623:
227:(or Archaediptera) which includes several Triassic fossil members. The family has characteristics associated with the Nematocera as well as the Brachycera. The antennae are shortened as in the Brachycera and these flies are long, having a snout with vestigeal mouthparts, non-differentiated abdominal segments with large cerci. The wings are narrow and hair-fringed and have very weak venation. They are known to form cloud-like swarms in summer and the short-lived non-feeding adults have wings that fracture at the base shortly after mating.
106:
2956:
222:
in northern regions of the world, including northeastern North
America, Japan, the Himalayas, and eastern Russia. Around a dozen extant species are known, with two fossil species found in amber, extending back to the Mid Cretaceous. Under an alternative classification, they are considered the only
282:
have atrophied mouthparts. Nymphomyiidae are unusual in that the adults are ventrally holoptic, meaning they possess two eyes that meet on the underside of the head. Adults form large swarms above water. One or two generations may breed in a single year depending on the region and climate.
278:, retaining various larval features. They have strap-like wings with a very reduced venation, and the wing margins have long fringes like those of the Thysanoptera. The wings break at the base after mating. The antennae are very reduced. Species in the genus
258:
so distinct that he put it in a separate superfamily
Nymphomyioidea related to Triassic Dictyodipteridae which are in a suborder Archidiptera. Modern classifications put all the species in a single genus
491:
Schneeberg, Katharina; Friedrich, Frank; Courtney, Gregory W.; Wipfler, Benjamin; Beutel, Rolf G. (2012). "The larvae of
Nymphomyiidae (Diptera, Insecta) – Ancestral and highly derived?".
538:
Harper, P.P.; Lauzon, Michel (2012). "Life Cycle of the Nymph Fly
Palaeodipteron Walkeri Ide 1965 (Diptera: Nymphomyiidae) in the White Mountains of Southern Québec".
359:
343:
319:
335:
367:
327:
379:
351:
235:
192:
311:
2876:
2574:
635:
Courtney, G. 1998. First records of the
Nymphomyiidae (Diptera) in Nepal. Proceedings of the Entomological Society of Washington 100: 595–597.
2902:
602:
Courtney, G.W. (1994). "Biosystematics of the
Nymphomyiidae (Insecta: Diptera): life history, morphology, and phylogenetic relationships".
303:
672:
Rüdiger Wagner; Patrick Müller (2020). "The oldest of the rare: A new species of the family
Nymphomyiidae (Diptera) from Burmese amber".
744:
2997:
2850:
2889:
3026:
2894:
447:
Ide, F.P. 1965. A fly of the archaic family
Nymphomyiidae (Diptera) from North America. Canadian Entomologist, 97: 496-507.
699:
Wagner, R.; Hoffeins, C.; Hoffeins, H. W. (2000). "A fossil nymphomyiid (Diptera) from the Baltic and
Bitterfeld amber".
519:
565:
D. Huang; A. Nel; C. Cai; Q. Lin & M. S. Engel (2013). "Amphibious flies and paedomorphism in the
Jurassic period".
518:
Pape, Thomas; Blagoderov, Vladimir; Mostovski, Michail B. (2011). "Order Diptera Linnaeus, 1758". In Z. Q. Zhang (ed.).
2907:
737:
2990:
3016:
230:
The family Nymphomyiidae has several species which were originally placed in separate genera of their own.
105:
3021:
2803:
2798:
3031:
730:
622:
2983:
2726:
403:
Tokunaga M. (1932). "A remarkable dipterous insect from Japan, Nymphomyia alba, gen. et sp. nov".
2779:
2933:
2741:
1905:
645:
Ide, F. P. (2012). "A Fly of the Archaic Family Nymphomyiidae Found in New Brunswick in 1961".
521:
Animal biodiversity: an outline of higher-level classification and survey of taxonomic richness
2928:
2881:
2920:
2837:
8:
712:
464:
alba(Diptera: Nymphomyiidae) from Japan with notes on the larvae and the micro-habitat".
234:, the type species for this family, was discovered in a fast-flowing stream in Japan by
2971:
2398:
966:
423:
246:
was described from the Himalayas in 1970. Hennig examined the pupal characteristics of
100:
2915:
2824:
1251:
584:
517:
264:
2829:
2626:
2532:
2515:
2393:
1835:
1438:
1195:
1106:
708:
681:
654:
611:
576:
567:
547:
500:
473:
207:
2243:
2238:
2103:
2002:
1951:
1788:
1469:
1334:
1160:
1152:
974:
933:
2967:
2764:
2524:
2450:
2364:
2301:
2217:
2152:
2134:
2119:
2025:
2020:
1869:
1647:
1568:
1524:
1485:
1259:
1178:
1119:
1114:
982:
880:
685:
263:. Based on larval morphology, the family has been suggested to be close to the
504:
477:
290:
from the Jurassic of Asia, which are thought to have had a similar lifestyle.
