1211:
1111:
944:
1456:
568:
817:
5950:
5961:
877:, which he estimated at over 35 m (115 ft) in length and 65–75 tonnes (64–74 long tons; 72–83 short tons) in weight. In 2020, Campione and Evans yielded a body mass estimate of approximately 55.7 tonnes (54.8 long tons; 61.4 short tons). In 2020, Otero and Carballido published a lower mean weight estimate of 57 tonnes (56 long tons; 63 short tons) using a revised scaling equation, with a margin of error of 42.5–71.4 tonnes (41.8–70.3 long tons; 46.8–78.7 short tons); this was close to the 2017 volumetric estimate.
5968:
7362:
8094:
144:
9329:
9323:
7367:
9404:
6824:
3147:
9318:
6816:
9312:
8086:
759:
6836:
5956:
6830:
8102:
118:
9392:
1640:; the two were united by the presence of long neural spines in the front back vertebrae. These two were placed within the Lognkosauria, based on the SDLs, the tall transverse processes, and small laminae that connect the neural spine to the prezygapophyses (i.e., spinoprezygapophyseal laminae or SPPLs) in the tail vertebrae of
655:. Based on overlapping elements, these include at least three individuals from FLV1, one from FLV2, and one from FLV3 other than the holotype. MPEF-PV 3399 (from FLV1) is a second skeleton consisting of six neck vertebrae, four back vertebrae, one front tail vertebra, sixteen rear tail vertebrae, ribs, chevrons, the left
1648:
to the exclusion of other titanosaurs, where the SPPLs of the tail vertebrae were much better-developed. Notably, this emergence of this grouping corresponded to a major increase in body size as it contained the largest titanosaurs, with maximum weights increasing from 20 to 60 tonnes (22 to 66 short
854:
size at 40 m (131 ft) long with a weight of 77 tonnes (85 short tons). In 2017, Carballido and colleagues published a length estimate of 37 m (121 ft), and provided two weight estimates: 69 tonnes (76 short tons) using a scaling equation, and 44.2–77.6 tonnes (43.5–76.4 long tons;
583:
The collected sauropod fossils were organized into six partial skeletons, which likely belonged to the same species due to uniformity in their morphology and size (with all individuals differing no more than 5% in length). Although the remains all came from the same quarry, the individuals likely did
495:
between only one pair of vertebrae at the level of the scapular blade, which was likely a weight-bearing adaptation not seen in any other sauropod (where they were either present between all pairs or between none). Several unique features in the limbs were also likely attachment scars for muscles. In
1154:(vertebral bodies). Yet another unique characteristic was that the tops of the neural spines were concave, making them somewhat bifurcated with the tips curling forwards. On the neural spine, longitudinal laminae separated from the prespinal and postspinal laminae, which was previously only seen in
2669:
was among the most phylogenetically stable titanosaurs, with a high recuperation index of 62% (meaning that it is stable enough to not be removed due to instability in 62% of analyses) due to its firm position within the
Lognkosauria. They found that the uncertainty regarding whether Lognkosauria
774:
are displayed in multiple museums. Research
Casting International digitally scanned the specimens, which was followed by the creation of foam molds, fiberglass casts, and 3D printing. One mount is exhibited in the Museum of Paleontology Egidio Feruglio. Another is exhibited in the
584:
not all die at the same time. Within the 3.43-metre (11.3 ft) sediment layer containing the fossils, there are three distinct but closely-spaced levels corresponding to three burial events, which are named FLV1, FLV2, and FLV3. One skeleton from FLV3 was designated as the
2595:
being closest to this group; unlike
Carballido and colleagues, however, they recovered the Lognkosauria within the Lithostrotia. In 2019, González Riga and colleagues recovered the same arrangement, and they named the group formed by the Lognkosauria and the Rinconsauria as
1054:
at the sides) were nearly vertical because the prezygapophyses were situated higher up on the vertebrae than the diapophyses. The PRDLs usually instead extended forward from the diapophyses in other sauropods. All of the neural spines in the back were tall and vertical in
2640:, and Lognkosauria being outside Lithostrotia; Federico Agnolin and colleagues found a similar arrangement in 2023. In 2021, Otero and colleagues recovered the same arrangement. Also in 2021, Pablo Gallina and colleagues recovered a similar arrangement, except with
536:(MPEF) in Trelew. One preliminary field expedition to the La Flecha site was carried out in late 2012 by the MPEF, followed by seven further expeditions that discovered new fossils between January 2013 and February 2015. The lead scientists on the excavation were
1085:
had short front neural spines lacking the arrow-shaped ends. Uniquely, in the first two back vertebrae, the prespinal laminae that ran alongside the front of the neural spines bulged outwards near the bottom. Also uniquely, the third back vertebra preserves a
1607:
on the inner surface of the femur was located a third of the way from the top instead of halfway down. Uniquely among sauropods, at the outer edge of the bottom of the femur, there were a series of coarse ridges right above the articulation known as the
1130:, the front articulating surface of the first tail vertebra was flat. Also uniquely, the neural spines of the front tail vertebrae were about four to six times as wide from side to side as they were long front to back, contrasting with the condition in
935:. He attributed this conclusion to a lack of proper measurement techniques. Also, he criticized the 3D model used for the volumetric estimate as having a flat-sided torso, and noted that the neck and tail were reconstructed as being fairly long.
4274:
González Riga, B.J.; Lamanna, M.C.; Otero, A.; Ortiz David, L.D.; Kellner, A.W.A.; Ibiricu, L.M. (2019). "An overview of the appendicular skeletal anatomy of South
American titanosaurian sauropods, with definition of a newly recognized clade".
4424:
Agnolin, F.L.; Gonzalez Riga, B.J.; Aranciaga
Rolando, A.M.; Rozadilla, S.; Motta, M.J.; Chimento, N.R.; Novas, F.E. (2023). "A new gigant titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Northwestern Patagonia, Argentina".
4199:
Carballido, J.L.; Otero, A.; Mannion, P.D.; Salgado, L.; Moreno, A.P. (2022). "Titanosauria: A critical appraisal of its systematics and the relevance of the South
American record". In Otero, A.; Carballido, J.L.; Pol, D. (eds.).
4064:
Carballido, J.L.; Salgado, L.; Pol, D.; Canudo, J.I.; Garrido, A. (2012). "A new basal rebbachisaurid (Sauropoda, Diplodocoidea) from the Early
Cretaceous of the Neuquén Basin; evolution and biogeography of the group".
1102:
at the front; the missing fourth back vertebra presumably had a corresponding hypantrum. Hyposphene-hypantrum articulations to stiffen the vertebrae were primitively present among sauropods and lost multiple times, but
2624:, and others depending on the exclusion of some other titanosaurs from the analysis; this exclusion also affected whether Lognkosauria was inside Lithostrotia. In 2020, Carballido and colleagues found again that
679:, the left pubic bone, and the left femur. MPEF-PV 3372 (from FLV1) is a tooth. MPEF-PV 3393 (not associated with a layer) is a rear tail vertebra. MPEF-PV 3395 and MPEV-PV 3396 (both from FLV1) are left
3514:
1293:. Furthermore, in addition to a ridge that ran along the midline of the blade parallel to the bottom, there was a second ridge that ran upwards diagonally to the top of the bone, which was unique to
2674:. Instead of using Lithostrotia, they suggested that Eutitanosauria be divided into Saltasauroidea and Colossosauria. They also refined Colossosauria as being the most inclusive group that includes
1107:
is the first known sauropod where they were retained between only one pair of vertebrae. Because the shoulder blade was located at this position, this may have been a weight-bearing adaptation.
687:, while MPEF-PV 3397 (from FLV2) is a right humerus. MPEF-PV 3375 (FLV3) is a left femur while MPEF-PV 3394 (FLV1) is a right femur. MPEF-PV 3391 and MPEF-PV 3392 (both from FLV1) represent two
1382:) on the front upper portion of the humerus, there was a pair of scars above each other, with the top one being wider than tall and the bottom one being taller than wide; this was where the
1247:. Furthermore, expansion was present at both ends such that the top end of the scapula was roughly at the same level as the bottom end, which was an uncommon characteristic also seen in
1418:. However, the inward-pointing anteromedial process of the ulna was relatively slender, being 3.5 times as long as it was wide, and was also relatively horizontal with an angle of 40°;
4468:
Otero, A.; Carballido, J.L.; Salgado, L.; Ignacio Canudo, J.; Garrido, A.C. (2021). "Report of a giant titanosaur sauropod from the Upper
Cretaceous of Neuquén Province, Argentina".
1521:, the bottom of the pubic "boot" was not particularly expanded. On the side of the ischium, there was also a well-developed ridge (seen to a lesser extent in the titanosauriforms
396:). Later research revised the length estimate down to 31 m (102 ft) and weight estimates down to approximately 50–57 tonnes (55–63 short tons), suggesting that
4874:
Leardi, J.M.; Pol, D. (2009). "The first crocodyliform from the Chubut Group (Chubut
Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia".
1669:. Lastly, unlike the results of prior analyses, the analysis of Carballido and colleagues placed Lognkosauria outside the Lithostrotia, as separate groups within the
715:
699:
4774:
Novas, F.E.; de Valais, S.; Vickers-Rich, P.; Rich, T. (2005). "A large
Cretaceous theropod from Patagonia, Argentina, and the evolution of carcharodontosaurids".
707:
3203:, was named from the fossilized wood in 2019; these trees were estimated as having been 15.3 metres (50 ft) tall. A new species of the angiosperm wood genus
3199:
719:
1434:. The radius was also fairly expanded, being 36% as broad at the top as it was long. On the inner edge of the radius, a bulging scar for the attachment of the
537:
525:
3522:
3488:
975:
had deep pleurocoels that opened from the sides into the interior of the bone; whether the neural spines of these vertebrae were horizontally expanded as in
1491:, and saltasaurids). The articulation between the pubic bones was clearly demarcated from their articulations with other bones by deflected surfaces, as in
1394:
on the outer side, there was one scar at the midpoint of the crest and one at the bottom; this was where the supracoracoideus, deltoideus clavicularis, and
1329:, in which it was strongly concave). There was a ridge running diagonally from the outer bottom corner of each sternal plate that was oriented similarly to
9493:
4753:"Estudio estratigráfico y sedimentolĂłgico de las sedimentitas portadoras de los titanosaurios gigantes del Albiano tardĂo de Patagonia Central, Argentina"
2658:. In 2023, AgustĂn PĂ©rez Moreno and colleagues performed an analysis based on that of Gallina and colleagues, and found a similar arrangement except with
843:
Given the size of these bones, which surpass any of the previously known giant animals, the new dinosaur is the largest animal known that walked on Earth.
9508:
711:
648:, and both femora. The skeleton was chosen to be the holotype because it was the best preserved and also the one showing the most distinguishing traits.
4233:
González Riga, B.J.; Mannion, P.D.; Poropat, S.F.; Ortiz David, L.; Coria, J.P. (2018). "Osteology of the Late Cretaceous Argentinean sauropod dinosaur
3240:, meaning that the individuals were preserved where they died. Animals from other localities in the Cerro Castaño Member include the theropod dinosaurs
4594:
3462:
1539:) extending down the ischial tuberosity (a common feature among titanosaurs) down to the blade, with the combination of these features being unique to
873:
at 31 m (102 ft) in length and 50–55 tonnes (49–54 long tons; 55–61 short tons) in weight using volumetric models. This made it smaller than
695:. Another quarry was excavated about 300 metres (980 ft) west, revealing a fourth sediment layer (FLV4) with a similarly-sized sauropod skeleton.
3205:
881:
of five femora and one humerus indicates that the individuals died while they were young adults, with growth having slowed but not ceased entirely.
1463:
In the hip girdle, the contact between the forward-pointing pubis and backward-pointing ischium was about half of the former's length (longer than
5003:
4330:"New information on the Cretaceous sauropod dinosaurs of Zhejiang Province, China: impact on Laurasian titanosauriform phylogeny and biogeography"
915:
has been estimated at 65–96.4 tonnes (71.7–106.3 short tons). In 2019, Paul noted that the articulated length of the back vertebrae was larger in
1297:. The articulating surface of the coracoid with the scapula was 80% as long front-to-back as it was tall, an uncommon characteristic shared with
3788:
2575:
in analyses including the three titanosaurs, with either one being its closest relative. In 2018, Bernardo González Riga and colleagues found
489:
439:
was a particularly large and robust titanosaur. It can be distinguished from its close relatives by a suite of unique characteristics in its
3690:"Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals"
3545:
3571:
4620:
3814:
1160:. At the sides of the front tail vertebrae, the transverse processes (on which the diapophyses were located) were high and thin, like
5052:
1259:
ridge was about 30% as long as the scapula. In addition to a projection at the bottom of the scapula just behind the shoulder joint (
1210:
1110:
943:
3161:
lived in what was then a forested region. Numerous plant fossils were found at a site about 200 metres (660 ft) away from the
2563:
Historically, the interrelationships of titanosaurs have been highly unstable and varied between different analyses. Despite this,
896:
was comparable in size with other known giant titanosaurs. However, almost every measurement that could be compared was larger in
2670:
belonged within Lithostrotia largely arose from the definition of Lithostrotia, which was based on the phylogenetically unstable
1267:), which was common among titanosaurs, there was a second one further back at the beginning of the blade, only seen otherwise in
1370:, it was about 70% as long as the femur. A number of bulges for muscle attachment on the humerus were unique characteristics of
9528:
4706:
from the Early Cretaceous of Patagonia (Chubut province, Argentina): the oldest record of angiosperm wood from South America".
4385:
Carballido, J.L.; Scheil, M.; Knötschke, N.; Sander, P.M. (2019). "The appendicular skeleton of the dwarf macronarian sauropod
3350:
Carballido, J.L.; Pol, D.; Otero, A.; Cerda, I.A.; Salgado, L.; Garrido, A.C.; Ramezani, J.; CĂşneo, N.R.; Krause, J.M. (2017).
1388:
1221:
had a robust, expanded scapula, with the blade being only 4.15 times as long as it was wide at its narrowest point; except for
1032:
533:
1202:, well-developed spinodiapophyseal laminae (SDLs) ran from the neural spines down the tranverse processes to the diapophyses.
590:
416:
is one of the most-known titanosaurs, and so its interrelationships with other titanosaurs have been relatively consistent in
4217:
1315:. Like other titanosauriforms, the sternal plates were crescent-shaped with the outer edge being slightly concave (more like
606:
598:
449:
441:
5073:
1040:
634:
4752:
9543:
3288:
957:
2686:. The following phylogenetic tree shows their suggested group definitions, including only the most stable titanosaurs.
1176:, where the transverse processes of the front tail vertebrae formed wing-like shapes, they were wider at the bottom in
1048:
965:
776:
763:
642:
1120:
1088:
931:
to have been larger, he reasoned that the rest of its body would have to have been improbably large, or vice versa in
588:
of a new species, and was catalogued as MPEF-PV 3400. It consists of a partial skeleton lacking the skull, with three
4975:
1096:
1253:
614:
457:
9523:
1261:
1010:
689:
3878:
1148:
681:
673:
665:
481:
473:
465:
5960:
1376:
786:
518:
126:
5967:
5949:
4852:
657:
622:
4925:, a New Eucryptodiran Turtle from the Early Cretaceous of Argentina, and the Relationships of the Meiolaniidae"
1613:
800:
377:
7361:
4647:"A new cupressaceous wood from the Lower Cretaceous of central Patagonia reveals possible clonal growth habit"
4546:
PĂ©rez Moreno, A.; Otero, A.; Carballido, J.L.; Salgado, L.; Calvo, J.O. (2023). "The appendicular skeleton of
785:
mount. Some of the original fossils, including a femur, were also on display briefly. One is exhibited in the
9518:
9513:
5045:
900:. Wedel also criticised the polygon-based method that Carballido and colleagues used to compare the sizes of
799:(which has been moved to another exhibit). From March 2023 until January 2024, one will be displayed at the
3236:, likely related to sporadic sheet flooding. This flooding was probably too weak to transport the bones of
1579:, a ridge ran along the back of the femur between these two features, but there was no ridge in front like
388:
and land animal overall, with an estimated length of 37 m (121 ft) and an estimated weight of 69
5025:
892:
cautioned against the reported estimates. In blog posts, Wedel noted that based on available measurements
855:
48.7–85.5 short tons) using volumetric method based on 3D skeletal models. These estimates suggested that
4511:
Gallina, P.A.; Canale, J.I.; Carballido, J.L. (2021). "The Earliest Known Titanosaur Sauropod Dinosaur".
