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Marine Biological Laboratory

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386:. He said of the MBL of that day, "Today, it stands as the uniquely national center for biology in this country; it is the National Biological Laboratory without being officially designated (or yet funded) as such. Its influence on the growth and development of biologic science has been equivalent to that of many of the country's universities combined, for it has had its pick of the world's scientific talent for each summer's research and teaching. Someone has counted thirty Nobel Laureates who have worked at the MBL at one time or another. It is amazing that such an institution, exerting so much influence on academic science, has been able to remain so absolutely autonomous. It has, to be sure, linkages of various kinds, arrangements with outside universities for certain graduate programs, and it adheres delicately, somewhat ambiguously, to the 359: 391:
evolve. It may shift soon into a new phase, with a year-round program for teaching and research and a year-round staff, but it will have to accomplish this without jeopardizing the immense power of its summer programs, or all institutional hell will break loose. It will have to find new ways for relating to the universities, if its graduate programs are to expand as they should. It will have to develop new symbiotic relations with the Oceanographic Institute, since both places have so much at stake. And it will have to find more money, much more — the kind of money that only federal governments possess — without losing any of its own initiative. It will be an interesting place to watch, in the years ahead."
315: 36: 422: 303: 27: 436: 880:, an emphasis maintained today. The Ecosystems Center has a year-round staff of more than 40 scientists who study a variety of ecosystems and their responses to human activities and environmental changes. The center is located in Woods Hole yet has a global reach, with active research sites in the Arctic tundra; in forest, coastal and marine sites in New England, Sweden and Brazil. The Ecosystems Center is home to two of the 26 U.S. Long Term Ecological Research (LTER) sites: 410: 863:(Nobel Laureate in 1937) conducted research at the MBL from 1947 to 1986, most significantly on the biochemical nature of muscular contraction. In the 1950s and 1960s, Frederik Bang and Jack Levin at the MBL discovered that the blood of the horseshoe crab clotted when exposed to bacterial endotoxins even in vanishingly small amounts. From this basic research, a reagent, Limulus amoebocyte lysate (LAL), was developed that can detect minute amounts of bacterial toxins. The 448: 756:
promise for translation to treatments for human conditions, including spinal cord injury, diabetes, organ failure, and degenerative neural diseases such as Alzheimer's. A cornerstone of the Bell Center is a national resource for research on the frog, Xenopus, which is a major animal model used in U.S. biomedical research. The National Xenopus Resource at the MBL is funded by the National Institutes of Health (MBL Facts).
331:, an embryologist, was retained to also serve as the first director of the MBL. Whitman believed "other things being equal, the investigator is always the best instructor," and emphasized the need to combine research and education at the new laboratory. The MBL's first summer course provided a six-week introduction to invertebrate zoology; facilities for visiting summer investigators were also offered. 673:, that are ideal for embryological analysis because they shed nearly transparent eggs which are fertilized and develop externally. In the first decades after the course was founded in 1893, its faculty pioneered research directions that remain central today, including the study of cytoplasmic localization in eggs; embryonic cell lineage (important in modern 529:. Whitman Center Scientists comprise more than 100 principal investigators from academic institutions around the world. Other resources include The Marine Resources Center, an advanced facility for maintaining, culturing, and providing aquatic and marine organisms essential to biological, biomedical, and ecological research; and The 621:(GFP) in the early 1960s, which led to the development of revolutionary techniques for imaging live cells and their components. Resident Distinguished Scientist Shinya Inoué's innovations in polarized light microscopy and video imaging since the 1950s have been instrumental in clarifying the cellular events of 947:
The MBL offers a range of courses, workshops, conferences, and internships throughout the year. Central to its programs are more than 20 Advanced Research Training Courses, graduate-level courses in topics ranging from physiology, embryology, neurobiology, and microbiology to imaging and computation
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Ecosystems research became a year-round commitment at the MBL in 1962 with the founding of the Systematics-Ecology program, under the direction of Melbourne R. Carriker. In 1975, the MBL's Ecosystems Center was established, with George Woodwell as director. The original research focus was on the
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just up the street. But it has never come under the domination of any outside institution or governmental agency, nor has it ever been told what to do by any outside group. There is no way of predicting what the future will be like for an institution such as the MBL. One way or another, it will
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The MBL has approximately 250 year-round employees, about half of which are scientists and scientific support staff. They are joined each year by more than 500 visiting scientists, summer staff, and research associates from hundreds of institutions around the world, as well as a large number of
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on July 1, 2013. In order to further scientific research and education, the affiliation builds on historical ties with the university, as MBL was led by University of Chicago faculty members in its first four decades. The president of the university chairs the MBL trustee's board and with their
