680:
3307:
5430:
3300:
1804:
2629:
2615:
4882:
4095:
4868:
2608:
5575:
4875:
1810:
3314:
148:
2622:
5443:
4113:
403:
4101:
595:
4120:
5437:
4862:
126:
4108:
5563:
1198:
394:. It has been used as a morphotaxon to describe other theropod teeth widely displaced in time and space from the type species. If all teeth assigned to the genus are truly reflective of the animals biology and taxonomic state (as some teeth go as far back as the Late Jurassic), it would have been one of the longest lasting dinosaur genera, perhaps also being the most widely distributed.
526:
age, about 75 million years ago. The jaws are slender and rather long, 193 millimeters, but the teeth are small and very finely serrated with five to six denticles per millimeter. The serration density is a distinctive trait of the species.
480:
being the normal latinisation of "Richard". However, except in one overlooked figure caption, the editors of the paper altered the spelling to include the 'h'. Ironically, in 1991 George
Olshevsky in a species list also used the spelling
493:
rules, he acted as "first revisor" choosing between the two spelling variants of the original publication and inadvertently made the misspelt name official. Subsequently, the original authors have adopted the spelling
636:
Because of the disparity in location and time of the many referred teeth, researchers have cast doubt on the idea that they all belong to the same genus or species, and the genus is best considered a
660:, roughly the same age. All other referred teeth most likely belong to different species, which have not been named due to the lack of body fossils for comparison.
472:, to honor his important work on small vertebrates and especially theropod teeth of the Late Cretaceous. The naming authors actually intended to use the spelling
1145:
Investigations on
Canadian dinosaurs: systematic palaeontology of isolated dinosaur teeth from the Latest Cretaceous of Dinosaur Provincial Park, Alberta, Canada
427:
5630:
458:
454:
756:
Currie PJ, Rigby KJ, Sloan RE (1990). "Theropod teeth from the Judith River
Formation of southern Alberta, Canada". In Currie PJ, Carpenter K (eds.).
531:
490:
5605:
898:
Sues HD, Averianov A (2013). "Enigmatic teeth of small theropod dinosaurs from the Upper
Cretaceous (Cenomanian–Turonian) of Uzbekistan".
671:
of northeastern Spain (Blasi 2 member). The oldest fossils that have been referred to
Richardoestesia come from the Jurassic of Portugal.
5655:
5660:
5507:
5670:
5665:
489:
to be the misspelling, unaware that the original authors actually intended the name to be spelled this way. As a result, under
3306:
3299:
1287:
5429:
1034:
886:(in Russian). St. Petersburg, Russia: Scientific Research Institute of the Earth's Crust, St. Petersburg State University.
1266:
1235:
1215:
786:
469:
5620:
5645:
1202:
1124:
Longrich N (2008). "Small theropod teeth from the Lance
Formation of Wyoming, USA". In Sankey JT, Baszio S (eds.).
977:"Multivariate Analyses of Small Theropod Dinosaur Teeth and Implications for Paleoecological Turnover through Time"
4094:
2628:
2614:
1182:
855:
824:
4867:
1152:
5308:
2607:
1803:
4112:
1809:
147:
5650:
4881:
1095:"Presence of a maniraptoriform dinosaur in the Late Cretaceous (Maastrichtian) of Chiapas, Southern Mexico"
625:(dated to about 66 million years ago). Similar teeth have been referred to this genus from as early as the
3313:
679:
5600:
2621:
936:
Averianov A, Sues HD (2019). "Morphometric analysis of the teeth and taxonomy of the enigmatic theropod
5640:
5635:
5553:
4874:
606:
415:
4119:
5615:
5610:
5442:
1164:"Small theropod and bird teeth from the Late Cretaceous (Late Campanian) Judith River Group, Alberta"
873:
5625:
1354:
630:
419:
114:
5436:
4100:
1259:
649:
499:
431:
5538:
5462:
5595:
5533:
5525:
3825:
3737:
586:
is CCMGE 460/12457, and two other teeth (ZIN PH 1110/ 16 and ZIN PH 1129/16) are also known.
5494:
5394:
3514:
3482:
1067:
988:
949:
907:
875:Динозавры северной Евразии: Новые данные о составе комплексов, экологии и палеобиогеографии
614:
4861:
726:. New Mexico Museum of Natural History and Science Bulletin. Vol. 14. pp. 79–89.
8:
5579:
3838:
3625:
737:
Gilmore CW (1924). "A new coelurid dinosaur from the Belly River
Cretaceous of Alberta".
637:
605:-like teeth have been found in many Late Cretaceous geological formations, including the
354:
4107:
1071:
992:
953:
911:
880:
Dinosaurs of
Northern Eurasia: new data about assemblages, ecology and paleobiogeography
5301:
3814:
3674:
3530:
3470:
2905:
2815:
2698:
1562:
1252:
1055:
1011:
976:
610:
567:
519:
317:
142:
1163:
805:
5520:
5183:
5100:
5074:
4913:
4900:
4778:
4328:
4280:
3112:
2462:
1541:
1148:
1094:
1016:
719:
1126:
Vertebrate microfossil assemblages: their role in paleoecology and paleobiogeography
5567:
5287:
5044:
4954:
4743:
4667:
4530:
4449:
4421:
4381:
4262:
4232:
3750:
3503:
3367:
3224:
3124:
2973:
2880:
2838:
2405:
1715:
1178:
1106:
1075:
1006:
996:
957:
915:
851:
838:
Company J, Suberbiola XP, Ruiz-Omeñaca JI, Buscalioni AD (2005). "A new species of
820:
684:
618:
450:
106:
961:
5350:
5088:
5051:
4983:
4838:
4831:
4651:
4593:
4566:
4556:
4516:
4351:
4194:
4012:
3984:
3962:
3849:
3694:
3401:
3175:
3058:
3036:
2957:
2786:
2763:
2756:
2734:
2720:
2713:
2674:
2525:
2372:
2299:
2288:
2224:
2112:
1999:
1576:
1485:
1339:
1314:
1239:
1219:
1001:
668:
657:
622:
539:
365:
30:
1056:"Small theropod teeth from the Upper Jurassic coal mine of Guimarota (Portugal)"
787:"Re: "Mega-raptors" and the high frequency of undescribed dromaeosaurs in lists"
5485:
5315:
5278:
5259:
5221:
5210:
5007:
4720:
4678:
4660:
4640:
4537:
4251:
3996:
3858:
3685:
3667:
3495:
3462:
3448:
3409:
3183:
3044:
3001:
2994:
2916:
2892:
2845:
2804:
2741:
2727:
2455:
2440:
2267:
2101:
1965:
1933:
1922:
1899:
1861:
1722:
1692:
1616:
1555:
1454:
446:
361:
5589:
5357:
5203:
5164:
5143:
5022:
5015:
4942:
4736:
4709:
4575:
4492:
4470:
4463:
4456:
4399:
4316:
4179:
4151:
4138:
4062:
3939:
3779:
3613:
3544:
3430:
3417:
3162:
3140:
3133:
3097:
3070:
3022:
2946:
2682:
2473:
2447:
2363:
2344:
2337:
2274:
2253:
2232:
2217:
2203:
2162:
2128:
2121:
2040:
1892:
1779:
1772:
1699:
1669:
1658:
1548:
1519:
1512:
1438:
1110:
888: – in Russian with short English, German, and French abstracts
775:. University of California Publications in Geological Sciences. Vol. 49.
551:
547:
436:
373:
231:
68:
1224:
1208:
790:
773:
Fossil vertebrates from the Late
Cretaceous Lance Formation, eastern Wyoming
640:. A comparative study of the teeth published in 2013 demonstrated that both
402:
5329:
5267:
5237:
5157:
5150:
5128:
5120:
5000:
4992:
4972:
4687:
4606:
4544:
4435:
4406:
4358:
4187:
4172:
3947:
3879:
3807:
3786:
3765:
3581:
3537:
3345:
3332:
3262:
3215:
3090:
2980:
2932:
2925:
2862:
2831:
2824:
2705:
2532:
2384:
2281:
2183:
2176:
2071:
2055:
1991:
1942:
1874:
1651:
1534:
1527:
1498:
1461:
1430:
1020:
562:
390:
303:
257:
5383:
5364:
5294:
5244:
5230:
5172:
5113:
5058:
4935:
4729:
4613:
4582:
4499:
4485:
4217:
4165:
4033:
3955:
3918:
3800:
3772:
3725:
3645:
3523:
3380:
3231:
3204:
3104:
3051:
2987:
2966:
2939:
2690:
2503:
2496:
2482:
2308:
2155:
2146:
2048:
2019:
1949:
1914:
1907:
1707:
1685:
1678:
1637:
1629:
1605:
1590:
1569:
1477:
1469:
1147:. Courier Forschungsinstitut Senckenberg. Vol. 196. pp. 33–77.
