386:
159:
109:
487:
particularly convoluted: it appears that the animals were arboreal at some time in the far distant past, moving afterward to the ground—gaining long kangaroo-like feet in the process — before returning to the trees, where they further developed a shortening and broadening of the hind feet and a novel
462:
Two key anatomical features, in combination, identify
Diprotodontia. Members of the order are, first, "diprotodont" (meaning "two front teeth"): they have a pair of large, procumbent incisors on the lower jaw, a common feature of many early groups of mammals and mammaliforms. The diprotodont jaw is
459:) became extinct when humans first arrived in Australia about 50,000 years ago. Their extinction possibly occurred as a direct result of hunting, but was more probably a result of widespread habitat changes brought about by human activities—notably the use of fire.
1440:Álvarez-Carretero, Sandra; Tamuri, Asif U.; Battini, Matteo; Nascimento, Fabrícia F.; Carlisle, Emily; Asher, Robert J.; Yang, Ziheng; Donoghue, Philip C.J.; dos Reis, Mario (2022). "A species-level timeline of mammal evolution integrating phylogenomic data".
482:
share it) and is generally posited as an adaptation to assist in climbing. Many modern diprotodonts, however, are strictly terrestrial, and have evolved further adaptations to their feet to better suit this lifestyle. This makes the history of the
1283:
475:", a fusing of the second and third digits of the foot up to the base of the claws, which leaves the claws themselves separate. Digit five is usually absent, and digit four is often greatly enlarged.
915:
families are split into two superfamilies. The
Macropodiformes are probably nested within the Phalangeriformes, though whether they are sister to Phalangeroidea or Petauroidea is debated.
1477:Álvarez-Carretero, Sandra; Tamuri, Asif U.; Battini, Matteo; Nascimento, Fabrícia F.; Carlisle, Emily; Asher, Robert J.; Yang, Ziheng; Donoghue, Philip C.J.; dos Reis, Mario (2022).
1405:
506:
1478:
448:, but their genesis certainly lies earlier than this, as large gaps occur in Australia's fossil record, with virtually no fossil record at all in geologically active
4579:
4631:
4695:
4553:
1341:
4592:
2087:
1554:
4610:
4597:
1516:"An emerging consensus in the evolution, phylogeny, and systematics of marsupials and their fossil relatives (Metatheria)"
4700:
429:, but these seem to have arisen as relatively recent adaptations from the mainstream herbivorous lifestyle. The extinct
3201:
1617:
1258:
4532:
3907:
1355:"Inferring the mammal tree: Species-level sets of phylogenies for questions in ecology, evolution and conservation"
1215:
4636:
4545:
4618:
3423:
3150:
3066:
3057:
3934:
3477:
3219:
452:. The great diversity of known Oligocene diprotodonts suggests the order began to diverge well beforehand.
158:
4605:
4475:
4452:
3450:
3441:
2253:
2244:
2080:
1220:
1248:
4033:
4006:
3943:
3790:
3405:
2992:
2983:
390:
4480:
1514:
Eldridge, Mark D B; Beck, Robin M D; Croft, Darin A; Travouillon, Kenny J; Fox, Barry J (2019-05-23).
4024:
3988:
3781:
3228:
2623:
2605:
1499:
1462:
4042:
3979:
3880:
3414:
3177:
3132:
2734:
2632:
2540:
4430:
3925:
3629:
3546:
3504:
3237:
3141:
3123:
3114:
3042:
3033:
2641:
2504:
4690:
4662:
4392:
3997:
3970:
3588:
3486:
3432:
3159:
3090:
3081:
3024:
2886:
2877:
2752:
2462:
2444:
2435:
2073:
1086:
4685:
4657:
4125:
3722:
3704:
3671:
3662:
3298:
3289:
3210:
3168:
2919:
2710:
2564:
2513:
2399:
1610:
1210:
1128:
1090:
394:
133:
4649:
4584:
471:
have only one pair), and no lower canines. The second trait distinguishing diprotodonts is "
4501:
4439:
4322:
4152:
4143:
4107:
4015:
3513:
2761:
2743:
2531:
2348:
523:
456:
141:
8:
4340:
4304:
4213:
4204:
4098:
3961:
3814:
3638:
3468:
3373:
3364:
2844:
2665:
2656:
2010:(Rats, guinea pigs, squirrels, beavers, chinchillas, porcupines, capybaras and relatives)
1188:
787:
907:
which contained all other families. Kirsch et al. (1997) split the families into three
4359:
4350:
4313:
4134:
3749:
3274:
3265:
2835:
2614:
2339:
1381:
1354:
1192:
1124:
1113:
153:
4444:
1284:"A phylogeny of Diprotodontia (Marsupialia) based on sequences for five nuclear genes"
455:
Many of the largest and least athletic diprotodonts (along with a wide range of other
4644:
4488:
4457:
3764:
2480:
1978:
1574:
1535:
1386:
1306:
1264:
1254:
385:
4272:
3740:
2588:
2426:
2417:
2285:
1999:
1740:
1677:
1603:
1566:
1527:
1494:
1486:
1457:
1449:
1420:
1376:
1366:
1298:
1026:
1009:
912:
610:
350:
276:
257:
1330:
531:
Cladogram of
Diprotodontia by Upham et al. 2019 and Álvarez-Carretero et al. 2022
4493:
4245:
4116:
3832:
3597:
3564:
3537:
3330:
2973:
2955:
2946:
2910:
2853:
2408:
2381:
2156:
1892:
1793:
1761:
1425:
1371:
1140:
1109:
1102:
1098:
993:
989:
971:
967:
944:
737:
656:
563:
479:
430:
334:
261:
125:
4623:
1570:
1302:
4540:
4527:
3823:
3713:
3555:
3105:
3007:
2862:
2573:
2522:
2065:
1938:
1884:
1785:
1769:
1490:
1453:
1282:
Meredith, Robert W.; Westerman, Michael; Springer, Mark S. (26 February 2009).