8093:
3181:. The plant fauna was dominated by conifers, with angiosperm wood being relatively rare. A new genus of
556:, and were specifically dated to approximately 101.62 million years in age (corresponding to the latest
143:
9538:
9382:
552:
teeth, were uncovered. The rock layers at the La Flecha site belong to the Cerro Castaño Member of the
490:
4110:"A new Cretaceous terrestrial ecosystem from Gondwana with the description of a new sauropod dinosaur"
1455:
1146:
where they were longer front to back. They were also tall, being 1.5 times taller than the underlying
9328:
9322:
3848:
3411:
947:
Back vertebra, showing the vertical neural spine to the right which bears the prespinal lamina on top
3906:
9533:
9365:
5140:
5038:
1609:
1126:, meaning that the front articulating surface was concave and the rear one was convex. Uniquely in
1016:, ridges of bone crossing the vertebrae, were thin and strongly developed in the back vertebrae of
7366:
1657:
represented independent increases in body size to a much lesser extent. The group of lognkosaurs,
366:, Argentina. The genus contains a single species known from at least six young adult individuals,
9498:
1395:
1383:
747:
553:
355:
9470:
487:
in the hindlimb. Among these was the presence of accessory vertebral articulations known as the
9420:
6835:
6823:
6815:
4702:
Nunes, C.I.; Pujana, R.R.; Escapa, I.H.; Gandolfo, M.A.; CĂşneo, N.R. (2018). "A new species of
3901:
1180:. These expanded transverse processes would have resulted in greater attachment areas for the
9465:
9355:
9317:
9046:
5328:
2794:
1625:
1426:
in both aspects. The bottom end of the bone was mildly expanded, though not to the extent of
417:
9311:
8085:
6829:
9503:
9452:
8101:
5955:
4883:
4783:
4658:
4559:
4477:
4434:
4341:
4074:
4012:
3950:
3893:
3427:
4919:
Gaffney, E.S.; Rich, T.H.; Vickers-Rich, P.; Constantine, A.; Vacca, R.; Kool, L. (2007).
3657:
3598:
3572:"'The sheer scale is extraordinary': meet the titanosaur that dwarfs Dippy the diplodocus"
3489:"Massive titanosaur, biggest dinosaur ever found, squeezes into Museum of Natural History"
8:
6031:
3515:"Gigantic Dinosaur, 'Titanosaur,' Going on Display at American Museum of Natural History"
567:
380:
by José Carballido and colleagues. Preliminary studies and press releases suggested that
4887:
4787:
4662:
4563:
4481:
4438:
4345:
4078:
4016:
3999:
González Riga, B.J.; Lamanna, M.C.; Ortiz David, L.D.; Calvo, J.O.; Coria, J.P. (2016).
3954:
3897:
3629:
3431:
1567:. Like other titanosauriforms, the head of the femur was angled upwards relative to the
1389:
1033:
9086:
6158:
4856:
4817:
4733:
4684:
4575:
4528:
4493:
4450:
4406:
4362:
4329:
4310:
4090:
4041:
4000:
3973:
3938:
3919:
3769:
3712:
3443:
3376:
3351:
3283:
2801:
1956:
1568:
1398:
inserted. The ulna was robust, being about 50% as broad at the top as it was long like
1335:
591:
138:
3296:
fossils over a two-year period. The programme is also known by the alternative title,
955:
were very long, being at least five times as long as they were wide at the back, with
816:
9222:
8763:
7510:
7151:
6880:
6068:
4860:
4809:
4688:
4579:
4532:
4497:
4454:
4410:
4389:
from the Late Jurassic of Germany and a re-evaluation of its systematic affinities".
4367:
4314:
4302:
4213:
4131:
4094:
4046:
4028:
3978:
3879:"Giants and bizarres: body size of some southern South American Cretaceous dinosaurs"
3773:
3761:
3753:
3716:
3447:
3381:
1674:
1604:
1506:
1064:
743:
607:
599:
450:
442:
4821:
4737:
4126:
4109:
4001:"A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot"
3923:
3352:"A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs"
1041:
635:
9408:
9396:
9093:
8938:
8878:
8520:
8412:
8318:
8209:
8011:
7991:
7930:
7049:
6902:
6749:
6466:
6420:
6400:
6344:
5647:
5424:
5404:
4932:
4920:
4899:
4891:
4848:
4799:
4791:
4723:
4715:
4674:
4666:
4567:
4520:
4485:
4442:
4398:
4357:
4349:
4292:
4284:
4254:
4246:
4205:
4176:
4166:
4121:
4082:
4036:
4020:
3968:
3958:
3911:
3743:
3704:
3435:
3371:
3363:
3166:
3022:
2734:
2727:
2341:
1555:
1529:
1346:
was on average 28% as broad as it was long, making it similarly robust to those of
977:
703:
4895:
4670:
4571:
4489:
4446:
4402:
4288:
3439:
958:
839:
and is notable for its large size. Carballido and colleagues stated in the media:
9443:
9360:
9230:
8963:
8856:
8552:
8532:
8513:
8471:
8398:
8384:
8267:
8018:
7938:
7585:
7121:
7112:
6973:
6965:
6892:
6652:
6530:
6351:
6207:
6038:
5767:
5760:
5752:
5739:
5656:
5202:
5189:
5152:
5125:
5100:
5061:
4524:
4086:
3963:
3915:
3256:
2432:
866:
861:
572:
402:
373:
359:
217:
4646:
4250:
4209:
1049:
966:
643:
524:
was discovered in 2010 by a farm laborer, Aurelio Hernández, in the desert near
9237:
9152:
9034:
8979:
8929:
8741:
8734:
8680:
8652:
8607:
8580:
8455:
8420:
8362:
8289:
8202:
7966:
7845:
7817:
7796:
7781:
7703:
7676:
7494:
7478:
7444:
7287:
7086:
6958:
6784:
6523:
6502:
6451:
6443:
6295:
6288:
6195:
6094:
6082:
5890:
5862:
5699:
5670:
5663:
5609:
5523:
5509:
5493:
5468:
5298:
3193:
2866:
2757:
2747:
1809:
1796:
1763:
1670:
1505:, there was a ridge running along the side of the pubis that vanished near the
1181:
1136:
1121:
1089:
971:(neurovascular openings) on the sides. By contrast, the rear neck vertebrae of
790:
585:
4795:
4719:
4595:"Researchers Discover Fossils of Largest Dino Believed to Ever Walk the Earth"
4550:(Sauropoda: Titanosauria) from the Upper Cretaceous of Patagonia, Argentina".
4423:
1097:
1063:
had front neural spines with arrow-shaped ends, but like other members of the
9487:
9280:
9265:
9244:
9160:
9120:
9108:
9078:
9027:
8999:
8972:
8893:
8870:
8818:
8749:
8673:
8639:
8628:
8566:
8485:
8448:
8427:
8377:
8355:
8347:
8340:
8333:
8274:
8239:
8216:
8158:
8025:
8004:
7872:
7863:
7852:
7831:
7810:
7788:
7731:
7608:
7545:
7429:
7252:
7228:
7194:
7165:
7158:
7137:
7072:
7065:
7058:
7023:
7014:
6951:
6943:
6936:
6866:
6853:
6770:
6694:
6637:
6616:
6565:
6558:
6551:
6544:
6537:
6494:
6473:
6392:
6369:
6216:
6173:
6075:
6052:
6024:
5855:
5848:
5796:
5692:
5629:
5620:
5502:
5376:
5344:
5312:
5273:
5255:
5234:
4032:
3757:
3309:
3260:
3253:
3046:
2968:
2924:
2852:
2814:
2784:
2771:
2597:
2310:
2167:
1990:
1922:
1905:
1830:
1746:
1473:
1326:
1321:
1305:
1289:
1254:
1233:
1073:
1004:
probably had 10. Several characteristics in the back vertebrae distinguished
989:
983:
911:
vertebrae, noting that the former was largely empty space. In other studies,
795:
615:
458:
421:
385:
68:
5020:
4950:
1262:
1011:
690:
9288:
9258:
9251:
9188:
9129:
9055:
8987:
8945:
8914:
8900:
8885:
8863:
8847:
8781:
8770:
8721:
8702:
8695:
8573:
8559:
8545:
8499:
8492:
8463:
8434:
8370:
8311:
8253:
8246:
8195:
8188:
8151:
8133:
8120:
8048:
8032:
7973:
7959:
7923:
7916:
7902:
7880:
7774:
7766:
7759:
7738:
7710:
7658:
7644:
7630:
7615:
7566:
7487:
7459:
7452:
7271:
7260:
7237:
7219:
7144:
7130:
7037:
6994:
6987:
6929:
6922:
6685:
6600:
6408:
6358:
6330:
6323:
6302:
6274:
6260:
6180:
6124:
6103:
5789:
5677:
5581:
5537:
5530:
5482:
5454:
5415:
5241:
4813:
4645:
Nunes, C.I.; Bodnar, J.; Escapa, I.H.; Gandolfo, M.A.; CĂşneo, N.R. (2019).
4371:
4306:
4259:
4135:
4050:
3982:
3765:
3732:"The accuracy and precision of body mass estimation in non-avian dinosaurs"
3689:
3576:
3385:
3367:
3242:
3182:
3008:
2937:
2900:
2886:
2693:
2408:
2391:
2358:
2296:
2283:
2143:
2129:
2055:
2014:
1857:
1843:
1698:
1684:
1666:
1633:
1535:
1501:
1448:
1354:
1348:
1311:
1299:
1198:
1173:
1149:
1142:
1082:
1026:
963:
on top that were horizontally flattened, and only small holes and no large
735:
723:
682:
674:
666:
505:
482:
474:
466:
432:
408:
311:
281:
265:
4171:
4150:
1438:
muscle extended towards the front of the bone as a crest, as also seen in
1377:
519:
9273:
9181:
9167:
9063:
8952:
8804:
8797:
8790:
8756:
8688:
8666:
8587:
8478:
8173:
7980:
7945:
7838:
7803:
7745:
7724:
7717:
7689:
7622:
7594:
7559:
7552:
7538:
7437:
7422:
7323:
7245:
7208:
7079:
7030:
6791:
6667:
6630:
6623:
6592:
6576:
6516:
6480:
6458:
6378:
6337:
6281:
6253:
6239:
6187:
6140:
6060:
6045:
6017:
5883:
5841:
5827:
5807:
5781:
5727:
5720:
5636:
5602:
5588:
5431:
5369:
5305:
5282:
5248:
5220:
4151:"Cranial anatomy and phylogenetic position of the titanosaurian sauropod
4108:
Calvo, J.O.; Porfiri, J.D.; González-Riga, B.J.; Kellner, A.W.A. (2007).
3708:
3463:"Here's How You Squeeze the Biggest Dinosaur Into a New York City Museum"
3248:
2951:
2821:
2259:
2208:
2191:
1973:
1939:
1479:
1269:
1239:
1156:
658:
623:
43:
4804:
4728:
4679:
4353:
4297:
4202:
South American Sauropodomorph Dinosaurs: Record, Diversity and Evolution
4181:
3292:, followed the excavation, research and reconstruction of the La Flecha
1644:. The last of these characteristics were used to group lognkosaurs with
9174:
9101:
9070:
8832:
8659:
8506:
8441:
8405:
8391:
8326:
8297:
8260:
8223:
8166:
7752:
7696:
7651:
7601:
7573:
7531:
7467:
7415:
7397:
7384:
7308:
7301:
7294:
7280:
7187:
7180:
7173:
7093:
7005:
6980:
6710:
6660:
6644:
6608:
6509:
6385:
6309:
6223:
6166:
6117:
5869:
5685:
5574:
5438:
5397:
5362:
5091:
4904:
3939:"March of the Titans: The Locomotor Capabilities of Sauropod Dinosaurs"
3270:
3265:
3233:
3178:
2707:
2642:
1881:
1715:
1549:
1523:
1244:
1000:
and other titanosaurs that preserved a complete set of back vertebrae,
889:
781:
561:
541:
501:
345:
249:
201:
88:
53:
4839:, a large-toothed ceratosaur (Dinosauria: Theropoda) from Patagonia".
4232:
4024:
3748:
3731:
3282:
A BBC television programme broadcast in January 2016 and presented by
3146:
2665:
Due to its completeness, Carballido and colleagues in 2022 found that
1030:. Uniquely, the prezygodiapophyseal laminae (PRDLs, which ran between
9137:
9019:
8825:
8304:
8282:
8231:
7952:
7909:
7895:
7637:
7330:
7316:
7201:
6915:
6763:
6756:
6487:
6436:
5999:
5986:
5901:
5876:
5834:
5713:
5553:
5320:
5262:
4936:
3225:
3217:
2038:
1547:
was robust, being on average 23% as broad as it was long, similar to
1283:
1227:
878:
739:
549:
545:
400:
was of a similar size to, if not smaller than, its closest relatives
393:
363:
348:
233:
155:
93:
37:
9414:
5026:
Raising the Dinosaur Giant, PBS, with transcript near bottom of page
4467:
4273:
3998:
3546:"New mega-dinosaur at Field Museum is named Maximo, unveiled June 1"
758:
9437:
9145:
8811:
8621:
7824:
7665:
7524:
6777:
6267:
6110:
5820:
5595:
5567:
5475:
5390:
5383:
5290:
5227:
5109:
5085:
3228:
sediments of the Cerro Castaño Member, from which the specimens of
3187:
2580:
1509:(which was elongated like other titanosauriforms) at the top. Like
1185:
652:
351:
188:
175:
117:
83:
78:
63:
58:
48:
5030:
4918:
4545:
3937:
Sellers, W.I.; Margetts, L.; Coria, R.A.B.; Manning, P.L. (2013).
9457:
8181:
6585:
6429:
6316:
5774:
5007:
3174:
2600:. Also in 2019, Philip Mannion and colleagues found the group of
629:
130:
98:
73:
4107:
1071:
represents a reversal to the condition in non-titanosauriforms.
8710:
7888:
5706:
5079:
4384:
1483:) but less than half of the latter's length (common, except in
1435:
1214:
Forelimb and scapula at Museo Egidio Feruglio de Trelew, Chubut
557:
529:
165:
30:
4773:
4204:. Springer Earth System Sciences. Springer. pp. 269–298.
3789:"Did Scientists Just Unveil the Biggest Dinosaur of All Time?"
951:
The vertebrae in the front and middle portions of the neck of
4198:
3232:
were uncovered, are indicative of low-energy water flow on a
389:
9012:
5516:
4853:
10.1671/0272-4634(2004)024[0894:PAAOGS]2.0.CO;2
3936:
3409:
3221:
3177:; and leaf and flower impressions and fossilized wood from
3170:
836:
832:
698:
In 2017, José Luis Carballido, Diego Pol, Alejandro Otero,
4757:
Latin American Journal of Sedimentology and Basin Analysis
4063:
3876:
746:
for the "strength and large size" of this titanosaur. The
6674:
4701:
4644:
4328:
Mannion, P.D.; Upchurch, P.; Jin, X.; Zheng, W. (2019).
1243:, this figure was usually considerably higher among the
4510:
4327:
3877:
Mazzetta, G.V.; Christiansen, P.; Farina, R.A. (2004).
3356:
Proceedings of the Royal Society B: Biological Sciences
3349:
1188:
muscles, and therefore a more muscular tail. Also like
923:(3.67 metres (12.0 ft)), making it impossible for
888:
in 2017, science writer Riley Black and paleontologist
3410:
Otero, A.; Carballido, J.L.; PĂ©rez Moreno, A. (2020).
420:. This led to its use in a re-definition of the group
9380:
9345:
2612:
in a polytomy with other titanosaurs, which included
1059:, and the first few had expanded, arrow-shaped ends.
528:, Argentina, about 250 km (160 mi) west of
4750:
4621:"Argentine fossil biggest dinosaur ever: scientists"
4237:: implications for basal titanosaur relationships".
1386:
inserted. On the other side of the bone, behind the
750:
honours the Mayo family, owners of La Flecha ranch.
318:
4751:Carmona, R.P.; Umazano, A.M.; Krause, J.M. (2016).
3994:
3992:
1118:In most titanosaurs, the front tail vertebrae were
770:Skeletal mounts based on all available material of
9214:
4835:Rauhut, O.W.M. (2004). "Provenance and anatomy of
2567:has been consistently found as being grouped with
1677:shows the results of their phylogenetic analysis.