1039:
739:
Canada
Department of Mines Geological Survey Bulletin (Geological Series)
688:
558:
441:
and referred the jaws to this species. In the 1980s it became clear that
43:
698:, due to its vast distribution as well as predominance in marine sites.
594:
5512:
5322:
5251:
5196:
4817:
4624:
4442:
4428:
4413:
4365:
4299:
4225:
4047:
4040:
4026:
3932:
3865:
3793:
3757:
3710:
3652:
3638:
3605:
3597:
3590:
3571:
3394:
3359:
3147:
2748:
2539:
2489:
2426:
2397:
2352:
2330:
2323:
2315:
2239:
2135:
2094:
2087:
2079:
2063:
1983:
1854:
1730:
1583:
1446:
1305:
1079:
837:
571:
377:
205:
88:
53:
919:
5343:
5336:
5036:
5029:
4702:
4694:
4633:
4287:
4005:
3551:
3455:
3190:
2775:
2660:
2647:
2574:
2510:
2433:
2412:
2169:
2032:
1840:
1827:
1745:
1597:
1416:
1403:
1366:
1275:
806:"Late Campanian southern dinosaurs, Aguja Formation, Big Bend, Texas"
626:
523:
218:
159:
110:
93:
37:
5456:
5479:
4927:
4344:
4337:
4309:
4202:
3702:
3083:
2419:
2246:
2007:
1644:
1323:
1299:
842:(Crocodyliformes: Ziphosuchia) from the Late Cretaceous of Spain".
694:
At least a few studies have speculated a piscivorous lifestyle for
511:
449:
to which the long jaws could not have belonged. Therefore, in 1990
358:
192:
179:
83:
78:
63:
58:
48:
125:
4477:
4239:
3969:
3872:
3197:
2518:
423:
98:
73:
5499:
5136:
4824:
2260:
1293:
1197:
369:
169:
518:(NMC 343) consists of a pair of lower jaws found in the upper
3976:
3717:
3659:
582:. A subsequent study confirmed this in 2019. The holotype of
1244:
1161:
542:, which is of a longer and less recurved type. The teeth of
5400:
4019:
1183:
10.1666/0022-3360(2002)076<0751:STABTF>2.0.CO;2
856:
10.1671/0272-4634(2005)025[0343:ANSODC]2.0.CO;2
825:
10.1666/0022-3360(2001)075<0208:LCSDAF>2.0.CO;2
1128:. Bloomington: Indiana University Press. pp. 135–158.
1092:
760:. Cambridge: Cambridge University Press. pp. 107–125.
1162:
Sankey JT, Brinkman DB, Guenther M, Currie PJ (2002).
5551:
538:, based on a tooth, LSUMGS 489:6238, from the Texan
299:
287:
264:
4809:
1093:Carbot-Chanona G, Rivera-Sylva HE (December 2011).
724:Lower and Middle Cretaceous Terrestrial Ecosystems
717:
652:, dated to between 76.5 and 75 million years ago.
758:Dinosaur Systematics: Perspectives and Approaches
5587:
4911:
4777:
789:(Mailing list). 22 November 1997. Archived from
755:
4260:
2784:
1338:
720:"Cretaceous dinosaurs of the Colorado Plateau"
557:In 2013 a study assumed that the teeth of the
1260:
935:
897:
722:. In Kirkland JI, Lucas SG, Estep JW (eds.).
4149:
974:
867:
865:
3343:
3261:
2383:
713:
711:
5631:Late Cretaceous dinosaurs of North America
1267:
1253:
940:from the Upper Cretaceous of Uzbekistan".
124:
1099:Boletin de la Sociedad Geologica Mexicana
1010:
1000:
968:
862:
2658:
2573:
1838:
1414:
1123:
1027:
718:Kirkland JI, Lucas SG, Estep JW (1998).
708:
678:
593:
401:
736:
5588:
1142:
871:
803:
648:were only definitively present in the
598:Referred teeth from the San Juan Basin
5461:
5460:
4899:
4898:
4136:
3330:
2645:
1825:
1401:
1286:
1248:
1053:
931:
929:
831:
770:
380:. It currently contains two species,
5606:Late Cretaceous dinosaurs of Europe
13:
1136:
942:Journal of Vertebrate Paleontology
926:
900:Canadian Journal of Earth Sciences
844:Journal of Vertebrate Paleontology
550:in shape, possibly belonging to a
14:
5682:
5656:Campanian genus first appearances
4137:
1190:
589:
5573:
5561:
5441:
5435:
5428:
4880:
4873:
4866:
4860:
4118:
4111:
4106:
4099:
4093:
3331:
3312:
3305:
3298:
2627:
2620:
2613:
2606:
1808:
1802:
1196:
364:, originally described from the
146:
41:
5661:Maastrichtian genus extinctions
1117:
1086:
1047:
674:
578:, and renamed the species into
414:The jaws were found in 1917 by
339:Currie, Rigby & Sloan, 1990
274:Currie, Rigby & Sloan, 1990
251:Currie, Rigby & Sloan, 1990
5671:Taxa named by Philip J. Currie
1225:First message of a passionate
891:
797:
779:
764:
749:
730:
566:, from the CBI-14 site of the
1:
5666:Fossil taxa described in 1990
2646:
1826:
1402:
1274:
975:Larson DW, Currie PJ (2013).
962:10.1080/02724634.2019.1614941
701:
574:, were identical to those of
546:have also been identified as
1060:Paläontologische Zeitschrift
1002:10.1371/journal.pone.0054329
667:have also been found in the
113:) records from teeth of the
7:
137:with close up of denticles
10:
5687:
633:, 125 million years ago).
607:Horseshoe Canyon Formation
505:
461:named a separate species:
416:Charles Hazelius Sternberg
410:with close up of denticles
397:
309:Averianov & Sues, 2013
16:Extinct genus of dinosaurs
5469:
5425:
5382:
5277:
5220:
5191:
5182:
5108:
5099:
5083:
5072:
4982:
4967:
4963:
4952:
4922:
4907:
4894:
4858:
4805:
4773:
4719:
4677:
4650:
4623:
4601:
4592:
4565:
4554:
4525:
4514:
4394:
4390:
4379:
4327:
4298:
4275:
4271:
4249:
4160:
4145:
4132:
4091:
4061:
3995:
3927:
3917:
3913:
3848:
3833:
3824:
3745:
3736:
3684:
3633:
3624:
3580:
3569:
3513:
3490:
3481:
3443:
3439:
3428:
3389:
3378:
3354:
3339:
3326:
3295:
3257:
3214:
3170:
3161:
3123:
3078:
3069:
3031:
3020:
2956:
2915:
2900:
2891:
2875:
2871:
2860:
2814:
2799:
2795:
2773:
2669:
2654:
2641:
2603:
2569:
2472:
2392:
2362:
2298:
2212:
2201:
2145:
2111:
2027:
2018:
1978:
1974:
1963:
1932:
1887:
1883:
1872:
1849:
1834:
1821:
1799:
1764:
1744:
1668:
1624:
1615:
1507:
1496:
1425:
1410:
1397:
1365:
1347:
1334:
1282:
576:Richardoestesia isosceles
536:Richardoestesia isosceles
323:
316:
284:
279:
263:
256:
143:Scientific classification
141:
132:
123:
23:
1111:10.18268/BSGM2011v63n3a2
1054:Zinke J (1 April 1998).
874:
656:was also present in the
631:Cedar Mountain Formation
580:Richardoestesia asiatica
534:named a second species:
516:Richardoestesia gilmorei
463:Richardoestesia gilmorei
420:Dinosaur Provincial Park
388:, and a possible third,
269:Richardoestesia gilmorei
115:Cedar Mountain Formation
5646:Paleontology in Alberta
1171:Journal of Paleontology
813:Journal of Paleontology
650:Dinosaur Park Formation
468:The genus is named for
432:Charles Whitney Gilmore
336:Ricardoestesia gilmorei
5621:La Huérguina Formation
691:
599:
411:
5534:Paleobiology Database
3826:Carcharodontosaurinae
3738:Carcharodontosauridae
682:
597:
563:Asiamericana asiatica
428:Little Sandhill Creek
405:
327:Asiamericana asiatica
4810:Dubious coelurosaurs
3515:Metriacanthosaurinae
3483:Metriacanthosauridae
1765:Dubious neotheropods
1205:at Wikimedia Commons
615:Hell Creek Formation
5651:Ojo Alamo Formation
3839:Carcharodontosaurus
3626:Carcharodontosauria
1072:1998PalZ...72..179Z
993:2013PLoSO...854329L
954:2019JVPal..39E4941A
912:2013CaJES..50..306S
5601:Maastrichtian life
5302:Archaeornithomimus
3815:Veterupristisaurus
3675:Veterupristisaurus
3531:Metriacanthosaurus
3471:Piatnitzkysauridae
2906:Eustreptospondylus
2816:Piatnitzkysauridae
2699:Chuandongocoelurus
1563:Notatesseraeraptor
1238:2020-08-06 at the
1218:2016-08-06 at the
1213:typo" is described
1080:10.1007/BF02987825
872:Nessov LA (1995).
