2611:
533:
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136:
409:
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118:
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561:
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498:, and, as regards osteoderms, two sets of co-ossified cervical plates and a number of elongate conical spines. A very unusual feature is the sagittal (midline) osteoderm on the first set of cervical plates; in most other ankylosaurs, these osteoderms are bilateral, i.e. paired with one on each side of the midline.
598:
to all other ankylosaurids (i.e., members of the
Ankylosauridae). These studies however, only utilized the skull, whereas many of the distinctive features of the family Polacanthidae are in the postcranial skeleton. In contrast, a phylogenetic analysis by Soto-Acuña and colleagues in 2021 recovered
1462:
Soto-Acuña, Sergio; Vargas, Alexander O.; Kaluza, Jonatan; Leppe, Marcelo A.; Botelho, Joao F.; Palma-Liberona, José; Simon-Gutstein, Carolina; Fernández, Roy A.; Ortiz, Héctor; Milla, Verónica; Aravena, Bárbara (2021).
436:. Besides the holotype, two other partial skeletons are known (although not yet described). The holotype consists of most of the skull and a partial postcranial skeleton. The specimen was originally described as
1372:""Dragons" on the landscape: Modeling the abundance of large carnivorous dinosaurs of the Upper Jurassic Morrison Formation (USA) and the Upper Cretaceous Dinosaur Park Formation (Canada)"
1533:
421:
1353:
Kilbourne, B. and
Carpenter, K. 2005. "Redescription of Gargoyleosaurus parkpinorum, a polacanthid ankylosaur from the Upper Jurassic of Albany County, Wyoming".
556:(DMNH 27726) Skull and cervical armor in side and front view. Insert shows a cheek tooth. The skull is slightly crushed changing the shape of the orbit.
463:
was a relatively small ankylosaur, reaching 3–3.5 m (9.8–11.5 ft) in length and 300–754 kg (661–1,662 lb) in body mass. Much of the
2712:
449:
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2699:
1528:
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2610:
1438:
1322:
486:
bosses. The taxon is further characterized by a narrow rostrum (in dorsal view), the presence of seven conical teeth in each
1586:
425:
387:
1370:
Farlow, James O.; Coroian, Dan; Currie, Philip J.; Foster, John R.; Mallon, Jordan C.; Therrien, François (2022-07-11).
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135:
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1287:(2nd edition), Weishampel, D. B., Dodson, P., and OsmĂłlska, H., editors. University of California Press.
1558:
1656:
1265:
Carpenter, K., Miles, C. and
Cloward, K. (1998). "Skull of a Jurassic ankylosaur (Dinosauria)."
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357:
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130:
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1800:
1512:
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1492:
1464:
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1401:
1393:
1318:
376:
2001:
1948:
1786:
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1383:
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564:
495:
2466:
2430:
2370:
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1848:
1808:
1641:
1537:
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837:
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333:
was discovered in 1995 at the Bone Cabin Quarry West locality, in Albany County,
2529:
2515:
2501:
2409:
2382:
2278:
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2174:
2120:
1985:
1955:
1925:
1888:
1864:
1488:
1294:, a polacanthid ankylosaur from the Upper Jurassic of Albany County, Wyoming".
1145:
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803:
762:
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1397:
939:
888:
714:
631:
479:
68:
30:
1465:"Bizarre tail weaponry in a transitional ankylosaur from subantarctic Chile"
2522:
2508:
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2190:
2141:
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2098:
2085:
2058:
1977:
1962:
1918:
1898:
1765:
1747:
1725:
1695:
1668:
1616:
1574:
1504:
1405:
989:
905:
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583:
364:
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241:
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196:
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2156:
2134:
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2015:
1815:
1793:
1625:
1129:
1055:
956:
922:
654:
617:
494:
nasal septum, a linearly arranged nasal cavity, the absence of an osseus
392:
326:
212:
43:
549:
532:
408:
2704:
2550:
2543:
2481:
2356:
2307:
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2197:
2183:
2149:
2127:
1993:
1837:
1772:
1105:
1024:
786:
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53:
1339:
Jurassic West: The
Dinosaurs of the Morrison Formation and Their World
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1306:
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93:
37:
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48:
1303:
117:
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2451:
491:
334:
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73:
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429:
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was present in stratigraphic zone 2 of the
Morrison Formation.
