4905:
7028:
4898:
3402:
4227:
4213:
1519:
6480:
5693:
6466:
4206:
6473:
3408:
1459:
4912:
144:
4220:
7041:
5711:
1405:
544:
5699:
5718:
7035:
1614:
6460:
1205:
827:
533:
1917:
119:
5706:
1601:, both in structure and distribution. Skin impressions were also preserved in the 2015 specimen, which indicated that, from mid-thigh to the feet, there was bare skin and that a flap of skin connecting the upper thigh to the torso. This latter structure is similar to that found in modern birds, including ostriches, but was positioned higher above the knee in
1435:. They would have been swift runners thanks to their very long limbs and hollow bones. They also had large brains and large eyes. The brains of ornithomimids in general were large for non-avialan dinosaurs, but this may not necessarily be a sign of high intelligence. Some paleontologists think that the enlarged portions of the brain were dedicated to
1937:
is still highly debated. As theropods, ornithomimids might have been carnivorous, but their body shape would have been suited for a largely omnivorous lifestyle. Suggested food in its diet includes insects, crustaceans, fruit, leaves, branches, eggs, lizards, and small mammals.
2378:
Brusatte, Choiniere, Benson, Carr and Norell, 2012. Theropod dinosaurs from the Late
Cretaceous of Eastern North America: Anatomy, systematics, biogeography and new information from historic specimens. Journal of Vertebrate Paleontology. Program and Abstracts 2012,
1557:
shafts, and a juvenile with impressions of feathers, which were up to five centimetres in length, in the form of hair-like filaments covering the rump, legs, and neck was also discovered). The fact that the feather imprints were found in
2546:"Dromiceiomimus." In: Dodson, Peter & Britt, Brooks & Carpenter, Kenneth & Forster, Catherine A. & Gillette, David D. & Norell, Mark A. & Olshevsky, George & Parrish, J. Michael & Weishampel, David B.
2748:
Xu, L.; Kobayashi, Y.; Lü, J.; Lee, Y. N.; Liu, Y.; Tanaka, K.; Zhang, X.; Jia, S.; Zhang, J. (2011). "A new ornithomimid dinosaur with North
American affinities from the Late Cretaceous Qiupa Formation in Henan Province of China".
2800:
Rothschild, B., Tanke, D. H., and Ford, T. L., 2001, Theropod stress fractures and tendon avulsions as a clue to activity: In: Mesozoic
Vertebrate Life, edited by Tanke, D. H., and Carpenter, K., Indiana University Press, p.
890:, respectively. At the same time, it was usual that workers referred to the entire ornithomimid material as simply "Struthiomimus". To solve this confusion by scientifically testing the separation between
1562:, previously thought to not be able to support such impressions, raised the possibility of finding similar structures with more careful preparation of future specimens. A study describing the fossils in
2614:
Zelenitsky, D. K.; Therrien, F.; Erickson, G. M.; Debuhr, C. L.; Kobayashi, Y.; Eberth, D. A.; Hadfield, F. (2012). "Feathered Non-Avian
Dinosaurs from North America Provide Insight into Wing Origins".
2721:
Van Der Reest, Aaron J.; Wolfe, Alexander P.; Currie, Philip J. (2016). "A densely feathered ornithomimid (Dinosauria: Theropoda) from the Upper
Cretaceous Dinosaur Park Formation, Alberta, Canada".
1236:
studied
Russell's 1972 proportional statistics to re-analyze ornithomimid relationships in light of newly discovered specimens. They concluded that there was no justification to separate
1570:
was covered in plumaceous feathers at all growth stages and that only adults had pennaceous wing-like structures, suggesting that wings may have evolved for mating displays. In
2824:
Reisdorf, A.G., and Wuttke, M. 2012. Re-evaluating Moodie's
Opisthotonic-Posture Hypothesis in fossil vertebrates. Part I: Reptiles - The taphonomy of the bipedal dinosaurs
2436:
Longrich, N. (2008). "A new, large ornithomimid from the
Cretaceous Dinosaur Park Formation of Alberta, Canada: Implications for the study of dissociated dinosaur remains."
1280:, though Longrich later suggested it may belong to a distinct, unnamed species from the Dinosaur Park Formation which has yet to be described. Longrich called the species
1309:. Dale Russell, in his 1972 revision of ornithomimids, could not determine which genus it actually belonged to, though he speculated that it may be intermediate between
2527:
1392:, which found that only the holotype specimen was confidently referable to that species. The authors of this study tentatively referred to the Kaiparowits specimen as
477:. Another seventeen species have been named since then, though almost all of them have been subsequently assigned to new genera or shown to be not directly related to
2271:
Russell, D.A. and
Chamney, T.P., 1967, "Notes on the biostratigraphy of dinosaurian and microfossil faunas in the Edmonton Formation (Cretaceous), Alberta",
577:
1427:
are characterized by feet with three weight-bearing toes, long slender arms, and long necks with birdlike, elongated, toothless, beaked skulls. They were
7199:
1225:
7254:
2528:
A preliminary report on the theropod dinosaur fauna of the late
Campanian Kaiparowits Formation, Grand Staircase-Escalante National Monument, Utah.
1119:. Like Russell, he considered the former two species indistinguishable from each other and noted that they both shared distinctive features with
726:
1052:
2341:
1455:, in having shorter torsos, long slender forearms, very slender, straight hand and foot claws, and hand bones and fingers of similar lengths.
1291:
dating to the early Maastrichtian, two other species are presently considered to be possibly valid and are also from the late Maastrichtian.
7249:
2567:
Makovicky, P.J., Kobayashi, Y., and Currie, P.J. (2004). "Ornithomimosauria." In Weishampel, D.B., Dodson, P., & Osmólska, H. (eds.),
2365:(Dinosauria: Theropoda: Ornithomimosauria), from the Upper Cretaceous Kirtland Formation (De-na-zin Member), San Juan Basin, New Mexico."
2146:
Gilmore, C.W., 1920, "Osteology of the carnivorous Dinosauria in the United States National Museum with special reference to the genera
7189:
1553:
specimens with evidence of feathers were found (two adults with carbonized traces on the lower arm, indicating the former presence of
1597:
specimen to preserve the feathers of its tail. The feathers, though crushed and distorted, bore numerous similarities to those of an
7194:
7120:
7244:
7239:
7234:
2785:
2598:
2414:
Kobayashi, Makovicky and Currie (2006). "Ornithomimids (Theropoda: Dinosauria) from the Late Cretaceous of Alberta, Canada."
906:
study. It showed that statistical differences in some proportions could be used to distinguish the two and he concluded that
2367:
New Mexico Geological Society Guidebook, 48th Field Conference, Mesozoic Geology and Paleontology of the Four Corners Region
2026:
4904:
4897:
2885:
1980:
7027:
2339:
Another Look at the Dinosaurs of the East Coast of North America. En (Colectivo Arqueológico-Paleontológico Salense, Ed.).
2864:
1478:
at 3.8 m (12 ft) and its weight at 170 kilograms (370 lb). One of its specimens, CMN 12228, preserves a
1269:
7214:
2555:
7229:
7219:
5692:
2821:
and new fossil discoveries. Journal of Vertebrate Paleontology, SVP Program and Abstracts Book, 2011, pp. 90.
1220:
Even after Russell's study, various researchers have found reasons to lump some or all of these species back into
7209:
4226:
2509:
Lillegraven and Eberle (1999). "Vertebrate faunal changes through Lancian and Puercan time in southern Wyoming."
4212:
1574:, Christian Foth and others argued that the evidence was insufficient to conclude that the forelimb feathers of
6465:
934:. However, Russell also interpreted the data as indicating that many specimens could not be referred to either
6906:
4205:
3401:
5710:
3407:
7179:
6479:
2204:
Parks, W.A., 1933, "New species of dinosaurs and turtles from the Upper Cretaceous formations of Alberta",
2091:
1685:
4911:
7184:
143:
4219:
2401:
Makovicky, Kobayashi and Currie (2004). "Ornithomimosauria." In Weishampel, Dodson and Osmolska (eds.),
1272:
annual meeting. Their opinion has been followed by most later authors. Makovicky's team also considered
7224:
6472:
1637:
studies indicate it having a derived position in the ornithomimids. These, however, have only included
1196:. This has found no acceptance among other workers and the name is not presently used by Paul himself.
490:
312:
5717:
7204:
7040:
811:
1952:. The large eye sockets suggest a keen visual sense and also suggest the possibility that they were
2952:
2338:
2492:(Theropoda, Ornithomimidae) from the terminal Cretaceous Kaiparowits Formation of southern Utah."
7034:
5698:
2857:
2037:
1750:
838:
At first, it had been common practice to name each newly discovered ornithomimid as a species of
706:
601:
560:
classification and the classification of ornithomimids in general has been very complicated. The
516:
2590:
7174:
7151:
7098:
7060:
2832:
from the Solnhofen Archipelago (Jurassic, Germany). Palaeobiodiversity and Palaeoenvironments,
1589:, this time from the lower portion of the Dinosaur Park Formation, was described in October of
1440:
698:
636:, but this changed when Hatcher found specimen USNM 4736, a partial ornithomimid skeleton, in
7146:
7138:
5423:
5335:
1714:
1365:
466:
263:
7107:
6992:
5112:
5080:
2679:
2624:
1960:
1805:
1661:
1622:
1590:
1571:
1563:
1546:
1542:
1530:
1522:
Life restoration of the plumage pattern suggested by specimens preserving feathers and skin
1413:
1385:
1373:
1318:
1265:
1229:
1161:
1142:
1128:
1100:
1088:
1048:
1040:
1031:
899:
871:
851:
780:
776:
762:
742:
734:
722:
718:
702:
688:
680:
672:
652:
647:
589:
462:
131:
6459:
1963:
study conducted by Bruce Rothschild and other paleontologists, 178 foot bones referred to
8:
5436:
5223:
1834:
1767:
1372:
was described in 1985. Re-evaluation of this specimen by Lindsay Zanno and colleagues in
1293:
642:
5705:
2683:
2628:
2102:
Lambe, L., 1902, "New genera and species from the Belly River Series (mid-Cretaceous)",
1439:
coordination. The bones of the hands are remarkably sloth-like in appearance, which led
818:
for a nearly complete skeleton from the Horseshoe Canyon Formation (specimen CMN 8632).
6899:
5412:
5272:
5128:
5068:
4503:
4413:
4296:
3160:
2850:
2703:
2648:
2583:
1578:
were necessarily pennaceous, citing the fact that the monofilamentous wing feathers in
1554:
1518:
1170:
1104:
1022:
944:
138:
2089:
Dinosaurs: The Most Complete, Up-to-Date Encyclopedia for Dinosaur Lovers of All Ages,
7133:
6781:
6698:
6672:
6511:
6498:
6376:
5926:
5878:
4710:
4060:
3139:
2781:
2695:
2640:
2594:
2551:
1630:
1420:
617:
493:. Additional species and specimens from other formations are sometimes classified as
405:
230:
2301:
Russell, D. (1972). "Ostrich dinosaurs from the Late Cretaceous of western Canada."
842:. In the sixties, this tendency was still very strong, as is shown by the fact that
481:. The best material of species still considered part of the genus has been found in
6885:
6642:
6552:
6341:
6265:
6128:
6047:
6019:
5979:
5860:
5830:
5348:
5101:
4965:
4822:
4722:
4571:
4478:
4436:
4003:
3313:
2833:
2758:
2730:
2707:
2687:
2665:
2652:
2632:
2470:
2010:
1353:
1124:
1075:, but Baird and Horner discovered that the name "Coelosaurus" was preoccupied by a
914:
were valid genera. In the latter, Russell recognised two species: the type species
754:
657:
581:
470:
347:
2762:
2734:
2474:
2014:
1529:, like many dinosaurs, was long thought to have been scaly. However, beginning in
6948:
6686:
6649:
6581:
6436:
6429:
6249:
6191:
6164:
6154:
6114:
5949:
5808:
5792:
5610:
5582:
5560:
5447:
5292:
4999:
4773:
4656:
4634:
4555:
4384:
4361:
4354:
4332:
4318:
4311:
4272:
4123:
3970:
3897:
3886:
3822:
3710:
3597:
3174:
3083:
2937:
2912:
2666:
Christian Foth; Helmut Tischlinger; Oliver W. M. Rauhut (2014). "New specimen of
2461:
from the upper Campanian of Alberta and implications for dinosaur biogeography".