1082:
1016:
982:
978:
501:
330:
98:
1989:(Pigs, camels, hippos, deer, buffalo, gazelles, giraffes, whales and dolphins)
4679:
4176:
4167:
3731:
3695:
3388:
3192:
2453:
2390:
2300:
2220:
2211:
2166:
2144:
1929:
1539:
1531:
1244:
1052:
1022:
926:
892:
549:
511:
484:
253:
223:
62:
1268:
4415:
3856:
3841:
3686:
3612:
3342:
2937:
2826:
2798:
2789:
2779:
2683:
2555:
2471:
1946:
1876:
1777:
1732:
1578:
1390:
1310:
1281:
1253:(3rd ed.). Baltimore: Johns Hopkins University Press. pp. 43–70.
1240:
1181:
1154:
1094:
1071:
1067:
1041:
1000:
904:
824:
700:
600:
504:(23.03 - 28.4 million years ago), and the earliest identifiable species is
362:
238:
4566:
4467:
4424:
3579:
3255:
2928:
2725:
2495:
2372:
2322:
2235:
1986:
1908:
1826:
1515:
1120:
1061:
1037:
678:
441:
414:
37:
1476:
1439:
4194:
3805:
2816:
2015:
1868:
1711:
1668:
1652:
1595:
1406:"DR_on4phylosCompared_linear_richCol_justScale_ownColors_withTips_80in"
1169:
1147:
1078:
933:
809:
722:
579:
472:
449:
422:
373:
367:
338:
82:
47:
108:
4571:
4514:
4081:
3528:
2363:
2128:
2023:
1970:
1847:
1839:
1719:
1685:
1048:
961:
955:
445:
434:
426:
406:
210:
170:
87:
31:
4558:
4386:
4519:
4409:
3898:
3653:
2901:
2701:
2116:
1962:
1818:
1646:
1158:
908:
888:
418:
342:
190:
77:
72:
57:
52:
42:
1555:"Of koalas and marsupial lions: the vombatiform radiation, part I"
4287:
4228:
3495:
3459:
3350:
2039:
1916:
1748:
1658:
1250:
Mammal
Species of the World: A Taxonomic and Geographic Reference
1177:
1173:
1162:
410:
346:
92:
67:
4263:
4254:
4066:
4057:
3889:
2308:
2193:
2122:
2110:
2031:
2007:
1855:
1702:
1640:
1627:
1030:
937:
896:
497:
468:
464:
433:("marsupial lions") are the only known group to have exhibited
358:
200:
180:
4506:
2175:
1900:
948:
900:
354:
117:
1513:
1404:
Upham, Nathan S.; Esselstyn, Jacob A.; Jetz, Walter (2019).
1353:
Upham, Nathan S.; Esselstyn, Jacob A.; Jetz, Walter (2019).
321:
318:
300:
294:
282:
309:
1954:
2283:
393:
characteristic of the diprotodonts are evident in this
425:
are almost unique among vertebrates in being largely
312:
285:
361:, and many others. Extinct diprotodonts include the
315:
306:
297:
291:
288:
279:
1546:
463:
short, usually with three pairs of upper incisors (
303:
2042:(lorises, lemurs, tarsiers, monkeys, apes, humans)
1403:
1352:
478:Syndactyly is not particularly common (though the
4677:
2095:
1676:
444:. The earliest known fossils date to the late
4348:
2081:
1611:
3328:
2164:
333:"two forward teeth") is the largest extant
2142:
2088:
2074:
1618:
1604:
107:
1565:(1). Scientific American, Inc.: 240–250.
1500:1983/de841853-d57b-40d9-876f-9bfcf7253f12
1498:
1463:1983/de841853-d57b-40d9-876f-9bfcf7253f12
1461:
1424:
1380:
1370:
1346:
1201:† means extinct family, genus or species
1949:(Hedgehogs, shrews, moles and relatives)
1625:
384:
341:, with about 155 species, including the
2787:
4678:
1239:
4391:
4390:
3327:
2282:
2141:
2069:
1599:
1559:Molecular Phylogenetics and Evolution
1552:
1291:Molecular Phylogenetics and Evolution
409:, as were most of those that are now
395:Kangaroo Island western grey kangaroo
4546:66ad7577-a578-4f66-b057-fb417b9526e1
1335:
1233:
500:of Diprotodontia dates back to the
405:Living diprotodonts are almost all
13:
3908:Eastern short-eared rock-wallaby (
380:
377:, the so-called "marsupial lion".