157:
2416:
591:
472:
464:
606:
Below is a reproduced phylogenetic analysis from Soto-Acuña
2578:
1461:
1290:
Killbourne, B. and
Carpenter, K. (2005). "Redescription of
560:
1276:. pp. 454–483. Indiana University Press, Bloomington.
1369:
511:
1317:. Edinburgh, Scotland: Dorling Kindersley. p. 278.
424:
in 1995 and is currently held in the collections of the
440:
by
Carpenter, Miles and Cloward in 1998, then renamed
1349:
1347:
125:
Fossil skeleton, Denver Museum of Nature and
Science
1235:
475:displays cranial sculpturing, including pronounced
329:known from reasonably complete fossil remains. The
1344:
2758:
396:, has been on display there since around 2002.
1640:
1331:
382:in 1998. A mounted skeletal reconstruction of
1559:
1296:Neues Jahrbuch für Geologie und Paläontologie
1283:, and Weishampel (2004). "Ankylosauria". in
1745:
1433:. Princeton University Press. p. 252.
1566:
1552:
390:and, alongside a couple skeletons of baby
116:
1387:
1341:. Indiana University Press. pp. 327–329.
586:and are in agreement with most previous
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548:
407:
2759:
1430:The Princeton Field Guide to Dinosaurs
1312:
2653:
2652:
1585:
1547:
1426:
2772:Dinosaurs of the Morrison Formation
426:Denver Museum of Nature and Science
388:Denver Museum of Nature and Science
13:
14:
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1522:
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2633:
2632:
2609:
2604:
1573:
1238:
531:
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325:lizard") is one of the earliest
134:
41:
2782:Taxa named by Kenneth Carpenter
1540:from the Natural History Museum
1337:Foster, J. (2007). "Appendix."
1254:Timeline of ankylosaur research
1455:
1420:
1363:
455:
1:
2777:Fossil taxa described in 1998
1259:
590:hypotheses, which place the
471:has been recovered, and the
448:in 2001, in accordance with
403:
7:
1313:Benton, Michael J. (2012).
1292:Gargoyleosaurus parkpinorum
1231:
576:Gargoyleosaurus parkpinorum
554:Gargoyleosaurus parkpinorum
540:Gargoyleosaurus parkpinorum
418:Gargoyleosaurus parkpinorum
384:Gargoyleosaurus parkpinorum
299:Gargoyleosaurus parkpinorum
10:
2808:
1489:10.1038/s41586-021-04147-1
1355:N. Jb. Geol. Palaont. Abh.
16:Extinct species of reptile
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1427:Paul, Gregory S. (2016).
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422:Western Paleontology Labs
416:The holotype specimen of
292:
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131:Scientific classification
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115:
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438:Gargoyleosaurus parkpini
2787:Paleontology in Wyoming
567:
557:
522:Museum of Ancient Life
413:
2739:Paleobiology Database
1376:The Anatomical Record
1274:The Armored Dinosaurs
563:
552:
411:
2792:Ornithischian genera
1272:Carpenter, K. (ed.)
524:, Thanksgiving Point
1481:2021Natur.600..259S
386:can be seen at the
375:) was described by
278:G. parkpinorum
1536:2008-09-10 at the
578:within the family
568:
558:
414:
358:Morrison Formation
2754:
2753:
2726:Open Tree of Life
2655:Taxon identifiers
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2645:
2618:
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2588:
2587:
2537:Nodocephalosaurus
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1882:
1881:
1801:Serendipaceratops
1737:
1736:
1733:
1732:
1718:
1717:
1714:
1713:
1475:(7888): 259–263.
1440:978-1-78684-190-2
1324:978-0-7566-9910-9
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1210:
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1165:
1164:
1156:
1155:
1084:
1083:
1000:
999:
752:
751:
743:
742:
603:as a nodosaurid.