804:Sankey JT (2001).
692:
611:Scollard Formation
600:
568:Bissekty Formation
522:, dating from the
520:Judith River Group
412:
5641:Fossils of Canada
5636:Cretaceous Canada
5549:
5548:
5521:Open Tree of Life
5463:Taxon identifiers
5454:
5453:
5450:
5449:
5423:
5422:
5419:
5418:
5415:
5414:
5411:
5410:
5378:
5377:
5374:
5373:
5184:Macrocheiriformes
5101:Ornithomimosauria
5075:Maniraptoriformes
5068:
5067:
4914:Maniraptoromorpha
4901:Maniraptoromorpha
4890:
4889:
4856:
4855:
4852:
4851:
4848:
4847:
4801:
4800:
4797:
4796:
4779:Maniraptoromorpha
4769:
4768:
4765:
4764:
4761:
4760:
4757:
4756:
4753:
4752:
4510:
4509:
4375:
4374:
4329:Proceratosauridae
4281:Chingkankousaurus
4128:
4127:
4089:
4088:
4085:
4084:
4081:
4080:
4057:
4056:
3909:
3908:
3905:
3904:
3901:
3900:
3897:
3896:
3893:
3892:
3889:
3888:
3565:
3564:
3561:
3560:
3322:
3321:
3293:
3292:
3289:
3288:
3285:
3284:
3281:
3280:
3253:
3252:
3249:
3248:
3245:
3244:
3241:
3240:
3157:
3156:
3113:Vallibonavenatrix
3016:
3015:
3012:
3011:
2856:
2855:
2637:
2636:
2601:
2600:
2597:
2596:
2593:
2592:
2565:
2564:
2561:
2560:
2557:
2556:
2553:
2552:
2549:
2548:
2197:
2196:
2193:
2192:
1959:
1958:
1817:
1816:
1797:
1796:
1793:
1792:
1789:
1788:
1740:
1739:
1393:
1392:
1389:
1388:
1385:
1384:
1201:Media related to
1143:Baszio S (1997).
920:10.1139/e2012-033
793:on 6 August 2016.
459:Robert Evan Sloan
346:
345:
340:
332:
310:
296:
252:
118:
5678:
5616:Fossils of Spain
5611:Cretaceous Spain
5578:
5577:
5576:
5566:
5565:
5564:
5557:
5542:
5541:
5529:
5528:
5516:
5515:
5503:
5502:
5490:
5489:
5488:
5458:
5457:
5445:
5439:
5432:
5288:Aepyornithomimus
5189:
5188:
5106:
5105:
5081:
5080:
5045:Sinocalliopteryx
4965:
4964:
4961:
4960:
4955:Neocoelurosauria
4920:
4919:
4909:
4908:
4896:
4895:
4884:
4877:
4870:
4864:
4807:
4806:
4775:
4774:
4744:Zhuchengtyrannus
4668:Thanatotheristes
4599:
4598:
4563:
4562:
4531:Appalachiosaurus
4523:
4522:
4392:
4391:
4388:
4387:
4382:Pantyrannosauria
4273:
4272:
4269:
4268:
4263:Tyrannosauroidea
4258:
4257:
4233:Xinjiangovenator
4158:
4157:
4147:
4146:
4134:
4133:
4122:
4115:
4110:
4103:
4097:
3925:
3924:
3915:
3914:
3831:
3830:
3751:Acrocanthosaurus
3743:
3742:
3631:
3630:
3578:
3577:
3504:Yangchuanosaurus
3488:
3487:
3441:
3440:
3437:
3436:
3387:
3386:
3368:Lourinhanosaurus
3352:
3351:
3341:
3340:
3328:
3327:
3316:
3309:
3302:
3259:
3258:
3225:Sigilmassasaurus
3168:
3167:
3125:Ceratosuchopsini
3076:
3075:
3029:
3028:
2974:Dubreuillosaurus
2898:
2897:
2881:Streptospondylus
2873:
2872:
2869:
2868:
2839:Piatnitzkysaurus
2797:
2796:
2793:
2792:
2782:
2781:
2667:
2666:
2656:
2655:
2643:
2642:
2631:
2624:
2617:
2610:
2571:
2570:
2406:Ekrixinatosaurus
2390:
2389:
2381:
2380:
2210:
2209:
2025:
2024:
1976:
1975:
1972:
1971:
1885:
1884:
1881:
1880:
1847:
1846:
1836:
1835:
1823:
1822:
1812:
1806:
1716:Procompsognathus
1622:
1621:
1505:
1504:
1423:
1422:
1412:
1411:
1399:
1398:
1345:
1344:
1336:
1335:
1329:
1328:
1284:
1283:
1269:
1262:
1255:
1246:
1245:
1200:
1186:
1168:
1158:
1130:
1129:
1121:
1115:
1114:
1090:
1084:
1083:
1051:
1045:
1044:
1031:
1025:
1024:
1014:
1004:
972:
966:
965:
933:
924:
923:
895:
889:
887:
885:
869:
860:
859:
835:
829:
828:
810:
801:
795:
794:
783:
777:
776:
771:Estes R (1964).
768:
762:
761:
753:
747:
746:
734:
728:
727:
715:
685:life restoration
619:Ferris Formation
502:honors Gilmore.
485:, and indicated
455:John Keith Rigby
418:and sons in the
338:
330:
308:
301:
294:
289:
266:
250:
243:
151:
150:
128:
107:Early Cretaceous
104:
103:
40:
29:Temporal range:
21:
20:
5686:
5685:
5681:
5680:
5679:
5677:
5676:
5675:
5626:Tremp Formation
5586:
5585:
5584:
5574:
5572:
5562:
5560:
5552:
5550:
5545:
5537:
5532:
5524:
5519:
5511:
5506:
5498:
5493:
5484:
5483:
5478:
5471:Richardoestesia
5465:
5455:
5446:
5433:
5407:
5370:
5351:Sinornithomimus
5273:
5216:
5178:
5095:
5089:Compsognathidae
5077:
5064:
5052:Sinosauropteryx
4984:Compsognathidae
4978:
4957:
4948:
4916:
4903:
4886:
4885:
4871:
4844:
4839:Shanyangosaurus
4832:Phaedrolosaurus
4793:
4749:
4715:
4673:
4652:Daspletosaurini
4646:
4619:
4594:Tyrannosaurinae
4588:
4567:Albertosaurinae
4559:
4557:Tyrannosauridae
4550:
4519:
4517:Eutyrannosauria
4506:
4384:
4371:
4352:Proceratosaurus
4323:
4294:
4265:
4254:
4245:
4211:Richardoestesia
4195:Chilantaisaurus
4154:
4141:
4124:
4123:
4104:
4077:
4053:
4013:Australovenator
3991:
3985:Chilantaisaurus
3963:Phuwiangvenator
3885:
3850:Giganotosaurini
3844:
3820:
3732:
3695:Chilantaisaurus
3680:
3620:
3574:
3557:
3509:
3477:
3433:
3424:
3402:Monolophosaurus
3383:
3374:
3348:
3335:
3318:
3317:
3303:
3277:
3237:
3210:
3176:Camarillasaurus
3153:
3119:
3065:
3059:Ostafrikasaurus
3037:Camarillasaurus
3025:
3008:
2958:Afrovenatorinae
2952:
2911:
2887:
2865:
2852:
2810:
2789:
2787:Megalosauroidea
2778:
2769:
2764:Vectaerovenator
2757:Szechuanosaurus
2735:Monolophosaurus
2721:Kaijiangosaurus
2714:Cryolophosaurus
2675:Calamospondylus
2663:
2650:
2633:
2632:
2625:
2618:
2611:
2589:
2545:
2526:Pycnonemosaurus
2468:
2379:
2373:Majungasaurinae
2358:
2300:Majungasaurinae
2294:
2289:Xenotarsosaurus
2225:Dryptosauroides
2206:
2189:
2141:
2113:Elaphrosaurinae
2107:
2014:
2000:Ornithomimoides
1968:
1955:
1928:
1877:
1868:
1843:
1830:
1813:
1807:
1785:
1760:
1736:
1664:
1611:
1577:Shuangbaisaurus
1501:
1492:
1419:
1406:
1381:
1361:
1355:Avemetatarsalia
1340:Avemetatarsalia
1330:
1315:Avemetatarsalia
1289:
1288:
1278:
1273:
1240:Wayback Machine
1231:Richardoestesia
1220:Wayback Machine
1203:Richardoestesia
1193:
1166:
1155:
1139:
1137:Further reading
1134:
1133:
1122:
1118:
1091:
1087:
1052:
1048:
1033:
1032:
1028:
973:
969:
948:(3): e1614941.