116:Clockwise from upper left: female
14:
4712:
4696:Extant Chattian first appearances
1981:(Horses, rhinoceroses and tapirs)
1479:"4705sp_colours_mammal-time.tree"
517:
1216:List of recently extinct mammals
491:
480:Australian omnivorous marsupials
421:diprotodonts are known, and the
399:Macropus fuliginosus fuliginosus
275:
157:
35:
3151:Herbert River ringtail possum (
3067:Rock-haunting ringtail possum (
1150:: (basal quadrupedal kangaroos)
440:Diprotodonts are restricted to
3478:Golden-mantled tree-kangaroo (
3220:Plush-coated ringtail possum (
3202:D'Albertis's ringtail possum (
1507:
1470:
1433:
1397:
1324:
1275:
1:
2254:Northern hairy-nosed wombat (
2245:Southern hairy-nosed wombat (
1226:
891:in Diprotodontia were noted:
4034:Purple-necked rock-wallaby (
4007:Black-flanked rock-wallaby (
3944:Yellow-footed rock-wallaby (
3791:Northern nail-tail wallaby (
3424:Goodfellow's tree-kangaroo (
2993:Lemur-like ringtail possum (
1426:10.1371/journal.pbio.3000494
1372:10.1371/journal.pbio.3000494
7:
4025:Brush-tailed rock-wallaby (
3935:Rothschild's rock-wallaby (
3782:Bridled nail-tail wallaby (
3229:Reclusive ringtail possum (
2624:Mountain brushtail possum (
2606:Northern brushtail possum (
1571:10.1016/j.ympev.2004.05.004
1303:10.1016/j.ympev.2009.02.009
1221:List of prehistoric mammals
1204:
507:Hypsiprymnodon bartholomaii
391:mandibular central incisors
10:
4717:
4701:Taxa named by Richard Owen
4043:Mount Claro rock-wallaby (
3881:Short-eared rock-wallaby (
3451:Matschie's tree-kangaroo (
3442:Lumholtz's tree-kangaroo (
3178:Vogelkop ringtail possum (
3133:Cinereus ringtail possum (
3043:Southern greater glider (
2735:Long-tailed pygmy possum (
2633:Coppery brushtail possum (
2541:Blue-eyed spotted cuscus (
2018:(Rabbits, hares and pikas)
1973:(Dogs, cats and relatives)
1879:(Tenrecs and golden moles)
1491:10.1038/s41586-021-04341-1
1454:10.1038/s41586-021-04341-1
521:
16:Order of marsupial mammals
4399:
4339:
4285:
4226:
4202:
4193:
4165:
4079:
4055:
3926:Proserpine rock-wallaby (
3854:
3803:
3762:
3684:
3651:
3630:Spectacled hare-wallaby (
3610:
3577:
3547:White-striped dorcopsis (
3526:
3406:Bennett's tree-kangaroo (
3386:
3362:
3341:
3337:
3323:
3287:
3263:
3254:
3238:Coppery ringtail possum (
3190:
3142:Painted ringtail possum (
3124:Weyland ringtail possum (
3115:Lowland ringtail possum (
3103:
3079:
3055:
3034:Northern greater glider (
3005:
2981:
2972:
2899:
2875:
2824:
2815:
2778:
2723:
2699:
2682:
2654:
2642:Common brushtail possum (
2586:
2553:
2505:Admiralty Island cuscus (
2493:
2361:
2320:
2299:
2295:
2278:
2233:
2209:
2192:
2155:
2151:
2137:
2105:
1998:
1937:
1928:
1867:
1838:
1817:
1804:(Kangaroos and relatives)
1760:
1731:
1710:
1701:
1667:
1635:
1342:The Paleobiology Database
887:Until recently, only two
822:
807:
800:
785:
778:
735:
720:
713:
698:
691:
676:
669:
661:
635:
622:
615:
604:
577:
561:
553:
543:
251:
246:
154:Scientific classification
152:
115:
106:
23:
3998:Unadorned rock-wallaby (
3989:Herbert's rock-wallaby (
3971:Cape York rock-wallaby (
3956:P. lateralis/penicillata
3487:Lowlands tree-kangaroo (
3433:Grizzled tree-kangaroo (
3160:Masked ringtail possum (
3091:Common ringtail possum (
3025:Central greater glider (
2753:Tasmanian pygmy possum (
2445:Northern common cuscus (
2436:Southern common cuscus (
1796:(Bilbies and bandicoots)
1247:; Reeder, D. M. (eds.).
