1466:
426:
709:
7778:
6545:
525:
7952:
5347:
161:
744:
7964:
1683:
783:
1595:. The model successfully flew with a combination of soaring and wing flapping. The model was based on a then-current weight estimate of around 80 kg (180 lb), far lower than more modern estimates of over 200 kg (440 lb). The method of flight in these pterosaurs depends largely on their weight, which has been controversial, and widely differing masses have been favored by different scientists. Some researchers have suggested that these animals employed slow,
5353:
961:
4730:
6561:
7785:
4735:
133:
4189:
1612:
7791:
1564:
4197:
1326:
4204:
6553:
685:
1640:. Henderson's work was also further criticized by Witton and Habib in another study, which pointed out that, although Henderson used excellent mass estimations, they were based on outdated pterosaur models, which caused Henderson's mass estimations to be more than double what Habib used in his estimations and that anatomical study of
1520:, catching fish during flight while cleaving the waves with its beak. While this skim-feeding view became widely accepted, it was not subjected to scientific research until 2007 when a study showed that, for such large pterosaurs, it was not a viable method because the energy costs would be too high due to excessive
1673:
had limited walking motion in its wings, with the limbs on each side of the body being moved together and the forelimbs being lifted out of the way of the hindlimbs. This study also suggests that the wings of pterosaurs were only attached to the body, with the legs and feet being tucked underneath,
1507:
In 1996, Lehman and
Langston rejected the scavenging hypothesis, pointing out that the lower jaw bent so strongly downwards that even when it closed completely a gap of over 5 cm (2.0 in) remained between it and the upper jaw, very different from the hooked beaks of specialized scavenging
1668:
was a very powerful flier, it had a large breastbone to support the necessary muscles for creating the flight stroke, and that it would probably have used its powerful hind legs to launch as high as 8 ft (2.4 m) when taking off, allowing it to gain enough height and momentum to begin the
1528:
published an examination of possible feeding habits and ecology of azhdarchids. Witton and Naish noted that most azhdarchid remains are found in inland deposits far from seas or other large bodies of water required for skimming. Additionally, the beak, jaw, and neck anatomy are unlike those of any
720:
More recent estimates based on greater knowledge of azhdarchid proportions place its wingspan at 10–11 m (33–36 ft). Remains found in Texas in 1971 indicate that this pterosaur had a minimum wingspan of about 11 m (36 ft). Generalized height in a bipedal stance, based on its
704:
fossils came from an individual with a wingspan as large as 15.9 m (52 ft). Choosing the middle of three extrapolations from the proportions of other pterosaurs gave an estimate of 11 m, 15.5 m, and 21 m, respectively (36 ft, 50.85 ft, 68.9 ft). In 1981, further advanced
463:
at the
University of Texas, Austin. He would then go on to become a doctoral student at the University of California, Berkeley. The specimen he discovered consisted of a partial wing (in pterosaurs composed of the forearms and elongated fourth finger) from an individual later estimated at over
724:
Body mass estimates for giant azhdarchids are extremely problematic because no existing species shares a similar size or body plan, and in consequence, published results vary widely. Generalized weight, based on some studies that have historically found extremely low weight estimates for
767:
had a very sharp and pointed beak. That is contrary to some earlier reconstructions that showed a blunter snout, based on the inadvertent inclusion of jaw material from another pterosaur species, as this material was named as the holotype of a genus of short-snouted azhdarchid,
536:
At first, it was assumed that the smaller specimens were juvenile or subadult forms of the larger type. Later, when more remains were found, it was realized they could have been a separate species. This possible second species from Texas was provisionally referred to as a
1533:, and that they probably hunted small vertebrates on land or in small streams. Additionally, Quetzalcoatlus had the ability to turn its head 180 degrees which would allow it to be more adept at finding prey and preventing ambushes from foes. Though
1631:, and Mark Witton, a British paleontologist, undertook further investigation into the claims of flightlessness in large pterosaurs. After factoring wingspan, body weight, and aerodynamics, computer modeling led the two researchers to conclude that
1761:'s 350th-anniversary exhibition. The models, which included both flying and standing individuals with wingspans of over 10 m (33 ft), were intended to help build public interest in science. The models were created by scientists from the
1644:
and other big pterosaur forelimbs showed a higher degree of robustness than would be expected if they were purely quadrupedal. This study proposed that large pterosaurs most likely utilized a short burst of powered flight to then transition to
1450:
had precursors in North
America and its apparent rise to widespreadness may represent the expansion of its preferred habitat rather than an immigration event, as some experts have suggested. It co-existed with another azhdarchid known as
1481:. Because the area of the fossil site was 400 km (250 mi) removed from the coastline and there were no indications of large rivers or deep lakes nearby at the end of the Cretaceous, Lawson, in 1975, rejected a
1741:
incorporates a "flight control system/autopilot which processes pilot commands and sensor inputs, implements several feedback loops, and delivers command signals to its various servo-actuators". It is on exhibit at the
1635:
was capable of flight up to 130 km/h (80 mph) for 7 to 10 days at altitudes of 4,600 m (15,000 ft). Habib further suggested a maximum flight range of 13,000–19,000 km (8,000–12,000 mi) for
1603:
had been underestimated, even the highest estimates, and that it was too massive to have achieved powered flight. He estimated it in his 2010 paper as 540 kg (1,190 lb). Henderson argued that it may have been
729:, was as low as 70 kg (150 lb) for a 10 m (32 ft 10 in) individual. A majority of estimates published since the 2000s have been substantially higher, around 200–250 kg (440–550 lb).
1626:
Other flight capability estimates have disagreed with
Henderson's research, suggesting instead an animal superbly adapted to long-range, extended flight. In 2010, Mike Habib, a professor of biomechanics at
665:, the vertebra shows no evidence that it was chewed on by the dinosaur. The bone came from an individual azhdarchid pterosaur estimated to have had a wingspan of 5–5.5 m (16–18 ft).
572:—represent elements which are typically considered undiagnostic to generic or specific level and that this complicates interpretations of azhdarchid taxonomy. For instance, Witton
3005:
1710:. Over the next twenty-five years prior to future discoveries, it would launch similar depictions colloquially known as "paleomemes" in various books, as noted by Darren Naish.
1729:(UAV). They produced a half-scale model weighing 18 kg (40 lb), with a wingspan of 5.5 m (18 ft). Coincidentally, Douglas A. Lawson, who discovered
1465:
620:
differ enough that they cannot be regarded as the same animal, but that the significance of this cannot be ascertained, given uncertainty over the relationships of
3024:
636:, could have made long, transcontinental flights, suggesting that locations as disparate as North America and Europe could have shared giant azhdarchid species.
580:
type material is of generalized enough morphology to be near identical to that of other giant azhdarchids, such as the overlapping elements of the contemporary
2781:
Lawson 1975 (Pterodactyloidea: Azhdarchoidea): A paleoenvironmental reconstruction of the
Javelina Formation (Upper Cretaceous), Big Bend National Park, Texas"
549:
holotype, and include four partial skulls, though they are much less massive, with an estimated wingspan of 5.5 m (18 ft). This species was named
7989:
545:
and
Langston in 1996, indicating that its status was too uncertain to give it a full new species name. The smaller specimens are more complete than the
1504:. Lawson had found the remains of the giant pterosaur while searching for the bones of this dinosaur, which formed an important part of its ecosystem.
3211:
7896:
1579:
and other giant azhdarchids was poorly understood until serious biomechanical studies were conducted in the 21st century. A 1984 experiment by
736:
would have weighed 20 kg (44 lb) for individuals with a wingspan of 4.5 m (15 ft). In 2022, Gregory S. Paul estimated that
471:
Lawson discovered a second site of the same age, about 40 km (25 mi) from the first, where between 1972 and 1974, he and
Professor
1910:
425:
413:, which was known for many years as an unnamed species, before being named by Brian Andres and Wann Langston Jr. (posthumously) in 2021.
2387:"On the size and flight diversity of giant pterosaurs, the use of birds as pterosaur analogues and comments on pterosaur flightlessness"
2651:: paleoenvironmental reconstruction of the Javelina Formation (Upper Cretaceous), Big Bend National Park, Texas". Abstracts of Papers.
2006:
Lawson, Douglas A. (1975). "Pterosaur from the Latest
Cretaceous of West Texas: Discovery of the largest flying creature". Reports.
8004:
7999:
7870:
17:
8009:
2908:
655:. The specimen, BMR P2002.2, was recovered accidentally when it was included in a field jacket prepared to transport part of a
3112:
1599:
flight, while others have concluded that their flight was fast and dynamic. In 2010, Donald
Henderson argued that the mass of
3190:
2337:
740:
had a wingspan of 5 m (16 ft), body length of 3.5 m (11 ft), and body mass of 65 kg (143 lb).
3054:
2598:"Late Maastrichtian pterosaurs from North Africa and mass extinction of Pterosauria at the Cretaceous-Paleogene boundary"
1529:
known skimming animal. Rather, they concluded that azhdarchids were more likely terrestrial stalkers, similar to modern
1457:, as well as an additional pterosaur taxon, suggesting a relatively high diversity of Late Cretaceous pterosaur genera.
3032:
479:
unearthed three fragmentary skeletons of much smaller individuals. Lawson in 1975 announced the find in an article in
2517:
2366:
1649:. However, a study from 2022 suggests that they would only have flown occasionally and for short distances, like the
148:
7777:
1870:
Lehman, Thomas M.; Mcdowell, Fred W.; Connelly, James N. (2006). "First isotopic (U-Pb) age for the Late
Cretaceous
632:
being investigated. An additional complication to these discussions is the likelihood that huge pterosaurs, such as
485:. That same year, in a subsequent letter to the same journal, he made the original large specimen, TMM 41450-3, the
2986:
2938:"How did extinct giant birds and pterosaurs fly? A comprehensive modeling approach to evaluate soaring performance"
2240:"An azhdarchid pterosaur cervical vertebra from the Hell Creek Formation (Maastrichtian) of southeastern Montana"
1779:
5352:
2258:
1887:
1743:
460:
2680:"Did pterosaurs feed by skimming? Physical modelling and anatomical evaluation of an unusual feeding method"
1493:(which will scavenge, but is more of a terrestrial predator of small animals), but then on the carcasses of
7994:
1545:
mammals than to their smaller cousins, implying that they were uniquely suited to a terrestrial lifestyle.
1303:
1114:
160:
6544:
5346:
1279:
7942:
2581:
Lehman, T. M. (2001). "Late Cretaceous dinosaur provinciality". In Tanke, D. H.; Carpenter, K. (eds.).
4729:
3183:
568:
and colleagues (2010) noted that the type species of the genus—the fragmentary wing bones comprising
2329:
2323:
807:, a group of advanced toothless pterosaurs with unusually long, stiffened necks and giant wingspans
7848:
3287:
1774:
1762:
1555:
is found in great numbers in alkaline lakes, and likely lived like modern gregarious wading birds.
955:
901:
4188:
3084:
4765:
4196:
1726:
1206:
1090:
1010:
560:
had not been properly described and diagnosed until 2021 and until then, the status of the genus
502:
4734:
7927:
7810:
7685:
5843:
1138:
452:
7901:
7784:
2115:(Pterosauria, Azhdarchidae) from Late Cretaceous sediments of Big Bend National Park, Texas".
1874:
vertebrate fauna of West Texas, and its significance as a link between two faunal provinces".
708:
8014:
7922:
7914:
7790:
1235:
860:
476:
3006:"Quetzalcoatlus: the evil, pin-headed, toothy nightmare monster that wants to eat your soul"
2829:
1653:(the world's heaviest bird that actively flies) and that they were not able to soar at all.
7984:
7968:
7857:
6552:
5938:
4377:
4287:
4203:
3176:
2873:
2792:
2741:
2464:
2398:
2289:
2168:
2124:
2066:
2015:
1957:
1826:
1619:
648:
510:
6560:
8:
7956:
7442:
7131:
6818:
6117:
6044:
3812:
3532:
2159:
1541:
and other azhdarchids have fore and hind limb proportions more similar to modern running
977:
884:
774:, in 2021. A skull crest was also present but its exact form and size are still unknown.
2877:
2796:
2745:
2468:
2402:
2293:
2217:"Clipping the wings of giant pterosaurs: comments on wingspan estimations and diversity"
2172:
2128:
2070:
2019:
1961:
1830:
700:
When it was first named as a new species in 1975, scientists estimated that the largest
6971:
5445:
5288:
4431:
4234:
4221:
4135:
3977:
3608:
3583:
2962:
2937:
2889:
2810:
2757:
2706:
2679:
2624:
2597:
2558:
2523:
2482:
2421:
2386:
2355:
2305:
2301:
2262:
2192:
2090:
2039:
1980:
1945:
1891:
1852:
1628:
1155:
448:
155:
2858:
2239:
2151:
7909:
7397:
6212:
6141:
5383:
5370:
5313:
5076:
4974:
4876:
4794:
4572:
4549:
4515:
4445:
3986:
3902:
3854:
3828:
3541:
3446:
2967:
2814:
2711:
2629:
2527:
2513:
2486:
2426:
2362:
2333:
2266:
2216:
2184:
2082:
2031:
1985:
1895:
1856:
1844:
1075:
1027:
542:
481:
472:
456:
390:
264:
2893:
2761:
2562:
2196:
2094:
2043:
7626:
7509:
7501:
7293:
7222:
7147:
6689:
6592:
6579:
6511:
6391:
6380:
5630:
5487:
5424:
5199:
5181:
5019:
4843:
4659:
4645:
4603:
4109:
4095:
4027:
3706:
3256:
2957:
2949:
2881:
2800:
2749:
2701:
2691:
2678:
Humphries, Stuart; Bonser, Richard H. C.; Witton, Mark P; Martill, David M (2007).