532:. This discovery was reported to P. Huerta at the
5004:Meet Patagotitan, the Biggest Dinosaur Ever Found
4194:
4192:
4148:
3345:
9485:
6741:
4943:
4267:
4226:
3989:
3343:
3341:
3339:
3337:
3335:
3333:
3331:
3329:
3327:
3325:
1067:the rear neural spines were inclined backwards;
734:one of the most completely-known members of the
4539:
2632:, with their grouping being in a polytomy with
5200:
5124:
5021:David Attenborough and the Giant Dinosaur, BBC
4378:
4189:
4057:
3730:Campione, Nicolás E.; Evans, David C. (2020).
5046:
3729:
3322:
1358:but less so than those of saltasaurids. Like
6864:
3870:
3623:
3621:
3619:
3506:
1673:and in turn the Titanosauria. The following
9494:Early Cretaceous dinosaurs of South America
8131:
4968:
4912:
4744:
4504:
4417:
4321:
4142:
3930:
3683:
3681:
3679:
3405:
3403:
3401:
3399:
3397:
3395:
847:In 2014 news reports provided estimates of
738:. The generic name combines a reference to
9509:Late Cretaceous dinosaurs of South America
5053:
5039:
4867:
4828:
4586:
3840:
3806:
3652:
3650:
3599:"Titanosaur: Life as the Biggest Dinosaur"
3537:
3173:; leaves, cones, and fossilized wood from
116:
7395:
4903:
4873:
4803:
4767:
4727:
4695:
4678:
4638:
4461:
4361:
4296:
4258:
4239:Zoological Journal of the Linnean Society
4180:
4170:
4125:
4101:
4040:
3972:
3962:
3905:
3780:
3747:
3616:
3512:
3375:
1205:
762:Reconstructed skeleton on display at the
540:and Diego Pol, with partial funding from
125:Reconstructed skeleton on display at the
5997:
4277:Anais da Academia Brasileira de CiĂŞncias
4114:Anais da Academia Brasileira de CiĂŞncias
3676:
3454:
3392:
3145:
1454:
1452:exhibited the bulge without the crest).
1209:
1109:
942:
831:was a quadrupedal herbivore with a long
815:
757:
566:
4618:
3647:
3543:
3480:
1251:. However, like other titanosaurs, the
730:. Collectively, the known remains made
651:Other specimens were designated as the
544:. More than 200 fossils, including 130
9486:
4834:
4592:
4149:Gallina, P.A.; ApesteguĂa, S. (2011).
3627:
534:Museum of Paleontology Egidio Feruglio
424:by Carballido and colleagues in 2022.
9419:
9418:
9344:
8118:
7382:
6851:
5984:
5187:
5072:
5034:
4976:"Attenborough and the Giant Dinosaur"
4951:"Attenborough and the Giant Dinosaur"
3857:Sauropod Vertebra Picture of the Week
3846:
3827:Sauropod Vertebra Picture of the Week
3812:
3786:
3569:
3460:
3277:
1665:was recovered as the sister group of
919:(4.47 metres (14.7 ft)) than in
884:Following the initial publication of
3687:
3630:"'Biggest dinosaur ever' discovered"
3563:
3486:
1628:by Carballido and colleagues placed
5188:
5060:
4391:Journal of Systematic Palaeontology
3289:Attenborough and the Giant Dinosaur
13:
4997:
4841:Journal of Vertebrate Paleontology
3591:
3420:Journal of Vertebrate Paleontology
1172:and other titanosaurs. Unlike the
777:American Museum of Natural History
764:American Museum of Natural History
491:hyposphene-hypantrum articulations
14:
9555:
9346:Topics in sauropodomorph research
6852:
5014:
3658:"Giant dinosaur slims down a bit"
3570:McKie, Robin (26 November 2022).
3513:Battaglia, A. (15 January 2016).
1619:
1543:. Like the humerus, the femur of
827:Like other titanosaur sauropods,
9402:
9390:
9327:
9321:
9316:
9310:
8119:
8100:
8092:
8084:
7365:
7360:
6834:
6828:
6822:
6814:
5966:
5959:
5954:
5948:
1114:Chevrons and rear tail vertebrae
927:to have had a larger torso. For
500:lived in a forested region on a
142:
41:
4612:
4127:10.1590/S0001-37652007000300013
3723:
3412:"The appendicular osteology of
3141:
787:Field Museum of Natural History
779:, where it replaced a juvenile
127:Field Museum of Natural History
7383:
1614:flexor digitorum longus muscle
806:
801:Natural History Museum, London
1:
9529:Fossil taxa described in 2017
5985:
4896:10.1016/j.cretres.2009.08.002
4671:10.1016/j.cretres.2019.02.013
4572:10.1016/j.cretres.2022.105389
4490:10.1016/j.cretres.2021.104754
4447:10.1016/j.cretres.2023.105487
4403:10.1080/14772019.2019.1683770
4289:10.1590/0001-3765201920180374
4159:Acta Palaeontologica Polonica
3697:Annals of the Carnegie Museum
3461:Black, R. (28 January 2016).
3440:10.1080/02724634.2020.1793158
3315:
640:in the shoulder girdle, both
4525:10.5710/AMGH.20.08.2020.3376
4087:10.1080/08912963.2012.672416
3964:10.1371/journal.pone.0078733
3916:10.1080/08912960410001715132
3847:Wedel, M. (10 August 2017).
3487:Mach, A. (16 January 2016).
2678:(instead of the less stable
2662:closer to the Rinconsauria.
1008:from other titanosaurs. The
938:
511:
372:, which was first announced
7:
4619:Harding, G. (17 May 2014).
4210:10.1007/978-3-030-95959-3_8
3813:Wedel, M. (9 August 2017).
3787:Black, R. (9 August 2017).
3544:Johnson, S. (11 May 2018).
3303:
3213:, was also named in 2018.
1571:on its outer top end. Like
1184:, ilio-ischiocaudalis, and
1081:, and other members of the
753:
722:named these remains as the
571:US ambassador to Argentina
10:
9560:
9544:Early Cretaceous sauropods
4593:Mohney, G. (17 May 2014).
4334:Royal Society Open Science
3849:"Some further thoughts on
3628:Morgan, J. (17 May 2014).
3298:Raising the Dinosaur Giant
1587:, and saltasaurines. Like
1094:at the rear end but not a
16:Extinct genus of dinosaurs
9427:
9351:
9340:
9308:
9210:
9119:
9045:
9007:
8998:
8962:
8928:
8846:
8780:
8729:
8720:
8647:
8638:
8616:
8605:
8540:
8531:
8146:
8142:
8127:
8114:
8081:
8047:
7999:
7990:
7862:
7684:
7675:
7584:
7519:
7508:
7477:
7410:
7406:
7391:
7378:
7358:
7270:
7227:
7218:
7120:
7111:
7048:
7013:
7004:
6910:
6901:
6890:
6875:
6860:
6847:
6811:
6737:
6709:
6693:
6684:
6575:
6419:
6368:
6248:
6237:
6206:
6153:
6149:
6138:
6093:
6012:
6008:
5993:
5980:
5946:
5900:
5815:
5806:
5747:
5738:
5646:
5619:
5562:
5551:
5492:
5463:
5452:
5414:
5357:
5353:
5342:
5272:
5215:
5211:
5196:
5183:
5151:
5133:
5120:
5068:
4929:American Museum Novitates
4796:10.1007/s00114-005-0623-3
4720:10.1163/22941932-20170206
4251:10.1093/zoolinnean/zlx103
3815:"Don't believe the hype:
3060:
3043:
3036:
3019:
3012:
2965:
2948:
2941:
2921:
2914:
2897:
2890:
2880:
2863:
2856:
2818:
2798:
2788:
2768:
2761:
2751:
2731:
2721:
2704:
2697:
2646:possibly being closer to
2446:
2429:
2422:
2405:
2388:
2381:
2355:
2338:
2331:
2324:
2307:
2300:
2280:
2273:
2256:
2249:
2205:
2188:
2181:
2164:
2157:
2140:
2133:
2123:
2052:
2035:
2028:
2011:
2004:
1987:
1970:
1953:
1936:
1919:
1902:
1895:
1878:
1871:
1854:
1847:
1827:
1820:
1813:
1793:
1786:
1760:
1743:
1736:
1729:
1712:
1695:
1688:
1636:to its closest relative,
431:and other members of the
317:
310:
139:Scientific classification
137:
124:
115:
23:
9366:List of sauropod species
3416:(Dinosauria, Sauropoda)"
3150:Life restoration of two
1396:latissimus dorsi muscles
789:, where it replaced the
386:largest known titanosaur
9524:Cerro Barcino Formation
4548:Rinconsaurus caudamirus
4235:Mendozasaurus neguyelap
1553:but somewhat less than
1384:coracobrachialis muscle
811:
700:Ignacio Alejandro Cerda
554:Cerro Barcino Formation
542:The Jurassic Foundation
356:Cerro Barcino Formation
4283:(suppl 2): e20180374.
3603:Natural History Museum
3368:10.1098/rspb.2017.1219
3284:Sir David Attenborough
3185:similar to the living
3155:
1460:
1215:
1206:Limbs and limb girdles
1115:
993:cannot be determined.
948:
845:
824:
767:
708:Alberto Carlos Garrido
580:
504:that was dominated by
9466:Paleobiology Database
9356:Sauropod neck posture
9047:Opisthocoelicaudiinae
4172:10.4202/app.2010.0011
3200:Austrocupressinoxylon
3165:specimens, including
3149:
2795:Opisthocoelicaudiinae
1626:phylogenetic analysis
1612:; this was where the
1458:
1213:
1182:caudofemoralis longus
1113:
946:
841:
823:compared with a human
819:
761:
720:Javier Marcelo Krause
628:, both plates of the
570:
471:in the forelimb, and
418:phylogenetic analyses
9519:Fossils of Argentina
9514:Cretaceous Argentina
4387:Europasaurus holgeri
4153:Bonitasaura salgadoi
3819:was not bigger than
3709:10.2992/007.085.0403
3467:Smithsonian Magazine
1387:
1375:
1260:
1252:
1147:
1119:
1095:
1087:
1047:
1046:at the front to the
1039:
1031:
1009:
964:
956:
859:was 10% larger than
688:
680:
672:
664:
656:
641:
633:
621:
613:
605:
597:
589:
538:Jose Luis Carballido
517:
488:
480:
472:
464:
456:
448:
440:
9215:Dubious titanosaurs
6032:Chinshakiangosaurus
4888:2009CrRes..30.1376L
4876:Cretaceous Research
4788:2005NW.....92..226N
4776:Naturwissenschaften
4663:2019CrRes..99..133N
4651:Cretaceous Research
4564:2023CrRes.14205389P
4552:Cretaceous Research
4482:2021CrRes.12204754O
4470:Cretaceous Research
4439:2023CrRes.14605487A
4427:Cretaceous Research
4354:10.1098/rsos.191057
4346:2019RSOS....691057M
4079:2012HBio...24..631C
4017:2016NatSR...619165G
3955:2013PLoSO...878733S
3898:2004HBio...16...71M
3688:Paul, G.S. (2019).
3519:Wall Street Journal
3432:2020JVPal..40E3158O
3414:Patagotitan mayorum
1390:deltopectoral crest
1374:. In a depression (
1034:articular processes
728:Patagotitan mayorum
369:Patagotitan mayorum
323:Patagotitan mayorum
9087:Opisthocoelicaudia
6159:Archaeodontosaurus
4067:Historical Biology
4005:Scientific Reports
3886:Historical Biology
3736:Biological Reviews
3362:(1860): 20171219.
3278:In popular culture
3156:
2802:Opisthocoelicaudia
1957:Opisthocoelicaudia
1569:greater trochanter
1561:Opisthocoelicaudia
1461:
1428:Opisthocoelicaudia
1412:Opisthocoelicaudia
1368:Opisthocoelicaudia
1336:Opisthocoelicaudia
1216:
1116:
949:
825:
768:
716:NĂ©stor Ruben CĂşneo
581:
516:A part of a lower
9539:2014 in Argentina
9481:
9480:
9421:Taxon identifiers
9378:
9377:
9374:
9373:
9336:
9335:
9306:
9305:
9302:
9301:
9298:
9297:
9223:Bruhathkayosaurus
9206:
9205:
9202:
9201:
9198:
9197:
8924:
8923:
8842:
8841:
8764:Panamericansaurus
8601:
8600:
8597:
8596:
8110:
8109:
8079:
8078:
8075:
8074:
8071:
8070:
8067:
8066:
8043:
8042:
7511:Titanosauriformes
7504:
7503:
7374:
7373:
7356:
7355:
7352:
7351:
7348:
7347:
7344:
7343:
7340:
7339:
7152:Brachytrachelopan
7107:
7106:
7103:
7102:
6881:Haplocanthosaurus
6843:
6842:
6809:
6808:
6805:
6804:
6801:
6800:
6742:Dubious sauropods
6733:
6732:
6729:
6728:
6725:
6724:
6233:
6232:
6134:
6133:
6069:Protognathosaurus
5976:
5975:
5944:
5943:
5940:
5939:
5936:
5935:
5932:
5931:
5928:
5927:
5924:
5923:
5920:
5919:
5547:
5546:
5448:
5447:
5338:
5337:
5179:
5178:
5175:
5174:
5171:
5170:
4837:Genyodectes serus
4219:978-3-030-95958-6
4025:10.1038/srep19165
3749:10.1111/brv.12638
3525:on 5 October 2021
3310:Largest dinosaurs
3216:The fine-grained
3138:
3137:
3129:
3128:
3120:
3119:
3111:
3110:
3102:
3101:
3093:
3092:
3084:
3083:
3075:
3074:
2998:
2997:
2989:
2988:
2980:
2979:
2842:
2841:
2833:
2832:
2560:
2559:
2551:
2550:
2542:
2541:
2533:
2532:
2524:
2523:
2515:
2514:
2506:
2505:
2497:
2496:
2488:
2487:
2479:
2478:
2470:
2469:
2461:
2460:
2370:
2369:
2238:
2237:
2229:
2228:
2220:
2219:
2112:
2111:
2103:
2102:
2094:
2093:
2085:
2084:
2076:
2075:
2067:
2066:
1775:
1774:
1675:phylogenetic tree
1605:fourth trochanter
1507:obturator foramen
1065:Titanosauriformes
712:Jahandar Ramezani
586:holotype specimen
337:
336:
306:
131:Chicago, Illinois
9551:
9474:
9473:
9461:
9460:
9448:
9447:
9446:
9416:
9415:
9407:
9406:
9405:
9395:
9394:
9393:
9386:
9342:
9341:
9331:
9325:
9320:
9314:
9212:
9211:
9094:Pellegrinisaurus
9005:
9004:
8960:
8959:
8939:Bustingorrytitan
8879:Futalognkosaurus
8727:
8726:
8645:
8644:
8614:
8613:
8538:
8537:
8521:Diamantinasauria
8413:Pellegrinisaurus
8319:Jiangshanosaurus
8210:Baotianmansaurus
8144:
8143:
8140:
8139:
8129:
8128:
8116:
8115:
8105:
8104:
8097:
8096:
8089:
8088:
8012:Diamantinasaurus
7997:
7996:
7992:Diamantinasauria
7931:Jiangshanosaurus
7682:
7681:
7517:
7516:
7408:
7407:
7404:
7403:
7393:
7392:
7380:
7379:
7369:
7364:
7225:
7224:
7118:
7117:
7050:Rebbachisaurinae
7011:
7010:
6908:
6907:
6903:Rebbachisauridae
6899:
6898:
6873:
6872:
6862:
6861:
6849:
6848:
6838:
6832:
6827:
6826:
6819:
6818:
6750:Bothriospondylus
6739:
6738:
6691:
6690:
6467:Eomamenchisaurus
6421:Mamenchisauridae
6401:Lapparentosaurus
6345:Spinophorosaurus
6246:
6245:
6151:
6150:
6147:
6146:
6010:
6009:
6006:
6005:
5995:
5994:
5982:
5981:
5971:
5970:
5963:
5958:
5952:
5813:
5812:
5745:
5744:
5648:Massospondylidae
5560:
5559:
5490:
5489:
5461:
5460:
5425:Jaklapallisaurus
5405:Thecodontosaurus
5355:
5354:
5351:
5350:
5213:
5212:
5209:
5208:
5198:
5197:
5185:
5184:
5131:
5130:
5122:
5121:
5115:
5114:
5070:
5069:
5055:
5048:
5041:
5032:
5031:
4991:
4990:
4988:
4986:
4972:
4966:
4965:
4963:
4961:
4947:
4941:
4940:
4916:
4910:
4909:
4907:
4882:(6): 1376–1386.