420:was collected by
314:
313:
266:
2799:
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2746:
2734:
2733:
2721:
2720:
2708:
2707:
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2650:
2649:
2636:
2635:
2613:
2608:
2388:
2387:
2284:
2283:
2091:
2090:
2002:Propanoplosaurus
1949:Glyptodontopelta
1904:
1903:
1895:
1894:
1787:Priodontognathus
1758:
1757:
1754:
1753:
1743:
1742:
1674:
1673:
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1637:
1631:
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1517:
1516:
1459:
1453:
1452:
1424:
1418:
1417:
1391:
1389:10.1002/ar.25024
1382:(7): 1669–1696.
1367:
1361:
1351:
1342:
1335:
1329:
1328:
1315:Prehistoric Life
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1248:
1246:Dinosaurs portal
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672:
648:
647:
624:
623:
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565:Life restoration
535:
514:
496:secondary palate
490:, an incomplete
301:
297:
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139:
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40:
29:Temporal range:
21:
20:
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2806:
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2801:
2800:
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2797:
2796:
2757:
2756:
2755:
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2737:
2729:
2724:
2716:
2711:
2703:
2698:
2690:
2685:
2676:
2675:
2670:
2663:Gargoyleosaurus
2657:
2647:
2642:
2614:
2584:
2487:
2467:Tianzhenosaurus
2431:Minotaurasaurus
2377:
2371:Zhongyuansaurus
2342:
2337:Zhejiangosaurus
2330:Liaoningosaurus
2265:
2223:Struthiosaurini
2217:
2169:
2164:Tatankacephalus
2080:
2036:Gargoyleosaurus
2021:
1933:Gargoyleosaurus
1891:
1878:
1849:Parankylosauria
1843:
1809:Sinankylosaurus
1750:
1729:
1710:
1690:
1663:
1657:Avemetatarsalia
1642:Avemetatarsalia
1632:
1588:
1587:
1577:
1572:
1538:Wayback Machine
1530:Gargoyleosaurus
1525:
1520:
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1425:
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1332:
1325:
1311:
1304:
1298:, 237, 111–160.
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1234:
1229:
1220:
1211:
1202:
1193:
1184:
1175:
1166:
1157:
1085:
1073:Liaoningosaurus
1001:
821:Gargoyleosaurus
753:
744:
677:Parankylosauria
601:Gargoyleosaurus
547:
546:
545:
544:
543:
536:
527:
526:
525:
520:skeleton cast,
518:Gargoyleosaurus
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461:Gargoyleosaurus
458:
412:Size comparison
406:
398:Gargoyleosaurus
318:Gargoyleosaurus
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86:
81:
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66:
61:
56:
51:
46:
35:
34:
27:
25:Gargoyleosaurus
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12:
11:
5:
2805:
2795:
2794:
2789:
2784:
2779:
2774:
2769:
2752:
2751:
2749:
2748:
2735:
2722:
2709:
2696:
2683:
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2658:
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2641:
2640:
2630:
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2601:
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2589:
2586:
2585:
2583:
2582:
2575:
2568:
2561:
2554:
2547:
2540:
2533:
2530:Euoplocephalus
2526:
2519:
2516:Anodontosaurus
2512:
2505:
2502:Akainacephalus
2497:
2495:
2489:
2488:
2486:
2485:
2478:
2471:
2463:
2456:
2448:
2441:
2434:
2427:
2420:
2413:
2410:Crichtonsaurus
2406:
2399:
2391:
2385:
2383:Ankylosaurinae
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2378:
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2375:
2367:
2360:
2352:
2350:
2344:
2343:
2341:
2340:
2333:
2326:
2318:
2311:
2304:
2296:
2287:
2281:
2279:Ankylosauridae
2275:
2274:
2271:
2270:
2267:
2266:
2264:
2263:
2260:Struthiosaurus
2256:
2249:
2242:
2235:
2227:
2225:
2219:
2218:
2216:
2215:
2208:
2201:
2194:
2187:
2179:
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2175:Panoplosaurini
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2170:
2168:
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2160:
2153:
2146:
2138:
2131:
2124:
2121:Niobrarasaurus
2117:
2110:
2102:
2094:
2088:
2082:
2081:
2079:
2078:
2070:
2063:
2055:
2048:
2040:
2031:
2029:
2023:
2022:
2020:
2019:
2012:
2005:
1998:
1990:
1986:Polacanthoides
1982:
1974:
1967:
1959:
1956:Horshamosaurus
1952:
1945:
1937:
1929:
1926:Dongyangopelta
1922:
1915:
1907:
1901:
1892:
1889:Euankylosauria
1887:
1884:
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1879:
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1865:Kunbarrasaurus
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1523:External links
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502:Classification
500:
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452:art. 31.1.2A.
