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Icaronycteris

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measured about 14 centimetres (5.5 in) long and had a wingspan of 37 centimetres (15 in). It closely resembled modern bats, but had some primitive traits. The tail was much longer and not connected to the hind legs with a skin membrane, the first wing finger bore a claw and the body was
320:, making it the earliest bat genus known from complete skeletons, and the earliest known bat from North America. Multiple exceptionally preserved specimens, among the best preserved bat fossils, are known from the 206: 415: 1295: 914:
Rietbergen, Tim B.; Ostende, Lars W. van den Hoek; Aase, Arvid; Jones, Matthew F.; Medeiros, Edward D.; Simmons, Nancy B. (12 April 2023).
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The Simon & Schuster Encyclopedia of Dinosaurs and Prehistoric Creatures: A Visual Who's Who of Prehistoric Life
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to extant bat lineages, with comments on the evolution of echolocation and foraging strategies in Microchiroptera"
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more flexible. Similarly, it had a full set of relatively unspecialised teeth, similar to those of a modern
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Russell, D. E.; Louis, P.; Savage, D. E. (1973). "Chiroptera and Dermoptera of the French early Eocene".
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slept while hanging upside down, holding onto a tree branch or stone ridge with its hind legs.
1300: 1170: 1164: 916:"The oldest known bat skeletons and their implications for Eocene chiropteran diversification" 1321: 1313: 321: 1354: 1269: 1009: 996:
Smith, T.; Rana, R. S.; Missiaen, P.; Rose, K. D.; Sahni, A.; Singh, H.; Singh, L. (2007).
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were proposed to be removed from the genus due to them not being closely related.
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is based on well-preserved fragments of dentaries and lower teeth found in
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The Marshall Illustrated Encyclopedia of Dinosaurs and Prehistoric Animals
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Fragmentary material from France has also been tentatively placed within
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Cox, Barry; Harrison, Colin; Savage, R. J. G.; Gardiner, Brian (1999).
81: 46: 315: 153: 86: 1231: 1254: 751: 173: 76: 71: 56: 51: 41: 91: 66: 1274: 746: 310: 183: 163: 998:"High bat (Chiroptera) diversity in the Early Eocene of India" 384: 977:
University of California Publications in Geological Sciences
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also from the Green River Formation was distinguished from
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Naish, Darren; Wyse, Liz (2001). Parsons, Jayne (ed.).