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of marine micro-organisms. Early results from this census in 2006 revealed some 10 to 100 times more types of marine microbes than expected, and the vast majority are previously unknown, low-abundance microorganisms now called the "rare biosphere". Other Bay Paul Center projects are focused on
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and sensory physiology are significant, fostered today by more than 65 visiting investigators and resident researchers in these fields, as well as five graduate- and post-graduate level Advanced Research Training courses. The MBL has been a magnet for the discipline since L.W. Williams in 1910
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In 2010, the MBL established the Eugene Bell Center for Regenerative Biology and Tissue Engineering, where researchers study the ability of marine and other animals to spontaneously regenerate damaged or aging body parts. An understanding of tissue and organ regeneration in lower animals holds
544:. The MBLWHOI Library holds print and electronic collections in the biological, biomedical, ecological, and oceanographic sciences, and houses a growing archival collection, including photograph and videos from the MBL's history. The library also conducts digitization and informatics projects. 793:
technique developed by Cole, laid the basis for the modern understanding of electrical activity in the nervous system by measuring quantitatively the flow of ions across the axonal membrane. Hodgkin and Huxley received the Nobel Prize in 1963 for their description of the ionic basis of nerve
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A large portion of the leading developmental biologists in the United States, both historically and today, have participated in the MBL's Embryology Course as directors, lecturers or students. One draw is the Woods Hole location and the availability of marine organisms, particularly the
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to highly acidic ecosystems, which may lead to a better understanding of life that could exist on other planets. Activities at the Bay Paul Center are supported by advanced DNA sequencing and other genomics equipment at the center's Keck Ecological and Evolutionary Genetics Facility.
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of terrestrial ecosystems, and John Hobbie, a microbial ecologist. The Ecosystems Center is founded on a vision of collaborative, interdisciplinary science; shared lab facilities and instrumentation; and a long-term, large-scale, systems-wide view of ecosystem processes.
884:, Alaska; and Plum Island, Massachusetts. Scientists in the Ecosystems Center study the effects of forest clearance and land-use change on atmospheric chemistry, watershed processes and coastal ecology, the global-scale anthropogenic enrichment of the 289:, land was purchased, a building was erected, and the MBL was incorporated with Hyatt as the first president of the board of trustees. The Fish Commission supplied crucial support, including marine organisms and running sea water. 827:, an MBL summer investigator in the 1920s and early 1930s, uncovered several basic mechanisms of photoreceptor function through his studies on the horseshoe crab. Hartline shared the 1967 Nobel Prize with summer MBL colleague 664:
began coming to the MBL to study the role that the protein ubiquitin plays in cell division. In 2004, Hershko won a Nobel Prize for his work to establish the basic mechanism of ubiquitin-mediated protein degradation.
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fundamental biological research, often using marine organisms as novel model systems, encompassing research in regenerative biology, neuroscience, sensory physiology, and comparative evolution and genomics;
1270: 781:, who in 1938 used it to record the resistance changes underlying the action potential, which provided evidence that ions flowing across the axonal membrane generate this electrical impulse. In 1938, 613:
and imaging. The availability of cutting-edge imaging instrumentation in the MBL's Advanced Research Courses puts faculty and students at the forefront of experimentation. MBL Distinguished Scientist
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Cell, developmental, and reproductive biology have been a central part of the MBL's programs since the 1890s. Important discoveries in these fields at the MBL reach back to 1899, when
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The Josephine Bay Paul Center for Comparative Molecular Biology and Evolution was founded at the MBL in 1997 and is currently directed by David Mark Welch. By comparing diverse
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1897 Embryology Class, six women sitting in front row, two rows of men standing behind them, many with hats on, most looking at camera. Gertrude Stein sitting on lookers left.
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Other marine organisms draw neuroscientists and neurobiologists to the MBL each summer, where a history of research into sensory physiology and behavior has been established.
2221: 1719: 350:, later well known as a novelist and art collector, took part in MBL's Embryology course in the summer of 1897, while her brother Leo took part in the Invertebrates course. 2266: 1694: 2296: 1740: 2166: 2161: 1780: 1341: 2286: 602:, considered by many to be a founder of modern cell biology due to his pioneering work on the fine structure of cells, including the discovery of 2226: 1245: 382: 859:
in the mid-1960s at Harvard Medical School and he also initiated instruction in neurobiology at the MBL (Barlow et al., 1993:175-234; 203-234).
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and others made connections between the chromosomes and Mendelian heredity, while Wilson's colleague at both the MBL and Columbia University,
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received the 2012 Lasker Award for Basic Medical Research for their discoveries related to molecular motors. In 1991 Israeli scientist
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column called "Notes of a Biology-Watcher", in an installment called "The MBL"; the essay was later collected into the volume titled
285:, off the coast of Woods Hole, Hyatt accepted the offer. With $ 10,000 raised by the Woman's Education Association of Boston and the 1498:
Kenney, Diana E. and Borisy, Gary G. (2009) Thomas Hunt Morgan at the Marine Biological Laboratory: Naturalist and Experimentalist.
