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Haplogroup

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2487:. The ground zero extinction event appears to be Toba during which haplogroup CDEF* appeared to diverge into C, DE and F. C and F have almost nothing in common while D and E have plenty in common. Extinction event #1 according to current estimates occurred after Toba, although older ancient DNA could push the ground zero extinction event to long before Toba, and push the first extinction event here back to Toba. Haplogroups with extinction event notes by them have a dubious origin and this is because extinction events lead to severe bottlenecks, so all notes by these groups are just guesses. Note that the SNP counting of ancient DNA can be highly variable meaning that even though all these groups diverged around the same time no one knows when. 1716: 2174: 231: 534: 2439:– Large group found within the N type. Populations contained therein can be divided geographically into West Eurasia and East Eurasia. Almost all European populations and a large number of Middle-Eastern population today are contained within this branch. A smaller percentage is contained in other N type groups (See above). Below are 4378:
Rootsi S, Magri C, Kivisild T, Benuzzi G, Help H, Bermisheva M, Kutuev I, Barać L, Pericić M, Balanovsky O, Pshenichnov A, Dion D, Grobei M, Zhivotovsky LA, Battaglia V, Achilli A, Al-Zahery N, Parik J, King R, Cinnioğlu C, Khusnutdinova E, Rudan P, Balanovska E, Scheffrahn W, Simonescu M, Brehm A,
2357:
Haplogroup R consists of two subgroups defined on the basis of their geographical distributions, one found in southeastern Asia and Oceania and the other containing almost all of the modern European populations. Haplogroup N(xR), i.e. mtDNA that belongs to the N group but not to its R subgroup, is
305: 326:
All of these molecules are part of the ancestral haplogroup, but at some point in the past a mutation occurred in the ancestral molecule, mutation A, which produced a new lineage; this is haplogroup A and is defined by mutation A. At some more recent point in the past, a new mutation, mutation B,
2353:
The M group comprises the first wave of human migration which is thought to have evolved outside of Africa, following an eastward route along southern coastal areas. Descendant lineages of haplogroup M are now found throughout Asia, the Americas, and Melanesia, as well as in parts of the Horn of
4622:
Raghavan M, Skoglund P, Graf KE, Metspalu M, Albrechtsen A, Moltke I, Rasmussen S, Stafford TW, Orlando L, Metspalu E, Karmin M, Tambets K, Rootsi S, Mägi R, Campos PF, Balanovska E, Balanovsky O, Khusnutdinova E, Litvinov S, Osipova LP, Fedorova SA, Voevoda MI, DeGiorgio M, Sicheritz-Ponten T,
454:
The special feature that both Y chromosomes and mtDNA display is that mutations can accrue along a certain segment of both molecules and these mutations remain fixed in place on the DNA. Furthermore, the historical sequence of these mutations can also be inferred. For example, if a set of ten Y
458:
Furthermore, all ten individuals who carry the chromosome with mutation A are the direct male line descendants of the same man who was the first person to carry this mutation. The first man to carry mutation B was also a direct male line descendant of this man, but is also the direct male line
1193:
K-M2313*, which as yet has no phylogenetic name, has been documented in two living individuals, who have ethnic ties to India and South East Asia. In addition, K-Y28299, which appears to be a primary branch of K-M2313, has been found in three living individuals from India. See: Poznik
2349:
The mitochondrial haplogroups are divided into three main groups, which are designated by the sequential letters L, M, N. Humanity first split within the L group between L0 and L1-6. L1-6 gave rise to other L groups, one of which, L3, split into the M and N group.
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Africa and North Africa; almost none have been found in Europe. The N haplogroup may represent another macrolineage that evolved outside of Africa, heading northward instead of eastward. Shortly after the migration, the large R group split off from the N.
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U – High frequency in West Eurasia, Indian sub-continent, and Algeria, found from India to the Mediterranean and to the rest of Europe; U5 in particular shows high frequency in Scandinavia and Baltic countries with the highest frequency in the
4379:
Goncalves R, Rosa A, Moisan JP, Chaventre A, Ferak V, Füredi S, Oefner PJ, Shen P, Beckman L, Mikerezi I, Terzić R, Primorac D, Cambon-Thomsen A, Krumina A, Torroni A, Underhill PA, Santachiara-Benerecetti AS, Villems R, Semino O (Jul 2004).
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at different chromosomal regions that are closely linked and that tend to be inherited together. As a haplogroup consists of similar haplotypes, it is usually possible to predict a haplogroup from haplotypes. Haplogroups pertain to a single
2373:
M1 – Ethiopian, Somali and Indian populations. Likely due to much gene flow between the Horn of Africa and the Arabian Peninsula (Saudi Arabia, Yemen, Oman), separated only by a narrow strait between the Red Sea and the Gulf of Aden.
1340:(M215) Associated with the spread of Afroasiatic languages; now concentrated in North Africa and the Horn of Africa, as well as parts of the Middle East, the Mediterranean, and the Balkans; formerly known as E3b 455:
chromosomes (derived from ten different individuals) contains a mutation, A, but only five of these chromosomes contain a second mutation, B, then it must be the case that mutation B occurred after mutation A.
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All humans carrying mutation A form a single haplogroup, and all humans carrying mutation B are part of this haplogroup, but mutation B also defines a more recent haplogroup (which is a subgroup or
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DNA (Y-DNA) haplogroups are named from A to T, and are further subdivided using numbers and lower case letters. Y chromosome haplogroup designations are established by the Y Chromosome Consortium.
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According to SNPS haplogroups which are the age of the first extinction event tend to be around 45–50 kya. Haplogroups of the second extinction event seemed to diverge 32–35 kya according to
466:) of its own to which humans carrying only mutation A do not belong. Both mtDNA and Y chromosomes are grouped into lineages and haplogroups; these are often presented as tree-like diagrams. 459:
ancestor of all men carrying mutation B. Series of mutations such as this form molecular lineages. Furthermore, each mutation defines a set of specific Y chromosomes called a haplogroup.
451:. Effectively this means that the genetic material from these chromosomes gets mixed up in every generation, and so any new mutations are passed down randomly from parents to offspring. 2459:
Pre-JT – Arose in the Levant (modern Lebanon area), found in 25% frequency in Bedouin populations; branch JT (branch J; branch T) – North, Eastern Europe, Indus, Mediterranean
424:; the remaining 95% of the Y chromosome does not recombine. Therefore, the Y chromosome and any mutations that arise in it are passed down in a direct male line of descent. 377:
of the cell, but mtDNA is an exception. An individual inherits their cytoplasm and the organelles contained by that cytoplasm exclusively from the maternal ovum (egg cell);
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Haplogroups are normally identified by an initial letter of the alphabet, and refinements consist of additional number and letter combinations, such as (for example)
637: 627: 1056:
Van Oven M, Van Geystelen A, Kayser M, Decorte R, Larmuseau HD (2014). "Seeing the wood for the trees: a minimal reference phylogeny for the human Y chromosome".
724: 707: 715: 1509:(M267) (Mostly associated with Semitic peoples in the Middle East but also found in; Mediterranean Europe, Ethiopia, North Africa, Pakistan, India and with 114: 874: 867: 860: 389:
arises in a mtDNA molecule, the mutation is therefore passed down in a direct female line of descent. Mutations are changes in the nitrogen bases of the
227:. As such, membership of a haplogroup, by any individual, relies on a relatively small proportion of the genetic material possessed by that individual. 254:
of a single broader set (as opposed, that is, to biparental models, such as human family trees). Haplogroups can be further divided into subclades.