3010:
2665:
2657:
2648:
2540:
2474:
2466:
2419:
2385:
2201:
2157:
2062:
2054:
1976:
1966:
1910:
1882:
1772:
1631:
1584:
1553:
1305:
1297:
1208:
1088:
1080:
1008:
909:
372:
59:
615:
2696:
2688:
2679:
2621:
2582:
2565:
2440:
2340:
2277:
2185:
2162:
2093:
2038:
2015:
1971:
1961:
1956:
1946:
1925:
1840:
1801:
1764:
1734:
1692:
1671:
1639:
1540:
1490:
1288:
1187:
1098:
1046:
1038:
941:
917:
855:
797:
588:
287:
224:
2790:
722:
2863:
2773:
2613:
2592:
2323:
2306:
2282:
2269:
2193:
2172:
2085:
2077:
2067:
2010:
1981:
1941:
1915:
1806:
1780:
1725:
1609:
1576:
1519:
1456:
1413:
1267:
1226:
1218:
1200:
1093:
1075:
888:
251:
34:
2816:
580:
551:
2963:
2605:
2548:
2482:
2458:
2432:
2377:
2209:
2180:
2147:
2124:
2046:
1933:
1900:
1887:
1861:
1796:
1684:
1511:
1498:
1443:
1393:
1350:
1342:
1059:
1025:
1017:
864:
837:
829:
813:
658:
167:
79:
44:
2868:
2955:
2631:
2587:
2424:
2414:
2348:
2332:
2256:
2248:
2230:
2142:
2129:
2111:
2098:
1877:
1827:
1814:
1751:
1743:
1655:
1558:
1545:
1532:
1477:
1422:
1358:
1318:
1127:
925:
785:
773:
137:
117:
84:
2855:
2735:
2811:
2758:
2356:
2264:
1989:
1892:
1848:
1700:
1663:
1448:
1430:
896:
791:
74:
69:
54:
49:
39:
490:
1622:
872:
275:
89:
64:
298:
Currently all species are treated as members of a single genus:
2406:
1464:
779:
767:
286:
Nymphomyiidae are thought to be closely related to the extinct
219:
147:
127:
2842:
421:
671:
215:
564:
754:
211:
157:
527:. Magnolia Press. Zootaxa, Vol. 3148. pp. 222–229.
698:
267:
while others place them in a separate infraorder, the
223:
living representatives of a separate, suborder called
424:"Insect mist-swarming of Nymphomyia species in Japan"
2962:
This article related to members of the fly suborder
1151:
965:
3008:
1250:
1666:(scuttle flies, coffin flies, humpbacked flies)
1203:(minute black scavenger flies, or dung midges)
601:
2991:
2656:
2523:
1177:
738:
460:Takemon, Y.; Tanida, K. (1994). "New data on
459:
2687:
1007:
828:
537:
1287:
854:
752:
402:
214:(Diptera). Larvae are found among aquatic
2998:
2984:
1412:
745:
731:
422:Saigusa, T.; T. Nakamura; S. Sato (2009).
323:Rohdendorf & Kalugina, 1974 - Siberia
558:
455:
453:
2388:(blow-flies: bluebottles, greenbottles)
1391:
811:
242:described by Ide in Quebec in 1965 and
3009:
1621:
210:of tiny (2 mm) slender, delicate
2740:
2739:
1390:
810:
726:
493:Arthropod Structure & Development
450:
2950:
713:10.1046/j.1365-3113.2000.025001115.x
604:Smithsonian Contributions to Zoology
644:
13:
1851:(cactus flies, banana stalk flies)
405:Annotationes Zoologicae Japonenses
14:
3043:
2954:
621:
104:
32:
692:
665:
638:
629:
595:
531:
511:
484:
441:
415:
396:
363:Courtney, 1994 - North America
238:in 1932. This was followed by
1:
1535:(dagger flies, balloon flies)
389:
3027:Monogeneric Diptera families
2970:. You can help Knowledge by
250:and placed it in the family
7:
2727:List of families of Diptera
2359:(house flies, stable flies)
347:Makarchenko, 1979 - Siberia
10:
3048:
2949:
1130:(dark-winged fungus gnats)
1122:(long-winged fungus gnats)
1109:(long-beaked fungus gnats)
686:10.11646/zootaxa.4763.2.11
293:
2748:
2724:
2678:
2647:
2604:
2573:
2564:
2514:
2510:
2449:
2376:
2331:
2322:
2294:
2229:
2204:(vinegar and fruit flies)
2171:
2114:(lekking, or druid flies)
2076:
2037:
2001:
1924:
1860:
1826:
1763:
1742:
1733:
1724:
1720:
1683:
1630:
1617:
1608:
1567:
1510:
1421:
1408:
1404:
1400:
1386:
1337:(long-bodied crane flies)
1317:
1296:
1283:
1246:
1217:
1186:
1173:
1147:
1058:
1037:
1016:
1003:
961:
908:
863:
850:
824:
820:
806:
762:
647:The Canadian Entomologist
540:The Canadian Entomologist
505:10.1016/j.asd.2012.01.002
478:10.1080/01650429409361544
339:Courtney 1994 - Hong Kong
331:(Kevan, 1970) - Himalayas
101:Scientific classification
99:
23:
1101:(predatory fungus gnats)
254:. Rohdendorf considered
1992:(black scavenger flies)
1353:(hairy-eyed craneflies)
1270:(primitive crane flies)
1028:(march flies, lovebugs)
616:10.5479/si.00810282.550
2634:(horse and deer flies)
1906:Strongylophthalmyiidae
1817:(picture-winged flies)
1345:(limoniid crane flies)
360:Nymphomyia dolichopeza
