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Megophryidae

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743: 767: 113: 137: 236: 551:(100-126 mya) via fossils of related frog groups, a study in early 2017 revealed that this was likely an overestimation. Using DNA sequencing, the study indicated the group more likely originated much later during the Cretaceous period, around 77 mya. The study also indicated that there are likely many more new species in the family that are currently unknown to science. 276:, and extending to the Philippines. Fossil remains are also known from North America. As of 2014 it encompasses 246 species of frogs divided between five genera. For lack of a better vernacular name, they are commonly called 547:
The origin of this group of frogs was largely unknown, due to the lack of members of this family in the fossil record. While the family was originally considered to have originated in the early-mid
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While the family is currently restricted to Asia, fossils indicate that it once had a much wider distribution extending to North America. The earliest known fossils of this family are from the
857:. Ed. Michael Hutchins, Arthur V. Evans, Jerome A. Jackson, Devra G. Kleiman, James B. Murphy, Dennis A. Thoney, et al. Vol. 6: Amphibians. 2nd ed. Detroit: Gale, 2004. p109-117. 296:, which often look like dead leaves. The camouflage is accurate to the point of some having skin folds that look like leaf veins, and at least one species, the 350: 1099: 991: 620: 457: 1017: 426: 468: 965: 1004: 1009: 307:
Megophryids range in size from 2 to 12.5 cm (0.79 to 4.92 in) in length. The adults' tongues are noticeably
826: 596: 840: 1104: 1056: 1022: 913: 669: 641: 337: 591:. Harold G. Cogger, Richard George Zweifel, David Kirshner (2nd ed.). San Diego, CA. 1998. p. 88. 1094: 1089: 931: 926: 136: 624: 1061: 918: 695: 213: 791: 495: 273: 122: 1048: 904: 866: 297: 1084: 1043: 996: 269: 1035: 952: 836: 342: 8: 304:) has sharp projections extending past the eye and nose, which disguise the frog shape. 614: 131: 1030: 939: 822: 602: 592: 112: 531: 719: 446: 345:
differ on the number of species per genera, leading to the variability in numbers:
257: 944: 666: 401: 357: 1078: 563: 515: 504: 421: 368: 70: 606: 889: 490: 435: 414: 198: 978: 898: 792:"Timetree dating in the absence of a fossil record in Asian Horned Frogs" 586: 45: 548: 379: 323:. The tadpoles are extremely diverse in form because of the variety of 289: 90: 55: 983: 526: 479: 390: 235: 178: 148: 95: 39: 970: 860: 883: 265: 168: 85: 80: 65: 60: 50: 559: 324: 312: 100: 75: 335:
The following genera are recognised in the family Megophryidae;
696:"Megophryidae Bonaparte, 1850 | Amphibian Species of the World" 555: 320: 308: 293: 158: 32: 28: 957: 676:. Version 6.0. American Museum of Natural History, New York 316: 261: 188: 315:
can be found in a variety of waters, but especially
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Amphibian Species of the World, an Online Reference
1076: 268:foothills eastwards, south to Indonesia and the 853:"Asian Toadfrogs (Megophryidae)". Amy Lathrop. 264:native to the warm southeast of Asia, from the 814:Cogger, H.G; Zweifel, R.G. & Kirschner, D. 431:(hidden-tympanum horned toads; 3-4 species) 619:: CS1 maint: location missing publisher ( 234: 111: 818:Encyclopedia of Reptiles & Amphibians 588:Encyclopedia of Reptiles & Amphibians 1100:Taxa named by Charles Lucien Bonaparte 1077: 500:(narrow-mouth horned toads; 6 species) 486:(Indonesian horned toads; 5-8 species) 288:The megophryids are notable for their 865: 864: 581: 579: 536:(strange-horned toads; 28-31 species) 375:(eastern spadefoot toads; 38 species) 353:(pseudomoustache toads; 216 species) 