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described the transmission properties at the squid giant synapse (Llinas 1999). The "scientific career" of the "Woods Hole squid",
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identified circulating hormones that influence sexual differentiation (Lillie, 1944). In the MBL's first two decades, cytologists
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winners have been affiliated with MBL as students, faculty members or researchers. In addition, since 1960, there have been, 137
537:(frog) genetic stocks; and provides training in Xenopus husbandry, cell biology, imaging, genetics, transgenesis, and genomics. 228:
winners have been affiliated with MBL as students, faculty members or researchers. In addition, since 1960, there have been 137
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and other colleges and universities, as well as workshops and conferences—accommodating more than 2,600 participants in 2016.
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advice appoints its members. The Laboratory is a non-profit Massachusetts corporation, whose sole member is the university.
2276: 2216: 2206: 2121: 2019: 1896: 1876: 1820: 1679: 606:, carried out research at the MBL starting in 1937 and directed the laboratory from 1975-77 (Barlow et al., 1993: 95-115). 541: 387: 1316: 2271: 2101: 1932: 1271:"University of Chicago President Robert J. Zimmer to serve as Chairman of Marine Biological Laboratory Board of Trustees" 974: 376: 2301: 1647: 678: 1187:
A boating party aboard The Vigilant at a dock in Falmouth, Massachusetts. Gertrude Stein is at the extreme right rear.
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faculty and students participating in MBL courses (in 2016, 550 students from 333 institutions and 58 countries).
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both explained and praised the nature of the MBL as a scientific institution. He wrote about it in his recurring
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has resulted in dramatic improvement in the quality of drugs and biological products for intravenous injection.
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first identified egg cytoplasmic regions that are programmed to form certain tissues or organs; to 1916, when
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and others at the MBL identified the first of a class of proteins that regulate the cycle of cell division (
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the study of microbiomes and microbial diversity and ecology in a variety of ocean and terrestrial habitats;
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research station in Woods Hole in 1882, and had ambitions to expand it into a major laboratory. He invited
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conduction (Barlow et al., 1993: 151-172). Following on Hodgkin and Huxley's work, in the 1960s and 1970s
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investigators, early career scientists, international researchers, and professors; 306 members of the
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investigators, early career scientists, international researchers, and professors; 306 members of the
2070: 618: 358: 214: 112: 35: 2261: 2096: 2065: 1886: 1684: 832: 514: 274: 518: 2111: 1616: 984: 617:, who joined the MBL in 1983, was awarded the 2008 Nobel Prize in Chemistry for his discovery of 340: 254: 860: 1927: 1795: 1785: 1640: 824: 785:
came to the MBL to learn about the squid giant axon from Cole. After World War II, Hodgkin and
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In September 2018, Nipam Patel became director of the Marine Biological Laboratory, succeeding
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The MBL has long been a center for the world's experts in cell division. In the early 1980s,
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that allow sodium and potassium ions to carry electric current across the cell membrane and
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One Hundred Years Exploring Life, 1888-1988: The Marine Biological Laboratory at Woods Hole
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ecosystems science and climate change, and organismal adaptation to changing environments.
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In addition, the MBL hosts courses for undergraduate and graduate students from the
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The Biological Century: Friday Evening Talks at the Marine Biological Laboratory.
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The Marine Biological Laboratory grew from the vision of several Bostonians and
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The MBL Library was established in 1889, with scientist and future MBL trustee
1488:, Marine Biological Laboratory, 1893-1993. Pamphlet, MBLWHOI Library Archives. 1177:(photograph). Falmouth, Massachusetts: Historic New England. 1897. GUSN-197262 2255: 1565: 1548: 897: 790: 786: 661: 657: 637: 435: 278: 269:
to move his marine biology laboratory and school which he had founded at the
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demonstrated artificial parthenogenesis in sea urchin eggs; to 1905, when
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The trustees of the Marine Biological Laboratory in 1934. The lone woman,
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Institute for the Study of Ancient Cultures, West Asia & North Africa
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Josephine Bay Paul Center for Comparative Molecular Biology and Evolution
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on July 1, 2013. It also collaborates with numerous other institutions.
