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Human Y-chromosome DNA haplogroup

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29: 4846:"we date the Y-chromosomal most recent common ancestor (MRCA) in Africa at 254 (95% CI 192–307) kya and detect a cluster of major non-African founder haplogroups in a narrow time interval at 47–52 kya, consistent with a rapid initial colonization model of Eurasia and Oceania after the out-of-Africa bottleneck. In contrast to demographic reconstructions based on mtDNA, we infer a second strong bottleneck in Y-chromosome lineages dating to the last 10 ky. We hypothesize that this bottleneck is caused by cultural changes affecting variance of reproductive success among males." 308: 265: 3866: 2909: 2563: 2279: 2233: 1887: 256:). Although Southeast Asia has been proposed as the origin for all non-African human Y chromosomes, this hypothesis is considered unlikely. Other bottlenecks occurred roughly 50,000 and 5,000 years ago, and the majority of Eurasian men are believed to be descended from four ancestors who lived 50,000 years ago, all of whom were descendants of an African lineage (Haplogroup E-M168). 6717:
P.A. Underhill, N.M. Myres, S. Rootsi, C.T. Chow, A.A. Lin, R.P. Otillar, R. King, L.A. Zhivotovsky, O. Balanovsky, A. Pshenichnov, K.H. Ritchie, L.L. Cavalli-Sforza, T. Kivisild, R. Villems, S.R. Woodward, New Phylogenetic Relationships for Y-chromosome Haplogroup I: Reappraising its Phylogeography
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Y-DNA haplogroup nomenclature is changing over time to accommodate the increasing number of SNPs being discovered and tested, and the resulting expansion of the Y-chromosome phylogenetic tree. This change in nomenclature has resulted in inconsistent nomenclature being used in different sources. This
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Hallast, P.; Batini, C.; Zadik, D.; Maisano Delser, P.; Wetton, J. H.; Arroyo-Pardo, E.; Cavalleri, G. L.; De Knijff, P.; Destro Bisol, G.; Dupuy, B. M.; Eriksen, H. A.; Jorde, L. B.; King, T. E.; Larmuseau, M. H.; Lopez De Munain, A.; Lopez-Parra, A. M.; Loutradis, A.; Milasin, J.; Novelletto, A.;
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Around 7000 years ago - all the way back in the Neolithic - something really peculiar happened to human genetic diversity. Over the next 2,000 years, and seen across Africa, Europe and Asia, the genetic diversity of the Y chromosome collapsed, becoming as though there was only one man for every 17
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Trombetta, Beniamino; d'Atanasio, Eugenia; Massaia, Andrea; Ippoliti, Marco; Coppa, Alfredo; Candilio, Francesca; Coia, Valentina; Russo, Gianluca; Dugoujon, Jean-Michel; Moral, Pedro; Akar, Nejat; Sellitto, Daniele; Valesini, Guido; Novelletto, Andrea; Scozzari, Rosaria; Cruciani, Fulvio (2015).
6792:
Jones, Eppie R.; Gonzalez-Fortes, Gloria; Connell, Sarah; Siska, Veronika; Eriksson, Anders; Martiniano, Rui; McLaughlin, Russell L.; Gallego Llorente, Marcos; Cassidy, Lara M.; Gamba, Cristina; Meshveliani, Tengiz; Bar-Yosef, Ofer; MĂĽller, Werner; Belfer-Cohen, Anna; Matskevich, Zinovi; Jakeli,
6588:
Batini, Chiara; Hallast, Pille; Zadik, Daniel; Delser, Pierpaolo Maisano; Benazzo, Andrea; Ghirotto, Silvia; Arroyo-Pardo, Eduardo; Cavalleri, Gianpiero L.; De Knijff, Peter; Dupuy, Berit Myhre; Eriksen, Heidi A.; King, Turi E.; De Munain, Adolfo LĂłpez; LĂłpez-Parra, Ana M.; Loutradis, Aphrodite;
6139:
Batini, Chiara; Hallast, Pille; Zadik, Daniel; Delser, Pierpaolo Maisano; Benazzo, Andrea; Ghirotto, Silvia; Arroyo-Pardo, Eduardo; Cavalleri, Gianpiero L.; De Knijff, Peter; Dupuy, Berit Myhre; Eriksen, Heidi A.; King, Turi E.; De Munain, Adolfo LĂłpez; LĂłpez-Parra, Ana M.; Loutradis, Aphrodite;
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Since the first studies, it has been found that extant Native American populations exhibit almost exclusively five "mtDNA haplogroups" (A–D and X)6 classified in the autochthonous haplogroups A2, B2, C1, D1, and X2a.7 Haplogroups A–D are found all over the New World and are frequent in Asia,
7015:
Underhill, Peter A.; Myres, Natalie M.; Rootsi, Siiri; Metspalu, Mait; Zhivotovsky, Lev A.; King, Roy J.; Lin, Alice A.; Chow, Cheryl-Emiliane T.; Semino, Ornella; Battaglia, Vincenza; Kutuev, Ildus; Järve, Mari; Chaubey, Gyaneshwer; Ayub, Qasim; Mohyuddin, Aisha; Mehdi, S Qasim; Sengupta,
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K-M2313*, which as yet has no phylogenetic name, has been documented in two living individuals, who have ethnic ties to India and South East Asia. In addition, K-Y28299, which appears to be a primary branch of K-M2313, has been found in three living individuals from India. See: Poznik
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Sanghamitra; Rogaev, Evgeny I.; Khusnutdinova, Elza K.; Pshenichnov, Andrey; Balanovsky, Oleg; Balanovska, Elena; Jeran, Nina; Augustin, Dubravka Havas; Baldovic, Marian; Herrera, Rene J.; Thangaraj, Kumarasamy; Singh, Vijay; Singh, Lalji; et al. (2010).
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The defining mutations separating CT (all haplogroups except for A and B) are M168 and M294. The site of origin is likely in Africa. Its age has been estimated at approximately 88,000 years old, and more recently at around 100,000 or 101,000 years old.
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Di Cristofaro, Julie; Pennarun, Erwan; Mazières, Stéphane; Myres, Natalie M.; Lin, Alice A.; Temori, Shah Aga; Metspalu, Mait; Metspalu, Ene; Witzel, Michael; King, Roy J.; Underhill, Peter A.; Villems, Richard; Chiaroni, Jacques (2013).
1868:, about 48,000 years BP, and remains largely prevalent there in the forms of H1 (M69) and H3 (Z5857). Its sub-clades are also found in lower frequencies in Iran, Central Asia, across the middle-east, and the Arabian peninsula. 6919: 5673:
Fagundes, Nelson J.R.; Kanitz, Ricardo; Eckert, Roberta; Valls, Ana C.S.; Bogo, Mauricio R.; Salzano, Francisco M.; Smith, David Glenn; Silva, Wilson A.; Zago, Marco A.; Ribeiro-dos-Santos, Andrea K.; Santos, Sidney E.B.;
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Mendez, Fernando; Krahn, Thomas; Schrack, Bonnie; Krahn, Astrid-Maria; Veeramah, Krishna; Woerner, August; Fomine, Forka Leypey Mathew; Bradman, Neil; Thomas, Mark; Karafet, Tatiana M.; Hammer, Michael F. (7 March 2013).
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tree are provided below. ss4 bp, rs41352448, is not represented in the ISOGG 2008 tree because it is a value for an STR. This low frequency value has been found as a novel Q lineage (Q5) in Indian populations
295:). YCC shorthand nomenclature names Y-DNA haplogroups and their subclades with the first letter of the major Y-DNA haplogroup followed by a dash and the name of the defining terminal SNP. 6731:
Sharma, Swarkar; Rai, Ekta; Sharma, Prithviraj; Jena, Mamata; Singh, Shweta; Darvishi, Katayoon; Bhat, Audesh K.; Bhanwer, A J S.; Tiwari, Pramod Kumar; Bamezai, Rameshwar N K. (2009).
7334: 6479:"Phylogeographic Refinement and Large Scale Genotyping of Human y Chromosome Haplogroup e Provide New Insights into the Dispersal of Early Pastoralists in the African Continent" 291:(YCC) developed a system of naming major Y-DNA haplogroups with the capital letters A through T, with further subclades named using numbers and lower case letters (YCC longhand 6416:"Origin, diffusion, and differentiation of Y-chromosome haplogroups E and J: inferences on the neolithization of Europe and later migratory events in the Mediterranean area" 6362:"Origin, Diffusion, and Differentiation of Y-Chromosome Haplogroups E and J: Inferences on the Neolithization of Europe and Later Migratory Events in the Mediterranean Area" 7343: 4361: 3763: 7449: 6360:
Semino, Ornella; Magri, Chiara; Benuzzi, Giorgia; Lin, Alice A.; Al-Zahery, Nadia; Battaglia, Vincenza; MacCioni, Liliana; Triantaphyllidis, Costas; et al. (2004).
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populations toward the northeast in the Nile Valley. BT is a subclade of haplogroup A, more precisely of the A1b clade (A2-T in Cruciani et al. 2011), as follows:
4388:
Van Oven M, Van Geystelen A, Kayser M, Decorte R, Larmuseau HD (2014). "Seeing the wood for the trees: a minimal reference phylogeny for the human Y chromosome".
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Tumonggor, Meryanne K; Karafet, Tatiana M; Downey, Sean; Lansing, J Stephen; Norquest, Peter; Sudoyo, Herawati; Hammer, Michael F; Cox, Murray P (31 July 2014).
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Cruciani (2007), "Tracing Past Human Male Movements in Northern/Eastern Africa and Western Eurasia: New Clues from Y-Chromosomal Haplogroups E-M78 and J-M12",
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Found among Han Chinese (Guangdong, Hunan, and Shaanxi), Dai people (Yunnan), Murut people (Brunei), Malay people (Singapore), and Aeta people (Philippines)
7501: 7327: 1675:(M201) originated some 48,000 years ago and its most recent common ancestor likely lived 26,000 years ago in the Middle East. It spread to Europe with the 124: 225:(SNPs). The Y-chromosome accumulates approximately two mutations per generation, and Y-DNA haplogroups represent significant branches of the Y-chromosome 4992:
Y-chromosome DNA (Y-DNA) haplogroups are the major branches on the human paternal family tree. Each haplogroup has many subbranches. These are subclades.
4206: 4199: 4192: 1089: 1638:. F1 (P91), F2 (M427) and F3 (M481; previously F5) are all highly rare and virtually exclusive to regions/ethnic minorities in Sri Lanka, India, Nepal, 2973:. Its subclade R1b1a2 (M269) is the haplogroup that is most commonly found among modern Western European populations, and has been associated with the 1808: 1602:. One study, which did not comprehensively screen for other subclades of F-M89 (including some subclades of GHIJK), found that Indonesian men with the 7424: 1798: 1793: 1654:. In such cases, however, the possibility of misidentification is considered to be relatively high and some may belong to misidentified subclades of 7429: 7403: 7387: 4640: 4315: 4310: 5871:
Pamjav, H.; Sajantila, A.; Schempp, W.; Sears, M.; Tolun, A.; Tyler-Smith, C.; Van Geystelen, A.; Watkins, S.; Winney, B.; Jobling, M. A. (2015).
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Nino; Higham, Thomas F. G.; Currat, Mathias; Lordkipanidze, David; Hofreiter, Michael; Manica, Andrea; Pinhasi, Ron; Bradley, Daniel G. (2015).
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The groups descending from haplogroup F are found in some 90% of the world's population, but almost exclusively outside of sub-Saharan Africa.
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Underhill and Kivisild; Kivisild, T (2007). "Use of Y Chromosome and Mitochondrial DNA Population Structure in Tracing Human Migrations".
7465: 4657: 176: 7204: 7199: 36:. 'kya' means 'thousand years ago'. (b) Geographical distributions of haplogroups are shown in color. (c) Geographical color legend. 7566: 7382: 5750:"High-Resolution SNPs and Microsatellite Haplotypes Point to a Single, Recent Entry of Native American Y Chromosomes into the Americas" 3768: 2717: 299:
inconsistency, and increasingly cumbersome longhand nomenclature, has prompted a move toward using the simpler shorthand nomenclature.
150: 914:) macrohaplogroup from which all modern paternal haplogroups descend. It is sparsely distributed in Africa, being concentrated among 7556: 6853: 5679: 6589:
Milasin, Jelena; Novelletto, Andrea; Pamjav, Horolma; Sajantila, Antti; Tolun, Aslıhan; Winney, Bruce; Jobling, Mark A. (2015).