2664:
2660:
2619:
2609:
2550:
2505:
2472:
2416:
2406:
2297:
2254:
2176:
2136:
2132:
2074:
2023:
1975:
1965:
1883:
1834:
1707:
933:
845:
640:
was found to be a valid taxon in 2021 and confirmed to belong to the same genus as
328:
244:
218:
2805:
2776:
2477:
2448:
1839:
1810:
524:
403:, named by Douglas Lawson in 1975 after the tailless fixed-wing aircraft designer
7715:
7558:
7407:
7192:
7096:
7071:
7064:
7051:
6984:
6870:
6768:
6734:
6630:
6610:
6464:
6322:
6263:
6250:
6221:
6097:
6008:
5867:
5740:
5695:
5567:
5496:
5475:
5301:
4962:
4942:
4914:
4890:
4826:
4818:
4811:
4803:
4558:
4538:
4368:
4344:
4294:
4102:
4085:
4075:
4052:
4008:
4000:
3968:
3957:
3936:
3890:
3882:
3821:
3737:
3677:
3596:
3565:
3455:
3434:
3387:
3272:
3247:
2913:
2696:
2614:
2411:
2152:"A new giant pterosaur with a robust skull from the latest Cretaceous of Romania"
2078:
2027:
1970:
1605:
383:
41:
2953:
2732:
Padian, K. (1983). "A functional analysis of flying and walking in pterosaurs".
721:
wingspan, would have been at least 3 m (9.8 ft) high at the shoulder.
7692:
7665:
7643:
7467:
7381:
7366:
7318:
7257:
7242:
7208:
7033:
7017:
7005:
6991:
6944:
6901:
6861:
6775:
6721:
6449:
6435:
6407:
6314:
6235:
6162:
6073:
5829:
5721:
5656:
5597:
5537:
5503:
5223:
5150:
5122:
5041:
4991:
4930:
4899:
4499:
4475:
4453:
4399:
4357:
4301:
4269:
3950:
3867:
3844:
3723:
3649:
3617:
3470:
3419:
3367:
3062:
1674:
much like how modern birds tuck their legs beneath their own bodies in flight.
1646:
1596:
1580:
1521:
1517:
792:
679:
3157:
3145:
3133:
3121:
2885:
2753:
2596:
Longrich, Nicholas R.; Martill, David M.; Andres, Brian; Penny, David (2018).
2554:
2543:
Earth and Environmental Science Transactions of the Royal Society of Edinburgh
2180:
1946:"A Reappraisal of azhdarchid pterosaur functional morphology and paleoecology"
417:
has gained fame as a candidate for the largest flying animal ever discovered.
7978:
7833:
7729:
7700:
7676:
7657:
7615:
7600:
7549:
7538:
7431:
7423:
7373:
7310:
7281:
7265:
7249:
7235:
7176:
7154:
7105:
6998:
6759:
6702:
6621:
6456:
6428:
6400:
6271:
6154:
5945:
5875:
5747:
5730:
5586:
5523:
5467:
5232:
5188:
5069:
5062:
5055:
5048:
5033:
4983:
4907:
4864:
4857:
4783:
4630:
4623:
4616:
4465:
4407:
4392:
4385:
4333:
4116:
3785:
3745:
3695:
3664:
3625:
3520:
3512:
3490:
3394:
3378:
2936:
Goto, Yusuke; Yoda, Ken; Weimerskirch, Henri; Sato, Katsufumi (May 2, 2022).
2357:
Dinosaurs of the Air: The Evolution and Loss of Flight in Dinosaurs and Birds
1848:
1758:
1734:
1718:
1691:
1669:
necessary downstrokes needed for takeoff. This same study also suggests that
1490:
1453:
826:
770:
743:
657:
586:
506:
445:
404:
386:
83:
45:
1737:, a developer of tailless flying wing aircraft in the 1940s. The replica of
1477:
There have been a number of different ideas proposed about the lifestyle of
7722:
7650:
7521:
7416:
7350:
7302:
7112:
7040:
6918:
6886:
6654:
6421:
6349:
6290:
6185:
5822:
5807:
5791:
5782:
5771:
5611:
5574:
5460:
5410:
5253:
5166:
5159:
5139:
5111:
5104:
5090:
5083:
4998:
4955:
4922:
4638:
4580:
4523:
4508:
4488:
4313:
4044:
4036:
4016:
3943:
3922:
3915:
3770:
3558:
3504:
3463:
2971:
2859:"Pterosaur body mass estimates from three-dimensional mathematical slicing"
2715:
2633:
2430:
2188:
2086:
2035:
1989:
1682:
1650:
1525:
1220:
804:
782:
661:
specimen. Despite this association with the remains of a large carnivorous
647:
An azhdarchid neck vertebra, discovered in 2002 from the Maastrichtian age
596:
498:
394:
376:
273:
231:
2987:"Vintage Dinosaur Art: The evolution and ecology of the Dinosaurs: Part 2"
2509:
389:
age of North America. Its name comes from the Aztec feathered serpent god
7883:
7842:
7608:
7568:
7487:
7451:
7185:
6960:
6893:
6836:
6782:
6639:
6307:
6178:
6170:
6130:
6089:
6081:
5956:
5923:
5891:
5883:
5854:
5815:
5799:
5758:
5687:
5679:
5672:
5618:
5559:
5551:
5212:
5174:
5097:
4752:
4685:
4652:
4588:
3929:
3798:
3753:
3730:
3405:
3085:"Pterosaurs Coming to London to Celebrate 350 Years of the Royal Society"
2111:
Kellner, Alexander W. A.; Langston, Wann Jr. (1996). "Cranial remains of
1588:
1434:
565:
514:
58:
2309:
1664:
published in 2021 by Kevin Padian and colleagues instead suggested that
7875:
7576:
7531:
7459:
7334:
7326:
7161:
7140:
7085:
6953:
6928:
6878:
6843:
6789:
6677:
6669:
6662:
6414:
6337:
6329:
6228:
6192:
6030:
5931:
5910:
5899:
5836:
5453:
5246:
5239:
4674:
4565:
3572:
3229:
2647:
Lehman, Thomas M.; Langston, Wann Jr. (1996). "Habitat and behavior of
1754:
1687:
1494:
103:
68:
7888:
1911:"Fleshing out the bones of Quetzalcoatlus, Earth's largest flier ever"
960:
7636:
7078:
7026:
6748:
6741:
6709:
6647:
6373:
6366:
6358:
6278:
6056:
6022:
5972:
5965:
5702:
5640:
5544:
5132:
4531:
4248:
3875:
3715:
3550:
3427:
3349:
3336:
3299:
3238:
3199:
1592:
1497:
1482:
811:
379:
205:
172:
132:
108:
52:
7804:
1702:
as a small-headed scavenger with an extremely long neck in the book
1611:
7827:
7708:
7592:
7584:
7358:
7342:
7273:
6850:
6200:
6105:
6037:
5709:
5664:
5437:
4666:
4326:
3223:
2447:
Padian, K.; Cunningham, J.R.; Langston, W. Jr.; Conway, J. (2021).
1563:
1551:
is found in plains deposits, being likely a solitary hunter, while
1542:
1509:
1325:
662:
486:
465:
192:
98:
93:
78:
73:
63:
1429:
705:
studies lowered these estimates to 11–12 m (36–39 ft).
594:
can be distinguished from other pterosaurs (i.e., if it is not a
581:
430:
113:
88:
7862:
6299:
6066:
5648:
4256:
4123:
3217:
628:
being demonstrated as a valid taxon and its relationships with
494:
182:
3168:
2446:
684:
343:
5397:
3993:
2057:
Lawson, Douglas A. (1975). "Could pterosaurs fly?". Letters.
1714:
1537:, like other pterosaurs, was a quadruped when on the ground,
1530:
441:
2677:
2385:
Witton, Mark P.; Habib, Michael B.; Laudet, Vincent (2010).
1713:
In 1985, the US Defense Advanced Research Projects Agency (
364:
349:
334:
2595:
2231:
2106:
2104:
2541:
Andres, B.; Myers, T. S. (2013). "Lone Star pterosaurs".
2259:
10.1671/0272-4634(2006)26[192:AAPCVF]2.0.CO;2
1888:
10.1671/0272-4634(2006)26[922:FIUAFT]2.0.CO;2
1446:
association probably represents semi-arid inland plains.
355:
2935:
2149:
732:
In 2021, Kevin Padian and his colleagues estimated that
2640:
2101:
27:
Genus of azhdarchid pterosaurs from the Late Cretaceous
4763:
2214:
1939:
1937:
1935:
1933:
1931:
1869:
7940:
2909:"Peerless pterosaur could fly long-distance for days"
2215:
Witton, M.P.; Martill, D.M.; Loveridge, R.F. (2010).
367:
346:
340:
331:
3055:"UAS Advanced Development: Quetzalcoatlus Northropi"
553:
in 2021, named after the genus' original describer.
361:
358:
337:
303:
280:
2380:
2378:
2238:Henderson, Michael D.; Peterson, Joseph E. (2006).
1928:
1587:. MacCready constructed a model flying machine or,
818:within Neoazhdarchia from Andres and Myers (2013).
459:, who was then a geology graduate student from the
352:
3811:
2354:
1583:used practical aerodynamics to test the flight of
455:(dated to around 68 million years ago) in 1971 by
5287:
4232:
2671:
2384:
2237:
2150:Buffetaut, E.; Grigorescu, D.; Csiki, Z. (2002).
7976:
5381:
2852:
2850:
2504:. Princeton University Press. pp. 161–162.
2375:
2143:
624:specimens. These issues can only be resolved by
6590:
6510:
5423:
2361:. Johns Hopkins University Press. p. 472.
2110:
1508:birds. They suggested that, with its long neck
604:should be regarded as a European occurrence of
3271:
2646:
2453:Lawson 1975 (Pterodactyloidea: Azhdarchoidea)"
1815:Lawson 1975 (Pterodactyloidea: Azhdarchoidea)"
1808:
513:, who drove the development of large tailless
407:. The genus also includes the smaller species
3184:
3047:
2847:
2821:
2727:
2725:
2585:. Indiana University Press. pp. 310–328.
2576:
2574:
2572:
2442:
2440:
1725:as the basis for an experimental ornithopter
1524:. In 2008, pterosaur workers Mark Witton and
7175:
2589:
2273:
2210:
2208:
2206:
2001:
1999:
1804:
1802:
1800:
1798:
1796:
1794:
7990:Late Cretaceous pterosaurs of North America
6289:
3016:
2540:
1943:
1591:, with a simple computer functioning as an
3347:
3191:
3177:
2731:
2722:
2569:
2534:
2437:
2050:
1704:The evolution and ecology of the Dinosaurs
1698:In 1975, artist Giovanni Caselli depicted
131:
3077:
2997:
2961:
2856:
2827:
2804:
2705:
2695:
2623:
2613:
2476:
2420:
2410:
2280:Langston, Wann Jr. (1981). "Pterosaurs".
2203:
1996:
1979:
1969:
1863:
1838:
1791:
1618:skeleton mounted in a flying pose at the
3158:"Going out on a limb for Quetzalcoatlus"
3022:
2978:
2346:
2328:. The Jim Pattison Group. 2014. p.
2279:
1681:
1677:
1610:
1562:
1464:
790:cervical vertebrae on the left with the
781:
742:
707:
683:
590:. This being the case and assuming that
523:
424:
420:
3010:Tetrapod Zoology at Scientific American
2984:
2906:
2900:
2502:The Princeton Field Guide to Pterosaurs
1944:Witton, Mark P.; Naish, Darren (2008).
1908:
14:
7977:
4220:
2774:
2580:
2056:
2005:
1749:In 2010, several life-sized models of
1485:lifestyle. Instead, he suggested that
814:showing the phylogenetic placement of
612:also noted that the skull material of
7809:
7808:
6577:
5369:
5368:
4750:
4219:
3334:
3210:
3172:
3061:. AeroVironment. 2011. Archived from
3023:Jacobsen, Annie (February 27, 2011).
3003:
1809:Andres, B.; Langston, W. Jr. (2021).
6578:
2499:
2352:
1909:Sanders, Robert (December 8, 2021).
3025:"When Flying Dinosaurs Were Drones"
2907:Frazier, Reid (November 22, 2010).
1757:as the centerpiece exhibit for the
24:
3134:"The Biggest Thing That Ever Flew"
3128:. Paleo Studio. December 29, 2013.