4871:
4865:
4864:
4832:
4826:
4825:
4807:
4771:
4765:
4764:
4748:
4742:
4741:
4731:
4699:
4693:
4692:
4682:
4642:
4636:
4635:
4633:
4631:
4616:
4610:
4609:
4607:
4605:
4590:
4584:
4583:
4543:
4537:
4536:
4508:
4502:
4501:
4465:
4459:
4458:
4421:
4415:
4414:
4382:
4376:
4375:
4365:
4325:
4319:
4318:
4300:
4271:
4265:
4264:
4262:
4230:
4224:
4223:
4196:
4187:
4186:
4184:
4174:
4146:
4140:
4139:
4129:
4105:
4099:
4098:
4061:
4055:
4054:
4044:
3996:
3987:
3986:
3976:
3966:
3934:
3928:
3927:
3909:
3883:
3874:
3868:
3867:
3865:
3863:
3844:
3838:
3837:
3835:
3833:
3810:
3804:
3803:
3801:
3799:
3784:
3778:
3777:
3751:
3742:(6): 1759–1797.
3727:
3721:
3720:
3694:
3685:
3674:
3673:
3671:
3669:
3664:. 10 August 2017
3654:
3645:
3644:
3642:
3640:
3625:
3614:
3613:
3611:
3609:
3595:
3589:
3588:
3586:
3584:
3567:
3561:
3560:
3558:
3556:
3541:
3535:
3534:
3532:
3530:
3521:. Archived from
3510:
3504:
3503:
3501:
3499:
3484:
3478:
3477:
3475:
3473:
3458:
3452:
3451:
3407:
3390:
3389:
3379:
3347:
3039:
3038:
3023:Futalognkosaurus
3015:
3014:
2944:
2943:
2917:
2916:
2893:
2892:
2883:
2882:
2859:
2858:
2791:
2790:
2764:
2763:
2754:
2753:
2735:Diamantinasaurus
2728:Diamantinasauria
2724:
2723:
2700:
2699:
2690:
2689:
2618:Futalognkosaurus
2425:
2424:
2384:
2383:
2342:Futalognkosaurus
2334:
2333:
2327:
2326:
2303:
2302:
2276:
2275:
2252:
2251:
2184:
2183:
2160:
2159:
2136:
2135:
2126:
2125:
2031:
2030:
2007:
2006:
1898:
1897:
1874:
1873:
1850:
1849:
1823:
1822:
1816:
1815:
1789:
1788:
1739:
1738:
1732:
1731:
1691:
1690:
1681:
1680:
1585:Diamantinasaurus
1556:Diamantinasaurus
1530:Jiangshanosaurus
1515:Futalognkosaurus
1469:Futalognkosaurus
1391:
1379:
1264:
1256:
1190:Futalognkosaurus
1162:Futalognkosaurus
1151:
1123:
1099:
1091:
1051:
1043:
1035:
1013:
998:Futalognkosaurus
978:Futalognkosaurus
968:
960:
910:
853:
726:of a new genus,
704:Leonardo Salgado
692:
684:
676:
668:
660:
645:
637:
625:
617:
609:
601:
593:
521:
506:coniferous trees
492:
484:
476:
468:
460:
452:
444:
320:
300:
293:
280:
264:
248:
232:
216:
147:
146:
120:
110:
40:
29:Temporal range:
21:
20:
9559:
9558:
9554:
9553:
9552:
9550:
9549:
9548:
9534:2014 in science
9484:
9483:
9482:
9477:
9469:
9464:
9456:
9451:
9442:
9441:
9436:
9423:
9413:
9403:
9401:
9391:
9389:
9381:
9379:
9370:
9361:Sauropod hiatus
9347:
9332:
9294:
9231:Campylodoniscus
9194:
9115:
9041:
8994:
8964:Nemegtosauridae
8958:
8920:
8857:Argentinosaurus
8838:
8776:
8716:
8634:
8610:
8593:
8553:Atsinganosaurus
8533:Lirainosaurinae
8527:
8514:Xianshanosaurus
8399:Normanniasaurus
8385:Narambuenatitan
8268:Dongyangosaurus
8136:
8123:
8106:
8099:
8091:
8083:
8063:
8039:
8019:Sarmientosaurus
7986:
7939:Phuwiangosaurus
7858:
7671:
7586:Brachiosauridae
7580:
7513:
7500:
7473:
7400:
7387:
7370:
7336:
7266:
7214:
7122:Dicraeosauridae
7113:Flagellicaudata
7099:
7044:
7000:
6974:Nopcsaspondylus
6966:Maraapunisaurus
6895:
6893:Diplodocimorpha
6886:
6869:
6856:
6839:
6821:
6813:
6797:
6721:
6705:
6680:
6653:Tehuelchesaurus
6571:
6531:Tienshanosaurus
6415:
6364:
6352:Tehuelchesaurus
6242:
6229:
6208:Vulcanodontidae
6202:
6143:
6130:
6089:
6039:Gongxianosaurus
6002:
5989:
5972:
5965:
5916:
5896:
5802:
5761:Jingshanosaurus
5753:Chuxiongosaurus
5740:Sauropodiformes
5734:
5657:Adeopapposaurus
5642:
5615:
5556:
5543:
5488:
5457:
5444:
5410:
5347:
5334:
5268:
5205:
5203:Sauropodomorpha
5192:
5190:Sauropodomorpha
5167:
5153:Sauropodomorpha
5147:
5141:Avemetatarsalia
5126:Avemetatarsalia
5116:
5101:Avemetatarsalia
5075:
5074:
5064:
5062:Sauropodomorpha
5059:
5017:
5000:
4998:Further reading
4995:
4994:
4984:
4982:
4974:
4973:
4969:
4959:
4957:
4949:
4948:
4944:
4917:
4913:
4872:
4868:
4833:
4829:
4772:
4768:
4749:
4745:
4704:Carlquistoxylon
4700:
4696:
4643:
4639:
4629:
4627:
4617:
4613:
4603:
4601:
4591:
4587:
4544:
4540:
4509:
4505:
4466:
4462:
4422:
4418:
4383:
4379:
4326:
4322:
4272:
4268:
4231:
4227:
4220:
4197:
4190:
4147:
4143:
4106:
4102:
4062:
4058:
3997:
3990:
3935:
3931:
3907:10.1.1.694.1650
3881:
3875:
3871:
3861:
3859:
3845:
3841:
3831:
3829:
3821:Argentinosaurus
3811:
3807:
3797:
3795:
3785:
3781:
3728:
3724:
3692:
3686:
3677:
3667:
3665:
3656:
3655:
3648:
3638:
3636:
3626:
3617:
3607:
3605:
3597:
3596:
3592:
3582:
3580:
3568:
3564:
3554:
3552:
3550:Chicago Tribune
3542:
3538:
3528:
3526:
3511:
3507:
3497:
3495:
3485:
3481:
3471:
3469:
3459:
3455:
3426:(4): e1793158.
3408:
3393:
3348:
3323:
3318:
3306:
3280:
3206:Carlquistoxylon
3144:
3139:
3130:
3121:
3112:
3103:
3094:
3085:
3076:
2999:
2990:
2981:
2843:
2834:
2652:Argentinosaurus
2630:Argentinosaurus
2614:Argentinosaurus
2593:Argentinosaurus
2569:Argentinosaurus
2561:
2552:
2543:
2534:
2525:
2516:
2507:
2498:
2489:
2480:
2471:
2462:
2433:Argentinosaurus
2371:
2239:
2230:
2221:
2113:
2104:
2095:
2086:
2077:
2068:
1776:
1638:Argentinosaurus
1622:
1610:lateral condyle
1393:
1381:
1342:The humerus of
1266:
1258:
1208:
1153:
1125:
1101:
1093:
1061:Argentinosaurus
1053:
1045:
1042:prezygapophyses
1037:
1022:Argentinosaurus
1015:
970:
962:
941:
933:Argentinosaurus
917:Argentinosaurus
913:Argentinosaurus
908:
906:Argentinosaurus
898:Argentinosaurus
875:Argentinosaurus
867:Gregory S. Paul
862:Argentinosaurus
851:
814:
809:
756:
694:
686:
678:
670:
662:
647:
639:
636:scapulocoracoid
627:
619:
611:
603:
595:
573:Marc R. Stanley
523:
514:
494:
486:
478:
470:
462:
454:
446:
429:Argentinosaurus
403:Argentinosaurus
376:and then named
360:Chubut Province
333:
326:
299:
291:
278:
262:
246:
230:
218:Sauropodomorpha
214:
141:
111:
109:
108:
107:
106:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
35:
34:
33:
27:
17:
12:
11:
5:
9557:
9547:
9546:
9541:
9536:
9531:
9526:
9521:
9516:
9511:
9506:
9501:
9499:Lithostrotians
9496:
9479:
9478:
9476:
9475:
9462:
9449:
9433:
9431:
9425:
9424:
9412:
9411:
9399:
9376:
9375:
9372:
9371:
9369:
9368:
9363:
9358:
9352:
9349:
9348:
9338:
9337:
9334:
9333:
9309:
9307:
9304:
9303:
9300:
9299:
9296:
9295:
9293:
9292:
9285:
9277:
9270:
9262:
9255:
9248:
9241:
9238:Clasmodosaurus
9234:
9227:
9218:
9216:
9208:
9207:
9204:
9203:
9200:
9199:
9196:
9195:
9193:
9192:
9185:
9178:
9171:
9164:
9157:
9153:Maxakalisaurus
9149:
9142:
9134:
9125:
9123:
9117:
9116:
9114:
9113:
9105:
9098:
9090:
9083:
9075:
9067:
9060:
9051:
9049:
9043:
9042:
9040:
9039:
9035:Quaesitosaurus
9031:
9024:
9016:
9008:
9002:
8996:
8995:
8993:
8992:
8984:
8980:Quaesitosaurus
8976:
8968:
8966:
8957:
8956:
8949:
8942:
8934:
8932:
8930:Saltasauroidea
8926:
8925:
8922:
8921:
8919:
8918:
8911:
8904:
8897:
8890:
8882:
8875:
8867:
8860:
8852:
8850:
8844:
8843:
8840:
8839:
8837:
8836:
8829:
8822:
8815:
8808:
8801:
8794:
8786:
8784:
8778:
8777:
8775:
8774:
8767:
8760:
8753:
8746:
8742:Maxakalisaurus
8738:
8735:Adamantisaurus
8730:
8724:
8718:
8717:
8715:
8714:
8707:
8699:
8692:
8685:
8681:Epachthosaurus
8677:
8670:
8663:
8656:
8653:Antarctosaurus
8648:
8642:
8636:
8635:
8633:
8632:
8625:
8617:
8611:
8608:Eutitanosauria
8606:
8603:
8602:
8599:
8598:
8595:
8594:
8592:
8591:
8584:
8581:Mansourasaurus
8577:
8570:
8563:
8556:
8549:
8541:
8535:
8529:
8528:
8526:
8525:
8517:
8510:
8503:
8496:
8489:
8482:
8475:
8468:
8460:
8456:Tastavinsaurus
8452:
8445:
8438:
8431:
8424:
8421:Petrobrasaurus
8417:
8409:
8402:
8395:
8388:
8381:
8374:
8367:
8363:Malarguesaurus
8359:
8352:
8344:
8337:
8330:
8323:
8315:
8308:
8301:
8294:
8290:Epachthosaurus
8286:
8279:
8271:
8264:
8257:
8250:
8243:
8236:
8228:
8220:
8213:
8206:
8203:Austroposeidon
8199:
8192:
8185:
8178:
8170:
8163:
8155:
8147:
8137:
8132:
8125:
8124:
8112:
8111:
8108:
8107:
8082:
8080:
8077:
8076:
8073:
8072:
8069:
8068:
8065:
8064:
8062:
8061:
8060:
8059:
8053:
8051:
8045:
8044:
8041:
8040:
8038:
8037:
8029:
8022:
8015:
8008:
8000:
7994:
7988:
7987:
7985:
7984:
7977:
7970:
7967:Tangvayosaurus
7963:
7956:
7949:
7942:
7935:
7927:
7920:
7913:
7906:
7899:
7892:
7885:
7877:
7868:
7866:
7860:
7859:
7857:
7856:
7849:
7846:Tastavinsaurus
7842:
7835:
7828:
7821:
7818:Pukyongosaurus
7814:
7807:
7800:
7797:Liubangosaurus
7793:
7785:
7782:Liaoningotitan
7778:
7771:
7763:
7756:
7749:
7742:
7735:
7728:
7721:
7714:
7707:
7704:Astrophocaudia
7700:
7693:
7685:
7679:
7677:Somphospondyli
7673:
7672:
7670:
7669:
7662:
7655:
7648:
7641:
7634:
7627:
7619:
7612:
7605:
7598:
7590:
7588:
7582:
7581:
7579:
7578:
7570:
7563:
7556:
7549:
7542:
7535:
7528:
7520:
7514:
7509:
7506:
7505:
7502:
7501:
7499:
7498:
7495:Lourinhasaurus
7491:
7483:
7481:
7479:Camarasauridae
7475:
7474:
7472:
7471:
7464:
7456:
7449:
7445:Dashanpusaurus
7441:
7434:
7426:
7419:
7411:
7401:
7396:
7389:
7388:
7376:
7375:
7372:
7371:
7359:
7357:
7354:
7353:
7350:
7349:
7346:
7345:
7342:
7341:
7338:
7337:
7335:
7334:
7327:
7320:
7313:
7305:
7298:
7291:
7288:Dinheirosaurus
7284:
7276:
7274:
7268:
7267:
7265:
7264:
7257:
7249:
7242:
7233:
7231:
7222:
7216:
7215:
7213:
7212:
7205:
7198:
7191:
7184:
7177:
7170:
7162:
7155:
7148:
7141:
7134:
7126:
7124:
7115:
7109:
7108:
7105:
7104:
7101:
7100:
7098:
7097:
7090:
7087:Rebbachisaurus
7083:
7076:
7069:
7062:
7054:
7052:
7046:
7045:
7043:
7042:
7034:
7027:
7019:
7017:
7008:
7002:
7001:
6999:
6998:
6991:
6984:
6977:
6970:
6962:
6959:Lavocatisaurus
6955:
6948:
6940:
6933:
6926:
6919:
6911:
6905:
6896:
6891:
6888:
6887:
6885:
6884:
6876:
6870:
6865:
6858:
6857:
6845:
6844:
6841:
6840:
6812:
6810:
6807:
6806:
6803:
6802:
6799:
6798:
6796:
6795:
6788:
6785:Qinlingosaurus
6781:
6774:
6767:
6760:
6753:
6745:
6743:
6735:
6734:
6731:
6730:
6727:
6726:
6723:
6722:
6720:
6719:
6718:(see below ↓ )
6715:
6713:
6707:
6706:
6704:
6703:
6702:(see below ↓ )
6699:
6697:
6688:
6682:
6681:
6679:
6678:
6671:
6664:
6657:
6649:
6641:
6634:
6627:
6620:
6613:
6605:
6597:
6589:
6581:
6579:
6573:
6572:
6570:
6569:
6562:
6555:
6548:
6541:
6534:
6527:
6524:Rhomaleopakhus
6520:
6513:
6506:
6503:Mamenchisaurus
6499:
6491:
6484:
6477:
6470:
6463:
6455:
6452:Chuanjiesaurus
6448:
6444:Cetiosauriscus
6440:
6433:
6425:
6423:
6417:
6416:
6414:
6413:
6405:
6397:
6389:
6382:
6374:
6372:
6366:
6365:
6363:
6362:
6355:
6348:
6341:
6334:
6327:
6320:
6313:
6306:
6299:
6296:Dashanpusaurus
6292:
6289:Cetiosauriscus
6285:
6278:
6271:
6264:
6257:
6249:
6243:
6238:
6235:
6234:
6231:
6230:
6228:
6227:
6220:
6212:
6210:
6204:
6203:
6201:
6200:
6196:Zizhongosaurus
6192:
6184:
6177:
6170:
6163:
6154:
6144:
6139:
6136:
6135:
6132:
6131:
6129:
6128:
6121:
6114:
6107:
6099:
6097:
6095:Lessemsauridae
6091:
6090:
6088:
6087:
6083:Tuebingosaurus
6079:
6072:
6065:
6057:
6049:
6042:
6035:
6028:
6021:
6013:
6003:
5998:
5991:
5990:
5978:
5977:
5974:
5973:
5947:
5945:
5942:
5941:
5938:
5937:
5934:
5933:
5930:
5929:
5926:
5925:
5922:
5921:
5918:
5917:
5915:
5914:
5913:
5912:
5906:
5904:
5898:
5897:
5895:
5894:
5891:Sefapanosaurus
5887:
5880:
5873:
5866:
5863:Melanorosaurus
5859:
5852:
5845:
5838:
5831:
5824:
5816:
5810:
5804:
5803:
5801:
5800:
5793:
5786:
5778:
5771:
5764:
5757:
5748:
5742:
5736:
5735:
5733:
5732:
5724:
5717:
5710:
5703:
5700:Massospondylus
5696:
5689:
5682:
5674:
5671:Glacialisaurus
5667:
5664:Coloradisaurus
5660:
5652:
5650:
5644:
5643:
5641:
5640:
5633:
5625:
5623:
5617:
5616:
5614:
5613:
5610:Tuebingosaurus
5606:
5599:
5592:
5585:
5578:
5571:
5563:
5557:
5552:
5549:
5548:
5545:
5544:
5542:
5541:
5534:
5527:
5524:Pachysauriscus
5520:
5513:
5510:Gresslyosaurus
5506:
5498:
5496:
5494:Plateosauridae
5487:
5486:
5479:
5472:
5469:Plateosauravus
5464:
5458:
5453:
5450:
5449:
5446:
5445:
5443:
5442:
5435:
5428:
5420:
5418:
5412:
5411:
5409:
5408:
5401:
5394:
5387:
5380:
5373:
5366:
5358:
5348:
5343:
5340:
5339:
5336:
5335:
5333:
5332:
5325:
5317:
5309:
5302:
5299:Chromogisaurus
5295:
5287:
5278:
5276:
5270:
5269:
5267:
5266:
5259:
5252:
5245:
5238:
5231:
5224:
5216:
5206:
5201:
5194:
5193:
5181:
5180:
5177:
5176:
5173:
5172:
5169:
5168:
5166:
5165:
5164:
5163:
5157:
5155:
5149:
5148:
5146:
5145:
5144:
5143:
5134:
5128:
5118:
5117:
5113:
5112:
5103:
5094:
5088:
5082:
5066:
5065:
5058:
5057:
5050:
5043:
5035:
5029:
5028:
5023:
5016:
5015:External links
5013:
5012:
5011:
4999:
4996:
4993:
4992:
4967:
4942:
4931:(3599): 1–35.