659: 636: 634: 475:(diminutive horned toads; 6 species) 13: 855:Grzimek's Animal Life Encyclopedia 847: 576: 442:(Chinese horned toads; 65 species) 14: 1116: 665: 631: 522:(Sarawak horned toads; 1 species) 292:, especially those that live in 135: 43: 768:"AmphibiaWeb -- Search Results" 744:"AmphibiaWeb -- Search Results" 784: 760: 736: 712: 688: 670:"Megophryidae Bonaparte, 1850" 511:(clay horned toads; 7 species) 462:Dubois, Ohler, and Pyron, 2021 453:(Karin Hills frogs; 8 species) 338:Amphibian Species of the World 1: 700:amphibiansoftheworld.amnh.org 569: 364:(Borneo frogs; 85-93 species) 283: 720:"AmphibiaWeb - Megophryidae" 542: 408:(cat-eyed toads; 24 species) 7: 820:(2nd ed.). Fog City Press. 642:"Fossilworks: Megophryidae" 10: 1121: 484:Kuhl and Van Hasselt, 1822 873: 330: 242: 233: 226: 221: 132:Scientific classification 130: 119: 110: 23: 843:. Retrieved 2006-MAY-08. 440:Fei, Ye, and Jiang, 2016 1105:Indomalayan realm fauna 395:Myers and Leviton, 1962 274:Maritime Southeast Asia 123:Leptobrachium hasseltii 298:long-nosed horned frog 27:Temporal range: Early 1044:Paleobiology Database 623:) CS1 maint: others ( 270:Greater Sunda Islands 243:Global range (black) 120:Java spadefoot toad ( 837:Animal Diversity Web 509:Beddard, 1908 "1907" 252:, commonly known as 16:Family of amphibians 451:Tian & Hu, 1983 1095:Amphibians of Asia 1090:Amphibian families 520:Lyu and Wang, 2023 473:Lyu and Wang, 2023 1072: 1071: 1031:Open Tree of Life 867:Taxon identifiers 829: 535: 521: 510: 499: 485: 474: 463: 452: 441: 430: 407: 396: 385: 374: 363: 302:Megophrys montana 247: 246: 217: 1112: 1065: 1064: 1052: 1051: 1039: 1038: 1026: 1025: 1013: 1012: 1000: 999: 987: 986: 974: 973: 961: 960: 948: 947: 935: 934: 922: 921: 909: 908: 907: 894: 893: 892: 862: 861: 834: 821: 815: 806: 805: 803: 802: 796:sciencedaily.com 788: 782: 781: 779: 778: 764: 758: 757: 755: 754: 740: 734: 733: 731: 730: 716: 710: 709: 707: 706: 692: 686: 685: 683: 681: 667:Frost, Darrel R. 663: 657: 656: 654: 652: 638: 629: 628: 618: 610: 583: 530: 519: 508: 494: 483: 472: 461: 450: 447:Brachytarsophrys 439: 425: 405: 394: 383: 372: 361: 238: 212: 140: 139: 115: 105: 42: 21: 20: 1120: 1119: 1115: 1114: 1113: 1111: 1110: 1109: 1075: 1074: 1073: 1068: 1060: 1055: 1047: 1042: 1034: 1029: 1021: 1016: 1008: 1003: 995: 990: 982: 977: 969: 964: 956: 951: 943: 938: 930: 925: 917: 912: 903: 902: 897: 888: 887: 882: 869: 850: 848:Further reading 832: 813: 810: 809: 800: 798: 790: 789: 785: 776: 774: 772:amphibiaweb.org 766: 765: 761: 752: 750: 748:amphibiaweb.org 742: 741: 737: 728: 726: 724:amphibiaweb.org 718: 717: 713: 704: 702: 694: 693: 689: 679: 677: 664: 660: 650: 648: 646:fossilworks.org 640: 639: 632: 612: 611: 599: 585: 584: 577: 572: 545: 351:Leptobrachiinae 333: 311:-shaped. Their 286: 211: 134: 106: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 37: 36: 25: 17: 12: 11: 5: 1118: 1108: 1107: 1102: 1097: 1092: 1087: 1070: 1069: 1067: 1066: 1053: 1040: 1027: 1014: 1001: 988: 975: 962: 949: 936: 923: 910: 895: 879: 877: 871: 870: 859: 858: 849: 846: 845: 844: 830: 808: 807: 783: 759: 735: 711: 687: 658: 630: 597: 574: 573: 571: 568: 544: 541: 540: 539: 538: 537: 523: 512: 501: 487: 476: 465: 454: 443: 432: 417:(124 species) 411: 410: 409: 406:Theobald, 1868 398: 387: 376: 365: 358:Leptobrachella 332: 329: 327:they inhabit. 