315:Tokunaga, 1932 - Japan
240:Palaeodipteron walkeri
2929:Paleobiology Database
2220:(frightful hairy fly)
2049:(beetle-backed flies)
1587:(tangle-veined flies)
1472:(hilarimorphid flies)
1433:(flower-loving flies)
1262:(phantom crane flies)
701:Systematic Entomology
244:Felicitomyia brundini
2551:(wood soldier flies)
2401:(New Zealand batfly)
2351:(little house flies)
1843:(stilt-legged flies)
1754:(thick-headed flies)
1650:(spear-winged flies)
1579:(small-headed flies)
1308:(winter crane flies)
883:(frog-biting midges)
355:(Ide, 1964) - Canada
3017:Nematocera families
2616:(water snipe flies)
2137:(upside-down flies)
1674:(flat-footed flies)
1658:(flat-footed flies)
1527:(long-legged flies)
1361:(large crane flies)
977:(net-winged midges)
944:(non-biting midges)
581:10.1038/nature11898
552:10.4039/Ent121603-7
368:Nymphomyia allissae
3022:Blephariceromorpha
2285:(freeloader flies)
2088:(leaf miner flies)
2028:(small dung flies)
1872:(stalk-eyed flies)
1695:(big-headed flies)
1153:Perissommatomorpha
983:Deuterophlebiidae
967:Blephariceromorpha
659:10.4039/Ent96119-1
380:Nymphomyia succina
274:Nymphomyiidae are
2979:
2978:
2944:
2943:
2916:Open Tree of Life
2742:Taxon identifiers
2733:
2732:
2720:
2719:
2716:
2715:
2712:
2711:
2708:
2707:
2700:
2674:
2673:
2643:
2642:
2635:
2617:
2596:
2560:
2559:
2552:
2544:
2536:
2533:Pantophthalmidae
2506:
2505:
2502:
2501:
2498:
2497:
2494:
2493:
2486:
2478:
2470:
2462:
2436:
2428:
2410:
2402:
2389:
2368:
2360:
2352:
2344:
2318:
2317:
2310:
2286:
2273:
2260:
2252:
2239:Acartophthalmidae
2221:
2213:
2205:
2197:
2189:
2188:(quasimodo flies)
2138:
2115:
2107:
2089:
2058:
2050:
2029:
1993:
1985:
1937:
1896:
1873:
1852:
1844:
1818:
1810:
1792:
1784:
1776:
1755:
1716:
1715:
1712:
1711:
1704:
1696:
1675:
1667:
1659:
1651:
1643:
1604:
1603:
1600:
1599:
1596:
1595:
1588:
1580:
1549:
1536:
1528:
1502:
1494:
1481:
1473:
1460:
1452:
1434:
1382:
1381:
1378:
1377:
1374:
1373:
1370:
1369:
1362:
1354:
1346:
1338:
1328:
1309:
1279:
1278:
1271:
1263:
1252:Ptychopteromorpha
1242:
1241:
1238:
1237:
1230:
1204:
1169:
1168:
1143:
1142:
1139:
1138:
1131:
1123:
1110:
1107:Lygistorrhinidae
1102:
1084:
1069:
1050:
1029:
999:
998:
986:
985:(mountain midges)
978:
957:
956:
953:
952:
945:
937:
929:
921:
920:(solitary midges)
900:
892:
884:
876:
875:(meniscus midges)
846:
845:
265:Deuterophlebiidae
200:
199:
196:
30:Cenomanian–Recent
3039:
3032:Nematocera stubs
3000:
2993:
2986:
2958:
2951:
2937:
2936:
2924:
2923:
2911:
2910:
2898:
2897:
2885:
2884:
2872:
2871:
2859:
2858:
2846:
2845:
2833:
2832:
2820:
2819:
2807:
2806:
2794:
2793:
2784:
2783:
2782:
2769:
2768:
2767:
2737:
2736:
2698:
2685:
2684:
2654:
2653:
2649:Vermileonomorpha
2633:
2627:Pelecorhynchidae
2615:
2594:
2571:
2570:
2550:
2542:
2534:
2521:
2520:
2516:Stratiomyomorpha
2512:
2511:
2484:
2476:
2468:
2460:
2434:
2426:
2408:
2400:
2399:Mystacinobiidae
2394:Mesembrinellidae
2387:
2366:
2358:
2350:
2342:
2329:
2328:
2308:
2284:
2271:
2258:
2250:
2219:
2211:
2203:
2195:
2187:
2136:
2113:
2105:
2104:Aulacigastridae
2087:
2056:
2048:
2027:
1991:
1983:
1935:
1911:Syringogastridae
1894:
1871:
1850:
1842:
1836:Cypselosomatidae
1816:
1808:
1790:
1789:Platystomatidae
1782:
1774:
1753:
1740:
1739:
1731:
1730:
1722:
1721:
1702:
1694:
1673:
1665:
1657:
1649:
1641:
1628:
1627:
1619:
1618:
1615:
1614:
1586:
1578:
1547:
1534:
1526:
1501:(stiletto flies)
1500:
1492:
1479:
1471:
1470:Hilarimorphidae
1458:
1450:
1439:Apsilocephalidae
1432:
1419:
1418:
1410:
1409:
1406:
1405:
1402:
1401:
1388:
1387:
1360:
1352:
1344:
1336:
1335:Cylindrotomidae
1327:
1326:
1322:
1307:
1294:
1293:
1285:
1284:
1269:
1261:
1248:
1247:
1228:
1202:
1196:Canthyloscelidae
1184:
1183:
1175:
1174:
1149:
1148:
1129:
1121:
1108:
1100:
1082:
1068:
1067:
1063:
1048:
1027:
1014:
1013:
1005:
1004:
984:
976:
975:Blephariceridae
963:
962:
943:
935:
934:Ceratopogonidae
927:
919:
898:
891:(phantom midges)
890:
882:
874:
861:
860:
852:
851:
826:
825:
822:
821:
808:
807:
747:
740:
733:
724:
723:
717:
716:
696:
690:
689:
669:
663:
662:
653:(1–2): 119–120.