4911:
4866:
4847:(4): 894–902.
4827:
4782:(5): 226–230.
4766:
4743:
4714:(4): 406–426.
4694:
4637:
4611:
4585:
4538:
4503:
4460:
4416:
4397:(9): 739–781.
4377:
4320:
4266:
4245:(1): 136–181.
4225:
4218:
4188:
4141:
4120:(3): 529–541.
4100:
4073:(6): 631–654.
4056:
3988:
3949:(10): e78733.
3929:
3869:
3839:
3805:
3779:
3722:
3703:(4): 335–358.
3675:
3646:
3615:
3590:
3562:
3536:
3505:
3479:
3453:
3391:
3320:
3319:
3317:
3314:
3313:
3312:
3305:
3302:
3279:
3276:
3194:Pilgerodendron
3143:
3140:
3136:
3135:
3132:
3131:
3127:
3126:
3123:
3122:
3118:
3117:
3114:
3113:
3109:
3108:
3105:
3104:
3100:
3099:
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3091:
3090:
3087:
3086:
3082:
3081:
3078:
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3073:
3072:
3069:
3068:
3059:
3056:
3055:
3052:
3051:
3042:
3037:
3035:
3032:
3031:
3028:
3027:
3018:
3013:
3011:
3005:
3004:
3001:
3000:
2996:
2995:
2992:
2991:
2987:
2986:
2983:
2982:
2978:
2977:
2974:
2973:
2964:
2961:
2960:
2957:
2956:
2947:
2942:
2940:
2934:
2933:
2930:
2929:
2920:
2915:
2913:
2910:
2909:
2906:
2905:
2896:
2891:
2889:
2881:
2879:
2876:
2875:
2872:
2871:
2867:Epachthosaurus
2862:
2857:
2855:
2849:
2848:
2845:
2844:
2840:
2839:
2836:
2835:
2831:
2830:
2827:
2826:
2817:
2811:
2810:
2807:
2806:
2797:
2789:
2787:
2781:
2780:
2777:
2776:
2767:
2762:
2760:
2758:Saltasauroidea
2752:
2750:
2748:Eutitanosauria
2744:
2743:
2740:
2739:
2730:
2722:
2720:
2717:
2716:
2713:
2712:
2703:
2698:
2696:
2688:
2558:
2557:
2554:
2553:
2549:
2548:
2545:
2544:
2540:
2539:
2536:
2535:
2531:
2530:
2527:
2526:
2522:
2521:
2518:
2517:
2513:
2512:
2509:
2508:
2504:
2503:
2500:
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2485:
2482:
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2477:
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2473:
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2468:
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2463:
2459:
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2454:
2445:
2442:
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2438:
2437:
2428:
2423:
2421:
2418:
2417:
2414:
2413:
2404:
2401:
2400:
2397:
2396:
2387:
2382:
2380:
2377:
2376:
2373:
2372:
2368:
2367:
2364:
2363:
2354:
2351:
2350:
2347:
2346:
2337:
2332:
2330:
2325:
2323:
2320:
2319:
2316:
2315:
2306:
2301:
2299:
2293:
2292:
2289:
2288:
2279:
2274:
2272:
2269:
2268:
2265:
2264:
2255:
2250:
2248:
2245:
2244:
2241:
2240:
2236:
2235:
2232:
2231:
2227:
2226:
2223:
2222:
2218:
2217:
2214:
2213:
2204:
2201:
2200:
2197:
2196:
2187:
2182:
2180:
2177:
2176:
2173:
2172:
2163:
2158:
2156:
2153:
2152:
2149:
2148:
2139:
2134:
2132:
2124:
2122:
2119:
2118:
2115:
2114:
2110:
2109:
2106:
2105:
2101:
2100:
2097:
2096:
2092:
2091:
2088:
2087:
2083:
2082:
2079:
2078:
2074:
2073:
2070:
2069:
2065:
2064:
2061:
2060:
2051:
2048:
2047:
2044:
2043:
2034:
2029:
2027:
2024:
2023:
2020:
2019:
2010:
2005:
2003:
2000:
1999:
1996:
1995:
1986:
1983:
1982:
1979:
1978:
1969:
1966:
1965:
1962:
1961:
1952:
1949:
1948:
1945:
1944:
1935:
1932:
1931:
1928:
1927:
1918:
1915:
1914:
1911:
1910:
1901:
1896:
1894:
1891:
1890:
1887:
1886:
1877:
1872:
1870:
1867:
1866:
1863:
1862:
1853:
1848:
1846:
1840:
1839:
1836:
1835:
1826:
1821:
1819:
1814:
1812:
1810:Eutitanosauria
1806:
1805:
1802:
1801:
1797:Epachthosaurus
1792:
1787:
1785:
1782:
1781:
1778:
1777:
1773:
1772:
1769:
1768:
1764:Malarguesaurus
1759:
1756:
1755:
1752:
1751:
1742:
1737:
1735:
1730:
1728:
1725:
1724:
1721:
1720:
1711:
1708:
1707:
1704:
1703:
1694:
1689:
1687:
1679:
1671:Eutitanosauria
1621:
1620:Classification
1618:
1436:biceps brachii
1422:differed from
1364:Epachthosaurus
1207:
1204:
1137:Epachthosaurus
940:
937:
813:
810:
808:
805:
755:
752:
632:and the right
608:tail vertebrae
600:back vertebrae
592:neck vertebrae
513:
510:
451:tail vertebrae
344:is a genus of
335:
334:
327:
315:
314:
308:
307:
289:
285:
284:
276:
269:
268:
260:
253:
252:
244:
237:
236:
228:
221:
220:
212:
205:
204:
199:
192:
191:
186:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
135:
134:
122:
121:
113:
112:
104:
103:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
28:
15:
9:
6:
4:
3:
2:
9556:
9545:
9542:
9540:
9537:
9535:
9532:
9530:
9527:
9525:
9522:
9520:
9517:
9515:
9512:
9510:
9507:
9505:
9502:
9500:
9497:
9495:
9492:
9491:
9489:
9472:
9467:
9463:
9459:
9454:
9450:
9445:
9439:
9435:
9434:
9432:
9430:
9426:
9422:
9417:
9410:
9400:
9398:
9388:
9387:
9384:
9367:
9364:
9362:
9359:
9357:
9354:
9353:
9350:
9343:
9339:
9330:
9326:
9324:
9319:
9313:
9291:
9290:
9286:
9283:
9282:
9281:Mongolosaurus
9278:
9276:
9275:
9271:
9268:
9267:
9266:Macrurosaurus
9263:
9261:
9260:
9256:
9254:
9253:
9249:
9247:
9246:
9245:Hypselosaurus
9242:
9240:
9239:
9235:
9233:
9232:
9228:
9225:
9224:
9220:
9219:
9217:
9213:
9209:
9191:
9190:
9186:
9184:
9183:
9179:
9177:
9176:
9172:
9170:
9169:
9165:
9163:
9162:
9161:Neuquensaurus
9158:
9155:
9154:
9150:
9148:
9147:
9143:
9140:
9139:
9135:
9132:
9131:
9127:
9126:
9124:
9122:
9121:Saltasaurinae
9118:
9111:
9110:
9109:Zhuchengtitan
9106:
9104:
9103:
9099:
9096:
9095:
9091:
9089:
9088:
9084:
9081:
9080:
9079:Dreadnoughtus
9076:
9073:
9072:
9068:
9066:
9065:
9061:
9058:
9057:
9053:
9052:
9050:
9048:
9044:
9037:
9036:
9032:
9030:
9029:
9028:Qingxiusaurus
9025:
9022:
9021:
9017:
9015:
9014:
9010:
9009:
9006:
9003:
9001:
9000:Saltasauridae
8997:
8990:
8989:
8985:
8982:
8981:
8977:
8975:
8974:
8973:Nemegtosaurus
8970:
8969:
8967:
8965:
8961:
8955:
8954:
8950:
8948:
8947:
8943:
8941:
8940:
8936:
8935:
8933:
8931:
8927:
8917:
8916:
8912:
8910:
8909:
8905:
8903:
8902:
8898:
8896:
8895:
8894:Mendozasaurus
8891:
8888:
8887:
8883:
8881:
8880:
8876:
8873:
8872:
8871:Dzharatitanis
8868:
8866:
8865:
8861:
8859:
8858:
8854:
8853:
8851:
8849:
8845:
8835:
8834:
8830:
8828:
8827:
8823:
8821:
8820:
8819:Gondwanatitan
8816:
8814:
8813:
8809:
8807:
8806:
8802:
8800:
8799:
8795:
8793:
8792:
8788:
8787:
8785:
8783:
8779:
8773:
8772:
8768:
8766:
8765:
8761:
8759:
8758:
8754:
8752:
8751:
8750:Muyelensaurus
8747:
8744:
8743:
8739:
8737:
8736:
8732:
8731:
8728:
8725:
8723:
8719:
8713:
8712:
8708:
8705:
8704:
8700:
8698:
8697:
8693:
8691:
8690:
8686:
8683:
8682:
8678:
8676:
8675:
8674:Chucarosaurus
8671:
8669:
8668:
8664:
8662:
8661:
8657:
8655:
8654:
8650:
8649:
8646:
8643:
8641:
8640:Colossosauria
8637:
8631:
8630:
8629:Menucocelsior
8626:
8624:
8623:
8619:
8618:
8615:
8612:
8609:
8604:
8590:
8589:
8585:
8583:
8582:
8578:
8576:
8575:
8571:
8569:
8568:
8567:Lirainosaurus
8564:
8562:
8561:
8557:
8555:
8554:
8550:
8548:
8547:
8543:
8542:
8539:
8536:
8534:
8530:
8523:
8522:
8518:
8516:
8515:
8511:
8509:
8508:
8504:
8502:
8501:
8497:
8495:
8494:
8490:
8488:
8487:
8486:Trigonosaurus
8483:
8481:
8480:
8476:
8474:
8473:
8469:
8466:
8465:
8461:
8458:
8457:
8453:
8451:
8450:
8449:Ruyangosaurus
8446:
8444:
8443:
8439:
8437:
8436:
8432:
8430:
8429:
8428:Pitekunsaurus
8425:
8423:
8422:
8418:
8415:
8414:
8410:
8408:
8407:
8403:
8401:
8400:
8396:
8394:
8393:
8389:
8387:
8386:
8382:
8380:
8379:
8378:Mnyamawamtuka
8375:
8373:
8372:
8368:
8365:
8364:
8360:
8358:
8357:
8356:Magyarosaurus
8353:
8350:
8349:
8348:Ligabuesaurus
8345:
8343:
8342:
8341:Laplatasaurus
8338:
8336:
8335:
8334:Karongasaurus
8331:
8329:
8328:
8324:
8321:
8320:
8316:
8314:
8313:
8309:
8307:
8306:
8302:
8300:
8299:
8295:
8292:
8291:
8287:
8285:
8284:
8280:
8277:
8276:
8275:Dreadnoughtus
8272:
8270:
8269:
8265:
8263:
8262:
8258:
8256:
8255:
8251:
8249:
8248:
8244:
8242:
8241:
8240:Borealosaurus
8237:
8234:
8233:
8229:
8226:
8225:
8221:
8219:
8218:
8217:Barrosasaurus
8214:
8212:
8211:
8207:
8205:
8204:
8200:
8198:
8197:
8193:
8191:
8190:
8186:
8184:
8183:
8179:
8176:
8175:
8171:
8169:
8168:
8164:
8161:
8160:
8159:Aegyptosaurus
8156:
8154:
8153:
8149:
8148:
8145:
8141:
8138:
8135:
8130:
8126:
8122:
8117:
8113:
8103:
8098:
8095:
8087:
8057:
8056:
8055:
8054:
8052:
8050:
8046:
8035:
8034:
8030:
8028:
8027:
8026:Savannasaurus
8023:
8021:
8020:
8016:
8014:
8013:
8009:
8007:
8006:
8005:Australotitan
8002:
8001:
7998:
7995:
7993:
7989:
7983:
7982:
7978:
7976:
7975:
7971:
7969:
7968:
7964:
7962:
7961:
7957:
7955:
7954:
7950:
7948:
7947:
7943:
7941:
7940:
7936:
7933:
7932:
7928:
7926:
7925:
7921:
7919:
7918:
7914:
7912:
7911:
7907:
7905:
7904:
7900:
7898:
7897:
7893:
7891:
7890:
7886:
7883:
7882:
7878:
7875:
7874:
7873:Australodocus
7870:
7869:
7867:
7865:
7864:Euhelopodidae
7861:
7855:
7854:
7853:Triunfosaurus
7850:
7848:
7847:
7843:
7841:
7840:
7836:
7834:
7833:
7832:Sauroposeidon
7829:
7827:
7826:
7822:
7820:
7819:
7815:
7813:
7812:
7811:Padillasaurus
7808:
7806:
7805:
7801:
7799:
7798:
7794:
7791:
7790:
7789:Ligabuesaurus
7786:
7784:
7783:
7779:
7777:
7776:
7772:
7769:
7768:
7764:
7762:
7761:
7757:
7755:
7754:
7750:
7748:
7747:
7743:
7741:
7740:
7736:
7734:
7733:
7732:Chubutisaurus
7729:
7727:
7726:
7722:
7720:
7719:
7715:
7713:
7712:
7708:
7706:
7705:
7701:
7699:
7698:
7694:
7692:
7691:
7687:
7686:
7683:
7680:
7678:
7674:
7668:
7667:
7663:
7661:
7660:
7656:
7654:
7653:
7649:
7647:
7646:
7642:
7640:
7639:
7635:
7633:
7632:
7628:
7625:
7624:
7620:
7618:
7617:
7613:
7611:
7610:
7609:Brachiosaurus
7606:
7604:
7603:
7599:
7597:
7596:
7592:
7591:
7589:
7587:
7583:
7576:
7575:
7571:
7569:
7568:
7564:
7562:
7561:
7557:
7555:
7554:
7550:
7548:
7547:
7546:Fushanosaurus
7543:
7541:
7540:
7536:
7534:
7533:
7529:
7527:
7526:
7522:
7521:
7518:
7515:
7512:
7507:
7497:
7496:
7492:
7490:
7489:
7485:
7484:
7482:
7480:
7476:
7470:
7469:
7465:
7462:
7461:
7457:
7455:
7454:
7450:
7447:
7446:
7442:
7440:
7439:
7435:
7432:
7431:
7430:Bashunosaurus
7427:
7425:
7424:
7420:
7418:
7417:
7413:
7412:
7409:
7405:
7402:
7399:
7394:
7390:
7386:
7381:
7377:
7368:
7363:
7333:
7332:
7328:
7326:
7325:
7321:
7319:
7318:
7314:
7311:
7310:
7306:
7304:
7303:
7299:
7297:
7296:
7292:
7290:
7289:
7285:
7283:
7282:
7278:
7277:
7275:
7273:
7269:
7263:
7262:
7258:
7255:
7254:
7253:Atlantosaurus
7250:
7248:
7247:
7243:
7240:
7239:
7235:
7234:
7232:
7230:
7229:Apatosaurinae
7226:
7223:
7221:
7217:
7211:
7210:
7206:
7204:
7203:
7199:
7197:
7196:
7195:Smitanosaurus
7192:
7190:
7189:
7185:
7183:
7182:
7178:
7176:
7175:
7171:
7168:
7167:
7166:Dyslocosaurus
7163:
7161:
7160:
7159:Dicraeosaurus
7156:
7154:
7153:
7149:
7147:
7146:
7142:
7140:
7139:
7138:Amargatitanis
7135:
7133:
7132:
7128:
7127:
7125:
7123:
7119:
7116:
7114:
7110:
7096:
7095:
7091:
7089:
7088:
7084:
7082:
7081:
7077:
7075:
7074:
7073:Katepensaurus
7070:
7068:
7067:
7066:Itapeuasaurus
7063:
7061:
7060:
7059:Demandasaurus
7056:
7055:
7053:
7051:
7047:
7040:
7039:
7035:
7033:
7032:
7028:
7026:
7025:
7024:Cathartesaura
7021:
7020:
7018:
7016:
7015:Limaysaurinae
7012:
7009:
7007:
7003:
6997:
6996:
6992:
6990:
6989:
6985:
6983:
6982:
6978:
6976:
6975:
6971:
6968:
6967:
6963:
6961:
6960:
6956:
6954:
6953:
6952:Histriasaurus
6949:
6946:
6945:
6944:Dzharatitanis
6941:
6939:
6938:
6937:Comahuesaurus
6934:
6932:
6931:
6927:
6925:
6924:
6920:
6918:
6917:
6913:
6912:
6909:
6906:
6904:
6900:
6897:
6894:
6889:
6883:
6882:
6878:
6877:
6874:
6871:
6868:
6867:Diplodocoidea
6863:
6859:
6855:
6854:Diplodocoidea
6850:
6846:
6837:
6833:
6831:
6825:
6817:
6794:
6793:
6789:
6787:
6786:
6782:
6780:
6779:
6775:
6773:
6772:
6771:Gigantosaurus
6768:
6766:
6765:
6761:
6759:
6758:
6754:
6752:
6751:
6747:
6746:
6744:
6740:
6736:
6717:
6716:
6714:
6712:
6708:
6701:
6700:
6698:
6696:
6695:Diplodocoidea
6692:
6689:
6687:
6683:
6677:
6676:
6672:
6670:
6669:
6665:
6663:
6662:
6658:
6655:
6654:
6650:
6647:
6646:
6642:
6640:
6639:
6638:Narindasaurus
6635:
6633:
6632:
6628:
6626:
6625:
6621:
6619:
6618:
6617:Losillasaurus
6614:
6611:
6610:
6606:
6603:
6602:
6598:
6595:
6594:
6590:
6588:
6587:
6583:
6582:
6580:
6578:
6574:
6568:
6567:
6566:Zigongosaurus
6563:
6561:
6560:
6559:Yuanmousaurus
6556:
6554:
6553:
6552:Xinjiangtitan
6549:
6547:
6546:
6545:Wamweracaudia
6542:
6540:
6539:
6538:Tonganosaurus
6535:
6533:
6532:
6528:
6526:
6525:
6521:
6519:
6518:
6514:
6512:
6511:
6507:
6505:
6504:
6500:
6497:
6496:
6495:Klamelisaurus
6492:
6490:
6489:
6485:
6483:
6482:
6478:
6476:
6475:
6474:Huangshanlong
6471:
6469:
6468:
6464:
6461:
6460:
6456:
6454:
6453:
6449:
6446:
6445:
6441:
6439:
6438:
6434:
6432:
6431:
6427:
6426:
6424:
6422:
6418:
6411:
6410:
6406:
6403:
6402:
6398:
6395:
6394:
6393:Ferganasaurus
6390:
6388:
6387:
6383:
6381:
6380:
6376:
6375:
6373:
6371:
6370:Cetiosauridae
6367:
6361:
6360:
6356:
6354:
6353:
6349:
6347:
6346:
6342:
6340:
6339:
6335:
6333:
6332:
6328:
6326:
6325:
6321:
6319:
6318:
6314:
6312:
6311:
6307:
6305:
6304:
6300:
6298:
6297:
6293:
6291:
6290:
6286:
6284:
6283:
6279:
6277:
6276:
6272:
6270:
6269:
6265:
6263:
6262:
6258:
6256:
6255:
6251:
6250:
6247:
6244:
6241:
6236:
6226:
6225:
6221:
6219:
6218:
6217:Tazoudasaurus
6214:
6213:
6211:
6209:
6205:
6198:
6197:
6193:
6190:
6189:
6185:
6183:
6182:
6178:
6176:
6175:
6174:Ohmdenosaurus
6171:
6169:
6168:
6164:
6161:
6160:
6156:
6155:
6152:
6148:
6145:
6142:
6137:
6127:
6126:
6122:
6120:
6119:
6115:
6113:
6112:
6108:
6106:
6105:
6101:
6100:
6098:
6096:
6092:
6085:
6084:
6080:
6078:
6077:
6076:Schleitheimia
6073:
6071:
6070:
6066:
6063:
6062:
6058:
6055:
6054:
6053:Leonerasaurus
6050:
6048:
6047:
6043:
6041:
6040:
6036:
6034:
6033:
6029:
6027:
6026:
6025:Blikanasaurus
6022:
6020:
6019:
6015:
6014:
6011:
6007:
6004:
6001:
5996:
5992:
5988:
5983:
5979:
5969:
5964:
5962:
5957:
5951:
5910:
5909:
5908:
5907:
5905:
5903:
5899:
5893:
5892:
5888:
5886:
5885:
5881:
5879:
5878:
5874:
5872:
5871:
5867:
5865:
5864:
5860:
5858:
5857:
5856:Leonerasaurus
5853:
5851:
5850:
5849:Lamplughsaura
5846:
5844:
5843:
5839:
5837:
5836:
5832:
5830:
5829:
5825:
5823:
5822:
5818:
5817:
5814:
5811:
5809:
5805:
5799:
5798:
5797:Yunnanosaurus
5794:
5792:
5791:
5787:
5784:
5783:
5779:
5777:
5776:
5772:
5770:
5769:
5765:
5763:
5762:
5758:
5755:
5754:
5750:
5749:
5746:
5743:
5741:
5737:
5730:
5729:
5725:
5723:
5722:
5718:
5716:
5715:
5711:
5709:
5708:
5704:
5702:
5701:
5697:
5695:
5694:
5693:Lufengosaurus
5690:
5688:
5687:
5683:
5680:
5679:
5675:
5673:
5672:
5668:
5666:
5665:
5661:
5659:
5658:
5654:
5653:
5651:
5649:
5645:
5639:
5638:
5634:
5632:
5631:
5630:Eucnemesaurus
5627:
5626:
5624:
5622:
5621:Riojasauridae
5618:
5612:
5611:
5607:
5605:
5604:
5600:
5598:
5597:
5593:
5591:
5590:
5586:
5584:
5583:
5579:
5577:
5576:
5572:
5570:
5569:
5565:
5564:
5561:
5558:
5555:
5550:
5540:
5539:
5535:
5533:
5532:
5528:
5526:
5525:
5521:
5519:
5518:
5514:
5512:
5511:
5507:
5505:
5504:
5503:Euskelosaurus
5500:
5499:
5497:
5495:
5491:
5485:
5484:
5480:
5478:
5477:
5473:
5471:
5470:
5466:
5465:
5462:
5459:
5456:
5451:
5441:
5440:
5436:
5434:
5433:
5429:
5427:
5426:
5422:
5421:
5419:
5417:
5413:
5407:
5406:
5402:
5400:
5399:
5395:
5393:
5392:
5388:
5386:
5385:
5381:
5379:
5378:
5377:Bagualosaurus
5374:
5372:
5371:
5367:
5365:
5364:
5360:
5359:
5356:
5352:
5349:
5346:
5345:Bagualosauria
5341:
5331:
5330:
5326:
5323:
5322:
5318:
5315:
5314:
5313:Pampadromaeus
5310:
5308:
5307:
5303:
5301:
5300:
5296:
5293:
5292:
5288:
5285:
5284:
5280:
5279:
5277:
5275:
5274:Saturnaliidae
5271:
5265:
5264:
5260:
5258:
5257:
5256:Pampadromaeus
5253:
5251:
5250:
5246:
5244:
5243:
5239:
5237:
5236:
5235:Gigantoscelus
5232:
5230:
5229:
5225:
5223:
5222:
5218:
5217:
5214:
5210:
5207:
5204:
5199:
5195:
5191:
5186:
5182:
5161:
5160:
5159:
5158:
5156:
5154:
5150:
5142:
5138:
5137:
5136:
5135:
5132:
5129:
5127:
5123:
5119:
5111:
5107:
5104:
5102:
5098:
5095:
5093:
5089:
5087:
5083:
5081:
5077:
5076:
5071:
5067:
5063:
5056:
5051:
5049:
5044:
5042:
5037:
5036:
5033:
5027:
5024:
5022:
5019:
5018:
5009:
5005:
5002:
5001:
4981:
4977:
4971:
4956:
4952:
4946:
4938:
4934:
4930:
4926:
4924:
4915:
4906:
4901:
4897:
4893:
4889:
4885:
4881:
4877:
4870:
4862:
4858:
4854:
4850:
4846:
4842:
4838:
4831:
4823:
4819:
4815:
4811:
4806:
4801:
4797:
4793:
4789:
4785:
4781:
4777:
4770:
4763:(2): 127–132.
4762:
4758:
4754:
4747:
4739:
4735:
4730:
4725:
4721:
4717:
4713:
4709:
4705:
4698:
4690:
4686:
4681:
4676:
4672:
4668:
4664:
4660:
4656:
4652:
4648:
4641:
4626:
4625:NY Daily News
4622:
4615:
4600:
4596:
4589:
4581:
4577:
4573:
4569:
4565:
4561:
4557:
4553:
4549:
4542:
4534:
4530:
4526:
4522:
4518:
4514:
4507:
4499:
4495:
4491:
4487:
4483:
4479:
4475:
4471:
4464:
4456:
4452:
4448:
4444:
4440:
4436:
4432:
4428:
4420:
4412:
4408:
4404:
4400:
4396:
4392:
4388:
4381:
4373:
4369:
4364:
4359:
4355:
4351:
4347:
4343:
4340:(8): 191057.
4339:
4335:
4331:
4324:
4316:
4312:
4308:
4304:
4299:
4294:
4290:
4286:
4282:
4278:
4270:
4261:
4260:10044/1/53967
4256:
4252:
4248:
4244:
4240:
4236:
4229:
4221:
4215:
4211:
4207:
4203:
4195:
4193:
4183:
4178:
4173:
4168:
4164:
4160:
4156:
4154:
4145:
4137:
4133:
4128:
4123:
4119:
4115:
4111:
4104:
4096:
4092:
4088:
4084:
4080:
4076:
4072:
4068:
4060:
4052:
4048:
4043:
4038:
4034:
4030:
4026:
4022:
4018:
4014:
4010:
4006:
4002:
3995:
3993:
3984:
3980:
3975:
3970:
3965:
3960:
3956:
3952:
3948:
3944:
3940:
3933:
3925:
3921:
3917:
3913:
3908:
3903:
3899:
3895:
3892:(2–4): 1–13.