2345:
1968:
1471:
1298:
1165:
661:
423:
30:
2526:
Zanno, L.E., Weirsma, J.P., Loewen, M.A., Sampson, S.D. and Getty, M.A. (2010).
2041:
6913:
6876:
6857:
6819:
6808:
6605:
6318:
6276:
6258:
6238:
6135:
5849:
5594:
5456:
5283:
5265:
5093:
5060:
5046:
5007:
4781:
4642:
4599:
4592:
4514:
4490:
4443:
4402:
4339:
4325:
4053:
4038:
3865:
3699:
3563:
3531:
3520:
3497:
3459:
3320:
3290:
3214:
3153:
3052:
1671:
1213:
1182:
1068:
1056:
966:
863:
804:
676:
625:
505:
243:
2837:
2221:
Gilmore, C.W., 1933, "On the dinosaurian fauna of the Iren Dabasu Formation",
576:
YPM 542 and YPM 548 (a partial hindlimb and forelimb, respectively), found by
7168:
7083:
6955:
6801:
6762:
6741:
6620:
6613:
6540:
6334:
6307:
6173:
6090:
6068:
6061:
6054:
5997:
5914:
5777:
5749:
5736:
5660:
5537:
5377:
5211:
5142:
5028:
5015:
4760:
4738:
4731:
4695:
4668:
4620:
4544:
4280:
4071:
4045:
3961:
3942:
3935:
3872:
3851:
3830:
3815:
3801:
3760:
3726:
3719:
3638:
3490:
3377:
3370:
3297:
3267:
3256:
3146:
3117:
3110:
3036:
2813:
Claessens, L., Loewen, M. and Lavender, Z. 2011. A reevaluation of the genus
1458:
1451:
1348:, at first known from very limited remains. Additional specimens referred to
1176:
789:
693:
550:
511:
433:
was a swift, bipedal dinosaur which fossil evidence indicates was covered in
419:
76:
38:
2636:
1404:
6865:
6835:
6755:
6748:
6726:
6718:
6598:
6590:
6570:
6285:
6204:
6142:
6033:
6004:
5956:
5785:
5770:
5545:
5477:
5405:
5384:
5363:
5179:
5135:
4943:
4930:
4860:
4813:
4688:
4578:
4530:
4523:
4460:
4429:
4422:
4303:
4130:
3982:
3879:
3781:
3774:
3669:
3653:
3589:
3540:
3472:
3249:
3132:
3125:
3096:
3059:
3028:
2699:
2644:
2193:
Second Bibliography and Catalogue of the Fossil Vertebrata of North America
1926:
1487:
1334:
1137:
1084:
1076:
1036:
867:
771:
561:
442:
272:
2778:
The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals
2318:
Baird D., and Horner, J., 1979, "Cretaceous dinosaurs of North Carolina",
2174:
Russell, L.S., 1930, "Upper Cretaceous dinosaur faunas of North America",
543:
7092:
6981:
6962:
6892:
6842:
6828:
6770:
6711:
6656:
6533:
6327:
6211:
6180:
6097:
6083:
5815:
5763:
5631:
5553:
5516:
5398:
5370:
5323:
5243:
5121:
4978:
4829:
4802:
4702:
4649:
4585:
4564:
4537:
4288:
4101:
4094:
4080:
3906:
3753:
3744:
3646:
3617:
3547:
3512:
3505:
3305:
3283:
3276:
3235:
3227:
3203:
3188:
3167:
3075:
3067:
1579:
1436:
1233:
903:
51:
2691:
7125:
6920:
6849:
6794:
6415:
6222:
6040:
6026:
6011:
5963:
5897:
5823:
5645:
5638:
5624:
5530:
5463:
5391:
5355:
5308:
5250:
5236:
5203:
5195:
5188:
5169:
4992:
4957:
4745:
4346:
4137:
4087:
4024:
3995:
3950:
3928:
3921:
3913:
3837:
3733:
3692:
3685:
3677:
3661:
3581:
3452:
3328:
3181:
3044:
2903:
2071:
O.C. Marsh, 1892, "Notice of new reptiles from the Laramie Formation",
1503:
1256:
in this analysis. The same team further supported the synonymy between
1188:
1026:
843:
795:
633:
592:. The generic name means "bird mimic", derived from Greek words ὄρνις (
569:
438:
201:
96:
61:
1613:
1593:
by Aaron van der Reest, Alex Wolfe, and Phil Currie. It was the first
6941:
6934:
6634:
6627:
6300:
6292:
6231:
5885:
5603:
5149:
5053:
4788:
4373:
4258:
4245:
4172:
4108:
4031:
4010:
3767:
3630:
3438:
3425:
3343:
3195:
3014:
3001:
2964:
2873:
1953:
1634:
1559:
1204:
1079:
taxon, which was based on a single vertebra. It was originally named
826:
532:
426:
415:
408:
214:
155:
101:
45:
34:
7054:
2001:
Claessens, L. & Mark A. Loewen, M.A. (2015). A redescription of
1396:, along with all of the specimens from the Dinosaur Park Formation.
1135:
to be indeterminate among ornithomimosaurs, resulting in it being a
7077:
6525:
5942:
5935:
5907:
5800:
5300:
4681:
4017:
3844:
3605:
3242:
2921:
2897:
2104:
Geological Survey of Canada Contributions to Canadian Palaeontology
1916:
1495:
1092:
585:
474:
411:
188:
175:
91:
86:
71:
66:
56:
118:
6075:
5837:
5567:
5470:
4795:
4116:
1944:
had legs that seem clearly suited for rapid locomotion, with the
1598:
1538:
1357:
1330:
1302:
855:
684:
637:
621:
573:
482:
434:
106:
81:
7112:
2613:
1443:
to suggest that they were used to hook branches during feeding.
6734:
6422:
3858:
2891:
2542:
2540:
2031:
1432:
165:
2670:
provides insights into the evolution of pennaceous feathers".
1470:
species today seen as possibly valid differ in size. In 2010,
437:
and equipped with a small toothless beak that may indicate an
5574:
5315:
5257:
2054:
O.C. Marsh, 1890, "Description of new dinosaurian reptiles",
1949:
1945:
1479:
1428:
1297:
was named by Marsh in 1892 from partial remains found in the
1091:. Baird referred several other specimens from New Jersey and
1018:
605:
401:
2842:
2537:
922:(even though he had trouble reliably distinguishing it from
389:
362:
353:
6998:
5617:
1369:
1352:
have been described from the Denver Formation and from the
392:
380:
374:
365:
16:
Ornithomimid dinosaur genus from the Late Cretaceous Period
2532:
Learning from the Land Symposium: Geology and Paleontology
2333:
2331:
974:
mimic". This comes from the old generic name for the emu:
942:. Therefore, he created two new genera. The first one was
2350:
Actas de las III Jornadas sobre Dinosaurios y su Entorno.
971:
359:
2720:
1494:, is based on material of a smaller animal. Whereas the
2395:
2328:
660:), but it has since been recognized as belonging to an
1337:), these have yet to be described and compared to the
679:
and was based on specimen CMN 930 (hindlimbs found in
2589:. Princeton, NJ: Princeton University Press. p.
2195:. Carnegie Institution of Washington. 390(II): 1-1074
1997:
1995:
1633:, a classification that is still very common. Modern
441:
diet. It is usually classified into two species: the
383:
356:
350:
2571:. University of California Press, Berkeley: 137-150.
1506:
eighty-four millimetres long, the same element with
600:), "mimic", in reference to the bird-like foot. The
319:
302:
279:
6407:
2714:
2503:
2258:Kuhn, O., 1965, "Saurischia (Supplementum 1)". In:
1325:. Although it has since been referred to mainly as
821:
377:
371:
368:
2582:
2223:Bulletin of the American Museum of Natural History
2132:Bulletin of the American Museum of Natural History
1992:
1284:in a faunal list for the Dinosaur Park Formation.
2488:DeCourten and Russell, D. (1985). "A specimen of
2176:Proceedings of the American Philosophical Society
1252:. However, they did not include the type species
982:species named during the 20th century, including
608:. Simultaneously, Marsh named two other species:
469:on the basis of a foot and partial hand from the
7166:
6509:
6375:
2747:
2273:National Museum of Canada Natural History Papers
2206:University of Toronto Studies, Geological Series
5858:
2794:
2432:
2430:
2428:
2352:129-168. Salas de los Infantes, Burgos, España.
1145:abstract agreed with Weishampel by noting that
687:), but this was renamed to a separate genus in
656:was also created based on specimen YPM 1049 (a
616:. Both consist of fragmentary fossils found by
4382:
2936:
2561:
2484:
2482:
2453:Longrich, N. R. (2014). "The horned dinosaurs
1482:that is 46.8 centimetres (18.4 in) long.
1009:
2858:
2520:
1641:in their analyses. The relationships between
5747:
2817:based on new preparation of the holotype of
2534:. Washington, DC: Bureau of Land Management.
2425:
2355:
2118:H.F. Osborn, 1916, "Skeletal adaptations of
956:were reassigned to this new genus, becoming
4941:
4859:
3981:
2741:
2550:. Publications International, LTD. p. 140.
2479:
2295:
1376:, however, cast doubt on its assignment to
7200:Late Cretaceous dinosaurs of North America
2865:
2851:
2780:. London: Marshall Editions. p. 109.
2775:
1449:differs from other ornithomimids, such as
1199:
117:
2405:. University of California Press. 861 pp.
2290:The Illustrated Encyclopedia of Dinosaurs
1216:deformed bones reconstructed on the right
4256:
4171:
3436:
3012:
2392:. Simon & Schuster: New York. 464 pp
2081:
1915:
1612:
1517:
1457:
1403:
1344:The other is the original type species:
1203:
825:
542:
531:
489:, known from several skeletons from the
7255:Multispecific non-avian dinosaur genera
2408:
1656:The following cladogram is based on Xu
1537:have been found preserving evidence of
1510:measures only fifty-three millimetres.
1083:by an anonymous author now known to be
721:), based on indeterminate material. In
522:
7167:
2585:The Princeton Field Guide to Dinosaurs
2160:United States National Museum Bulletin
1924:being restrained while preyed upon by
1321:, he considered it to be a species of
1043:. The tibiae were first attributed to
527:
503:(alternately classified in the genera
7059:
7058:
6497:
6496:
5734:
4928:
4243:
3423:
2999:
2884:
2846:
2005:Marsh, 1890 (Dinosauria, Theropoda).
1384:. This conclusion was supported by a
2769:
2580:
1981:Timeline of ornithomimosaur research
1513:
1333:and some fragments from Alberta and
862:(which is understood today to be an
753:, respectively. The very same year,
7250:Taxa named by Othniel Charles Marsh
1582:would likely leave similar traces.
13:
2807:
2416:Journal of Vertebrate Paleontology
2303:Canadian Journal of Earth Sciences
2242:with complete abdominal cuirass",
2007:Journal of Vertebrate Paleontology
1329:(based on complete specimens from
1270:Society of Vertebrate Paleontology
1224:in various combinations. In 2004,
1168:classified the various species of
978:. Russell assigned several former
628:material. At first, Marsh assumed
612:(based on specimen USNM 5814) and
14:
7266:
7190:Campanian genus first appearances
5735:
2292:, Crescent Books, New York, p. 48
1608:
1063:. Normally, this would have made
7039:
7033:
7026:
6478:
6471:
6464:
6458:
5716:
5709:
5704:
5697:
5691:
4929:
4910:
4903:
4896:
4225:
4218:
4211:
4204:
3406:
3400:
2390:Predatory Dinosaurs of the World
1948:being about 20% longer than the
1431:and superficially resemblant to
1412:found in 1995 with quill knobs,
1305:a year after the description of
822:Reclassification by Dale Russell
799:. That same year, Gilmore named
787:, which is today seen as either
346:
142:
49:
7195:Maastrichtian genus extinctions
2659:
2607:
2574:
2569:The Dinosauria (second edition)
2447:
2382:
2372:
2312:
2282:
2265:
2260:Fossilium Catalogus 1. Animalia
2252:
2232:
2215:
2198:
2185:
2056:The American Journal of Science
1911:
1585:A fourth feathered specimen of
1502:, CMN 8632, preserves a second
1212:skull RTMP 1995.110.0001, with
1157:in the features of the tibiae.