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Milasin, Jelena; Novelletto, Andrea; Pamjav, Horolma; Sajantila, Antti; Tolun, Aslıhan; Winney, Bruce; Jobling, Mark A. (2015).
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and Prehistory, in P. Mellars, K. Boyle, O. Bar-Yosef and C. Stringer (eds.), Rethinking the Human Evolution (2007), pp. 33–42.
6311:"A Rare Deep-Rooting D0 African Y-chromosomal Haplogroup and its Implications for the Expansion of Modern Humans Out of Africa" 5256:"A Rare Deep-Rooting D0 African Y-chromosomal Haplogroup and its Implications for the Expansion of Modern Humans Out of Africa" 4366: 5680:"Mitochondrial Population Genomics Supports a Single Pre-Clovis Origin with a Coastal Route for the Peopling of the Americas" 4788: 7534: 5431:"Different genetic components in the Norwegian population revealed by the analysis of mtDNA and Y chromosome polymorphisms" 4961: 7157: 6246: 7066: 6733:"The Indian origin of paternal haplogroup R1a1* substantiates the autochthonous origin of Brahmins and the caste system" 2574: 2290: 2244: 1898: 7561: 7438: 7103: 5435: 3788: 2826: 2094: 95: 17: 7366: 2716:
approximately 32,000 years ago. The subclades of Haplogroup Q with their defining mutation(s), according to the 2008
7618: 5030:"Understanding Results: Y-DNA Single Nucleotide Polymorphism (SNP): How are haplogroups and their subclades named?" 4980:"Understanding Results: Y-DNA Single Nucleotide Polymorphism (SNP): What is a Y-chromosome DNA (Y-DNA) haplogroup?" 2012: 4507:
Haplogroup K2b1 (P397/P399) is also known as Haplogroup MS, but has a broader and more complex internal structure.
7623: 7522: 7222: 7096:"An African American paternal lineage adds an extremely ancient root to the human Y chromosome phylogenetic tree" 3840: 273: 222: 169: 5626:"Improved phylogenetic resolution and rapid diversification of Y-chromosome haplogroup K-M526 in Southeast Asia" 268:
Schematic illustration of Y-DNA haplogroups naming convention. Haplogroups are defined through mutations (SNPs).
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Wen B, Li H, Lu D, et al. (September 2004). "Genetic evidence supports demic diffusion of Han culture".
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is the dominant haplogroup of Western Europe and is also found sparsely distributed among various peoples of
2004: 1270: 1103: 7307: 4553:
Haplogroup S, as of 2017, is also known as K2b1a. (Previously the name Haplogroup S was assigned to K2b1a4.)
6939: 6531:"Genetic Evidence of an East Asian Origin and Paleolithic Northward Migration of Y-chromosome Haplogroup N" 5873:"The Y-Chromosome Tree Bursts into Leaf: 13,000 High-Confidence SNPs Covering the Majority of Known Clades" 5811:
Sharma, Swarkar; Rai, Ekta; Bhat, Audesh K; Bhanwer, Amarjit S; Bamezaicorresponding, Rameshwar NK (2007).
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Haplogroup M, as of 2017, is also known as K2b1b. (Previously the name Haplogroup M was assigned to K2b1d.)
911: 3818: 7597: 7544: 7213: 6651:"Phylogeography of Y-Chromosome Haplogroup I Reveals Distinct Domains of Prehistoric Gene Flow in Europe" 2940: 237: 84: 6309:
Haber M, Jones AL, Connel BA, Asan, Arciero E, Huanming Y, Thomas MG, Xue Y, Tyler-Smith C (June 2019).
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Haber M, Jones AL, Connel BA, Asan, Arciero E, Huanming Y, Thomas MG, Xue Y, Tyler-Smith C (June 2019).
2646:. Considered together, these distributions tend to suggest that P* emerged from K2b in South East Asia. 7312: 244:. Y-chromosomal Adam is estimated to have lived around 236,000 years ago in Africa. By examining other 162: 44: 7539: 5152:"New binary polymorphisms reshape and increase resolution of the human Y chromosomal haplogroup tree" 3773: 3266: 1959:
save for I2B1 (m223) which is found at a moderate frequency in Western, Central, and Northern Europe.
1813: 1518: 1489: 1478: 1472: 1370: 1202: 1013: 461: 5017:
A terminal SNP determines the terminal (final) subbranch on the Y-DNA Tree to which someone belongs.
28: 7551: 7302: 6691: 5415:
This was, for instance, the case with the original subclade F3 (M96), which has since been renamed
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Semino O, Passarino G, Oefner PJ, et al. (November 2000). "The genetic legacy of Paleolithic
7018:"Separating the post-Glacial coancestry of European and Asian y chromosomes within haplogroup R1a" 5675: 4684: 4232: 2501: 2443: 2439: 2412: 1871:
However, H2 (P96) is present in Europe since the Neolithic and H1a1 (M82) spread westward in the
1788: 1457: 710: 288: 7194: 4221: 4213: 3953: 3003: 2994: 2974: 2944: 2749: 2269: 2117: 2105: 1827: 1822: 1451: 1442: 1186: 700: 523: 245: 6529:
Shi, Hong; Qi, Xuebin; Zhong, Hua; Peng, Yi; Zhang, Xiaoming; Ma, Runlin Z.; Su, Bing (2013).
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Hallast, Pille; Agdzhoyan, Anastasia; Balanovsky, Oleg; Xue, Yali; Tyler-Smith, Chris (2021).
4661: 4463:
F-Y27277, sometimes known as F2'4, is both the parent clade of F2 and F4 and a child of F-M89.
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Passarino G, Cavalleri GL, Lin AA, Cavalli-Sforza LL, Børresen-Dale AL, Underhill PA (2002).
4634: 4283: 4129: 3808: 3783: 3176: 3140: 3100: 3082: 3029: 2985: 2820: 2608: 2523: 2447: 2419: 2390: 2330:
Haplogroup N possibly originated in eastern Asia and spread both northward and westward into
2222: 2218: 2026: 2000: 1952: 1943: 1778: 1773: 1559: 1436: 1141: 1033: 992: 972: 960: 651: 641: 533: 448: 399: 389: 364: 206: 130: 7255: 6966: 6806: 6602: 6542: 6153: 6044: 5989: 5934: 5824: 5321: 5125: 5004: 4322: 4241: 4176: 4120: 4105: 4091: 4012: 3984: 3340: 3230: 3212: 3194: 3158: 3118: 2917: 2903: 2704: 2659: 2653: 2604: 2599: 2547: 2537: 2347: 2320: 2315: 2140: 2077: 1974: 1969: 1923: 1851: 1676: 1672: 1667: 1574: 1423: 1417: 1314: 1254: 1248: 1007: 966: 926: 787: 772: 759: 738: 723: 679: 664: 620: 605: 582: 561: 546: 511: 486: 473: 427: 412: 377: 352: 218: 5029: 4979: 4588: 1232: 8: 7208: 6955:"Wave-of-Advance Models of the Diffusion of the y Chromosome Haplogroup R1b1b2 in Europe" 6591:"Large-scale recent expansion of European patrilineages shown by population resequencing" 6262:"A recent bottleneck of Y chromosome diversity coincides with a global change in culture" 6142:"Large-scale recent expansion of European patrilineages shown by population resequencing" 5472:
Karlsson, Andreas O; Wallerström, Thomas; Götherström, Anders; Holmlund, Gunilla (2006).
5204:"A recent bottleneck of Y chromosome diversity coincides with a global change in culture" 4806:"A recent bottleneck of Y chromosome diversity coincides with a global change in culture" 3833: 3798: 2435: 2408: 1293: 145: 135: 66: 7259: 7189: 6970: 6810: 6606: 6546: 6157: 6048: 5993: 5938: 5828: 5325: 3879:
Please help update this article to reflect recent events or newly available information.
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men trace their descent from a man who lived in Africa approximately 69,000 years ago (
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Karafet, TM; Mendez, FL; Meilerman, MB; Underhill, PA; Zegura, SL; Hammer, MF (2008).
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This article is about the human Y-DNA haplogroup. For the human mtDNA haplogroup, see
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Found in Central Asia, South Asia, and Central, Northern and Eastern Europe, Balkans
2709: 2630:(or Agta) people of Luzon. While, P1* is now more common among living individuals in 2478: 2352: 2023:
and also found in Mediterranean Europe in smaller frequencies much like haplogroup T.
1986: 1928: 1536: 1362: 1328: 1306: 1283: 1260: 1152: 226: 140: 52: 33: 7138: 4417: 2137:(M20) is found in South Asia, Central Asia, South-West Asia, and the Mediterranean. 7263: 7236: 7120: 7112: 7037: 7029: 6984: 6974: 6900: 6822: 6814: 6744: 6673: 6665: 6618: 6610: 6560: 6550: 6498: 6490: 6435: 6427: 6381: 6373: 6332: 6322: 6281: 6273: 6218: 6210: 6169: 6161: 6111: 6103: 6092:"Y-chromosome evidence for differing ancient demographic histories in the Americas" 6062: 6052: 6017: 5997: 5952: 5942: 5892: 5884: 5842: 5832: 5761: 5702: 5694: 5645: 5637: 5596: 5588: 5526: 5515:"Phylogeographic analysis of paternal lineages in NE Portuguese Jewish communities" 5485: 5444: 5388: 5380: 5339: 5329: 5306:
Chiaroni, Jacques; Underhill, Peter A.; Cavalli-Sforza, Luca L. (1 December 2009).
5275: 5267: 5223: 5215: 5171: 5163: 5121: 4827: 4819: 4749: 4733: 4692: 4612: 4602: 4397: 4305: 4063: 4017: 3529: 3322: 2631: 2541: 2335: 2324: 2134: 1655: 1499: 1344: 1224: 1115: 1054: 7287: 7267: 2662:(R-M207). These share the common marker M45 in addition to at least 18 other SNPs. 2451: 7226: 6979: 6555: 5947: 4810: 4077: 3660: 2761: 2639: 2172: 2082: 1735: 1723: 1591: 1278: 1042: 498: 307: 6327: 5271: 2626:. In particular, P* and P1* are found at significant rates among members of the 264: 7292: 7161: 7116: 6214: 5698: 5577:"Isolation, contact and social behavior shaped genetic diversity in West Timor" 5369:"Isolation, contact and social behavior shaped genetic diversity in West Timor" 5313:
Proceedings of the National Academy of Sciences of the United States of America
4737: 4607: 4293: 4086: 3919: 3826: 3793: 3746: 3730: 3714: 3609: 3593: 3472: 3453: 2962: 2956: 2933: 2929: 2786: 2404: 2396: 2360: 2144: 2030: 1524: 1336: 1324: 1302: 1099: 931: 276: 241: 190: 1626:. F* (F xF1,F2,F3) has been reported among 10% of males in Sri Lanka and 7612: 6926: 6033:"The Eurasian heartland: a continental perspective on Y-chromosome diversity" 4745: 4538: 4486: 4168: 4162: 4148: 4137: 4100: 4003: 3998: 3989: 3975: 3936: 3396: 2921: 2886: 2882: 2871: 2842: 2838: 2753: 2515: 2208: 2192: 2188: 2180: 2160: 2129: 1731: 1631: 1382: 1228: 1212: 1027: 942: 936: 592: 253: 232:
The Y-chromosomal most recent common ancestor (Y-MRCA), often referred to as
6905: 5888: 5837: 5766: 5749: 5556: 5490: 5473: 5449: 5430: 5334: 7275: 7134: 7051: 6998: 6940:"The Evatt Clan: A Worldwide Historical Review of the Evatt Family Surname" 6914: 6836: 6758: 6687: 6632: 6574: 6512: 6449: 6395: 6346: 6295: 6232: 6183: 6125: 6076: 6057: 6009: 5966: 5906: 5856: 5775: 5716: 5659: 5610: 5540: 5499: 5458: 5402: 5353: 5289: 5237: 5185: 5133: 4841: 4763: 4626: 4409: 3848: 3625: 3017: 3013: 2948: 2846: 2790: 2713: 2688: 2635: 2376: 2368: 2062: 2020: 1872: 1857: 1766: 1528: 1398: 292: 214: 7219: 7033: 6277: 5641: 5219: 4962:"Something Weird Happened to Men 7,000 Years Ago, And We Finally Know Why" 4823: 4722:"A Southeast Asian origin for present-day non-African human Y chromosomes" 4697: 4591:"A Southeast Asian origin for present-day non-African human Y chromosomes" 7303:
Y-DNA Ethnographic and Genographic Atlas and Open-Source Data Compilation
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The Prehistory of Language: A Triangulated Y-Chromosome-Based Perspective
3698: 3679: 3644: 3386: 3367: 3330: 3294: 3275: 3238: 3184: 3166: 3126: 2773: 2757: 2627: 2623: 2527: 2484: 2156: 1978: 1876: 1639: 1627: 1532: 1429: 1366: 1332: 7342: 7174:(chart highlighting new branches added to the A phylotree in March 2013) 6001: 5592: 5384: 5308:"Y chromosome diversity, human expansion, drift, and cultural evolution" 4445:
Haplogroup A0-T is also known as A-L1085 (and previously as A0'1'2'3'4).