983: 978: 1572:(M70, M184/USP9Y+3178, M193, M272) (North Africa, Horn of Africa, Southwest Asia, the Mediterranean, South Asia); formerly known as Haplogroup K2 1304:(M130) (Oceania, North/Central/East Asia, North America and a minor presence in South America, Southeast Asia, South Asia, West Asia, and Europe) 4806: 4380: 2358:
typical of Australian aboriginal populations, while also being present at low frequencies among many populations of Eurasia and the Americas.
350:, such as those of humans. Their primary function is to provide energy to the cell. Mitochondria are thought to be reduced descendants of 2377:
CZ – Many Siberians; branch C – Some Amerindian; branch Z – Many Saami, some Korean, some North Chinese, some Central Asian populations.
166: 4756: 140: 4884: 4879: 296:, and thus Y-DNA and mtDNA change only by chance mutation at each generation with no intermixture between parents' genetic material. 3048:
East Africa (First Extinction Event is the split between E1b1 and E1a, second extinction event is the split between E1b1b and E1b1a)
1254:
Major Y-chromosome haplogroups, and their geographical regions of occurrence (prior to the recent European colonization), include:
4741: 1322:(CTS3946) (Tibet, Nepal, Japan, the Andaman Islands, Central Asia, and a sporadic presence in Nigeria, Syria, and Saudi Arabia) 1034: 4145: 4122: 2456:
R0 – Found in Arabia and among Ethiopians and Somalis; branch HV (branch H; branch V) – Europe, Western Asia, North Africa;
2189:
populations and are often geographically oriented. For example, the following are common divisions for mtDNA haplogroups:
1453:(M170, P19, M258) (widespread in Europe, found infrequently in parts of the Middle East, and virtually absent elsewhere) 277: 4500:
Tolk HV, Barac L, Pericic M, Klaric IM, Janicijevic B, Campbell H, Rudan I, Kivisild T, Villems R, Rudan P (Sep 2001).
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B – Some Chinese, Tibetans, Mongolians, Central Asians, Koreans, Amerindians, South Siberians, Japanese, Austronesians
1893: 1724: 273: 85: 518: 475: 90: 357:
that were once free living. One indication that mitochondria were once free living is that each contains a circular
4896: 4117: 1175:
Haplogroup K2b1 (P397/P399) is also known as Haplogroup MS, but has a broader and more complex internal structure.
508: 159: 4687:"Improved phylogenetic resolution and rapid diversification of Y-chromosome haplogroup K-M526 in Southeast Asia" 1376:(M201) (present among many ethnic groups in Eurasia, usually at low frequency; most common in the Caucasus, the 4165: 394: 212: 1510: 4769: 4600: 4250: 1221:
Haplogroup S, as of 2017, is also known as K2b1a. (Previously the name Haplogroup S was assigned to K2b1a4.)
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happened in a person carrying haplogroup A; mutation B defined haplogroup B. Haplogroup B is a subgroup, or
4926: 4078: 2475:
Here is a list of Y-chromosome and MtDNA geographic haplogroup assignation proposed by Bekada et al. 2013.
1884:. The most up-to-date version of the mtDNA tree is maintained by Mannis van Oven on the PhyloTree website. 1230:
Haplogroup M, as of 2017, is also known as K2b1b. (Previously the name Haplogroup M was assigned to K2b1d.)
4853: 486: 4936: 4931: 4865: 4381:"Phylogeography of Y-chromosome haplogroup I reveals distinct domains of prehistoric gene flow in europe" 4102: 4093: 2484: 2162: 1248: 405: 74: 17: 4543:"Introducing the Algerian mitochondrial DNA and Y-chromosome profiles into the North African landscape" 1909: 152: 34: 4841: 4839:
MitoTool – a web server for the analysis and retrieval of human mitochondrial DNA sequence variations
1782: 1651: 1577: 1301: 421: 4916: 4816: 4421: 1523:(M172) (Mainly found in West Asia, Central Asia, Iran, Italy, Greece, the Balkans and North Africa) 1337: 1331: 697: 4874: 4623:
Brunak S, Demeshchenko S, Kivisild T, Villems R, Nielsen R, Jakobsson M, Willerslev E (Jan 2014).
2433:; also found with low frequency in some other populations of Siberia, East Asia, and Central Asia 4277:"Punctuated bursts in human male demography inferred from 1,244 worldwide Y-chromosome sequences" 4088: 1715: 1657: 1566:(M11, M20, M22, M61, M185, M295) (South Asia, Central Asia, Southwestern Asia, the Mediterranean) 900: 262: 4502:"The evidence of mtDNA haplogroup F in a European population and its ethnohistoric implications" 4459:
Loogväli EL, Roostalu U, Malyarchuk BA, Derenko MV, Kivisild T, Metspalu E, et al. (2004).
4921: 4149: 1683: 1677: 1438: 1425: 889: 881: 621: 4169: 1420:
H2 (P96) Formerly known as haplogroup F3. Found with low frequency in Europe and western Asia.
1131:
F-Y27277, sometimes known as F2'4, is both the parent clade of F2 and F4 and a child of F-M89.
4112: 2218: 2214: 2210: 2206: 2202: 2198: 2123: 2118: 2085: 1992: 1970: 1965: 1960: 1949: 1941: 1936: 1849: 1821: 1817: 1813: 1809: 1805: 1801: 1789: 1770: 1746: 1691: 1671: 1600: 1557: 1520: 1506: 1476: 1462: 1456: 1444: 1313: 951: 797: 436: 417: 293: 120: 2173: 4636: 4554: 4541:
Bekada A, Fregel R, Cabrera VM, Larruga JM, Pestano J, Benhamamouch S, González AM (2013).
2384: 2343: 2339: 2335: 2326: 2322: 2318: 2314: 2310: 2303: 2299: 2295: 2291: 2287: 2283: 2279: 2275: 2271: 2267: 2260: 2256: 2252: 2248: 2244: 2240: 2236: 2232: 2228: 2224: 2181:, with key (coloured) indicating periods in numbered thousands of years before the present. 2150: 2145: 2140: 2135: 2128: 2111: 2103: 2080: 2075: 2065: 2058: 2053: 2048: 2043: 2035: 2030: 2025: 2020: 2012: 2007: 2002: 1997: 1982: 1977: 1881: 1877: 1873: 1869: 1865: 1861: 1857: 1853: 1845: 1841: 1837: 1833: 1829: 1825: 1793: 1778: 1774: 1766: 1762: 1758: 1754: 1750: 1742: 1738: 1734: 1701: 1665: 1645: 1612: 1606: 1594: 1569: 1563: 1552: 1530: 1470: 1450: 1387: 1373: 1365: 1359: 1325: 1319: 1277: 1263: 990: 909: 844: 788: 773: 759: 680: 652: 409: 366: 8: 4331:"A recent bottleneck of Y chromosome diversity coincides with a global change in culture" 4083: 1621: 501: 281: 135: 125: 56: 4851:
HaploGrep – automatic classification of mitochondrial DNA haplogroups based on PhyloTree
4799: 4640: 4558: 547:
Please help update this article to reflect recent events or newly available information.
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Each haplogroup originates from, and remains part of, a preceding single haplogroup (or
4711: 4686: 4657: 4624: 4577: 4542: 4408: 4355: 4330: 4303: 4276: 1244: 1081: 919: 593: 576: 4227: 4194: 1098: 243: 4863:
HaploFind – fast automatic haplogroup assignment pipeline for human mitochondrial DNA
4716: 4662: 4582: 4523: 4482: 4413: 4360: 4308: 4232: 4214: 4161: 4141: 4107: 2178: 2158: 2094: 1905: 1730: 1635: 1473:(M304) (the Middle East, Turkey, Caucasus, Italy, Greece, the Balkans, North Africa) 1381: 1150: 1073: 1011: 1006: 997: 940: 931: 851: 822: 782: 740: 692: 610: 362: 247: 130: 42: 1085: 365:(mtDNA), whose structure is more similar to bacteria than eukaryotic organisms (see 331:
of haplogroup A; both haplogroups A and B are subclades of the ancestral haplogroup.