642:
636:
633:
627:
626:
625:
619:
599:
593:
592:
562:
556:
555:
535:
529:
528:
526:
515:
509:
508:
488:
482:
481:
457:
448:
445:
439:
438:
428:
419:
413:
412:
400:
383:† (Baltic amber)
269:Nymphomyiomorpha
236:Masaaki Tokunaga
218:in small, rapid
191:
178:Nymphomyiomorpha
109:
108:
94:
31:
27:Temporal range:
21:
20:
3047:
3046:
3042:
3041:
3040:
3038:
3037:
3036:
3007:
3006:
3005:
3004:
2947:
2945:
2940:
2932:
2927:
2919:
2914:
2906:
2901:
2893:
2888:
2880:
2875:
2867:
2862:
2854:
2849:
2841:
2836:
2828:
2823:
2815:
2810:
2802:
2797:
2789:
2787:
2778:
2777:
2772:
2763:
2762:
2757:
2744:
2734:
2729:
2704:
2680:Xylophagomorpha
2670:
2639:
2600:
2556:
2543:(soldier flies)
2490:
2445:
2435:(tachina flies)
2372:
2365:Scathophagidae
2343:(cabbage flies)
2314:
2290:
2244:Australimyzidae
2225:
2218:Mormotomyiidae
2167:
2135:Neurochaetidae
2072:
2033:
2026:Sphaeroceridae
2003:Sphaeroceroidea
1997:
1957:Heterocheilidae
1952:Helosciomyzidae
1920:
1856:
1822:
1809:(peacock flies)
1775:(flutter flies)
1759:
1708:
1679:
1648:Lonchopteridae
1592:
1563:
1541:Homalocnemiidae
1525:Dolichopodidae
1506:
1396:
1366:
1324:
1323:
1321:
1313:
1275:
1260:Ptychopteridae
1234:
1213:
1165:
1161:Perissommatidae
1135:
1120:Rangomaramidae
1065:
1064:
1062:
1054:
1033:
995:
949:
936:(biting midges)
904:
881:Corethrellidae
842:
816:
802:
758:
751:
721:
720:
697:
693:
670:
666:
643:
639:
634:
630:
620:
600:
596:
575:(7439): 94–97.