3980:Godman's rock-wallaby (
3723:Western brush wallaby (
3705:Black-striped wallaby (
3672:Eastern grey kangaroo (
3663:Western grey kangaroo (
3415:Doria's tree-kangaroo (
3299:Feather-tailed possum (
3211:Green ringtail possum (
3169:Pygmy ringtail possum (
2920:Yellow-bellied glider (
2711:Mountain pygmy possum (
2565:Sulawesi dwarf cuscus (
2514:Common spotted cuscus (
2400:Eastern common cuscus (
1688:(Platypus and echidnas)
911:. In addition, the six
4153:Red-necked pademelon (
4144:Red-legged pademelon (
4016:Mareeba rock-wallaby (
3514:Ursine tree-kangaroo (
3505:Seri's tree-kangaroo (
2762:Eastern pygmy possum (
2744:Western pygmy possum (
2532:Black-spotted cuscus (
2349:Sulawesi bear cuscus (
1919:(Dugongs and manatees)
1858:(Anteaters and sloths)
1553:Naish, Darren (2004).
1532:10.1093/jmammal/gyz018
895:which encompassed the
402:
122:Phascolarctos cinereus
4658:Paleobiology Database
4305:Long-footed potoroo (
4214:Rufous rat-kangaroo (
4099:Tasmanian pademelon (
3962:Allied rock-wallaby (
3815:Antilopine kangaroo (
3772:(nail-tail wallabies)
3639:Rufous hare-wallaby (
3589:Macleay's dorcopsis (
3374:Banded hare-wallaby (
2887:Leadbeater's possum (
2845:Long-fingered triok (
2666:Scaly-tailed possum (
2463:Rothschild's cuscus (
1788:(Quolls and dunnarts)
1211:List of mammal genera
1129:feather-tailed possum
1091:yellow-bellied glider
388:
134:eastern grey kangaroo
4541:Fauna Europaea (new)
4360:Musky rat-kangaroo (
4314:Long-nosed potoroo (
4135:Mountain pademelon (
4126:Calaby's pademelon (
3750:Red-necked wallaby (
3275:Feathertail glider (
2836:Great-tailed triok (
2615:Short-eared possum (
2340:Talaud bear cuscus (
1520:Journal of Mammalogy
559:Phascolarctimorphia
524:List of diprotodonts
457:Australian megafauna
142:Sulawesi bear cuscus
4341:Hypsiprymnodontidae
4323:Gilbert's potoroo (
4108:Brown's pademelon (
2596:(brushtail possums)
2402:P. intercastellanus
1189:Hypsiprymnodontidae
1114:ring-tailed possums
1087:Leadbeater's possum
919:Order Diprotodontia
788:Hypsiprymnodontidae
496:The earliest known
4273:Northern bettong (
3741:Whiptail wallaby (
2427:Telefomin cuscus (
2418:Blue-eyed cuscus (
1772:(Monito del monte)
1193:musky rat-kangaroo
1125:feathertail glider
437:on a large scale.
403:
138:Macropus giganteus
4673:
4672:
4645:Open Tree of Life
4393:Taxon identifiers
4384:
4383:
4380:
4379:
4376:
4375:
4372:
4371:
4335:
4334:
4298:
4246:Eastern bettong (
4239:
4189:
4188:
4117:Dusky pademelon (
4092:
4036:P. purpureicollis
3867:
3833:Common wallaroo (
3775:
3632:L. conspicillatus
3623:
3598:Small dorcopsis (
3565:Brown dorcopsis (
3538:Black dorcopsis (
3399:
3356:
3319:
3318:
3315:
3314:
3311:
3310:
3250:
3249:
3018:
3015:(greater gliders)
2968:
2967:
2956:Squirrel glider (
2947:Mahogany glider (
2911:Northern glider (
2811:
2810:
2774:
2773:
2693:
2678:
2677:
2599:
2409:Woodlark cuscus (
2382:Mountain cuscus (
2333:
2314:
2289:
2286:Phalangeriformes
2274:
2273:
2270:
2269:
2266:
2265:
2203:
2188:
2187:
2063:
2062:
2059:
2058:
2055:
2054:
2051:
2050:
2043:
2035:
2027:
2019:
2011:
1990:
1982:
1974:
1966:
1958:
1950:
1920:
1912:
1904:
1896:
1888:
1887:(Elephant shrews)
1880:
1859:
1851:
1832:
1813:
1812:
1805:
1797:
1789:
1781:
1780:(Marsupial moles)
1778:Notoryctemorphia
1773:
1752:
1744:
1741:Paucituberculata
1725:
1697:
1696:
1689:
1448:(7896): 263–267.
1027:brushtail possums
885:
884:
880:
879:
871:
870:
862:
861:
853:
852:
844:
843:
835:
834:
766:
765:
757:
756:
748:
747:
646:
645:
590:
589:
488:climbing method.