3891:
3887:
3880:
3873:
3858:
3854:
3852:
3843:
3828:
3824:
3822:
3818:
3809:
3794:
3790:
3783:
3775:
3771:
3767:
3763:
3759:
3755:
3750:
3745:
3741:
3737:
3733:
3726:
3718:
3714:
3710:
3706:
3702:
3698:
3691:
3684:
3682:
3680:
3663:
3659:
3653:
3651:
3635:
3631:
3624:
3622:
3620:
3604:
3600:
3594:
3579:
3578:
3573:
3566:
3551:
3547:
3540:
3524:
3520:
3516:
3509:
3494:
3490:
3483:
3468:
3464:
3457:
3449:
3445:
3441:
3437:
3433:
3429:
3425:
3421:
3417:
3415:
3406:
3404:
3402:
3400:
3398:
3396:
3387:
3383:
3378:
3373:
3369:
3365:
3361:
3357:
3353:
3346:
3344:
3342:
3340:
3338:
3336:
3334:
3332:
3330:
3328:
3326:
3321:
3311:
3308:
3307:
3301:
3299:
3295:
3291:
3290:
3285:
3275:
3273:
3272:
3267:
3263:
3262:
3261:Barcinosuchus
3258:
3257:crocodyliform
3255:
3251:
3250:
3245:
3244:
3239:
3235:
3231:
3227:
3223:
3219:
3214:
3212:
3208:
3207:
3202:
3201:
3196:
3195:
3190:
3189:
3184:
3180:
3176:
3172:
3168:
3164:
3160:
3153:
3148:
3134:
3133:
3125:
3124:
3116:
3115:
3107:
3106:
3098:
3097:
3089:
3088:
3080:
3079:
3071:
3070:
3067:
3066:
3065:
3058:
3057:
3054:
3053:
3050:
3049:
3048:
3047:Mendozasaurus
3041:
3040:
3034:
3033:
3030:
3029:
3026:
3025:
3024:
3017:
3016:
3010:
3007:
3006:
3003:
3002:
2994:
2993:
2985:
2984:
2976:
2975:
2972:
2971:
2970:
2969:Gondwanatitan
2963:
2962:
2959:
2958:
2955:
2954:
2953:
2946:
2945:
2939:
2936:
2935:
2932:
2931:
2928:
2927:
2926:
2925:Muyelensaurus
2919:
2918:
2912:
2911:
2908:
2907:
2904:
2903:
2902:
2895:
2894:
2888:
2885:
2884:
2878:
2877:
2874:
2873:
2870:
2869:
2868:
2861:
2860:
2854:
2853:Colossosauria
2851:
2850:
2847:
2846:
2838:
2837:
2829:
2828:
2825:
2824:
2823:
2816:
2815:Saltasaurinae
2813:
2812:
2809:
2808:
2805:
2804:
2803:
2796:
2793:
2792:
2786:
2785:Saltasauridae
2783:
2782:
2779:
2778:
2775:
2774:
2773:
2772:Nemegtosaurus
2766:
2765:
2759:
2756:
2755:
2749:
2746:
2745:
2742:
2741:
2738:
2737:
2736:
2729:
2726:
2725:
2719:
2718:
2715:
2714:
2711:
2710:
2709:
2702:
2701:
2695:
2692:
2691:
2687:
2685:
2681:
2680:Mendozasaurus
2677:
2673:
2668:
2663:
2661:
2657:
2653:
2649:
2645:
2644:
2639:
2635:
2631:
2628:grouped with
2627:
2623:
2622:Mendozasaurus
2619:
2615:
2611:
2607:
2603:
2599:
2598:Colossosauria
2594:
2590:
2586:
2582:
2578:
2574:
2570:
2566:
2556:
2555:
2547:
2546:
2538:
2537:
2529:
2528:
2520:
2519:
2511:
2510:
2502:
2501:
2493:
2492:
2484:
2483:
2475:
2474:
2466:
2465:
2457:
2456:
2453:
2452:
2451:
2444:
2443:
2440:
2439:
2436:
2435:
2434:
2427:
2426:
2420:
2419:
2416:
2415:
2412:
2411:
2410:
2403:
2402:
2399:
2398:
2395:
2394:
2393:
2386:
2385:
2379:
2378:
2375:
2374:
2366:
2365:
2362:
2361:
2360:
2353:
2352:
2349:
2348:
2345:
2344:
2343:
2336:
2335:
2329:
2328:
2322:
2321:
2318:
2317:
2314:
2313:
2312:
2311:Mendozasaurus
2305:
2304:
2298:
2295:
2294:
2291:
2290:
2287:
2286:
2285:
2278:
2277:
2271:
2270:
2267:
2266:
2263:
2262:
2261:
2254:
2253:
2247:
2246:
2243:
2242:
2234:
2233:
2225:
2224:
2216:
2215:
2212:
2211:
2210:
2203:
2202:
2199:
2198:
2195:
2194:
2193:
2186:
2185:
2179:
2178:
2175:
2174:
2171:
2170:
2169:
2168:Muyelensaurus
2162:
2161:
2155:
2154:
2151:
2150:
2147:
2146:
2145:
2138:
2137:
2131:
2128:
2127:
2121:
2120:
2117:
2116:
2108:
2107:
2099:
2098:
2090:
2089:
2081:
2080:
2072:
2071:
2063:
2062:
2059:
2058:
2057:
2050:
2049:
2046:
2045:
2042:
2041:
2040:
2033:
2032:
2026:
2025:
2022:
2021:
2018:
2017:
2016:
2009:
2008:
2002:
2001:
1998:
1997:
1994:
1993:
1992:
1991:Neuquensaurus
1985:
1984:
1981:
1980:
1977:
1976:
1975:
1968:
1967:
1964:
1963:
1960:
1959:
1958:
1951:
1950:
1947:
1946:
1943:
1942:
1941:
1934:
1933:
1930:
1929:
1926:
1925:
1924:
1923:Trigonosaurus
1917:
1916:
1913:
1912:
1909:
1908:
1907:
1906:Nemegtosaurus
1900:
1899:
1893:
1892:
1889:
1888:
1885:
1884:
1883:
1876:
1875:
1869:
1868:
1865:
1864:
1861:
1860:
1859:
1852:
1851:
1845:
1842:
1841:
1838:
1837:
1834:
1833:
1832:
1831:Dreadnoughtus
1825:
1824:
1818:
1817:
1811:
1808:
1807:
1804:
1803:
1800:
1799:
1798:
1791:
1790:
1784:
1783:
1780:
1779:
1771:
1770:
1767:
1766:
1765:
1758:
1757:
1754:
1753:
1750:
1749:
1748:
1747:Ruyangosaurus
1741:
1740:
1734:
1733:
1727:
1726:
1723:
1722:
1719:
1718:
1717:
1710:
1709:
1706:
1705:
1702:
1701:
1700:
1693:
1692:
1686:
1683:
1682:
1678:
1676:
1672:
1668:
1664:
1660:
1656:
1652:
1651:Dreadnoughtus
1647:
1643:
1639:
1635:
1631:
1627:
1617:
1615:
1611:
1606:
1602:
1598:
1594:
1593:Neuquensaurus
1590:
1586:
1582:
1578:
1574:
1573:Neuquensaurus
1570:
1566:
1562:
1558:
1557:
1552:
1551:
1546:
1542:
1538:
1537:
1532:
1531:
1526:
1525:
1520:
1516:
1512:
1508:
1504:
1503:
1498:
1494:
1490:
1486:
1482:
1481:
1476:
1475:
1474:Muyelensaurus
1470:
1466:
1465:Dreadnoughtus
1457:
1453:
1451:
1450:
1445:
1441:
1437:
1433:
1429:
1425:
1424:Neuquensaurus
1421:
1417:
1413:
1409:
1408:Neuquensaurus
1405:
1401:
1400:Dreadnoughtus
1397:
1392:
1385:
1380:
1373:
1369:
1365:
1361:
1360:Dreadnoughtus
1357:
1356:
1351:
1350:
1345:
1340:
1338:
1337:
1332:
1331:Dreadnoughtus
1328:
1327:Saltasauridae
1324:
1323:
1322:Savannasaurus
1318:
1317:Dreadnoughtus
1314:
1313:
1308:
1307:
1306:Ruyangosaurus
1302:
1301:
1296:
1292:
1291:
1290:Sauroposeidon
1286:
1285:
1280:
1276:
1275:Dreadnoughtus
1272:
1271:
1265:
1257:
1250:
1246:
1242:
1241:
1236:
1235:
1234:Neuquensaurus
1230:
1229:
1224:
1220:
1212:
1203:
1201:
1200:
1195:
1194:Mendozasaurus
1191:
1187:
1183:
1179:
1175:
1171:
1170:Dreadnoughtus
1167:
1166:Mendozasaurus
1163:
1159:
1158:
1152:
1145:
1144:
1139:
1138:
1133:
1132:Dreadnoughtus
1129:
1124:
1112:
1108:
1106:
1100:
1092:
1084:
1080:
1076:
1075:
1074:Dreadnoughtus
1070:
1066:
1062:
1058:
1052:
1044:
1036:
1029:
1028:
1023:
1019:
1014:
1007:
1003:
999:
994:
992:
991:
990:Mendozasaurus
986:
985:
984:Ligabuesaurus
980:
979:
974:
969:
961:
959:neural spines
954:
945:
936:
934:
930:
926:
922:
918:
914:
907:
903:
899:
895:
891:
887:
882:
880:
876:
872:
868:
864:
863:
858:
850:
844:
840:
838:
834:
830:
822:
818:
804:
802:
798:
797:
796:Tyrannosaurus
792:
788:
784:
783:
778:
773:
765:
760:
751:
749:
748:specific name
745:
742:with a Greek
741:
737:
733:
729:
725:
721:
717:
713:
709:
705:
701:
696:
693:
685:
677:
669:
661:
654:
649:
646:
638:
631:
626:
618:
610:
602:
594:
587:
578:
574:
569:
565:
563:
559:
555:
551:
548:bones and 57
547:
543:
539:
535:
531:
527:
522:
509:
507:
503:
499:
493:
485:
477:
469:
461:
453:
445:
438:
434:
430:
425:
423:
422:Colossosauria
419:
415:
411:
410:
405:
404:
399:
395:
391:
387:
383:
379:
375:
371:
370:
365:
361:
357:
353:
350:
347:
346:titanosaurian
343:
342:
331:
325:
324:
316:
313:
309:
304:
298:
297:
290:
287:
286:
283:
277:
274:
271:
270:
267:
261:
258:
255:
254:
251:
245:
242:
239:
238:
235:
229:
226:
223:
222:
219:
213:
210:
207:
206:
203:
200:
197:
194:
193:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
145:
140:
136:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
32:
26:
22:
19:
9428:
9315:
9289:Titanosaurus
9287:
9279:
9272:
9264:
9259:Loricosaurus
9257:
9252:Iuticosaurus
9250:
9243:
9236:
9229:
9221:
9189:Yamanasaurus
9187:
9180:
9173:
9166:
9159:
9151:
9144:
9136:
9130:Abditosaurus
9128:
9107:
9100:
9092:
9085:
9077:
9069:
9062:
9056:Abditosaurus
9054:
9033:
9026:
9018:
9011:
8988:Tapuiasaurus
8986:
8978:
8971:
8951:
8946:Titanomachya
8944:
8937:
8915:Puertasaurus
8913:
8907:
8906:
8901:Notocolossus
8899:
8892:
8886:Jiangxititan
8884:
8877:
8869:
8864:Drusilasaura
8862:
8855:
8848:Lognkosauria
8831:
8824:
8817:
8810:
8803:
8796:
8789:
8782:Aeolosaurini
8771:Rinconsaurus
8769:
8762:
8755:
8748:
8740:
8733:
8722:Rinconsauria
8709:
8703:Tengrisaurus
8701:
8696:Quetecsaurus
8694:
8687:
8679:
8672:
8665:
8658:
8651:
8627:
8620:
8586:
8579:
8574:Lohuecotitan
8572:
8565:
8560:Garrigatitan
8558:
8551:
8546:Ampelosaurus
8544:
8519:
8512:
8505:
8500:Uberabatitan
8498:
8493:Sonidosaurus
8491:
8484:
8477:
8470:
8464:Tengrisaurus
8462:
8454:
8447:
8440:
8435:Rapetosaurus
8433:
8426:
8419:
8411:
8404:
8397:
8390:
8383:
8376:
8371:Malawisaurus
8369:
8361:
8354:
8346:
8339:
8332:
8325:
8317:
8312:Huabeisaurus
8310:
8303:
8296:
8288:
8281:
8273:
8266:
8259:
8254:Choconsaurus
8252:
8247:Brasilotitan
8245:
8238:
8230:
8222:
8215:
8208:
8201:
8196:Atacamatitan
8194:
8189:Argyrosaurus
8187:
8180:
8172:
8165:
8157:
8152:Abdarainurus
8150:
8134:Titanosauria
8121:Titanosauria
8090:
8049:Titanosauria
8033:Wintonotitan
8031:
8024:
8017:
8010:
8003:
7979:
7974:Yongjinglong
7972:
7965:
7960:Tambatitanis
7958:
7951:
7944:
7937:
7929:
7924:Huanghetitan
7922:
7917:Huabeisaurus
7915:
7908:
7903:Gannansaurus
7901:
7894:
7887:
7881:Chiayusaurus
7879:
7871:
7851:
7844:
7837:
7830:
7823:
7816:
7809:
7802:
7795:
7787:
7780:
7775:Jiutaisaurus
7773:
7767:Jiangxititan
7765:
7760:Garumbatitan
7758:
7751:
7744:
7739:Dongbeititan
7737:
7730:
7723:
7716:
7711:Austrosaurus
7709:
7702:
7695:
7688:
7664:
7659:Venenosaurus
7657:
7650:
7645:Sonorasaurus
7643:
7636:
7631:Giraffatitan
7629:
7621:
7616:Cedarosaurus
7614:
7607:
7600:
7593:
7572:
7567:Pelorosaurus
7565:
7558:
7551:
7544:
7537:
7530:
7523:
7493:
7488:Camarasaurus
7486:
7466:
7460:Haestasaurus
7458:
7453:Europasaurus
7451:
7443:
7436:
7428:
7421:
7414:
7329:
7322:
7315:
7307:
7300:
7293:
7286:
7279:
7272:Diplodocinae
7261:Brontosaurus
7259:
7251:
7244:
7238:Amphicoelias
7236:
7220:Diplodocidae
7207:
7200:
7193:
7186:
7179:
7172:
7164:
7157:
7150:
7145:Bajadasaurus
7143:
7136:
7131:Amargasaurus
7129:
7092:
7085:
7078:
7071:
7064:
7057:
7038:Rayososaurus
7036:
7029:
7022:
6995:Zapalasaurus
6993:
6988:Xenoposeidon
6986:
6979:
6972:
6964:
6957:
6950:
6942:
6935:
6930:Campananeyen
6928:
6923:Amazonsaurus
6921:
6914:
6879:
6820:
6790:
6783:
6776:
6769:
6762:
6755:
6748:
6686:Neosauropoda
6673:
6666:
6659:
6651:
6643:
6636:
6629:
6622:
6615:
6607:
6601:Haestasaurus
6599:
6591:
6584:
6564:
6557:
6550:
6543:
6536:
6529:
6522:
6515:
6508:
6501:
6493:
6486:
6479:
6472:
6465:
6457:
6450:
6442:
6435:
6428:
6409:Patagosaurus
6407:
6399:
6391:
6384:
6377:
6359:Volkheimeria
6357:
6350:
6343:
6336:
6331:Perijasaurus
6329:
6324:Nebulasaurus
6322:
6315:
6308:
6303:Dystrophaeus
6301:
6294:
6287:
6280:
6275:Barapasaurus
6273:
6266:
6261:Asiatosaurus
6259:
6252:
6222:
6215:
6194:
6186:
6181:Rhoetosaurus
6179:
6172:
6165:
6157:
6125:Lessemsaurus
6123:
6116:
6109:
6104:Antetonitrus
6102:
6081:
6074:
6067:
6059:
6051:
6044:
6037:
6030:
6023:
6016:
5953:
5889:
5882:
5875:
5868:
5861:
5854:
5847:
5840:
5833:
5826:
5819:
5795:
5790:Yizhousaurus
5788:
5780:
5773:
5766:
5759:
5751:
5726:
5719:
5712:
5705:
5698:
5691:
5684:
5678:Ignavusaurus
5676:
5669:
5662:
5655:
5635:
5628:
5608:
5601:
5594:
5587:
5582:Ignavusaurus
5580:
5573:
5566:
5538:Yimenosaurus
5536:
5531:Plateosaurus
5529:
5522:
5515:
5508:
5501:
5483:Xixiposaurus
5481:
5474:
5467:
5455:Plateosauria
5437:
5430:
5423:
5416:Unaysauridae
5403:
5396:
5389:
5382:
5375:
5368:
5361:
5327:
5319:
5311:
5304:
5297:
5289:
5281:
5261:
5254:
5247:
5242:Guaibasaurus
5240:
5233:
5226:
5219:
5105:
5096:
4983:. Retrieved
4979:
4970:
4958:. Retrieved
4954:
4945:
4928:
4922:
4914:
4879:
4875:
4869:
4844:
4840:
4836:
4830:
4805:11336/103474
4779:
4775:
4769:
4760:
4756:
4746:
4729:11336/117578
4711:
4708:IAWA Journal
4707:
4703:
4697:
4680:11336/127834
4654:
4650:
4640:
4628:. Retrieved
4624:
4614:
4602:. Retrieved
4598:
4588:
4555:
4551:
4547:
4541:
4519:(1): 35–51.
4516:
4512:
4506:
4473:
4469:
4463:
4430:
4426:
4419:
4394:
4390:
4386:
4380:
4337:
4333:
4323:
4298:11336/106658
4280:
4276:
4269:
4242:
4238:
4234:
4228:
4201:
4182:11336/192915
4165:(1): 45–60.