4785: 4706: 4698: 4652: 4644: 4572: 4562: 4513: 4472: 4403: 4395: 4350: 4342: 4298: 4290: 4222: 4206: 2426: 2414:
X – Some Amerindians, Southern Siberians, Southwest Asians, and Southern Europeans
1627: 1065: 973: 731: 685: 4791: 4869: 4857: 4845: 4567: 4461:"Disuniting uniformity: a pied cladistic canvas of mtDNA haplogroup H in Eurasia" 4281: 1686:(M343) (Europe, Caucasus, Central Asia, South Asia, North Africa, Central Africa) 1377: 745: 230: 1615:(M175) (East Asia, Southeast Asia, the South Pacific, South Asia, Central Asia) 381:
only pass on the chromosomal DNA, all paternal mitochondria are digested in the
4098: 1545: 1500: 1496: 1307: 961: 754: 587: 494: 269: 196: 4625:"Upper Palaeolithic Siberian genome reveals dual ancestry of Native Americans" 4195:"A Nomenclature System for the Tree of Human Y-Chromosomal Binary Haplogroups" 412:
male. Although Y chromosomes are situated in the cell nucleus and paired with
4905: 4829:
PhyloTree – The phylogenetic tree of global human mitochondrial DNA variation
4218: 2430: 2194: 1923: 1797: 1206: 1154: 836: 830: 816: 805: 768: 671: 666: 657: 643: 604: 444: 432: 4792:
A Nomenclature System for the Tree of Human Y-Chromosomal Binary Haplogroups
4518: 4501: 4477: 4460: 242:). As such, any related group of haplogroups may be precisely modelled as a 4911: 4720: 4666: 4586: 4527: 4486: 4417: 4364: 4312: 4236: 2470: 2070: 1680:(M17) (Central Asia, South Asia, and Central, Northern, and Eastern Europe) 1403: 1077: 516: 481: 413: 401: 390: 374: 335: 4807:
The Genetic Legacy of Paleolithic Homo sapiens sapiens in Extant Europeans
4702: 4346: 4817:
Y-DNA Ethnographic and Genographic Atlas and Open-Source Data Compilation
2464: 2331: 289: 285: 4885:
Do the Four Clades of the mtDNA Haplogroup L2 Evolve at Different Rates?
4774: 4648: 2259:(all listed West Eurasian haplogroups are derived from macro-haplogroup 1113:
Haplogroup A0-T is also known as A-L1085 (and previously as A0'1'2'3'4).
4140:
By C. Barry Cox, Peter D. Moore, Richard Ladle. Wiley-Blackwell, 2016.
2390:
G – Many Northeast Siberians, northern East Asians, and Central Asians
1069: 370: 211:) is a group of similar haplotypes that share a common ancestor with a 187:
in an organism that are inherited together from a single parent, and a
4275:
Poznik, G. David; Xue, Yali; Mendez, Fernando L.; et al. (2016).
4210: 4181: 1399:
H1a (M69/Page45) India, Sri Lanka, Nepal, Pakistan, Iran, Central Asia
1199: 1153:
was known as "Haplogroup K2". That name has since been re-assigned to
1210:, 2017, "Details of the Y-SNP markers included in the minimal Y tree" 428: 351: 347: 343: 339: 239: 180: 65: 4294: 1654:(M242) (Occurred ~15,000–20,000 bp. Found in Asia and the Americas) 304: 4685:
Karafet TM, Mendez FL, Sudoyo H, Lansing JS, Hammer MF (Mar 2015).
4399: 2411:
W – Some Eastern Europeans, South Asians, and southern East Asians
2408:
S – Some Indigenous Australian (First Nations People of Australia)
2186: 1514: 1484: 1166:
Haplogroup K2b (M1221/P331/PF5911) is also known as Haplogroup MPS.
463: 386: 354: 328: 215: 97: 69: 1055: 369:). The overwhelming majority of a human's DNA is contained in the 4458: 4329:
Karmin, Monika; Saag, Lauri; Vicente, Mário; et al. (2015).
4254: 2450: 2402:
A – Found in many Amerindians and some East Asians and Siberians
1480: 1459:(M253, M307, P30, P40) (Northern Europe, dominant in Scandinavia) 1407: 1285: 1281: 1271: 1267: 440: 224: 192: 4850: 4833: 4601:"Common genetic ancestors lived during roughly same time period" 1334:(V38) West Africa and surrounding regions; formerly known as E3a 2418: 1492: 448: 382: 251: 219: 184: 4862: 4779: 2380:
D – Some Amerindians, many Siberians and northern East Asians
4828: 1704:(M230, P202, P204) (New Guinea, Melanesia, eastern Indonesia) 378: 261:. The alphabetical nomenclature was published in 2002 by the 4621: 4445: 1886: 4838: 2422: 1488: 4834:
PhyloD3 – D3.js-based phylogenetic tree based on PhyloTree
4684: 4540: 4377: 4186: 1099:
International Society of Genetic Genealogy (ISOGG; 2015),
435:(when another X chromosome is available to pair with it), 2161:
is the name given by researchers to the woman who is the
1140:
Haplogroup LT (L298/P326) is also known as Haplogroup K1.