624:, which is today understood as
2361:Sullivan, (1997). "A juvenile
2238:Sternberg, C.M., 1933, "A new
2168:
2140:
2112:
2096:
2065:
2048:
2019:
2009:(advance online publication).
1399:
1:
7245:Fossil taxa described in 1890
4244:
3424:
3000:
2872:
2763:10.1016/j.cretres.2010.12.004
2735:10.1016/j.cretres.2015.10.004
2475:10.1016/j.cretres.2014.06.011
2403:The Dinosauria Second Edition
2244:The Canadian Field-Naturalist
2015:10.1080/02724634.2015.1034593
1986:
1462:Size of the two valid species
1360:. One specimen attributed to
1035:("antique hollow lizard") by
7240:Paleontology in North Dakota
7235:Paleontology in South Dakota
2087:Holtz, Thomas R. Jr. (2011)
1686:Archaeornithomimus asiaticus
958:Archaeornithomimus asiaticus
7:
2073:American Journal of Science
1974:
1967:were examined for signs of
992:Dromiceiomimus brevitertius
930:to be a younger synonym of
769:, which is today seen as a
485:, representing the species
10:
7271:
1276:to be a junior synonym of
962:Archaeornithomimus affinis
731:Struthiomimus brevetertius
491:Horseshoe Canyon Formation
451:, and a referred species,
7067:
7023:
6980:
6875:
6818:
6789:
6780:
6706:
6697:
6681:
6670:
6580:
6565:
6561:
6550:
6520:
6505:
6492:
6456:
6403:
6371:
6317:
6275:
6248:
6221:
6199:
6190:
6163:
6152:
6123:
6112:
5992:
5988:
5977:
5925:
5896:
5873:
5869:
5847:
5758:
5743:
5730:
5689:
5659:
5593:
5525:
5515:
5511:
5446:
5431:
5422:
5343:
5334:
5282:
5231:
5222:
5178:
5167:
5111:
5088:
5079:
5041:
5037:
5026:
4987:
4976:
4952:
4937:
4924:
4893:
4855:
4812:
4768:
4759:
4721:
4676:
4667:
4629:
4618:
4554:
4513:
4498:
4489:
4473:
4469:
4458:
4412:
4397:
4393:
4371:
4267:
4252:
4239:
4201:
4167:
4070:
3990:
3960:
3896:
3810:
3799:
3743:
3709:
3625:
3616:
3576:
3572:
3561:
3530:
3485:
3481:
3470:
3447:
3432:
3419:
3397:
3362:
3342:
3266:
3222:
3213:
3105:
3094:
3023:
3008:
2995:
2963:
2945:
2932:
2880:
2838:10.1007/s12549-011-0068-y
2581:Paul, Gregory S. (2010).
2337:Weishampel, D.B.(2004). "
1852:Ornithomimus edmontonicus
1848:
1831:
1824:
1802:
1795:
1764:
1747:
1740:
1733:
1711:
1704:
1682:
1675:
1653:have not been published.
1551:Ornithomimus edmontonicus
1410:Ornithomimus edmontonicus
932:Ornithomimus edmontonicus
920:Ornithomimus edmontonicus
816:Ornithomimus edmontonicus
812:Charles Mortram Sternberg
747:Ornithomimus brevitertius
454:Ornithomimus edmontonicus
299:
294:
278:
271:
139:Scientific classification
137:
125:
116:
23:
7215:Paleontology in Colorado
2776:Palmer, D., ed. (1999).
1474:estimated the length of
1364:(MNA P1 1762A) from the
572:in 1890 and is based on
7230:Paleontology in Montana
7220:Paleontology in Alberta
2637:10.1126/science.1225376
2511:Journal of Paleontology
2494:Journal of Paleontology
2038:Oxford University Press
1971:, but none were found.
1751:Anserimimus planinychus
1533:, several specimens of
1200:Present interpretations
990:, to this new genus as
928:Struthiomimus currellii
876:Struthiomimus currellii
860:Ornithomimus lonzeensis
848:Megalosaurus lonzeensis
707:Charles Whitney Gilmore
517:Dinosaur Park Formation
7210:Horseshoe Canyon fauna
2826:Compsognathus longipes
1930:
1618:
1523:
1463:
1441:Henry Fairfield Osborn
1416:
1274:Dromiceiomimus samueli
1217:
1115:as junior synonyms of
1004:Dromiceiomimus samueli
950:Ornithomimus asiaticus
884:Ornithomimus currellii
835:
803:for material found in
801:Ornithomimus asiaticus
699:Henry Fairfield Osborn
596:), "bird", and μῖμος (
553:
540:
7147:Paleobiology Database
5424:Carcharodontosaurinae
5336:Carcharodontosauridae
2363:Ornithomimus antiquus
2092:Winter 2010 Appendix.
2034:UK English Dictionary
1919:
1715:Sinornithomimus dongi
1616:
1521:
1461:
1407:
1380:and possibly even to
1366:Kaiparowits Formation
1207:
1061:Ornithomimus antiquus
964:. The second one was
829:
767:Ornithomimus mirandus
739:Struthiomimus samueli
568:, was first named by
546:
536:Holotype material of
535:
467:Othniel Charles Marsh
427:Western North America
400:; "bird mimic") is a
6408:Dubious coelurosaurs
5113:Metriacanthosaurinae
5081:Metriacanthosauridae
3363:Dubious neotheropods
2548:The Age of Dinosaurs
1806:Qiupalong henanensis
1414:Royal Tyrrell Museum
1327:Struthiomimus sedens
1282:Ornithomimus samueli
1230:Yoshitsugu Kobayashi
1032:Coelosaurus antiquus
996:Ornithomimus samueli
954:Ornithomimus affinis
880:Struthiomimus ingens
785:Ornithomimus elegans
783:created the species
781:William Arthur Parks
751:Ornithomimus samueli
711:Ornithomimus affinis
653:Ornithomimus minutus
650:. On that occasion,
614:Ornithomimus grandis
523:History of discovery
500:Ornithomimus samueli
132:Royal Ontario Museum
7180:Feathered dinosaurs
5437:Carcharodontosaurus
5224:Carcharodontosauria
2751:Cretaceous Research
2723:Cretaceous Research
2692:10.1038/nature13467
2684:2014Natur.511...79F
2629:2012Sci...338..510Z
2463:Cretaceous Research
1835:Struthiomimus altus
1768:Gallimimus bullatus
888:Ornithomimus ingens
834:sp. (RTMP 95.110.1)
759:Aublysodon mirandus
643:Ornithomimus sedens
610:Ornithomimus tenuis
578:George Lyman Cannon
528:First species named
515:) from the earlier
7185:Maastrichtian life
6900:Archaeornithomimus
5413:Veterupristisaurus
5273:Veterupristisaurus
5129:Metriacanthosaurus
5069:Piatnitzkysauridae
4504:Eustreptospondylus
4414:Piatnitzkysauridae
4297:Chuandongocoelurus
3161:Notatesseraeraptor
2830:Juravenator starki
2490:Ornithomimus velox
2388:Paul, G.S., 1988,
2344:2012-09-04 at the
2288:Norman, D., 1985,
2003:Ornithomimus velox
1931:
1619:
1555:pennaceous feather
1524:
1464:
1417:
1388:re-description of
1218:
1171:Archaeornithomimus
1131:) considered "C."
1105:Robert M. Sullivan
1023:Navesink Formation
994:. He also renamed
945:Archaeornithomimus
916:Ornithomimus velox
898:, Dale Russell in
882:(Parks 1933) into
836:
830:Skull and neck of
814:named the species
715:Dryosaurus grandis
669:Ornithomimus altus
566:Ornithomimus velox
554:
548:Ornithomimus velox
541:
479:Ornithomimus velox
448:Ornithomimus velox
284:Ornithomimus velox
7225:Laramie Formation
7162:
7161:
7134:Open Tree of Life
7061:Taxon identifiers
7052:
7051:
7048:
7047:
7021:
7020:
7017:
7016:
7013:
7012:
7009:
7008:
6976:
6975:
6972:
6971:
6782:Macrocheiriformes
6699:Ornithomimosauria
6673:Maniraptoriformes
6666:
6665:
6512:Maniraptoromorpha
6499:Maniraptoromorpha
6488:
6487:
6454:
6453:
6450:
6449:
6446:
6445:
6399:
6398:
6395:
6394:
6377:Maniraptoromorpha
6367:
6366:
6363:
6362:
6359:
6358:
6355:
6354:
6351:
6350:
6108:
6107:
5973:
5972:
5927:Proceratosauridae
5879:Chingkankousaurus
5726:
5725:
5687:
5686:
5683:
5682:
5679:
5678:
5655:
5654:
5507:
5506:
5503:
5502:
5499:
5498:
5495:
5494:
5491:
5490:
5487:
5486:
5163:
5162:
5159:
5158:
4920:
4919:
4891:
4890:
4887:
4886:
4883:
4882:
4879:
4878:
4851:
4850:
4847:
4846:
4843:
4842:
4839:
4838:
4755:
4754:
4711:Vallibonavenatrix
4614:
4613:
4610:
4609:
4454:
4453:
4235:
4234:
4199:
4198:
4195:
4194:
4191:
4190:
4163:
4162:
4159:
4158:
4155:
4154:
4151:
4150:
4147:
4146:
3795:
3794:
3791:
3790:
3557:
3556:
3415:
3414:
3395:
3394:
3391:
3390:
3387:
3386:
3338:
3337:
2991:
2990:
2987:
2986:
2983:
2982:
2787:978-1-84028-152-1
2623:(6106): 510–514.
2600:978-0-691-13720-9
2191:Hay, O.P., 1930,
1908:
1907:
1899:
1898:
1890:
1889:
1881:
1880:
1872:
1871:
1863:
1862:
1779:
1778:
1631:Ornithomimosauria
1625:, Marsh assigned
1514:Feathers and skin
926:). He considered
878:(Parks 1933) and
667:A sixth species,
640:. Marsh named it
618:John Bell Hatcher
604:means "swift" in
337:
336:
331:
316:
267:
231:Ornithomimosauria
7262:
7205:Hell Creek fauna
7155:
7154:
7142:
7141:
7129:
7128:
7116:
7115:
7103:
7102:
7101:
7088:
7087:
7086:
7056:
7055:
7043:
7037:
7030:
6886:Aepyornithomimus
6787:
6786:
6704:
6703:
6679:
6678:
6643:Sinocalliopteryx
6563:
6562:
6559:
6558:
6553:Neocoelurosauria
6518:
6517:
6507:
6506:
6494:
6493:
6482:
6475:
6468:
6462:
6405:
6404:
6373:
6372:
6342:Zhuchengtyrannus
6266:Thanatotheristes
6197:
6196:
6161:
6160:
6129:Appalachiosaurus
6121:
6120:
5990:
5989:
5986:
5985:
5980:Pantyrannosauria
5871:
5870:
5867:
5866:
5861:Tyrannosauroidea
5856:
5855:
5831:Xinjiangovenator
5756:
5755:
5745:
5744:
5732:
5731:
5720:
5713:
5708:
5701:
5695:
5523:
5522:
5513:
5512:
5429:
5428:
5349:Acrocanthosaurus
5341:
5340:
5229:
5228:
5176:
5175:
5102:Yangchuanosaurus
5086:
5085:
5039:
5038:
5035:
5034:
4985:
4984:
4966:Lourinhanosaurus
4950:
4949:
4939:
4938:
4926:
4925:
4914:
4907:
4900:
4857:
4856:
4823:Sigilmassasaurus
4766:
4765:
4723:Ceratosuchopsini
4674:
4673:
4627:
4626:
4572:Dubreuillosaurus
4496:
4495:
4479:Streptospondylus
4471:
4470:
4467:
4466:
4437:Piatnitzkysaurus
4395:
4394:
4391:
4390:
4380:
4379:
4265:
4264:
4254:
4253:
4241:
4240:
4229:
4222:
4215:
4208:
4169:
4168:
4004:Ekrixinatosaurus
3988:
3987:
3979:
3978:
3808:
3807:
3623:
3622:
3574:
3573:
3570:
3569:
3483:
3482:
3479:
3478:
3445:
3444:
3434:
3433:
3421:
3420:
3410:
3404:
3314:Procompsognathus
3220:
3219:
3103:
3102:
3021:
3020:
3010:
3009:
2997:
2996:
2943:
2942:
2934:
2933:
2927:
2926:
2882:
2881:
2867:
2860:
2853:
2844:
2843:
2802:
2798:
2792:
2791:
2773:
2767:
2766:
2745:
2739:
2738:
2718:
2712:
2711:
2663:
2657:
2656:
2611:
2605:
2604:
2588:
2578:
2572:
2565:
2559:
2544:
2535:
2524:
2518:
2507:
2501:
2486:
2477:
2451:
2445:
2434:
2423:
2412:
2406:
2399:
2393:
2386:
2380:
2376:
2370:
2359:
2353:
2335:
2326:
2316:
2310:
2299:
2293:
2286:
2280:
2269:
2263:
2256:
2250:
2236:
2230:
2219:
2213:
2202:
2196:
2189:
2183:
2172:
2166:
2144:
2138:
2116:
2110:
2100:
2094:
2085:
2079:
2069:
2063:
2052:
2046:
2045:
2040:. Archived from
2023:
2017:
1999:
1827:
1826:
1821:
1798:
1797:
1792:
1743:
1742:
1736:
1735:
1730:
1707:
1706:
1701:
1678:
1677:
1668:
1667:
1394:Ornithomimus sp.
