129:
147:
115:
397:(schizomids), which has been recovered subsequently in a number of other phylogenies to include fossils. All share the character of a subdivided tarsus (or foot) which gives the group its name. A 2014 analysis found that
667:
460:" (= foot) and refers to its quite long front pair of legs with their subdivided tips which look as though they might have been used to 'feel' their way around in front of the animal.
609:
1254:
629:
1396:
378:
Revisions have confirmed that
Haptopoda should be treated as a separate and independent order. A 2007 study tentatively recognised a group named
796:
Garwood, Russell J.; Dunlop, Jason A.; Selden, Paul A.; Spencer, Alan R. T.; Atwood, Robert C.; Vo, Nghia T.; Drakopoulos, Michael (2016).
1460:
637:
1470:
1383:
545:
Dunlop, Jason A. (2011). "A redescription of the
Carboniferous arachnid Plesiosiro madeleyi Pocock, 1911 (Arachnida: Haptopoda)".
1480:
863:
1455:
666:
Wang, Bo; Dunlop, Jason A.; Selden, Paul A.; Garwood, Russell J.; Shear, William A.; Mรผller, Patrick; Lei, Xiaojie (2018).
902:
1475:
1450:
1465:
1427:
1361:
867:
339:
in his important 1911 monograph on
British Carboniferous arachnids. It is the only known member of the order
350:
in 1949 and Dunlop in 1999. A supposed example from the Coal
Measures of Lancashire is a misidentification.
871:
146:
17:
895:
855:
604:
442:
347:
1414:
1401:
1323:
1409:
580:
375:). These harvestmen do, in some ways, resemble the reconstructed body plan of the haptopods.
336:
294:
275:
251:
229:
590:
1370:
742:
679:
8:
405:
consisting of two elements, a basal paturon and a fang, which suggests a position within
746:
683:
888:
832:
797:
773:
730:
711:
517:
482:
417:
was found to be the most basal member of a clade containing
Amblypygi, Schizomida, and
141:
1214:
837:
819:
778:
760:
703:
695:
668:"Cretaceous arachnid Chimerarachne yingi gen. et sp. nov. illuminates spider origins"
651:
584:
562:
522:
504:
410:
363:
1297:
827:
809:
768:
750:
715:
687:
646:
554:
512:
494:
729:
Garwood, Russell J.; Dunlop, Jason A.; Knecht, Brian J.; Hegna, Thomas A. (2017).
630:"A phylogenetic analysis of the arachnid orders based on morphological characters"
483:"Three-dimensional reconstruction and the phylogeny of extinct chelicerate orders"
1302:
1206:
1196:
1191:
1051:
1026:
961:
406:
326:
211:
128:
755:
691:
558:
1444:
823:
764:
699:
566:
508:
379:
368:
322:
314:
72:
798:"Almost a spider: a 305-million-year-old fossil arachnid and spider origins"
441:
The order has also been called
Haptopodida; the ending -ida originated when
1346:
1232:
841:
814:
782:
707:
526:
1355:
1219:
1183:
936:
188:
114:
47:
1388:
1110:
969:
586:
A Monograph of the
Terrestrial Carboniferous Arachnida of Great Britain
430:
402:
394:
92:
57:
499:
1151:
1088:
1034:
999:
982:
930:
386:
372:
310:
178:
158:
97:
1317:
1340:
1076:
1007:
911:
880:
198:
87:
82:
67:
62:
52:
1422:
1375:
1118:
589:. Monographs of the Palaeontographical Society, London. pp.
418:
390:
318:
102:
77:
445:(1955) tried to standardize the endings of the arachnid orders.
1063:
924:
426:
168:
547:
Transactions of the Royal
Society of Edinburgh: Earth Sciences
1164:
610:
Transactions of the
Connecticut Academy of Arts and Sciences
1224:
313:
genus known exclusively from nine specimens from the Upper
795:
329:. The genus is monotypic, represented only by the species
728:
346:
The original fossils have been redescribed in detail by
854:
802:
Proceedings of the Royal
Society B: Biological Sciences
665:
603:
371:); the most primitive group of the living harvestmen (
579:
476:
474:
472:
627:
452:" (= tangible, subject to the sense of touch) + "
1442:
789:
722:
469:
896:
659:
480:
540:
538:
536:
425:), which was sister to the clade containing
858:(1955). "Arachnida". In R. C. Moore (ed.).
607:(1949). "A study of Palaeozoic Arachnida".
481:Garwood, Russell J.; Dunlop, Jason (2014).
