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269:, which their ancestors would have found most easy to reach by crossing the central Atlantic Ocean, rather than taking the longer route via Europe and North America or the Pacific Ocean. It is also possible that the group originated in the
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Spread-winged skippers bask with their wings held wide open. The wings are held closed when they are at rest. They are usually brown, black, or checkered, but some are brilliantly coloured. Some have long tailed hind wings.
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Males maintain territories and frequently perch on overhanging branches and tree trunks. They do not visit flowers for sustenance, but rather drink fluids from
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It still is the second-largest subfamily of skipper butterflies, although of its over 1,000 species a considerable number are no longer in that classification.
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Li, Wenlin; Cong, Qian; Shen, Jinhui; Zhang, Jing; Hallwachs, Winnie; Janzen, Daniel H.; Grishin, Nick V. (2019-03-26).
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A large and successful subfamily, the spread-winged skippers are found essentially worldwide, except in very cold or
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has changed considerably in recent years with the most recent review being in 2019.
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389:"Genomes of skipper butterflies reveal extensive convergence of wing patterns"
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Most of the more advanced tribes, on the other hand, are very diverse in the
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Catalogue of the
Butterflies of the United States and Canada
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of the
Pyrginae in their modern circumscription are, in
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and subsequently dispersed eastwards to Africa via the
464:. Adventure Publications, Inc., Cambridge, MN, USA.
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537:Reference photographs: Skippers of North America
258:and subsequently extended their range into more
187:(Hesperiidae). The subfamily was established by
393:Proceedings of the National Academy of Sciences
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254:regions. They probably originated in the
191:in 1878. Their delimitation and internal
486:Butterflies through Binoculars, The West
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542:Butterflies and Moths of North America
500:Life Histories of Cascadia Butterflies
491:Guppy, Crispin S. and Shepard, Jon H.
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498:James, David G. and Nunnallee, David
685:1b740b6b-9d4b-4885-94e0-8ae70695b16a
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354:- now placed within the subfamily
345:- now placed within the subfamily
336:- now placed within the subfamily
327:- now placed within the subfamily
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765:Taxa named by Hermann Burmeister
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493:Butterflies of British Columbia
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275:Intertropical Convergence Zone
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462:Butterflies of the Carolinas
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202:Description and distribution
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514:The Butterflies of Cascadia
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210:Typical resting positions
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64:Scientific classification
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27:Subfamily of butterflies
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414:10.1073/pnas.1821304116
35:Spread-winged skippers
547:Butterflies of America
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174:spread-winged skippers
512:Pyle, Robert Michael
247:, and rotting fruit.
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680:Fauna Europaea (new)
377:Daniels (2003): p.12
172:, commonly known as
484:Glassberg, Jeffrey
405:2019PNAS..116.6232L
527:Subfamily Pyrginae
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189:Hermann Burmeister
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558:Taxon identifiers
505:Pelham, Jonathan
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458:Daniels, Jaret C.
399:(13): 6232–6237.
182:skipper butterfly
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56:Ngorongoro Crater
16:(Redirected from
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319:Former tribes
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352:Pyrrhopygini
303:Achlyodidini
298:Carcharodini
291:phylogenetic
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51:Spialia mafa
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693:iNaturalist
590:Wikispecies
193:systematics
137:Subfamily:
131:Hesperiidae
121:Lepidoptera
58:, Tanzania
754:Categories
452:References
347:Tagiadinae
329:Tagiadinae
293:sequence:
271:Neotropics
267:Neotropics
262:regions.
146:Burmeister
101:Arthropoda
423:0027-8424
362:Footnotes
343:Tagiadini
338:Eudaminae
285:The four
260:temperate
178:subfamily
155:Diversity
87:Kingdom:
81:Eukaryota
760:Pyrginae
659:59285139
628:BugGuide
617:BioLib:
610:Pyrginae
596:Pyrginae
581:Q1306572
575:Wikidata
566:Pyrginae
460:(2003):
441:30877254
334:Eudamini
308:Erynnini
176:, are a
170:Pyrginae
162:4 tribes
141:Pyrginae
127:Family:
97:Phylum:
91:Animalia
77:Domain:
18:Pyrginae
432:6442542
401:Bibcode
313:Pyrgini
256:tropics
245:carrion
229:Pyrgini
225:Florida
180:of the
117:Order:
111:Insecta
107:Class:
54:in the
711:694116
672:440742
516:(2002)
509:(2008)
502:(2011)
495:(2001)
488:(2001)
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439:
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287:tribes
281:Tribes
252:desert
185:family
148:, 1878
737:40100
698:51312
646:93MZN
633:12853
620:51218
529:from
223:from
732:NCBI
706:ITIS
467:ISBN
437:PMID
419:ISSN
241:dung
719:NBN
654:EoL
641:CoL
605:AFD
427:PMC
409:doi
397:116
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370:^
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227:(
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Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.