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and other MBL researchers described a number of the properties of the
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Controlling Life: Jacques Loeb and the Engineering Ideal in Biology
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Eugene Bell Center for Regenerative Biology and Tissue Engineering
1419: 1317:"Introducing Nipam Patel: Director, Marine Biological Laboratory" 914: 840: 653: 622: 598:, launched the field of experimental genetics (Pauly, 2000:158). 534: 84: 1366: 909:
Comparative genomics, molecular evolution, and microbial ecology
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Research institution in Woods Hole, Massachusetts, United States
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Barlow, Robert B., John E. Dowling, and Gerald Weissmann, eds.
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Graham School of Continuing Liberal and Professional Studies
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Rainger, Ronald, Keith R. Benson and Jane Maienschein, eds.
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The Squid Giant Synapse: A Model for Chemical Transmission
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found in a wide range of ecosystems, including the human
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The MBL is also a proving ground for new technologies in
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serving as librarian. In 1899, the MBL began publishing
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Woods Hole: The Marine Biological Laboratory, 1993.
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The MBL shares a library, the MBLWHOI Library, with
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Biological research institutes in the United States
892:. The interim director of the Ecosystems Center is 1617:"Cornelia Clapp and the Earliest Years of the MBL" 760:Neuroscience, neurobiology, and sensory physiology 934:microbes that live in extreme environments, from 2297:Research institutes of the University of Chicago 2253: 1509:. Chicago: University Press, 1944. Reprinted in 1491:Hunt, Tim (2004) "The Discovery of Cyclin (I)." 1246:The Lives of a Cell: Notes of a Biology Watcher 1202:(1972), "Notes of a biology-watcher: the MBL", 1192: 383:The Lives of a Cell: Notes of a Biology Watcher 277:, to Woods Hole. Inspired by Harvard biologist 1129:"Women of Science, Gertrude Stein (1874-1946)" 320:The laboratory, hatchery, aquarium, and museum 1641: 1444:"The College at Marine Biological Laboratory" 1391:"MBL – University of Chicago Affiliation FAQ" 855:, is credited with "founding" the science of 789:, working in Plymouth, England and using the 570:Cell, developmental, and reproductive biology 1235: 1233: 552:Research at the MBL focuses on four themes: 513:The MBL's resident research centers are the 460:The MBL became formally affiliated with the 1604:History of the Marine Biological Laboratory 1570:Biologists and the Promise of American Life 1555:. New York: Oxford University Press, 1987. 1507:The Woods Hole Marine Biological Laboratory 1648: 1634: 366:, stands near the center of the front row. 34: 25: 1655: 1572:. Princeton, NJ: University Press, 2000. 1230: 1081:"Nobel Laureates Affiliated with the MBL" 851:. Another long-term summer investigator, 1933:Francis and Rose Yuen (Hong Kong) campus 1715:Pritzker School of Molecular Engineering 1054:"Affiliation with University of Chicago" 357: 2287:Research institutes established in 1888 1761:Chicago Project on Security and Threats 1720:School of Social Service Administration 1484:Davidson, Eric (1993). "Introduction", 831:, who described the molecular basis of 652:, Joe DeGiorgis, and others discovered 505:who have been affiliated with the lab. 240:who have been affiliated with the lab. 2254: 1756:Center for Research in Security Prices 1700:Harris School of Public Policy Studies 1275:Affiliation with University of Chicago 1239: 1198: 1058:Affiliation with University of Chicago 1629: 1414: 1412: 1410: 948:integrated with biological research. 