938:Richardoestesia
934:
927:
896:
892:
883:
876:
870:
863:
836:
832:
808:
802:
798:
785:
784:
780:
769:
765:
754:
750:
735:
731:
716:
709:
704:
696:Richardoestesia
677:
669:Tremp Formation
665:Richardoestesia
658:Aguja Formation
623:Lance Formation
603:Richardoestesia
592:
540:Aguja Formation
508:
496:Richardoestesia
483:Richardoestesia
400:
368:of what is now
366:Late Cretaceous
350:Richardoestesia
293:
275:
272:
249:
246:Richardoestesia
241:
145:
119:
102:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
31:Late Cretaceous
27:
25:Richardoestesia
17:
12:
11:
5:
5684:
5674:
5673:
5668:
5663:
5658:
5653:
5648:
5643:
5638:
5633:
5628:
5623:
5618:
5613:
5608:
5603:
5598:
5583:
5582:
5570:
5547:
5546:
5544:
5543:
5530:
5517:
5504:
5491:
5475:
5473:
5467:
5466:
5452:
5451:
5448:
5447:
5427:
5426:
5424:
5421:
5420:
5417:
5416:
5413:
5412:
5409:
5408:
5406:
5405:
5404:
5403:
5397:
5388:
5386:
5380:
5379:
5376:
5375:
5372:
5371:
5369:
5368:
5361:
5354:
5347:
5340:
5333:
5326:
5319:
5316:Dromiceiomimus
5312:
5305:
5298:
5291:
5283:
5281:
5279:Ornithomimidae
5275:
5274:
5272:
5271:
5264:
5260:Paraxenisaurus
5256:
5248:
5241:
5234:
5226:
5224:
5222:Deinocheiridae
5218:
5217:
5215:
5214:
5211:Shenzhousaurus
5207:
5200:
5192:
5186:
5180:
5179:
5177:
5176:
5169:
5161:
5154:
5147:
5140:
5133:
5125:
5117:
5109:
5103:
5097:
5096:
5094:
5093:
5084:
5078:
5073:
5070:
5069:
5066:
5065:
5063:
5062:
5055:
5048:
5041:
5033:
5026:
5019:
5012:
5008:Beipiaognathus
5004:
4997:
4988:
4986:
4980:
4979:
4977:
4976:
4968:
4958:
4953:
4950:
4949:
4947:
4946:
4939:
4932:
4923:
4917:
4912:
4905:
4904:
4892:
4891:
4888:
4887:
4879:
4865:
4859:
4857:
4854:
4853:
4850:
4849:
4846:
4845:
4843:
4842:
4835:
4828:
4821:
4813:
4811:
4803:
4802:
4799:
4798:
4795:
4794:
4792:
4791:
4790:
4789:
4783:
4781:
4771:
4770:
4767:
4766:
4763:
4762:
4759:
4758:
4755:
4754:
4751:
4750:
4748:
4747:
4740:
4733:
4725:
4723:
4721:Tyrannosaurini
4717:
4716:
4714:
4713:
4706:
4699:
4691:
4683:
4681:
4679:Teratophoneini
4675:
4674:
4672:
4671:
4664:
4661:Daspletosaurus
4656:
4654:
4648:
4647:
4645:
4644:
4641:Qianzhousaurus
4637:
4629:
4627:
4621:
4620:
4618:
4617:
4610:
4602:
4596:
4590:
4589:
4587:
4586:
4579:
4571:
4569:
4560:
4555:
4552:
4551:
4549:
4548:
4541:
4538:Bistahieversor
4534:
4526:
4520:
4515:
4512:
4511:
4508:
4507:
4505:
4504:
4496:
4489:
4482:
4474:
4467:
4460:
4453:
4446:
4439:
4432:
4425:
4418:
4410:
4403:
4395:
4385:
4380:
4377:
4376:
4373:
4372:
4370:
4369:
4362:
4355:
4348:
4341:
4333:
4331:
4325:
4324:
4322:
4321:
4313:
4305:
4303:
4296:
4295:
4293:
4292:
4284:
4276:
4266:
4261:
4255:
4252:Tyrannoraptora
4250:
4247:
4246:
4244:
4243:
4236:
4229:
4222:
4214:
4207:
4199:
4191:
4184:
4176:
4169:
4161:
4155:
4150:
4143:
4142:
4130:
4129:
4126:
4125:
4117:
4098:
4092:
4090:
4087:
4086:
4083:
4082:
4079:
4078:
4076:
4075:
4074:
4073:
4067:
4065:
4059:
4058:
4055:
4054:
4052:
4051:
4044:
4037:
4030:
4023:
4016:
4009:
4001:
3999:
3997:Megaraptoridae
3993:
3992:
3990:
3989:
3981:
3973:
3966:
3959:
3952:
3944:
3936:
3928:
3922:
3911:
3910:
3907:
3906:
3903:
3902:
3899:
3898:
3895:
3894:
3891:
3890:
3887:
3886:
3884:
3883:
3876:
3869:
3862:
3859:Giganotosaurus
3854:
3852:
3846:
3845:
3843:
3842:
3834:
3828:
3822:
3821:
3819:
3818:
3811:
3804:
3797:
3790:
3783:
3776:
3769:
3762:
3754:
3746:
3740:
3734:
3733:
3731:
3730:
3722:
3714:
3707:
3699:
3690:
3688:
3686:Neovenatoridae
3682:
3681:
3679:
3678:
3671:
3668:Ulughbegsaurus
3664:
3656:
3649:
3642:
3634:
3628:
3622:
3621:
3619:
3618:
3610:
3602:
3594:
3586:
3584:
3575:
3570:
3567:
3566:
3563:
3562:
3559:
3558:
3556:
3555:
3548:
3541:
3534:
3527:
3519:
3517:
3511:
3510:
3508:
3507:
3500:
3496:Xuanhanosaurus
3491:
3485:
3479:
3478:
3476:
3475:
3467:
3463:Xuanhanosaurus
3459:
3452:
3449:Asfaltovenator
3444:
3434:
3429:
3426:
3425:
3423:
3422:
3414:
3410:Megalosauridae
3406:
3398:
3390:
3384:
3379:
3376:
3375:
3373:
3372:
3364:
3355:
3349:
3344:
3337:
3336:
3324:
3323:
3320:
3319:
3311:
3297:
3296:
3294:
3291:
3290:
3287:
3286:
3283:
3282:
3279:
3278:
3276:
3275:
3274:
3273:
3267:
3265:
3255:
3254:
3251:
3250:
3247:
3246:
3243:
3242:
3239:
3238:
3236:
3235:
3228:
3220:
3218:
3212:
3211:
3209:
3208:
3201:
3194:
3187:
3184:Ichthyovenator
3180:
3171:
3165:
3159:
3158:
3155:
3154:
3152:
3151:
3144:
3137:
3129:
3127:
3121:
3120:
3118:
3117:
3109:
3101:
3094:
3087:
3079:
3073:
3067:
3066:
3064:
3063:
3055:
3048:
3045:Cristatusaurus
3041:
3032:
3026:
3021:
3018:
3017:
3014:
3013:
3010:
3009:
3007:
3006:
3002:Poekilopleuron
2998:
2995:Piveteausaurus
2991:
2984:
2977:
2970:
2962:
2960:
2954:
2953:
2951:
2950:
2943:
2936:
2929:
2921:
2919:
2917:Megalosaurinae
2913:
2912:
2910:
2909:
2901:
2895:
2893:Megalosauridae
2889:
2888:
2886:
2885:
2876:
2866:
2861:
2858:
2857:
2854:
2853:
2851:
2850:
2846:Xuanhanosaurus
2842:
2835:
2828:
2820:
2818:
2812:
2811:
2809:
2808:
2805:Yunyangosaurus
2800:
2790:
2785:
2779:
2774:
2771:
2770:
2768:
2767:
2760:
2753:
2745:
2742:Pandoravenator
2738:
2731:
2728:Kayentavenator
2724:
2717:
2710:
2702:
2695:
2687:
2679:
2670:
2664:
2659:
2652:
2651:
2639:
2638:
2635:
2634:
2626:
2619:
2612:
2605:
2604:
2602:
2599:
2598:
2595:
2594:
2591:
2590:
2588:
2587:
2586:
2585:
2579:
2577:
2567:
2566:
2563:
2562:
2559:
2558:
2555:
2554:
2551:
2550:
2547:
2546:
2544:
2543:
2536:
2529:
2522:
2515:
2507:
2500:
2493:
2486:
2478:
2476:
2470:
2469:
2467:
2466:
2459:
2456:Skorpiovenator
2452:
2444:
2437:
2430:
2423:
2416:
2409:
2402:
2393:
2387:
2378:
2377:
2368:
2366:
2360:
2359:
2357:
2356:
2349:
2341:
2334:
2327:
2320:
2312:
2304:
2302:
2296:
2295:
2293:
2292:
2285:
2278:
2271:
2268:Spectrovenator
2264:
2257:
2250:
2243:
2236:
2229:
2221:
2213:
2207:
2202:
2199:
2198:
2195:
2194:
2191:
2190:
2188:
2187:
2180:
2173:
2166:
2159:
2151:
2149:
2143:
2142:
2140:
2139:
2132:
2125:
2117:
2115:
2109:
2108:
2106:
2105:
2102:Spinostropheus
2098:
2091:
2084:
2076:
2068:
2060:
2052:
2045:
2037:
2028:
2022:
2016:
2015:
2013:
2012:
2004:
1996:
1988:
1979:
1969:
1966:Abelisauroidea
1964:
1961:
1960:
1957:
1956:
1954:
1953:
1946:
1938:
1936:
1934:Ceratosauridae
1930:
1929:
1927:
1926:
1923:Saltriovenator
1919:
1911:
1904:
1900:Fosterovenator
1896:
1888:
1878:
1873:
1870:
1869:
1867:
1866:
1862:Lophostropheus
1858:
1850:
1844:
1839:
1832:
1831:
1819:
1818:
1815:
1814:
1801:
1800:
1798:
1795:
1794:
1791:
1790:
1787:
1786:
1784:
1783:
1776:
1768:
1766:
1762:
1761:
1759:
1758:
1757:
1756:
1750:
1748:
1742:
1741:
1738:
1737:
1735:
1734:
1727:
1723:Pterospondylus
1719:
1712:
1704:
1696:
1693:Lucianovenator
1689:
1682:
1674:
1672:
1666:
1665:
1663:
1662:
1655:
1648:
1641:
1634:
1625:
1619:
1617:Coelophysoidea
1613:
1612:
1610:
1609:
1602:
1594:
1587:
1580:
1573:
1566:
1559:
1556:Lophostropheus
1552:
1545:
1538:
1531:
1524:
1516:
1508:
1502:
1497:
1494:
1493:
1491:
1490:
1482:
1474:
1466:
1458:
1455:Erythrovenator
1451:
1443:
1435:
1426:
1420:
1415:
1408:
1407:
1395:
1394:
1391:
1390:
1387:
1386:
1383:
1382:
1380:
1379:
1378:
1377:
1371:
1369:
1363:
1362:
1360:
1359:
1358:
1357:
1348:
1342:
1332:
1331:
1327:
1326:
1317:
1308:
1302:
1296:
1280:
1279:
1272:
1271:
1264:
1257:
1249:
1243:
1242:
1227:Ricardoestesia
1222:
1211:Ricardoestesia
1206:
1192:
1191:External links
1189:
1188:
1187:
1177:(4): 751–763.