425:, in a series leading to extant microchiropteran bats. 1169:. Smithsonian Handbooks. Dorling Kindersley. p.  995: 733: 286: 628: 278: 265: 253: 241: 974: 273: 678: 676: 1341: 782: 387:. Its anatomy suggests that, like modern bats, 1063: 816: 776: 721: 682: 673: 437:to be sister to the North American species of 400: 1046: 800: 788: 324:of North America. The best known species is 111:Cast of Icaronycteris index holotype at the 1162: 791:, Discussion of Phylogenetic Relationships 119: 102: 1137: 1029: 957: 939: 858:(1966). "Early Eocene bat from Wyoming". 845: 704: 819:"Fossil evidence and the origin of bats" 367: 1166:Dinosaurs and Other Prehistoric Animals 1064:Simmons, N. B.; Geisler, J. H. (1998). 817:Gunnell, G. F.; Simmons, N. B. (2005). 1342: 1203: 894: 854: 764: 745:The name relates the mythic flight of 710: 260: 226: 1236: 1235: 794: 695: 693: 691: 1047:Simmons, N. B.; Conway, T. (1998). 758: 755:, the genus of "hollow-faced bats". 715: 13: 1097: 113:American Museum of Natural History 14: 1381: 688: 631: 407:is the first genus, followed by 274:Russell, Louis & Savage 1973 140: 34: 1365:Eocene mammals of North America 1066:"Phylogenetic relationships of 1208:. Welcome Books. p. 119. 826:Journal of Mammalian Evolution 770: 739: 727: 363: 1: 1360:Fossil taxa described in 1966 899:. London: Marshall Editions. 880:10.1126/science.154.3754.1333 661: 941:10.1371/journal.pone.0283505 394: 7: 624: 10: 1386: 1163:Richardson, Hazel (2003). 809: 777:Simmons & Geisler 1998 722:Simmons & Geisler 1998 683:Gunnell & Simmons 2005 401:Simmons & Geisler 1998 1244: 1206:The Big Book Of Dinosaurs 1022:10.1007/s00114-007-0280-9 838:10.1007/s10914-005-6945-2 801:Simmons & Conway 1998 789:Simmons & Conway 1998 560: 542: 534: 526: 508: 500: 482: 474: 458: 450: 238: 233: 137:Scientific classification 135: 127: 118: 110: 101: 23: 895:Palmer, D., ed. (1999). 666: 309:that lived in the early 344:. In 2023, the species 318: million years ago 301:is an extinct genus of 1350:Prehistoric bat genera 1204:Norman, David (2001). 700:Rietbergen et al. 2023 429:Rietbergen et al. 2023 376: 332:as the second species 249:Rietbergen et al. 2023 245:Icaronycteris gunnelli 1370:Green River Formation 1322:Paleobiology Database 1141:Dinosaur Encyclopedia 685:, Fossil Bats, p. 214 371: 322:Green River Formation 128:Holotype specimen of 16:Extinct genus of bats 1113:Simon & Schuster 1089:Bulletin of the AMNH 565:(Echolocating bats) 1014:2007NW.....94.1003S 1002:Naturwissenschaften 932:2023PLoSO..1883505R 872:1966Sci...154.1333J 639:Paleontology portal 282:Icaronycteris sigei 269:Icaronycteris menui 257:Icaronycteris index 1146:Dorling Kindersley 377: 1337: 1336: 1309:Open Tree of Life 1238:Taxon identifiers 1122:978-0-684-86411-2 1080:Palaeochiropteryx 1008:(12): 1003–1009. 906:978-1-84028-152-1 734:Smith et al. 2007 619: 618: 610: 609: 601: 600: 592: 591: 583: 582: 574: 573: 546:Palaeochiropteryx 422:Palaeochiropteryx 294: 293: 287:Smith et al. 2007 229: 1377: 1330: 1329: 1317: 1316: 1304: 1303: 1291: 1290: 1278: 1277: 1265: 1264: 1263: 1233: 1232: 1227: 1200: 1159: 1134: 1092: 1086: 1060: 1058: 1056: 1043: 1033: 992: 971: 961: 943: 910: 891: 866:(3754): 1333–9. 851: 849: 832:(1–2): 209–246. 823: 803: 798: 792: 786: 780: 774: 768: 762: 756: 743: 737: 731: 725: 719: 713: 708: 702: 697: 686: 680: 655:Australonycteris 641: 636: 635: 634: 537: 536: 529: 528: 503: 502: 477: 476: 453: 452: 445: 444: 319: 313:, approximately 303:microchiropteran 280: 267: 255: 243: 225: 218: 205: 145: 144: 123: 106: 96: 33: 29:Temporal range: 21: 20: 1385: 1384: 1380: 1379: 1378: 1376: 1375: 1374: 1340: 1339: 1338: 1333: 1325: 1320: 1312: 1307: 1299: 1294: 1286: 1281: 1273: 1268: 1259: 1258: 1253: 1240: 1230: 1216: 1181: 1156: 1123: 1100: 1098:Further reading 1095: 1084: 1072:Archaeonycteris 1054: 1052: 926:(4): e0283505. 