358: 288:
line, from father to son, while mtDNA is passed down the
4782:
A minimal reference phylogeny for the human Y-chromosome
2471:
Y-chromosome and MtDNA geographic haplogroup assignation
2163:
most recent common matrilineal (female-lineage) ancestor
1660:(M3) (North America, Central America, and South America) 1517:; J1 with DYS388=13 is associated with eastern Anatolia) 1247:
is the name given by researchers to the male who is the
408:; nearly all humans that possess a Y chromosome will be 1465:(S31) (Central and Southeast Europe, Sardinia, Balkans) 393:. Single changes from the original sequence are called 4499: 1710: 1432: 1249:
most recent common patrilineal (male-lineage) ancestor
292:
line, from mother to offspring of both sexes. Neither
4150:
Biogeography: An Ecological and Evolutionary Approach
1415:
H3 (Z5857) India, Sri Lanka, Pakistan, Bahrain, Qatar
1362:(M89) Oceania, Europe, Asia, North and South America 469: 218:. More specifically, a haplotype is a combination of 2393:
Q – Melanesian, Polynesian, New Guinean populations
2302:(note: C, D, E, G, and Z belong to macro-haplogroup 4798:(2), 339–48, February 2002. (Detailed hierarchical 2757:West of Burma in Eurasia (First Extinction Event?) 1380:, and Anatolia; in Europe mainly in Greece, Italy, 2449:F – Mainly found in southeastern Asia, especially 4328: 1597:(P256) (New Guinea, Melanesia, eastern Indonesia) 4903: 4822: 4180:International Society of Genetic Genealogy 2015 2102: 2099: 1945: 4763: 4680: 4678: 4676: 4274: 2177:Map of human haplotype migration, according to 1257: 4813:(paper which introduced the "Eu" haplogroups). 2405:I – 10% frequency in Northern, Eastern Europe 1719:Human migrations and mitochondrial haplogroups 1384:, the Tyrol, Bohemia; rare in Northern Europe) 1212:(Access date of these pages: 9 December 2017) 4802:has conversions from previous naming schemes) 4438: 4324: 4322: 4205:(2). Cold Spring Harbor Laboratory: 339–348. 2752:M172, M410, M158, M319, DYS445=6, M339, M340 502: 160: 4673: 4452: 4154: 1586: 1292: 272:, the haplogroups most commonly studied are 4193:Consortium, The Y Chromosome (2002-02-01). 4160:Editorial Board, V&S Publishers, 2012, 2779:M11, M20, M27, M76, M317, M274, M349, M357 1894:human mitochondrial DNA (mtDNA) haplogroups 1694:(M124) (South Asia, Caucasus, Central Asia) 1350: 1184:Haplogroup P (P295) is also klnown as K2b2. 4319: 4268: 4192: 1539: 1049: 509: 495: 167: 153: 4880:The Making of the African mtDNA Landscape 4710: 4656: 4576: 4566: 4517: 4476: 4407: 4354: 4302: 4226: 3010:M60, M181, SRY10831.1, M150, M109, M112 2828:Middle East, Europe, Siberia, South Asia 2172: 1714: 1125: 1122:Haplogroup A1 is also known as A1'2'3'4. 1092: 303: 229: 4174: 2168: 1368:(P14, M213) (China, Vietnam, Singapore) 1187: 299: 14: 4904: 4747: 4134: 2795:East Asia, South East Asia, South Asia 2034: 1986: 1953: 1948: 1930: 1927: 1914: 1904: 280:, each of which can be used to define 115:Y-chromosome haplogroups by population 4123:List of haplogroups of notable people 3037:West Africa (First Extinction Event?) 1981: 1976: 490: 278:mitochondrial DNA (mtDNA) haplogroups 4790:The Y Chromosome Consortium (2002), 2762:Indonesia (First Extinction Event?) 2127: 2122: 2117: 2107: 2092: 2089: 2084: 2079: 2074: 2069: 2064: 2057: 2052: 2047: 2042: 2039: 2029: 2024: 2019: 2016: 2011: 2006: 2001: 1996: 1991: 1969: 1964: 1959: 1956: 1940: 1935: 1922: 1917: 1901: 1297:(mutation M168 occurred ~50,000 bp) 1169: 1035:Y-DNA haplogroups of historic people 1010: 1005: 996: 994: 989: 987: 982: 977: 972: 966: 959: 949: 947: 937: 929: 917: 915: 906: 898: 856: 849: 843: 841: 835: 829: 827: 821: 815: 803:       793: 787: 781: 772: 767: 758: 753: 744: 739: 703: 689: 572: 527: 4875:graphical mtDNA haplogroup skeleton 1711:Human mitochondrial DNA haplogroups 1433:Groups with mutations L15 & L16 1355:(mutation M89 occurred ~45,000 bp) 1160: 1107: 677: 617: 600: 284:. Y-DNA is passed solely along the 24: 4691:European Journal of Human Genetics 4506:European Journal of Human Genetics 4388:American Journal of Human Genetics 3021:M1, YAP, M174, M40, M96, M75, M98 2839:Anatolia, South East Europe ? 2741:M304, M267, P58, M365, M368, M369 2453:; 8.3% in Hvar Island in Croatia. 2185:Haplogroups can be used to define 1725:Human mitochondrial DNA haplogroup 1143: 1134: 470:Human Y-chromosome DNA haplogroups 234:Y-DNA haplogroups map of the world 25: 4948: 4730: 2504:Europe (Second Extinction Event?) 2364:consists of nearly all Africans. 1224: 1215: 1178: 1116: 1104:. (Access date: 1 February 2015.) 476:Human Y-chromosome DNA haplogroup 207:, "onefold, simple" and English: 4118:List of genetic genealogy topics 2383:E – Malay, Borneo, Philippines, 1648:(M45) (M45 occurred ~35,000 bp) 532: 274:Y-chromosome (Y-DNA) haplogroups 4615: 4593: 4534: 4493: 4371: 2521:P215, M438, P37.2, M359, P41.2 2478: 1157:, the sibling of Haplogroup LT. 395:single nucleotide polymorphisms 4243: 2790:M231, M214, LLY22g, Tat, M178 1544:(mutation M9 occurred ~40,000 1280:(M60) (Africa, especially the 1266:(M91) (Africa, especially the 213:single-nucleotide polymorphism 13: 1: 4823:Mitochondrial DNA haplogroups 4170:Concise Dictionary of Science 4128: 2823:MEH2, M242, P36.2, M25, M346 2510:M170, M253, P259, M227, M507 1609:(M231) (northernmost Eurasia) 1483:, with a few observations in 420:with the X chromosome at the 4775:ISOGG Y-DNA Haplogroup Tree 4764:Y chromosome DNA haplogroups 4568:10.1371/journal.pone.0056775 4079:International HapMap Project 2834:M420, M17, M198, M204, M458 1258:Groups without mutation M168 519:Y-chromosome DNA haplogroups 437:share their genetic