1354:Ferris Formation
1248:as a synonym of
1125:David Weishampel
755:Oliver Perry Hay
582:Denver Formation
471:Denver Formation
399:
398:
395:
394:
391:
386:
385:
382:
379:
376:
373:
370:
367:
364:
361:
358:
355:
352:
329:
321:
311:
304:
281:
262:
255:
242:
229:
147:
146:
121:
111:
48:
29:Temporal range:
21:
20:
7270:
7269:
7265:
7264:
7263:
7261:
7260:
7259:
7165:
7164:
7163:
7158:
7150:
7145:
7137:
7132:
7124:
7119:
7111:
7106:
7097:
7096:
7091:
7082:
7081:
7076:
7063:
7053:
7044:
7031:
7005:
6968:
6949:Sinornithomimus
6871:
6814:
6776:
6693:
6687:Compsognathidae
6675:
6662:
6650:Sinosauropteryx
6582:Compsognathidae
6576:
6555:
6546:
6514:
6501:
6484:
6483:
6469:
6442:
6437:Shanyangosaurus
6430:Phaedrolosaurus
6391:
6347:
6313:
6271:
6250:Daspletosaurini
6244:
6217:
6192:Tyrannosaurinae
6186:
6165:Albertosaurinae
6157:
6155:Tyrannosauridae
6148:
6117:
6115:Eutyrannosauria
6104:
5982:
5969:
5950:Proceratosaurus
5921:
5892:
5863:
5852:
5843:
5809:Richardoestesia
5793:Chilantaisaurus
5752:
5739:
5722:
5721:
5702:
5675:
5651:
5611:Australovenator
5589:
5583:Chilantaisaurus
5561:Phuwiangvenator
5483:
5448:Giganotosaurini
5442:
5418:
5330:
5293:Chilantaisaurus
5278:
5218:
5172:
5155:
5107:
5075:
5031:
5022:
5000:Monolophosaurus
4981:
4972:
4946:
4933:
4916:
4915:
4901:
4875:
4835:
4808:
4774:Camarillasaurus
4751:
4717:
4663:
4657:Ostafrikasaurus
4635:Camarillasaurus
4623:
4606:
4556:Afrovenatorinae
4550:
4509:
4485:
4463:
4450:
4408:
4387:
4385:Megalosauroidea
4376:
4367:
4362:Vectaerovenator
4355:Szechuanosaurus
4333:Monolophosaurus
4319:Kaijiangosaurus
4312:Cryolophosaurus
4273:Calamospondylus
4261:
4248:
4231:
4230:
4223:
4216:
4209:
4187:
4143:
4124:Pycnonemosaurus
4066:
3977:
3971:Majungasaurinae
3956:
3898:Majungasaurinae
3892:
3887:Xenotarsosaurus
3823:Dryptosauroides
3804:
3787:
3739:
3711:Elaphrosaurinae
3705:
3612:
3598:Ornithomimoides
3566:
3553:
3526:
3475:
3466:
3441:
3428:
3411:
3405:
3383:
3358:
3334:
3262:
3209:
3175:Shuangbaisaurus
3099:
3090:
3017:
3004:
2979:
2959:
2953:Avemetatarsalia
2938:Avemetatarsalia
2928:
2913:Avemetatarsalia
2887:
2886:
2876:
2871:
2810:
2808:Further reading
2805:
2799:
2795:
2788:
2774:
2770:
2746:
2742:
2719:
2715:
2678:(7507): 79–82.
2664:
2660:
2612:
2608:
2601:
2579:
2575:
2566:
2562:
2545:
2538:
2525:
2521:
2508:
2504:
2500:(5): 1091-1099.
2487:
2480:
2452:
2448:
2435:
2426:
2413:
2409:
2400:
2396:
2387:
2383:
2377:
2373:
2360:
2356:
2346:Wayback Machine
2336:
2329:
2317:
2313:
2300:
2296:
2287:
2283:
2270:
2266:
2257:
2253:
2237:
2233:
2220:
2216:
2203:
2199:
2190:
2186:
2173:
2169:
2145:
2141:
2117:
2113:
2101:
2097:
2086:
2082:
2070:
2066:
2053:
2049:
2025:
2024:
2020:
2000:
1993:
1989:
1977:
1969:stress fracture
1914:
1909:
1900:
1891:
1882:
1873:
1864:
1822:
1819:
1793:
1790:
1780:
1731:
1728:
1702:
1699:
1643:O. edmontonicus
1639:O. edmontonicus
1611:
1605:than in birds.
1568:O. edmontonicus
1566:concluded that
1516:
1500:O. edmontonicus
1476:O. edmontonicus
1472:Gregory S. Paul
1402:
1299:Lance Formation
1289:O. edmontonicus
1278:O. edmontonicus
1268:lecture at the
1262:O. edmontonicus
1250:O. edmontonicus
1226:Peter Makovicky
1210:O. edmontonicus
1202:
1166:Gregory S. Paul
1113:O. edmontonicus
1099:. Beginning in
1015:
1010:Misassigned to
984:O. brevitertius
824:
671:, was named in
556:The history of
530:
525:
487:O. edmontonicus
424:Late Cretaceous
388:
349:
345:
328:
310:
307:O. edmontonicus
290:
287:
261:
253:
240:
227:
141:
128:O. edmontonicus
112:
110:
109:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
43:
42:
31:Late Cretaceous
27:
17:
12:
11:
5:
7268:
7258:
7257:
7252:
7247:
7242:
7237:
7232:
7227:
7222:
7217:
7212:
7207:
7202:
7197:
7192:
7187:
7182:
7177:
7160:
7159:
7157:
7156:
7143:
7130:
7117:
7104:
7089:
7073:
7071:
7065:
7064:
7050:
7049:
7046:
7045:
7025:
7024:
7022:
7019:
7018:
7015:
7014:
7011:
7010:
7007:
7006:
7004:
7003:
7002:
7001:
6995:
6986:
6984:
6978:
6977:
6974:
6973:
6970:
6969:
6967:
6966:
6959:
6952:
6945:
6938:
6931:
6924:
6917:
6914:Dromiceiomimus
6910:
6903:
6896:
6889:
6881:
6879:
6877:Ornithomimidae
6873:
6872:
6870:
6869:
6862:
6858:Paraxenisaurus
6854:
6846:
6839:
6832:
6824:
6822:
6820:Deinocheiridae
6816:
6815:
6813:
6812:
6809:Shenzhousaurus
6805:
6798:
6790:
6784:
6778:
6777:
6775:
6774:
6767:
6759:
6752:
6745:
6738:
6731:
6723:
6715:
6707:
6701:
6695:
6694:
6692:
6691:
6682:
6676:
6671:
6668:
6667:
6664:
6663:
6661:
6660:
6653:
6646:
6639:
6631:
6624:
6617:
6610:
6606:Beipiaognathus
6602:
6595:
6586:
6584:
6578:
6577:
6575:
6574:
6566:
6556:
6551:
6548:
6547:
6545:
6544:
6537:
6530:
6521:
6515:
6510:
6503:
6502:
6490:
6489:
6486:
6485:
6477:
6463:
6457:
6455:
6452:
6451:
6448:
6447:
6444:
6443:
6441:
6440:
6433:
6426:
6419:
6411:
6409:
6401:
6400:
6397:
6396:
6393:
6392:
6390:
6389:
6388:
6387:
6381:
6379:
6369:
6368:
6365:
6364:
6361:
6360:
6357:
6356:
6353:
6352:
6349:
6348:
6346:
6345:
6338:
6331:
6323:
6321:
6319:Tyrannosaurini
6315:
6314:
6312:
6311:
6304:
6297:
6289:
6281:
6279:
6277:Teratophoneini
6273:
6272:
6270:
6269:
6262:
6259:Daspletosaurus
6254:
6252:
6246:
6245:
6243:
6242:
6239:Qianzhousaurus
6235:
6227:
6225:
6219:
6218:
6216:
6215:
6208:
6200:
6194:
6188:
6187:
6185:
6184:
6177:
6169:
6167:
6158:
6153:
6150:
6149:
6147:
6146:
6139:
6136:Bistahieversor
6132:
6124:
6118:
6113:
6110:
6109:
6106:
6105:
6103:
6102:
6094:
6087:
6080:
6072:
6065:
6058:
6051:
6044:
6037:
6030:
6023:
6016:
6008:
6001:
5993:
5983:
5978:
5975:
5974:
5971:
5970:
5968:
5967:
5960:
5953:
5946:
5939:
5931:
5929:
5923:
5922:
5920:
5919:
5911:
5903:
5901:
5894:
5893:
5891:
5890:
5882:
5874:
5864:
5859:
5853:
5850:Tyrannoraptora
5848:
5845:
5844:
5842:
5841:
5834:
5827:
5820:
5812:
5805:
5797:
5789:
5782:
5774:
5767:
5759:
5753:
5748:
5741:
5740:
5728:
5727:
5724:
5723:
5715:
5696:
5690:
5688:
5685:
5684:
5681:
5680:
5677:
5676:
5674:
5673:
5672:
5671:
5665:
5663:
5657:
5656:
5653:
5652:
5650:
5649:
5642:
5635:
5628:
5621:
5614:
5607:
5599:
5597:
5595:Megaraptoridae
5591:
5590:
5588:
5587:
5579:
5571:
5564:
5557:
5550:
5542:
5534:
5526:
5520:
5509:
5508:
5505:
5504:
5501:
5500:
5497:
5496:
5493:
5492:
5489:
5488:
5485:
5484:
5482:
5481:
5474:
5467:
5460:
5457:Giganotosaurus
5452:
5450:
5444:
5443:
5441:
5440:
5432:
5426:
5420:
5419:
5417:
5416:
5409:
5402:
5395:
5388:
5381:
5374:
5367:
5360:
5352:
5344:
5338:
5332:
5331:
5329:
5328:
5320:
5312:
5305:
5297:
5288:
5286:
5284:Neovenatoridae
5280:
5279:
5277:
5276:
5269:
5266:Ulughbegsaurus
5262:
5254:
5247:
5240:
5232:
5226:
5220:
5219:
5217:
5216:
5208:
5200:
5192:
5184:
5182:
5173:
5168:
5165:
5164:
5161:
5160:
5157:
5156:
5154:
5153:
5146:
5139:
5132:
5125:
5117:
5115:
5109:
5108:
5106:
5105:
5098:
5094:Xuanhanosaurus
5089:
5083:
5077:
5076:
5074:
5073:
5065:
5061:Xuanhanosaurus
5057:
5050:
5047:Asfaltovenator
5042:
5032:
5027:
5024:
5023:
5021:
5020:
5012:
5008:Megalosauridae
5004:
4996:
4988:
4982:
4977:
4974:
4973:
4971:
4970:
4962:
4953:
4947:
4942:
4935:
4934:
4922:
4921:
4918:
4917:
4909:
4895:
4894:
4892:
4889:
4888:
4885:
4884:
4881:
4880:
4877:
4876:
4874:
4873:
4872:
4871:
4865:
4863:
4853:
4852:
4849:
4848:
4845:
4844:
4841:
4840:
4837:
4836:
4834:
4833:
4826:
4818:
4816:
4810:
4809:
4807:
4806:
4799:
4792:
4785:
4782:Ichthyovenator
4778:
4769:
4763:
4757:
4756:
4753:
4752:
4750:
4749:
4742:
4735:
4727:
4725:
4719:
4718:
4716:
4715:
4707:
4699:
4692:
4685:
4677:
4671:
4665:
4664:
4662:
4661:
4653:
4646:
4643:Cristatusaurus
4639:
4630:
4624:
4619:
4616:
4615:
4612:
4611:
4608:
4607:
4605:
4604:
4600:Poekilopleuron
4596:
4593:Piveteausaurus
4589:
4582:
4575:
4568:
4560:
4558:
4552:
4551:
4549:
4548:
4541:
4534:
4527:
4519:
4517:
4515:Megalosaurinae
4511:
4510:
4508:
4507:
4499:
4493:
4491:Megalosauridae
4487:
4486:
4484:
4483:
4474:
4464:
4459:
4456:
4455:
4452:
4451:
4449:
4448:
4444:Xuanhanosaurus
4440:
4433:
4426:
4418:
4416:
4410:
4409:
4407:
4406:
4403:Yunyangosaurus
4398:
4388:
4383:
4377:
4372:
4369:
4368:
4366:
4365:
4358:
4351:
4343:
4340:Pandoravenator
4336:
4329:
4326:Kayentavenator
4322:
4315:
4308:
4300:
4293:
4285:
4277:
4268:
4262:
4257:
4250:
4249:
4237:
4236:
4233:
4232:
4224:
4217:
4210:
4203:
4202:
4200:
4197:
4196:
4193:
4192:
4189:
4188:
4186:
4185:
4184:
4183:
4177:
4175:
4165:
4164:
4161:
4160:
4157:
4156:
4153:
4152:
4149:
4148:
4145:
4144:
4142:
4141:
4134:
4127:
4120:
4113:
4105:
4098:
4091:
4084:
4076:
4074:
4068:
4067:
4065:
4064:
4057:
4054:Skorpiovenator
4050:
4042:
4035:
4028:
4021:
4014:
4007:
4000:
3991:
3985:
3976:
3975:
3966:
3964:
3958:
3957:
3955:
3954:
3947:
3939:
3932:
3925:
3918:
3910:
3902:
3900:
3894:
3893:
3891:
3890:
3883:
3876:
3869:
3866:Spectrovenator
3862:
3855:
3848:
3841:
3834:
3827:
3819:
3811:
3805:
3800:
3797:
3796:
3793:
3792:
3789:
3788:
3786:
3785:
3778:
3771:
3764:
3757:
3749:
3747:
3741:
3740:
3738:
3737:
3730:
3723:
3715:
3713:
3707:
3706:
3704:
3703:
3700:Spinostropheus
3696:
3689:
3682:
3674:
3666:
3658:
3650:
3643:
3635:
3626:
3620:
3614:
3613:
3611:
3610:
3602:
3594:
3586:
3577:
3567:
3564:Abelisauroidea
3562:
3559:
3558:
3555:
3554:
3552:
3551:
3544:
3536:
3534:
3532:Ceratosauridae
3528:
3527:
3525:
3524:
3521:Saltriovenator
3517:
3509:
3502:
3498:Fosterovenator
3494:
3486:
3476:
3471:
3468:
3467:
3465:
3464:
3460:Lophostropheus
3456:
3448:
3442:
3437:
3430:
3429:
3417:
3416:
3413:
3412:
3399:
3398:
3396:
3393:
3392:
3389:
3388:
3385:
3384:
3382:
3381:
3374:
3366:
3364:
3360:
3359:
3357:
3356:
3355:
3354:
3348:
3346:
3340:
3339:
3336:
3335:
3333:
3332:
3325:
3321:Pterospondylus
3317:
3310:
3302:
3294:
3291:Lucianovenator
3287:
3280:
3272:
3270:
3264:
3263:
3261:
3260:
3253:
3246:
3239:
3232:
3223:
3217:
3215:Coelophysoidea
3211:
3210:
3208:
3207:
3200:
3192:
3185:
3178:
3171:
3164:
3157:
3154:Lophostropheus
3150:
3143:
3136:
3129:
3122:
3114:
3106:
3100:
3095:
3092:
3091:
3089:
3088:
3080:
3072:
3064:
3056:
3053:Erythrovenator
3049:
3041:
3033:
3024:
3018:
3013:
3006:
3005:
2993:
2992:
2989:
2988:
2985:
2984:
2981:
2980:
2978:
2977:
2976:
2975:
2969:
2967:
2961:
2960:
2958:
2957:
2956:
2955:
2946:
2940:
2930:
2929:
2925:
2924:
2915:
2906:
2900:
2894:
2878:
2877:
2870:
2869:
2862:
2855:
2847:
2841:
2840:
2822:
2809:
2806:
2804:
2803:
2793:
2786:
2768:
2740:
2713:
2658:
2606:
2599:
2573:
2560:
2536:
2519:
2502:
2478:
2446:
2424:
2407:
2394:
2381:
2371:
2354:
2327:
2311:
2294:
2281:
2264:
2251:
2231:
2214:
2197:
2184:
2167:
2139:
2111:
2095:
2080:
2064:
2047:
2044:on 2020-08-06.
2027:"Ornithomimus"
2018:
1990:
1988:
1985:
1984:
1983:
1976:
1973:
1913:
1910:
1906:
1905:
1902:
1901:
1897:
1896:
1893:
1892:
1888:
1887:
1884:
1883:
1879:
1878:
1875:
1874:
1870:
1869:
1866:
1865:
1861:
1860:
1857:
1856:
1847:
1844:
1843:
1840:
1839:
1830:
1825:
1823:
1818:
1815:
1814:
1811:
1810:
1801:
1796:
1794:
1789:
1786:
1785:
1782:
1781:
1777:
1776:
1773:
1772:
1763:
1760:
1759:
1756:
1755:
1746:
1741:
1739:
1734:
1732:
1727:
1724:
1723:
1720:
1719:
1710:
1705:
1703:
1698:
1695:
1694:
1691:
1690:
1681:
1676:
1674:
1672:Ornithomimidae
1666:
1610:
1609:Classification
1607:
1515:
1512:
1401:
1398:
1315:Dromiceiomimus
1258:Dromiceiomimus
1246:Dromiceiomimus
1238:Dromiceiomimus
1214:taphonomically
1201:
1198:
1183:Dromiceiomimus
1069:junior synonym
1057:John R. Horner
1014:
1008:
1000:Dromiceiomimus
998:into a second
967:Dromiceiomimus
823:
820:
810:Also in 1933,
805:Inner Mongolia
677:Lawrence Lambe
626:tyrannosauroid
529:
526:
524:
521:
506:Dromiceiomimus
335:
334:
333:
332:
317:
297:
296:
295:Other species
292:
291:
288:
276:
275:
269:
268:
251:
247:
246:
244:Ornithomimidae
238:
234:
233:
225:
218:
217:
212:
205:
204:
199:
192:
191:
186:
179:
178:
173:
169:
168:
163:
159:
158:
153:
149:
148:
135:
134:
123:
122:
114:
113:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
28:
15:
9:
6:
4:
3:
2:
7267:
7256:
7253:
7251:
7248:
7246:
7243:
7241:
7238:
7236:
7233:
7231:
7228:
7226:
7223:
7221:
7218:
7216:
7213:
7211:
7208:
7206:
7203:
7201:
7198:
7196:
7193:
7191:
7188:
7186:
7183:
7181:
7178:
7176:
7175:Ornithomimids
7173:
7172:
7170:
7153:
7148:
7144:
7140:
7135:
7131:
7127:
7122:
7118:
7114:
7109:
7105:
7100:
7094:
7090:
7085:
7079:
7075:
7074:
7072:
7070:
7066:
7062:
7057:
7042:
7038:
7036:
7029:
7000:
6996:
6994:
6990:
6989:
6988:
6987:
6985:
6983:
6979:
6965:
6964:
6960:
6958:
6957:
6956:Struthiomimus
6953:
6951:
6950:
6946:
6944:
6943:
6939:
6937:
6936:
6932:
6930:
6929:
6925:
6923:
6922:
6918:
6916:
6915:
6911:
6909:
6908:
6907:"Coelosaurus"
6904:
6902:
6901:
6897:
6895:
6894:
6890:
6888:
6887:
6883:
6882:
6880:
6878:
6874:
6868:
6867:
6863:
6860:
6859:
6855:
6852:
6851:
6847:
6845:
6844:
6840:
6838:
6837:
6833:
6831:
6830:
6826:
6825:
6823:
6821:
6817:
6811:
6810:
6806:
6804:
6803:
6802:Pelecanimimus
6799:
6797:
6796:
6792:
6791:
6788:
6785:
6783:
6779:
6773:
6772:
6768:
6765:
6764:
6763:Thecocoelurus
6760:
6758:
6757:
6753:
6751:
6750:
6746:
6744:
6743:
6742:Kinnareemimus
6739:
6737:
6736:
6732:
6729:
6728:
6724:
6721:
6720:
6716:
6714:
6713:
6709:
6708:
6705:
6702:
6700:
6696:
6689:
6688:
6684:
6683:
6680:
6677:
6674:
6669:
6659:
6658:
6654:
6652:
6651:
6647:
6645:
6644:
6640:
6637:
6636:
6632:
6630:
6629:
6625:
6623:
6622:
6621:Huaxiagnathus
6618:
6616:
6615:
6614:Compsognathus
6611:
6608:
6607:
6603:
6601:
6600:
6596:
6593:
6592:
6588:
6587:
6585:
6583:
6579:
6573:
6572:
6568:
6567:
6564:
6560:
6557:
6554:
6549:
6543:
6542:
6541:Ornitholestes
6538:
6536:
6535:
6531:
6528:
6527:
6523:
6522:
6519:
6516:
6513:
6508:
6504:
6500:
6495:
6491:
6481:
6476:
6474:
6467:
6461:
6439:
6438:
6434:
6432:
6431:
6427:
6425:
6424:
6420:
6418:
6417:
6413:
6412:
6410:
6406:
6402:
6385:
6384:
6383:
6382:
6380:
6378:
6374:
6370:
6344:
6343:
6339:
6337:
6336:
6335:Tyrannosaurus
6332:
6330:
6329:
6325:
6324:
6322:
6320:
6316:
6310:
6309:
6308:Teratophoneus
6305:
6303:
6302:
6298:
6295:
6294:
6290:
6288:
6287:
6283:
6282:
6280:
6278:
6274:
6268:
6267:
6263:
6261:
6260:
6256:
6255:
6253:
6251:
6247:
6241:
6240:
6236:
6234:
6233:
6229:
6228:
6226:
6224:
6220:
6214:
6213:
6209:
6207:
6206:
6202:
6201:
6198:
6195:
6193:
6189:
6183:
6182:
6178:
6176:
6175:
6174:Albertosaurus
6171:
6170:
6168:
6166:
6162:
6159:
6156:
6151:
6145:
6144:
6140:
6138:
6137:
6133:
6131:
6130:
6126:
6125:
6122:
6119:
6116:
6111:
6100:
6099:
6095:
6093:
6092:
6091:Xiongguanlong
6088:
6086:
6085:
6081:
6078:
6077:
6073:
6071:
6070:
6069:Suskityrannus
6066:
6064:
6063:
6062:Stokesosaurus
6059:
6057:
6056:
6055:Santanaraptor
6052:
6050:
6049:
6045:
6043:
6042:
6038:
6036:
6035:
6031:
6029:
6028:
6024:
6022:
6021:
6017:
6014:
6013:
6009:
6007:
6006:
6002:
6000:
5999:
5998:Alectrosaurus
5995:
5994:
5991:
5987:
5984:
5981:
5976:
5966:
5965:
5961:
5959:
5958:
5954:
5952:
5951:
5947:
5945:
5944:
5940:
5938:
5937:
5933:
5932:
5930:
5928:
5924:
5917:
5916:
5915:Tanycolagreus
5912:
5910:
5909:
5905:
5904:
5902:
5899:
5895:
5888:
5887:
5883:
5881:
5880:
5876:
5875:
5872:
5868:
5865:
5862:
5857:
5854:
5851:
5846:
5840:
5839:
5835:
5833:
5832:
5828:
5826:
5825:
5821:
5818:
5817:
5813:
5811:
5810:
5806:
5803:
5802:
5798:
5795:
5794:
5790:
5788:
5787:
5783:
5780:
5779:
5778:Bahariasaurus
5775:
5773:
5772:
5768:
5766:
5765:
5761:
5760:
5757:
5754:
5751:
5750:Coelurosauria
5746:
5742:
5738:
5737:Coelurosauria
5733:
5729:
5719:
5714:
5712:
5707:
5700:
5694:
5669:
5668:
5667:
5666:
5664:
5662:
5661:Coelurosauria
5658:
5648:
5647:
5643:
5641:
5640:
5636:
5634:
5633:
5629:
5627:
5626:
5622:
5620:
5619:
5615:
5613:
5612:
5608:
5606:
5605:
5601:
5600:
5598:
5596:
5592:
5586:
5584:
5580:
5578:
5576:
5572:
5570:
5569:
5565:
5563:
5562:
5558:
5556:
5555:
5551:
5548:
5547:
5543:
5540:
5539:
5538:Bahariasaurus
5535:
5533:
5532:
5528:
5527:
5524:
5521:
5518:
5514:
5510:
5480:
5479:
5475:
5473:
5472:
5468:
5466:
5465:
5461:
5459:
5458:
5454:
5453:
5451:
5449:
5445:
5439:
5438:
5434:
5433:
5430:
5427:
5425:
5421:
5415:
5414:
5410:
5408:
5407:
5403:
5401:
5400:
5396:
5394:
5393:
5389:
5387:
5386:
5382:
5380:
5379:
5378:Kelmayisaurus
5375:
5373:
5372:
5368:
5366:
5365:
5361:
5358:
5357:
5353:
5351:
5350:
5346:
5345:
5342:
5339:
5337:
5333:
5326:
5325:
5321:
5318:
5317:
5313:
5311:
5310:
5306:
5303:
5302:
5298:
5295:
5294:
5290:
5289:
5287:
5285:
5281:
5275:
5274:
5270:
5268:
5267:
5263:
5260:
5259:
5255:
5253:
5252:
5248:
5246:
5245:
5241:
5239:
5238:
5234:
5233:
5230:
5227:
5225:
5221:
5214:
5213:
5212:Saurophaganax
5209:
5206:
5205:
5201:
5198:
5197:
5193:
5191:
5190:
5186:
5185:
5183:
5181:
5177:
5174:
5171:
5166:
5152:
5151:
5147:
5145:
5144:
5143:Siamotyrannus
5140:
5138:
5137:
5133:
5131:
5130:
5126:
5124:
5123:
5119:
5118:
5116:
5114:
5110:
5104:
5103:
5099:
5096:
5095:
5091:
5090:
5087:
5084:
5082:
5078:
5071:
5070:
5066:
5063:
5062:
5058:
5056:
5055:
5051:
5049:
5048:
5044:
5043:
5040:
5036:
5033:
5030:
5029:Allosauroidea
5025:
5018:
5017:
5016:Spinosauridae
5013:
5010:
5009:
5005:
5002:
5001:
4997:
4995:
4994:
4990:
4989:
4986:
4983:
4980:
4975:
4968:
4967:
4963:
4960:
4959:
4955:
4954:
4951:
4948:
4945:
4940:
4936:
4932:
4927:
4923:
4913:
4908:
4906:
4899:
4869:
4868:
4867:
4866:
4864:
4862:
4858:
4854:
4832:
4831:
4827:
4825:
4824:
4820:
4819:
4817:
4815:
4811:
4805:
4804:
4800:
4798:
4797:
4793:
4791:
4790:
4786:
4784:
4783:
4779:
4776:
4775:
4771:
4770:
4767:
4764:
4762:
4761:Spinosaurinae
4758:
4748:
4747:
4743:
4741:
4740:
4739:Riparovenator
4736:
4734:
4733:
4732:Ceratosuchops
4729:
4728:
4726:
4724:
4720:
4713:
4712:
4708:
4705:
4704:
4700:
4698:
4697:
4696:Riojavenatrix
4693:
4691:
4690:
4686:
4684:
4683:
4679:
4678:
4675:
4672:
4670:
4669:Baryonychinae
4666:
4659:
4658:
4654:
4652:
4651:
4647:
4645:
4644:
4640:
4637:
4636:
4632:
4631:
4628:
4625:
4622:
4621:Spinosauridae
4617:
4602:
4601:
4597:
4595:
4594:
4590:
4588:
4587:
4583:
4581:
4580:
4576:
4574:
4573:
4569:
4567:
4566:
4562:
4561:
4559:
4557:
4553:
4547:
4546:
4545:Wiehenvenator
4542:
4540:
4539:
4535:
4533:
4532:
4528:
4526:
4525:
4521:
4520:
4518:
4516:
4512:
4506:
4505:
4501:
4500:
4497:
4494:
4492:
4488:
4481:
4480:
4476:
4475:
4472:
4468:
4465:
4462:
4457:
4446:
4445:
4441:
4439:
4438:
4434:
4432:
4431:
4427:
4425:
4424:
4420:
4419:
4417:
4415:
4411:
4405:
4404:
4400:
4399:
4396:
4392:
4389:
4386:
4381:
4378:
4375:
4370:
4364:
4363:
4359:
4357:
4356:
4352:
4349:
4348:
4344:
4342:
4341:
4337:
4335:
4334:
4330:
4328:
4327:
4323:
4321:
4320:
4316:
4314:
4313:
4309:
4306:
4305:
4301:
4299:
4298:
4294:
4291:
4290:
4286:
4283:
4282:
4281:Chienkosaurus
4278:
4275:
4274:
4270:
4269:
4266:
4263:
4260:
4255:
4251:
4247:
4242:
4238:
4228:
4221:
4214:
4207:
4181:
4180:
4179:
4178:
4176:
4174:
4170:
4166:
4140:
4139:
4135:
4133:
4132:
4128:
4126:
4125:
4121:
4119:
4118:
4114:
4111:
4110:
4106:
4104:
4103:
4099:
4097:
4096:
4092:
4090:
4089:
4085:
4083:
4082:
4078:
4077:
4075:
4073:
4072:Furileusauria
4069:
4063:
4062:
4058:
4056:
4055:
4051:
4048:
4047:
4046:Rahiolisaurus
4043:
4041:
4040:
4036:
4034:
4033:
4029:
4027:
4026:
4022:
4020:
4019:
4015:
4013:
4012:
4008:
4006:
4005:
4001:
3998:
3997:
3993:
3992:
3989:
3986:
3984:
3980:
3973:
3972:
3968:
3967:
3965:
3963:
3962:Carnotaurinae
3959:
3953:
3952:
3948:
3945:
3944:
3943:Rahiolisaurus
3940:
3938:
3937:
3936:Majungasaurus
3933:
3931:
3930:
3926:
3924:
3923:
3919:
3916:
3915:
3911:
3909:
3908:
3904:
3903:
3901:
3899:
3895:
3889:
3888:
3884:
3882:
3881:
3877:
3875:
3874:
3873:Tarascosaurus
3870:
3868:
3867:
3863:
3861:
3860:
3856:
3854:
3853:
3849:
3847:
3846:
3842:
3840:
3839:
3835:
3833:
3832:
3831:Eoabelisaurus
3828:
3825:
3824:
3820:
3818:
3817:
3816:Chenanisaurus
3813:
3812:
3809:
3806:
3803:
3802:Abelisauridae
3798:
3784:
3783:
3779:
3777:
3776:
3772:
3770:
3769:
3765:
3763:
3762:
3761:Masiakasaurus
3758:
3756:
3755:
3751:
3750:
3748:
3746:
3742:
3736:
3735:
3731:
3729:
3728:
3727:Huinculsaurus
3724:
3722:
3721:
3720:Elaphrosaurus
3717:
3716:
3714:
3712:
3708:
3702:
3701:
3697:
3695:
3694:
3690:
3688:
3687:
3683:
3680:
3679:
3675:
3672:
3671:
3667:
3664:
3663:
3659:
3656:
3655:
3651:
3649:
3648:
3644:
3641:
3640:
3639:Austrocheirus
3636:
3633:
3632:
3628:
3627:
3624:
3621:
3619:
3615:
3608:
3607:
3603:
3600:
3599:
3595:
3592:
3591:
3587:
3584:
3583:
3579:
3578:
3575:
3571:
3568:
3565:
3560:
3550:
3549:
3545:
3543:
3542:
3538:
3537:
3535:
3533:
3529:
3523:
3522:
3518:
3515:
3514:
3510:
3508:
3507:
3503:
3500:
3499:
3495:
3493:
3492:
3491:Berberosaurus
3488:
3487:
3484:
3480:
3477:
3474:
3469:
3462:
3461:
3457:
3455:
3454:
3450:
3449:
3446:
3443:
3440:
3435:
3431:
3427:
3422:
3418:
3409:
3403:
3380:
3379:
3378:Halticosaurus
3375:
3373:
3372:
3371:Teinurosaurus
3368:
3367:
3365:
3361:
3352:
3351:
3350:
3349:
3347:
3345:
3341:
3331:
3330:
3326:
3323:
3322:
3318:
3316:
3315:
3311:
3308:
3307:
3303:
3300:
3299:
3298:Megapnosaurus
3295:
3293:
3292:
3288:
3286:
3285:
3281:
3279:
3278:
3274:
3273:
3271:
3269:
3268:Coelophysidae
3265:
3259:
3258:
3257:Powellvenator
3254:
3252:
3251:
3247:
3245:
3244:
3240:
3238:
3237:
3233:
3230:
3229:
3225:
3224:
3221:
3218:
3216:
3212:
3206:
3205:
3201:
3198:
3197:
3193:
3191:
3190:
3186:
3184:
3183:
3179:
3177:
3176:
3172:
3170:
3169:
3165:
3163:
3162:
3158:
3156:
3155:
3151:
3149:
3148:
3147:Liliensternus
3144:
3142:
3141:
3137:
3135:
3134:
3130:
3128:
3127:
3123:
3120:
3119:
3118:Dolichosuchus
3115:
3113:
3112:
3111:Dilophosaurus
3108:
3107:
3104:
3101:
3098:
3093:
3086:
3085:
3084:Herrerasauria
3081:
3078:
3077:
3073:
3070:
3069:
3065:
3062:
3061:
3057:
3055:
3054:
3050:
3047:
3046:
3042:
3039:
3038:
3037:Daemonosaurus
3034:
3031:
3030:
3026:
3025:
3022:
3019:
3016:
3011:
3007:
3003:
2998:
2994:
2973:
2972:
2971:
2970:
2968:
2966:
2962:
2954:
2950:
2949:
2948:
2947:
2944:
2941:
2939:
2935:
2931:
2923:
2919:
2916:
2914:
2910:
2907:
2905:
2901:
2899:
2895:
2893:
2889:
2888:
2883:
2879:
2875:
2868:
2863:
2861:
2856:
2854:
2849:
2848:
2845:
2839:
2835:
2831:
2827:
2823:
2820:
2816:
2812:
2811:
2797:
2789:
2783:
2779:
2772:
2764:
2760:
2756:
2752:
2744:
2736:
2732:
2728:
2724:
2717:
2709:
2705:
2701:
2697:
2693:
2689:
2685:
2681:
2677:
2673:
2669:
2668:Archaeopteryx
2662:
2654:
2650:
2646:
2642:
2638:
2634:
2630:
2626:
2622:
2618:
2610:
2602:
2596:
2592:
2587:
2586:
2577:
2570:
2564:
2557:
2556:0-7853-0443-6
2553:
2549:
2543:
2541:
2533:
2529:
2523:
2517:(4): 691-710.