907: 821: 812: 807: 806: 799: 795: 787: 783: 775: 771: 763: 759: 744: 740: 732: 728: 720: 716: 709: 705: 698: 689: 681: 674: 669: 664: 637: 632: 630: 627: 620: 611: 602: 593: 584: 575: 563:Microchiroptera 512:Archaeonycteris 410:Archaeonycteris 397: 372:Restoration of 366: 314: 305:(echolocating) 285: 272: 259: 247: 224: 216: 207:Icaronycteridae 203: 139: 97: 95: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 39: 31: 30: 27: 17: 12: 11: 5: 1383: 1373: 1372: 1367: 1362: 1357: 1352: 1335: 1334: 1332: 1331: 1318: 1305: 1292: 1279: 1266: 1250: 1248: 1242: 1241: 1229: 1228: 1215:978-0941807487 1214: 1201: 1180:978-0789493613 1179: 1160: 1154: 1135: 1121: 1101: 1099: 1096: 1094: 1093: 1076:Hassianycteris 1061: 1051:. Tree of Life 1044: 1031:1854/LU-385394 993: 972: 911: 905: 892: 852: 813: 811: 808: 805: 804: 793: 781: 769: 757: 738: 726: 714: 703: 687: 671: 670: 668: 665: 663: 660: 659: 658: 651: 648:Onychonycteris 643: 642: 626: 623: 617: 616: 613: 612: 608: 607: 604: 603: 599: 598: 595: 594: 590: 589: 586: 585: 581: 580: 577: 576: 572: 571: 568: 567: 559: 555: 554: 551: 550: 541: 535: 533: 527: 525: 521: 520: 517: 516: 507: 501: 499: 495: 494: 491: 490: 481: 475: 473: 469: 468: 465: 464: 461:Megachiroptera 457: 451: 449: 443: 434:Onychonycteris 416:Hassianycetris 396: 393: 365: 362: 292: 291: 290: 289: 276: 263: 251: 236: 235: 231: 230: 214: 210: 209: 201: 197: 196: 191: 187: 186: 181: 177: 176: 171: 167: 166: 161: 157: 156: 151: 147: 146: 133: 132: 125: 124: 116: 115: 108: 107: 99: 98: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 40: 35: 28: 15: 9: 6: 4: 3: 2: 1382: 1371: 1368: 1366: 1363: 1361: 1358: 1356: 1353: 1351: 1348: 1347: 1345: 1328: 1323: 1319: 1315: 1310: 1306: 1302: 1297: 1293: 1289: 1284: 1280: 1276: 1271: 1267: 1262: 1256: 1252: 1251: 1249: 1247: 1246:Icaronycteris 1243: 1239: 1234: 1225: 1221: 1217: 1211: 1207: 1202: 1198: 1194: 1190: 1186: 1182: 1176: 1172: 1168: 1167: 1161: 1157: 1155:9780789479358 1151: 1147: 1143: 1142: 1136: 1132: 1128: 1124: 1118: 1114: 1110: 1109: 1103: 1102: 1090: 1083: 1081: 1077: 1073: 1069: 1068:Icaronycteris 1062: 1050: 1045: 1041: 1037: 1032: 1027: 1023: 1019: 1015: 1011: 1007: 1003: 999: 994: 990: 986: 982: 978: 973: 969: 965: 960: 955: 951: 947: 942: 937: 933: 929: 925: 921: 917: 912: 908: 902: 898: 893: 889: 885: 881: 877: 873: 869: 865: 861: 857: 856:Jepsen, G. L. 853: 848: 847:2027.42/44972 843: 839: 835: 831: 827: 820: 815: 814: 802: 797: 790: 785: 778: 773: 767:, p. 211 766: 761: 754: 753: 748: 742: 735: 730: 723: 718: 712: 707: 701: 696: 694: 692: 684: 679: 677: 672: 657: 656: 652: 650: 649: 645: 644: 640: 629: 622: 615: 614: 606: 605: 597: 596: 588: 587: 579: 578: 570: 569: 566: 564: 557: 556: 553: 552: 549: 548: 547: 539: 538: 531: 530: 523: 522: 519: 518: 515: 514: 513: 505: 504: 497: 496: 493: 492: 489: 488: 487: 486:Icaronycteris 479: 478: 471: 470: 467: 466: 463: 462: 455: 454: 447: 446: 442: 440: 439:Icaronycteris 436: 435: 430: 426: 424: 423: 418: 417: 412: 411: 406: 405:Icaronycteris 