material 7: 4890: 4103:Race (human categorization) 4094:Human evolutionary genetics 4072: 2719:M201, M285, P15, P16, M406 1393:H1'3 (Z4221/M2826, Z13960) 569: 517:Phylogenetic tree of human 75:Most recent common ancestor 27:Group of similar haplotypes 10: 4953: 4735: 2784:East Asia, South East Asia 2730:M69, M52, M82, M197, M370 2683:C1(formerly known as CxC3) 1889: 1733:haplogroups are lettered: 1722: 1101:Y-DNA Haplogroup Tree 2015 1003: 970: 927: 896: 887: 879: 872: 865: 858: 813: 779: 765: 751: 737: 721: 705: 574: 473: 312: Ancestral haplogroup 4251:"Y Chromosome Consortium" 2993:West Africa, North Africa 2694:C2 (formerly known as C3) 1587:Groups with mutation M526 1293:Groups with mutation M168 1024: 802: 795: 729: 712: 696: 691: 684: 679: 670: 665: 663: 656: 651: 649: 642: 635: 633: 626: 619: 609: 602: 592: 585: 583: 567: 541:This article needs to be 525: 324: Haplogroup B (Hg B) 318: Haplogroup A (Hg A) 208: 200: 4897:Y-DNA Haplogroup Browser 4794:, Genome Research, Vol. 3071: 2149: 2144: 2139: 2134: 2110: 1351:Groups with mutation M89 422:ends of the Y chromosome 4770:Y Chromosome Consortium 4742:The Genographic Project 4519:10.1038/sj.ejhg.5200709 4089:Molecular phylogenetics 2702:Indonesia or South Asia 2680:South Asia or Melanesia 1540:Groups with mutation M9 1149:Between 2002 and 2008, 263:Y Chromosome Consortium 250:(haplogroup) is also a 4805:Semino et al. (2000), 4757:World Haplogroups Maps 2812:92R7, M207, M173, M45 2806:Indonesia, Philippines 2587:M12, M102, M280, M241 2182: 2165:of all living humans. 1720: 1533:(M9, P128, P131, P132) 1402:H1b (B108) Found in a 1251:of all living humans. 332: 235: 4703:10.1038/ejhg.2014.106 4478:10.1093/molbev/msh209 4347:10.1101/gr.186684.114 4113:Genealogical DNA test 2845:M173, M343, P25, M73 2746:Europe or Middle East 2735:Europe or Middle East 2543:M223, M284, P78, P95 2176: 1892:Phylogenetic tree of 1718: 1618:Haplogroup O1 (F265) 307: 233: 121:Genealogical DNA test 4786:Haplogroup Predictor 2385:Taiwanese aborigines 2169:Defining populations 367:endosymbiotic theory 300:Haplogroup formation 4927:Population genetics 4748:all DNA haplogroups 4649:10.1038/nature12736 4641:2014Natur.505...87R 4559:2013PLoSO...856775B 4084:Molecular evolution 3054:P2, M2, U175, M191 2820:R and Q (QR) split 2817:South Asia, Siberia 2387:, Papua New Guinea 2332:Australo-Melanesian 1511:Northeast Caucasian 1030:Y-DNA by population 945:    427:Other chromosomes, 282:genetic populations 136:Genographic Project 126:Surname DNA project 86:Human mitochondrial 57:Population genetics 4937:Types of groupings 4932:Classical genetics 4868:2016-06-11 at the 4856:2016-06-12 at the 4844:2016-06-19 at the 4780:PhyloTree's Y-tree 2183: 1721: 1245:Y-chromosomal Adam 1070:10.1002/humu.22468 796:    577:Y-chromosomal Adam 404:are male-specific 333: 236: 91:Human Y-chromosome 4211:10.1101/gr.217602 4182:Genetics Glossary 4146:978-1-118-96858-1 4108:Genetic genealogy 4070: 4069: 3069: 3068: 2713:Europe (Caucasus) 2631:U198, P312, S116 2309:Native American: 2179:mitochondrial DNA 2159:Mitochondrial Eve 2156: 2155: 2097: 1906:Mitochondrial Eve 1785: 1652:Haplogroup Q-M242 1578:Haplogroup K(xLT) 1479:(Mainly found in 1316:(M1, M145, M203) 1241: 1240: 1203:, 2017, "K-M2335" 1201:YFull YTree v5.08 1151:Haplogroup T-M184 1019: 1018: 964: 954: 944: 935: 923: 912: 904: 892: 884: 877: 870: 863: 808: 727: 719: 710: 640: 624: 607: 590: 562: 561: 443:, the process of 363:mitochondrial DNA 177: 176: 131:Personal genomics 43:Genetic genealogy 16:(Redirected from 4944: 4725: 4724: 4714: 4682: 4671: 4670: 4660: 4619: 4613: 4612: 4610: 4608: 4597: 4591: 4590: 4580: 4570: 4538: 4532: 4531: 4521: 4497: 4491: 4490: 4480: 4456: 4450: 4449: 4442: 4436: 4435: 4433: 4432: 4426: 4420:. Archived from 4411: 4385: 4375: 4369: 4368: 4358: 4326: 4317: 4316: 4306: 4272: 4266: 4265: 4263: 4262: 4253:. Archived from 4247: 4241: 4240: 4230: 4190: 4184: 4178: 4172: 4158: 4152: 4138: 3076: 3075: 3026:East Asia, Nepal 2490: 2489: 2093: 1887: 1783: 1646:Haplogroup P-M45 1580:(rs2033003/M526) 1396:H1 (L902/M3061) 1338:Haplogroup E1b1b 1332:Haplogroup E1b1a 1231: 1228: 1222: 1219: 1213: 1191: 1185: 1182: 1176: 1173: 1167: 1164: 1158: 1147: 1141: 1138: 1132: 1129: 1123: 1120: 1114: 1111: 1105: 1096: 1090: 1089: 1053: 960: 950: 938: 930: 918: 908: 899: 888: 880: 873: 866: 859: 804: 723: 714: 706: 636: 620: 603: 586: 570: 557: 554: 548: 536: 535: 528: 511: 504: 497: 488: 487: 348:eukaryotic cells 342:that lie in the 323: 317: 311: 260: 246:, in which each 244:nested hierarchy 210: 202: 169: 162: 155: 30: 29: 21: 4952: 4951: 4947: 4946: 4945: 4943: 4942: 4941: 4917:Human evolution 4902: 4901: 4893: 4870:Wayback Machine 4858:Wayback Machine 4846:Wayback Machine 4825: 4809:, Science, Vol 4766: 4750: 4738: 4733: 4728: 4683: 4674: 4635:(7481): 87–91. 4620: 4616: 4606: 4604: 4599: 4598: 4594: 4539: 4535: 4498: 4494: 4471:(11): 2012–21. 4465:Mol. Biol. Evol 4457: 4453: 4446:"PhyloTree.org" 4444: 4443: 4439: 4430: 4428: 4424: 4383: 4376: 4372: 4335:Genome Research 4327: 4320: 4295:10.1038/ng.3559 4282:Nature Genetics 4273: 4269: 4260: 4258: 4249: 4248: 4244: 4199:Genome Research 4191: 4187: 4179: 4175: 4159: 4155: 4139: 4135: 4131: 4075: 3074: 2872:Pakistan, India 2481: 2473: 2427:Tungusic people 2266:East Eurasian: 2223:West Eurasian: 2171: 1727: 1713: 1658:Haplogroup Q-M3 1589: 1542: 1435: 1378:Iranian plateau 1353: 1295: 1260: 1242: 1237: 1236: 1235: 1234: 1229: 1225: 1220: 1216: 1192: 1188: 1183: 1179: 1174: 1170: 1165: 1161: 1148: 1144: 1139: 1135: 1130: 1126: 1121: 1117: 1112: 1108: 1097: 1093: 1054: 1050: 1045: 1039: 1020: 563: 558: 552: 549: 546: 537: 533: 521: 515: 478: 472: 447:which produces 410:morphologically 406:sex chromosomes 325: 321: 319: 315: 313: 309: 302: 258: 225:line of descent 173: 117: 94: 89: 83: 81:DNA haplogroups 28: 23: 22: 15: 12: 11: 5: 4950: 4940: 4939: 4934: 4929: 4924: 4919: 4914: 4900: 4899: 4892: 4889: 4888: 4887: 4882: 4877: 4872: 4860: 4848: 4836: 4831: 4824: 4821: 4820: 4819: 4814: 4803: 4788: 4783: 4777: 4772: 4765: 4762: 4761: 4760: 4749: 4746: 4745: 4744: 4737: 4734: 4732: 4731:External links 4729: 4727: 4726: 4672: 4614: 4592: 4533: 4492: 4451: 4437: 4400:10.1086/422196 4370: 4341:(4): 459–466. 