903:
889:
127:
113:
831:
813:
772:
754:
650:
638:Zoological Journal of the Linnean Society
533:
516:
498:
367:", which is a genus of cyphophthalmid (
14:
1443:
731:"The phylogeny of fossil whip spiders"
544:
1322:
1321:
1281:
949:
884:
864:Treatise on Invertebrate Paleontology
1428:5C6CDEFD-D48F-4FCB-B58D-1109ABC951CA
910:
24:
1461:Carboniferous arthropods of Europe
25:
1492:
448:Haptopoda originates from Greek "
1262:are possibly paraphyletic groups
652:10.1111/j.1096-3642.2007.00284.x
353:
145:
45:
1471:Pennsylvanian first appearances
848:
672:Nature Ecology & Evolution
621:
597:
573:
13:
1:
1481:Fossil taxa described in 1911
868:Geological Society of America
463:
1253:Classification is based on
7:
1456:Prehistoric arachnid genera
1282:
1113:(shorttailed whipscorpions)
10:
1497:
950:
872:University of Kansas Press
628:Jeffrey W. Shultz (2007).
27:Extinct genus of arachnids
1476:Pennsylvanian extinctions
1330:
1288:
1277:
1249:
1205:
1182:
1163:
1147:
1138:
1100:
1091:(tailless whip scorpions)
1084:
1075:
1059:
1050:
1019:
992:
981:
960:
956:
945:
919:
756:10.1186/s12862-017-0931-1
692:10.1038/s41559-017-0449-3
559:10.1017/S0263593300002492
281:
274:
142:Scientific classification
140:
135:
126:
121:
112:
34:
735:BMC Evolutionary Biology
436:
1451:Carboniferous arachnids
856:Alexander Petrunkevitch
605:Alexander Petrunkevitch
409:. Under both equal and
348:Alexander Petrunkevitch
815:10.1098/rspb.2016.0125
385:comprising Haptopoda,
1466:Carboniferous England
1410:Paleobiology Database
972:(microwhip scorpions)
581:Reginald Innes Pocock
393:(whip scorpions) and
337:Reginald Innes Pocock
1154:(hooded tickspiders)
860:Part P, Arthropoda 2
1362:Plesiosiro madeleyi
1332:Plesiosiro madeleyi
747:2017BMCEE..17..105G
684:2018NatEE...2..614W
332:Plesiosiro madeleyi
288:Plesiosiro madeleyi
43:Upper Carboniferous
1027:Pseudoscorpionida
874:. pp. 42โ162.
808:(1827): 20160125.
1438:
1437:
1324:Taxon identifiers
1315:
1314:
1311:
1310:
1273:
1272:
1269:
1268:
1245:
1244:
1241:
1240:
1228:
1215:Opilioacariformes
1175:
1155:
1134:
1133:
1130:
1129:
1122:
1114:
1092:
1067:
1046:
1045:
1038:
1030:
1029:(pseudoscorpions)
1011:
1003:
973:
500:10.7717/peerj.641
384:
302:
301:
255:
233:
136:Life restoration
16:(Redirected from
1488:
1431:
1430:
1418:
1417:
1405:
1404:
1392:
1391:
1379:
1378:
1366:
1365:
1364:
1351:
1350:
1349:
1319:
1318:
1298:Phalangiotarbida
1279:
1278:
1261:
1226:
1180:
1179:
1174:
1173:
1169:
1167:
1153:
1145:
1144:
1141:
1120:
1112:
1090:
1082:
1081:
1065:
1057:
1056:
1036:
1028:
1009:
1001:
990:
989:
985:
971:
958:
957:
947:
946:
905:
898:
891:
882:
881:
876:
875:
852:
846:
845:
835:
817:
793:
787:
786:
776:
758:
726:
720:
719:
663:
657:
656:
654:
634:
625:
619:
618:
601:
595:
594:
577:
571:
570:
542:
531:
530:
520:
502:
478:
389:(whip spiders),
382:
361:means "close to
290:
286:
267:P. madeleyi
250:
243:
228:
223:
150:
149:
131:
117:
107:
44:
40:Temporal range:
32:
31:
21:
1496:
1495:
1491:
1490:
1489:
1487:
1486:
1485:
1441:
1440:
1439:
1434:
1426:
1421:
1413:
1408:
1400:
1395:
1387:
1382:
1374:
1369:
1360:
1359:
1354:
1345:
1344:
1339:
1326:
1316:
1307:
1284:
1265:
1259:
1237:
1201:
1171:
1170:
1168:
1165:
1159:
1139:
1126:
1096:
1071:
1042:
1037:(camel spiders)
1015:
983:
977:
952:
941:
915:
909:
879:
853:
849:
794:
790:
727:
723:
664:
660:
632:
626:
622:
602:
598:
578:
574:
543:
534:
479:
470:
466:
439:
411:implied weights
356:
298:
292:
284:
283:
270:
249:
241:
227:
221:
144:
108:
106:
105:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
42:
41:
38:
28:
23:
22:
15:
12:
11:
5:
1494:
1484:
1483:
1478:
1473:
1468:
1463:
1458:
1453:
1436:
1435:
1433:
1432:
1419:
1406:
1393:
1380:
1367:
1352:
1336:
1334:
1328:
1327:
1313:
1312:
1309:
1308:
1306:
1305:
1303:Trigonotarbida
1300:
1295:
1289:
1286:
1285:
1275:
1274:
1271:
1270:
1267:
1266:
1264:
1263:
1260:Items in green
1257:
1250:
1247:
1246:
1243:
1242:
1239:
1238:
1236:
1235:
1230:
1222:
1217:
1211:
1209:
1207:Parasitiformes
1203:
1202:
1200:
1199:
1197:Trombidiformes
1194:
1192:Sarcoptiformes
1188:
1186:
1177:
1172:(ticks, mites)
1161:
1160:
1158:
1157:
1148:
1142:
1136:
1135:
1132:
1131:
1128:
1127:
1125:
1124:
1116:
1107:
1105:
1098:
1097:
1095:
1094:
1085:
1079:
1073:
1072:
1070:
1069:
1060:
1054:
1052:Tetrapulmonata
1048:
1047:
1044:
1043:
1041:
1040:
1032:
1023:
1021:
1017:
1016:
1014:
1013:
1005:
996:
994:
987:
979:
978:
976:
975:
966:
964:
962:Incertae sedis
954:
953:
943:
942:
940:
939:
933:
927:
920:
917:
916:
908:
907:
900:
893:
885:
878:
877:
847:
788:
721:
678:(4): 614โ622.
658:
645:(2): 221โ265.
620:
596:
572:
532:
467:
465:
462:
438:
435:
421:(Thelyphonida
407:Tetrapulmonata
355:
352:
327:United Kingdom
309:is an extinct
300:
299:
293:
279:
278:
272:
271:
263:
261:
257:
256:
239:
235:
234:
219:
215:
214:
212:Tetrapulmonata
209:
202:
201:
196:
192:
191:
186:
182:
181:
176:
172:
171:
166:
162:
161:
156:
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151:
138:
137:
133:
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123:
119:
118:
110:
109:
101:
96:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
39:
26:
9:
6:
4:
3:
2:
1493:
1482:
1479:
1477:
1474:
1472:
1469:
1467:
1464:
1462:
1459:
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1429:
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1420:
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1411:
1407:
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1377:
1372:
1368:
1363:
1357:
1353:
1348:
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1333:
1329:
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1320:
1304:
1301:
1299:
1296:
1294:
1291:
1290:
1287:
1280:
1276:
1258:
1256:
1255:Shultz (2007)
1252:
1251:
1248:
1234:
1231:
1229:
1223:
1221:
1218:
1216:
1213:
1212:
1210:
1208:
1204:
1198:
1195:
1193:
1190:
1189:
1187:
1185:
1181:
1178:
1176:
1162:
1156:
1150:
1149:
1146:
1143:
1137:
1123:
1121:(vinegaroons)
1117:
1115:
1109:
1108:
1106:
1104:
1101:Thelyphonida
1099:
1093:
1087:
1086:
1083:
1080:
1078:
1074:
1068:
1062:
1061:
1058:
1055:
1053:
1049:
1039:
1033:
1031:
1025:
1024:
1022:
1018:
1012:
1006:
1004:
998:
997:
995:
991:
988:
986:
980:
974:
968:
967:
965:
963:
959:
955:
948:
944:
938:
934:
932:
928:
926:
922:
921:
918:
913:
906:
901:
899:
894:
892:
887:
886:
883:
873:
869:
865:
861:
857:
851:
843:
839:
834:
829:
825:
821:
816:
811:
807:
803:
799:
792:
784:
780:
775:
770:
766:
762:
757:
752:
748:
744:
740:
736:
732:
725:
717:
713:
709:
705:
701:
697:
693:
689:
685:
681:
677:
673:
669:
662:
653:
648:
644:
640:
639:
631:
624:
616:
612:
611:
606:
600:
592:
588:
587:
582:
576:
568:
564:
560:
556:
552:
548:
541:
539:
537:
528:
524:
519:
514:
510:
506:
501:
496:
492:
488:
484:
477:
475:
473:
468:
461:
459:
455:
451:
446:
444:
443:Petrunkevitch
434:
432:
428:
424:
420:
416:
412:
408:
404:
400:
396:
392:
388:
381:
380:Schizotarsata
376:
374:
370:
369:Cyphophthalmi
366:
365:
360:
354:Relationships
351:
349:
344:
342:
338:
335:described by
334:
333:
328:
324:
323:Staffordshire
320:
316:
315:Carboniferous
312:
308:
307:
296:
291:
289:
280:
277:
276:Binomial name
273:
269:
268:
262:
259:
258:
253:
248:
247:
240:
237:
236:
231:
226:
220:
217:
216:
213:
210:
207:
204:
203:
200:
197:
194:
193:
190:
187:
184:
183:
180:
177:
174:
173:
170:
167:
164:
163:
160:
157:
154:
153:
148:
143:
139:
134:
130:
125:
120:
116:
111:
104:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
37:
33:
30:
19:
1331:
1292:
1233:Mesostigmata
1102:
1020:Haplocnemata
1002:(harvestmen)
993:Stomothecata
866:. Lawrence:
859:
850:
805:
801:
791:
738:
734:
724:
675:
671:
661:
642:
636:
623:
614:
608:
599:
585:
575:
553:(1): 29โ47.