3122:"Quetzalcoatlus - Flying monsters"
2866:Journal of Vertebrate Paleontology
2785:Journal of Vertebrate Paleontology
2653:Journal of Vertebrate Paleontology
2457:Journal of Vertebrate Paleontology
2302:10.1038/scientificamerican0281-122
2247:Journal of Vertebrate Paleontology
2117:Journal of Vertebrate Paleontology
1876:Journal of Vertebrate Paleontology
1819:Journal of Vertebrate Paleontology
1469:Artist's impression of a group of
25:
8026:
3146:"Quetzalcoatlus - Ancient Animal"
3106:
3004:Naish, Darren (August 20, 2013).
1733:in Texas in 1971, named it after
1428:was abundant in Texas during the
777:
716:with a Cessna 172 light aircraft.
149:Houston Museum of Natural Science
7962:
7950:
7789:
7783:
7776:
6559:
6551:
6543:
5351:
5345:
4733:
4728:
4202:
4195:
4187:
2777:"Habitat of the giant pterosaur
1753:were put on display on London's
1324:
959:
327:
159:
146:displayed in quadrupedal stance,
56:
8005:Taxa named by Douglas A. Lawson
8000:Maastrichtian genus extinctions
4751:
3335:
3198:
2991:Love in the Time of Chasmosaurs
2929:
2830:"The Great Pterodactyl Project"
2768:
2493:
2316:
1694:'s 350th anniversary exhibition
1420:
564:was identified as problematic.
493:. The genus name refers to the
3164:. thebrainscoop. June 9, 2020.
2985:Vincent, Mark (May 14, 2012).
2665:10.1080/02724634.1996.10011371
2137:10.1080/02724634.1996.10011310
1902:
1780:Timeline of pterosaur research
688:Size comparison of a human to
668:
13:
1:
8010:Fossil taxa described in 1975
3152:. North 02. October 25, 2019.
2857:Henderson, Donald M. (2010).
2806:10.1080/02724634.2019.1593184
2478:10.1080/02724634.2020.1780247
1840:10.1080/02724634.2021.1907587
1785:
1744:National Air and Space Museum
461:Jackson School of Geosciences
312:Andres and Langston Jr., 2021
3140:. PBS Eons. August 14, 2017.
2697:10.1371/journal.pbio.0050204
2615:10.1371/journal.pbio.2001663
2412:10.1371/journal.pone.0013982
2079:10.1126/science.188.4189.676
2028:10.1126/science.187.4180.947
1971:10.1371/journal.pone.0002271
1811:"Morphology and taxonomy of
1304:Arambourgiania philadelphiae
1115:Shenzhoupterus chaoyangensis
517:aircraft designs resembling
489:of a new genus and species,
7:
2325:Guinness World Records 2014
1768:
1280:Zhejiangopterus linhaiensis
528:Skeletal reconstruction of
440:fossils were discovered in
10:
8031:
3029:Los Angeles Times Magazine
2775:Lehman, Thomas M. (2021).
2449:"Functional morphology of
1706:by British paleontologist
1489:scavenged, similar to the
1460:
677:
576:(2010) suggested that the
464:10 m (33 ft) in
7817:
7773:
7675:
7625:
7557:
7548:
7520:
7496:
7485:
7441:
7406:
7396:
7292:
7230:
7221:
7217:
7206:
7171:
7130:
7095:
7059:
7050:
7016:
6979:
6970:
6943:
6939:
6916:
6860:
6831:
6827:
6816:
6758:
6729:
6720:
6697:
6688:
6629:
6620:
6605:
6601:
6586:
6573:
6540:
6506:
6390:
6348:
6258:
6249:
6211:
6149:
6140:
6125:
6116:
6055:
6017:
6006:
5955:
5918:
5909:
5862:
5853:
5781:
5766:
5757:
5729:
5720:
5629:
5606:
5595:
5584:
5532:
5521:
5486:
5432:
5419:
5408:
5392:
5377:
5364:
5342:
5312:
5296:
5283:
5222:
5207:
5198:
5149:
5121:
5028:
5017:
4973:
4950:
4941:
4889:
4885:
4874:
4852:
4841:
4793:
4778:
4774:
4759:
4746:
4726:
4684:
4611:
4602:
4548:
4498:
4483:
4474:
4463:
4440:
4429:
4367:
4352:
4343:
4321:
4312:
4282:
4278:
4267:
4243:
4228:
4215:
4184:
4134:
4084:
4074:
4026:
3976:
3967:
3910:
3901:
3862:
3853:
3842:
3807:
3796:
3780:
3768:
3705:
3690:
3686:
3675:
3659:
3647:
3607:
3591:
3582:
3540:
3531:
3499:
3488:
3445:
3414:
3404:
3377:
3362:
3358:
3343:
3330:
3298:
3280:
3267:
3206:
2954:10.1093/pnasnexus/pgac023
2886:10.1080/02724631003758334
2837:Engineering & Science
2754:10.1017/S009483730000765X
2555:10.1017/S1755691013000303
2500:Paul, Gregory S. (2022).
2353:Paul, Gregory S. (2002).
2181:10.1007/s00114-002-0307-1
1558:
1512:and long toothless jaws,
1339:
1317:
1300:
1293:
1276:
1269:
1256:
1249:
1232:
1225:
1203:
1196:
1152:
1135:
1128:
1111:
1104:
1087:
1080:
1068:
1024:
1007:
1000:
974:
952:
945:
938:
898:
881:
874:
857:
850:
838:
831:
556:The holotype specimen of
300:
295:
279:
272:
156:Scientific classification
154:
139:
130:
34:
2583:Mesozoic Vertebrate Life
2221:Acta Geoscientica Sinica
1775:List of pterosaur genera
1763:University of Portsmouth
1723:Quetzalcoatlus northropi
1575:The nature of flight in
1549:Quetzalcoatlus northropi
1432:in a fauna dominated by
1321:Quetzalcoatlus northropi
956:Domeykodactylus ceciliae
902:Tupuxuara longicristatus
754:
747:Skull reconstruction of
491:Quetzalcoatlus northropi
285:Quetzalcoatlus northropi
18:Quetzalcoatlus northropi
4766:Archaeopterodactyloidea
1727:unmanned aerial vehicle
1207:Radiodactylus langstoni
1091:Eoazhdarcho liaoxiensis
1011:Noripterus complicidens
673:
497:feathered serpent god,
2828:MacCready, P. (1985).
1695:
1623:
1572:
1474:
1343:Quetzalcoatlus lawsoni
1139:Chaoyangopterus zhangi
803:belongs to the family
797:
751:
717:
697:
533:
453:Big Bend National Park
433:
7923:Paleobiology Database
2510:10.1515/9780691232218
1685:
1678:Cultural significance
1614:
1566:
1473:feeding on the ground
1468:
1236:Azhdarcho lancicollis
861:Thalassodromeus sethi
796:holotype on the right
785:
746:
711:
687:
651:, may also belong to
527:
507:John Knudsen Northrop
477:Texas Memorial Museum
429:Cast of the holotype
428:
421:Discovery and species
4378:Archaeoistiodactylus
4288:Archaeoistiodactylus
3117:at EarthArchives.org
1735:John "Jack" Northrop
1620:Royal Ontario Museum
1516:fed like modern-day
759:Skull material from
649:Hell Creek Formation
511:Northrop Corporation
7995:Ojo Alamo Formation
7443:Targaryendraconidae
7132:Ornithocheiriformes
6819:Ornithocheiromorpha
6118:Neopterodactyloidea
6045:Dsungaripteromorpha
3813:Campylognathoididae
3533:Eudimorphodontoidea
3089:Everything Dinosaur
2878:2010JVPal..30..768H
2797:2021JVPal..41S..21L
2746:1983Pbio....9..218P
2469:2021JVPal..41S.218P
2403:2010PLoSO...513982W
2294:1981SciAm.244b.122L
2282:Scientific American
2173:2002NW.....89..180B
2160:Naturwissenschaften
2129:1996JVPal..16..222K
2071:1975Sci...188..676G
2020:1975Sci...187..947L
1962:2008PLoSO...3.2271W
1831:2021JVPal..41S..46A
1686:Models on London's
978:Dsungaripterus weii
885:Tupuxuara leonardii
6972:Istiodactyliformes
5446:Lonchognathosaurus
5289:Eupterodactyloidea
4432:Pterodactyliformes
4235:Pterodactylomorpha
4222:Pterodactylomorpha
4136:Pterodactylomorpha
3978:Angustinaripterini
3609:Eudimorphodontinae
3584:Eudimorphodontidae
3065:on January 7, 2012
1696:
1629:Chatham University
1624:
1573:
1475:
1156:Jidapterus edentus
798:
752:
718:
698:
608:. However, Witton
534:
449:Javelina Formation
434:
7938:
7937:
7910:Open Tree of Life
7811:Taxon identifiers
7802:
7801:
7798:
7797:
7771:
7770:
7767:
7766:
7763:
7762:
7759:
7758:
7755:
7754:
7751:
7750:
7747:
7746:
7743:
7742:
7739:
7738:
7481:
7480:
7477:
7476:
7398:Targaryendraconia
7392:
7391:
7202:
7201:
7126:
7125:
7122:
7121:
6912:
6911:
6812:
6811:
6808:
6807:
6804:
6803:
6800:
6799:
6569:
6568:
6538:
6537:
6534:
6533:
6530:
6529:
6502:
6501:
6498:
6497:
6494:
6493:
6490:
6489:
6486:
6485:
6482:
6481:
6478:
6477:
6474:
6473:
6245:
6244:
6213:Chaoyangopterinae
6142:Chaoyangopteridae
6002:
6001:
5998:
5997:
5994:
5993:
5990:
5989:
5986:
5985:
5982:
5981:
5517:
5516:
5513:
5512:
5384:Ornithocheiroidea
5371:Ornithocheiroidea
5360:
5359:
5340:
5339:
5336:
5335:
5332:
5331:
5314:Ornithocheiroidea
5279:
5278:
5275:
5274:
5271:
5270:
5267:
5266:
5263:
5262:
5077:Gladocephaloideus
5013:
5012:
5009:
5008:
4975:Aurorazhdarchidae
4877:Ctenochasmatoidea
4837:
4836:
4795:Germanodactylidae
4742:
4741:
4724:
4723:
4720:
4719:
4716:
4715:
4712:
4711:
4708:
4707:
4704:
4703:
4598:
4597:
4573:Dendrorhynchoides
4550:Batrachognathinae
4516:Dendrorhynchoides
4446:Changchengopterus
4425:
4424:
4421:
4420:
4417:
4416:
4211:
4210:
4182:
4181:
4178:
4177:
4174:
4173:
4170:
4169:
4166:
4165:
4162:
4161:
4158:
4157:
4154:
4153:
4070:
4069:
4066:
4065:
4062:
4061:
3987:Angustinaripterus
3903:Rhamphorhynchinae
3855:Rhamphorhynchidae
3838:
3837:
3829:Campylognathoides
3764:
3763:
3643:
3642:
3639:
3638:
3635:
3634:
3542:Raeticodactylidae
3484:
3483:
3480:
3479:
3447:Austriadraconidae
3326:
3325:
3322:
3321:
3318:
3317:
2463:(sup1): 218–251.
2339:978-1-908843-15-9
2065:(4189): 676–677.
2014:(4180): 947–948.
1567:Restoration of a
1417:
1416:
1408:
1407:
1399:
1398:
1390:
1389:
1381:
1380:
1372:
1371:
1363:
1362:
1354:
1353:
1185:
1184:
1176:
1175:
1167:
1166:
1076:Chaoyangopteridae
1057:
1056:
1048:
1047:
1039:
1038:
1028:Noripterus parvus
989:
988:
922:
921:
913:
912:
658:Tyrannosaurus rex
543:Alexander Kellner
509:, the founder of
473:Wann Langston Jr.