4318: 4289:(6): 593–599. 4267: 4242: 4185: 4173: 4153: 4132: 4130: 4127: 4126: 4125: 4120: 4115: 4110: 4105: 4099:Race (biology) 4096: 4091: 4086: 4081: 4074: 4071: 4068: 4067: 4064: 4060: 4059: 4056: 4052: 4051: 4048: 4044: 4043: 4040: 4036: 4035: 4032: 4028: 4027: 4024: 4020: 4019: 4016: 4012: 4011: 4008: 4004: 4003: 4000: 3996: 3995: 3992: 3988: 3987: 3984: 3980: 3979: 3976: 3972: 3971: 3968: 3964: 3963: 3960: 3956: 3955: 3952: 3948: 3947: 3944: 3940: 3939: 3936: 3932: 3931: 3928: 3924: 3923: 3920: 3916: 3915: 3912: 3908: 3907: 3904: 3900: 3899: 3896: 3892: 3891: 3888: 3884: 3883: 3880: 3876: 3875: 3874:N* / M* / L3* 3872: 3868: 3867: 3864: 3860: 3859: 3856: 3852: 3851: 3848: 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2497: 2494: 2480: 2477: 2472: 2469: 2441:subclades of R 2425:; many Ainus, 2421:and people of 2347: 2346: 2329: 2307: 2264: 2221: 2170: 2167: 2154: 2153: 2148: 2143: 2138: 2132: 2131: 2126: 2121: 2115: 2114: 2109: 2106: 2101: 2098: 2091: 2088: 2083: 2078: 2073: 2068: 2062: 2061: 2056: 2051: 2046: 2041: 2038: 2033: 2028: 2023: 2018: 2015: 2010: 2005: 2000: 1995: 1989: 1988: 1985: 1980: 1974: 1973: 1968: 1963: 1958: 1955: 1952: 1947: 1944: 1939: 1933: 1932: 1929: 1926: 1920: 1919: 1916: 1913: 1903: 1899: 1898: 1723:Main article: 1712: 1709: 1708: 1707: 1706: 1705: 1699: 1698: 1697: 1696: 1695: 1689: 1688: 1687: 1684:Haplogroup R1b 1681: 1678:Haplogroup R1a 1663: 1662: 1661: 1643: 1642: 1641: 1640: 1639: 1633: 1632: 1631: 1628:Haplogroup O1b 1625: 1622:Haplogroup O1a 1610: 1598: 1588: 1585: 1584: 1583: 1582: 1581: 1575: 1574: 1573: 1567: 1541: 1538: 1537: 1536: 1535: 1534: 1528: 1527: 1526: 1525: 1524: 1518: 1504: 1501:Turkic peoples 1497:Czech Republic 1468: 1467: 1466: 1460: 1439:Haplogroup IJK 1434: 1431: 1430: 1429: 1426:Haplogroup IJK 1423: 1422: 1421: 1418: 1417: 1416: 1413: 1412: 1411: 1406:individual in 1400: 1385: 1371: 1370: 1369: 1352: 1349: 1348: 1347: 1346: 1345: 1344: 1343: 1342: 1341: 1335: 1323: 1305: 1294: 1291: 1290: 1289: 1275: 1259: 1256: 1239: 1238: 1233: 1232: 1223: 1214: 1186: 1177: 1168: 1159: 1142: 1133: 1124: 1115: 1106: 1091: 1058:Human Mutation 1047: 1046: 1041: 1040: 1038: 1037: 1032: 1026: 1025: 1022: 1021: 1017: 1016: 1014: 1009: 1004: 1001: 1000: 995: 993: 988: 986: 981: 976: 971: 968: 967: 965: 958: 956: 948: 946: 936: 928: 925: 924: 916: 914: 905: 897: 894: 893: 886: 878: 871: 864: 857: 855: 848: 842: 840: 834: 828: 826: 820: 814: 811: 810: 801: 794: 792: 786: 780: 777: 776: 771: 766: 763: 762: 757: 752: 749: 748: 743: 738: 735: 734: 728: 720: 711: 704: 701: 700: 695: 690: 688: 683: 678: 675: 674: 669: 664: 661: 660: 655: 650: 647: 646: 641: 634: 631: 630: 625: 618: 615: 614: 608: 601: 598: 597: 591: 584: 581: 580: 573: 568: 565: 564: 560: 559: 540: 538: 531: 526: 523: 522: 514: 513: 506: 499: 491: 474:Main article: 471: 468: 320: 314: 308: 301: 298: 270:human genetics 183:is a group of 175: 174: 172: 171: 164: 157: 149: 146: 145: 144: 143: 138: 133: 128: 123: 118: 113: 108: 107: 106:Related topics 103: 102: 101: 100: 95: 79: 77: 72: 59: 51: 50: 46: 45: 39: 38: 26: 9: 6: 4: 3: 2: 4949: 4938: 4935: 4933: 4930: 4928: 4925: 4923: 4922:Phylogenetics 4920: 4918: 4915: 4913: 4910: 4909: 4907: 4898: 4895: 4894: 4886: 4883: 4881: 4878: 4876: 4873: 4871: 4867: 4864: 4861: 4859: 4855: 4852: 4849: 4847: 4843: 4840: 4837: 4835: 4832: 4830: 4827: 4826: 4818: 4815: 4812: 4808: 4804: 4801: 4797: 4793: 4789: 4787: 4784: 4781: 4778: 4776: 4773: 4771: 4768: 4767: 4758: 4755: 4754: 4753: 4752:Y-Chromosome 4743: 4740: 4739: 4722: 4718: 4713: 4708: 4704: 4700: 4697:(3): 369–73. 4696: 4692: 4688: 4681: 4679: 4677: 4668: 4664: 4659: 4654: 4650: 4646: 4642: 4638: 4634: 4630: 4626: 4618: 4602: 4596: 4588: 4584: 4579: 4574: 4569: 4564: 4560: 4556: 4553:(2): e56775. 4552: 4548: 4544: 4537: 4529: 4525: 4520: 4515: 4512:(9): 717–23. 4511: 4507: 4503: 4496: 4488: 4484: 4479: 4474: 4470: 4466: 4462: 4455: 4447: 4441: 4427:on 2009-06-24 4423: 4419: 4415: 4410: 4405: 4401: 4397: 4394:(1): 128–37. 4393: 4389: 4382: 4374: 4366: 4362: 4357: 4352: 4348: 4344: 4340: 4336: 4332: 4325: 4323: 4314: 4310: 4305: 4300: 4296: 4292: 4288: 4284: 4283: 4278: 4271: 4257:on 2017-01-16 4256: 4252: 4246: 4238: 4234: 4229: 4224: 4220: 4216: 4212: 4208: 4204: 4200: 4196: 4189: 4183: 4177: 4171: 4167: 4163: 4157: 4151: 4147: 4143: 4137: 4133: 4124: 4121: 4119: 4116: 4114: 4111: 4109: 4106: 4104: 4100: 4097: 4095: 4092: 4090: 4087: 4085: 4082: 4080: 4077: 4076: 4065: 4062: 4061: 4057: 4054: 4053: 4049: 4046: 4045: 4041: 4038: 4037: 4033: 4030: 4029: 4025: 4022: 4021: 4017: 4014: 4013: 4009: 4006: 4005: 4001: 3998: 3997: 3993: 3990: 3989: 3985: 3982: 3981: 3977: 3974: 3973: 3969: 3966: 3965: 3961: 3958: 3957: 3953: 3950: 3949: 3945: 3942: 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1999: 1994: 1990: 1984: 1979: 1975: 1972: 1967: 1962: 1951: 1943: 1938: 1934: 1925: 1921: 1911: 1907: 1900: 1897: 1896: 1895: 1888: 1885: 1883: 1879: 1875: 1871: 1867: 1863: 1859: 1855: 1851: 1847: 1843: 