870: 809:Doryteuthis (formerly Loligo) pealeii 503:American Academy of Arts and Sciences 238:American Academy of Arts and Sciences 2292:1888 establishments in Massachusetts 1897:Housing at the University of Chicago 1610:WHOAS: Woods Hole Open Access Server 1075: 1073: 1048: 1046: 1021: 1019: 1017: 1015: 1013: 1011: 1009: 1007: 1005: 942: 835:by showing that the light-sensitive 542:Woods Hole Oceanographic Institution 388:Woods Hole Oceanographic Institution 1584:The American Development of Biology 975:Scripps Institution of Oceanography 812:, continues today, with studies on 13: 1464: 1407: 1383: 679:evolutionary developmental biology 14: 2313: 2061:Kenwood Astrophysical Observatory 1746:Center for Middle Eastern Studies 1597: 1070: 1043: 1002: 751:Regenerative biology and medicine 625:, including his discovery of the 327:University of Chicago professor, 287:Boston Society of Natural History 1841:Sonia Shankman Orthogenic School 1826:National Opinion Research Center 718:(1980–1982) (see Davidson, 1993) 446: 434: 420: 408: 313: 301: 2127:University Athletic Association 2031:Cannon v. University of Chicago 1918:George Herbert Jones Laboratory 1751:Center for Population Economics 1436: 1359: 1334: 1309: 1289: 1263: 495:Howard Hughes Medical Institute 394: 377:New England Journal of Medicine 353: 248: 230:Howard Hughes Medical Institute 1923:Gerald Ratner Athletics Center 1851:Toyota Technological Institute 1537:Marine Biological Laboratory, 1165: 1146: 1121: 1097: 843:, a slightly modified form of 1: 2056:University of Chicago sit-ins 1953:Joe and Rika Mansueto Library 1621:Woods Hole Historical Museum. 995: 970:Cold Spring Harbor Laboratory 888:, and ecosystem responses to 547: 533:, which breeds and maintains 508: 263:United States Fish Commission 2167:Contemporary Chamber Players 1816:Marine Biological Laboratory 1486:Embryology Course Centennial 499:National Academy of Sciences 234:National Academy of Sciences 207:Marine Biological Laboratory 20:Marine Biological Laboratory 7: 2277:Oceanographic organizations 1856:University of Chicago Press 1781:Comprehensive Cancer Center 1776:Committee on Social Thought 1766:Chicago school of economics 1736:Argonne National Laboratory 1710:Pritzker School of Medicine 1216:10.1056/NEJM197206082862308 958: 764:The MBL's contributions to 475: 453:Swope Building (residences) 10: 2318: 2272:Marine biological stations 1970:Lorado Taft Midway Studios 773:in 1936 rediscovered, the 734:Alejandro Sánchez Alvarado 243: 2302:Woods Hole, Massachusetts 2205: 2135: 2084: 2071:Old University of Chicago 2018: 1905: 1875: 1831:Obama Presidential Center 1771:Comer Children's Hospital 1741:Becker Friedman Institute 1728: 1672: 1665: 619:green fluorescent protein 531:National Xenopus Resource 501:; and 236 Members of the 236:; and 236 Members of the 215:Woods Hole, Massachusetts 187: 177: 169: 157: 142:41.5262222°N 70.6728889°W 108: 100: 90: 78: 64: 45: 33: 24: 2066:Metallurgical Laboratory 1685:Booth School of Business 562:imaging and computation; 480: 275:Annisquam, Massachusetts 2112:Lascivious Costume Ball 2051:Graduate Library School 1543:The Biological Bulletin 985:Statue of Rachel Carson 341:The Biological Bulletin 255:Spencer Fullerton Baird 147:41.5262222; -70.6728889 2117:Latke–Hamantash Debate 1928:Henry Crown Fieldhouse 1796:James Franck Institute 1786:Enrico Fermi Institute 825:Haldan Keffer Hartline 712:(1972–1974; 1988–1996) 367: 261:). Baird had set up a 80:Field of research 2282:University of Chicago 2136:Student organizations 2085:Sports and traditions 1806:Institute of Politics 1658:University of Chicago 1495:, Vol. S116, S63-S64. 