457:Douglas A. Lawson
319:
318:
313:
268:
16:(Redirected from
8022:
7967:
7966:
7965:
7955:
7954:
7953:
7946:
7931:
7930:
7918:
7917:
7905:
7904:
7892:
7891:
7879:
7878:
7866:
7865:
7853:
7852:
7851:
7838:
7837:
7836:
7806:
7805:
7793:
7787:
7781:
7780:
7627:Coloborhynchinae
7555:
7554:
7510:Ornithocheiridae
7502:Brasileodactylus
7494:
7493:
7404:
7403:
7294:Ornithocheirinae
7228:
7227:
7223:Ornithocheiridae
7219:
7218:
7215:
7214:
7173:
7172:
7169:
7168:
7148:Brasileodactylus
7057:
7056:
6977:
6976:
6945:Lonchodraconidae
6941:
6940:
6937:
6936:
6829:
6828:
6825:
6824:
6727:
6726:
6695:
6694:
6690:Nyctosauromorpha
6627:
6626:
6603:
6602:
6599:
6598:
6593:Pteranodontoidea
6588:
6587:
6580:Pteranodontoidea
6575:
6574:
6564:
6563:
6556:
6555:
6548:
6547:
6512:Pteranodontoidea
6508:
6507:
6392:Quetzalcoatlinae
6381:Mistralazhdarcho
6346:
6345:
6287:
6286:
6256:
6255:
6147:
6146:
6123:
6122:
6015:
6014:
5916:
5915:
5860:
5859:
5779:
5778:
5764:
5763:
5727:
5726:
5631:Thalassodromidae
5604:
5603:
5593:
5592:
5530:
5529:
5488:Dsungaripterinae
5430:
5429:
5425:Dsungaripteridae
5421:
5420:
5417:
5416:
5390:
5389:
5379:
5378:
5366:
5365:
5355:
5350:
5349:
5294:
5293:
5285:
5284:
5205:
5204:
5200:Ctenochasmatinae
5182:Plataleorhynchus
5026:
5025:
5020:Ctenochasmatidae
4948:
4947:
4887:
4886:
4883:
4882:
4850:
4849:
4844:Euctenochasmatia
4776:
4775:
4772:
4771:
4761:
4760:
4748:
4747:
4737:
4732:
4660:Propterodactylus
4646:Ningchengopterus
4609:
4608:
4604:Pterodactyloidea
4481:
4480:
4472:
4471:
4438:
4437:
4350:
4349:
4319:
4318:
4280:
4279:
4276:
4275:
4241:
4240:
4230:
4229:
4217:
4216:
4206:
4200:
4199:
4192:
4191:
4110:Jianchangopterus
4096:Fenghuangopterus
4082:
4081:
4028:Rhamphorhynchini
3974:
3973:
3908:
3907:
3860:
3859:
3851:
3850:
3809:
3808:
3805:
3804:
3778:
3777:
3707:Dimorphodontidae
3688:
3687:
3684:
3683:
3657:
3656:
3589:
3588:
3538:
3537:
3497:
3496:
3412:
3411:
3360:
3359:
3356:
3355:
3345:
3344:
3332:
3331:
3278:
3277:
3269:
3268:
3262:
3261:
3257:Pterosauromorpha
3208:
3207:
3193:
3186:
3179:
3170:
3169:
3165:
3153:
3141:
3129:
3101:
3100:
3098:
3096:
3081:
3075:
3074:
3072:
3070:
3051:
3045:
3044:
3042:
3040:
3035:on March 8, 2013
3031:. Archived from
3020:
3014:
3013:
3001:
2995:
2994:
2982:
2976:
2975:
2965:
2933:
2927:
2926:
2924:
2922:
2904:
2898:
2897:
2863:
2854:
2845:
2844:
2834:
2825:
2819:
2818:
2808:
2772:
2766:
2765:
2729:
2720:
2719:
2709:
2699:
2675:
2669:
2668:
2644:
2638:
2637:
2627:
2617:
2593:
2587:
2586:
2578:
2567:
2566:
2538:
2532:
2531:
2497:
2491:
2490:
2480:
2444:
2435:
2434:
2424:
2414:
2382:
2373:
2372:
2360:
2350:
2344:
2343:
2320:
2314:
2313:
2277:
2271:
2270:
2244:
2235:
2229:
2228:
2212:
2201:
2200:
2156:
2147:
2141:
2140:
2108:
2099:
2098:
2054:
2048:
2047:
2003:
1994:
1993:
1983:
1973:
1941:
1926:
1925:
1923:
1921:
1906:
1900:
1899:
1867:
1861:
1860:
1842:
1806:
1708:Beverly Halstead
1328:
1296:
1295:
1272:
1271:
1252:
1251:
1228:
1227:
1199:
1198:
1131:
1130:
1107:
1106:
1083:
1082:
1071:
1070:
1003:
1002:
963:
948:
947:
941:
940:
934:Dsungaripteridae
877:
876:
853:
852:
846:Thalassodrominae
841:
840:
834:
833:
822:
821:
786:A comparison of
584:giant azhdarchid
374:
373:
370:
369:
366:
363:
360:
357:
354:
351:
348:
345:
342:
339:
336:
333:
311:
305:
282:
263:
256:
245:Quetzalcoatlinae
243:
230:
219:Pterodactyloidea
217:
204:
164:
163:
135:
125:
55:
40:Temporal range:
32:
31:
21:
8030:
8029:
8025:
8024:
8023:
8021:
8020:
8019:
7975:
7974:
7973:
7963:
7961:
7951:
7949:
7941:
7939:
7934:
7926:
7921:
7913:
7908:
7900:
7895:
7887:
7882:
7874:
7869:
7861:
7856:
7847:
7846:
7841:
7832:
7831:
7826:
7813:
7803:
7794:
7775:
7735:
7716:Liaoningopterus
7671:
7621:
7559:Tropeognathinae
7544:
7516:
7490:
7473:
7437:
7408:Cimoliopteridae
7388:
7288:
7211:
7198:
7193:Zhenyuanopterus
7167:
7118:
7097:Istiodactylinae
7091:
7072:Longchengpterus
7065:Lingyuanopterus
7052:Istiodactylidae
7046:
7012:
6985:Hongshanopterus
6966:
6935:
6921:
6908:
6871:Hongshanopterus
6856:
6821:
6796:
6769:Barbaridactylus
6754:
6716:
6684:
6631:Pteranodontidae
6616:
6611:Santanadactylus
6595:
6582:
6565:
6558:
6550:
6542:
6526:
6470:
6465:Zhejiangopterus
6386:
6344:
6323:Palaeocursornis
6285:
6264:Montanazhdarcho
6251:Azhdarchiformes
6241:
6222:Chaoyangopterus
6207:
6136:
6112:
6098:Montanazhdarcho
6051:
6011:
6009:Azhdarchomorpha
5978:
5951:
5905:
5868:Aymberedactylus
5849:
5777:
5753:
5741:Aymberedactylus
5716:
5696:Thalassodromeus
5625:
5600:
5589:
5580:
5568:Montanazhdarcho
5526:
5509:
5497:Domeykodactylus
5482:
5476:Tendaguripterus
5413:
5404:
5386:
5373:
5356:
5344:
5328:
5308:
5302:Altmuehlopterus
5259:
5218:
5194:
5145:
5117:
5022:
5005:
4969:
4963:Gallodactylidae
4943:Aurorazhdarchia
4937:
4915:Normannognathus
4891:Gallodactylidae
4879:
4870:
4846:
4833:
4827:Tendaguripterus
4819:Normannognathus
4812:Germanodactylus
4804:Altmuehlopterus
4789:
4768:
4755:
4738:
4700:
4680:
4594:
4559:Batrachognathus
4544:
4539:Vesperopterylus
4494:
4468:
4459:
4434:
4413:
4369:Wukongopterinae
4363:
4345:Wukongopteridae
4339:
4308:
4295:Normannognathus
4272:
4263:
4237:
4224:
4207:
4194:
4186:
4150:
4130:
4103:Jianchangnathus
4086:Scaphognathidae
4076:Digibrevisauria
4058:
4053:Rhamphorhynchus
4022:
4009:Qinglongopterus
4001:Harpactognathus
3969:Rhamphorhynchae
3963:
3958:Qinglongopterus
3937:Harpactognathus
3897:
3891:Scaphognathinae
3883:Parapsicephalus
3847:
3834:
3822:Bergamodactylus
3801:
3792:
3774:
3760:
3738:Parapsicephalus
3701:
3680:
3678:Macronychoptera
3671:
3653:
3631:
3603:
3597:Arcticodactylus
3578:
3566:Raeticodactylus
3527:
3493:
3476:
3456:Arcticodactylus
3441:
3435:Raeticodactylus
3400:
3388:Austriadactylus
3373:
3352:
3339:
3314:
3294:
3288:Avemetatarsalia
3273:Avemetatarsalia
3263:
3248:Avemetatarsalia
3213:
3212:
3202:
3197:
3156:
3144:
3132:
3120:
3109:
3104:
3094:
3092:
3091:. June 22, 2010
3083:
3082:
3078:
3068:
3066:
3053:
3052:
3048:
3038:
3036:
3021:
3017:
3002:
2998:
2983:
2979:
2934:
2930:
2920:
2918:
2914:Morning Edition
2905:
2901:
2861:
2855:
2848:
2832:
2826:
2822:
2791:(sup1): 21–45.
2773:
2769:
2730:
2723:
2676:
2672:
2659:(Sup. 3): 48A.
2645:
2641:
2608:(3): e2001663.
2594:
2590:
2579:
2570:
2539:
2535:
2520:
2498:
2494:
2445:
2438:
2383:
2376:
2369:
2351:
2347:
2340:
2322:
2321:
2317:
2278:
2274:
2242:
2236:
2232:
2213:
2204:
2154:
2148:
2144:
2109:
2102:
2055:
2051:
2004:
1997:
1942:
1929:
1919:
1917:
1907:
1903:
1868:
1864:
1807:
1792:
1788:
1771:
1680:
1647:thermal soaring
1561:
1463:
1423:
1418:
1409:
1400:
1391:
1382:
1373:
1364:
1355:
1186:
1177:
1168:
1058:
1049:
1040:
990:
923:
914:
780:
757:
682:
676:
671:
423:
384:Late Cretaceous
382:known from the
330:
326:
291:
288:
262:
254:
241:
228:
215:
202:
158:
147:
145:
126:
124:
123:
122:
121:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
50:
49:
42:Late Cretaceous
38:
28:
23:
22:
15:
12:
11:
5:
8028:
8018:
8017:
8012:
8007:
8002:
7997:
7992:
7987:
7972:
7971:
7959:
7936:
7935:
7933:
7932:
7919:
7906:
7893:
7880:
7867:
7854:
7849:Quetzalcoatlus
7839:
7823:
7821:
7819:Quetzalcoatlus
7815:
7814:
7800:
7799:
7796:
7795:
7774:
7772:
7769:
7768:
7765:
7764:
7761:
7760:
7757:
7756:
7753:
7752:
7749:
7748:
7745:
7744:
7741:
7740:
7737:
7736:
7734:
7733:
7726:
7719:
7712:
7705:
7697:
7693:Caulkicephalus
7689:
7681:
7679:
7673:
7672:
7670:
7669:
7666:Uktenadactylus
7662:
7654:
7647:
7644:Coloborhynchus
7640:
7632:
7630:
7623:
7622:
7620:
7619:
7612:
7605:
7597:
7589:
7581:
7573:
7564:
7562:
7552:
7546:
7545:
7543:
7542:
7535:
7527:
7525:
7518:
7517:
7515:
7514:
7506:
7497:
7491:
7486:
7483:
7482:
7479:
7478:
7475:
7474:
7472:
7471:
7468:Targaryendraco
7464:
7456:
7447:
7445:
7439:
7438:
7436:
7435:
7428:
7420:
7412:
7410:
7401:
7394:
7393:
7390:
7389:
7387:
7386:
7382:Uktenadactylus
7378:
7370:
7367:Ornithocheirus
7363:
7355:
7347:
7339:
7331:
7323:
7319:Coloborhynchus
7315:
7307:
7298:
7296:
7290:
7289:
7287:
7286:
7278:
7270:
7262:
7258:Caulkicephalus
7254:
7246:
7243:Arthurdactylus
7239:
7231:
7225:
7212:
7209:Ornithocheirae
7207:
7204:
7203:
7200:
7199:
7197:
7196:
7189:
7181:
7179:
7166:
7165:
7158:
7151:
7144:
7136:
7134:
7128:
7127:
7124:
7123:
7120:
7119:
7117:
7116:
7109:
7101:
7099:
7093:
7092:
7090:
7089:
7082:
7075:
7068:
7060:
7054:
7048:
7047:
7045:
7044:
7037:
7034:Linlongopterus
7030:
7022:
7020:
7018:Mimodactylidae
7014:
7013:
7011:
7010:
7006:Lonchodectidae
7002:
6995:
6992:Linlongopterus
6988:
6980:
6974:
6968:
6967:
6965:
6964:
6957:
6949:
6947:
6934:
6933:
6924:
6922:
6917:
6914:
6913:
6910:
6909:
6907:
6906:
6902:Targaryendraco
6898:
6890:
6883:
6875:
6866:
6864:
6862:Lonchodectidae
6858:
6857:
6855:
6854:
6847:
6840:
6832:
6822:
6817:
6814:
6813:
6810:
6809:
6806:
6805:
6802:
6801:
6798:
6797:
6795:
6794:
6786:
6779:
6776:Muzquizopteryx
6772:
6764:
6762:
6756:
6755:
6753:
6752:
6745:
6738:
6730:
6724:
6722:Aponyctosauria
6718:
6717:
6715:
6714:
6706:
6698:
6692:
6686:
6685:
6683:
6682:
6674:
6666:
6659:
6651:
6644:
6635:
6633:
6624:
6618:
6617:
6615:
6614:
6606:
6596:
6591:
6584:
6583:
6571:
6570:
6567:
6566:
6541:
6539:
6536:
6535:
6532:
6531:
6528:
6527:
6525:
6524:
6523:
6522:
6516:
6514:
6504:
6503:
6500:
6499:
6496:
6495:
6492:
6491:
6488:
6487:
6484:
6483:
6480:
6479:
6476:
6475:
6472:
6471:
6469:
6468:
6461:
6453:
6450:Thanatosdrakon
6446:
6443:Quetzalcoatlus
6439:
6436:Phosphatodraco
6432:
6425:
6418:
6411:
6408:Arambourgiania
6404:
6396:
6394:
6388:
6387:
6385:
6384:
6377:
6370:
6363:
6354:
6352:
6343:
6342:
6334:
6326:
6319:
6315:Navajodactylus
6311:
6304:
6295:
6293:
6284:
6283:
6275:
6268:
6259:
6253:
6247:
6246:
6243:
6242:
6240:
6239:
6236:Shenzhoupterus
6232:
6225:
6217:
6215:
6209:
6208:
6206:
6205:
6197:
6189:
6182:
6175:
6167:
6163:Argentinadraco
6159:
6150:
6144:
6138:
6137:
6135:
6134:
6126:
6120:
6114:
6113:
6111:
6110:
6102:
6094:
6086:
6078:
6074:Argentinadraco
6070:
6062:
6060:
6053:
6052:
6050:
6049:
6041:
6034:
6027:
6018:
6012:
6007:
6004:
6003:
6000:
5999:
5996:
5995:
5992:
5991:
5988:
5987:
5984:
5983:
5980:
5979:
5977:
5976:
5969:
5961:
5959:
5953:
5952:
5950:
5949:
5942:
5935:
5928:
5919:
5913:
5907:
5906:
5904:
5903:
5896:
5888:
5880:
5872:
5863:
5857:
5851:
5850:
5848:
5847:
5840:
5833:
5830:Nemicolopterus
5826:
5819:
5812:
5804:
5796:
5787:
5785:
5776:
5775:
5767:
5761:
5755:
5754:
5752:
5751:
5744:
5736:
5734:
5724:
5722:Tapejariformes
5718:
5717:
5715:
5714:
5706:
5699:
5692:
5684:
5676:
5669:
5661:
5657:Argentinadraco
5653:
5645:
5636:
5634:
5627:
5626:
5624:
5623:
5615:
5607:
5601:
5598:Tapejaromorpha
5596:
5590:
5585:
5582:
5581:
5579:
5578:
5571:
5564:
5556:
5548:
5541:
5538:Argentinadraco
5533:
5527:
5522:
5519:
5518:
5515:
5514:
5511:
5510:
5508:
5507:
5504:Dsungaripterus
5500:
5492:
5490:
5484:
5483:
5481:
5480:
5472:
5464:
5457:
5450:
5442:
5433:
5427:
5414:
5409:
5406:
5405:
5403:
5402:
5393:
5387:
5382:
5375:
5374:
5362:
5361:
5358:
5357:
5343:
5341:
5338:
5337:
5334:
5333:
5330:
5329:
5327:
5326:
5325:
5324:
5318:
5316:
5310:
5309:
5307:
5306:
5297:
5291:
5281:
5280:
5277:
5276:
5273:
5272:
5269:
5268:
5265:
5264:
5261:
5260:
5258:
5257:
5250:
5243:
5236:
5228:
5226:
5224:Pterodaustrini
5220:
5219:
5217:
5216:
5208:
5202:
5196:
5195:
5193:
5192:
5185:
5178:
5171:
5163:
5155:
5153:
5151:Gnathosaurinae
5147:
5146:
5144:
5143:
5136:
5128:
5126:
5123:Moganopterinae
5119:
5118:
5116:
5115:
5108:
5101:
5094:
5087:
5080:
5073:
5066:
5059:
5052:
5045:
5042:Balaenognathus
5038:
5029:
5023:
5018:
5015:
5014:
5011:
5010:
5007:
5006:
5004:
5003:
4995:
4992:Aurorazhdarcho
4988:
4979:
4977:
4971:
4970:
4968:
4967:
4959:
4951:
4945:
4939:
4938:
4936:
4935:
4931:Moganopterinae
4927:
4919:
4911:
4904:
4900:Aurorazhdarcho
4895:
4893:
4880:
4875:
4872:
4871:
4869:
4868:
4861:
4853:
4847:
4842:
4839:
4838:
4835:
4834:
4832:
4831:
4823:
4815:
4808:
4799:
4797:
4791:
4790:
4788:
4787:
4779:
4769:
4764:
4757:
4756:
4744:
4743:
4740:
4739:
4727:
4725:
4722:
4721:
4718:
4717:
4714:
4713:
4710:
4709:
4706:
4705:
4702:
4701:
4699:
4698:
4697:
4696:
4690:
4688:
4682:
4681:
4679:
4678:
4671:
4663:
4656:
4649:
4642:
4635:
4627:
4620:
4612:
4606:
4600:
4599:
4596:
4595:
4593:
4592:
4585:
4577:
4569:
4562:
4554:
4552:
4546:
4545:
4543:
4542:
4535:
4528:
4520:
4512:
4504:
4502:
4500:Anurognathinae
4496:
4495:
4493:
4492:
4484:
4478:
4476:Anurognathidae
4469:
4464:
4461:
4460:
4458:
4457:
4454:Douzhanopterus
4450:
4441:
4435:
4430:
4427:
4426:
4423:
4422:
4419:
4418:
4415:
4414:
4412:
4411:
4404:
4400:Douzhanopterus
4396:
4389:
4382:
4373:
4371:
4365:
4364:
4362:
4361:
4358:Kunpengopterus
4353:
4347:
4341:
4340:
4338:
4337:
4330:
4322:
4316:
4310:
4309:
4307:
4306:
4302:Anurognathidae
4298:
4291:
4283:
4273:
4270:Monofenestrata
4268:
4265:
4264:
4262:
4261:
4253:
4244:
4238:
4233:
4226:
4225:
4213:
4212:
4209:
4208:
4185:
4183:
4180:
4179:
4176:
4175:
4172:
4171:
4168:
4167:
4164:
4163:
4160:
4159:
4156:
4155:
4152:
4151:
4149:
4148:
4147:
4146:
4140:
4138:
4132:
4131:
4129:
4128:
4120:
4113:
4106:
4099:
4091:
4089:
4079:
4072:
4071:
4068:
4067:
4064:
4063:
4060:
4059:
4057:
4056:
4049:
4041:
4032:
4030:
4024:
4023:
4021:
4020:
4013:
4005:
3997:
3990:
3982:
3980:
3971:
3965:
3964:
3962:
3961:
3954:
3951:Orientognathus
3947:
3940:
3933:
3926:
3919:
3911:
3905:
3899:
3898:
3896:
3895:
3887:
3879:
3872:
3868:Dolicorhamphus
3863:
3857:
3848:
3845:Breviquartossa
3843:
3840:
3839:
3836:
3835:
3833:
3832:
3825:
3817:
3815:
3802:
3797:
3794:
3793:
3791:
3790:
3781:
3775:
3769:
3766:
3765:
3762:
3761:
3759:
3758:
3750:
3742:
3734:
3727:
3724:Caelestiventus
3720:
3711:
3709:
3703:
3702:
3700:
3699:
3691:
3681:
3676:
3673:
3672:
3670:
3669:
3660:
3654:
3650:Zambellisauria
3648:
3645:
3644:
3641:
3640:
3637:
3636:
3633:
3632:
3630:
3629:
3622:
3618:Carniadactylus
3613:
3611:
3605:
3604:
3602:
3601:
3592:
3586:
3580:
3579:
3577:
3576:
3569:
3562:
3555:
3546:
3544:
3535:
3529:
3528:
3526:
3525:
3517:
3509:
3500:
3494:
3489:
3486:
3485:
3482:
3481:
3478:
3477:
3475:
3474:
3471:Seazzadactylus
3467:
3460:
3451:
3449:
3443:
3442:
3440:
3439:
3431:
3424:
3420:Carniadactylus
3415:
3409:
3402:
3401:
3399:
3398:
3391:
3383:
3381:
3375:
3374:
3372:
3371:
3368:Daohugoupterus
3363:
3353:
3348:
3341:
3340:
3328:
3327:
3324:
3323:
3320:
3319:
3316:
3315:
3313:
3312:
3311:
3310:
3304:
3302:
3296:
3295:
3293:
3292:
3291:
3290:
3281:
3275:
3265:
3264:
3260:
3259:
3250:
3241:
3232:
3226:
3220:
3204:
3203:
3196:
3195:
3188:
3181:
3173:
3167:
3166:
3154:
3142:
3130:
3118:
3114:Quetzalcoatlus
3108:
3107:External links
3105:
3103:
3102:
3076:
3046:
3015:
2996:
2977:
2948:(1): pgac023.
2928:
2899:
2872:(3): 768–785.
2846:
2820:
2779:Quetzalcoatlus
2767:
2740:(3): 218–239.
2721:
2670:
2649:Quetzalcoatlus
2639:
2588:
2568:
2533:
2518:
2492:
2451:Quetzalcoatlus
2436:
2397:(11): e13982.
2374:
2367:
2345:
2338:
2315:
2288:(2): 122–137.
2272:
2253:(1): 192–195.
2230:
2202:
2167:(4): 180–184.
2142:
2123:(2): 222–231.
2113:Quetzalcoatlus
2100:
2049:
1995:
1927:
1901:
1882:(4): 922–928.
1862:
1813:Quetzalcoatlus
1789:
1787:
1784:
1783:
1782:
1777:
1770:
1767:
1700:Quetzalcoatlus
1679:
1676:
1671:Quetzalcoatlus
1666:Quetzalcoatlus
1616:Quetzalcoatlus
1585:Quetzalcoatlus
1581:Paul MacCready
1577:Quetzalcoatlus
1569:Quetzalcoatlus
1560:
1557:
1539:Quetzalcoatlus
1535:Quetzalcoatlus
1514:Quetzalcoatlus
1487:Quetzalcoatlus
1479:Quetzalcoatlus
1471:Quetzalcoatlus
1462:
1459:
1448:Quetzalcoatlus
1444:Quetzalcoatlus
1426:Quetzalcoatlus
1422:
1419:
1415:
1414:
1411:
1410:
1406:
1405:
1402:
1401:
1397:
1396:
1393:
1392:
1388:
1387:
1384:
1383:
1379:
1378:
1375:
1374:
1370:
1369:
1366:
1365:
1361:
1360:
1357:
1356:
1352:
1351:
1348:
1347:
1338:
1335:
1334:
1331:
1330:
1316:
1313:
1312:
1309:
1308:
1299:
1294:
1292:
1289:
1288:
1285:
1284:
1275:
1270:
1268:
1265:
1264:
1261:
1260:
1255:
1250:
1248:
1245:
1244:
1241:
1240:
1231:
1226:
1224:
1216:
1215:
1212:
1211:
1202:
1197:
1195:
1192:
1191:
1188:
1187:
1183:
1182:
1179:
1178:
1174:
1173:
1170:
1169:
1165:
1164:
1161:
1160:
1151:
1148:
1147:
1144:
1143:
1134:
1129:
1127:
1124:
1123:
1120:
1119:
1110:
1105:
1103:
1100:
1099:
1096:
1095:
1086:
1081:
1079:
1069:
1067:
1064:
1063:
1060:
1059:
1055:
1054:
1051:
1050:
1046:
1045:
1042:
1041:
1037:
1036:
1033:
1032:
1023:
1020:
1019:
1016:
1015:
1006:
1001:
999:
996:
995:
992:
991:
987:
986:
983:
982:
973:
970:
969:
966:
965:
951:
946:
944:
939:
937:
929:
928:
925:
924:
920:
919:
916:
915:
911:
910:
907:
906:
897:
894:
893:
890:
889:
880:
875:
873:
870:
869:
866:
865:
856:
851:
849:
839:
837:
832:
830:
820:
816:Quetzalcoatlus
801:Quetzalcoatlus
793:Phosphatodraco
788:Quetzalcoatlus
779:
778:Classification
776:
765:Quetzalcoatlus
756:
753:
727:Quetzalcoatlus
712:Comparison of
702:Quetzalcoatlus
680:Pterosaur size
675:
672:
670:
667:
653:Quetzalcoatlus
622:Quetzalcoatlus
606:Quetzalcoatlus
562:Quetzalcoatlus
539:Quetzalcoatlus
519:Quetzalcoatlus
438:Quetzalcoatlus
422:
419:
375:is a genus of
323:Quetzalcoatlus
317:
316:
315:
314:
298:
297:
296:Other species
293:
292:
289:
277:
276:
270:
269:
259:Quetzalcoatlus
252:
248:
247:
239:
235:
234:
226:
222:
221:
213:
209:
208:
200:
196:
195:
190:
186:
185:
180:
176:
175:
170:
166:
165:
152:
151:
142:Quetzalcoatlus
137:
136:
128:
127:
119:
118:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
39:
36:Quetzalcoatlus
26:
9:
6:
4:
3:
2:
8027:
8016:
8013:
8011:
8008:
8006:
8003:
8001:
7998:
7996:
7993:
7991:
7988:
7986:
7983:
7982:
7980:
7970:
7969:United States
7960:
7958:
7948:
7947:
7944:
7929:
7924:
7920:
7916:
7911:
7907:
7903:
7898:
7894:
7890:
7885:
7881:
7877:
7872:
7868:
7864:
7859:
7855:
7850:
7844:
7840:
7835:
7829:
7825:
7824:
7822:
7820:
7816:
7812:
7807:
7792:
7788:
7786:
7779:
7732:
7731:
7730:Maaradactylus
7727:
7725:
7724:
7720:
7718:
7717:
7713:
7711:
7710:
7706:
7703:
7702:
7701:Cearadactylus
7698:
7695:
7694:
7690:
7688:
7687:
7683:
7682:
7680:
7678:
7677:Anhanguerinae
7674:
7668:
7667:
7663:
7660:
7659:
7658:Siroccopteryx
7655:
7653:
7652:
7648:
7646:
7645:
7641:
7639:
7638:
7634:
7633:
7631:
7628:
7624:
7618:
7617:
7616:Tropeognathus
7613:
7611:
7610:
7606:
7603:
7602:
7601:Siroccopteryx
7598:
7595:
7594:
7590:
7587:
7586:
7582:
7579:
7578:
7574:
7571:
7570:
7566:
7565:
7563:
7560:
7556:
7553:
7551:
7550:Anhangueridae
7547:
7541:
7540:
7539:Iberodactylus
7536:
7534:
7533:
7529:
7528:
7526:
7523:
7519:
7512:
7511:
7507:
7504:
7503:
7499:
7498:
7495:
7492:
7489:
7484:
7470:
7469:
7465:
7462:
7461:
7457:
7454:
7453:
7449:
7448:
7446:
7444:
7440:
7434:
7433:
7432:Cimoliopterus
7429:
7426:
7425:
7424:Camposipterus
7421:
7419:
7418:
7414:
7413:
7411:
7409:
7405:
7402:
7399:
7395:
7384:
7383:
7379:
7376:
7375:
7374:Siroccopteryx
7371:
7369:
7368:
7364:
7361:
7360:
7356:
7353:
7352:
7348:
7345:
7344:
7340:
7337:
7336:
7332:
7329:
7328:
7324:
7321:
7320:
7316:
7313:
7312:
7311:Camposipterus
7308:
7305:
7304:
7300:
7299:
7297:
7295:
7291:
7284:
7283:
7282:Tropeognathus
7279:
7276:
7275:
7271:
7268:
7267:
7266:Cimoliopterus
7263:
7260:
7259:
7255:
7252:
7251:
7250:Camposipterus
7247:
7245:
7244:
7240:
7238:
7237:
7236:Araripesaurus
7233:
7232:
7229:
7226:
7224:
7220:
7216:
7213:
7210:
7205:
7195:
7194:
7190:
7188:
7187:
7183:
7182:
7180:
7178:
7177:Boreopteridae
7174:
7170:
7164:
7163:
7159:
7157:
7156:
7155:Cearadactylus
7152:
7150:
7149:
7145:
7143:
7142:
7138:
7137:
7135:
7133:
7129:
7115:
7114:
7110:
7108:
7107:
7106:Istiodactylus
7103:
7102:
7100:
7098:
7094:
7088:
7087:
7083:
7081:
7080:
7076:
7074:
7073:
7069:
7067:
7066:
7062:
7061:
7058:
7055:
7053:
7049:
7043:
7042:
7038:
7036:
7035:
7031:
7029:
7028:
7024:
7023:
7021:
7019:
7015:
7008:
7007:
7003:
7001:
7000:
6999:Yixianopterus
6996:
6994:
6993:
6989:
6987:
6986:
6982:
6981:
6978:
6975:
6973:
6969:
6963:
6962:
6958:
6956:
6955:
6951:
6950:
6948:
6946:
6942:
6938:
6931:
6930:
6926:
6925:
6923:
6920:
6915:
6904:
6903:
6899:
6896:
6895:
6891:
6889:
6888:
6884:
6881:
6880:
6876:
6873:
6872:
6868:
6867:
6865:
6863:
6859:
6853:
6852:
6848:
6846:
6845:
6841:
6839:
6838:
6834:
6833:
6830:
6826:
6823:
6820:
6815:
6792:
6791:
6787:
6785:
6784:
6780:
6778:
6777:
6773:
6771:
6770:
6766:
6765:
6763:
6761:
6760:Nyctosauridae
6757:
6751:
6750:
6746:
6744:
6743:
6739:
6737:
6736:
6732:
6731:
6728:
6725:
6723:
6719:
6712:
6711:
6707:
6705:
6704:
6703:Alamodactylus
6700:
6699:
6696:
6693:
6691:
6687:
6680:
6679:
6675:
6672:
6671:
6667:
6665:
6664:
6660:
6657:
6656:
6652:
6650:
6649:
6645:
6642:
6641:
6637:
6636:
6634:
6632:
6628:
6625:
6623:
6622:Pteranodontia
6619:
6613:
6612:
6608:
6607:
6604:
6600:
6597:
6594:
6589:
6585:
6581:
6576:
6572:
6562:
6557:
6554:
6546:
6520:
6519:
6518:
6517:
6515:
6513:
6509:
6505:
6467:
6466:
6462:
6459:
6458:
6457:Wellnhopterus
6454:
6452:
6451:
6447:
6445:
6444:
6440:
6438:
6437:
6433:
6431:
6430:
6429:Hatzegopteryx
6426:
6424:
6423:
6419:
6417:
6416:
6412:
6410:
6409:
6405:
6403:
6402:
6401:Aralazhdarcho
6398:
6397:
6395:
6393:
6389:
6383:
6382:
6378:
6376:
6375:
6371:
6369:
6368:
6364:
6361:
6360:
6356:
6355:
6353:
6351:
6347:
6340:
6339:
6335:
6332:
6331:
6327:
6325:
6324:
6320:
6317:
6316:
6312:
6310:
6309:
6305:
6302:
6301:
6297:
6296:
6294:
6292:
6288:
6281:
6280:
6276:
6274:
6273:
6272:Radiodactylus
6269:
6266:
6265:
6261:
6260:
6257:
6254:
6252:
6248:
6238:
6237:
6233:
6231:
6230:
6226:
6224:
6223:
6219:
6218:
6216:
6214:
6210:
6203:
6202:
6198:
6195:
6194:
6190:
6188:
6187:
6183:
6181:
6180:
6176:
6173:
6172:
6168:
6165:
6164:
6160:
6157:
6156:
6155:Apatorhamphus
6152:
6151:
6148:
6145:
6143:
6139:
6133:
6132:
6128:
6127:
6124:
6121:
6119:
6115:
6108:
6107:
6103:
6100:
6099:
6095:
6092:
6091:
6087:
6084:
6083:
6079:
6076:
6075:
6071:
6069:
6068:
6064:
6063:
6061:
6058:
6054:
6047:
6046:
6042:
6040:
6039:
6035:
6033:
6032:
6028:
6025:
6024:
6020:
6019:
6016:
6013:
6010:
6005:
5975:
5974:
5970:
5968:
5967:
5963:
5962:
5960:
5958:
5954:
5948:
5947:
5946:Tupandactylus
5943:
5941:
5940:
5936:
5934:
5933:
5929:
5926:
5925:
5921:
5920:
5917:
5914:
5912:
5908:
5902:
5901:
5897:
5894:
5893:
5889:
5886:
5885:
5881:
5878:
5877:
5876:Caupedactylus
5873:
5870:
5869:
5865:
5864:
5861:
5858:
5856:
5852:
5846:
5845:
5841:
5839:
5838:
5834:
5832:
5831:
5827:
5825:
5824:
5820:
5818:
5817:
5813:
5810:
5809:
5805:
5802:
5801:
5797:
5794:
5793:
5789:
5788:
5786:
5784:
5780:
5774:
5773:
5769:
5768:
5765:
5762:
5760:
5756:
5750:
5749:
5748:Caupedactylus
5745:
5743:
5742:
5738:
5737:
5735:
5732:
5731:Caupedactylia
5728:
5725:
5723:
5719:
5712:
5711:
5707:
5705:
5704:
5700:
5698:
5697:
5693:
5690:
5689:
5685:
5682:
5681:
5677:
5675:
5674:
5670:
5667:
5666:
5662:
5659:
5658:
5654:
5651:
5650:
5646:
5643:
5642:
5638:
5637:
5635:
5632:
5628:
5621:
5620:
5616:
5614:
5613:
5609:
5608:
5605:
5602:
5599:
5594:
5591:
5588:
5587:Neoazhdarchia
5583:
5577:
5576:
5572:
5570:
5569:
5565:
5562:
5561:
5557:
5554:
5553:
5549:
5547:
5546:
5542:
5540:
5539:
5535:
5534:
5531:
5528:
5525:
5524:Azhdarchoidea
5520:
5506:
5505:
5501:
5499:
5498:
5494:
5493:
5491:
5489:
5485:
5478:
5477:
5473:
5470:
5469:
5468:Puntanipterus
5465:
5463:
5462:
5458:
5456:
5455:
5451:
5448:
5447:
5443:
5440:
5439:
5435:
5434:
5431:
5428:
5426:
5422:
5418:
5415:
5412:
5407:
5400:
5399:
5395:
5394:
5391:
5388:
5385:
5380:
5376:
5372:
5367:
5363:
5354:
5348:
5322:
5321:
5320:
5319:
5317:
5315:
5311:
5304:
5303:
5299:
5298:
5295:
5292:
5290:
5286:
5282:
5256:
5255:
5251:
5249:
5248:
5244:
5242:
5241:
5237:
5235:
5234:
5233:Beipiaopterus
5230:
5229:
5227:
5225:
5221:
5215:
5214:
5210:
5209:
5206:
5203:
5201:
5197:
5191:
5190:
5189:Tacuadactylus
5186:
5184:
5183:
5179:
5177:
5176:
5172:
5169:
5168:
5164:
5162:
5161:
5157:
5156:
5154:
5152:
5148:
5142:
5141:
5137:
5135:
5134:
5130:
5129:
5127:
5124:
5120:
5114:
5113:
5109:
5107:
5106:
5102:
5100:
5099:
5095:
5093:
5092:
5088:
5086:
5085:
5081:
5079:
5078:
5074:
5072:
5071:
5070:Forfexopterus
5067:
5065:
5064:
5063:Elanodactylus
5060:
5058:
5057:
5056:Cratonopterus
5053:
5051:
5050:
5049:Cathayopterus
5046:
5044:
5043:
5039:
5036:
5035:
5034:Ardeadactylus
5031:
5030:
5027:
5024:
5021:
5016:
5001:
5000:
4996:
4994:
4993:
4989:
4986:
4985:
4984:Ardeadactylus
4981:
4980:
4978:
4976:
4972:
4965:
4964:
4960:
4958:
4957:
4953:
4952:
4949:
4946:
4944:
4940:
4933:
4932:
4928:
4925:
4924:
4920:
4917:
4916:
4912:
4910:
4909:
4908:Cycnorhamphus
4905:
4902:
4901:
4897:
4896:
4894:
4892:
4888:
4884:
4881:
4878:
4873:
4867:
4866:
4865:Pterodactylus
4862:
4860:
4859:
4858:Diopecephalus
4855:
4854:
4851:
4848:
4845:
4840:
4829:
4828:
4824:
4821:
4820:
4816:
4814:
4813:
4809:
4806:
4805:
4801:
4800:
4798:
4796:
4792:
4786:
4785:
4784:Prejanopterus
4781:
4780:
4777:
4773:
4770:
4767:
4762:
4758:
4754:
4749:
4745:
4736:
4731:
4694:
4693:
4692:
4691:
4689:
4687:
4683:
4677:
4676:
4672:
4669:
4668:
4664:
4662:
4661:
4657:
4655:
4654:
4650:
4648:
4647:
4643:
4641:
4640:
4636:
4633:
4632:
4631:Herbstosaurus
4628:
4626:
4625:
4624:Eurolimnornis
4621:
4619:
4618:
4617:Dermodactylus
4614:
4613:
4610:
4607:
4605:
4601:
4591:
4590:
4586:
4583:
4582:
4578:
4575:
4574:
4570:
4568:
4567:
4563:
4561:
4560:
4556:
4555:
4553:
4551:
4547:
4541:
4540:
4536:
4534:
4533:
4529:
4526:
4525:
4521:
4518:
4517:
4513:
4511:
4510:
4506:
4505:
4503:
4501:
4497:
4491:
4490:
4486:
4485:
4482:
4479:
4477:
4473:
4470:
4467:
4466:Caelidracones
4462:
4456:
4455:
4451:
4448:
4447:
4443:
4442:
4439:
4436:
4433:
4428:
4410:
4409:
4408:Wukongopterus
4405:
4402:
4401:
4397:
4395:
4394:
4393:Darwinopterus
4390:
4388:
4387:
4386:Cuspicephalus
4383:
4380:
4379:
4375:
4374:
4372:
4370:
4366:
4360:
4359:
4355:
4354:
4351:
4348:
4346:
4342:
4336:
4335:
4334:Pterorhynchus
4331:
4329:
4328:
4324:
4323:
4320:
4317:
4315:
4311:
4304:
4303:
4299:
4297:
4296:
4292:
4290:
4289:
4285:
4284:
4281:
4277:
4274:
4271:
4266:
4259:
4258:
4254:
4251:
4250:
4246:
4245:
4242:
4239:
4236:
4231:
4227:
4223:
4218:
4214:
4205:
4201:
4198:
4190:
4144:
4143:
4142:
4141:
4139:
4137:
4133:
4126:
4125:
4121:
4119:
4118:
4117:Scaphognathus
4114:
4112:
4111:
4107:
4105:
4104:
4100:
4098:
4097:
4093:
4092:
4090:
4087:
4083:
4080:
4077:
4073:
4055:
4054:
4050:
4047:
4046:
4042:
4039:
4038:
4034:
4033:
4031:
4029:
4025:
4019:
4018:
4014:
4011:
4010:
4006:
4003:
4002:
3998:
3996:
3995:
3991:
3989:
3988:
3984:
3983:
3981:
3979:
3975:
3972:
3970:
3966:
3960:
3959:
3955:
3953:
3952:
3948:
3946:
3945:
3941:
3939:
3938:
3934:
3932:
3931:
3927:
3925:
3924:
3920:
3918:
3917:
3913:
3912:
3909:
3906:
3904:
3900:
3893:
3892:
3888:
3885:
3884:
3880:
3878:
3877:
3873:
3870:
3869:
3865:
3864:
3861:
3858:
3856:
3852:
3849:
3846:
3841:
3831:
3830:
3826:
3824:
3823:
3819:
3818:
3816:
3814:
3810:
3806:
3803:
3800:
3795:
3788:
3787:
3786:Eudimorphodon
3783:
3782:
3779:
3776:
3772:
3767:
3756:
3755:
3751:
3748:
3747:
3746:Peteinosaurus
3743:
3740:
3739:
3735:
3733:
3732:
3728:
3726:
3725:
3721:
3718:
3717:
3713:
3712:
3710:
3708:
3704:
3698:
3697:
3696:Herbstosaurus
3693:
3692:
3689:
3685:
3682:
3679:
3674:
3667:
3666:
3665:Peteinosaurus
3662:
3661:
3658:
3655:
3651:
3646:
3628:
3627:
3626:Eudimorphodon
3623:
3620:
3619:
3615:
3614:
3612:
3610:
3606:
3599:
3598:
3594:
3593:
3590:
3587:
3585:
3581:
3575:
3574:
3570:
3568:
3567:
3563:
3561:
3560:
3556:
3553:
3552:
3548:
3547:
3545:
3543:
3539:
3536:
3534:
3530:
3523:
3522:
3521:Preondactylia
3518:
3515:
3514:
3513:Peteinosaurus
3510:
3507:
3506:
3502:
3501:
3498:
3495:
3492:
3491:Eopterosauria
3487:
3473:
3472:
3468:
3466:
3465:
3461:
3458:
3457:
3453:
3452:
3450:
3448:
3444:
3437:
3436:
3432:
3430:
3429:
3425:
3422:
3421:
3417:
3416:
3413:
3410:
3407:
3403:
3397:
3396:
3395:Preondactylus
3392:
3390:
3389:
3385:
3384:
3382:
3380:
3379:Preondactylia
3376:
3370:
3369:
3365:
3364:
3361:
3357:
3354:
3351:
3346:
3342:
3338:
3333:
3329:
3308:
3307:
3306:
3305:
3303:
3301:
3297:
3289:
3285:
3284:
3283:
3282:
3279:
3276:
3274:
3270:
3266:
3258:
3254:
3251:
3249:
3245:
3242:
3240:
3236:
3233:
3231:
3227:
3225:
3221:
3219:
3215:
3214:
3209:
3205:
3201:
3194:
3189:
3187:
3182:
3180:
3175:
3174:
3171:
3163:
3159:
3155:
3151:
3147:
3143:
3139:
3135:
3131:
3127:
3123:
3119:
3116:
3115:
3111:
3110:
3090:
3086:
3080:
3064:
3060:
3056:
3050:
3034:
3030:
3026:
3019:
3011:
3007:
3000:
2992:
2988:
2981:
2973:
2969:
2964:
2959:
2955:
2951:
2947:
2943:
2939:
2932:
2916:
2915:
2910:
2903:
2895:
2891:
2887:
2883:
2879:
2875:
2871:
2867:
2860:
2853:
2851:
2842:
2838:
2831:
2824:
2816:
2812:
2807:
2802:
2798:
2794:
2790:
2786:
2782:
2780:
2771:
2763:
2759:
2755:
2751:
2747:
2743:
2739:
2735:
2728:
2726:
2717:
2713:
2708:
2703:
2698:
2693:
2689:
2685:
2681:
2674:
2666:
2662:
2658:
2654:
2650:
2643:
2635:
2631:
2626:
2621:
2616:
2611:
2607:
2603:
2599:
2592:
2584:
2577:
2575:
2573:
2564:
2560:
2556:
2552:
2548:
2544:
2537:
2529:
2525:
2521:
2519:9780691232218
2515:
2511:
2507:
2503:
2496:
2488:
2484:
2479:
2474:
2470:
2466:
2462:
2458:
2454:
2452:
2443:
2441:
2432:
2428:
2423:
2418:
2413:
2408:
2404:
2400:
2396:
2392:
2388:
2381:
2379:
2370:
2368:0-8018-6763-0
2364:
2359:
2358:
2349:
2341:
2335:
2331:
2327:
2326:
2319:
2311:
2307:
2303:
2299:
2295:
2291:
2287:
2283:
2276:
2268:
2264:
2260:
2256:
2252:
2248:
2241:
2234:
2226:
2222:
2218:
2211:
2209:
2207:
2198:
2194:
2190:
2186:
2182:
2178:
2174:
2170:
2166:
2162:
2161:
2153:
2146:
2138:
2134:
2130:
2126:
2122:
2118:
2114:
2107:
2105:
2096:
2092:
2088:
2084:
2080:
2076:
2072:
2068:
2064:
2060:
2053:
2045:
2041:
2037:
2033:
2029:
2025:
2021:
2017:
2013:
2009:
2002:
2000:
1991:
1987:
1982:
1977:
1972:
1967:
1963:
1959:
1955:
1951:
1947:
1940:
1938:
1936:
1934:
1932:
1916:
1915:Berkeley News
1912:
1905:
1897:
1893:
1889:
1885:
1881:
1877:
1873:
1866:
1858:
1854:
1850:
1846:
1841:
1836:
1832:
1828:
1825:(sup1): 142.
1824:
1820:
1816:
1814:
1805:
1803:
1801:
1799:
1797:
1795:
1790:
1781:
1778:
1776:
1773:
1772:
1766:
1764:
1760:
1759:Royal Society
1756:
1752:
1747:
1745:
1740:
1736:
1732:
1728:
1724:
1720:
1719:AeroVironment
1716:
1711:
1709:
1705:
1701:
1693:
1692:Royal Society
1689:
1684:
1675:
1672:
1667:
1663:
1659:
1654:
1652:
1648:
1643:
1639:
1634:
1630:
1621:
1617:
1613:
1609:
1607:
1602:
1598:
1594:
1590:
1586:
1582:
1578:
1570:
1565:
1556:
1554:
1550:
1546:
1544:
1540:
1536:
1532:
1527:
1523:
1519:
1515:
1511:
1505:
1503:
1499:
1496:
1492:
1491:marabou stork
1488:
1484:
1480:
1472:
1467:
1458:
1456:
1455:
1454:Wellnhopterus
1449:
1445:
1441:
1437:
1436:
1431:
1427:
1413:
1412:
1404:
1403:
1395:
1394:
1386:
1385:
1377:
1376:
1368:
1367:
1359:
1358:
1350:
1349:
1346:
1345:
1344:
1337:
1336:
1333:
1332:
1329:
1327:
1323:
1322:
1315:
1314:
1311:
1310:
1307:
1306:
1305:
1298:
1297:
1291:
1290:
1287:
1286:
1283:
1282:
1281:
1274:
1273:
1267:
1266:
1263:
1262:
1259:
1254:
1253:
1247:
1246:
1243:
1242:
1239:
1238:
1237:
1230:
1229:
1222:
1218:
1217:
1214:
1213:
1210:
1209:
1208:
1201:
1200:
1194:
1193:
1190:
1189:
1181:
1180:
1172:
1171:
1163:
1162:
1159:
1158:
1157:
1150:
1149:
1146:
1145:
1142:
1141:
1140:
1133:
1132:
1126:
1125:
1122:
1121:
1118:
1117:
1116:
1109:
1108:
1102:
1101:
1098:
1097:
1094:
1093:
1092:
1085:
1084:
1077:
1073:
1072:
1066:
1065:
1062:
1061:
1053:
1052:
1044:
1043:
1035:
1034:
1031:
1030:
1029:
1022:
1021:
1018:
1017:
1014:
1013:
1012:
1005:
1004:
998:
997:
994:
993:
985:
984:
981:
980:
979:
972:
971:
968:
967:
964:
962:
958:
957:
950:
949:
943:
942:
935:
931:
930:
927:
926:
918:
917:
909:
908:
905:
904:
903:
896:
895:
892:
891:
888:
887:
886:
879:
878:
872:
871:
868:
867:
864:
863:
862:
855:
854:
847:
843:
842:
836:
835:
828:
827:Neoazhdarchia
824:
823:
819:
817:
813:
808:
806:
802:
795:
794:
789:
784:
775:
773:
772:
771:Wellnhopterus
766:
762:
750:
745:
741:
739:
735:
730:
728:
722:
715:
710:
706:
703:
695:
691:
686:
681:
666:
664:
660:
659:
654:
650:
645:
643:
639:
635:
631:
627:
623:
619:
615:
614:Hatzegopteryx
611:
607:
603:
602:Hatzegopteryx
599:
598:
593:
589:
588:
587:Hatzegopteryx
583:
579:
575:
571:
567:
563:
559:
554:
552:
548:
544:
540:
531:
526:
522:
520:
516:
512:
508:
504:
503:specific name
500:
496:
492:
488:
484:
483:
478:
474:
469:
467:
462:
458:
454:
450:
447:
446:Maastrichtian
443:
439:
432:
427:
418:
416:
412:
411:
406:
405:Jack Northrop
402:
401:
396:
392:
388:
387:Maastrichtian
385:
381:
378:
372:
325:
324:
310:
309:
302:
301:
299:
294:
287:
286:
278:
275:
271:
266:
261:
260:
253:
250:
249:
246:
240:
237:
236:
233:
227:
224:
223:
220:
214:
211:
210:
207:
201:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
162:
157:
153:
150:
143:
138:
134:
129:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
54:
47:
46:Maastrichtian
43:
37:
33:
30:
19:
8015:Quetzalcoatl
7957:Paleontology
7818:
7782:
7728:
7723:Ludodactylus
7721:
7714:
7707:
7699:
7691:
7684:
7664:
7656:
7651:Nicorhynchus
7649:
7642:
7635:
7614:
7607:
7599:
7591:
7583:
7575:
7567:
7537:
7530:
7522:Hamipteridae
7508:
7500:
7466:
7458:
7450:
7430:
7422:
7417:Aetodactylus
7415:
7380:
7372:
7365:
7357:
7351:Ludodactylus
7349:
7341:
7333:
7325:
7317:
7309:
7303:Aetodactylus
7301:
7280:
7272:
7264:
7256:
7248:
7241:
7234:
7191:
7184:
7160:
7153:
7146:
7139:
7113:Liaoxipterus
7111:
7104:
7084:
7077:
7070:
7063:
7041:Mimodactylus
7039:
7032:
7025:
7004:
6997:
6990:
6983:
6959:
6952:
6927:
6919:Lanceodontia
6900:
6892:
6887:Lonchodectes
6885:
6877:
6869:
6849:
6842:
6835:
6788:
6781:
6774:
6767:
6747:
6740:
6733:
6708:
6701:
6676:
6668:
6661:
6655:Ornithostoma
6653:
6646:
6638:
6609:
6549:
6463:
6455:
6448:
6442:
6441:
6434:
6427:
6422:Eurazhdarcho
6420:
6413:
6406:
6399:
6379:
6372:
6365:
6357:
6350:Azhdarchinae
6336:
6328:
6321:
6313:
6306:
6298:
6291:Azhdarchidae
6277:
6270:
6262:
6234:
6227:
6220:
6199:
6191:
6186:Meilifeilong
6184:
6177:
6169:
6161:
6153:
6129:
6104:
6096:
6088:
6080:
6072:
6065:
6043:
6036:
6029:
6021:
5971:
5964:
5944:
5937:
5930:
5922:
5898:
5890:
5882:
5874:
5866:
5842:
5835:
5828:
5823:Huaxiapterus
5821:
5814:
5808:Eopteranodon
5806:
5798:
5792:Afrotapejara
5790:
5783:Sinopterinae
5772:Afrotapejara
5770:
5746:
5739:
5708:
5701:
5694:
5686:
5678:
5671:
5663:
5655:
5647:
5639:
5617:
5612:Bennettazhia
5610:
5575:Ornithostoma
5573:
5566:
5558:
5550:
5543:
5536:
5502:
5495:
5474:
5466:
5461:Ordosipterus
5459:
5452:
5444:
5436:
5411:Tapejaroidea
5396:
5300:
5254:Pterodaustro
5252:
5245:
5238:
5231:
5211:
5187:
5180:
5173:
5167:Huanhepterus
5165:
5160:Gnathosaurus
5158:
5140:Moganopterus
5138:
5131:
5112:Pterofiltrus
5110:
5105:Petrodactyle
5103:
5096:
5091:Liaodactylus
5089:
5084:Kepodactylus
5082:
5075:
5068:
5061:
5054:
5047:
5040:
5032:
4999:Huanhepterus
4997:
4990:
4982:
4961:
4956:Aerodactylus
4954:
4929:
4923:Petrodactyle
4921:
4913:
4906:
4898:
4863:
4856:
4825:
4817:
4810:
4802:
4782:
4673:
4665:
4658:
4651:
4644:
4639:Kryptodrakon
4637:
4629:
4622:
4615:
4587:
4581:Jeholopterus
4579:
4571:
4564:
4557:
4537:
4530:
4524:Jeholopterus
4522:
4514:
4509:Anurognathus
4507:
4489:Mesadactylus
4487:
4452:
4444:
4406:
4398:
4391:
4384:
4376:
4356:
4332:
4325:
4314:Darwinoptera
4300:
4293:
4286:
4255:
4247:
4193:
4122:
4115:
4108:
4101:
4094:
4051:
4045:Nesodactylus
4043:
4037:Cacibupteryx
4035:
4017:Sericipterus
4015:
4007:
3999:
3992:
3985:
3956:
3949:
3944:Nesodactylus
3942:
3935:
3928:
3923:Cacibupteryx
3921:
3916:Bellubrunnus
3914:
3889:
3881:
3874:
3866:
3827:
3820:
3784:
3771:Lonchognatha
3752:
3744:
3736:
3729:
3722:
3714:
3694:
3663:
3624:
3616:
3595:
3571:
3564:
3559:Pachagnathus
3557:
3549:
3519:
3511:
3505:Austriadraco
3503:
3469:
3464:Austriadraco
3462:
3454:
3433:
3426:
3418:
3393:
3386:
3366:
3252:
3243:
3234:
3161:
3149:
3137:
3125:
3113:
3093:. Retrieved
3088:
3079:
3067:. Retrieved
3063:the original
3058:
3049:
3037:. Retrieved
3033:the original
3028:
3018:
3009:
2999:
2990:
2980:
2945:
2941:
2931:
2919:. Retrieved
2912:
2902:
2869:
2865:
2840:
2836:
2823:
2788:
2784:
2778:
2770:
2737:
2734:Paleobiology
2733:
2687:
2684:PLOS Biology
2683:
2673:
2656:
2652:
2648:
2642:
2605:
2602:PLOS Biology
2601:
2591:
2582:
2546:
2542:
2536:
2501:
2495:
2460:
2456:
2450:
2394:
2390:
2356:
2348:
2324:
2318:
2285:
2281:
2275:
2250:
2246:
2233:
2224:
2220:
2164:
2158:
2145:
2120:
2116:
2112:
2062:
2058:
2052:
2011:
2007:
1956:(5): e2271.