563:
559:
536:
532:
524:
516:
512:
489:
485:
466:Aquatic Insects
458:
451:
446:
442:
426:
420:
416:
401:
397:
392:
307:Tokunaga, 1932
296:
232:Nymphomyia alba
190:
103:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
16:Family of flies
12:
11:
5:
3045:
3035:
3034:
3029:
3024:
3019:
3003:
3002:
2995:
2988:
2980:
2977:
2976:
2959:
2942:
2941:
2939:
2938:
2925:
2912:
2899:
2886:
2873:
2860:
2847:
2834:
2821:
2808:
2795:
2785:
2770:
2754:
2752:
2746:
2745:
2731:
2730:
2725:
2722:
2721:
2718:
2717:
2714:
2713:
2710:
2709:
2706:
2705:
2703:
2702:
2693:
2691:
2682:
2676:
2675:
2672:
2671:
2669:
2668:
2662:
2660:
2658:Vermileonoidea
2651:
2645:
2644:
2641:
2640:
2638:
2637:
2629:
2624:
2619:
2610:
2608:
2602:
2601:
2599:
2598:
2590:
2585:
2583:Austroleptidae
2579:
2577:
2568:
2562:
2561:
2558:
2557:
2555:
2554:
2546:
2541:Stratiomyidae
2538:
2535:(timber flies)
2529:
2527:
2525:Stratiomyoidea
2518:
2508:
2507:
2504:
2503:
2500:
2499:
2496:
2495:
2492:
2491:
2489:
2488:
2480:
2475:Nycteribiidae
2472:
2467:Hippoboscidae
2464:
2461:(tsetse flies)
2455:
2453:
2451:Hippoboscoidea
2447:
2446:
2444:
2443:
2438:
2430:
2425:Sarcophagidae
2422:
2417:
2412:
2404:
2396:
2391:
2386:Calliphoridae
2382:
2380:
2374:
2373:
2371:
2370:
2362:
2354:
2346:
2337:
2335:
2326:
2320:
2319:
2316:
2315:
2313:
2312:
2304:
2302:Cryptochetidae
2298:
2296:
2292:
2291:
2289:
2288:
2280:
2275:
2267:
2262:
2254:
2246:
2241:
2235:
2233:
2227:
2226:
2224:
2223:
2215:
2207:
2202:Drosophilidae
2199:
2191:
2183:
2177:
2175:
2169:
2168:
2166:
2165:
2160:
2155:
2153:Periscelididae
2150:
2145:
2140:
2132:
2127:
2122:
2120:Fergusoninidae
2117:
2109:
2101:
2096:
2091:
2082:
2080:
2074:
2073:
2071:
2070:
2065:
2060:
2055:Chamaemyiidae
2052:
2043:
2041:
2035:
2034:
2032:
2031:
2023:
2021:Nannodastiidae
2018:
2013:
2007:
2005:
1999:
1998:
1996:
1995:
1987:
1979:
1974:
1969:
1964:
1959:
1954:
1949:
1944:
1939:
1930:
1928:
1922:
1921:
1919:
1918:
1913:
1908:
1903:
1898:
1890:
1885:
1880:
1875:
1866:
1864:
1858:
1857:
1855:
1854:
1846:
1838:
1832:
1830:
1824:
1823:
1821:
1820:
1812:
1804:
1799:
1794:
1791:(signal flies)
1786:
1783:(cheese flies)
1778:
1773:Pallopteridae
1769:
1767:
1761:
1760:
1758:
1757:
1748:
1746:
1737:
1728:
1718:
1717:
1714:
1713:
1710:
1709:
1707:
1706:
1698:
1689:
1687:
1681:
1680:
1678:
1677:
1669:
1661:
1653:
1645:
1642:(ironic flies)
1636:
1634:
1625:
1612:
1606:
1605:
1602:
1601:
1598:
1597:
1594:
1593:
1591:
1590:
1585:Nemestrinidae
1582:
1573:
1571:
1569:Nemestrinoidea
1565:
1564:
1562:
1561:
1556:
1551:
1543:
1538:
1530:
1522:
1516:
1514:
1508:
1507:
1505:
1504:
1496:
1493:(window flies)
1488:
1486:Mythicomyiidae
1483:
1475:
1467:
1462:
1454:
1451:(robber flies)
1446:
1441:
1436:
1427:
1425:
1416:
1398:
1397:
1384:
1383:
1380:
1379:
1376:
1375:
1372:
1371:
1368:
1367:
1365:
1364:
1356:
1348:
1340:
1331:
1329:
1315:
1314:
1312:
1311:
1306:Trichoceridae
1302:
1300:
1298:Trichoceroidea
1291:
1281:
1280:
1277:
1276:
1274:
1273:
1265:
1256:
1254:
1244:
1243:
1240:
1239:
1236:
1235:
1233:
1232:
1223:
1221:
1215:
1214:
1212:
1211:
1206:
1198:
1192:
1190:
1181:
1179:Psychodomorpha
1171:
1170:
1167:
1166:
1164:
1163:
1157:
1155:
1145:
1144:
1141:
1140:
1137:
1136:
1134:
1133:
1125:
1117:
1115:Mycetophilidae
1112:
1104:
1096:
1091:
1086:
1081:Cecidomyiidae
1078:
1076:Bolitophilidae
1072:
1070:
1066:(fungus gnats)
1056:
1055:
1053:
1052:
1043:
1041:
1035:
1034:
1032:
1031:
1022:
1020:
1011:
1001:
1000:
997:
996:
994:
993:
988:
980:
971:
969:
959:
958:
955:
954:
951:
950:
948:
947:
939:
931:
923:
914:
912:
906:
905:
903:
902:
894:
886:
878:
869:
867:
858:
848:
847:
844:
843:
841:
840:
834:
832:
818:
817:
804:
803:
801:
800:
794:
788:
782:
776:
770:
763:
760:
759:
750:
749:
742:
735:
727:
719:
718:
691:
680:(2): 287–293.