268:
267:
242:
130:Petaurus gracilis
4708:
4666:
4665:
4653:
4652:
4640:
4639:
4627:
4626:
4624:NHMSYS0000066025
4614:
4613:
4601:
4600:
4588:
4587:
4575:
4574:
4562:
4561:
4549:
4548:
4536:
4535:
4523:
4522:
4510:
4509:
4497:
4496:
4484:
4483:
4471:
4470:
4461:
4460:
4448:
4447:
4435:
4434:
4433:
4420:
4419:
4418:
4388:
4387:
4346:
4345:
4297:
4296:
4292:
4238:
4237:
4233:
4200:
4199:
4091:
4090:
4086:
3866:
3865:
3864:(rock-wallabies)
3861:
3824:Black wallaroo (
3774:
3773:
3769:
3714:Tammar wallaby (
3622:
3621:
3620:(hare-wallabies)
3617:
3556:Gray dorcopsis (
3398:
3397:
3396:(tree-kangaroos)
3393:
3360:
3359:
3355:
3354:
3346:
3339:
3338:
3325:
3324:
3261:
3260:
3017:
3016:
3012:
2979:
2978:
2863:Striped possum (
2822:
2821:
2785:
2784:
2697:
2696:
2692:
2691:
2687:
2668:W. squamicaudata
2598:
2597:
2593:
2574:Banggai cuscus (
2523:Waigeou cuscus (
2481:Stein's cuscus (
2332:
2331:
2327:
2318:
2317:
2313:
2312:
2304:
2297:
2296:
2287:
2280:
2279:
2207:
2206:
2202:
2201:
2197:
2162:
2161:
2153:
2152:
2139:
2138:
2090:
2083:
2076:
2067:
2066:
2041:
2033:
2025:
2017:
2009:
2000:Euarchontoglires
1988:
1980:
1972:
1964:
1956:
1948:
1935:
1934:
1918:
1910:
1902:
1894:
1886:
1878:
1857:
1849:
1836:
1835:
1831:
1830:
1822:
1803:
1795:
1794:Peramelemorphia
1787:
1779:
1771:
1750:
1749:Didelphimorphia
1743:(Shrew opossums)
1742:
1729:
1728:
1724:
1723:
1715:
1708:
1707:
1687:
1678:Australosphenida
1674:
1673:
1620:
1613:
1606:
1597:
1596:
1590:
1589:
1587:
1585:
1550:
1544:
1543:
1511:
1505:
1504:
1502:
1485:(602): 263–267.
1474:
1468:
1467:
1465:
1437:
1431:
1430:
1428:
1410:
1401:
1395:
1394:
1384:
1374:
1365:(12): e3000494.
1350:
1344:
1339:
1333:
1328:
1322:
1321:
1319:
1317:
1288:
1279:
1273:
1272:
1237:
1010:Phalangeriformes
983:marsupial tapirs
913:Phalangeriformes
803:
802:
781:
780:
716:
715:
694:
693:
672:
671:
664:
663:
618:
617:
611:Phalangeriformes
607:
606:
556:
555:
546:
545:
538:
537:
528:
527:
328:
327:
324:
323:
320:
317:
314:
311:
308:
305:
302:
299:
296:
293:
290:
287:
284:
281:
258:Phalangeriformes
237:
162:
161:
146:Ailurops ursinus
111:
97:
34:
27:Temporal range:
21:
20:
4716:
4715:
4711:
4710:
4709:
4707:
4706:
4705:
4676:
4675:
4674:
4669:
4661:
4656:
4648:
4643:
4635:
4630:
4622:
4617:
4609:
4604:
4596:
4591:
4583:
4578:
4570:
4565:
4557:
4552:
4544:
4539:
4531:
4526:
4518:
4513:
4505:
4500:
4492:
4487:
4479:
4474:
4466:
4464:
4456:
4451:
4443:
4438:
4429:
4428:
4423:
4414:
4413:
4408:
4395:
4385:
4368:
4331:
4294:
4293:
4291:
4281:
4235:
4234:
4232:
4222:
4185:
4177:Swamp wallaby (
4161:
4101:T. billardierii
4088:
4087:
4085:
4075:
4051:
3863:
3862:
3860:
3850:
3799:
3771:
3770:
3768:
3758:
3732:Parma wallaby (
3696:Agile wallaby (
3680:
3647:
3619:
3618:
3616:
3606:
3573:
3522:
3480:D. pulcherrimus
3408:D. bennettianus
3395:
3394:
3392:
3382:
3348:
3347:
3345:
3333:
3331:Macropodiformes
3307:
3283:
3246:
3186:
3153:P. herbertensis
3099:
3075:
3051:
3014:
3013:
3011:
3001:
2974:Pseudocheiridae
2964:
2958:P. norfolcensis
2895:
2871:
2807:
2770:
2719:
2690:(pygmy possums)
2689:
2688:
2686:
2674:
2650:
2595:
2594:
2592:
2582:
2549:
2489:
2454:Ornate cuscus (
2391:Ground cuscus (
2357:
2330:(bear cuscuses)
2329:
2328:
2326:
2306:
2305:
2303:
2291:
2262:
2229:
2221:Common wombat (
2199:
2198:
2196:
2184:
2157:Phascolarctidae
2147:
2133:
2101:
2094:
2064:
2047:
1994:
1979:Perissodactyla
1924:
1863:
1824:
1823:
1821:
1809:
1786:Dasyuromorphia
1770:Microbiotheria
1762:Australidelphia
1756:
1717:
1716:
1714:
1693:
1663:
1631:
1624:
1594:
1593:
1583:
1581:
1551:
1547:
1512:
1508:
1475:
1471:
1438:
1434:
1408:
1402:
1398:
1351:
1347:
1340:
1336:
1329:
1325:
1315:
1313:
1286:
1280:
1276:
1261:
1238:
1234:
1229:
1207:
1141:Macropodiformes
1110:Pseudocheiridae
1103:squirrel glider
1099:mahogany glider
994:marsupial lions
990:Thylacoleonidae
968:Diprotodontidae
945:Phascolarctidae
940:(three species)
881:
872:
863:
854:
845:
836:
767:
758:
749:
738:Pseudocheiridae
657:Macropodiformes
647:
591:
575:Vombatomorphia
564:Phascolarctidae
526:
520:
494:
383:
381:Characteristics
278:
274:
262:Macropodiformes
260:
256:
236:
156:
126:mahogany glider
102:
96:
95:
90:
85:
80:
75:
70:
65:
60:
55:
50:
45:
40:
29:
28:
25:
17:
12:
11:
5:
4714:
4704:
4703:
4698:
4693:
4688:
4671:
4670:
4668:
4667:
4654:
4641:
4628:
4615:
4602:
4589:
4576:
4563:
4550:
4537:
4528:Fauna Europaea
4524:
4511:
4498:
4485:
4472:
4462:
4449:
4436:
4421:
4405:
4403:
4397:
4396:
4382:
4381:
4378:
4377:
4374:
4373:
4370:
4369:
4367:
4366:
4356:
4354:
4351:Hypsiprymnodon
4343:
4337:
4336:
4333:
4332:
4330:
4329:
4320:
4316:P. tridactylus
4311:
4301:
4299:
4283:
4282:
4280:
4279:
4270:
4266:B. penicillata
4261:
4252:
4242:
4240:
4224:
4223:
4221:
4220:
4210:
4208:
4197:
4191:
4190:
4187:
4186:
4184:
4183:
4173:
4171:
4163:
4162:
4160:
4159:
4150:
4141:
4132:
4123:
4114:
4105:
4095:
4093:
4077:
4076:
4074:
4073:
4063:
4061:
4053:
4052:
4050:
4049:
4040:
4031:
4027:P. penicillata
4022:
4013:
4004:
3995:
3986:
3977:
3968:
3951:
3950:
3941:
3937:P. rothschildi
3932:
3915:
3914:
3905:
3896:
3887:
3870:
3868:
3852:
3851:
3849:
3848:
3842:Red kangaroo (
3839:
3830:
3821:
3817:O. antilopinus
3811:
3809:
3801:
3800:
3798:
3797:
3788:
3778:
3776:
3760:
3759:
3757:
3756:
3752:N. rufogriseus
3747:
3738:
3729:
3720:
3711:
3702:
3692:
3690:
3682:
3681:
3679:
3678:
3669:
3665:M. fuliginosus
3659:
3657:
3649:
3648:
3646:
3645:
3636:
3626:
3624:
3608:
3607:
3605:
3604:
3595:
3585:
3583:
3575:
3574:
3572:
3571:
3562:
3553:
3544:
3534:
3532:
3524:
3523:
3521:
3520:
3511:
3502:
3493:
3484:
3475:
3466:
3457:
3448:
3439:
3430:
3426:D. goodfellowi
3421:
3412:
3402:
3400:
3384:
3383:
3381:
3380:
3370:
3368:
3357:
3335:
3334:
3321:
3320:
3317:
3316:
3313:
3312:
3309:
3308:
3306:
3305:
3295:
3293:
3285:
3284:
3282:
3281:
3271:
3269:
3258:
3252:
3251:
3248:
3247:
3245:
3244:
3235:
3226:
3217:
3208:
3198:
3196:
3188:
3187:
3185:
3184:
3175:
3166:
3157:
3148:
3139:
3130:
3121:
3111:
3109:
3106:Pseudochirulus
3101:
3100:
3098:
3097:
3087:
3085:
3077:
3076:
3074:
3073:
3063:
3061:
3053:
3052:
3050:
3049:
3040:
3031:
3021:
3019:
3003:
3002:
3000:
2999:
2989:
2987:
2976:
2970:
2969:
2966:
2965:
2963:
2962:
2953:
2944:
2938:Sugar glider (
2935:
2926:
2917:
2907:
2905:
2897:
2896:
2894:
2893:
2889:G. leadbeateri
2883:
2881:
2873:
2872:
2870:
2869:
2860:
2854:Tate's triok (
2851:
2842:
2832:
2830:
2819:
2813:
2812:
2809:
2808:
2806:
2805:
2799:Honey possum (
2795:
2793:
2782:
2776:
2775:
2772:
2771:
2769:
2768:
2759:
2750:
2741:
2731:
2729:
2721:
2720:
2718:
2717:
2707:
2705:
2694:
2680:
2679:
2676:
2675:
2673:
2672:
2662:
2660:
2652:
2651:
2649:
2648:
2639:
2630:
2626:T. cunninghami