1953:
1949:
1918:. Retrieved
1914:
1904:
1879:
1875:
1871:
1865:
1822:
1818:
1812:
1751:Q. northropi
1750:
1748:
1739:Q. northropi
1738:
1731:Q. northropi
1730:
1722:
1712:
1703:
1699:
1697:
1670:
1665:
1661:
1658:Q. northropi
1657:
1655:
1651:Kori bustard
1642:Q. northropi
1641:
1638:Q. northropi
1637:
1633:Q. northropi
1632:
1625:
1615:
1601:Q. northropi
1600:
1584:
1576:
1574:
1568:
1552:
1548:
1547:
1538:
1534:
1526:Darren Naish
1513:
1506:
1501:
1486:
1478:
1476:
1470:
1452:
1447:
1443:
1439:
1433:
1425:
1424:
1421:Paleobiology
1342:
1341:
1340:
1320:
1319:
1318:
1302:
1301:
1278:
1277:
1257:
1234:
1233:
1221:Azhdarchidae
1205:
1204:
1154:
1153:
1137:
1136:
1113:
1112:
1089:
1088:
1026:
1025:
1009:
1008:
976:
975:
954:
953:
900:
899:
883:
882:
859:
858:
815:
809:
805:Azhdarchidae
800:
799:
791:
787:
769:
764:
760:
758:
748:
737:
733:
731:
726:
723:
719:
714:Q. northropi
713:
701:
699:
693:
692:(green) and
690:Q. northropi
689:
656:
652:
646:
642:Q. northropi
641:
637:
634:Q. northropi
633:
629:
626:Q. northropi
625:
621:
617:
613:
609:
605:
601:
597:nomen dubium
595:
592:Q. northropi
591:
585:
578:Q. northropi
577:
573:
570:Q. northropi
569:
561:
558:Q. northropi
557:
555:
550:
547:Q. northropi
546:
538:
535:
529:
518:
499:Quetzalcoatl
490:
480:
470:
437:
435:
415:Q. northropi
414:
409:
408:
400:Q. northropi
399:
398:
395:type species
391:Quetzalcoatl
322:
321:
320:
307:
306:
290:Lawson, 1975
284:
283:
274:Type species
258:
257:
232:Azhdarchidae
141:
35:
29:
7985:Azhdarchids
7884:iNaturalist
7843:Wikispecies
7609:Thapunngaka
7569:Amblydectes
7488:Anhangueria
7452:Aussiedraco
7186:Boreopterus
6961:Lonchodraco
6894:Lonchodraco
6837:Aussiedraco
6783:Nyctosaurus
6640:Bogolubovia
6308:Bogolubovia
6179:Lacusovagus
6171:Eoazhdarcho
6131:Eoazhdarcho
6090:Leptostomia
6082:Keresdrakon
5957:Caiuajarina
5924:Bakonydraco
5892:Lacusovagus
5884:Keresdrakon
5855:Tapejarinae
5816:Huaxiadraco
5800:Bakonydraco
5759:Tapejaridae
5688:Leptostomia
5680:Lacusovagus
5673:Kariridraco
5619:Keresdrakon
5560:Leptostomia
5552:Keresdrakon
5213:Ctenochasma
5175:Lusognathus
5098:Otogopterus
4753:Lophocratia
4686:Lophocratia
4653:Pangupterus
4589:Sinomacrops
3930:Dorygnathus
3799:Novialoidea
3754:Rhamphinion
3731:Dimorphodon
3406:Caviramidae
3350:Pterosauria
3337:Pterosauria
3300:Pterosauria
3239:Archosauria
3200:Pterosauria
3039:November 1,
2921:November 1,
2843:(2): 18–24.
2690:(8): e204.
1920:December 7,
1872:Alamosaurus
1656:Studies of
1589:ornithopter
1502:Alamosaurus
1483:fish-eating
1440:Alamosaurus
1435:Alamosaurus
810:Below is a
763:shows that
669:Description
600:), perhaps
566:Mark Witton
515:flying wing
238:Subfamily:
206:Pterosauria
51:68–66
7979:Categories
7686:Anhanguera
7577:Ferrodraco
7532:Hamipterus
7460:Barbosania
7335:Ferrodraco
7327:Draigwenia
7162:Hamipterus
7141:Barbosania
7086:Nurhachius
6954:Ikrandraco
6929:Draigwenia
6879:Ikrandraco
6844:Serradraco
6790:Volgadraco
6678:Volgadraco
6670:Tethydraco
6663:Pteranodon
6521:see below↓
6415:Cryodrakon
6338:Volgadraco
6330:Tethydraco
6229:Jidapterus
6193:Microtuban
6031:Microtuban
5932:Europejara
5911:Tapejarini
5900:Vectidraco
5837:Sinopterus
5454:Noripterus
5323:see below↓
5247:Gegepterus
5240:Eosipterus
4695:see below↓
4675:Wenupteryx
4566:Cascocauda
4145:see below↓
3573:Yelaphomte
3309:see below↓
3230:Sauropsida
3095:August 20,
2942:PNAS Nexus
2549:(3–4): 1.
1786:References
1755:South Bank
1688:South Bank
1662:Q. lawsoni
1606:flightless
1553:Q. lawsoni
1500:, such as
1495:titanosaur
1258:TMM 42489
761:Q. lawsoni
749:Q. lawsoni
738:Q. lawsoni
734:Q. lawsoni
694:Q. lawsoni
678:See also:
638:Q. lawsoni
630:Q. lawsoni
618:Q. lawsoni
551:Q. lawsoni
530:Q. lawsoni
436:The first
410:Q. lawsoni
377:azhdarchid
308:Q. lawsoni
212:Suborder:
7637:Aerodraco
7079:Luchibang
7027:Haopterus
6749:Simurghia
6742:Epapatelo
6710:Cretornis
6648:Dawndraco
6374:Azhdarcho
6367:Albadraco
6359:Aerotitan
6279:Alanqidae
6057:Alanqidae
6023:Cretornis
5973:Torukjara
5966:Caiuajara
5703:Tupuxuara
5641:Aerotitan
5545:Inabtanin
5133:Feilongus
4532:Luopterus
4249:Allkaruen
3876:Klobiodon
3716:Allkaruen
3551:Caviramus
3428:Caviramus
3216:Kingdom:
3059:avinc.com
2815:245009158
2528:249332375
2487:245125427
2267:130751879
1896:130280606
1857:245125409
1849:0272-4634
1593:autopilot
1571:in flight
1510:vertebrae
1498:sauropods
812:cladogram
444:from the
380:pterosaur
179:Kingdom:
173:Eukaryota
140:Restored
7828:Wikidata
7709:Guidraco
7593:Mythunga
7585:Haliskia
7359:Mythunga
7343:Guidraco
7274:Haliskia
6851:Unwindia
6201:Xericeps
6106:Xericeps
6038:Xericeps
5939:Tapejara
5710:Xericeps
5665:Banguela
5438:Banguela
4667:Samrukia
4327:Ceoptera
3224:Chordata
3222:Phylum:
3218:Animalia
3069:July 28,
2972:36712794
2894:85679231
2762:88434056
2716:17676976
2634:29534059
2563:84617119
2431:21085624
2391:PLOS ONE
2310:24964287
2227:: 79–81.
2197:15423666
2189:12061403
2095:44400704
2087:17755159
2044:46396417
2036:17745279
1990:18509539
1950:PLOS ONE
1769:See also
1690:for the
1543:ungulate
1518:skimmers
663:dinosaur
582:Romanian
487:holotype
466:wingspan
225:Family:
193:Chordata
189:Phylum:
183:Animalia
169:Domain:
144:skeleton
7943:Portals
7915:4946961
7902:1042058
7876:4818447
7863:4532239
7834:Q131205
6735:Alcione
5844:Wightia
3228:Class:
3162:YouTube
3150:YouTube
3138:YouTube
3126:YouTube
2963:9802081
2874:Bibcode
2793:Bibcode
2742:Bibcode
2707:1925135
2625:5849296
2465:Bibcode
2422:2981443
2399:Bibcode
2290:Bibcode
2169:Bibcode
2125:Bibcode
2067:Bibcode
2059:Science
2016:Bibcode
2008:Science
1981:2386974
1958:Bibcode
1827:Bibcode
1597:soaring
1461:Feeding
1430:Lancian
1223:
1078:
936:
848:
829:
541:sp. by
505:honors
482:Science
475:of the
431:humerus
251:Genus:
199:Order:
120:↓
7889:523016
6300:Alanqa
6067:Alanqa
5649:Alanqa
4257:Sordes
4124:Sordes
2970:
2960:
2892:
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2760:
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2632:
2622:
2561:
2526:
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2093:
2085:
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1988:
1978:
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1717:) and
1559:Flight
1531:storks
1438:. The
1219:
1074:
932:
844:
825:
696:(blue)
610:et al.
574:et al.
501:. The
393:. The
267:, 1975
265:Lawson
7928:38504
7897:IRMNG
5398:Piksi
3994:Dearc
3253:Clade
3244:Clade
3235:Clade
2917:. NPR
2890:S2CID
2862:(PDF)
2833:(PDF)
2811:S2CID
2758:S2CID
2559:S2CID
2524:S2CID
2483:S2CID
2306:JSTOR
2263:S2CID
2243:(PDF)
2193:S2CID
2155:(PDF)
2091:S2CID
2040:S2CID
1892:S2CID
1853:S2CID
1721:used
1715:DARPA
755:Skull
495:Aztec
442:Texas
7871:GBIF
3286:see
3097:2020
3071:2011
3041:2017
2968:PMID
2923:2017
2712:PMID
2630:PMID
2514:ISBN
2427:PMID
2363:ISBN
2334:ISBN
2185:PMID
2083:PMID
2032:PMID
1986:PMID
1922:2023
1845:ISSN
1660:and
1522:drag
674:Size
616:and
59:PreꞒ
7858:EoL
2958:PMC
2950:doi
2882:doi
2801:doi
2750:doi
2702:PMC
2692:doi
2661:doi
2620:PMC
2610:doi
2551:doi
2547:103
2506:doi
2473:doi
2417:PMC
2407:doi
2298:doi
2286:244
2255:doi
2177:doi
2133:doi
2075:doi
2063:188
2024:doi
2012:187
1976:PMC
1966:doi
1884:doi
1835:doi
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451:at
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