442:G. parkpinorum
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369:G. parkpinorum
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15:
9:
6:
4:
3:
2:
2804:
2793:
2790:
2788:
2785:
2783:
2780:
2778:
2775:
2773:
2770:
2768:
2765:
2764:
2762:
2745:
2740:
2736:
2732:
2727:
2723:
2719:
2714:
2710:
2706:
2701:
2697:
2693:
2688:
2684:
2679:
2673:
2669:
2668:
2666:
2664:
2660:
2656:
2651:
2639:
2631:
2629:
2625:
2624:
2621:
2612:
2607:
2581:
2580:
2576:
2574:
2573:
2569:
2567:
2566:
2562:
2560:
2559:
2555:
2553:
2552:
2548:
2546:
2545:
2541:
2539:
2538:
2534:
2532:
2531:
2527:
2525:
2524:
2520:
2518:
2517:
2513:
2511:
2510:
2506:
2504:
2503:
2499:
2498:
2496:
2494:
2493:Ankylosaurini
2490:
2484:
2483:
2479:
2477:
2476:
2472:
2469:
2468:
2464:
2462:
2461:
2457:
2454:
2453:
2449:
2447:
2446:
2442:
2440:
2439:
2435:
2433:
2432:
2428:
2426:
2425:
2421:
2419:
2418:
2414:
2412:
2411:
2407:
2405:
2404:
2403:Crichtonpelta
2400:
2398:
2397:
2396:Bissektipelta
2393:
2392:
2389:
2386:
2384:
2380:
2373:
2372:
2368:
2366:
2365:
2361:
2359:
2358:
2354:
2353:
2351:
2349:
2348:Shamosaurinae
2345:
2339:
2338:
2334:
2332:
2331:
2327:
2324:
2323:
2319:
2317:
2316:
2312:
2310:
2309:
2305:
2302:
2301:
2297:
2294:
2293:
2292:Ahshislepelta
2289:
2288:
2285:
2282:
2280:
2276:
2262:
2261:
2257:
2255:
2254:
2250:
2248:
2247:
2243:
2241:
2240:
2239:Hungarosaurus
2236:
2234:
2233:
2229:
2228:
2226:
2224:
2220:
2214:
2213:
2209:
2207:
2206:
2205:Panoplosaurus
2202:
2200:
2199:
2195:
2193:
2192:
2188:
2186:
2185:
2181:
2180:
2178:
2176:
2172:
2166:
2165:
2161:
2159:
2158:
2154:
2152:
2151:
2147:
2144:
2143:
2139:
2137:
2136:
2132:
2130:
2129:
2125:
2123:
2122:
2118:
2116:
2115:
2111:
2108:
2107:
2106:Ahshislepelta
2103:
2101:
2100:
2096:
2095:
2092:
2089:
2087:
2083:
2076:
2075:
2071:
2069:
2068:
2064:
2061:
2060:
2056:
2054:
2053:
2052:Hoplitosaurus
2049:
2046:
2045:
2041:
2038:
2037:
2033:
2032:
2030:
2028:
2027:Polacanthinae
2024:
2018:
2017:
2013:
2011:
2010:
2009:Rhadinosaurus
2006:
2004:
2003:
1999:
1996:
1995:
1991:
1988:
1987:
1983:
1980:
1979:
1975:
1973:
1972:
1968:
1965:
1964:
1960:
1958:
1957:
1953:
1951:
1950:
1946:
1943:
1942:
1938:
1935:
1934:
1930:
1928:
1927:
1923:
1921:
1920:
1916:
1914:
1913:
1912:Acanthopholis
1909:
1908:
1905:
1902:
1900:
1896:
1893:
1890:
1885:
1875:
1874:
1870:
1867:
1866:
1862:
1860:
1859:
1858:Antarctopelta
1855:
1854:
1852:
1850:
1846:
1840:
1839:
1835:
1833:
1832:
1831:Tianchisaurus
1828:
1825:
1824:
1823:Stegosaurides
1820:
1818:
1817:
1813:
1811:
1810:
1806:
1803:
1802:
1798:
1796:
1795:
1791:
1789:
1788:
1784:
1782:
1781:
1777:
1775:
1774:
1770:
1768:
1767:
1763:
1762:
1759:
1755:
1752:
1749:
1744:
1740:
1728:
1727:
1721:
1704:
1703:
1702:
1701:
1699:
1697:
1693:
1685:
1681:
1680:
1679:
1678:
1675:
1672:
1670:
1666:
1658:
1654:
1653:
1652:
1651:
1648:
1645:
1643:
1639:
1635:
1627:
1623:
1620:
1618:
1614:
1611:
1609:
1605:
1602:
1600:
1596:
1594:
1590:
1589:
1584:
1580:
1576:
1569:
1564:
1562:
1557:
1555:
1550:
1549:
1546:
1539:
1535:
1532:
1531:
1527:
1526:
1514:
1510:
1506:
1502:
1498:
1494:
1490:
1486:
1482:
1478:
1474:
1470:
1466:
1458:
1450:
1446:
1442:
1436:
1432:
1431:
1423:
1415:
1411:
1407:
1403:
1399:
1395:
1390:
1385:
1381:
1377:
1373:
1366:
1359:
1356:
1350:
1348:
1340:
1334:
1326:
1320:
1316:
1309:
1307:
1302:
1297:
1293:
1289:
1286:
1282:
1278:
1275:
1271:
1269:393: 782–783.
1268:
1264:
1263:
1255:
1252:
1251:
1247:
1236:
1224:
1223:
1215:
1214:
1206:
1205:
1197:
1196:
1188:
1187:
1179:
1178:
1170:
1169:
1161:
1160:
1152:
1151:
1148:
1147:
1141:
1140:
1137:
1136:
1133:
1132:
1131:
1124:
1123:
1117:
1116:
1113:
1112:
1109:
1108:
1107:
1100:
1099:
1093:
1092:
1089:
1088:
1080:
1079:
1076:
1075:
1074:
1067:
1066:
1063:
1062:
1059:
1058:
1057:
1050:
1049:
1043:
1042:
1036:
1035:
1032:
1031:
1028:
1027:
1026:
1019:
1018:
1012:
1009:
1008:
1005:
1004:
996:
995:
992:
991:
985:
984:
981:
980:
977:
976:
975:
968:
967:
964:
963:
960:
959:
958:
951:
950:
947:
946:
943:
942:
941:
940:Hoplitosaurus
934:
933:
930:
929:
926:
925:
924:
917:
916:
913:
912:
909:
908:
907:
900:
899:
896:
895:
892:
891:
890:
889:Panoplosaurus
883:
882:
879:
878:
875:
874:
873:
866:
865:
862:
861:
858:
857:
856:
849:
848:
845:
844:
841:
840:
839:
832:
831:
828:
827:
824:
823:
822:
815:
814:
811:
810:
807:
806:
805:
798:
797:
794:
793:
790:
789:
788:
781:
780:
774:
771:
770:
764:
761:
760:
757:
756:
748:
747:
739:
738:
735:
734:
733:
726:
725:
722:
721:
718:
717:
716:
715:Antarctopelta
709:
708:
702:
701:
698:
697:
694:
693:
692:
685:
684:
678:
675:
674:
668:
665:
664:
661:
660:
657:
656:
650:
649:
643:
642:
639:
638:
635:
634:
633:
632:Scelidosaurus
626:
625:
619:
616:
615:
611:
609:
604:
602:
597:
593:
589:
585:
581:
577:
574:(2004) place
573:
566:
562:
555:
551:
541:
534:
523:
519:
513:
499:
497:
493:
489:
485:
481:
480:quadratojugal
478:
474:
470:
466:
462:
453:
451:
447:
444:by Carpenter
443:
439:
435:
431:
427:
423:
419:
410:
401:
399:
395:
394:
389:
385:
381:
378:
377:Ken Carpenter
374:
370:
366:
361:
359:
355:
352:
348:
344:
341:of the Upper
340:
336:
332:
328:
324:
320:
319:
308:
302:
300:
291:
288:
287:Binomial name