2562: 2278: 2232: 1886: 229:, each characterized by hundreds or even thousands of unique mutations. 6818: 6749: 6732: 6614: 6165: 5531: 5514: 5167: 4401: 2952: 2908: 2692: 2643: 2551: 2387:
Haplogroup O1 (F265/M1354, CTS2866, F75/M1297, F429/M1415, F465/M1422)
2372: 2196: 2152: 2058: 1861: 1750: 1607: 1583: 1173: 1165: 1095: 198: 90: 71: 6475: 5471: 5249: 5247: 5197: 5195: 4531: 4485:
was known as "Haplogroup K2". That name has since been re-assigned to
2323:(M231) is found in northern Eurasia, especially among speakers of the 7297: 7077:
A Nomenclature system for the Tree of Human Y-Chromosomal Haplogroups
5923:"Afghan Hindu Kush: Where Eurasian Sub-Continent Gene Flows Converge" 4542:, 2017, "Details of the Y-SNP markers included in the minimal Y tree" 3009: 2684: 2511: 2507: 2488: 2467: 2356: 2221:. Major studies published in 2014 and 2015 suggest that up to 27% of 2214: 2113: 2101: 1865: 1758: 1587: 1310: 1190: 1177: 1169: 1161: 1064: 316: 202: 75: 6669: 6431: 6377: 6107: 6090:
Bortolini MC, Salzano FM, Thomas MG, et al. (September 2003).
5428: 5244: 5192: 4589:
Hallast P, Agdzhoyan A, Balanovsky O, Xue Y, Tyler-Smith C (2020).
4498:
Haplogroup K2b (M1221/P331/PF5911) is also known as Haplogroup MPS.
3021: 2806: 2777: 2673: 2100:
K(xLT,K2a,K2b) – that is, K*, K2c, K2d or K2e – is found mainly in
2054: 2034: 2016: 2008: 1982: 1936: 1762: 1754: 1719: 1699: 1691: 1683: 1643: 1619: 1595: 1495: 287:, the SNP furthest down in the Y-chromosome phylogenetic tree. The 280: 272:
Y-DNA haplogroups are defined by the presence of a series of Y-DNA
249: 107: 79: 7232:
Video tutorial on Discovering Paternal Ancestry with Y-Chromosomes
6795:"Upper Palaeolithic genomes reveal deep roots of modern Eurasians" 5919: 5813:"A novel subgroup Q5 of human Y-chromosomal haplogroup Q in India" 5005:"myFTDNA 2.0 User Guide: Y-DNA: What is the Y-DNA – Matches page?" 4387: 217:(Y-DNA). Individuals within a haplogroup share similar numbers of 6031:
Wells RS, Yuldasheva N, Ruzibakiev R, et al. (August 2001).
3481: 3462: 3148: 2925: 2794: 2765: 2669: 2364: 2331: 2164: 2090: 2086: 2042: 1994: 1727: 1703: 1682:
It is found in many ethnic groups in Eurasia; most common in the
1651: 1623: 1615: 1606:
P14/PF2704 (which is equivalent to M89), comprise 1.8% of men in
1599: 1540: 1394: 1220: 1216: 1208: 1068: 1023: 919: 915: 7195:
Chart of the speed of different Y chromosomal STR mutation rates
6791: 5869: 5748:
Zegura, S. L.; Karafet, TM; Zhivotovsky, LA; Hammer, MF (2003).
4783:. Stuttgart, Germany: Genetic–Linguistic Interface. p. 29. 2912:
The hypothetical divergence of Haplogroup R and its descendants.
5305: 5149: 4652: 4650: 3561: 3545: 3513: 3497: 3443: 3424: 3256: 3108: 3090: 3072: 3034:
Found in South Asia, Caucasus, Central Asia, and Eastern Europe
2970: 2680: 2400: 2168: 2038: 1932: 1707: 1695: 1611: 1060: 7184: 5789: 5747: 5111: 4676: 7293:
Kerchner's YDNA Haplogroup Descriptions, Projects & Links
6199:"The Divergence of Neandertal and Modern Human Y Chromosomes" 5574: 4939: 4719: 3057: 2814: 2810: 2785:
Found at low to moderate frequency among some populations of
2712:
is defined by the SNP M242. It is believed to have arisen in
2619: 2463: 2431: 2184: 2109: 2046: 1746: 1742: 1715: 1711: 1698:. Found in almost all European countries, but most common in 1647: 1582:
F xG,H,I,J,K is rare in modern populations and peaks in
1402: 1350: 1274: 1266: 1072: 1048: 7014: 5474:"Y-chromosome diversity in Sweden – A long-time perspective" 4647: 3764:
List of Y-chromosome haplogroups in populations of the world
2191:. It is also found at low frequencies in other parts of the 1347:(M55, M57, M64.1, M179, P12, P37.1, P41.1 (M359.1), 12f2.2) 4658:"Understanding Haplogroups: How are the haplogroups named?" 3577: 3405: 3348: 3312: 3220: 3202: 3130: 3039: 2966: 2176: 2050: 1687: 1635: 1019: 4431:
International Society of Genetic Genealogy (ISOGG; 2015),
6471: 6469: 6467: 6465: 6463: 6461: 6459: 6414:
Semino, O; Magri, C; Benuzzi, G; et al. (May 2004).
6030: 5672: 4472:
Haplogroup LT (L298/P326) is also known as Haplogroup K1.
210: 6587: 6138: 5736:
supporting a northeastern Asian origin of these lineages
2618:
P*, P1* and P2 are found together only on the island of
7092: 6260:
Kamin M, Saag L, Vincente M, et al. (April 2015).
5202:
Kamin M, Saag L, Vincente M, et al. (April 2015).
2143:(M184, M70, M193, M272) is found at high levels in the 6456: 6089: 2540:(B254) also known as K2b1a: found in the highlands of 2263: 2225:
males carry K2*, while others carry a subclade of K2.
2029:
Syrid/Nahrainid Arabids (M172) is found mainly in the
1993:
Haplogroup J* (J-M304*) is rare outside the island of
1957:
Found mainly in Balkans, southeast Europe and Sardinia
1741:
G-M201 is also found in small numbers in northwestern
311:
Migration route of Human Y-chromosome DNA haplogroups.
32:
Human Y-DNA phylogeny and haplogroup distribution.(a)
7241: 6359: 2916:
Haplogroup R is defined by the SNP M207. The bulk of
2003:
Semitid/Bedouinid Arabids (M267) are associated with
7502:
List of haplogroups of historical and famous figures
7344:
Y-chromosome haplogroups in populations of the world
6644: 6642: 6524: 6522: 6308: 5810: 5253: 5105: 2163:. T-M184 is also found in significant minorities of 1630:, 5% in Pakistan, as well as lower levels among the 240:
from whom all currently living humans are descended
6730: 4582: 7246:in extant Europeans: a Y chromosome perspective". 7010: 7008: 6413: 2649:P1 is also the parent node of two primary clades: 2213:The only living males reported to carry the basal 7298:Y-DNA Testing Company STR Marker Comparison Chart 6639: 6519: 6259: 5623: 5201: 2533:Its primary subclades are two major haplogroups: 1132:Found with low frequency in Saudi Arabia and Iraq 7610: 5913: 5078: 2483:Found throughout East Asia, Southeast Asia, and 7005: 5301: 5299: 1590:. It also appears to have long been present in 6952: 6528: 5145: 5143: 4544:(Access date of these pages: 9 December 2017) 2495: 2355:(M175) is found with its highest frequency in 7593:List of Y-DNA single-nucleotide polymorphisms 7328: 5422: 4934: 4932: 4930: 4928: 4926: 4924: 4922: 4920: 4918: 4916: 4914: 4912: 4910: 4908: 4906: 4904: 4902: 4900: 4898: 4896: 4894: 4892: 4890: 4888: 4886: 4884: 4882: 4880: 4878: 4876: 4874: 4872: 4710:"one mutation in every 30 million base pairs" 3834: 3814:Conversion table for Y chromosome haplogroups 3779:List of Y-DNA single-nucleotide polymorphisms 2638:, it is also found at low levels in mainland 170: 7072: 6886: 6884: 6882: 6880: 6878: 6247:"The father of all men is 340,000 years old" 5792:. International Society of Genetic Genealogy 5617: 5570: 5568: 5552: 5550: 5465: 5296: 5074: 5072: 4870: 4868: 4866: 4864: 4862: 4860: 4858: 4856: 4854: 4852: 4639:: CS1 maint: multiple names: authors list ( 2334:, being the most common group found in some 1977:(M304, S6, S34, S35) is found mainly in the 5863: 5559:Y-DNA Haplogroup P and its Subclades – 2016 5140: 5097:: CS1 maint: numeric names: authors list ( 4516:Haplogroup P (P295) is also klnown as K2b2. 4381: 2947:peoples, and is now found predominantly in 2510:K2b1* have been identified. Males carrying 2123: 1594:; it has been reported at rates of 4–5% in 1535:, and Europe (especially in areas near the 7335: 7321: 7062: 6848: 6846: 6353: 6302: 6190: 4954: 3841: 3827: 3769:Y-DNA haplogroups in populations of Europe 2920:is represented in the descendant subclade 900: 177: 163: 7124: 7041: 6988: 6978: 6904: 6875: 6826: 6748: 6677: 6622: 6564: 6554: 6502: 6439: 6409: 6407: 6405: 6385: 6336: 6326: 6285: 6222: 6173: 6115: 6066: 6056: 5956: 5946: 5896: 5846: 5836: 5765: 5706: 5649: 5600: 5565: 5547: 5530: 5519:American Journal of Physical Anthropology 5489: 5448: 5392: 5366: 5343: 5333: 5279: 5227: 5175: 5069: 4849: 4831: 4797: 4776: 4753: 4696: 4616: 4606: 4424: 2434:, China, Taiwan, Southeast Asia, and the 2202: 1242: 302: 6890: 5666: 5512: 5081:"ISOGG 2015 Y-DNA Haplogroup Tree Trunk" 5053:"New Y DNA Haplogroup Naming Convention" 4457: 4454:Haplogroup A1 is also known as A1'2'3'4. 3040:Chronological development of haplogroups 2907: 2881:Found at low frequency among samples of 2395:Found in eastern, central, and southern 306: 263: 27: 7288:Y-DNA Haplogroup and Sub-clade Projects 6953:Sjödin, Per; François, Olivier (2011). 6843: 5979: 5624:Tatiana M Karafet; et al. (2015). 