664:
637:
628:
594:
557:
530:
510:
483:
449:
440:
414:
394:
393:
391:
388:
387:
386:
385:
384:
376:
364:
356:
348:
344:N. rohdendorfi
340:
332:
324:
320:N. levanidovae
316:
295:
292:
198:
197:
185:
181:
180:
175:
171:
170:
165:
161:
160:
155:
151:
150:
145:
141:
140:
135:
131:
130:
125:
121:
120:
115:
111:
110:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
3044:
3033:
3030:
3028:
3025:
3023:
3020:
3018:
3015:
3014:
3012:
3001:
2996:
2994:
2989:
2987:
2982:
2981:
2975:
2973:
2969:
2965:
2960:
2957:
2953:
2952:
2948:
2935:
2930:
2926:
2922:
2917:
2913:
2909:
2904:
2900:
2896:
2891:
2887:
2883:
2878:
2874:
2870:
2865:
2861:
2857:
2852:
2848:
2844:
2839:
2835:
2831:
2826:
2822:
2818:
2813:
2809:
2805:
2800:
2796:
2792:
2786:
2781:
2780:Nymphomyiidae
2775:
2771:
2766:
2760:
2756:
2755:
2753:
2751:
2750:Nymphomyiidae
2747:
2743:
2738:
2728:
2723:
2701:
2697:Xylophagidae
2695:
2694:
2692:
2690:
2689:Xylophagoidea
2686:
2683:
2681:
2677:
2667:
2666:Vermileonidae
2664:
2663:
2661:
2659:
2655:
2652:
2650:
2646:
2636:
2630:
2628:
2625:
2623:
2620:
2618:
2612:
2611:
2609:
2607:
2603:
2597:
2595:(snipe flies)
2591:
2589:
2586:
2584:
2581:
2580:
2578:
2576:
2572:
2569:
2567:
2563:
2553:
2547:
2545:
2539:
2537:
2531:
2530:
2528:
2526:
2522:
2519:
2517:
2513:
2509:
2487:
2481:
2479:
2473:
2471:
2469:(louse flies)
2465:
2463:
2457:
2456:
2454:
2452:
2448:
2442:
2439:
2437:
2431:
2429:
2427:(flesh flies)
2423:
2421:
2420:Rhinophoridae
2418:
2416:
2413:
2411:
2405:
2403:
2397:
2395:
2392:
2390:
2384:
2383:
2381:
2379:
2375:
2369:
2363:
2361:
2355:
2353:
2347:
2345:
2341:Anthomyiidae
2339:
2338:
2336:
2334:
2330:
2327:
2325:
2321:
2311:
2309:(lance flies)
2305:
2303:
2300:
2299:
2297:
2293:
2287:
2281:
2279:
2276:
2274:
2268:
2266:
2263:
2261:
2259:(beach flies)
2255:
2253:
2247:
2245:
2242:
2240:
2237:
2236:
2234:
2232:
2228:
2222:
2216:
2214:
2212:(shore flies)
2208:
2206:
2200:
2198:
2192:
2190:
2186:Curtonotidae
2184:
2182:
2179:
2178:
2176:
2174:
2170:
2164:
2161:
2159:
2158:Teratomyzidae
2156:
2154:
2151:
2149:
2146:
2144:
2141:
2139:
2133:
2131:
2128:
2126:
2123:
2121:
2118:
2116:
2110:
2108:
2102:
2100:
2097:
2095:
2092:
2090:
2084:
2083:
2081:
2079:
2075:
2069:
2066:
2064:
2063:Cremifaniidae
2061:
2059:
2057:(aphid flies)
2053:
2051:
2045:
2044:
2042:
2040:
2036:
2030:
2024:
2022:
2019:
2017:
2014:
2012:
2009:
2008:
2006:
2004:
2000:
1994:
1988:
1986:
1984:(marsh flies)
1980:
1978:
1977:Ropalomeridae
1975:
1973:
1970:
1968:
1967:Natalimyzidae
1965:
1963:
1960:
1958:
1955:
1953:
1950:
1948:
1945:
1943:
1940:
1938:
1932:
1931:
1929:
1927:
1923:
1917:
1914:
1912:
1909:
1907:
1904:
1902:
1899:
1897:
1891:
1889:
1886:
1884:
1883:Megamerinidae
1881:
1879:
1876:
1874:
1868:
1867:
1865:
1863:
1859:
1853:
1847:
1845:
1841:Micropezidae
1839:
1837:
1834:
1833:
1831:
1829:
1825:
1819:
1813:
1811:
1805:
1803:
1800:
1798:
1795:
1793:
1787:
1785:
1779:
1777:
1771:
1770:
1768:
1766:
1762:
1756:
1750:
1749:
1747:
1745:
1741:
1738:
1736:
1732:
1729:
1727:
1723:
1719:
1705:
1699:
1697:
1693:Pipunculidae
1691:
1690:
1688:
1686:
1682:
1676:
1672:Platypezidae
1670:
1668:
1662:
1660:
1654:
1652:
1646:
1644:
1640:Ironomyiidae
1638:
1637:
1635:
1633:
1632:Platypezoidea
1629:
1626:
1624:
1620:
1616:
1613:
1611:
1607:
1589:
1583:
1581:
1575:
1574:
1572:
1570:
1566:
1560:
1557:
1555:
1554:Oreogetonidae
1552:
1550:
1548:(dance flies)
1544:
1542:
1539:
1537:
1531:
1529:
1523:
1521:
1518:
1517:
1515:
1513:
1509:
1503:
1497:
1495:
1491:Scenopinidae
1489:
1487:
1484:
1482:
1480:(mydas flies)
1476:
1474:
1468:
1466:
1463:
1461:
1455:
1453:
1447:
1445:
1444:Apystomyiidae
1442:
1440:
1437:
1435:
1429:
1428:
1426:
1424:
1420:
1417:
1415:
1411:
1407:
1403:
1399:
1395:
1389:
1385:
1363:
1357:
1355:
1349:
1347:
1341:
1339:
1333:
1332:
1330:
1325:(crane flies)
1320:
1316:
1310:
1304:
1303:
1301:
1299:
1295:
1292:
1290:
1286:
1282:
1272:
1266:
1264:
1258:
1257:
1255:
1253:
1249:
1245:
1231:
1225:
1224:
1222:
1220:
1216:
1210:
1209:Valeseguyidae
1207:
1205:
1199:
1197:
1194:
1193:
1191:
1189:
1185:
1182:
1180:
1176:
1172:
1162:
1159:
1158:
1156:
1154:
1150:
1146:
1132:
1126:
1124:
1118:
1116:
1113:
1111:
1105:
1103:
1099:Keroplatidae
1097:
1095:
1092:
1090:
1089:Diadocidiidae
1087:
1085:
1083:(gall midges)
1079:
1077:
1074:
1073:
1071:
1061:
1057:
1051:
1047:Anisopodidae
1045:
1044:
1042:
1040:
1039:Anisopodoidea
1036:
1030:
1024:
1023:
1021:
1019:
1015:
1012:
1010:
1009:Bibionomorpha
1006:
1002:
992:
991:Nymphomyiidae
989:
987:
981:
979:
973:
972:
970:
968:
964:
960:
946:
942:Chironomidae
940:
938:
932:
930:
928:(black flies)
924:
922:
918:Thaumaleidae
916:
915:
913:
911:
910:Chironomoidea
907:
901:
895:
893:
887:
885:
879:
877:
871:
870:
868:
866:
862:
859:
857:
853:
849:
839:
836:
835:
833:
831:
830:Axymyiomorpha
827:
823:
819:
815:
809:
805:
799:
795:
793:
789:
787:
783:
781:
777:
775:
771:
769:
765:
764:
761:
756:
748:
743:
741:
736:
734:
729:
728:
725:
714:
710:
706:
702:
695:
687:
683:
679:
675:
668:
660:
656:
652:
648:
641:
632:
624:
617:
613:
610:(550): 1–41.
609:
605:
598:
590:
586:
582:
578:
574:
570:
569:
561:
553:
549:
545:
541:
534:
523:
522:
514:
506:
502:
498:
494:
487:
479:
475:
471:
467:
463:
456:
454:
444:
436:
432:
425:
418:
410:
406:
399:
395:
382:
381:
377:
374:
373:Burmese amber
370:
369:
365:
362:
361:
357:
354:
353:
349:
346:
345:
341:
338:
337:
333:
330:
329:
325:
322:
321:
317:
314:
313:
309:
308:
306:
305:
301:
300:
299:
291:
289:
284:
281:
277:
272:
270:
266:
262:
257:
253:
249:
245:
241:
237:
233:
228:
226:
221:
217:
213:
209:
205:
204:Nymphomyiidae
194:
189:
188:Nymphomyiidae
186:
183:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
139:
136:
133:
132:
129:
126:
123:
122:
119:
116:
113:
112:
107:
102:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
24:Nymphomyiidae
22:
19:
2972:expanding it
2961:
2946:
2749:
2622:Oreoleptidae
2614:Athericidae
2593:Rhagionidae
2588:Bolbomyiidae
2575:Rhagionoidea
2566:Tabanomorpha
2459:Glossinidae
2441:Ulurumyiidae
2367:(dung flies)
2307:Lonchaeidae
2295:Lonchaeoidea
2283:Milichiidae
2278:Inbiomyiidae
2272:(frit flies)
2270:Chloropidae
2194:Diastatidae
2163:Xenasteiidae
2094:Anthomyzidae
2086:Agromyzidae
2039:Lauxanioidea
2016:Heleomyzidae
1982:Sciomyzidae
1972:Phaeomyiidae
1962:Huttoninidae
1947:Helcomyzidae
1936:(kelp flies)
1926:Sciomyzoidea
1895:(rust flies)
1807:Tephritidae
1802:Richardiidae
1781:Piophilidae
1765:Tephritoidea
1735:Acalyptratae
1703:(hoverflies)
1577:Acroceridae
1457:Bombyliidae
1431:Apioceridae
1289:Tipulomorpha
1268:Tanyderidae
1229:(moth flies)
1227:Psychodidae
1219:Psychodoidea
1201:Scatopsidae
1188:Scatopsoidea
1049:(wood gnats)
990:
899:(mosquitoes)
889:Chaoboridae
856:Culicomorpha
798:Holometabola
796:Superorder:
790:Infraclass:
704:
700:
694:
677:
673:
667:
650:
646:
640:
631:
607:
603:
597:
572:
566:
560:
543:
539:
533:
520:
513:
496:
492:
486:
469:
465:
461:
443:
434:
430:
417:
408:
404:
398:
378:
366:
358:
350:
342:
336:N. holoptica
334:
326:
318:
310:
302:
297:
288:Strashilidae
285:
279:
273:
268:
260:
255:
247:
243:
239:
231:
229:
225:Archidiptera
203:
201:
187:
177:
174:Infraorder:
18:
2864:iNaturalist
2774:Wikispecies
2699:(awl flies)
2549:Xylomyidae
2485:(bat flies)
2483:Streblidae
2477:(bat flies)
2433:Tachinidae
2324:Calyptratae
2210:Ephydridae
2196:(bog flies)
2173:Ephydroidea
2106:(sap flies)
2078:Opomyzoidea
2068:Lauxaniidae
2047:Celyphidae
2011:Chyromyidae
1942:Dryomyzidae
1934:Coelopidae
1916:Tanypezidae
1726:Schizophora
1610:Muscomorpha
1520:Atelestidae
1499:Therevidae
1459:(bee flies)
1414:Asilomorpha
1351:Pediciidae
1343:Limoniidae
1094:Ditomyiidae
1026:Bibionidae
1018:Bibionoidea
926:Simuliidae
707:: 115–120.