1155:Embryology Class 1897 965:University of Chicago 953:University of Chicago 849:photoreceptor protein 588:Edmund Beecher Wilson 470:Huntington F. Willard 462:University of Chicago 441:C.V. Starr Laboratory 361: 219:University of Chicago 183:University of Chicago 1887:Burton–Judson Courts 1513:(1988) 174 (suppl.). 1448:college.uchicago.edu 861:Albert Szent-Györgyi 692:Frank Rattray Lillie 686:Charles Otis Whitman 584:Frank Rattray Lillie 329:Charles Otis Whitman 2192:Voices in Your Head 2005:Smart Museum of Art 1985:Renaissance Society 1958:John Crerar Library 1539:First Annual Report 1530:Maienschein, Jane. 1511:Biological Bulletin 1243:(1974), "The MBL", 980:Catherine N. Norton 818:squid giant synapse 580:Edwin Grant Conklin 271:Norwood-Hyatt House 138: /  21: 2239:School of Business 2222:School of Business 2187:Student Government 2149:The Chicago Maroon 2107:Women's basketball 2076:Yerkes Observatory 1995:Rockefeller Chapel 1963:Regenstein Library 1938:Hutchinson Commons 1811:Laboratory Schools 1277:. 27 November 2013 936:hydrothermal vents 900:, who studies the 871:Ecosystems science 853:Stephen W. Kuffler 783:Alan Lloyd Hodgkin 771:John Zachary Young 596:Thomas Hunt Morgan 402:Facilities in 2023 368: 295:Facilities in 1899 65:Research type 19: 2249: 2248: 1871: 1870: 1846:Stieglitz Lecture 1836:Paulson Institute 1516:Llinas, Rodolfo. 1210:(23): 1254–1256, 943:Education program 738:Richard Behringer 519:Ecosystems Center 370:Writing in 1972, 203: 202: 40:Lillie Laboratory 2309: 2209: 2102:Men's basketball 2022: 1975:Midway Plaisance 1879: 1670: 1669: 1659: 1650: 1643: 1636: 1627: 1626: 1505:Lillie, Frank R. 1458: 1457: 1455: 1454: 1440: 1434: 1433: 1431: 1430: 1416: 1405: 1404: 1402: 1401: 1387: 1381: 1380: 1378: 1377: 1363: 1357: 1356: 1354: 1352: 1338: 1332: 1331: 1329: 1327: 1313: 1307: 1306: 1304: 1303: 1293: 1287: 1286: 1284: 1282: 1267: 1261: 1260: 1249:, Bantam Books, 1237: 1228: 1227: 1196: 1190: 1189: 1184: 1182: 1169: 1163: 1162: 1150: 1144: 1143: 1141: 1139: 1125: 1119: 1118: 1116: 1115: 1101: 1095: 1094: 1092: 1091: 1077: 1068: 1067: 1065: 1064: 1050: 1041: 1040: 1038: 1037: 1023: 814:axonal transport 775:squid giant axon 769:discovered, and 728:Marianne Bronner 710:Eric H. Davidson 698:Viktor Hamburger 450: 438: 424: 412: 317: 305: 199: 196: 194: 153: 152: 150: 149: 148: 143: 139: 136: 135: 134: 131: 73:applied research 60: 58: 38: 29: 22: 18: 2317: 2316: 2312: 2311: 2310: 2308: 2307: 2306: 2262:1888 in biology 2252: 2251: 2250: 2245: 2207: 2201: 2162:CMAC (UChicago) 2131: 2080: 2020: 2014: 1980:Quadrangle Club 1901: 1892:Snell–Hitchcock 1878:Residence halls 1877: 1867: 1724: 1690:Divinity School 1661: 1657: 1654: 1615:Clapp, Pamela. 1600: 1467: 1465:Further reading 1462: 1461: 1452: 1450: 1442: 1441: 1437: 1428: 1426: 1418: 1417: 1408: 1399: 1397: 1389: 1388: 1384: 1375: 1373: 1367:"About the MBL" 1365: 1364: 1360: 1350: 1348: 1340: 1339: 1335: 1325: 1323: 1315: 1314: 1310: 1301: 1299: 1297:"About the MBL" 1295: 1294: 1290: 1280: 1278: 1269: 1268: 1264: 1257: 1238: 1231: 1197: 1193: 1180: 1178: 1171: 1170: 1166: 1152: 1151: 1147: 1137: 1135: 1127: 1126: 1122: 1113: 1111: 1103: 1102: 1098: 1089: 1087: 1079: 1078: 1071: 1062: 1060: 1052: 1051: 1044: 1035: 1033: 1027:"About the MBL" 1025: 1024: 1003: 998: 961: 945: 925:, are studied. 