4519: 2837:Found among some indigenous peoples of 2393:(M119, CTS31, F589/Page20, L246, L466) 1856:Haplogroup H (M69) probably emerged in 986: 951:Haplogroup A1b (also A2-T; P108, V221) 7611: 6726: 6724: 6713: 6711: 6648: 6402: 6366:The American Journal of Human Genetics 6203:The American Journal of Human Genetics 6196: 5741: 5126:10.1146/annurev.genet.41.110306.130407 4803: 4682: 2897: 2593: 1963: 1917: 1661: 1379:Haplogroup D1b (L1366, L1378, M226.2) 1001: 905: 125:Y-chromosome haplogroups by population 7316: 6937: 5050: 3822: 2550:(P256) also known as K2b1b: found in 1845: 1568: 1411: 4501: 4439: 4367:Y-DNA haplogroups of historic people 4342: 4337: 4328: 4326: 4321: 4319: 4314: 4309: 4304: 4298: 4291: 4281: 4279: 4269: 4261: 4249: 4247: 4238: 4230: 4188: 4181: 4175: 4173: 4167: 4161: 4159: 4153: 4147: 4135:       4125: 4119: 4113: 4104: 4099: 4090: 4085: 4076: 4071: 4035: 4021: 3904: 3859: 2698: 2557: 2273: 2227: 2071: 1881: 1730:with highest concentrations on some 259: 7284:Paper that defined "Eu" haplogroups 7067:2005 Y-chromosome Phylogenetic Tree 6721: 6708: 6649:Rootsi, Siiri; et al. (2004). 4492: 4009: 3949: 3932: 3123:65,200, 69,000, or 73,000 years ago 2928:. R1 has two descendant subclades: 2870:Found in a significant minority of 2264:Haplogroups K2a, K2a1, NO & NO1 13: 7158:"Y-Haplogroup A Phylogenetic Tree" 7104:American Journal of Human Genetics 7086: 7022:European Journal of Human Genetics 6658:American Journal of Human Genetics 5687:American Journal of Human Genetics 5630:European Journal of Human Genetics 5478:European Journal of Human Genetics 5436:European Journal of Human Genetics 4475: 4466: 4448: 3789:Human mitochondrial DNA haplogroup 2827:indigenous peoples of the Americas 2514:of K2b1 are found primarily among 1477:Found in Africa, especially among 18:Human mitochondrial DNA haplogroup 14: 7635: 7588:Human Y-chromosome DNA haplogroup 7178: 6854:"Mesolithic Western Eurasian DNA" 4660:. Family Tree DNA. Archived from 4556: 4547: 4510: 4436:. (Access date: 1 February 2015.) 2607:(P295) has two primary branches: 2013:Semitic languages speaking people 1428:Found in Africa and parts of the 918:populations in the southwest and 195:human Y-chromosome DNA haplogroup 7190:Family Tree DNA Public Haplotree 6197:Mendez, L.; et al. (2016). 5057:DNAeXplained - Genetic Genealogy 3864: 2990:Found throughout western Eurasia 2561: 2363:, with lower frequencies in the 2277: 2231: 1955:Dinarid/Dinaric Europids (P215) 1931:(M170, M258) is found mainly in 1885: 954:Haplogroup A1b1a1 (also A2; M14) 7220:DNA Heritage's Y-haplogroup map 6946: 6931: 6893:Molecular Biology and Evolution 6785: 6776: 6765: 6581: 6253: 6239: 6132: 6083: 6024: 5973: 5877:Molecular Biology and Evolution 5804: 5782: 5754:Molecular Biology and Evolution 5506: 5409: 5360: 5044: 5022: 4997: 4489:, the sibling of Haplogroup LT. 3105:88,000 or 101-100,000 years ago 2748:Found with low frequency among 2666:Haplogroup Q (MEH2, M242, P36) 2611:(P-M45) and the extremely rare 2341: 2309: 1948:Found mainly in northern Europe 1494:Found in Africa, especially in 1083:Haplogroup C1b1 (AM00694/K281) 1080:Haplogroup C1b (F1370, Z16480) 957:Haplogroup A1b1b (also A3; M32) 274:single-nucleotide polymorphisms 223:single-nucleotide polymorphisms 5367:Tumonggor, Meryanne K (2014). 4972: 4770: 4713: 4685:"Human mutation rate revealed" 1946:Nordid/Nordic Europids (M253) 1183:Haplogroup C1b3b (M347, P309) 1086:Haplogroup C1b1a (B66/Z16458) 1: 7268:10.1126/science.290.5494.1155 5678:; Bonatto, Sandro L. (2008). 5051:Estes, Roberta (2013-03-31). 4576: 2151:-speaking peoples), parts of 7160:. March 2013. Archived from 6980:10.1371/journal.pone.0021592 6556:10.1371/journal.pone.0066102 6483:Genome Biology and Evolution 6037:Proc. Natl. Acad. Sci. U.S.A 5948:10.1371/journal.pone.0076748 5790:"Y-DNA Haplogroup Tree 2010" 4804:Karmin; et al. (2015). 3851:Y-chromosome DNA haplogroups 3069:235,900 or 275,000 years ago 2924:(M173), which originated in 1094:Found with low frequency in 1059:Found with low frequency in 1047:Found with low frequency in 963:(M91, M42, M94, M139, M299) 7: 7598:Paternal mtDNA transmission 7545:African admixture in Europe 7185:ISOGG Y-DNA Haplogroup Tree 6938:Evatt, Danny (2013-11-01). 6328:10.1534/genetics.119.302368 5272:10.1534/genetics.119.302368 4777:St. Clair, Michael (2021). 3901: 3849:Phylogenetic tree of human 3757: 2939:R1a is associated with the 2677:Haplogroup R (M207, M306): 2496:Haplogroups K2b1, M & S 1130:Haplogroup C1b1a3 (Z16582) 238:most recent common ancestor 85:Most recent common ancestor 10: 7640: 7205:Atlas of the Human Journey 7117:10.1016/j.ajhg.2013.02.002 6215:10.1016/j.ajhg.2016.02.023 5699:10.1016/j.ajhg.2007.11.013 4738:10.1007/s00439-020-02204-9 4608:10.1007/s00439-020-02204-9 4433:Y-DNA Haplogroup Tree 2015 4335: 4302: 4259: 4228: 4219: 4211: 4204: 4197: 4190: 4145: 4111: 4097: 4083: 4069: 4053: 4037: 3906: 2901: 2805:Found at low frequency in 2702: 2597: 2499: 2383:Madagascar, the Comoros). 2345: 2313: 2267: 2206: 2127: 2075: 1967: 1921: 1875:with the migration of the 1849: 1665: 1572: 1415: 1290:Haplogroup D1a (CTS11577) 1281: 1246: 1149:Haplogroup C1b2 (C-Z16582) 1122:Haplogroup C1b1a2b (F725) 1039:Haplogroup C1a (CTS11043) 1005: 990: 319:tree of Y-DNA haplogroups 15: 7575: 7510: 7494: 7474: 7458: 7417: 7396: 7375: 7359: 7350: 6737:Journal of Human Genetics 5581:Journal of Human Genetics 5373:Journal of Human Genetics 4356: 4134: 4127: 4061: 4044: 4028: 4023: 4016: 4011: 4002: 3997: 3995: 3988: 3983: 3981: 3974: 3967: 3965: 3958: 3951: 3941: 3934: 3924: 3917: 3915: 3899: 3873:This article needs to be 3857: 3774:Genetic history of Europe 3053:Possible place of origin 3008:Found in Western Europe, 2855:Q1a3a3 (M199, P106, P292) 2506:No examples of the basal 1830:(formerly Haplogroup G5) 1296:(Z27276, Z27283, Z29263) 1112:Haplogroup C1b1a2a (B67) 910:Haplogroup A is the NRY ( 785: 770: 763: 736: 721: 714: 704: 677: 662: 655: 645: 618: 603: 596: 586: 559: 544: 537: 527: 509: 502: 484: 477: 459: 452: 425: 410: 403: 393: 375: 368: 350: 343: 328: 7069:, from FamilyTreeDNA.com 5676:Petzl-Erler, Maria Luiza 3050:Possible time of origin 2124:Haplogroups L and T (K1) 2085:(M9) is spread all over 2031:Semitic-speaking peoples 1391:Haplogroup D2 (A5580.2) 1109:Haplogroup C1b1a2 (B65) 7619:Human Y-DNA haplogroups 7450:East and Southeast Asia 7308:Y Chromosome Consortium 5838:10.1186/1471-2148-7-232 5513:Nogueiro, InĂŞs (2009). 5491:10.1038/sj.ejhg.5201651 5450:10.1038/sj.ejhg.5200834 5335:10.1073/pnas.0910803106 4481:Between 2002 and 2008, 2502:Haplogroup K2b1 (Y-DNA) 2336:Uralic-speaking peoples 1321:Haplogroup D1a1b (P99) 1299:Haplogroup D1a1a (M15) 1215:, but especially among 1158:Haplogroup C1b3a (M38) 1138:Haplogroup C1b1b (B68) 901:Major Y-DNA haplogroups 289:Y Chromosome Consortium 7624:Genetics-related lists 7435:Central and North Asia 6058:10.1073/pnas.171305098 5079:Copyright 2015 ISOGG. 2913: 2746:Q1a1 (M120, M265/N14) 2524:indigenous Australians 2270:Haplogroup K2a (Y-DNA) 2219:indigenous Australians 2203:Haplogroup K2 (K-M526) 2118:Island South East Asia 2106:Aboriginal Australians 1502:-speaking populations. 1481:-speaking populations. 1243:Haplogroup D (CTS3946) 312: 303:Phylogenetic structure 269: 246:population bottlenecks 37: 7511:Genetic history of... 7034:10.1038/ejhg.2009.194 6906:10.1093/molbev/msm049 6799:Nature Communications 6595:Nature Communications 6278:10.1101/gr.186684.114 6146:Nature Communications 5889:10.1093/molbev/msu327 5767:10.1093/molbev/msh009 5642:10.1038/ejhg.2014.106 5220:10.1101/gr.186684.114 4824:10.1101/gr.186684.114 4698:10.1038/news.2009.864 4683:Dolgin, Elie (2009). 