285: 282: 245: 244: 240: 239: 231: 230: 224: 223: 219: 218: 206: 202: 201: 196: 192: 191: 186: 182: 181: 176: 172: 171: 166: 162: 161: 156: 152: 151: 146: 142: 141: 128: 127: 117: 116: 108: 107: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 26: 15: 9: 6: 4: 3: 2: 1117: 1106: 1103: 1101: 1098: 1096: 1093: 1091: 1088: 1086: 1083: 1082: 1080: 1063: 1058: 1054: 1050: 1045: 1041: 1037: 1032: 1028: 1024: 1019: 1015: 1011: 1006: 1002: 998: 993: 989: 985: 980: 976: 972: 967: 963: 959: 954: 950: 946: 941: 937: 933: 928: 924: 920: 915: 911: 906: 900: 896: 891: 885: 881: 880: 878: 876: 872: 868: 863: 856: 852: 851: 842: 838: 831: 828: 827:1-877019-69-0 824: 819: 812: 811: 797: 793: 787: 773: 769: 763: 749: 745: 739: 725: 721: 715: 701: 697: 691: 675: 671: 668: 662: 647: 643: 637: 635: 626: 622: 616: 608: 604: 600: 598:0-12-178560-2 594: 590: 589: 582: 580: 575: 567: 565: 564:United States 561: 557: 552: 550: 533: 529: 528: 524: 518: 517: 516:Sarawakiphrys 513: 507: 506: 505:Pelobatrachus 502: 497: 493: 492: 488: 482: 481: 477: 471: 470: 466: 460: 459: 455: 449: 448: 444: 438: 437: 433: 428: 424: 423: 422:Atympanophrys 419: 418: 416: 412: 404: 403: 399: 393: 392: 388: 382: 381: 377: 373:Tschudi, 1838 371: 370: 369:Leptobrachium 366: 360: 359: 355: 354: 352: 348: 347: 346: 344: 340: 339: 328: 326: 322: 318: 314: 310: 305: 303: 299: 295: 291: 281: 279: 275: 271: 267: 263: 259: 256:, is a large 255: 251: 241: 237: 232: 229: 225: 220: 215: 210: 207: 204: 203: 200: 197: 195:Superfamily: 194: 193: 190: 187: 184: 183: 180: 177: 174: 173: 170: 167: 164: 163: 160: 157: 154: 153: 150: 147: 144: 143: 138: 133: 129: 125: 124: 118: 114: 109: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 41: 34: 30: 22: 19: 1085:Megophryidae 919:Megophryidae 905:Megophryidae 875:Megophryidae 874: 854: 841:Megophryidae 817: 799:. Retrieved 795: 786: 775:. Retrieved 771: 762: 751:. Retrieved 747: 738: 727:. Retrieved 723: 714: 703:. Retrieved 699: 690: 678:. Retrieved 673: 661: 649:. Retrieved 645: 587: 553: 546: 525: 514: 503: 491:Ophryophryne 489: 478: 467: 458:Grillitschia 456: 445: 436:Boulenophrys 434: 429:and Hu, 1983 420: 415:Megophryinae 400: 397:(19 species) 389: 386:(50 species) 384:Dubois, 1980 378: 367: 356: 336: 334: 306: 301: 287: 277: 253: 250:Megophryidae 249: 248: 227: 209:Megophryidae 208: 199:Pelobatoidea 121: 38:50.3–0  24:Megophryidae 18: 979:iNaturalist 899:Wikispecies 833:Heying, H. 651:17 December 464:(2 species) 362:Smith, 1925 343:AmphibiaWeb 278:megophryids 254:goose frogs 1079:Categories 801:2017-02-26 777:2021-03-25 753:2021-03-25 729:2022-09-03 705:2022-09-03 570:References 549:Cretaceous 469:Jingophrys 413:Subfamily 380:Leptolalax 349:Subfamily 290:camouflage 284:Morphology 680:29 August 615:cite book 543:Evolution 527:Xenophrys 496:Boulenger 480:Megophrys 391:Oreolalax 266:Himalayan 228:See text 214:Bonaparte 155:Kingdom: 149:Eukaryota 884:Wikidata 839:– 835:(2003): 816:(2004): 607:39559811 402:Scutiger 325:habitats 313:tadpoles 205:Family: 179:Amphibia 169:Chordata 165:Phylum: 159:Animalia 145:Domain: 1062:1397874 562:in the 560:Wyoming 532:Günther 321:streams 294:forests 222:Genera 185:Order: 175:Class: 1049:249610 1036:828160 1023:161696 1010:550192 997:106097 932:110762 890:Q56056 825:  605:  595:  556:Eocene 534:, 1864 498:, 1903 331:Genera 309:paddle 258:family 216:, 1850 33:Recent 29:Eocene 1057:WoRMS 992:IRMNG 984:24601 317:ponds 262:frogs 189:Anura 1018:NCBI 1005:ITIS 971:3087 966:GBIF 958:7503 927:BOLD 823:ISBN 682:2020 653:2021 625:link 621:link 603:OCLC 593:ISBN 427:Tian 341:and 319:and 46:PreꞒ 953:EoL 945:CJ6 940:CoL 914:ASW 558:of 272:in 260:of 31:to 1081:: 1059:: 1046:: 1033:: 1020:: 1007:: 994:: 981:: 968:: 955:: 942:: 929:: 916:: 901:: 886:: 794:. 770:. 746:. 722:. 698:. 672:. 644:. 633:^ 617:}} 613:{{ 601:. 578:^ 566:. 280:. 126:) 96:Pg 40:Ma 35:, 804:. 780:. 756:. 732:. 708:. 684:. 655:. 627:) 609:. 300:( 101:N 91:K 86:J 81:T 76:P 71:C 66:D 61:S 56:O 51:Ꞓ

Index

Eocene
Recent
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Leptobrachium hasseltii
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Amphibia
Anura
Pelobatoidea
Megophryidae
Bonaparte

family
frogs

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