328:N. brundini
252:Psychodidae
3011:Categories
2964:Nematocera
2632:Tabanidae
2606:Tabanoidea
2409:(botflies)
2407:Oestridae
2378:Oestroidea
2349:Fanniidae
2257:Canacidae
2251:(bee lice)
2249:Braulidae
2181:Camillidae
2148:Opomyzidae
2125:Marginidae
2112:Clusiidae
1901:Somatiidae
1888:Nothybidae
1870:Diopsidae
1862:Diopsoidea
1815:Ulidiidae
1797:Pyrgotidae
1752:Conopidae
1744:Conopoidea
1701:Syrphidae
1685:Syrphoidea
1656:Opetiidae
1546:Hybotidae
1533:Empididae
1512:Empidoidea
1394:Brachycera
1359:Tipulidae
1319:Tipuloidea
1128:Sciaridae
1060:Sciaroidea
897:Culicidae
865:Culicoidea
838:Axymyiidae
814:Nematocera
784:Subclass:
774:Arthropoda
546:(7): 603.
499:(3): 293.
472:(2): 119.
462:Nymphomyia
411:: 559–569.
390:References
352:N. walkeri
304:Nymphomyia
280:Nymphomyia
261:Nymphomyia
256:Nymphomyia
248:Nymphomyia
168:Nematocera
164:Suborder:
138:Arthropoda
2415:Rhiniidae
2357:Muscidae
2333:Muscoidea
2231:Carnoidea
2143:Odiniidae
2130:Neminidae
2099:Asteiidae
1990:Sepsidae
1893:Psilidae
1878:Gobryidae
1849:Neriidae
1828:Nerioidea
1664:Phoridae
1559:Ragadidae
1465:Evocoidae
1449:Asilidae
1423:Asiloidea
1392:Suborder
812:Suborder
786:Pterygota
766:Kingdom:
431:Fly Times
124:Kingdom:
118:Eukaryota
2812:BugGuide
2788:BioLib:
2765:Q1936442
2759:Wikidata
2265:Carnidae
1478:Mydidae
873:Dixidae
792:Neoptera
772:Phylum:
768:Animalia
757:families
589:23426262
276:neotenic
193:Tokunaga
184:Family:
134:Phylum:
128:Animalia
114:Domain:
1623:Aschiza
780:Insecta
778:Class:
755:Diptera
753:Extant
674:Zootaxa
312:N. alba
294:Species
220:streams
158:Diptera
154:Order:
148:Insecta
144:Class:
2934:140835
2908:560809
2895:121292
2882:109702
2869:557991
2817:172895
2804:449536
2791:779144
587:
568:Nature
437:: 2–8.
216:mosses
208:family
206:are a
195:, 1932
2966:is a
2921:53947
2877:IRMNG
2830:626HL
525:(PDF)
427:(PDF)
212:flies
2968:stub
2903:NCBI
2890:ITIS
2856:5574
2851:GBIF
2843:8976
2799:BOLD
678:4763
585:PMID
202:The
35:PreꞒ
2838:EoL
2825:CoL
709:doi
682:doi
655:doi
612:doi
608:550
577:doi
573:495
548:doi
544:121
501:doi
474:doi
371:† (
3013::
2931::
2918::
2905::
2892::
2879::
2866::
2853::
2840::
2827::
2814::
2801::
2776::
2761::
705:25
703:.
676:.
651:96
649:.
606:.
583:.
571:.
542:.
497:41
495:.
470:16
468:.
452:^
435:43
433:.
429:.
409:13
407:.
271:.
85:Pg
2999:e
2992:t
2985:v
2974:.
746:e
739:t
732:v
715:.
711::
688:.
684::
661:.
657::
618:.
614::
591:.
579::
554:.
550::
507:.
503::
480:.
476::
375:)
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:Ꞓ
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