3809:Genealogical DNA test 3784:List of Y-STR markers 2911: 2375:, and islands in the 2223:Aboriginal Australian 1858:Southern Central Asia 1734:islands; uncommon in 1142:Dusun people (Brunei) 993:Haplogroup CT (Y-DNA) 310: 267: 213:on the male-specific 131:Genealogical DNA test 31: 7353:in populations of... 7244:Homo sapiens sapiens 7237:Haplogroup Predictor 7200:Map of Y Haplogroups 7144:on 24 September 2019 2904:Haplogroup R (Y-DNA) 2724:The 2008 ISOGG tree 2705:Haplogroup Q (Y-DNA) 2600:Haplogroup P (Y-DNA) 2470:, and Southeast Asia 2348:Haplogroup O (Y-DNA) 2316:Haplogroup N (Y-DNA) 2078:Haplogroup K (Y-DNA) 2015:in the Middle East, 1970:Haplogroup J (Y-DNA) 1924:Haplogroup I (Y-DNA) 1852:Haplogroup H (Y-DNA) 1677:Neolithic Revolution 1668:Haplogroup G (Y-DNA) 1575:Haplogroup F (Y-DNA) 1418:Haplogroup E (Y-DNA) 1249:Haplogroup D (Y-DNA) 1118:in Borneo, Indonesia 1008:Haplogroup C (Y-DNA) 987:Haplogroup CT (P143) 948:Haplogroup A1a (M31) 219:short tandem repeats 201:defined by specific 7260:2000Sci...290.1155S 7214:National Geographic 7209:Genographic Project 6971:2011PLoSO...621592S 6811:2015NatCo...6.8912J 6607:2015NatCo...6.7152B 6547:2013PLoSO...866102S 6158:2015NatCo...6.7152B 6049:2001PNAS...9810244W 6002:10.1038/nature02878 5994:2004Natur.431..302W 5939:2013PLoSO...876748D 5829:2007BMCEE...7..232S 5593:10.1038/jhg.2014.62 5385:10.1038/jhg.2014.62 5326:2009PNAS..10620174C 4362:Y-DNA by population 4277:    3799:Molecular phylogeny 2898:Haplogroup R (M207) 2594:Haplogroup P (K2b2) 2520:Micronesian peoples 2446:-speaking peoples, 2438:, especially among 2436:Indian subcontinent 2407:, especially among 2005:Northeast Caucasian 1964:Haplogroup J (M304) 1918:Haplogroup I (M170) 1662:Haplogroup G (M201) 1002:Haplogroup C (M130) 939:(formerly also A1b) 906:Haplogroups A and B 146:Genographic Project 136:Surname DNA project 96:Human mitochondrial 67:Population genetics 25:Human DNA groupings 7583:Y-chromosomal Adam 7409:Sub-Saharan Africa 7225:2006-02-18 at the 6920:Supplementary Data 6858:Ancestral Journeys 6819:10.1038/ncomms9912 6750:10.1038/jhg.2008.2 6615:10.1038/ncomms8152 6495:10.1093/gbe/evv118 6166:10.1038/ncomms8152 5532:10.1002/ajpa.21154 5168:10.1101/gr.7172008 4402:10.1002/humu.22468 4128:    3909:Y-chromosomal Adam 3077:235,900 years ago 2941:proto-Indo-Iranian 2914: 2573:. You can help by 2289:. You can help by 2243:. You can help by 2187:(Eivissencs), and 1897:. You can help by 1846:Haplogroup H (M69) 1569:Haplogroup F (M89) 1412:Haplogroup E (M96) 1363:Andamanese peoples 1187:indigenous peoples 331:Y-chromosomal Adam 313: 270: 234:Y-chromosomal Adam 101:Human Y-chromosome 38: 7606: 7605: 7557:Iberian Peninsula 7495:Famous haplotypes 7490: 7489: 7351:Y-DNA haplogroups 7254:(5494): 1155–59. 7164:on 18 August 2018 6420:Am. J. Hum. Genet 6096:Am. J. Hum. Genet 5032:. Family Tree DNA 5007:. Family Tree DNA 4982:. Family Tree DNA 4790:978-3-00-071045-2 4573: 4572: 4535:, 2017, "K-M2335" 4533:YFull YTree v5.08 4483:Haplogroup T-M184 4351: 4350: 4296: 4286: 4276: 4267: 4255: 4244: 4236: 4224: 4216: 4209: 4202: 4195: 4140: 4059: 4051: 4042: 3972: 3956: 3939: 3922: 3894: 3893: 3804:Genetic genealogy 3755: 3754: 3582:11,900 years ago 3566:12,100 years ago 3550:13,500 years ago 3534:13,400 years ago 3518:18,500 years ago 3502:21,100 years ago 3486:20,400 years ago 3467:18,300 years ago 3448:21,800 years ago 3410:22,800 years ago 3391:27,800 years ago 3372:18,500 years ago 3353:28,200 years ago 3335:24,100 years ago 3317:22,100 years ago 3299:40,100 years ago 3280:34,800 years ago 3261:27,500 years ago 3243:31,600 years ago 3225:31,900 years ago 3207:45,400 years ago 3189:42,900 years ago 3171:26,200 years ago 3153:48,800 years ago 3135:53,100 years ago 3113:68,500 years ago 3095:88,000 years ago 3087:130,700 years ago 2783:Q1a2 (M25, M143) 2699:Haplogroup Q M242 2591: 2590: 2307: 2306: 2261: 2260: 2072:Haplogroup K (M9) 1987:South-East Europe 1915: 1914: 1814:Haplogroup G2a3b1 1519:Haplogroup E1b1b1 1490:Haplogroup E1b1a2 1473:Haplogroup E1b1a1 1284:Haplogroup D-M174 1235:-speaking peoples 1207:Found throughout 1090:Haplogroup C1b1a1 1045:(M8, M105, M131) 912:non-recombining Y 897: 896: 888: 887: 879: 878: 870: 869: 861: 860: 852: 851: 843: 842: 834: 833: 825: 824: 816: 815: 807: 806: 798: 797: 749: 748: 690: 689: 631: 630: 572: 571: 438: 437: 283:are defined by a 260:Naming convention 227:phylogenetic tree 187: 186: 141:Personal genomics 53:Genetic genealogy 34:Phylogenetic tree 7631: 7357: 7356: 7337: 7330: 7323: 7314: 7313: 7279: 7173: 7171: 7169: 7153: 7151: 7149: 7143: 7137:. Archived from 7128: 7100: 7075: 7065: 7056: 7055: 7045: 7012: 7003: 7002: 6992: 6982: 6950: 6944: 6943: 6935: 6929: 6917: 6908: 6888: 6873: 6872: 6870: 6869: 6860:. Archived from 6850: 6841: 6840: 6830: 6789: 6783: 6780: 6774: 6769: 6763: 6762: 6752: 6728: 6719: 6715: 6706: 6705: 6703: 6702: 6696: 6690:. Archived from 6681: 6655: 6646: 6637: 6636: 6626: 6585: 6579: 6578: 6568: 6558: 6526: 6517: 6516: 6506: 6473: 6454: 6453: 6443: 6411: 6400: 6399: 6389: 6357: 6351: 6350: 6340: 6330: 6321:(4): 1421–1428. 6306: 6300: 6299: 6289: 6257: 6251: 6250: 6243: 6237: 6236: 6226: 6194: 6188: 6187: 6177: 6136: 6130: 6129: 6119: 6087: 6081: 6080: 6070: 6060: 6043:(18): 10244–49. 6028: 6022: 6021: 5988:(7006): 302–05. 5977: 5971: 5970: 5960: 5950: 5917: 5911: 5910: 5900: 5867: 5861: 5860: 5850: 5840: 5808: 5802: 5801: 5799: 5797: 5786: 5780: 5779: 5769: 5745: 5739: 5738: 5732: 5731: 5725: 5719:. Archived from 5710: 5684: 5670: 5664: 5663: 5653: 5621: 5615: 5614: 5604: 5572: 5563: 5554: 5545: 5544: 5534: 5510: 5504: 5503: 5493: 5469: 5463: 5462: 5452: 5426: 5420: 5413: 5407: 5406: 5396: 5364: 5358: 5357: 5347: 5337: 5320:(48): 20174–79. 5303: 5294: 5293: 5283: 5251: 5242: 5241: 5231: 5199: 5190: 5189: 5179: 5147: 5138: 5137: 5114:Annu. Rev. Genet 5109: 5103: 5102: 5096: 5088: 5076: 5067: 5066: 5064: 5063: 5048: 5042: 5041: 5039: 5037: 5026: 5020: 5019: 5014: 5012: 5001: 4995: 4994: 4989: 4987: 4976: 4970: 4969: 4958: 4952: 4951: 4949: 4947: 4936: 4847: 4845: 4835: 4801: 4795: 4794: 4774: 4768: 4767: 4757: 4717: 4711: 4709: 4707: 4705: 4700: 4680: 4674: 4673: 4671: 4669: 4654: 4645: 4644: 4638: 4630: 4620: 4610: 4586: 4563: 4560: 4554: 4551: 4545: 4523: 4517: 4514: 4508: 4505: 4499: 4496: 4490: 4479: 4473: 4470: 4464: 4461: 4455: 4452: 4446: 4443: 4437: 4428: 4422: 4421: 4385: 4292: 4282: 4270: 4262: 4250: 4240: 4231: 4220: 4212: 4205: 4198: 4191: 4136: 4055: 4046: 4038: 3968: 3952: 3935: 3918: 3902: 3889: 3886: 3880: 3868: 3867: 3860: 3843: 3836: 3829: 3820: 3819: 3751:4,700 years ago 3735:4,600 years ago 3719:4,800 years ago 3703:5,300 years ago 3684:6,400 years ago 3665:4,800 years ago 3649:8,500 years ago 3630:9,900 years ago 3622:11,800 years ago 3614:6,400 years ago 3606:13,300 years ago 3598:8,500 years ago 3590:14,100 years ago 3574:14,200 years ago 3558:17,400 years ago 3542:18,500 years ago 3530:Northeast Africa 3526:19,800 years ago 3510:21,100 years ago 3494:21,800 years ago 3478:22,800 years ago 3459:22,800 years ago 3440:27,500 years ago 3429:4,600 years ago 3421:27,500 years ago 3402:28,200 years ago 3383:31,600 years ago 3364:31,600 years ago 3345:31,900 years ago 3327:34,800 years ago 3309:36,800 years ago 3291:42,300 years ago 3272:42,300 years ago 3253:42,900 years ago 3235:42,900 years ago 3217:45,400 years ago 3199:47,200 years ago 3181:47,200 years ago 3163:48,500 years ago 3145:65,900 years ago 3044: 3043: 2586: 2583: 2565: 2558: 2542:Papua New Guinea 2462:Found in Japan, 2325:Uralic languages 2302: 2299: 2281: 2274: 2256: 2253: 2235: 2228: 2095:Native Americans 1910: 1907: 1889: 1882: 1809:Haplogroup G2a3b 1804:Haplogroup G2a3a 1656:Haplogroup GHIJK 1634:(Nepal), and in 1513:Haplogroup E1b1b 1467:Haplogroup E1b1a 1349:Found mainly in 1323:Found mainly in 1301:Found mainly in 1225:Tungusic peoples 1185:Found among the 766: 765: 717: 716: 707: 706: 658: 657: 648: 647: 599: 598: 589: 588: 540: 539: 530: 529: 505: 504: 480: 479: 455: 454: 406: 405: 396: 395: 371: 370: 346: 345: 324: 323: 179: 172: 165: 40: 39: 7639: 7638: 7634: 7633: 7632: 7630: 7629: 7628: 7609: 7608: 7607: 7602: 7571: 7506: 7486: 7470: 7454: 7413: 7392: 7371: 7352: 7346: 7341: 7227:Wayback Machine 7181: 7167: 7165: 7156: 7147: 7145: 7141: 7098: 7089: 7087:Further reading 7059: 7013: 7006: 6951: 6947: 6936: 6932: 6889: 6876: 6867: 6865: 6852: 6851: 6844: 6790: 6786: 6781: 6777: 6770: 6766: 6729: 6722: 6716: 6709: 6700: 6698: 6694: 6653: 6647: 6640: 6586: 6582: 6527: 6520: 6474: 6457: 6412: 6403: 6358: 6354: 6307: 6303: 6266:Genome Research 6258: 6254: 6245: 6244: 6240: 6195: 6191: 6137: 6133: 6088: 6084: 6029: 6025: 5978: 5974: 5918: 5914: 5868: 5864: 5809: 5805: 5795: 5793: 5788: 5787: 5783: 5746: 5742: 5729: 5727: 5723: 5682: 5671: 5667: 5622: 5618: 5573: 5566: 5562:(20 June 2016). 5555: 5548: 5511: 5507: 5470: 5466: 5427: 5423: 5414: 5410: 5365: 5361: 5304: 5297: 5252: 5245: 5208:Genome Research 5200: 5193: 5156:Genome Research 5148: 5141: 5110: 5106: 5090: 5089: 5077: 5070: 5061: 5059: 5049: 5045: 5035: 5033: 5028: 5027: 5023: 5010: 5008: 5003: 5002: 4998: 4985: 4983: 4978: 4977: 4973: 4964:. 31 May 2018. 