1839: 1835: 1831: 1827: 1823: 1819: 1815: 1811: 1807: 1803: 1799: 1795: 1791: 1787: 1780: 1776: 1772: 1768: 1764: 1760: 1756: 1752: 1748: 1744: 1740: 1736: 1732: 1726: 1717: 1703: 1700: 1693: 1692:Haplogroup R2 1690: 1685: 1682: 1679: 1676: 1675: 1673: 1672:Haplogroup R1 1670: 1669: 1667: 1664: 1659: 1656: 1655: 1653: 1650: 1649: 1647: 1644: 1637: 1636:Haplogroup O2 1634: 1629: 1626: 1623: 1620: 1619: 1617: 1616: 1614: 1611: 1608: 1605: 1604: 1602: 1601:Haplogroup NO 1599: 1596: 1593: 1592: 1591: 1590: 1579: 1576: 1571: 1568: 1565: 1562: 1561: 1559: 1558:Haplogroup LT 1556: 1555: 1554: 1551: 1550: 1549: 1547: 1532: 1529: 1522: 1521:Haplogroup J2 1519: 1516: 1512: 1508: 1507:Haplogroup J1 1505: 1502: 1498: 1494: 1490: 1486: 1482: 1478: 1477:Haplogroup J* 1475: 1474: 1472: 1469: 1464: 1463:Haplogroup I2 1461: 1458: 1457:Haplogroup I1 1455: 1454: 1452: 1449: 1448: 1446: 1445:Haplogroup IJ 1443: 1442: 1440: 1437: 1436: 1427: 1424: 1419: 1414: 1409: 1405: 1401: 1398: 1397: 1395: 1394: 1392: 1391: 1390:(L901/M2939) 1389: 1386: 1383: 1379: 1375: 1372: 1367: 1366:Haplogroup FT 1364: 1363: 1361: 1358: 1357: 1356: 1339: 1336: 1333: 1330: 1329: 1327: 1324: 1321: 1318: 1317: 1315: 1314:Haplogroup DE 1312: 1311: 1309: 1306: 1303: 1300: 1299: 1298: 1287: 1283: 1279: 1276: 1273: 1269: 1265: 1262: 1261: 1255: 1252: 1250: 1246: 1227: 1218: 1211: 1209: 1204: 1202: 1197: 1190: 1181: 1172: 1163: 1156: 1152: 1146: 1137: 1128: 1119: 1110: 1103: 1102: 1095: 1087: 1083: 1079: 1075: 1071: 1067: 1064:(2): 187–91. 1063: 1059: 1052: 1048: 1044: 1036: 1033: 1031: 1028: 1027: 1023: 1015: 1013: 1008: 1002: 999: 992: 985: 980: 975: 969: 963: 957: 955:   953: 942: 933: 926: 921: 911: 902: 895: 891: 885:   883: 876: 869: 862: 853: 847:   846: 838: 833:   832: 824: 819:   818: 812: 807: 799: 790: 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Retrieved 4255:the original 4245: 4202: 4198: 4188: 4176: 4156: 4136: 4034:L3b(16124!) 3970:L2a1(16189) 3743:North Africa 3735:North Africa 3727:North Africa 3719:North Africa 3711:North Africa 3703:North Africa 3015:Asia, Africa 2982:North Africa 2971:North Africa 2960:North Africa 2949:North Africa 2938:North Africa 2927:North Africa 2905:North Africa 2894:North Africa 2661:R1b1b1a1b3,4 2482: 2479:Y-chromosome 2474: 2461: 2458: 2455: 2448: 2445: 2440: 2436: 2435: 2416: 2413: 2410: 2407: 2404: 2401: 2396: 2395: 2392: 2389: 2382: 2379: 2376: 2372: 2367: 2366: 2361: 2360: 2356: 2352: 2348: 2184: 2157: 1891: 1890: 1728: 1702:Haplogroup S 1666:Haplogroup R 1613:Haplogroup O 1607:Haplogroup N 1595:Haplogroup M 1570:Haplogroup T 1564:Haplogroup L 1560:(L298/P326) 1553:Haplogroup K 1543: 1531:Haplogroup K 1499:, and among 1471:Haplogroup J 1451:Haplogroup I 1388:Haplogroup H 1374:Haplogroup G 1360:Haplogroup F 1354: 1326:Haplogroup E 1320:Haplogroup D 1310:haplogroups 1302:Haplogroup C 1296: 1278:Haplogroup B 1264:Haplogroup A 1253: 1243: 1226: 1217: 1207: 1200: 1195: 1189: 1180: 1171: 1162: 1145: 1136: 1127: 1118: 1109: 1100: 1094: 1061: 1057: 1051: 1042: 907:   850:   550: 542: 482:Y chromosome 479: 461: 457: 453: 426: 416:, they only 399: 391:DNA sequence 336:Mitochondria 334: 267: 259:A → A1 → A1a 256: 237: 204: 188: 178: 80: 61: 4063:West Africa 4055:West Africa 4047:West Africa 4039:West Africa 4031:West Africa 4023:West Africa 4015:West Africa 4007:West Africa 3999:West Africa 3991:West Africa 3983:West Africa 3975:West Africa 3967:West Africa 3959:West Africa 3951:West Africa 3943:West Africa 3938:L2a1a2'3'4 3935:West Africa 3927:West Africa 3919:West Africa 3911:West Africa 3903:West Africa 3895:West Africa 3887:West Africa 3879:West Africa 3871:East Africa 3863:East Africa 3855:East Africa 3847:East Africa 3839:East Africa 3831:East Africa 3823:East Africa 3815:East Africa 3807:East Africa 3799:East Africa 3791:East Africa 3783:East Africa 3775:East Africa 3767:East Africa 3759:East Africa 3751:East Africa 3695:Middle East 3687:Middle East 3679:Middle East 3671:Middle East 3663:Middle East 3655:Middle East 3647:Middle East 3639:Middle East 3623:Middle East 3615:Middle East 3607:Middle East 3599:Middle East 3591:Middle East 3583:Middle East 3567:Middle East 3559:Middle East 3551:Middle East 3543:Middle East 3535:Middle East 3527:Middle East 3519:Middle East 3511:Middle East 3503:Middle East 3495:Middle East 3487:Middle East 3479:Middle East 3471:Middle East 3463:Middle East 3455:Middle East 3447:Middle East 3439:Middle East 3431:Middle East 3423:Middle East 3415:Middle East 3407:Middle East 3399:Middle East 3391:Middle East 3383:Middle East 3375:Middle East 3367:Middle East 3359:Middle East 3351:Middle East 3343:Middle East 3335:Middle East 3319:Middle East 3311:Middle East 3303:Middle East 3295:Middle East 3287:Middle East 3162:HV0/HV0a/V 3082:Haplogroup 3059:Middle East 3004:East Africa 2883:Middle East 2878:M479, M124 2801:M175, M119 2598:M412, P310 2496:Haplogroup 2465:Sami people 1630:(P31, M268) 1513:peoples in 1441:(L15, L16) 371:chromosomes 290:matrilineal 286:patrilineal 18:Haplogroups 4906:Categories 4431:2007-03-08 4261:2005-07-27 4166:9381588643 4129:References 3450:J1d / J2b 2988:M123, M34 2773:South Asia 2724:South Asia 2708:M89, M282 2691:North Asia 2672:R1b1b1a1b5 2664:M65, M153 2650:R1b1b1a1b2 2639:R1b1b1a1b1 2565:M67, M166 2397:The N type 2368:The M type 2362:The L type 1447:(S2, S22) 1428:(L15, L16) 340:organelles 338:are small 294:recombines 189:haplogroup 62:Haplogroup 4219:1088-9051 4058:L3e2'3'4 3962:L2a1c1'2 3786:L3c/L4/M 3738:U6a1'2'3 3575:West Asia 3322:D/G/M9/E 2999:M91, M13 2941:E1b1b1a1b 2916:West Asia 2765:K2 (NOPS) 2628:R1b1b1a1b 2617:R1b1b1a1a 2417:Y – Most 2193:African: 1208:PhyloTree 1043:Footnotes 429:autosomes 418:recombine 385:. When a 361:, called 352:symbiotic 344:cytoplasm 240:paragroup 195:from the 181:haplotype 66:Haplotype 4891:Software 4866:Archived 4854:Archived 4842:Archived 4721:24896152 4667:24256729 4587:23431392 4547:PLOS ONE 4528:11571562 4487:15254257 4418:15162323 4365:25770088 4313:27111036 4237:11827954 4168:p. 137. 