284:
280:
279:
273:
270:
269:
264:
258:
257:
250:
247:
246:
243:
237:
234:
233:
230:
224:
221:
218:
217:
214:
208:
205:
202:
201:
198:
192:
189:
186:
185:
182:
179:
176:
173:
172:
169:
166:
163:
162:
159:
156:
153:
152:
149:
146:
143:
142:
137:
132:
128:
123:
119:
114:
100:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
39:
36:154–150
32:
31:Late Jurassic
26:
22:
19:
2662:
2577:
2570:
2563:
2556:
2549:
2542:
2535:
2528:
2523:Dyoplosaurus
2521:
2514:
2509:Ankylosaurus
2507:
2500:
2480:
2473:
2465:
2458:
2450:
2443:
2438:Pinacosaurus
2436:
2429:
2422:
2415:
2408:
2401:
2394:
2369:
2362:
2355:
2335:
2328:
2322:Hylaeosaurus
2320:
2313:
2306:
2298:
2290:
2258:
2251:
2246:Pawpawsaurus
2244:
2237:
2230:
2210:
2203:
2196:
2191:Denversaurus
2189:
2182:
2162:
2155:
2148:
2142:Peloroplites
2140:
2133:
2126:
2119:
2114:Borealopelta
2112:
2104:
2099:Acantholipan
2097:
2086:Nodosaurinae
2072:
2065:
2059:Peloroplites
2057:
2050:
2042:
2035:
2034:
2014:
2007:
2000:
1992:
1984:
1978:Mymoorapelta
1976:
1969:
1963:Hylaeosaurus
1961:
1954:
1947:
1939:
1932:
1931:
1924:
1919:Anoplosaurus
1917:
1910:
1899:Nodosauridae
1871:
1863:
1856:
1836:
1829:
1821:
1814:
1807:
1799:
1792:
1785:
1778:
1771:
1766:Cryptosaurus
1764:
1748:Ankylosauria
1726:Ankylosauria
1724:
1696:Ankylosauria
1684:Ornithischia
1669:Ornithischia
1621:
1617:Ornithischia
1612:
1603:
1575:Ankylosauria
1529:
1472:
1468:
1457:
1429:
1422:
1379:
1375:
1365:
1357:
1354:
1338:
1333:
1314:
1295:
1291:
1284:
1279:Vickaryous,
1273:
1266:
1144:
1128:
1127:
1104:
1103:
1071:
1070:
1054:
1053:
1023:
1022:
990:Nodosaurinae
988:
972:
971:
955:
954:
938:
937:
921:
920:
906:Peloroplites
904:
903:
887:
886:
872:Mymoorapelta
870:
869:
855:Hylaeosaurus
853:
852:
836:
835:
820:
819:
818:
802:
801:
785:
784:
773:Nodosauridae
730:
729:
713:
712:
689:
688:
667:Ankylosauria
653:
630:
629:
607:
605:
600:
596:sister group
588:phylogenetic
584:Ankylosauria
575:
571:
569:
553:
539:
538:Skeleton of
517:
460:
459:
445:
441:
437:
417:
415:
397:
391:
383:
379:
372:
371:(originally
368:
365:type species
362:
347:Kimmeridgian
317:
316:
315:
306:
298:
293:
277:
276:
262:
255:
254:
242:Nodosauridae
229:Ankylosauria
219:
203:
197:Ornithischia
187:
174:
24:
18:
2767:Nodosaurids
2558:Scolosaurus
2475:Tsagantegia
2424:Jinyunpelta
2364:Shamosaurus
2315:Chuanqilong
2157:Silvisaurus
2135:Patagopelta
2067:Polacanthus
2016:Sauroplites
1816:Spicomellus
1794:Sarcolestes
1626:Thyreophora
1130:Shamosaurus
1056:Chuanqilong
957:Sauroplites
923:Polacanthus
655:Stegosauria
618:Thyreophora
570:Vickaryous