4960: 4959: 4955: 4945: 4943: 4938: 4937: 4850: 4811:Genome Research 4802: 4798: 4791: 4775: 4771: 4718: 4714: 4703: 4701: 4681: 4677: 4667: 4665: 4664:on 21 June 2012 4656: 4655: 4648: 4632: 4631: 4587: 4583: 4579: 4574: 4569: 4568: 4567: 4566: 4561: 4557: 4552: 4548: 4524: 4520: 4515: 4511: 4506: 4502: 4497: 4493: 4480: 4476: 4471: 4467: 4462: 4458: 4453: 4449: 4444: 4440: 4429: 4425: 4386: 4382: 4377: 4371: 4352: 3895: 3890: 3884: 3881: 3878: 3869: 3865: 3853: 3847: 3760: 3743:4,700 years ago 3727:5,000 years ago 3711:5,800 years ago 3695:6,400 years ago 3676:7,400 years ago 3661:Southern Europe 3657:8,100 years ago 3641:9,200 years ago 3042: 3020:, and northern 2906: 2900: 2707: 2701: 2640:South East Asia 2632:Eastern Siberia 2602: 2596: 2587: 2581: 2578: 2571:needs expansion 2504: 2498: 2460:(SRY465, M176) 2458:Haplogroup O1b2 2426:Haplogroup O1b1 2350: 2344: 2318: 2312: 2303: 2297: 2294: 2287:needs expansion 2272: 2266: 2257: 2251: 2248: 2241:needs expansion 2211: 2205: 2132: 2126: 2080: 2074: 1972: 1966: 1926: 1920: 1911: 1905: 1902: 1895:needs expansion 1854: 1848: 1834:Haplogroup G2c1 1799:Haplogroup G2a3 1794:Haplogroup G2a2 1789:Haplogroup G2a1 1736:Northern Europe 1702:, southeastern 1670: 1664: 1592:South East Asia 1577: 1571: 1460:(P2, DYS391p); 1458:Haplogroup E1b1 1420: 1414: 1357:Haplogroup D1a3 1345:Haplogroup D1a2 1329:Qiangic peoples 1307:Qiangic peoples 1294:Haplogroup D1a1 1286: 1279:Andaman Islands 1251: 1245: 1153:Haplogroup C1b3 1102:, and northern 1055:Haplogroup C1a2 1043:Haplogroup C1a1 1010: 1004: 995: 989: 908: 903: 898: 889: 880: 871: 862: 853: 844: 835: 826: 817: 808: 799: 750: 691: 632: 573: 439: 305: 277:genetic markers 262: 183: 127: 104: 99: 93: 91:DNA haplogroups 26: 21: 12: 11: 5: 7637: 7627: 7626: 7621: 7604: 7603: 7601: 7600: 7595: 7590: 7585: 7579: 7577: 7573: 7572: 7570: 7569: 7564: 7559: 7554: 7549: 7548: 7547: 7537: 7532: 7527: 7526: 7525: 7514: 7512: 7508: 7507: 7505: 7504: 7498: 7496: 7492: 7491: 7488: 7487: 7485: 7484: 7478: 7476: 7472: 7471: 7469: 7468: 7462: 7460: 7456: 7455: 7453: 7452: 7447: 7442: 7432: 7427: 7421: 7419: 7415: 7414: 7412: 7411: 7406: 7400: 7398: 7394: 7393: 7391: 7390: 7385: 7379: 7377: 7373: 7372: 7370: 7369: 7363: 7361: 7354: 7348: 7347: 7340: 7339: 7332: 7325: 7317: 7311: 7310: 7305: 7300: 7295: 7290: 7285: 7239: 7234: 7229: 7217: 7202: 7197: 7192: 7187: 7180: 7179:External links 7177: 7176: 7175: 7154: 7088: 7085: 7084: 7083: 7070: 7058: 7057: 7004: 6945: 6930: 6925:2012-12-05 at 6899:(6): 1300–11, 6874: 6842: 6784: 6775: 6764: 6720: 6707: 6670:10.1086/422196 6638: 6580: 6518: 6489:(7): 1940–50. 6455: 6432:10.1086/386295 6426:(5): 1023–34. 6401: 6378:10.1086/386295 6372:(5): 1023–34. 6352: 6301: 6272:(4): 459–466. 6252: 6238: 6189: 6131: 6108:10.1086/377588 6082: 6023: 5972: 5933:(10): e76748. 5912: 5862: 5803: 5781: 5740: 5665: 5616: 5587:(9): 494–503. 5564: 5546: 5505: 5464: 5421: 5408: 5379:(9): 494–503. 5359: 5295: 5266:(4): 1421–28. 5243: 5191: 5139: 5104: 5068: 5043: 5021: 4996: 4971: 4953: 4848: 4796: 4789: 4769: 4732:(2): 299–307. 4726:Human Genetics 4712: 4675: 4646: 4601:(2): 299–307. 4595:Human Genetics 4580: 4578: 4575: 4571: 4570: 4565: 4564: 4555: 4546: 4518: 4509: 4500: 4491: 4474: 4465: 4456: 4447: 4438: 4423: 4390:Human Mutation 4379: 4378: 4373: 4372: 4370: 4369: 4364: 4358: 4357: 4354: 4353: 4349: 4348: 4346: 4341: 4336: 4333: 4332: 4327: 4325: 4320: 4318: 4313: 4308: 4303: 4300: 4299: 4297: 4290: 4288: 4280: 4278: 4268: 4260: 4257: 4256: 4248: 4246: 4237: 4229: 4226: 4225: 4218: 4210: 4203: 4196: 4189: 4187: 4180: 4174: 4172: 4166: 4160: 4158: 4152: 4146: 4143: 4142: 4133: 4126: 4124: 4118: 4112: 4109: 4108: 4103: 4098: 4095: 4094: 4089: 4084: 4081: 4080: 4075: 4070: 4067: 4066: 4060: 4052: 4043: 4036: 4033: 4032: 4027: 4022: 4020: 4015: 4010: 4007: 4006: 4001: 3996: 3993: 3992: 3987: 3982: 3979: 3978: 3973: 3966: 3963: 3962: 3957: 3950: 3947: 3946: 3940: 3933: 3930: 3929: 3923: 3916: 3913: 3912: 3905: 3900: 3897: 3896: 3892: 3891: 3872: 3870: 3863: 3858: 3855: 3854: 3846: 3845: 3838: 3831: 3823: 3817: 3816: 3811: 3806: 3801: 3796: 3794:* (haplogroup) 3791: 3786: 3781: 3776: 3771: 3766: 3759: 3756: 3753: 3752: 3749: 3747:Eastern Europe 3744: 3741: 3737: 3736: 3733: 3731:Eastern Europe 3728: 3725: 3721: 3720: 3717: 3715:Eastern Europe 3712: 3709: 3705: 3704: 3701: 3696: 3693: 3686: 3685: 3682: 3677: 3674: 3667: 3666: 3663: 3658: 3655: 3651: 3650: 3647: 3642: 3639: 3632: 3631: 3628: 3623: 3620: 3616: 3615: 3612: 3610:Eastern Europe 3607: 3604: 3600: 3599: 3596: 3594:Eastern Europe 3591: 3588: 3584: 3583: 3580: 3575: 3572: 3568: 3567: 3564: 3559: 3556: 3552: 3551: 3548: 3543: 3540: 3536: 3535: 3532: 3527: 3524: 3520: 3519: 3516: 3511: 3508: 3504: 3503: 3500: 3495: 3492: 3488: 3487: 3484: 3479: 3476: 3469: 3468: 3465: 3460: 3457: 3450: 3449: 3446: 3441: 3438: 3431: 3430: 3427: 3422: 3419: 3412: 3411: 3408: 3403: 3400: 3393: 3392: 3389: 3384: 3381: 3374: 3373: 3370: 3365: 3362: 3355: 3354: 3351: 3346: 3343: 3337: 3336: 3333: 3328: 3325: 3319: 3318: 3315: 3310: 3307: 3301: 3300: 3297: 3292: 3289: 3282: 3281: 3278: 3273: 3270: 3263: 3262: 3259: 3254: 3251: 3245: 3244: 3241: 3236: 3233: 3227: 3226: 3223: 3218: 3215: 3209: 3208: 3205: 3200: 3197: 3191: 3190: 3187: 3182: 3179: 3173: 3172: 3169: 3164: 3161: 3155: 3154: 3151: 3146: 3143: 3137: 3136: 3133: 3124: 3121: 3115: 3114: 3111: 3106: 3103: 3097: 3096: 3093: 3088: 3085: 3079: 3078: 3075: 3070: 3067: 3061: 3060: 3054: 3051: 3048: 3041: 3038: 3037: 3036: 3027: 3026: 3025: 3004:Haplogroup R1b 3001: 2995:Haplogroup R1a 2963:Haplogroup R1b 2957:Eastern Europe 2902:Main article: 2899: 2896: 2895: 2894: 2893: 2892: 2891: 2890: 2877: 2876: 2875: 2866: 2863: 2860: 2859: 2858: 2857: 2856: 2853: 2850: 2841:, such as the 2833: 2818: 2798: 2787:Southwest Asia 2781: 2744: 2738: 2732: 2703:Main article: 2700: 2697: 2664: 2663: 2657: 2598:Main article: 2595: 2592: 2589: 2588: 2582:September 2016 2568: 2566: 2556: 2555: 2554:and Melanesia. 2545: 2516:Papuan peoples 2500:Main article: 2497: 2494: 2493: 2492: 2476: 2475: 2474: 2473: 2472: 2455: 2420:Haplogroup O1b 2417: 2405:Southeast Asia 2397:Mainland China 2391:Haplogroup O1a 2361:Southeast Asia 2346:Main article: 2343: 2340: 2314:Main article: 2311: 2308: 2305: 2304: 2298:September 2016 2284: 2282: 2268:Main article: 2265: 2262: 2259: 2258: 2252:September 2016 2238: 2236: 2207:Main article: 2204: 2201: 2145:Horn of Africa 2128:Main article: 2125: 2122: 2076:Main article: 2073: 2070: 2069: 2068: 2067: 2066: 2024: 1968:Main article: 1965: 1962: 1961: 1960: 1950: 1922:Main article: 1919: 1916: 1913: 1912: 1906:September 2016 1892: 1890: 1850:Main article: 1847: 1844: 1843: 1842: 1841: 1840: 1839: 1838: 1837: 1836: 1828:Haplogroup G2c 1825: 1823:Haplogroup G2b 1820: 1819: 1818: 1817: 1816: 1806: 1796: 1791: 1784:Haplogroup G2a 1776: 1666:Main article: 1663: 1660: 1573:Main article: 1570: 1567: 1566: 1565: 1564: 1563: 1557: 1556: 1555: 1554: 1553: 1552: 1551: 1550: 1549: 1525:Horn of Africa 1510: 1509: 1508: 1487: 1452:Haplogroup E1b 1449: 1443:Haplogroup E1a 1416:Main article: 1413: 1410: 1409: 1408: 1407: 1406: 1389: 1388: 1387: 1377: 1376: 1375: 1354: 1342: 1341: 1340: 1337:Turkic peoples 1319: 1282:Main article: 1247:Main article: 1244: 1241: 1240: 1239: 1238: 1237: 1229:Paleosiberians 1200: 1199: 1198: 1197: 1196: 1195: 1194: 1181: 1155:(B477/Z31885) 1150: 1147: 1146: 1145: 1136: 1135: 1134: 1128: 1127: 1126: 1120: 1107: 1100:Southwest Asia 1078: 1077: 1076: 1052: 1036:(F3393/Z1426) 1006:Main article: 1003: 1000: 991:Main article: 988: 985: 984: 983: 982: 981: 980: 979: 978: 977: 976: 975: 970: 958: 955: 949: 940: 934: 932:Haplogroup A00 907: 904: 902: 899: 895: 894: 891: 890: 886: 885: 882: 881: 877: 876: 873: 872: 868: 867: 864: 863: 859: 858: 855: 854: 850: 849: 846: 845: 841: 840: 837: 836: 832: 831: 828: 827: 823: 822: 819: 818: 814: 813: 810: 809: 805: 804: 801: 800: 796: 795: 792: 791: 784: 781: 780: 777: 776: 769: 764: 762: 756: 755: 752: 751: 747: 746: 743: 742: 735: 732: 731: 728: 727: 720: 715: 713: 705: 703: 697: 696: 693: 692: 688: 687: 684: 683: 676: 673: 672: 669: 668: 661: 656: 654: 646: 644: 638: 637: 634: 633: 629: 628: 625: 624: 617: 614: 613: 610: 609: 602: 597: 595: 587: 585: 579: 578: 575: 574: 570: 569: 566: 565: 558: 555: 554: 551: 550: 543: 538: 536: 528: 526: 520: 519: 516: 515: 508: 503: 501: 495: 494: 491: 490: 483: 478: 476: 470: 469: 466: 465: 458: 453: 451: 445: 444: 441: 440: 436: 435: 432: 431: 424: 421: 420: 417: 416: 409: 404: 402: 394: 392: 386: 385: 382: 381: 374: 369: 367: 361: 360: 357: 356: 349: 344: 342: 339: 338: 335: 334: 327: 322: 321: 320: 304: 301: 261: 258: 191:human genetics 185: 184: 182: 181: 174: 167: 159: 156: 155: 154: 153: 148: 143: 138: 133: 128: 123: 118: 117: 116:Related topics 113: 112: 111: 110: 105: 89: 87: 82: 69: 61: 60: 56: 55: 49: 48: 24: 9: 6: 4: 3: 2: 7636: 7625: 7622: 7620: 7617: 7616: 7614: 7599: 7596: 7594: 7591: 7589: 7586: 7584: 7581: 7580: 7578: 7576:Main articles 7574: 7568: 7565: 7563: 7560: 7558: 7555: 7553: 7550: 7546: 7543: 7542: 7541: 7538: 7536: 7535:British Isles 7533: 7531: 7528: 7524: 7521: 7520: 7519: 7516: 7515: 7513: 7509: 7503: 7500: 7499: 7497: 7493: 7483: 7480: 7479: 7477: 7473: 7467: 7464: 7463: 7461: 7457: 7451: 7448: 7446: 7443: 7440: 7436: 7433: 7431: 7428: 7426: 7423: 7422: 7420: 7416: 7410: 7407: 7405: 7402: 7401: 7399: 7395: 7389: 7386: 7384: 7381: 7380: 7378: 7374: 7368: 7365: 7364: 7362: 7358: 7355: 7349: 7345: 7338: 7333: 7331: 7326: 7324: 7319: 7318: 7315: 7309: 7306: 7304: 7301: 7299: 7296: 7294: 7291: 7289: 7286: 7283: 7277: 7273: 7269: 7265: 7261: 7257: 7253: 7249: 7245: 7240: 7238: 7235: 7233: 7230: 7228: 7224: 7221: 7218: 7216: 7215: 7210: 7206: 7203: 7201: 7198: 7196: 7193: 7191: 7188: 7186: 7183: 7182: 7163: 7159: 7155: 7140: 7136: 7132: 7127: 7122: 7118: 7114: 7111:(3): 454–59. 