2516:
2512:
2506:
2499:
2495:
2491:
2485:
2483:
2476:
2472:
2468:
2464:
2460:
2459:Kosmoceratops
2456:
2455:Pentaceratops
2450:
2444:(4): 983-997.
2443:
2439:
2438:Palaeontology
2433:
2431:
2429:
2421:
2417:
2411:
2404:
2398:
2391:
2385:
2375:
2368:
2364:
2358:
2351:
2347:
2343:
2340:
2334:
2332:
2324:
2321:
2315:
2308:
2304:
2298:
2291:
2285:
2278:
2274:
2268:
2261:
2255:
2248:
2245:
2241:
2235:
2228:
2224:
2218:
2211:
2207:
2201:
2194:
2188:
2181:
2177:
2171:
2164:
2161:
2157:
2153:
2149:
2143:
2137:(43): 733-771
2136:
2133:
2129:
2128:Tyrannosaurus
2125:
2124:Struthiomimus
2121:
2120:Ornitholestes
2115:
2108:
2105:
2099:
2093:
2090:
2084:
2077:
2074:
2068:
2061:
2057:
2051:
2043:
2039:
2035:
2033:
2028:
2022:
2016:
2012:
2008:
2004:
1998:
1996:
1991:
1982:
1979:
1978:
1972:
1970:
1966:
1962:
1957:
1955:
1951:
1947:
1943:
1939:
1936:
1929:
1928:
1923:
1918:
1904:
1903:
1895:
1894:
1886:
1885:
1877:
1876:
1868:
1867:
1859:
1858:
1855:
1854:
1853:
1846:
1845:
1842:
1841:
1838:
1837:
1836:
1829:
1828:
1817:
1816:
1813:
1812:
1809:
1808:
1807:
1800:
1799:
1788:
1787:
1784:
1783:
1775:
1774:
1771:
1770:
1769:
1762:
1761:
1758:
1757:
1754:
1753:
1752:
1745:
1744:
1738:
1737:
1726:
1725:
1722:
1721:
1718:
1717:
1716:
1709:
1708:
1697:
1696:
1693:
1692:
1689:
1688:
1687:
1680:
1679:
1673:
1670:
1669:
1665:
1663:
1659:
1654:
1652:
1648:
1644:
1640:
1636:
1632:
1629:to the clade
1628:
1624:
1615:
1606:
1604:
1600:
1596:
1592:
1588:
1583:
1581:
1577:
1573:
1569:
1565:
1561:
1556:
1552:
1548:
1544:
1540:
1536:
1532:
1528:
1520:
1511:
1509:
1505:
1501:
1497:
1493:
1489:
1485:
1481:
1477:
1473:
1469:
1460:
1456:
1454:
1453:
1452:Struthiomimus
1448:
1444:
1442:
1438:
1434:
1430:
1426:
1423:, species of
1422:
1421:ornithomimids
1415:
1411:
1406:
1397:
1395:
1391:
1387:
1383:
1379:
1375:
1371:
1367:
1363:
1359:
1355:
1351:
1347:
1342:
1340:
1336:
1332:
1328:
1324:
1320:
1316:
1312:
1311:Struthiomimus
1308:
1304:
1300:
1296:
1295:
1290:
1285:
1283:
1279:
1275:
1271:
1267:
1263:
1259:
1255:
1251:
1247:
1243:
1239:
1235:
1231:
1227:
1223:
1215:
1211:
1206:
1197:
1195:
1192:to the genus
1191:
1190:
1185:
1184:
1179:
1178:
1177:Struthiomimus
1173:
1172:
1167:
1163:
1158:
1156:
1152:
1149:differs from
1148:
1144:
1140:
1139:
1134:
1130:
1126:
1122:
1118:
1114:
1110:
1106:
1102:
1098:
1094:
1090:
1086:
1082:
1078:
1074:
1070:
1066:
1062:
1058:
1054:
1050:
1046:
1042:
1038:
1034:
1033:
1028:
1024:
1020:
1013:
1007:
1005:
1001:
997:
993:
989:
985:
981:
977:
973:
969:
968:
963:
959:
955:
951:
948:
946:
941:
940:Struthiomimus
937:
933:
929:
925:
921:
917:
913:
909:
908:Struthiomimus
905:
901:
897:
896:Struthiomimus
893:
889:
885:
881:
877:
873:
869:
865:
861:
857:
853:
849:
845:
841:
833:
828:
819:
817:
813:
808:
806:
802:
798:
797:
792:
791:
790:Chirostenotes
786:
782:
778:
774:
773:
768:
764:
760:
756:
752:
748:
744:
740:
736:
732:
728:
727:Loris Russell
724:
720:
716:
712:
708:
704:
700:
696:
695:
694:Struthiomimus
690:
686:
682:
678:
674:
670:
665:
663:
662:alvarezsaurid
659:
655:
654:
649:
645:
644:
639:
635:
631:
627:
623:
619:
615:
611:
607:
603:
602:specific name
599:
595:
591:
587:
583:
579:
575:
571:
567:
563:
559:
552:
551:type specimen
549:
545:
539:
534:
520:
518:
514:
513:
512:Struthiomimus
508:
507:
502:
501:
496:
492:
488:
484:
480:
476:
472:
468:
464:
461:was named in
460:
456:
455:
450:
449:
444:
440:
436:
432:
428:
425:
421:
420:Maastrichtian
417:
413:
410:
407:
403:
397:
343:
342:
330:(Parks, 1928)
326:
325:
318:
314:
309:
308:
301:
300:
298:
293:
286:
285:
277:
274:
270:
265:
260:
259:
252:
249:
248:
245:
239:
236:
235:
232:
226:
223:
220:
219:
216:
213:
210:
207:
206:
203:
200:
197:
194:
193:
190:
187:
184:
181:
180:
177:
174:
171:
170:
167:
164:
161:
160:
157:
154:
151:
150:
145:
140:
136:
133:
129:
124:
120:
115:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
47:
44:76.5–66
40:
39:Maastrichtian
36:
32:
26:
22:
19:
7099:Ornithomimus
7069:Ornithomimus
7068:
7032:
6961:
6954:
6947:
6940:
6933:
6928:Ornithomimus
6927:
6926:
6919:
6912:
6905:
6898:
6891:
6884:
6866:Tyrannomimus
6864:
6856:
6848:
6841:
6836:Deinocheirus
6834:
6827:
6807:
6800:
6793:
6769:
6761:
6756:Nqwebasaurus
6754:
6749:Nedcolbertia
6747:
6740:
6733:
6727:Calamosaurus
6725:
6719:Aviatyrannis
6717:
6710:
6685:
6655:
6648:
6641:
6633:
6626:
6619:
6612:
6604:
6599:Aristosuchus
6597:
6591:Aniksosaurus
6589:
6571:Aniksosaurus
6569:
6539:
6532:
6524:
6470:
6435:
6428:
6421:
6414:
6340:
6333:
6326:
6306:
6299:
6291:
6286:Dynamoterror
6284:
6264:
6257:
6237:
6230:
6210:
6205:Asiatyrannus
6203:
6179:
6172:
6143:Dryptosaurus
6141:
6134:
6127:
6096:
6089:
6082:
6074:
6067:
6060:
6053:
6046:
6039:
6034:Jinbeisaurus
6032:
6025:
6018:
6010:
6005:Aviatyrannis
6003:
5996:
5962:
5957:Sinotyrannus
5955:
5948:
5941:
5934:
5913:
5906:
5884:
5877:
5836:
5829:
5822:
5814:
5807:
5799:
5791:
5786:Bicentenaria
5784:
5776:
5771:Asiamericana
5769:
5762:
5703:
5644:
5637:
5630:
5623:
5616:
5609:
5602:
5581:
5573:
5566:
5559:
5552:
5546:Deltadromeus
5544:
5536:
5529:
5478:Tyrannotitan
5476:
5469:
5462:
5455:
5435:
5411:
5406:Taurovenator
5404:
5397:
5390:
5385:Lajasvenator
5383:
5376:
5369:
5364:Concavenator
5362:
5354:
5347:
5322:
5314:
5307:
5299:
5291:
5271:
5264:
5256:
5249:
5242:
5235:
5210:
5202:
5194:
5187:
5180:Allosauridae
5148:
5141:
5136:Shidaisaurus
5134:
5127:
5120:
5100:
5092:
5067:
5059:
5052:
5045:
5014:
5006:
4998:
4991:
4964:
4956:
4944:Avetheropoda
4931:Avetheropoda
4902:
4861:Avetheropoda
4828:
4821:
4814:Spinosaurini
4801:
4794:
4787:
4780:
4772:
4744:
4737:
4730:
4709:
4701:
4694:
4689:Protathlitis
4687:
4680:
4655:
4648:
4641:
4633:
4598:
4591:
4584:
4579:Leshansaurus
4577:
4570:
4563:
4543:
4536:
4531:Megalosaurus
4529:
4524:Duriavenator
4522:
4502:
4477:
4461:Megalosauria
4442:
4435:
4430:Marshosaurus
4428:
4423:Condorraptor
4421:
4401:
4360:
4353:
4345:
4338:
4331:
4324:
4317:
4310:
4304:Cruxicheiros
4302:
4295:
4287:
4279:
4271:
4136:
4131:Quilmesaurus
4129:
4122:
4115:
4107:
4100:
4093:
4086:
4079:
4059:
4052:
4044:
4037:
4030:
4023:
4016:
4009:
4002:
3994:
3983:Brachyrostra
3969:
3949:
3941:
3934:
3927:
3920:
3912:
3905:
3885:
3880:Tralkasaurus
3878:
3871:
3864:
3857:
3850:
3843:
3836:
3829:
3821:
3814:
3782:Vespersaurus
3780:
3775:Velocisaurus
3773:
3766:
3759:
3752:
3732:
3725:
3718:
3698:
3691:
3684:
3676:
3670:Deltadromeus
3668:
3660:
3654:Compsosuchus
3652:
3645:
3637:
3629:
3604:
3596:
3590:Lametasaurus
3588:
3580:
3546:
3541:Ceratosaurus
3539:
3519:
3511:
3504:
3496:
3489:
3473:Ceratosauria
3458:
3451:
3376:
3369:
3327:
3319:
3312:
3304:
3296:
3289:
3282:
3275:
3255:
3250:Podokesaurus
3248:
3241:
3234:
3226:
3202:
3194:
3187:
3180:
3173:
3166:
3159:
3152:
3145:
3138:
3133:Gojirasaurus
3131:
3126:Dracovenator
3124:
3116:
3109:
3097:Neotheropoda
3082:
3074:
3066:
3060:Guaibasaurus
3058:
3051:
3043:
3035:
3029:Chindesaurus
3027:
2917:
2908:
2829:
2825:
2818:
2815:Ornithomimus
2814:
2796:
2777:
2771:
2754:
2750:
2743:
2726:
2722:
2716:
2675:
2671:
2667:
2661:
2620:
2616:
2609:
2584:
2576:
2568:
2563:
2547:
2531:
2522:
2514:
2510:
2505:
2497:
2493:
2489:
2466:
2462:
2458:
2454:
2449:
2441:
2437:
2419:
2415:
2410:
2402:
2397:
2389:
2384:
2374:
2366:
2362:
2357:
2349:
2322:
2319:
2314:
2306:
2302:
2297:
2289:
2284:
2276:
2272:
2267:
2259:
2254:
2246:
2243:
2240:Ornithomimus
2239:
2234:
2226:
2222:
2217:
2209:
2205:
2200:
2192:
2187:
2182:(4): 133-159
2179:
2175:
2170:
2162:
2159:
2156:Ceratosaurus
2155:
2151:
2147:
2142:
2134:
2131:
2127:
2123:
2119:
2114:
2106:
2103:
2098:
2088:
2083:
2075:
2072:
2067:
2059:
2055:
2050:
2042:the original
2030:
2021:
2006:
2002:
1965:Ornithomimus
1964:
1958:
1942:Ornithomimus
1941:
1940:
1935:Ornithomimus
1934:
1933:The diet of
1932:
1927:Dakotaraptor
1925:
1922:Ornithomimus
1921:
1912:Paleobiology
1851:
1850:
1849:
1833:
1832:
1804:
1803:
1766:
1765:
1749:
1748:
1713:
1712:
1684:
1683:
1657:
1655:
1650:
1646:
1642:
1638:
1627:Ornithomimus
1626:
1620:
1603:Ornithomimus
1602:
1595:Ornithomimus
1594:
1587:Ornithomimus
1586:
1584:
1576:Ornithomimus
1575:
1567:
1550:
1535:Ornithomimus
1534:
1527:Ornithomimus
1526:
1525:
1507:
1499:
1492:Ornithomimus
1491:
1488:type species
1483:
1475:
1468:Ornithomimus
1467:
1465:
1450:
1447:Ornithomimus
1446:
1445:
1425:Ornithomimus
1424:
1418:
1409:
1408:Specimen of
1393:
1389:
1382:Ornithomimus
1381:
1377:
1361:
1349:
1345:
1343:
1338:
1335:Saskatchewan
1326:
1323:Ornithomimus
1322:
1314:
1310:
1306:
1292:
1288:
1286:
1281:
1277:
1273:
1261:
1257:
1253:
1249:
1245:
1242:Ornithomimus
1241:
1237:
1222:Ornithomimus
1221:
1219:
1209:
1194:Ornithomimus
1193:
1187:
1181:
1175:
1169:
1159:
1155:Ornithomimus
1154:
1150:
1146:
1138:nomen dubium
1136:
1132:
1120:
1116:
1112:
1108:
1096:
1085:Richard Owen
1080:
1072:
1065:Ornithomimus
1064:
1060:
1053:Donald Baird
1045:Ornithomimus
1044:
1037:Joseph Leidy
1030:
1016:
1012:Ornithomimus
1011:
1003:
999:
995:
991:
987:
983:
980:Ornithomimus
979:
975:
965:
961:
957:
953:
949:
943:
939:
936:Ornithomimus
935:
931:
927:
923:
919:
915:
912:Ornithomimus
911:
907:
904:morphometric
902:published a
895:
892:Ornithomimus
891:
887:
883:
879:
875:
868:Dale Russell
864:abelisauroid
859:
847:
840:Ornithomimus
839:
837:
832:Ornithomimus
831:
815:
809:
800:
794:
788:
784:
772:nomen dubium
770:
766:
758:
750:
746:
738:
730:
714:
710:
692:
668:
666:
651:
641:
630:Ornithomimus
629:
613:
609:
597:
593:
588:on June 30,
565:
562:type species
558:Ornithomimus
557:
555:
547:
537:
510:
504:
499:
498:
495:Ornithomimus
494:
486:
478:
458:
453:
452:
447:
446:
443:type species
431:Ornithomimus
430:
406:ornithomimid
341:Ornithomimus
340:
339:
338:
323:
322:
306:
305:
283:
282:
273:Type species
258:Ornithomimus
257:
256:
221:
208:
195:
182:
127:
25:Ornithomimus
24:
18:
7093:Wikispecies
6993:Maniraptora
6982:Maniraptora
6963:Tototlmimus
6893:Anserimimus
6843:Garudimimus
6829:Beishanlong
6771:Valdoraptor
6712:Arkansaurus
6657:Xunmenglong
6534:Juravenator
6328:Tarbosaurus
6212:Nanuqsaurus
6181:Gorgosaurus
6098:Megaraptora
6084:Timurlengia
5816:Sciurumimus
5764:Aratasaurus
5632:Murusraptor
5554:Fukuiraptor
5517:Megaraptora
5399:Shaochilong
5371:Eocarcharia
5324:Megaraptora
5244:Lusovenator
5122:Alpkarakush
4979:Carnosauria
4830:Spinosaurus
4803:Siamosaurus
4703:Suchosaurus
4650:Iberospinus
4586:Magnosaurus
4565:Afrovenator
4538:Torvosaurus
4289:Chilesaurus
4102:Carnotaurus
4095:Caletodraco
4081:Abelisaurus
3907:Arcovenator
3754:Laevisuchus
3745:Noasaurinae
3647:Berthasaura
3618:Noasauridae
3548:Genyodectes
3513:Lukousaurus
3506:Jubbulpuria
3306:Panguraptor
3284:Coelophysis
3277:Camposaurus
3236:Panguraptor
3228:Dracoraptor
3204:Zupaysaurus
3189:Tachiraptor
3168:Sarcosaurus
3068:Nhandumirim
2729:: 108–117.
2262:. 109: 1-94
2058:, series 3
1580:cassowaries
1541:. In 1995,
1437:kinesthetic
1419:Like other
1400:Description
1287:Apart from
1234:Phil Currie
1147:Coelosaurus
1123:. However,
1121:O. antiquus
1117:O. antiquus
1097:O. antiquus
1081:Coelosaurus
1073:Coelosaurus
1029:were named
970:, meaning "
289:Marsh, 1890
7169:Categories
6921:Gallimimus
6850:Harpymimus
6795:Harpymimus
6416:Iliosuchus
6386:see below↓
6223:Alioramini
6041:Juratyrant
6027:Eotyrannus
6012:Bagaraatan
5964:Yutyrannus
5898:Coeluridae
5824:Vayuraptor
5670:see below↓
5646:Tratayenia
5639:Orkoraptor
5625:Megaraptor
5531:Aoniraptor
5464:Mapusaurus
5392:Sauroniops
5356:Altispinax
5309:Neovenator
5251:Siamraptor
5237:Datanglong
5204:Epanterias
5196:Antrodemus
5189:Allosaurus
5170:Allosauria
4993:Altispinax
4958:Gasosaurus
4870:see below↓
4746:Suchomimus
4347:Sinosaurus
4182:see below↓
4138:Viavenator
4088:Aucasaurus
4025:Ilokelesia
3996:Dahalokely
3951:Rajasaurus
3929:Indosaurus
3922:Genusaurus
3914:Dahalokely
3838:Indosuchus
3734:Limusaurus
3693:Ligabueino
3686:Kiyacursor
3678:Genusaurus
3662:Dahalokely
3582:Betasuchus
3453:Dornraptor
3353:see below↓
3329:Segisaurus
3182:Sinosaurus
3045:Eodromaeus
2974:see below↓
2922:Dinosauria
2904:Sauropsida
2757:(2): 213.
2369:. 249-254.
2320:Brimleyana
2309:: 375-402.
2249:(5): 79-83
2152:Allosaurus
2148:Antrodemus
2109:(2): 25-81
1987:References
1504:metacarpal
1341:holotype.
1244:, sinking
1208:T scan of
1189:Gallimimus
1151:Gallimimus
1027:New Jersey
976:Dromiceius
866:claw) and
844:Oskar Kuhn
796:Elmisaurus
658:metatarsus
634:ornithopod
570:O.C. Marsh
497:, such as
439:omnivorous
324:O. samueli
202:Saurischia
189:Dinosauria
130:skeleton,
6997:includes
6942:Rativates
6935:Qiupalong
6635:Scipionyx
6628:Mirischia
6301:Lythronax
6293:Labocania
6232:Alioramus
5886:Labocania
5604:Aerosteon
5150:Sinraptor
5054:Erectopus
4789:Irritator
4374:Orionides
4259:Tetanurae
4246:Tetanurae
4173:Tetanurae
4109:Elemgasem
4032:Llukalkan
4011:Elemgasem
3768:Noasaurus
3631:Afromimus
3439:Averostra
3426:Averostra
3344:Averostra
3196:Velocipes
3015:Theropoda
3002:Theropoda
2965:Theropoda
2890:Kingdom:
2874:Theropoda
2422:(3): 86A.
2078:: 449-453
1954:nocturnal
1651:O. sedens
1635:cladistic
1617:Claw bone
1560:sandstone
1339:O. sedens
1294:O. sedens
1141:. An SVP
1107:regarded
1021:from the
1002:species:
988:O. ingens
416:Campanian
414:from the
412:dinosaurs
313:Sternberg
215:Theropoda
162:Kingdom:
156:Eukaryota
35:Campanian
7078:Wikidata
6526:Coelurus
5943:Kileskus
5936:Guanlong
5908:Coelurus
5801:Gualicho
5301:Gualicho
4682:Baryonyx
4018:Guemesia
3845:Kryptops
3606:Ozraptor
3243:Pendraig
2898:Chordata
2896:Phylum:
2892:Animalia
2819:O. velox
2801:331-336.
2700:24990749
2645:23112330
2342:Archived
1975:See also
1647:O. velox
1549:, three
1539:feathers
1508:O. velox
1496:holotype
1484:O. velox
1466:The two
1390:O. velox
1378:O. velox
1362:O. velox
1350:O. velox
1346:O. velox
1307:O. velox
1254:O. velox
1133:antiquus
1109:O. velox
1093:Maryland
924:O. velox
874:renamed
846:renamed
757:renamed
729:renamed
586:Colorado
574:syntypes
538:O. velox
475:Colorado
459:O. velox
435:feathers
422:ages of
409:theropod
237:Family:
176:Chordata
172:Phylum:
166:Animalia
152:Domain:
126:Mounted
7139:4129013
7126:4822900
7113:4433735
7084:Q131067
6076:Timimus
5838:Zuolong
5568:Rapator
5471:Meraxes
4796:Oxalaia
4117:Koleken
3140:Lepidus
2902:Class:
2708:4464659
2680:Bibcode
2653:2057698
2625:Bibcode
2617:Science
2469:: 292.
2229:: 23-78
2165:: l-154
2062:: 81-86
1820:unnamed
1791:unnamed
1729:unnamed
1700:unnamed
1599:ostrich
1433:ratites
1429:bipedal
1358:Wyoming
1331:Montana
1303:Wyoming
1077:dubious
856:Belgium
854:) from
850:(Dollo
765:) into
761:(Leidy
745:) into
741:(Parks
733:(Parks
685:Alberta
638:Wyoming
632:was an
622:Montana
580:in the
483:Alberta
250:Genus:
6735:Hexing
6423:Kakuru
6020:Dilong
4061:Thanos
4039:Niebla
3859:Rugops
3852:Kurupi
2784:
2706:
2698:
2672:Nature
2651:
2643:
2597:
2554:
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2279:: 1-22
2212:: 1-33
2154:) and
2032:Lexico
1658:et al.
1649:, and
1545:, and
1486:, the
1232:, and
1186:, and
1019:tibiae
737:) and
717:(Lull
709:named
315:, 1933
266:, 1890
7152:38544
6999:birds
6048:Moros
5575:Siats
5316:Siats
5258:Siats
2918:Clade
2909:Clade
2704:S2CID
2649:S2CID
1959:In a
1950:femur
1946:tibia
1480:femur
1317:. In
1264:in a
1240:from
858:into
775:. In
701:. In
697:, by
606:Latin
598:mimos
594:ornis
402:genus
264:Marsh
222:Clade
209:Clade
196:Clade
183:Clade
7121:GBIF
6991:see
5618:Maip
3076:Tawa
2951:see
2828:and
2782:ISBN
2696:PMID
2641:PMID
2595:ISBN
2552:ISBN
2457:and
1961:2001
1662:2011
1623:1890
1591:2015
1572:2014
1564:2012
1547:2009
1543:2008
1531:1995
1386:2015
1374:2010
1370:Utah
1319:1985
1313:and
1266:2006
1260:and
1162:1988
1153:and
1143:2012
1129:2004
1111:and
1101:1997
1089:1854
1055:and
1049:1979
1041:1865
1017:Two
986:and
960:and
952:and
918:and
910:and
900:1972
894:and
886:and
872:1967
852:1903
777:1933
763:1868
749:and
743:1928
735:1926
723:1930
719:1911
713:for
703:1920
689:1916
681:1901
673:1902
648:1892
590:1889
463:1890
418:and
52:PreꞒ
7108:EoL
2834:doi
2759:doi
2731:doi
2688:doi
2676:511
2633:doi
2621:338
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2471:doi
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2163:110
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2011:doi
1920:An
1621:In
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1368:of
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1160:In
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1071:of
1059:as
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1047:in
1039:in
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972:Emu
938:or
870:in
793:or
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675:by
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