542:in top view
456:Description
393:Stegosaurus
373:G. parkpini
327:ankylosaurs
213:Thyreophora
2761:Categories
2626:See also:
2551:Platypelta
2544:Oohkotokia
2482:Zaraapelta
2357:Gobisaurus
2308:Cedarpelta
2300:Aletopelta
2253:Stegopelta
2198:Edmontonia
2184:Animantarx
2150:Sauropelta
2128:Nodosaurus
1994:Priconodon
1838:Vectipelta
1773:Dracopelta
1705:see below↓
1608:Dinosauria
1260:References
1106:Gobisaurus
1025:Aletopelta
787:Cedarpelta
488:premaxilla
321:(meaning "
305:Carpenter
261:Carpenter
181:Dinosauria
2565:Talarurus
2445:Saichania
2232:Europelta
2212:Texasetes
2074:Taohelong
1971:Invictarx
1873:Stegouros
1591:Kingdom:
1513:244799975
1497:1476-4687
1449:985402380
1414:250422526
1398:1932-8486
1360:: 111–160
1281:Maryanska
974:Taohelong
732:Stegouros
484:squamosal
404:Discovery
351:Tithonian
339:exposures
271:Species:
154:Kingdom:
148:Eukaryota
2672:Wikidata
2638:Category
2628:Timeline
2572:Ziapelta
2044:Gastonia
1941:Gastonia
1599:Chordata
1597:Phylum:
1593:Animalia
1534:Archived
1505:34853468
1406:35815600
1232:See also
838:Gastonia
610:(2021):
469:skeleton
434:Colorado
343:Jurassic
331:holotype
323:gargoyle
235:Family:
168:Chordata
164:Phylum:
158:Animalia
144:Domain:
2731:4946560
2718:1233089
2705:4823339
2692:4530841
2678:Q134736
2460:Tarchia
2452:Shanxia
1477:Bibcode
594:as the
582:of the
492:osseous
477:deltoid
335:Wyoming
296:†
275:†
248:Genus:
105:↓
1511:
1503:
1495:
1469:Nature
1447:
1437:
1412:
1404:
1396:
1321:
1267:Nature
608:et al.
572:et al.
446:et al.
430:Denver
380:et al.
354:stages
307:et al.
263:et al.
2744:63703
2713:IRMNG
2417:Datai
1780:Minmi
1622:Clade
1613:Clade
1604:Clade
1509:S2CID
1410:S2CID
592:genus
473:taxon
465:skull
220:Clade
204:Clade
188:Clade
175:Clade
2700:GBIF
2579:Zuul
1682:see
1655:see
1501:PMID
1493:ISSN
1445:OCLC
1435:ISBN
1402:PMID
1394:ISSN
1319:ISBN
482:and
467:and
450:ICZN
363:The
309:1998
265:1998
44:Preęž’
2687:EoL
1485:doi
1473:600
1384:doi
1380:306
1358:237
349:to
337:in
2763::
2741::
2728::
2715::
2702::
2689::
2674::
1624::
1615::
1606::
1507:.
1499:.
1491:.
1483:.
1471:.
1467:.
1443:.
1408:.
1400:.
1392:.
1378:.
1374:.
1346:^
1305:^
432:,
428:,
367:,
360:.
356:)
222::
206::
190::
177::
94:Pg
38:Ma
33:,
2470:?
2455:?
2374:?
2325:?
2303:?
2295:?
2145:?
2109:?
2077:?
2062:?
2047:?
2039:?
1997:?
1989:?
1981:?
1966:?
1944:?
1936:?
1868:?
1826:?
1804:?
1567:e
1560:t
1553:v
1515:.
1487::
1479::
1451:.
1416:.
1386::
1327:.
345:(
252:â€
239:â€
226:â€
210:â€
194:â€
99:N
89:K
84:J
79:T
74:P
69:C
64:D
59:S
54:O
49:ęž’
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