7110: 7106: 7105: 7097: 7091: 7090: 7082: 7078: 7074: 7071: 7068: 7064: 7061: 7060: 7053: 7049: 7044: 7039: 7035: 7031: 7028:(4): 479–84. 7027: 7023: 7019: 7011: 7009: 7000: 6996: 6991: 6986: 6981: 6976: 6972: 6968: 6965:(6): e21592. 6964: 6960: 6956: 6949: 6941: 6934: 6928: 6927:archive.today 6924: 6921: 6916: 6912: 6907: 6902: 6898: 6894: 6887: 6885: 6883: 6881: 6879: 6864:on 2016-04-07 6863: 6859: 6855: 6849: 6847: 6838: 6834: 6829: 6824: 6820: 6816: 6812: 6808: 6804: 6800: 6796: 6788: 6779: 6773: 6768: 6760: 6756: 6751: 6746: 6742: 6738: 6734: 6727: 6725: 6714: 6712: 6697:on 2009-06-19 6693: 6689: 6685: 6680: 6675: 6671: 6667: 6664:(1): 128–37. 6663: 6659: 6652: 6645: 6643: 6634: 6630: 6625: 6620: 6616: 6612: 6608: 6604: 6600: 6596: 6592: 6584: 6576: 6572: 6567: 6562: 6557: 6552: 6548: 6544: 6541:(6): e66102. 6540: 6536: 6532: 6525: 6523: 6514: 6510: 6505: 6500: 6496: 6492: 6488: 6484: 6480: 6472: 6470: 6468: 6466: 6464: 6462: 6460: 6451: 6447: 6442: 6437: 6433: 6429: 6425: 6421: 6417: 6410: 6408: 6406: 6397: 6393: 6388: 6383: 6379: 6375: 6371: 6367: 6363: 6356: 6348: 6344: 6339: 6334: 6329: 6324: 6320: 6316: 6312: 6305: 6297: 6293: 6288: 6283: 6279: 6275: 6271: 6267: 6263: 6256: 6248: 6242: 6234: 6230: 6225: 6220: 6216: 6212: 6209:(4): 728–34. 6208: 6204: 6200: 6193: 6185: 6181: 6176: 6171: 6167: 6163: 6159: 6155: 6151: 6147: 6143: 6135: 6127: 6123: 6118: 6113: 6109: 6105: 6102:(3): 524–39. 6101: 6097: 6093: 6086: 6078: 6074: 6069: 6064: 6059: 6054: 6050: 6046: 6042: 6038: 6034: 6027: 6019: 6015: 6011: 6007: 6003: 5999: 5995: 5991: 5987: 5983: 5976: 5968: 5964: 5959: 5954: 5949: 5944: 5940: 5936: 5932: 5928: 5924: 5916: 5908: 5904: 5899: 5894: 5890: 5886: 5883:(3): 661–73. 5882: 5878: 5874: 5866: 5858: 5854: 5849: 5844: 5839: 5834: 5830: 5826: 5822: 5818: 5817:BMC Evol Biol 5814: 5807: 5791: 5785: 5777: 5773: 5768: 5763: 5760:(1): 164–75. 5759: 5755: 5751: 5744: 5737: 5726:on 2009-03-25 5722: 5718: 5714: 5709: 5704: 5700: 5696: 5693:(3): 583–92. 5692: 5688: 5681: 5677: 5669: 5661: 5657: 5652: 5647: 5643: 5639: 5636:(3): 369–73. 5635: 5631: 5627: 5620: 5612: 5608: 5603: 5598: 5594: 5590: 5586: 5582: 5578: 5571: 5569: 5561: 5560: 5557:ISOGG, 2016, 5553: 5551: 5542: 5538: 5533: 5528: 5525:(3): 373–81. 5524: 5520: 5516: 5509: 5501: 5497: 5492: 5487: 5484:(8): 963–70. 5483: 5479: 5475: 5468: 5460: 5456: 5451: 5446: 5443:(9): 521–29. 5442: 5438: 5437: 5432: 5425: 5418: 5417:Haplogroup H2 5412: 5404: 5400: 5395: 5390: 5386: 5382: 5378: 5374: 5370: 5363: 5355: 5351: 5346: 5341: 5336: 5331: 5327: 5323: 5319: 5315: 5314: 5309: 5302: 5300: 5291: 5287: 5282: 5277: 5273: 5269: 5265: 5261: 5257: 5250: 5248: 5239: 5235: 5230: 5225: 5221: 5217: 5214:(4): 459–66. 5213: 5209: 5205: 5198: 5196: 5187: 5183: 5178: 5173: 5169: 5165: 5162:(5): 830–38. 5161: 5157: 5153: 5146: 5144: 5135: 5131: 5127: 5123: 5120:(1): 539–64. 5119: 5115: 5108: 5100: 5094: 5086: 5082: 5075: 5073: 5058: 5054: 5047: 5031: 5025: 5018: 5006: 5000: 4993: 4981: 4975: 4968: 4963: 4957: 4941: 4940:"YFull YTree" 4935: 4933: 4931: 4929: 4927: 4925: 4923: 4921: 4919: 4917: 4915: 4913: 4911: 4909: 4907: 4905: 4903: 4901: 4899: 4897: 4895: 4893: 4891: 4889: 4887: 4885: 4883: 4881: 4879: 4877: 4875: 4873: 4871: 4869: 4867: 4865: 4863: 4861: 4859: 4857: 4855: 4853: 4843: 4839: 4834: 4829: 4825: 4821: 4818:(4): 459–66. 4817: 4813: 4812: 4807: 4800: 4792: 4786: 4782: 4781: 4773: 4765: 4761: 4756: 4751: 4747: 4743: 4739: 4735: 4731: 4727: 4723: 4716: 4699: 4694: 4690: 4686: 4679: 4663: 4659: 4653: 4651: 4642: 4636: 4628: 4624: 4619: 4614: 4609: 4604: 4600: 4596: 4592: 4585: 4581: 4559: 4550: 4543: 4541: 4536: 4534: 4529: 4522: 4513: 4504: 4495: 4488: 4484: 4478: 4469: 4460: 4451: 4442: 4435: 4434: 4427: 4419: 4415: 4411: 4407: 4403: 4399: 4396:(2): 187–91. 4395: 4391: 4384: 4380: 4376: 4368: 4365: 4363: 4360: 4359: 4355: 4347: 4345: 4340: 4334: 4331: 4324: 4317: 4312: 4307: 4301: 4295: 4289: 4287:   4285: 4274: 4265: 4258: 4253: 4243: 4234: 4227: 4223: 4217:   4215: 4208: 4201: 4194: 4185: 4179:   4178: 4170: 4165:   4164: 4156: 4151:   4150: 4144: 4139: 4131: 4122: 4117:   4116: 4110: 4107: 4102: 4096: 4093: 4088: 4082: 4079: 4074: 4068: 4065: 4058: 4049: 4041: 4034: 4031: 4026: 4019: 4014: 4008: 4005: 4000: 3994: 3991: 3986: 3980: 3977: 3971: 3964: 3961: 3955: 3948: 3944: 3938: 3931: 3927: 3921: 3914: 3910: 3903: 3898: 3888: 3885:February 2021 3876: 3871: 3862: 3861: 3856: 3852: 3844: 3839: 3837: 3832: 3830: 3825: 3824: 3821: 3815: 3812: 3810: 3807: 3805: 3802: 3800: 3797: 3795: 3792: 3790: 3787: 3785: 3782: 3780: 3777: 3775: 3772: 3770: 3767: 3765: 3762: 3761: 3750: 3748: 3745: 3742: 3739: 3738: 3734: 3732: 3729: 3726: 3723: 3722: 3718: 3716: 3713: 3710: 3707: 3706: 3702: 3700: 3697: 3694: 3691: 3688: 3687: 3683: 3681: 3678: 3675: 3672: 3669: 3668: 3664: 3662: 3659: 3656: 3653: 3652: 3648: 3646: 3643: 3640: 3637: 3634: 3633: 3629: 3627: 3624: 3621: 3618: 3617: 3613: 3611: 3608: 3605: 3602: 3601: 3597: 3595: 3592: 3589: 3586: 3585: 3581: 3579: 3576: 3573: 3570: 3569: 3565: 3563: 3560: 3557: 3554: 3553: 3549: 3547: 3544: 3541: 3538: 3537: 3533: 3531: 3528: 3525: 3522: 3521: 3517: 3515: 3512: 3509: 3506: 3505: 3501: 3499: 3496: 3493: 3491:I2-L460 (I2a) 3490: 3489: 3485: 3483: 3480: 3477: 3474: 3471: 3470: 3466: 3464: 3461: 3458: 3455: 3452: 3451: 3447: 3445: 3442: 3439: 3436: 3433: 3432: 3428: 3426: 3423: 3420: 3417: 3414: 3413: 3409: 3407: 3404: 3401: 3398: 3395: 3394: 3390: 3388: 3385: 3382: 3379: 3376: 3375: 3371: 3369: 3366: 3363: 3360: 3357: 3356: 3352: 3350: 3347: 3344: 3342: 3339: 3338: 3334: 3332: 3329: 3326: 3324: 3321: 3320: 3316: 3314: 3311: 3308: 3306: 3303: 3302: 3298: 3296: 3293: 3290: 3287: 3284: 3283: 3279: 3277: 3274: 3271: 3268: 3265: 3264: 3260: 3258: 3255: 3252: 3250: 3247: 3246: 3242: 3240: 3237: 3234: 3232: 3229: 3228: 3224: 3222: 3219: 3216: 3214: 3211: 3210: 3206: 3204: 3201: 3198: 3196: 3193: 3192: 3188: 3186: 3183: 3180: 3178: 3175: 3174: 3170: 3168: 3165: 3162: 3160: 3157: 3156: 3152: 3150: 3147: 3144: 3142: 3139: 3138: 3134: 3132: 3128: 3125: 3122: 3120: 3117: 3116: 3112: 3110: 3107: 3104: 3102: 3099: 3098: 3094: 3092: 3089: 3086: 3084: 3081: 3080: 3076: 3074: 3071: 3068: 3066: 3063: 3062: 3059: 3055: 3052: 3049: 3046: 3045: 3035: 3031: 3030:Haplogroup R2 3028: 3024: 3023: 3019: 3015: 3011: 3005: 3002: 3000: 2996: 2993: 2992: 2991: 2987: 2986:Haplogroup R1 2984: 2983: 2982: 2980: 2976: 2972: 2968: 2964: 2960: 2958: 2954: 2950: 2946: 2942: 2937: 2935: 2931: 2927: 2923: 2919: 2910: 2905: 2889: 2888: 2884: 2878: 2874: 2873: 2867: 2864: 2861: 2854: 2852:Q1a3a2 (M194) 2851: 2849: 2848: 2844: 2840: 2839:South America 2835:Q1a3a1 (M19) 2834: 2831: 2830: 2829: 2828: 2822: 2819: 2817: 2816: 2812: 2808: 2802: 2801: 2799: 2797: 2796: 2792: 2788: 2782: 2780: 2779: 2775: 2771: 2767: 2763: 2759: 2755: 2751: 2745: 2742: 2741: 2739: 2736: 2735: 2733: 2730: 2729: 2727: 2726: 2725: 2722: 2719: 2715: 2711: 2706: 2696: 2695: 2694: 2690: 2686: 2682: 2676: 2675: 2671: 2661: 2658: 2656:(Q-M242) and; 2655: 2652: 2651: 2650: 2647: 2645: 2641: 2637: 2633: 2629: 2625: 2621: 2616: 2614: 2610: 2606: 2601: 2585: 2576: 2572: 2569:This section 2567: 2564: 2560: 2559: 2553: 2549: 2546: 2543: 2539: 2536: 2535: 2534: 2531: 2529: 2525: 2521: 2517: 2513: 2509: 2503: 2491: 2490: 2486: 2480: 2479:Haplogroup O2 2477: 2471: 2469: 2465: 2459: 2456: 2454: 2453: 2449: 2445: 2441: 2440:Austroasiatic 2437: 2433: 2427: 2424: 2423: 2421: 2418: 2416: 2414: 2410: 2406: 2402: 2398: 2392: 2389: 2388: 2386: 2385: 2384: 2382: 2378: 2374: 2370: 2366: 2365:South Pacific 2362: 2358: 2354: 2349: 2339: 2337: 2333: 2328: 2326: 2322: 2317: 2301: 2292: 2288: 2285:This section 2283: 2280: 2276: 2275: 2271: 2255: 2246: 2242: 2239:This section 2237: 2234: 2230: 2229: 2226: 2224: 2220: 2216: 2210: 2209:Haplogroup K2 2200: 2198: 2194: 2193:Mediterranean 2190: 2186: 2182: 2181:Sephardi Jews 2178: 2174: 2170: 2166: 2162: 2161:Mediterranean 2158: 2154: 2150: 2146: 2142: 2138: 2136: 2131: 2130:Haplogroup LT 2121: 2119: 2115: 2111: 2107: 