4148:p. 106. 4073:See also 3954:L2a1b'f 3746:U6b'c'd 3426:J1a'b'e 2963:E1b1b1a4 2952:E1b1b1a3 2930:E1b1b1a1 2919:E1b1b1a2 2675:SRY2627 2606:R1b1b1a1 1515:Dagestan 1485:Pakistan 1205:, and; 1196:op. cit. 1086:23291764 1078:24166809 716:F-Y27277 464:subclade 397:(SNPs). 387:mutation 355:bacteria 329:subclade 216:mutation 98:Genomics 70:Subclade 49:Concepts 35:a series 33:Part of 4736:General 4712:4326703 4658:4105016 4637:Bibcode 4578:3576335 4555:Bibcode 4409:1181996 4356:4381518 4304:4884158 4066:L3f1b* 4026:L3b1a3 3994:L2c1'2 3850:L4a'b* 3826:L3h1b* 3794:L3d1a1 3631:Eurasia 3474:J2a2a1 3434:J1b1a1 3258:U5b1b* 3242:U5a1b1 3079:Origin 2985:E1b1b1c 2974:E1b1b1b 2908:E1b1b1a 2809:P (xQR) 2595:R1b1b1a 2499:Marker 2493:Origin 2451:Vietnam 2187:genetic 2108:  2100:  2090:  2040:  2017:  1987:  1957:  1954:  1946:  1931:  1918:  1915:  1902:  1674:(M173) 1668:(M207) 1603:(M214) 1481:Socotra 1408:Myanmar 1404:Burmese 1286:Hadzabe 1282:Pygmies 1272:Nilotes 1268:Khoisan 920:K-M2313 913:  854:  839:  825:  809:  800:  791:  613:  596:  543:updated 449:gametes 441:meiosis 439:during 375:nucleus 373:in the 220:alleles 205:haploûs 193:haploid 185:alleles 4719:  4709:  4665:  4655:  4629:Nature 4607:23 Jan 4585:  4575:  4526:  4485:  4416:  4406:  4363:  4353:  4311:  4301:  4235:  4228:155271 4225:  4217:  4164:  4144:  4042:L3b2a 3946:L2a1b 3930:L2a1* 3810:L3e1* 3802:L3d1d 3778:L0a2* 3770:L0a1b 3498:K1b1* 3442:J1b2a 3346:H13a1 3282:X2c'e 3279:Europe 3271:Europe 3266:U5b1c 3263:Europe 3255:Europe 3247:Europe 3239:Europe 3231:Europe 3223:Europe 3215:Europe 3207:Europe 3199:Europe 3191:Europe 3183:Europe 3175:Europe 3167:Europe 3159:Europe 3151:Europe 3143:Europe 3135:Europe 3127:Europe 3119:Europe 3111:Europe 3103:Europe 3095:Europe 3087:Europe 2897:E1b1b1 2864:R1b1b1 2861:Europe 2850:Europe 2697:M217+ 2686:Z1426 2669:Europe 2658:Europe 2647:Europe 2636:Europe 2625:Europe 2614:Europe 2603:Europe 2592:Europe 2581:Europe 2570:Europe 2559:Europe 2548:Europe 2537:Europe 2526:Europe 2515:Europe 2485:Mal'ta 2429:, and 2419:Nivkhs 2095:pre-JT 1880:, and 1729:Human 1638:(M122) 1624:(M119) 1495:, the 1493:Greece 1382:Iberia 1328:(M96) 1155:K-M526 1084:  1076:  943:  939:  934:  922:  903:  730:  722:  718:  713:  480:Human 400:Human 383:oocyte 322:  316:  310:  252:subset 201:ἁπλοῦς 4800:chart 4759:(PDF) 4425:(PDF) 4384:(PDF) 4050:L3d* 3986:L2b* 3978:L2a2 3906:L1c2 3898:L1c* 3890:L1b3 3882:L1b* 3842:L3x* 3834:L3i* 3818:L3f* 3762:L0a1 3722:M1a1 3706:L3e5 3698:X2b1 3650:U4a* 3506:N1a* 3490:K1a* 3458:J1d1 3370:H2a1 3354:H14a 3327:India 3274:U5b3 3250:U5b* 3218:U4c1 3202:T2b4 3194:T2b* 3186:J2b1 3178:J1c7 3098:H11a 3072:mtDNA 3032:M174 2856:M269 2853:R1b1b 2768:M526 2653:M529 2642:U152 2620:U106 2573:J2a2a 1928:L1–6 1731:mtDNA 1082:S2CID 732:GHIJK 379:sperm 209:group 197:Greek 141:ISOGG 4717:PMID 4663:PMID 4609:2015 4583:PMID 4524:PMID 4483:PMID 4414:PMID 4361:PMID 4309:PMID 4233:PMID 4215:ISSN 4162:ISBN 4142:ISBN 4018:L3b 4010:L2e 4002:L2d 3922:L2a 3914:L2* 3858:L5* 3754:L0* 3730:U6a 3690:X1a 3674:U9a 3666:U8* 3634:U3* 3626:U2e 3618:U2* 3610:U1* 3602:T2i 3594:T2c 3578:T1a 3570:T1* 3554:R0a 3522:N1c 3514:N1b 3466:J2a 3402:HV1 3386:H6b 3362:H20 3314:C/Z 3234:U5a 3226:U5* 3210:T2e 3146:H6a 3138:H5a 3122:H2a 3114:H1b 3106:H1a 3065:P58 3051:E1b1 3043:M33 2977:M81 2966:V65 2955:V22 2944:V32 2933:V12 2922:V13 2911:M78 2900:M35 2889:M70 2867:L23 2831:R1a1 2609:L11 2576:M92 2562:J2a2 2554:M47 2551:J2a1 2532:M26 2529:I1b2 2423:Nias 1784:pre- 1489:Oman 1308:YAP+ 1284:and 1270:and 1074:PMID 901:K2b1 746:HIJK 638:A1b1 594:A0-T 431:and 276:and 4912:DNA 4811:290 4707:PMC 4699:doi 4653:PMC 4645:doi 4633:505 4573:PMC 4563:doi 4514:doi 4473:doi 4404:PMC 4396:doi 4351:PMC 4343:doi 4299:PMC 4291:doi 4223:PMC 4207:doi 3866:L6 3714:M1 3658:U7 3642:U4 3586:T2 3546:R* 3538:N9 3530:N2 3482:K* 3410:I1 3394:H8 3378:H4 3338:H* 3170:I4 3154:H7 3130:H3 3090:H1 3040:E1a 2842:R1b 2584:J2b 2540:I1c 2518:I1b 1066:doi 984:P1a 979:P1b 974:P1c 962:NO1 890:K2a 882:K2b 875:K2c 868:K2d 861:K2e 755:IJK 628:A1b 622:A1a 588:A00 359:DNA 346:of 268:In 248:set 4908:: 4796:12 4715:. 4705:. 4695:23 4693:. 4689:. 4675:^ 4661:. 4651:. 4643:. 4631:. 4627:. 4581:. 4571:. 4561:. 4549:. 4545:. 4522:. 4508:. 4504:. 4481:. 4469:21 4467:. 4463:. 4412:. 4402:. 4392:75 4390:. 4386:. 4359:. 4349:. 4339:25 4337:. 4333:. 4321:^ 4307:. 4297:. 4287:48 4285:. 4279:. 4231:. 4221:. 4213:. 4203:12 4201:. 4197:. 4101:/ 3682:X 3562:T 3330:F 3306:B 3298:A 3290:I 3062:J1 3018:DE 2875:R2 2749:J2 2738:J1 2467:. 2443:: 2342:, 2338:, 2334:: 2325:, 2321:, 2317:, 2313:, 2298:, 2294:, 2290:, 2286:, 2282:, 2278:, 2274:, 2270:, 2255:, 2251:, 2247:, 2243:, 2239:, 2235:, 2231:, 2227:, 2219:L6 2217:, 2215:L5 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Index

Haplogroups
a series
Genetic genealogy
Population genetics
Haplogroup
Haplotype
Subclade
Most recent common ancestor
DNA haplogroups
Human mitochondrial
Human Y-chromosome
Genomics
Y-chromosome haplogroups by population
Genealogical DNA test
Surname DNA project
Personal genomics
Genographic Project
ISOGG
v
t
e
haplotype
alleles
haploid
Greek
single-nucleotide polymorphism
mutation
alleles
line of descent

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