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Spread-winged skipper

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69: 42: 215: 207: 269:, which their ancestors would have found most easy to reach by crossing the central Atlantic Ocean, rather than taking the longer route via Europe and North America or the Pacific Ocean. It is also possible that the group originated in the 235:
Spread-winged skippers bask with their wings held wide open. The wings are held closed when they are at rest. They are usually brown, black, or checkered, but some are brilliantly coloured. Some have long tailed hind wings.
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Males maintain territories and frequently perch on overhanging branches and tree trunks. They do not visit flowers for sustenance, but rather drink fluids from
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It still is the second-largest subfamily of skipper butterflies, although of its over 1,000 species a considerable number are no longer in that classification.
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Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; Hallwachs, Winnie; Janzen, Daniel H.; Grishin, Nick V. (2019-03-26).
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A large and successful subfamily, the spread-winged skippers are found essentially worldwide, except in very cold or
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has changed considerably in recent years with the most recent review being in 2019.
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Most of the more advanced tribes, on the other hand, are very diverse in the
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Catalogue of the Butterflies of the United States and Canada
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of the Pyrginae in their modern circumscription are, in
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and subsequently dispersed eastwards to Africa via the
464:. Adventure Publications, Inc., Cambridge, MN, USA. 751: 537:Reference photographs: Skippers of North America 258:and subsequently extended their range into more 187:(Hesperiidae). The subfamily was established by 393:Proceedings of the National Academy of Sciences 386: 201: 373: 371: 368: 40: 430: 412: 254:regions. They probably originated in the 191:in 1878. Their delimitation and internal 486:Butterflies through Binoculars, The West 213: 205: 14: 752: 542:Butterflies and Moths of North America 500:Life Histories of Cascadia Butterflies 491:Guppy, Crispin S. and Shepard, Jon H. 556: 555: 498:James, David G. and Nunnallee, David 685:1b740b6b-9d4b-4885-94e0-8ae70695b16a 24: 478: 354:- now placed within the subfamily 345:- now placed within the subfamily 336:- now placed within the subfamily 327:- now placed within the subfamily 25: 776: 520: 765:Taxa named by Hermann Burmeister 318: 67: 493:Butterflies of British Columbia 380: 275:Intertropical Convergence Zone 13: 1: 451: 462:Butterflies of the Carolinas 361: 202:Description and distribution 7: 514:The Butterflies of Cascadia 161: 10: 781: 564: 280: 210:Typical resting positions 160: 153: 64:Scientific classification 62: 48: 39: 34: 27:Subfamily of butterflies 535:Cirrus Digital Imaging 414:10.1073/pnas.1821304116 35:Spread-winged skippers 547:Butterflies of America 232: 211: 174:spread-winged skippers 512:Pyle, Robert Michael 247:, and rotting fruit. 217: 209: 680:Fauna Europaea (new) 377:Daniels (2003): p.12 172:, commonly known as 484:Glassberg, Jeffrey 405:2019PNAS..116.6232L 527:Subfamily Pyrginae 233: 212: 189:Hermann Burmeister 747: 746: 558:Taxon identifiers 505:Pelham, Jonathan 473: 458:Daniels, Jaret C. 399:(13): 6232–6237. 182:skipper butterfly 167: 166: 149: 56:Ngorongoro Crater 16:(Redirected from 772: 740: 739: 727: 726: 724:NHMSYS0000504010 714: 713: 701: 700: 688: 687: 675: 674: 662: 661: 649: 648: 636: 635: 623: 622: 613: 612: 600: 599: 598: 585: 584: 583: 553: 552: 465: 459: 445: 444: 434: 416: 384: 378: 375: 144: 72: 71: 44: 32: 31: 21: 780: 779: 775: 774: 773: 771: 770: 769: 750: 749: 748: 743: 735: 730: 722: 717: 709: 704: 696: 691: 683: 678: 670: 665: 657: 652: 644: 639: 631: 626: 618: 616: 608: 603: 594: 593: 588: 579: 578: 573: 560: 523: 481: 479:Further reading 457: 454: 449: 448: 385: 381: 376: 369: 364: 325:Celaenorrhinini 321: 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267:Neotropics 262:regions. 146:Burmeister 101:Arthropoda 423:0027-8424 362:Footnotes 343:Tagiadini 338:Eudaminae 285:The four 260:temperate 178:subfamily 155:Diversity 87:Kingdom: 81:Eukaryota 760:Pyrginae 659:59285139 628:BugGuide 617:BioLib: 610:Pyrginae 596:Pyrginae 581:Q1306572 575:Wikidata 566:Pyrginae 460:(2003): 441:30877254 334:Eudamini 308:Erynnini 176:, are a 170:Pyrginae 162:4 tribes 141:Pyrginae 127:Family: 97:Phylum: 91:Animalia 77:Domain: 18:Pyrginae 432:6442542 401:Bibcode 313:Pyrgini 256:tropics 245:carrion 229:Pyrgini 225:Florida 180:of the 117:Order: 111:Insecta 107:Class: 54:in the 711:694116 672:440742 516:(2002) 509:(2008) 502:(2011) 495:(2001) 488:(2001) 469:  439:  429:  421:  287:tribes 281:Tribes 252:desert 185:family 148:, 1878 737:40100 698:51312 646:93MZN 633:12853 620:51218 529:from 223:from 732:NCBI 706:ITIS 467:ISBN 437:PMID 419:ISSN 241:dung 719:NBN 654:EoL 641:CoL 605:AFD 427:PMC 409:doi 397:116 756:: 734:: 721:: 708:: 695:: 682:: 669:: 656:: 643:: 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Index

Pyrginae

Spialia mafa
Ngorongoro Crater
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Lepidoptera
Hesperiidae
Pyrginae
Burmeister
Diversity
4 tribes
subfamily
skipper butterfly
family
Hermann Burmeister
systematics


Pyrgus oileus
Florida
Pyrgini
dung
carrion
desert
tropics

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