402: 399:According to 392: 390: 389:Icaronycteris 386: 381: 380:Icaronycteris 375: 370: 361: 359: 355: 351: 347: 343: 342:Western India 339: 335: 331: 330:Icaronycteris 327: 323: 317: 312: 308: 304: 300: 299: 298:Icaronycteris 288: 283: 277: 275: 270: 264: 262: 258: 252: 250: 246: 240: 239: 237: 232: 228: 223: 222: 221:Icaronycteris 215: 212: 211: 208: 202: 199: 198: 195: 192: 189: 188: 185: 182: 179: 178: 175: 172: 169: 168: 165: 162: 159: 158: 155: 152: 149: 148: 143: 138: 134: 131: 126: 122: 117: 114: 109: 105: 100: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 38: 26: 25:Icaronycteris 22: 19: 1245: 1205: 1165: 1140: 1107: 1088: 1079: 1075: 1071: 1067: 1053:. Retrieved 1049:"Chiroptera" 1005: 1001: 980: 976: 923: 919: 896: 863: 859: 829: 825: 796: 784: 772: 760: 750: 741: 729: 724:, p. 40 717: 706: 653: 646: 621: 561: 544: 543: 510: 509: 485: 484: 483: 459: 438: 432: 427: 420: 414: 408: 404: 398: 388: 379: 378: 373: 357: 353: 349: 345: 337: 333: 329: 325: 297: 296: 295: 281: 268: 256: 244: 220: 219: 129: 32:Early Eocene 24: 18: 1355:Eocene bats 1055:1 September 765:Palmer 1999 711:Jepsen 1966 364:Description 346:I. gunnelli 261:Jepsen 1966 227:Jepsen 1966 130:I. gunnelli 1344:Categories 779:, Abstract 736:, Abstract 662:References 194:Chiroptera 989:691130847 950:1932-6203 395:Phylogeny 326:I. index. 160:Kingdom: 154:Eukaryota 1261:Q1062126 1255:Wikidata 1224:47522057 1197:8156728M 1189:52532157 1131:40943525 1040:17671774 983:: 1–57. 968:37043445 959:10096270 920:PLOS ONE 888:17770307 752:Nycteris 625:See also 374:I. index 358:I. sigei 354:I. menui 350:I. index 338:I. sigei 334:I. menui 234:Species 200:Family: 184:Mammalia 174:Chordata 170:Phylum: 164:Animalia 150:Domain: 1314:3614170 1301:1204623 1288:4832028 1275:4440253 1010:Bibcode 928:Bibcode 868:Bibcode 860:Science 810:Sources 352:, and 213:Genus: 190:Order: 180:Class: 1222:  1212:  1195:  1187:  1177:  1152:  1129:  1119:  1091:(235). 1078:, and 1038:  987:  966:  956:  948:  903:  886:  747:Icarus 431:found 419:, and 311:Eocene 1327:40632 1296:IRMNG 1085:(PDF) 822:(PDF) 667:Notes 385:shrew 1283:GBIF 1220:OCLC 1210:ISBN 1185:OCLC 1175:ISBN 1150:ISBN 1127:OCLC 1117:ISBN 1057:2014 1036:PMID 985:OCLC 964:PMID 946:ISSN 901:ISBN 884:PMID 356:and 316:52.2 37:Preκž’ 1270:EoL 1171:170 1026:hdl 1018:doi 954:PMC 936:doi 876:doi 864:154 842:hdl 834:doi 749:to 307:bat 1346:: 1324:: 1311:: 1298:: 1285:: 1272:: 1257:: 1218:. 1193:OL 1191:. 1183:. 1173:. 1148:. 1144:. 1125:. 1115:. 1111:. 1087:. 1074:, 1070:, 1034:. 1024:. 1016:. 1006:94 1004:. 1000:. 981:95 979:. 962:. 952:. 944:. 934:. 924:18 922:. 918:. 882:. 874:. 862:. 840:. 830:12 828:. 824:. 690:^ 675:^ 441:. 413:, 403:, 336:. 87:Pg 1226:. 1199:. 1158:. 1133:. 1059:. 1042:. 1028:: 1020:: 1012:: 991:. 970:. 938:: 930:: 909:. 890:. 878:: 870:: 850:. 844:: 836:: 284:? 279:† 271:? 266:† 254:† 242:† 217:† 204:† 92:N 82:K 77:J 72:T 67:P 62:C 57:D 52:S 47:O 42:κž’

Index

Preκž’
κž’
O
S
D
C
P
T
J
K
Pg
N

American Museum of Natural History

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Chiroptera
Icaronycteridae
Icaronycteris
Jepsen 1966
Rietbergen et al. 2023
Jepsen 1966
Russell, Louis & Savage 1973
Smith et al. 2007
microchiropteran

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