1159:
1153:
1138:
1135:
1132:
1131:
1116:
1105:(3): 393–398.
1085:
1066:(1): 179–189.
1046:
1026:
967:
925:
906:(3): 306–314.
890:
861:
850:(2): 343–353.
830:
819:(1): 208–215.
796:
778:
763:
748:
729:
706:
705:
703:
700:
676:
673:
591:
590:Classification
588:
507:
504:
487:Ricardoestesia
474:Ricardoestesia
451:Phillip Currie
447:oviraptorosaur
430:site. In 1924
399:
396:
362:dinosaur teeth
344:
343:
342:
341:
333:
321:
320:
314:
313:
312:
311:
297:
282:
281:
280:Other species
277:
276:
273:
261:
260:
254:
253:
239:
235:
234:
229:
222:
221:
216:
209:
208:
203:
196:
195:
190:
183:
182:
177:
173:
172:
167:
163:
162:
157:
153:
152:
139:
138:
130:
129:
121:
120:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
28:
15:
9:
6:
4:
3:
2:
5683:
5672:
5669:
5667:
5664:
5662:
5659:
5657:
5654:
5652:
5649:
5647:
5644:
5642:
5639:
5637:
5634:
5632:
5629:
5627:
5624:
5622:
5619:
5617:
5614:
5612:
5609:
5607:
5604:
5602:
5599:
5597:
5594:
5593:
5591:
5581:
5571:
5569:
5559:
5558:
5555:
5540:
5535:
5531:
5527:
5522:
5518:
5514:
5509:
5505:
5501:
5496:
5492:
5487:
5481:
5477:
5476:
5474:
5472:
5468:
5464:
5459:
5444:
5440:
5438:
5431:
5402:
5398:
5396:
5392:
5391:
5390:
5389:
5387:
5385:
5381:
5367:
5366:
5362:
5360:
5359:
5358:Struthiomimus
5355:
5353:
5352:
5348:
5346:
5345:
5341:
5339:
5338:
5334:
5332:
5331:
5327:
5325:
5324:
5320:
5318:
5317:
5313:
5311:
5310:
5309:"Coelosaurus"
5306:
5304:
5303:
5299:
5297:
5296:
5292:
5290:
5289:
5285:
5284:
5282:
5280:
5276:
5270:
5269:
5265:
5262:
5261:
5257:
5254:
5253:
5249:
5247:
5246:
5242:
5240:
5239:
5235:
5233:
5232:
5228:
5227:
5225:
5223:
5219:
5213:
5212:
5208:
5206:
5205:
5204:Pelecanimimus
5201:
5199:
5198:
5194:
5193:
5190:
5187:
5185:
5181:
5175:
5174:
5170:
5167:
5166:
5165:Thecocoelurus
5162:
5160:
5159:
5155:
5153:
5152:
5148:
5146:
5145:
5144:Kinnareemimus
5141:
5139:
5138:
5134:
5131:
5130:
5126:
5123:
5122:
5118:
5116:
5115:
5111:
5110:
5107:
5104:
5102:
5098:
5091:
5090:
5086:
5085:
5082:
5079:
5076:
5071:
5061:
5060:
5056:
5054:
5053:
5049:
5047:
5046:
5042:
5039:
5038:
5034:
5032:
5031:
5027:
5025:
5024:
5023:Huaxiagnathus
5020:
5018:
5017:
5016:Compsognathus
5013:
5010:
5009:
5005:
5003:
5002:
4998:
4995:
4994:
4990:
4989:
4987:
4985:
4981:
4975:
4974:
4970:
4969:
4966:
4962:
4959:
4956:
4951:
4945:
4944:
4943:Ornitholestes
4940:
4938:
4937:
4933:
4930:
4929:
4925:
4924:
4921:
4918:
4915:
4910:
4906:
4902:
4897:
4893:
4883:
4878:
4876:
4869:
4863:
4841:
4840:
4836:
4834:
4833:
4829:
4827:
4826:
4822:
4820:
4819:
4815:
4814:
4812:
4808:
4804:
4787:
4786:
4785:
4784:
4782:
4780:
4776:
4772:
4746:
4745:
4741:
4739:
4738:
4737:Tyrannosaurus
4734:
4732:
4731:
4727:
4726:
4724:
4722:
4718:
4712:
4711:
4710:Teratophoneus
4707:
4705:
4704:
4700:
4697:
4696:
4692:
4690:
4689:
4685:
4684:
4682:
4680:
4676:
4670:
4669:
4665:
4663:
4662:
4658:
4657:
4655:
4653:
4649:
4643:
4642:
4638:
4636:
4635:
4631:
4630:
4628:
4626:
4622:
4616:
4615:
4611:
4609:
4608:
4604:
4603:
4600:
4597:
4595:
4591:
4585:
4584:
4580:
4578:
4577:
4576:Albertosaurus
4573:
4572:
4570:
4568:
4564:
4561:
4558:
4553:
4547:
4546:
4542:
4540:
4539:
4535:
4533:
4532:
4528:
4527:
4524:
4521:
4518:
4513:
4502:
4501:
4497:
4495:
4494:
4493:Xiongguanlong
4490:
4488:
4487:
4483:
4480:
4479:
4475:
4473:
4472:
4471:Suskityrannus
4468:
4466:
4465:
4464:Stokesosaurus
4461:
4459:
4458:
4457:Santanaraptor
4454:
4452:
4451:
4447:
4445:
4444:
4440:
4438:
4437:
4433:
4431:
4430:
4426:
4424:
4423:
4419:
4416:
4415:
4411:
4409:
4408:
4404:
4402:
4401:
4400:Alectrosaurus
4397:
4396:
4393:
4389:
4386:
4383:
4378:
4368:
4367:
4363:
4361:
4360:
4356:
4354:
4353:
4349:
4347:
4346:
4342:
4340:
4339:
4335:
4334:
4332:
4330:
4326:
4319:
4318:
4317:Tanycolagreus
4314:
4312:
4311:
4307:
4306:
4304:
4301:
4297:
4290:
4289:
4285:
4283:
4282:
4278:
4277:
4274:
4270:
4267:
4264:
4259:
4256:
4253:
4248:
4242:
4241:
4237:
4235:
4234:
4230:
4228:
4227:
4223:
4220:
4219:
4215:
4213:
4212:
4208:
4205:
4204:
4200:
4197:
4196:
4192:
4190:
4189:
4185:
4182:
4181:
4180:Bahariasaurus
4177:
4175:
4174:
4170:
4168:
4167:
4163:
4162:
4159:
4156:
4153:
4152:Coelurosauria
4148:
4144:
4140:
4139:Coelurosauria
4135:
4131:
4121:
4116:
4114:
4109:
4102:
4096:
4071:
4070:
4069:
4068:
4066:
4064:
4063:Coelurosauria
4060:
4050:
4049:
4045:
4043:
4042:
4038:
4036:
4035:
4031:
4029:
4028:
4024:
4022:
4021:
4017:
4015:
4014:
4010:
4008:
4007:
4003:
4002:
4000:
3998:
3994:
3988:
3986:
3982:
3980:
3978:
3974:
3972:
3971:
3967:
3965:
3964:
3960:
3958:
3957:
3953:
3950:
3949:
3945:
3942:
3941:
3940:Bahariasaurus
3937:
3935:
3934:
3930:
3929:
3926:
3923:
3920:
3916:
3912:
3882:
3881:
3877:
3875:
3874:
3870:
3868:
3867:
3863:
3861:
3860:
3856:
3855:
3853:
3851:
3847:
3841:
3840:
3836:
3835:
3832:
3829:
3827:
3823:
3817:
3816:
3812:
3810:
3809:
3805:
3803:
3802:
3798:
3796:
3795:
3791:
3789:
3788:
3784:
3782:
3781:
3780:Kelmayisaurus
3777:
3775:
3774:
3770:
3768:
3767:
3763:
3760:
3759:
3755:
3753:
3752:
3748:
3747:
3744:
3741:
3739:
3735:
3728:
3727:
3723:
3720:
3719:
3715:
3713:
3712:
3708:
3705:
3704:
3700:
3697:
3696:
3692:
3691:
3689:
3687:
3683:
3677:
3676:
3672:
3670:
3669:
3665:
3662:
3661:
3657:
3655:
3654:
3650:
3648:
3647:
3643:
3641:
3640:
3636:
3635:
3632:
3629:
3627:
3623:
3616:
3615:
3614:Saurophaganax
3611:
3608:
3607:
3603:
3600:
3599:
3595:
3593:
3592:
3588:
3587:
3585:
3583:
3579:
3576:
3573:
3568:
3554:
3553:
3549:
3547:
3546:
3545:Siamotyrannus
3542:
3540:
3539:
3535:
3533:
3532:
3528:
3526:
3525:
3521:
3520:
3518:
3516:
3512:
3506:
3505:
3501:
3498:
3497:
3493:
3492:
3489:
3486:
3484:
3480:
3473:
3472:
3468:
3465:
3464:
3460:
3458:
3457:
3453:
3451:
3450:
3446:
3445:
3442:
3438:
3435:
3432:
3431:Allosauroidea
3427:
3420:
3419:
3418:Spinosauridae
3415:
3412:
3411:
3407:
3404:
3403:
3399:
3397:
3396:
3392:
3391:
3388:
3385:
3382:
3377:
3370:
3369:
3365:
3362:
3361:
3357:
3356:
3353:
3350:
3347:
3342:
3338:
3334:
3329:
3325:
3315:
3310:
3308:
3301:
3271:
3270:
3269:
3268:
3266:
3264:
3260:
3256:
3234:
3233:
3229:
3227:
3226:
3222:
3221:
3219:
3217:
3213:
3207:
3206:
3202:
3200:
3199:
3195:
3193:
3192:
3188:
3186:
3185:
3181:
3178:
3177:
3173:
3172:
3169:
3166:
3164:
3163:Spinosaurinae
3160:
3150:
3149:
3145:
3143:
3142:
3141:Riparovenator
3138:
3136:
3135:
3134:Ceratosuchops
3131:
3130:
3128:
3126:
3122:
3115:
3114:
3110:
3107:
3106:
3102:
3100:
3099:
3098:Riojavenatrix
3095:
3093:
3092:
3088:
3086:
3085:
3081:
3080:
3077:
3074:
3072:
3071:Baryonychinae
3068:
3061:
3060:
3056:
3054:
3053:
3049:
3047:
3046:
3042:
3039:
3038:
3034:
3033:
3030:
3027:
3024:
3023:Spinosauridae
3019:
3004:
3003:
2999:
2997:
2996:
2992:
2990:
2989:
2985:
2983:
2982:
2978:
2976:
2975:
2971:
2969:
2968:
2964:
2963:
2961:
2959:
2955:
2949:
2948:
2947:Wiehenvenator
2944:
2942:
2941:
2937:
2935:
2934:
2930:
2928:
2927:
2923:
2922:
2920:
2918:
2914:
2908:
2907:
2903:
2902:
2899:
2896:
2894:
2890:
2883:
2882:
2878:
2877:
2874:
2870:
2867:
2864:
2859:
2848:
2847:
2843:
2841:
2840:
2836:
2834:
2833:
2829:
2827:
2826:
2822:
2821:
2819:
2817:
2813:
2807:
2806:
2802:
2801:
2798:
2794:
2791:
2788:
2783:
2780:
2777:
2772:
2766:
2765:
2761:
2759:
2758:
2754:
2751:
2750:
2746:
2744:
2743:
2739:
2737:
2736:
2732:
2730:
2729:
2725:
2723:
2722:
2718:
2716:
2715:
2711:
2708:
2707:
2703:
2701:
2700:
2696:
2693:
2692:
2688:
2685:
2684:
2683:Chienkosaurus
2680:
2677:
2676:
2672:
2671:
2668:
2665:
2662:
2657:
2653:
2649:
2644:
2640:
2630:
2623:
2616:
2609:
2583:
2582:
2581:
2580:
2578:
2576:
2572:
2568:
2542:
2541:
2537:
2535:
2534:
2530:
2528:
2527:
2523:
2521:
2520:
2516:
2513:
2512:
2508:
2506:
2505:
2501:
2499:
2498:
2494:
2492:
2491:
2487:
2485:
2484:
2480:
2479:
2477:
2475:
2474:Furileusauria
2471:
2465:
2464:
2460:
2458:
2457:
2453:
2450:
2449:
2448:Rahiolisaurus
2445:
2443:
2442:
2438:
2436:
2435:
2431:
2429:
2428:
2424:
2422:
2421:
2417:
2415:
2414:
2410:
2408:
2407:
2403:
2400:
2399:
2395:
2394:
2391:
2388:
2386:
2382:
2375:
2374:
2370:
2369:
2367:
2365:
2364:Carnotaurinae
2361:
2355:
2354:
2350:
2347:
2346:
2345:Rahiolisaurus
2342:
2340:
2339:
2338:Majungasaurus
2335:
2333:
2332:
2328:
2326:
2325:
2321:
2318:
2317:
2313:
2311:
2310:
2306:
2305:
2303:
2301:
2297:
2291:
2290:
2286:
2284:
2283:
2279:
2277:
2276:
2275:Tarascosaurus
2272:
2270:
2269:
2265:
2263:
2262:
2258:
2256:
2255:
2251:
2249:
2248:
2244:
2242:
2241:
2237:
2235:
2234:
2233:Eoabelisaurus
2230:
2227:
2226:
2222:
2220:
2219:
2218:Chenanisaurus
2215:
2214:
2211:
2208:
2205:
2204:Abelisauridae
2200:
2186:
2185:
2181:
2179:
2178:
2174:
2172:
2171:
2167:
2165:
2164:
2163:Masiakasaurus
2160:
2158:
2157:
2153:
2152:
2150:
2148:
2144:
2138:
2137:
2133:
2131:
2130:
2129:Huinculsaurus
2126:
2124:
2123:
2122:Elaphrosaurus
2119:
2118:
2116:
2114:
2110:
2104:
2103:
2099:
2097:
2096:
2092:
2090:
2089:
2085:
2082:
2081:
2077:
2074:
2073:
2069:
2066:
2065:
2061:
2058:
2057:
2053:
2051:
2050:
2046:
2043:
2042:
2041:Austrocheirus
2038:
2035:
2034:
2030:
2029:
2026:
2023:
2021:
2017:
2010:
2009:
2005:
2002:
2001:
1997:
1994:
1993:
1989:
1986:
1985:
1981:
1980:
1977:
1973:
1970:
1967:
1962:
1952:
1951:
1947:
1945:
1944:
1940:
1939:
1937:
1935:
1931:
1925:
1924:
1920:
1917:
1916:
1912:
1910:
1909:
1905:
1902:
1901:
1897:
1895:
1894:
1893:Berberosaurus
1890:
1889:
1886:
1882:
1879:
1876:
1871:
1864:
1863:
1859:
1857:
1856:
1852:
1851:
1848:
1845:
1842:
1837:
1833:
1829:
1824:
1820:
1811:
1805:
1782:
1781:
1780:Halticosaurus
1777:
1775:
1774:
1773:Teinurosaurus
1770:
1769:
1767:
1763:
1754:
1753:
1752:
1751:
1749:
1747:
1743:
1733:
1732:
1728:
1725:
1724:
1720:
1718:
1717:
1713:
1710:
1709:
1705:
1702:
1701:
1700:Megapnosaurus
1697:
1695:
1694:
1690:
1688:
1687:
1683:
1681:
1680:
1676:
1675:
1673:
1671:
1670:Coelophysidae
1667:
1661:
1660:
1659:Powellvenator
1656:
1654:
1653:
1649:
1647:
1646:
1642:
1640:
1639:
1635:
1632:
1631:
1627:
1626:
1623:
1620:
1618:
1614:
1608:
1607:
1603:
1600:
1599:
1595:
1593:
1592:
1588:
1586:
1585:
1581:
1579:
1578:
1574:
1572:
1571:
1567:
1565:
1564:
1560:
1558:
1557:
1553:
1551:
1550:
1549:Liliensternus
1546:
1544:
1543:
1539:
1537:
1536:
1532:
1530:
1529:
1525:
1522:
1521:
1520:Dolichosuchus
1517:
1515:
1514:
1513:Dilophosaurus
1510:
1509:
1506:
1503:
1500:
1495:
1488:
1487:
1486:Herrerasauria
1483:
1480:
1479:
1475:
1472:
1471:
1467:
1464:
1463:
1459:
1457:
1456:
1452:
1449:
1448:
1444:
1441:
1440:
1439:Daemonosaurus
1436:
1433:
1432:
1428:
1427:
1424:
1421:
1418:
1413:
1409:
1405:
1400:
1396:
1375:
1374:
1373:
1372:
1370:
1368:
1364:
1356:
1352:
1351:
1350:
1349:
1346:
1343:
1341:
1337:
1333:
1325:
1321:
1318:
1316:
1312:
1309:
1307:
1303:
1301:
1297:
1295:
1291:
1290:
1285:
1281:
1277:
1270:
1265:
1263:
1258:
1256:
1251:
1250:
1247:
1241:
1237:
1234:
1232:
1228:
1223:
1221:
1217:
1214:
1212:
1207:
1204:
1199:
1195:
1194:
1184:
1180:
1176:
1172:
1165:
1160:
1156:
1150:
1146:
1141:
1140:
1127:
1120:
1112:
1108:
1104:
1100:
1096:
1089:
1081:
1077:
1073:
1069:
1065:
1061:
1057:
1050:
1042:
1041:
1036:
1030:
1022:
1018:
1013:
1008:
1003:
998:
994:
990:
987:(1): e54329.