911: 902:biogeochemistry 873: 847:, coupled to a 762: 753: 677:research); and 615:Osamu Shimomura 600:Keith R. Porter 572: 550: 511: 489:As of 2022, 60 483: 478: 458: 457: 456: 455: 454: 451: 443: 442: 439: 430: 429: 428: 427:Loeb Laboratory 425: 417: 416: 415:Rowe Laboratory 413: 404: 403: 397: 356: 325: 324: 323: 322: 321: 318: 310: 309: 306: 297: 296: 283:Penikese Island 251: 246: 224:As of 2023, 60 191: 162: 146: 144: 140: 137: 132: 129: 127: 125: 124: 123: 81: 54: 52: 41: 17: 12: 11: 5: 2315: 2305: 2304: 2299: 2294: 2289: 2284: 2279: 2274: 2269: 2264: 2247: 2246: 2244: 2243: 2242: 2241: 2231: 2230: 2229: 2224: 2213: 2211: 2203: 2202: 2200: 2199: 2194: 2189: 2184: 2179: 2177:Off-Off Campus 2174: 2169: 2164: 2159: 2156:Chicago Review 2152: 2145: 2139: 2137: 2133: 2132: 2130: 2129: 2124: 2122:Scavenger Hunt 2119: 2114: 2109: 2104: 2099: 2094: 2088: 2086: 2082: 2081: 2079: 2078: 2073: 2068: 2063: 2058: 2053: 2048: 2047: 2046: 2043:Nuclear Energy 2037:Chicago Pile-1 2034: 2026: 2024: 2016: 2015: 2013: 2012: 2007: 2002: 2000:Seminary Co-op 1997: 1992: 1987: 1982: 1977: 1972: 1967: 1966: 1965: 1960: 1955: 1945: 1943:Ida Noyes Hall 1940: 1935: 1930: 1925: 1920: 1915: 1909: 1907: 1903: 1902: 1900: 1899: 1894: 1889: 1883: 1881: 1873: 1872: 1869: 1868: 1866: 1865: 1864: 1863: 1853: 1848: 1843: 1838: 1833: 1828: 1823: 1821:Medical Center 1818: 1813: 1808: 1803: 1798: 1793: 1788: 1783: 1778: 1773: 1768: 1763: 1758: 1753: 1748: 1743: 1738: 1732: 1730: 1726: 1725: 1723: 1722: 1717: 1712: 1707: 1702: 1697: 1692: 1687: 1682: 1676: 1674: 1667: 1663: 1662: 1653: 1652: 1645: 1638: 1630: 1624: 1623: 1613: 1607: 1599: 1598:External links 1596: 1595: 1594: 1580: 1563: 1546: 1535: 1528: 1514: 1503: 1496: 1489: 1482: 1466: 1463: 1460: 1459: 1435: 1406: 1382: 1358: 1333: 1308: 1288: 1262: 1255: 1229: 1191: 1164: 1145: 1133:Inside the MBL 1120: 1109:embryo.asu.edu 1096: 1069: 1042: 1000: 999: 997: 994: 993: 992: 990:Cornelia Clapp 987: 982: 977: 972: 967: 960: 957: 944: 941: 927:Mitchell Sogin 919:Microorganisms 910: 907: 890:global warming 886:nitrogen cycle 872: 869: 833:photoreception 804:Rodolfo Llinas 796:Clay Armstrong 761: 758: 752: 749: 748: 747: 746:(2020–present) 744:Carole LaBonne 741: 731: 725: 722:Michael Levine 719: 713: 707: 704:James D. Ebert 701: 695: 689: 650:Michael Sheetz 627:spindle fibers 592:Nettie Stevens 571: 568: 567: 566: 563: 560: 557: 549: 546: 527:Whitman Center 510: 507: 482: 479: 477: 474: 452: 445: 444: 440: 433: 432: 431: 426: 419: 418: 414: 407: 406: 405: 401: 400: 399: 398: 396: 393: 364:Cornelia Clapp 355: 352: 348:Gertrude Stein 336:Cornelia Clapp 319: 312: 311: 307: 300: 299: 298: 294: 293: 292: 291: 250: 247: 245: 242: 201: 200: 189: 185: 184: 181: 175: 174: 171: 167: 166: 163: 158: 155: 154: 110: 106: 105: 102: 98: 97: 94: 88: 87: 82: 79: 76: 75: 66: 62: 61: 47: 43: 42: 39: 31: 30: 15: 9: 6: 4: 3: 2: 2314: 2303: 2300: 2298: 2295: 2293: 2290: 2288: 2285: 2283: 2280: 2278: 2275: 2273: 2270: 2268: 2265: 2263: 2260: 2259: 2257: 2240: 2237: 2236: 2235: 2232: 2228: 2225: 2223: 2220: 2219: 2218: 2215: 2214: 2212: 2210: 2204: 2198: 2195: 2193: 2190: 2188: 2185: 2183: 2180: 2178: 2175: 2173: 2170: 2168: 2165: 2163: 2160: 2158: 2157: 2153: 2151: 2150: 2146: 2144: 2141: 2140: 2138: 2134: 2128: 2125: 2123: 2120: 2118: 2115: 2113: 2110: 2108: 2105: 2103: 2100: 2098: 2095: 2093: 2090: 2089: 2087: 2083: 2077: 2074: 2072: 2069: 2067: 2064: 2062: 2059: 2057: 2054: 2052: 2049: 2045: 2044: 2040: 2039: 2038: 2035: 2033: 2032: 2028: 2027: 2025: 2023: 2017: 2011: 2008: 2006: 2003: 2001: 1998: 1996: 1993: 1991: 1988: 1986: 1983: 1981: 1978: 1976: 1973: 1971: 1968: 1964: 1961: 1959: 1956: 1954: 1951: 1950: 1949: 1946: 1944: 1941: 1939: 