2103: 2098: 2096: 2092: 2088: 2084: 2079: 2064: 2060: 2056: 2052: 2048: 2044: 2040: 2036: 2032: 2028: 2027:Haplogroup J2 2025: 2022: 2018: 2014: 2010: 2006: 2002: 2001:Haplogroup J1 1999: 1998: 1996: 1992: 1991: 1990: 1988: 1984: 1980: 1976: 1971: 1958: 1954: 1953:Haplogroup I2 1951: 1949: 1945: 1944:Haplogroup I1 1942: 1941: 1940: 1938: 1934: 1930: 1925: 1909: 1900: 1896: 1893:This section 1891: 1888: 1884: 1883: 1880: 1878: 1874: 1869: 1867: 1863: 1859: 1853: 1835: 1832: 1831: 1829: 1826: 1824: 1821: 1815: 1812: 1811: 1810: 1807: 1805: 1802: 1801: 1800: 1797: 1795: 1792: 1790: 1787: 1786: 1785: 1782: 1781: 1780: 1779:Haplogroup G2 1777: 1775: 1774:Haplogroup G1 1772: 1771: 1770: 1768: 1764: 1760: 1756: 1752: 1748: 1744: 1739: 1737: 1733: 1732:Mediterranean 1729: 1725: 1721: 1717: 1713: 1709: 1705: 1701: 1697: 1693: 1689: 1685: 1680: 1678: 1674: 1669: 1659: 1657: 1653: 1649: 1645: 1641: 1637: 1633: 1632:Tamang people 1629: 1625: 1621: 1617: 1613: 1609: 1605: 1601: 1597: 1593: 1589: 1586:, especially 1585: 1580: 1576: 1561: 1560:Haplogroup E2 1558: 1548: 1544: 1542: 1538: 1537:Mediterranean 1534: 1530: 1526: 1520: 1517: 1516: 1514: 1511: 1507: 1503: 1501: 1497: 1491: 1488: 1486: 1482: 1480: 1474: 1471: 1470: 1468: 1465: 1464: 1463: 1459: 1456: 1455: 1453: 1450: 1448: 1444: 1441: 1440: 1438: 1437:Haplogroup E1 1435: 1434: 1433: 1431: 1425: 1422: 1421: 1419: 1405: 1404: 1400: 1396: 1390: 1386: 1385:, Philippines 1384: 1383:Mactan Island 1378: 1374: 1372: 1368: 1364: 1358: 1355: 1353: 1352: 1346: 1343: 1339: 1338: 1334: 1330: 1326: 1320: 1318: 1316: 1312: 1308: 1304: 1298: 1297: 1295: 1292: 1291: 1289: 1288: 1287: 1285: 1280: 1276: 1272: 1268: 1262: 1261:Haplogroup D1 1259: 1258: 1256: 1253: 1252: 1250: 1236: 1234: 1230: 1226: 1222: 1218: 1214: 1213:North America 1210: 1204: 1203:Haplogroup C2 1201: 1193: 1192: 1188: 1182: 1180: 1179: 1175: 1171: 1167: 1163: 1157: 1156: 1154: 1151: 1148: 1144: 1143: 1137: 1133: 1129: 1125: 1121: 1119: 1117: 1116:Lebbo' people 1111: 1110: 1108: 1106: 1105: 1101: 1097: 1091: 1088: 1087: 1085: 1084: 1082: 1081: 1079: 1075: 1074: 1070: 1066: 1062: 1056: 1053: 1051: 1050: 1044: 1041: 1040: 1038: 1037: 1035: 1034:Haplogroup C1 1032: 1031: 1030: 1029: 1028:North America 1025: 1021: 1016:(M130, M216) 1015: 1012: 1011: 1009: 999: 994: 974: 973:Haplogroup CT 971: 968: 965: 964: 962: 961:Haplogroup BT 959: 956: 953: 952: 950: 947: 946: 945:(also A1a-T) 944: 943:Haplogroup A1 941: 938: 937:Haplogroup A0 935: 933: 930: 929: 928: 925: 924: 923: 921: 917: 913: 893: 892: 884: 883: 875: 874: 866: 865: 857: 856: 848: 847: 839: 838: 830: 829: 821: 820: 812: 811: 803: 802: 794: 793: 790: 789: 783: 782: 779: 778: 775: 774: 768: 767: 761: 758: 757: 754: 753: 745: 744: 741: 740: 734: 733: 730: 729: 726: 725: 719: 718: 712: 709: 708: 702: 699: 698: 695: 694: 686: 685: 682: 681: 675: 674: 671: 670: 667: 666: 660: 659: 653: 650: 649: 643: 640: 639: 636: 635: 627: 626: 623: 622: 616: 615: 612: 611: 608: 607: 601: 600: 594: 591: 590: 584: 581: 580: 577: 576: 568: 567: 564: 563: 557: 556: 553: 552: 549: 548: 542: 541: 535: 532: 531: 525: 522: 521: 518: 517: 514: 513: 507: 506: 500: 497: 496: 493: 492: 489: 488: 482: 481: 475: 472: 471: 468: 467: 464: 463: 457: 456: 450: 447: 446: 443: 442: 434: 433: 430: 429: 423: 422: 419: 418: 415: 414: 408: 407: 401: 398: 397: 391: 388: 387: 384: 383: 380: 379: 373: 372: 366: 363: 362: 359: 358: 355: 354: 348: 347: 341: 340: 337: 336: 333: 332: 326: 325: 318: 315: 314: 309: 300: 296: 294: 290: 286: 282: 278: 275: 266: 257: 255: 254:Haplogroup CT 251: 247: 243: 242:patrilineally 239: 235: 230: 228: 224: 220: 216: 212: 208: 204: 200: 196: 192: 180: 175: 173: 168: 166: 161: 160: 158: 157: 152: 149: 147: 144: 142: 139: 137: 134: 132: 129: 126: 122: 121: 120: 119: 115: 114: 109: 106: 102: 97: 92: 88: 86: 83: 81: 77: 73: 70: 68: 65: 64: 63: 62: 58: 57: 54: 51: 50: 46: 42: 41: 35: 30: 23: 19: 7587: 7523:North Africa 7404:North Africa 7251: 7247: 7243: 7212: 7166:. Retrieved 7162:the original 7146:. Retrieved 7139:the original 7108: 7102: 7080: 7073: 7063: 7025: 7021: 6962: 6958: 6948: 6933: 6896: 6892: 6866:. Retrieved 6862:the original 6857: 6802: 6798: 6787: 6778: 6767: 6743:(1): 47–55. 6740: 6736: 6699:. Retrieved 6692:the original 6661: 6657: 6598: 6594: 6583: 6538: 6534: 6486: 6482: 6423: 6419: 6369: 6365: 6355: 6318: 6314: 6304: 6269: 6265: 6255: 6241: 6206: 6202: 6192: 6149: 6145: 6134: 6099: 6095: 6085: 6040: 6036: 6026: 5985: 5981: 5975: 5930: 5926: 5915: 5880: 5876: 5865: 5820: 5816: 5806: 5794:. Retrieved 5784: 5757: 5753: 5743: 5734: 5728:. Retrieved 5721:the original 5690: 5686: 5668: 5633: 5629: 5619: 5584: 5580: 5558: 5522: 5518: 5508: 5481: 5477: 5467: 5440: 5434: 5424: 5411: 5376: 5372: 5362: 5317: 5311: 5263: 5259: 5211: 5207: 5159: 5155: 5117: 5113: 5107: 5084: 5060:. Retrieved 5056: 5046: 5034:. Retrieved 5024: 5016: 5009:. Retrieved 4999: 4991: 4984:. Retrieved 4974: 4965: 4956: 4946:15 September 4944:. Retrieved 4815: 4809: 4799: 4779: 4772: 4729: 4725: 4715: 4704:18 September 4702:. Retrieved 4688: 4678: 4666:. Retrieved 4662:the original 4635:cite journal 4598: 4594: 4584: 4558: 4549: 4539: 4532: 4527: 4521: 4512: 4503: 4494: 4477: 4468: 4459: 4450: 4441: 4432: 4426: 4393: 4389: 4383: 4374: 4239:   4182:   3882: 3874: 3850: 3692:(E1b1a-U174) 3626:North Africa 3033: 3018:North Africa 3014:Central Asia 3007: 2998: 2989: 2975:Italo-Celtic 2961: 2949:Central Asia 2945:Balto-Slavic 2938: 2918:Haplogroup R 2915: 2880: 2869: 2868:Q1a6 (M323) 2836: 2824: 2804: 2800:Q1a3 (M346) 2791:Central Asia 2784: 2747: 2723: 2714:Central Asia 2708: 2689:Central Asia 2678: 2667: 2665: 2660:Haplogroup R 2654:Haplogroup Q 2648: 2636:Central Asia 2617: 2605:Haplogroup P 2603: 2579: 2575:adding to it 2570: 2548:Haplogroup M 2538:Haplogroup S 2532: 2505: 2482: 2461: 2429: 2422:(P31, M268) 2409:Austronesian 2394: 2380: 2377:Indian Ocean 2369:Central Asia 2353:Haplogroup O 2351: 2342:Haplogroup O 2329: 2321:Haplogroup N 2319: 2310:Haplogroup N 2295: 2291:adding to it 2286: 2249: 2245:adding to it 2240: 2212: 2141:Haplogroup T 2139: 2135:Haplogroup L 2133: 2099: 2083:Haplogroup K 2081: 2063:Central Asia 2021:North Africa 1975:Haplogroup J 1973: 1956: 1947: 1929:Haplogroup I 1927: 1903: 1899:adding to it 1894: 1873:Medieval era 1870: 1855: 1767:North Africa 1740: 1681: 1673:Haplogroup G 1671: 1622:and 0.2% in 1581: 1578: 1547:formerly E3b 1546: 1529:North Africa 1522: 1506:formerly E3* 1505: 1493: 1485:formerly E3a 1484: 1476: 1461: 1446: 1445:(M33, M132) 1427: 1424:Haplogroup E 1399:Saudi Arabia 1392: 1380: 1360: 1348: 1322: 1300: 1273:(especially 1264: 1255:Haplogroup D 1206: 1205:(M217, P44) 1184: 1159: 1140:Found among 1139: 1131: 1123: 1114:Found among 1113: 1093: 1058: 1046: 1017: 1014:Haplogroup C 996: 967:Haplogroup B 927:Haplogroup A 909: 788:Haplogroup R 786: 773:Haplogroup Q 771: 739:Haplogroup M 737: 724:Haplogroup S 722: 680:Haplogroup O 678: 665:Haplogroup N 663: 621:Haplogroup T 619: 606:Haplogroup L 604: 562:Haplogroup J 560: 547:Haplogroup I 545: 512:Haplogroup H 510: 487:Haplogroup G 485: 462:Haplogroup C 460: 428:Haplogroup E 426: 413:Haplogroup D 411: 378:Haplogroup B 376: 353:Haplogroup A 351: 329: 317:Phylogenetic 297: 293:nomenclature 285:terminal SNP 284: 271: 231: 215:Y chromosome 209:portions of 194: 188: 100: 22: 7562:Middle East 7207:, from the 3699:East Africa 3680:East Africa 3645:East Africa 3387:Middle East 3368:Middle East 3331:East Africa 3295:East Africa 3276:East Africa 3239:Middle East 3185:Middle East 3167:Middle East 3127:East Africa 3047:Haplogroup 2879:Q1b (M378) 2872:Yemeni Jews 2825:Typical of 2758:Han Chinese 2740:Q1a (MEH2) 2734:Q1 (P36.2) 2624:Philippines 2528:Polynesians 2485:Austronesia 2452:Indonesians 2157:Middle East 2007:peoples in 1979:Middle East 1877:Roma people 1640:South China 1628:South India 1533:Middle East 1479:Niger–Congo 1462:formerly E3 1447:formerly E1 1430:Middle East 1426:(M40, M96) 221:(STRs) and 207:recombining 205:in the non- 7613:Categories 7567:South Asia 7445:South Asia 7081:Genome.org 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Index

Human mitochondrial DNA haplogroup

Phylogenetic tree
a series
Genetic genealogy
Population genetics
Haplogroup
Haplotype
Subclade
Most recent common ancestor
DNA haplogroups
Human mitochondrial
Human Y-chromosome
Genomics
Y-chromosome haplogroups by population
Genealogical DNA test
Surname DNA project
Personal genomics
Genographic Project
ISOGG
v
t
e
human genetics
haplogroup
mutations
recombining
DNA
Y chromosome
short tandem repeats

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