986:
982:
978:
971:
963:
959:
955:
951:
947:
943:
939:
932:
930:
921:
917:
913:
909:
905:
901:
894:
881:
877:
868:
866:
857:
853:
849:
845:
841:
834:
826:
822:
818:
814:
807:
800:
792:
788:
782:
774:
767:
759:
752:
744:
740:
733:
725:
721:
714:
712:
707:
699:
697:
690:
686:
683:Hypothetical
681:
672:
670:
666:
661:
659:
655:
651:
647:
643:
639:
634:
632:
628:
624:
620:
616:
612:
608:
604:
596:
587:
585:
581:
577:
573:
569:
565:
564:
560:
555:
553:
552:sebecosuchian
549:
548:crocodyliform
545:
541:
537:
533:
528:
525:
521:
517:
513:
503:
501:
500:specific name
497:
492:
488:
484:
479:
475:
471:
470:Richard Estes
466:
464:
460:
456:
452:
448:
444:
443:Chirostenotes
440:
438:
437:Chirostenotes
433:
429:
425:
421:
417:
409:
406:Tooth of cf.
404:
395:
393:
392:
387:
383:
379:
375:
374:United States
371:
367:
363:
360:
356:
352:
351:
337:
334:
328:
325:
324:
322:
319:
315:
306:
305:
298:
292:
286:
285:
283:
278:
271:
270:
262:
259:
255:
248:
247:
240:
237:
236:
233:
232:Coelurosauria
230:
227:
224:
223:
220:
217:
214:
211:
210:
207:
204:
201:
198:
197:
194:
191:
188:
185:
184:
181:
178:
175:
174:
171:
168:
165:
164:
161:
158:
155:
154:
149:
144:
140:
136:
133:Tooth of cf.
131:
127:
122:
116:
112:
108:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
5596:Coelurosaurs
5580:Paleontology
5470:
5434:
5363:
5356:
5349:
5342:
5335:
5330:Ornithomimus
5328:
5321:
5314:
5307:
5300:
5293:
5286:
5268:Tyrannomimus
5266:
5258:
5250:
5243:
5238:Deinocheirus
5236:
5229:
5209:
5202:
5195:
5171:
5163:
5158:Nqwebasaurus
5156:
5151:Nedcolbertia
5149:
5142:
5135:
5129:Calamosaurus
5127:
5121:Aviatyrannis
5119:
5112:
5087:
5057:
5050:
5043:
5035:
5028:
5021:
5014:
5006:
5001:Aristosuchus
4999:
4993:Aniksosaurus
4991:
4973:Aniksosaurus
4971:
4941:
4934:
4926:
4872:
4837:
4830:
4823:
4816:
4742:
4735:
4728:
4708:
4701:
4693:
4688:Dynamoterror
4686:
4666:
4659:
4639:
4632:
4612:
4607:Asiatyrannus
4605:
4581:
4574:
4545:Dryptosaurus
4543:
4536:
4529:
4498:
4491:
4484:
4476:
4469:
4462:
4455:
4448:
4441:
4436:Jinbeisaurus
4434:
4427:
4420:
4412:
4407:Aviatyrannis
4405:
4398:
4364:
4359:Sinotyrannus
4357:
4350:
4343:
4336:
4315:
4308:
4286:
4279:
4238:
4231:
4224:
4216:
4210:
4209:
4201:
4193:
4188:Bicentenaria
4186:
4178:
4173:Asiamericana
4171:
4164:
4105:
4046:
4039:
4032:
4025:
4018:
4011:
4004:
3983:
3975:
3968:
3961:
3954:
3948:Deltadromeus
3946:
3938:
3931:
3880:Tyrannotitan
3878:
3871:
3864:
3857:
3837:
3813:
3808:Taurovenator
3806:
3799:
3792:
3787:Lajasvenator
3785:
3778:
3771:
3766:Concavenator
3764:
3756:
3749:
3724:
3716:
3709:
3701:
3693:
3673:
3666:
3658:
3651:
3644:
3637:
3612:
3604:
3596:
3589:
3582:Allosauridae
3550:
3543:
3538:Shidaisaurus
3536:
3529:
3522:
3502:
3494:
3469:
3461:
3454:
3447:
3416:
3408:
3400:
3393:
3366:
3358:
3346:Avetheropoda
3333:Avetheropoda
3304:
3263:Avetheropoda
3230:
3223:
3216:Spinosaurini
3203:
3196:
3189:
3182:
3174:
3146:
3139:
3132:
3111:
3103:
3096:
3091:Protathlitis
3089:
3082:
3057:
3050:
3043:
3035:
3000:
2993:
2986:
2981:Leshansaurus
2979:
2972:
2965:
2945:
2938:
2933:Megalosaurus
2931:
2926:Duriavenator
2924:
2904:
2879:
2863:Megalosauria
2844:
2837:
2832:Marshosaurus
2830:
2825:Condorraptor
2823:
2803:
2762:
2755:
2747:
2740:
2733:
2726:
2719:
2712:
2706:Cruxicheiros
2704:
2697:
2689:
2681:
2673:
2538:
2533:Quilmesaurus
2531:
2524:
2517:
2509:
2502:
2495:
2488:
2481:
2461:
2454:
2446:
2439:
2432:
2425:
2418:
2411:
2404:
2396:
2385:Brachyrostra
2371:
2351:
2343:
2336:
2329:
2322:
2314:
2307:
2287:
2282:Tralkasaurus
2280:
2273:
2266:
2259:
2252:
2245:
2238:
2231:
2223:
2216:
2184:Vespersaurus
2182:
2177:Velocisaurus
2175:
2168:
2161:
2154:
2134:
2127:
2120:
2100:
2093:
2086:
2078:
2072:Deltadromeus
2070:
2062:
2056:Compsosuchus
2054:
2047:
2039:
2031:
2006:
1998:
1992:Lametasaurus
1990:
1982:
1948:
1943:Ceratosaurus
1941:
1921:
1913:
1906:
1898:
1891:
1875:Ceratosauria
1860:
1853:
1778:
1771:
1729:
1721:
1714:
1706:
1698:
1691:
1684:
1677:
1657:
1652:Podokesaurus
1650:
1643:
1636:
1628:
1604:
1596:
1589:
1582:
1575:
1568:
1561:
1554:
1547:
1540:
1535:Gojirasaurus
1533:
1528:Dracovenator
1526:
1518:
1511:
1499:Neotheropoda
1484:
1476:
1468:
1462:Guaibasaurus
1460:
1453:
1445:
1437:
1431:Chindesaurus
1429:
1319:
1310:
1230:
1226:
1210:
1174:
1170:
1144:
1125:
1119:
1102:
1098:
1088:
1063:
1059:
1049:
1038:
1029:
984:
980:
970:
945:
941:
937:
903:
899:
893:
879:
847:
843:
839:
833:
816:
812:
799:
791:the original
781:
772:
766:
757:
751:
742:
738:
732:
723:
695:
693:
675:Paleobiology
664:
662:
654:R. isosceles
653:
646:R. isosceles
645:
641:
635:
602:
601:
583:
579:
575:
561:
556:
544:R. isosceles
543:
535:
532:Julia Sankey
529:
515:
514:specimen of
509:
495:
486:
482:
477:
473:
467:
462:
442:
435:
413:
408:R. isosceles
407:
389:
386:R. isosceles
385:
381:
349:
348:
347:
335:
326:
302:
295:Sankey, 2001
291:R. isosceles
290:
268:
267:
258:Type species
245:
244:
225:
212:
199:
186:
134:
24:
18:
5395:Maniraptora
5384:Maniraptora
5365:Tototlmimus
5295:Anserimimus
5245:Garudimimus
5231:Beishanlong
5173:Valdoraptor
5114:Arkansaurus
5059:Xunmenglong
4936:Juravenator
4730:Tarbosaurus
4614:Nanuqsaurus
4583:Gorgosaurus
4500:Megaraptora
4486:Timurlengia
4218:Sciurumimus
4166:Aratasaurus
4034:Murusraptor
3956:Fukuiraptor
3919:Megaraptora
3801:Shaochilong
3773:Eocarcharia
3726:Megaraptora
3646:Lusovenator
3524:Alpkarakush
3381:Carnosauria
3232:Spinosaurus
3205:Siamosaurus
3105:Suchosaurus
3052:Iberospinus
2988:Magnosaurus
2967:Afrovenator
2940:Torvosaurus
2691:Chilesaurus
2504:Carnotaurus
2497:Caletodraco
2483:Abelisaurus
2309:Arcovenator
2156:Laevisuchus
2147:Noasaurinae
2049:Berthasaura
2020:Noasauridae
1950:Genyodectes
1915:Lukousaurus
1908:Jubbulpuria
1708:Panguraptor
1686:Coelophysis
1679:Camposaurus
1638:Panguraptor
1630:Dracoraptor
1606:Zupaysaurus
1591:Tachiraptor
1570:Sarcosaurus
1470:Nhandumirim
1040:Fossilworks
745:(43): 1–12.