550:
546:
490:
486:
457:
453:
449:
447:
440:
422:
414:
398:
383:Shultz, 2007
377:
362:
358:
357:
345:
340:
331:
330:
305:
304:
303:
287:
282:
266:
265:
245:
244:
224:
205:
35:
29:
1356:Wikispecies
1220:Holothyrida
1184:Acariformes
1140:Acaromorpha
1111:Schizomida
1010:(scorpions)
1008:Scorpiones
970:Palpigradi
937:Chelicerata
935:Subphylum:
189:Chelicerata
185:Subphylum:
1445:Categories
1152:Ricinulei
1089:Amblypygi
1035:Solifugae
1000:Opiliones
931:Arthropoda
741:(1): 105.
464:References
431:Uraraneida
403:chelicerae
399:Plesiosiro
395:Schizomida
359:Plesiosiro
306:Plesiosiro
246:Plesiosiro
179:Arthropoda
36:Plesiosiro
1293:Haptopoda
1077:Pedipalpi
1066:(spiders)
984:Dromopoda
912:Arachnida
824:0962-8452
765:1471-2148
700:2397-334X
617:: 69โ315.
567:0263-5933
509:2167-8359
415:Plesisiro
387:Amblypygi
373:Opiliones
341:Haptopoda
260:Species:
225:Haptopoda
199:Arachnida
165:Kingdom:
159:Eukaryota
18:Haptopoda
1402:11671678
1376:10577876
1341:Wikidata
1225:Ixodida
1119:Uropygi
1064:Araneae
929:Phylum:
925:Animalia
923:Kingdom
842:27030415
783:28431496
708:29403075
583:(1911).
527:25405073
493:: e641.
311:arachnid
175:Phylum:
169:Animalia
155:Domain:
122:CT scan
1423:ZooBank
1389:6895181
1283:Extinct
1227:(ticks)
833:4822468
774:5399839
743:Bibcode
716:4239867
680:Bibcode
518:4232842
427:Araneae
419:Uropygi
391:Uropygi
319:Coseley
285:†
264:†
238:Genus:
218:Order:
195:Class:
1415:272594
1347:Q19129
951:Extant
914:orders
840:
830:
822:
781:
771:
763:
714:
706:
698:
565:
525:
515:
507:
450:haptos
297:, 1911
295:Pocock
254:, 1911
252:Pocock
232:, 1911
230:Pocock
1397:IRMNG
1166:Acari
712:S2CID
633:(PDF)
487:PeerJ
458:podos
437:Names
206:Clade
1384:GBIF
1103:s.l.
870:and
838:PMID
820:ISSN
779:PMID
761:ISSN
704:PMID
696:ISSN
593:โ84.
563:ISSN
523:PMID
505:ISSN
454:pous
429:and
423:s.s.
401:had
364:Siro
48:Pre๊
1371:EoL
828:PMC
810:doi
806:283
769:PMC
751:doi
688:doi
647:doi
643:150
555:doi
513:PMC
495:doi
317:of
1447::
1425::
1412::
1399::
1386::
1373::
1358::
1343::
862:.
836:.
826:.
818:.
804:.
800:.
777:.
767:.
759:.
749:.
739:17
737:.
733:.
710:.
702:.
694:.
686:.
674:.
670:.
641:.
635:.
615:37
613:.
561:.
551:90
549:.
535:^
521:.
511:.
503:.
489:.
485:.
471:^
456:,
433:.
413:,
343:.
325:,
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242:โ
222:โ
103:N
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88:J
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78:P
73:C
68:D
63:S
58:O
53:๊
20:)
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