157:References
1090:(Malaysia)
890:— Thailand
832:— Thailand
755:— Thailand
700:— Sulawesi
501:(Malaysia)
458:Kalimantan
241:New Guinea
189:Laniatores
185:harvestmen
100:Laniatores
95:Suborder:
64:Arthropoda
1116:— Sumatra
1038:— Sarawak
1005:— Sarawak
967:— Sumatra
945:— Sarawak
934:— Sumatra
867:Singapore
806:— Sarawak
768:Halmahera
593:— Sarawak
534:— Sumatra
421:— Sumatra
249:Himalayan
160:Diversity
88:Opiliones
76:Arachnida
47:Kingdom:
1272:Q3471872
1266:Wikidata
1201:86046606
1027:— Penang
986:Sandokan
879:— Penang
677:Thailand
558:Mindanao
434:Malaysia
363:Gnomulus
251:region.
225:Gnomulus
201:Sandokan
147:Species
131:Family:
59:Phylum:
52:Animalia
1364:1012459
1325:7906794
1312:6877540
1075:— Sabah
922:Vietnam
903:Sichuan
819:Palawan
744:— Luzon
722:— Leyte
711:— Burma
571:— Luzon
545:— Sabah
523:— Luzon
512:— Luzon
486:— Luzon
432:— West
404:Sarawak
319:Sumatra
267:Species
197:Oncopus
83:Order:
71:Class:
1377:648777
1351:120739
1338:535008
1286:718926
1223:et al.
1199:
956:— Java
663:Borneo
634:Penang
620:Brunei
408:Borneo
245:Waigeo
1346:IRMNG
1197:S2CID
907:China
606:Nepal
499:Sabah
473:Luzon
389:India
296:Leyte
221:amber
1372:NCBI
1359:ITIS
1320:GBIF
1281:BOLD
649:Java
207:Name
1307:EoL
1299:FXK
1294:CoL
1189:doi
243:on
1396::
1374::
1361::
1348::
1335::
1322::
1309::
1296::
1283::
1268::
1209:^
1195:.
1185:35
1183:.
1179:.
1165:^
1086:—
1049:—
920:—
901:—
817:—
781:—
766:—
675:—
661:—
647:—
632:—
618:—
604:—
556:—
497:—
471:—
456:—
402:—
383:—
317:—
294:—
263:.
1203:.
1191::
909:)
905:(
410:)
406:(
391:)
302:)
298:(
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