1936: 1934: 1931: 1929: 1926: 1924: 1921: 1919: 1916: 1914: 1913:Bartlett Hall 1911: 1910: 1908: 1904: 1898: 1895: 1893: 1890: 1888: 1885: 1884: 1882: 1880: 1874: 1862: 1859: 1858: 1857: 1854: 1852: 1849: 1847: 1844: 1842: 1839: 1837: 1834: 1832: 1829: 1827: 1824: 1822: 1819: 1817: 1814: 1812: 1809: 1807: 1804: 1802: 1799: 1797: 1794: 1792: 1789: 1787: 1784: 1782: 1779: 1777: 1774: 1772: 1769: 1767: 1764: 1762: 1759: 1757: 1754: 1752: 1749: 1747: 1744: 1742: 1739: 1737: 1734: 1733: 1731: 1727: 1721: 1718: 1716: 1713: 1711: 1708: 1706: 1703: 1701: 1698: 1696: 1693: 1691: 1688: 1686: 1683: 1681: 1678: 1677: 1675: 1671: 1668: 1664: 1660: 1651: 1646: 1644: 1639: 1637: 1632: 1631: 1628: 1622: 1618: 1614: 1611: 1608: 1605: 1602: 1601: 1593: 1592:0-8122-8092-X 1589: 1585: 1581: 1579: 1578:0-691-04977-7 1575: 1571: 1567: 1566:Pauly, Philip 1564: 1562: 1561:0-19-504244-1 1558: 1554: 1550: 1549:Pauly, Philip 1547: 1544: 1540: 1536: 1533: 1529: 1527: 1526:0-19-511652-6 1523: 1519: 1515: 1512: 1508: 1504: 1502:181: 841-846. 1501: 1497: 1494: 1490: 1487: 1483: 1481: 1480:0-674-07403-3 1477: 1473: 1469: 1468: 1449: 1445: 1439: 1425: 1421: 1415: 1413: 1411: 1396: 1392: 1386: 1372: 1368: 1362: 1347: 1343: 1337: 1322: 1318: 1312: 1298: 1292: 1276: 1272: 1266: 1258: 1256:0-553-13972-X 1252: 1248: 1247: 1242: 1241:Thomas, Lewis 1236: 1234: 1225: 1221: 1217: 1213: 1209: 1205: 1201: 1200:Thomas, Lewis 1195: 1188: 1176: 1175: 1168: 1161: 1157: 1156: 1149: 1134: 1130: 1124: 1110: 1106: 1100: 1086: 1082: 1076: 1074: 1059: 1055: 1049: 1047: 1032: 1028: 1022: 1020: 1018: 1016: 1014: 1012: 1010: 1008: 1006: 1001: 991: 988: 986: 983: 981: 978: 976: 973: 971: 968: 966: 963: 962: 956: 954: 949: 940: 937: 932: 928: 924: 920: 916: 906: 903: 899: 898:Jerry Melillo 895: 891: 887: 883: 879: 868: 866: 862: 858: 854: 850: 846: 842: 838: 834: 830: 826: 821: 819: 815: 811: 810: 805: 801: 797: 792: 791:voltage clamp 788: 787:Andrew Huxley 784: 780: 776: 772: 767: 757: 745: 742: 739: 735: 732: 729: 726: 723: 720: 717: 714: 711: 708: 705: 702: 699: 696: 693: 690: 687: 684: 683: 682: 680: 676: 672: 666: 663: 662:Avram Hershko 659: 658:James Spudich 655: 651: 647: 643: 639: 638:Joan Ruderman 635: 630: 628: 624: 620: 616: 612: 607: 605: 601: 597: 593: 589: 585: 581: 577: 564: 561: 558: 555: 554: 553: 545: 543: 538: 536: 532: 528: 524: 520: 516: 506: 504: 500: 496: 492: 487: 473: 471: 466: 463: 449: 437: 423: 411: 392: 389: 385: 384: 379: 378: 373: 365: 360: 351: 349: 345: 343: 342: 337: 332: 330: 316: 308:The residence 304: 290: 288: 284: 280: 279:Louis Agassiz 276: 272: 268: 267:Alpheus Hyatt 264: 260: 256: 241: 239: 235: 231: 227: 222: 220: 216: 212: 208: 198: 190: 186: 182: 180: 176: 172: 168: 164: 161: 156: 151: 133:70°40′22.40″W 130:41°31′34.40″N 122: 121:United States 118: 117:Massachusetts 114: 111: 107: 103: 99: 95: 93: 89: 86: 83: 77: 74: 70: 67: 63: 57: 51: 48: 44: 37: 32: 28: 23: 2154: 2147: 2041: 2029: 1815: 1620: 1583: 1569: 1552: 1542: 1538: 1531: 1517: 1510: 1506: 1499: 1492: 1485: 1471: 1451:. 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Index



1888
1888
Pure
applied research
Biology
Director
Woods Hole
Massachusetts
United States
41°31′34.40″N 70°40′22.40″W / 41.5262222°N 70.6728889°W / 41.5262222; -70.6728889
ZIP code
Affiliations
www.mbl.edu
Woods Hole, Massachusetts
University of Chicago
Nobel Prize
Howard Hughes Medical Institute
National Academy of Sciences
American Academy of Arts and Sciences
Spencer Fullerton Baird
fisheries
United States Fish Commission
Alpheus Hyatt
Norwood-Hyatt House
Annisquam, Massachusetts
Louis Agassiz
Penikese Island
Boston Society of Natural History

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