689:dromaeosaur
663:Fossils of
642:R. gilmorei
584:R. asiatica
559:coelurosaur
439:pergracilis
391:R. asiatica
382:R. gilmorei
355:morphogenus
331:Nesov, 1995
304:R. asiatica
135:R. gilmorei
117:, but see .
36:90–66
5590:Categories
5323:Gallimimus
5252:Harpymimus
5197:Harpymimus
4818:Iliosuchus
4788:see below↓
4625:Alioramini
4443:Juratyrant
4429:Eotyrannus
4414:Bagaraatan
4366:Yutyrannus
4300:Coeluridae
4226:Vayuraptor
4072:see below↓
4048:Tratayenia
4041:Orkoraptor
4027:Megaraptor
3933:Aoniraptor
3866:Mapusaurus
3794:Sauroniops
3758:Altispinax
3711:Neovenator
3653:Siamraptor
3639:Datanglong
3606:Epanterias
3598:Antrodemus
3591:Allosaurus
3572:Allosauria
3395:Altispinax
3360:Gasosaurus
3272:see below↓
3148:Suchomimus
2749:Sinosaurus
2584:see below↓
2540:Viavenator
2490:Aucasaurus
2427:Ilokelesia
2398:Dahalokely
2353:Rajasaurus
2331:Indosaurus
2324:Genusaurus
2316:Dahalokely
2240:Indosuchus
2136:Limusaurus
2095:Ligabueino
2088:Kiyacursor
2080:Genusaurus
2064:Dahalokely
1984:Betasuchus
1855:Dornraptor
1755:see below↓
1731:Segisaurus
1584:Sinosaurus
1447:Eodromaeus
1376:see below↓
1324:Dinosauria
1306:Sauropsida
1154:3929907380
702:References
638:form taxon
621:, and the
572:Kazakhstan
378:Kazakhstan
206:Saurischia
193:Dinosauria
5568:Dinosaurs
5399:includes
5344:Rativates
5337:Qiupalong
5037:Scipionyx
5030:Mirischia
4703:Lythronax
4695:Labocania
4634:Alioramus
4288:Labocania
4006:Aerosteon
3552:Sinraptor
3456:Erectopus
3191:Irritator
2776:Orionides
2661:Tetanurae
2648:Tetanurae
2575:Tetanurae
2511:Elemgasem
2434:Llukalkan
2413:Elemgasem
2170:Noasaurus
2033:Afromimus
1841:Averostra
1828:Averostra
1746:Averostra
1598:Velocipes
1417:Theropoda
1404:Theropoda
1367:Theropoda
1292:Kingdom:
1276:Theropoda
1035:"Blasi 2"
840:Doratodon
627:Barremian
530:In 2001,
524:Campanian
219:Theropoda
166:Kingdom:
160:Eukaryota
111:Barremian
105:Possible
5486:Q2029022
5480:Wikidata
4928:Coelurus
4345:Kileskus
4338:Guanlong
4310:Coelurus
4203:Gualicho
3703:Gualicho
3084:Baryonyx
2420:Guemesia
2247:Kryptops
2008:Ozraptor
1645:Pendraig
1300:Chordata
1298:Phylum:
1294:Animalia
1236:Archived
1216:Archived
1021:23372708
981:PLOS ONE
512:holotype
478:Ricardus
359:theropod
318:Synonyms
180:Chordata
176:Phylum:
170:Animalia
156:Domain:
5554:Portals
5526:4946100
5513:4822949
5500:4531529
4478:Timimus
4240:Zuolong
3970:Rapator
3873:Meraxes
3198:Oxalaia
2519:Koleken
1542:Lepidus
1304:Class:
1229:versus
1068:Bibcode
1012:3553132
989:Bibcode
950:Bibcode
908:Bibcode
506:Species
445:was an
426:at the
424:Alberta
398:History
238:Genus:
5137:Hexing
4825:Kakuru
4422:Dilong
2463:Thanos
2441:Niebla
2261:Rugops
2254:Kurupi
1233:debate
1151:
1019:
1009:
882:]
609:, the
498:. The
434:named
372:, the
370:Canada
5539:58873
5401:birds
4450:Moros
3977:Siats
3718:Siats
3660:Siats
1320:Clade
1311:Clade
1209:The "
1167:(PDF)
884:(PDF)
878:[
809:(PDF)
687:as a
629:age (
353:is a
226:Clade
213:Clade
200:Clade
187:Clade
5508:GBIF
5393:see
4020:Maip
1478:Tawa
1353:see
1149:ISBN
1017:PMID
644:and
510:The
491:ICZN
457:and
384:and
376:and
44:PreꞒ
5495:EoL
1179:doi
1107:doi
1076:doi
1007:PMC
997:doi
958:doi
916:doi
852:doi
821:doi
570:in
465:.
422:in
357:of
5592::
5536::
5523::
5510::
5497::
5482::
1322::
1313::
1175:76
1173:.
1169:.
1103:63
1101:.
1097:.
1074:.
1064:72
1062:.
1058:.
1037:.
1015:.
1005:.
995:.
983:.
979:.
956:.
946:39
944:.
928:^
914:.
904:50
902:.
864:^
848:25
846:.
817:75
815:.
811:.
743:38
741:.
710:^
617:,
613:,
554:.
476:,
453:,
329:?
307:?
228::
215::
202::
189::
94:Pg
38:Ma
33:,
5556::
5263:?
5255:?
5168:?
5132:?
5124:?
5092:?
5040:?
5011:?
4996:?
4931:?
4698:?
4503:?
4481:?
4417:?
4320:?
4302:?
4291:?
4221:?
4206:?
4198:?
4183:?
3987:?
3979:?
3951:?
3943:?
3921:?
3761:?
3729:?
3721:?
3706:?
3698:?
3663:?
3617:?
3609:?
3601:?
3499:?
3474:?
3466:?
3421:?
3413:?
3405:?
3371:?
3363:?
3179:?
3116:?
3108:?
3062:?
3040:?
3005:?
2884:?
2849:?
2752:?
2709:?
2694:?
2686:?
2678:?
2514:?
2451:?
2401:?
2376:?
2348:?
2319:?
2228:?
2083:?
2075:?
2067:?
2059:?
2044:?
2036:?
2011:?
2003:?
1995:?
1987:?
1918:?
1903:?
1865:?
1726:?
1711:?
1703:?
1633:?
1601:?
1523:?
1489:?
1481:?
1473:?
1465:?
1450:?
1442:?
1434:?
1268:e
1261:t
1254:v
1185:.
1181::
1157:.
1113:.
1109::
1082:.
1078::
1070::
1043:.
1023:.
999::
991::
985:8
964:.
960::
952::
922:.
918::
910::
858:.
854::
827:.
823::
300:†
288:†
265:†
242:†
109:(
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:Ꞓ
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