4162:
4158:
4152:
4144:
4117:
4113:
4103:
4070:
4066:
4059:
4008:
4004:
3946:
3942:
3932:
3889:
3885:
3872:
3860:. Retrieved
3856:
3850:
3842:
3830:. Retrieved
3826:
3820:
3816:
3808:
3796:. Retrieved
3792:
3782:
3739:
3735:
3725:
3700:
3696:
3666:. Retrieved
3661:
3637:. Retrieved
3633:
3606:. Retrieved
3602:
3593:
3581:. Retrieved
3577:The Guardian
3575:
3565:
3553:. Retrieved
3549:
3539:
3527:. Retrieved
3523:the original
3518:
3508:
3496:. Retrieved
3492:
3482:
3470:. Retrieved
3466:
3456:
3423:
3419:
3413:
3359:
3355:
3297:
3293:
3287:
3281:
3269:
3259:
3247:
3243:Tyrannotitan
3241:
3237:
3229:
3215:
3210:
3204:
3198:
3192:
3186:
3162:
3158:
3157:
3151:
3142:Paleoecology
3063:
3062:
3061:
3045:
3044:
3021:
3020:
3009:Lognkosauria
2967:
2966:
2950:
2949:
2938:Aeolosaurini
2923:
2922:
2901:Rinconsaurus
2899:
2898:
2887:Rinconsauria
2865:
2864:
2820:
2819:
2800:
2799:
2770:
2769:
2733:
2732:
2706:
2705:
2694:Titanosauria
2683:
2679:
2675:
2672:Malawisaurus
2671:
2666:
2664:
2659:
2656:Puertasaurus
2655:
2651:
2647:
2641:
2638:Drusilasaura
2637:
2634:Puertasaurus
2633:
2629:
2625:
2621:
2617:
2613:
2610:Notocolossus
2609:
2606:Puertasaurus
2605:
2601:
2592:
2589:Notocolossus
2588:
2585:Puertasaurus
2584:
2576:
2573:Puertasaurus
2572:
2568:
2564:
2562:
2449:
2448:
2447:
2431:
2430:
2409:Drusilasaura
2407:
2406:
2392:Puertasaurus
2390:
2389:
2359:Quetecsaurus
2357:
2356:
2340:
2339:
2309:
2308:
2297:Lognkosauria
2284:Notocolossus
2282:
2281:
2258:
2257:
2207:
2206:
2190:
2189:
2166:
2165:
2144:Rinconsaurus
2142:
2141:
2130:Rinconsauria
2056:Tapuiasaurus
2054:
2053:
2037:
2036:
2015:Rapetosaurus
2013:
2012:
1989:
1988:
1972:
1971:
1955:
1954:
1938:
1937:
1921:
1920:
1904:
1903:
1880:
1879:
1858:Malawisaurus
1856:
1855:
1844:Lithostrotia
1829:
1828:
1795:
1794:
1762:
1761:
1745:
1744:
1714:
1713:
1699:Wintonotitan
1697:
1696:
1685:Titanosauria
1667:Rinconsauria
1662:
1659:Notocolossus
1658:
1654:
1650:
1646:Notocolossus
1645:
1641:
1637:
1634:sister group
1629:
1623:
1600:
1597:Rapetosaurus
1596:
1592:
1588:
1584:
1580:
1577:Rapetosaurus
1576:
1572:
1564:
1560:
1554:
1548:
1544:
1540:
1536:Venenosaurus
1534:
1528:
1522:
1518:
1514:
1510:
1502:Uberabatitan
1500:
1496:
1492:
1489:Malawisaurus
1488:
1484:
1478:
1472:
1468:
1464:
1462:
1449:Haestasaurus
1447:
1444:Giraffatitan
1443:
1439:
1431:
1427:
1423:
1419:
1415:
1411:
1407:
1403:
1399:
1371:
1367:
1363:
1359:
1355:Rapetosaurus
1353:
1349:Notocolossus
1347:
1343:
1341:
1334:
1330:
1320:
1316:
1312:Tapuiasaurus
1310:
1304:
1300:Giraffatitan
1298:
1294:
1288:
1282:
1279:Drusilasaura
1278:
1274:
1268:
1248:
1238:
1232:
1226:
1223:Drusilasaura
1222:
1218:
1217:
1199:Drusilasaura
1197:
1193:
1189:
1177:
1174:Diplodocidae
1169:
1165:
1161:
1155:
1143:Malawisaurus
1141:
1135:
1131:
1127:
1117:
1104:
1083:Lognkosauria
1079:Puertasaurus
1078:
1072:
1068:
1060:
1056:
1027:Puertasaurus
1025:
1021:
1017:
1005:
1001:
997:
995:
988:
982:
976:
972:
952:
950:
932:
928:
924:
920:
916:
912:
905:
901:
897:
893:
885:
883:
874:
870:
860:
856:
848:
846:
842:
828:
826:
820:
794:
793:specimen of
780:
771:
769:
736:Titanosauria
731:
727:
724:type species
697:
650:
604:, six front
582:
576:
515:
497:
436:
433:Lognkosauria
428:
426:
413:
409:Puertasaurus
407:
401:
397:
381:
368:
367:
340:
339:
338:
329:
322:
321:
312:Type species
302:
295:
294:
282:Lognkosauria
272:
266:Titanosauria
256:
240:
224:
208:
195:
182:
36:101.62
24:
18:
9504:Albian life
9429:Patagotitan
9274:Microcoelus
9182:Saltasaurus
9168:Paralititan
9064:Alamosaurus
8953:Udelartitan
8908:Patagotitan
8805:Bravasaurus
8798:Arrudatitan
8791:Aeolosaurus
8757:Overosaurus
8689:Jainosaurus
8667:Bonitasaura
8588:Paludititan
8479:Traukutitan
8174:Angolatitan
7981:Yunmenglong
7946:Qiaowanlong
7839:Sibirotitan
7804:Oceanotitan
7746:Europatitan
7725:Brontomerus
7718:Brohisaurus
7690:Angolatitan
7623:Galvesaurus
7595:Abydosaurus
7560:Ornithopsis
7553:Fusuisaurus
7539:Eucamerotus
7438:Daanosaurus
7423:Aragosaurus
7324:Supersaurus
7246:Apatosaurus
7209:Tharosaurus
7080:Nigersaurus
7031:Limaysaurus
6792:Ultrasaurus
6668:Turiasaurus
6631:Moabosaurus
6624:Mierasaurus
6593:Bellusaurus
6577:Turiasauria
6517:Qijianglong
6481:Hudiesaurus
6459:Datousaurus
6379:Cetiosaurus
6338:Shunosaurus
6282:Bellusaurus
6254:Algoasaurus
6240:Eusauropoda
6188:Sanpasaurus
6141:Gravisauria
6061:Pulanesaura
6046:Isanosaurus
6018:Amygdalodon
5884:Pulanesaura
5842:Irisosaurus
5828:Anchisaurus
5808:Anchisauria
5782:Xingxiulong
5728:Xingxiulong
5721:Sarahsaurus
5637:Riojasaurus
5603:Sarahsaurus
5589:Kholumolumo
5432:Macrocollum
5370:Asylosaurus
5306:Nhandumirim
5283:Buriolestes
5249:Mbiresaurus
5221:Buriolestes
4980:BBC iPlayer
4905:11336/90736
4657:: 133–148.
4513:Ameghiniana
3851:Patagotitan
3817:Patagotitan
3793:Smithsonian
3583:26 November
3294:Patagotitan
3254:peirosaurid
3249:Genyodectes
3238:Patagotitan
3230:Patagotitan
3211:C. australe
3179:angiosperms
3163:Patagotitan
3159:Patagotitan
3152:Patagotitan
3064:Patagotitan
2952:Aeolosaurus
2822:Saltasaurus
2684:Saltasaurus
2676:Patagotitan
2667:Patagotitan
2648:Patagotitan
2626:Patagotitan
2602:Patagotitan
2577:Patagotitan
2565:Patagotitan
2450:Patagotitan
2260:Bonitasaura
2209:Overosaurus
2192:Aeolosaurus
1974:Saltasaurus
1940:Alamosaurus
1663:Bonitasaura
1655:Alamosaurus
1642:Patagotitan
1630:Patagotitan
1624:In 2017, a
1601:Saltasaurus
1589:Bonitasaura
1581:Alamosaurus
1565:Saltasaurus
1545:Patagotitan
1541:Patagotitan
1519:Alamosaurus
1517:but unlike
1493:Alamosaurus
1480:Qiaowanlong
1432:Saltasaurus
1420:Patagotitan
1416:Saltasaurus
1372:Patagotitan
1344:Patagotitan
1295:Patagotitan
1270:Alamosaurus
1240:Saltasaurus
1219:Patagotitan
1178:Patagotitan
1168:but unlike
1157:Bonitasaura
1128:Patagotitan
1105:Patagotitan
1069:Patagotitan
1057:Patagotitan
1050:diapophyses
1038:, from the
1018:Patagotitan
1006:Patagotitan
1002:Patagotitan
973:Patagotitan
967:pleurocoels
953:Patagotitan
929:Patagotitan
925:Patagotitan
921:Patagotitan
902:Patagotitan
894:Patagotitan
886:Patagotitan
871:Patagotitan
857:Patagotitan
849:Patagotitan
829:Patagotitan
807:Description
772:Patagotitan
732:Patagotitan
644:pubic bones
577:Patagotitan
560:age of the
498:Patagotitan
437:Patagotitan
414:Patagotitan
398:Patagotitan
382:Patagotitan
341:Patagotitan
328:Carballido
301:Carballido
296:Patagotitan
25:Patagotitan
9488:Categories
9175:Rocasaurus
9102:Qunkasaura
9071:Baurutitan
8833:Shingopana
8660:Baalsaurus
8507:Volgatitan
8442:Rukwatitan
8406:Nullotitan
8392:Ninjatitan
8327:Kaijutitan
8298:Gandititan
8261:Daxiatitan
8224:Baurutitan
8167:Andesaurus
8058:see below↓
7753:Fukuititan
7697:Arkharavia
7652:Soriatitan
7602:Atlasaurus
7574:Rugocaudia
7532:Duriatitan
7468:Yuzhoulong
7416:Abrosaurus
7398:Macronaria
7385:Macronaria
7309:Kaatedocus
7302:Galeamopus
7295:Diplodocus
7281:Barosaurus
7188:Pilmatueia
7181:Lingwulong
7174:Kaatedocus
7094:Tataouinea
7006:Khebbashia
6981:Sidersaura
6711:Macronaria
6661:Tendaguria
6645:Oplosaurus
6609:Janenschia
6510:Omeisaurus
6386:Chebsaurus
6310:Janenschia
6224:Vulcanodon
6167:Kotasaurus
6118:Ledumahadi
5911:see below↓
5870:Meroktenos
5686:Leyesaurus
5575:Gyposaurus
5439:Unaysaurus
5398:Pantydraco
5363:Arcusaurus
5329:Saturnalia
5162:see below↓
5110:Dinosauria
5092:Sauropsida
4923:Chubutemys
4558:: 105389.
4476:: 104754.
4433:: 105487.
3316:References
3271:Chubutemys
3266:cryptodire
3264:, and the
3234:floodplain
2708:Andesaurus
2682:) but not
2660:Ninjatitan
2643:Ninjatitan
2579:to form a
1882:Baurutitan
1716:Andesaurus
1616:inserted.
1550:Rocasaurus
1524:Andesaurus
1245:Macronaria
1122:procoelous
1090:hyposphene
890:Matt Wedel
869:estimated
865:. In 2019
821:P. mayorum
782:Barosaurus
766:, New York
562:Cretaceous
502:floodplain
394:short tons
250:Macronaria
202:Saurischia
189:Dinosauria
9444:Q17103736
9409:Argentina
9397:Dinosaurs
9138:Bonatitan
9020:Isisaurus
8826:Punatitan
8305:Hamititan
8283:Elaltitan
8232:Bonatitan
7953:Silutitan
7910:Gobititan
7896:Euhelopus
7638:Lusotitan
7331:Tornieria
7317:Leinkupal
7202:Suuwassea
6916:Agustinia
6764:Dinodocus
6757:Cardiodon
6488:Jingiella
6437:Anhuilong
6000:Sauropoda
5987:Sauropoda
5902:Sauropoda
5877:Mussaurus
5835:Camelotia
5714:Pradhania
5554:Massopoda
5321:Panphagia
5263:Panphagia
5078:Kingdom:
4937:2246/5895
4861:131178611
4689:135268573
4580:252799392
4533:226680080
4498:233582290
4455:256559829
4411:213155599
4315:198493761
4095:130423764
4033:2045-2322
4011:: 19165.
3902:CiteSeerX
3862:8 January
3832:8 January
3798:8 January
3774:221404013
3758:1469-185X
3717:210840060
3639:8 January
3448:225012747
3226:siltstone
3224:-bearing
3218:sandstone
2039:Isisaurus
1511:Elaltitan
1497:Elaltitan
1485:Isisaurus
1440:Elaltitan
1404:Isisaurus
1325:than the
1284:Elaltitan
1249:Isisaurus
1228:Isisaurus
1098:hypantrum
1020:, unlike
939:Vertebrae
879:Histology
740:Patagonia
653:paratypes
526:La Flecha
512:Discovery
412:. Still,
364:Patagonia
354:from the
234:Sauropoda
162:Kingdom:
156:Eukaryota
9458:47458376
9438:Wikidata
9146:Ibirania
8812:Caieiria
8622:Inawentu
7825:Ruixinia
7666:Vouivria
7525:Astrodon
6778:Neosodon
6268:Bagualia
6111:Ingentia
5821:Aardonyx
5768:Qianlong
5596:Musankwa
5568:Gryponyx
5476:Ruehleia
5391:Nambalia
5384:Efraasia
5291:Eoraptor
5228:Eoraptor
5086:Chordata
5084:Phylum:
5080:Animalia
4822:24015414
4814:15834691
4738:92383900
4599:ABC News
4372:31598266
4307:31340217
4136:17768539
4051:26777391
3983:24348896
3943:PLOS ONE
3924:56028251
3766:32869488
3662:BBC News
3634:BBC News
3386:28794222
3304:See also
3188:Fitzroya
3175:conifers
3167:pinnules
2581:polytomy
1255:acromion
1186:spinalis
754:Exhibits
616:chevrons
612:, three
550:theropod
546:sauropod
459:scapulae
384:was the
352:dinosaur
349:sauropod
176:Chordata
172:Phylum:
166:Animalia
152:Domain:
9383:Portals
8182:Arackar
6586:Amanzia
6430:Analong
6317:Jobaria
5775:Seitaad
5090:Class:
5008:Ed Yong
4884:Bibcode
4784:Bibcode
4659:Bibcode
4560:Bibcode
4478:Bibcode
4435:Bibcode
4363:6731702
4342:Bibcode
4075:Bibcode
4042:4725985
4013:Bibcode
3974:3864407
3951:Bibcode
3894:Bibcode
3668:24 July
3555:24 July
3428:Bibcode
3377:5563814
3268:turtle
3183:cypress
3154:at dawn
2591:, with
1649:tons).
1632:as the
1499:. Like
1263:glenoid
1012:laminae
691:fibulae
671:, both
630:sternum
378:in 2017
374:in 2014
288:Genus:
133:, U.S.
105:↓
9471:358623
8711:Vahiny
8472:Tiamat
7889:Erketu
5707:Ngwevu
5010:, 2017
4985:4 July
4960:4 July
4859:
4820:
4812:
4736:
4687:
4630:17 May
4604:17 May
4578:
4531:
4496:
4453:
4409:
4370:
4360:
4313:
4305:
4216:
4134:
4093:
4049:
4039:
4031:
3981:
3971:
3922:
3904:
3772:
3764:
3756:
3715:
3608:4 July
3529:2 July
3498:2 July
3472:3 July
3446:
3384:
3374:
3252:, the
2608:, and
1661:, and
1603:, the
1599:, and
1563:, and
1533:, and
1477:, and
1414:, and
1366:, and
1309:, and
1287:, and
1237:, and
1196:, and
1150:centra
1140:, and
987:, and
852:'s
683:humeri
675:ischia
667:radius
596:, six
558:Albian
530:Trelew
496:life,
483:femora
475:ischia
467:humeri
390:tonnes
332:, 2017
330:et al.
305:, 2017
303:et al.
31:Albian
5106:Clade
5097:Clade
4857:S2CID
4818:S2CID
4734:S2CID
4685:S2CID
4576:S2CID
4529:S2CID
4494:S2CID
4451:S2CID
4407:S2CID
4311:S2CID
4091:S2CID
3920:S2CID
3882:(PDF)
3770:S2CID
3713:S2CID
3693:(PDF)
3444:S2CID
3171:ferns
3169:from
2654:than
2583:with
1459:Femur
1378:fossa
996:Like
909:'
744:Titan
579:bones
575:with
520:femur
427:Like
273:Clade
257:Clade
241:Clade
225:Clade
209:Clade
196:Clade
183:Clade
9013:Igai
5517:Issi
5139:see
4987:2023
4962:2023
4810:PMID
4632:2014
4606:2014
4368:PMID
4303:PMID
4214:ISBN
4132:PMID
4047:PMID
4029:ISSN
3979:PMID
3890:2004
3864:2019
3834:2019
3800:2019
3762:PMID
3754:ISSN
3670:2022
3641:2019
3610:2023
3585:2022
3557:2022
3531:2022
3500:2022
3474:2022
3382:PMID
3246:and
3222:tuff
3220:and
3191:and
2650:and
2636:and
2587:and
2571:and
1653:and
1575:and
1513:and
1495:and
1446:and
1430:and
1352:and
1333:and
1319:and
1164:and
1024:and
904:and
837:tail
835:and
833:neck
812:Size
718:and
663:and
659:ulna
624:ribs
479:and
463:and
447:and
443:back
406:and
392:(76
44:Preęž’
9453:EoL
6675:Zby
5006:by
4955:BBC
4933:hdl
4900:hdl
4892:doi
4849:doi
4800:hdl
4792:doi
4724:hdl
4716:doi
4675:hdl
4667:doi
4568:doi
4556:142
4521:doi
4486:doi
4474:122
4443:doi
4431:146
4399:doi
4358:PMC
4350:doi
4293:hdl
4285:doi
4255:hdl
4247:doi
4243:184
4206:doi
4177:hdl
4167:doi
4122:doi
4083:doi
4037:PMC
4021:doi
3969:PMC
3959:doi
3912:doi
3744:doi
3705:doi
3493:PBS
3436:doi
3372:PMC
3364:doi
3360:284
791:Sue
564:).
358:in
9490::
9468::
9455::
9440::
5108::
5099::
4978:.
4953:.
4927:.
4898:.
4890:.
4880:30
4878:.
4855:.
4845:24
4843:.
4816:.
4808:.
4798:.
4790:.
4780:92
4778:.
4761:23
4759:.
4755:.
4732:.
4722:.
4712:39
4710:.
4683:.
4673:.
4665:.
4655:99
4653:.
4649:.
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