2621:
2612:
2608:T. arnhemensis
2602:
2600:
2584:
2583:
2581:
2580:
2571:
2561:
2559:
2551:
2550:
2548:
2547:
2538:
2529:
2520:
2511:
2501:
2499:
2491:
2490:
2488:
2487:
2478:
2472:Silky cuscus (
2469:
2465:P. rothschildi
2460:
2451:
2442:
2433:
2424:
2415:
2406:
2397:
2388:
2379:
2369:
2367:
2359:
2358:
2356:
2355:
2346:
2336:
2334:
2315:
2293:
2292:
2276:
2275:
2272:
2271:
2268:
2267:
2264:
2263:
2261:
2260:
2251:
2241:
2239:
2231:
2230:
2228:
2227:
2217:
2215:
2204:
2190:
2189:
2186:
2185:
2183:
2182:
2172:
2170:
2159:
2149:
2148:
2135:
2134:
2132:
2131:
2125:
2119:
2113:
2106:
2103:
2102:
2093:
2092:
2085:
2078:
2070:
2061:
2060:
2057:
2056:
2053:
2052:
2049:
2048:
2046:
2045:
2037:
2029:
2021:
2013:
2004:
2002:
1996:
1995:
1993:
1992:
1984:
1976:
1968:
1960:
1952:
1943:
1941:
1939:Laurasiatheria
1932:
1926:
1925:
1923:
1922:
1914:
1906:
1898:
1893:Tubulidentata
1890:
1885:Macroscelidea
1882:
1873:
1871:
1865:
1864:
1862:
1861:
1853:
1844:
1842:
1833:
1815:
1814:
1811:
1810:
1808:
1807:
1802:Diprotodontia
1799:
1791:
1783:
1775:
1766:
1764:
1758:
1757:
1755:
1754:
1746:
1737:
1735:
1726:
1705:
1699:
1698:
1695:
1694:
1692:
1691:
1682:
1680:
1671:
1665:
1664:
1662:
1661:
1655:
1649:
1643:
1636:
1633:
1632:
1623:
1622:
1615:
1608:
1600:
1592:
1591:
1545:
1526:(3): 802–837.
1506:
1469:
1432:
1396:
1345:
1334:
1323:
1297:(3): 554–571.
1274:
1259:
1231:
1230:
1228:
1225:
1224:
1223:
1218:
1213:
1206:
1203:
1199:
1198:
1197:
1196:
1185:
1166:
1151:
1136:
1135:
1134:
1133:
1132:
1117:
1106:
1083:striped possum
1075:
1058:
1057:
1056:
1053:sprite possums
1049:Ektopodontidae
1045:
1034:
1017:Phalangeroidea
1005:
1004:
1003:
997:
986:
979:Palorchestidae
975:
964:
958:
952:
941:
883:
882:
878:
877:
874:
873:
869:
868:
865:
864:
860:
859:
856:
855:
851:
850:
847:
846:
842:
841:
838:
837:
833:
832:
829:
828:
821:
818:
817:
814:
813:
806:
801:
799:
796:
795:
792:
791:
784:
779:
777:
776:Macropodoidea
773:
772:
769:
768:
764:
763:
760:
759:
755:
754:
751:
750:
746:
745:
742:
741:
734:
731:
730:
727:
726:
719:
714:
712:
709:
708:
705:
704:
697:
692:
690:
687:
686:
683:
682:
675:
670:
668:
662:
660:
653:
652:
649:
648:
644:
643:
640:
639:
637:Phalangeridae
634:
631:
630:
627:
626:
621:
616:
614:
605:
603:
597:
596:
593:
592:
588:
587:
584:
583:
576:
572:
571:
568:
567:
560:
554:
552:
544:
542:
541:Diprotodontia
536:
533:
532:
522:Main article:
519:
518:Classification
516:
502:Late Oligocene
493:
490:
485:tree-kangaroos
431:thylacoleonids
389:The prominent
382:
379:
266:
265:
249:
248:
244:
243:
231:
227:
226:
221:
214:
213:
208:
204:
203:
198:
194:
193:
188:
184:
183:
178:
174:
173:
168:
164:
163:
150:
149:
113:
112:
104:
103:
99:Late Oligocene
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
26:
15:
9:
6:
4:
3:
2:
4713:
4702:
4699:
4697:
4694:
4692:
4691:Mammal orders
4689:
4687:
4684:
4683:
4681:
4664:
4659:
4655:
4651:
4646:
4642:
4638:
4633:
4629:
4625:
4620:
4616:
4612:
4607:
4603:
4599:
4594:
4590:
4586:
4581:
4577:
4573:
4568:
4564:
4560:
4555:
4551:
4547:
4542:
4538:
4534:
4529:
4525:
4521:
4516:
4512:
4508:
4503:
4499:
4495:
4490:
4486:
4482:
4477:
4473:
4469:
4463:
4459:
4458:Diprotodontia
4454:
4450:
4446:
4445:Diprotodontia
4441:
4437:
4432:
4431:Diprotodontia
4426:
4422:
4417:
4411:
4407:
4406:
4404:
4402:
4401:Diprotodontia
4398:
4394:
4389:
4365:
4363:
4358:
4357:
4355:
4353:
4352:
4347:
4344:
4342:
4338:
4328:
4326:
4321:
4319:
4317:
4312:
4310:
4308:
4303:
4302:
4300:
4290:
4289:
4284:
4278:
4276:
4271:
4269:
4267:
4262:
4260:
4258:
4253:
4251:
4249:
4244:
4243:
4241:
4231:
4230:
4225:
4219:
4217:
4212:
4211:
4209:
4207:
4206:
4201:
4198:
4196:
4192:
4182:
4180:
4175:
4174:
4172:
4170:
4169:
4164:
4158:
4156:
4151:
4149:
4147:
4146:T. stigmatica
4142:
4140:
4138:
4133:
4131:
4129:
4124:
4122:
4120:
4115:
4113:
4111:
4106:
4104:
4102:
4097:
4096:
4094:
4084:
4083:
4078:
4072:
4070:
4069:S. brachyurus
4065:
4064:
4062:
4060:
4059:
4054:
4048:
4046:
4041:
4039:
4037:
4032:
4030:
4028:
4023:
4021:
4019:
4014:
4012:
4010:
4005:
4003:
4001:
3996:
3994:
3992:
3987:
3985:
3983:
3978:
3976:
3974:
3969:
3967:
3965:
3959:
3958:species group
3957:
3953:
3952:
3949:
3947:
3942:
3940:
3938:
3933:
3931:
3929:
3928:P. persephone
3923:
3922:species group
3921:
3917:
3916:
3913:
3911:
3906:
3904:
3902:
3897:
3895:
3893:
3888:
3886:
3884:
3883:P. brachyotis
3878:
3877:species group
3876:
3875:P. brachyotis
3872:
3871:
3869:
3859:
3858:
3853:
3847:
3845:
3840:
3838:
3836:
3831:
3829:
3827:
3822:
3820:
3818:
3813:
3812:
3810:
3808:
3807:
3802:
3796:
3794:
3789:
3787:
3785:
3780:
3779:
3777:
3767:
3766:
3761:
3755:
3753:
3748:
3746:
3744:
3739:
3737:
3735:
3730:
3728:
3726:
3721:
3719:
3717:
3712:
3710:
3708:
3703:
3701:
3699:
3694:
3693:
3691:
3689:
3688:
3683:
3677:
3675:
3670:
3668:
3666:
3661:
3660:
3658:
3656:
3655:
3650:
3644:
3642:
3637:
3635:
3633:
3628:
3627:
3625:
3615:
3614:
3609:
3603:
3601:
3596:
3594:
3592:
3587:
3586:
3584:
3582:
3581:
3576:
3570:
3568:
3563:
3561:
3559:
3554:
3552:
3550:
3545:
3543:
3541:
3536:
3535:
3533:
3531:
3530:
3525:
3519:
3517:
3512:
3510:
3508:
3503:
3501:
3499:
3494:
3492:
3490:
3485:
3483:
3481:
3476:
3474:
3472:
3467:
3465:
3463:
3458:
3456:
3454:
3449:
3447:
3445:
3440:
3438:
3436:
3431:
3429:
3427:
3422:
3420:
3418:
3413:
3411:
3409:
3404:
3403:
3401:
3391:
3390:
3385:
3379:
3377:
3372:
3371:
3369:
3367:
3366:
3361:
3358:
3352:
3344:
3340:
3336:
3332:
3326:
3322:
3304:
3302:
3297:
3296:
3294:
3292:
3291:
3286:
3280:
3278:
3273:
3272:
3270:
3268:
3267:
3262:
3259:
3257:
3253:
3243:
3241:
3236:
3234:
3232:
3227:
3225:
3223:
3218:
3216:
3214:
3209:
3207:
3205:
3204:P. albertisii
3200:
3199:
3197:
3195:
3194:
3193:Pseudochirops
3189:
3183:
3181:
3176:
3174:
3172:
3167:
3165:
3163:
3158:
3156:
3154:
3149:
3147:
3145:
3140:
3138:
3136:
3131:
3129:
3127:
3122:
3120:
3118:
3113:
3112:
3110:
3108:
3107:
3102:
3096:
3094:
3093:P. peregrinus
3089:
3088:
3086:
3084:
3083:
3082:Pseudocheirus
3078:
3072:
3070:
3065:
3064:
3062:
3060:
3059:
3054:
3048:
3046:
3041:
3039:
3037:
3032:
3030:
3028:
3027:P. armillatus
3023:
3022:
3020:
3010:
3009:
3004:
2998:
2996:
2995:H. lemuroides
2991:
2990:
2988:
2986:
2985:
2980:
2977:
2975:
2971:
2961:
2959:
2954:
2952:
2950:
2945:
2943:
2941:
2936:
2934:
2932:
2929:Biak glider (
2927:
2925:
2923:
2918:
2916:
2914:
2909:
2908:
2906:
2904:
2903:
2898:
2892:
2890:
2885:
2884:
2882:
2880:
2879:
2878:Gymnobelideus
2874:
2868:
2866:
2865:D. trivirgata
2861:
2859:
2857:
2852:
2850:
2848:
2843:
2841:
2839:
2834:
2833:
2831:
2829:
2828:
2823:
2820:
2818:
2814:
2804:
2802:
2797:
2796:
2794:
2792:
2791:
2786:
2783:
2781:
2777:
2767:
2765:
2760:
2758:
2756:
2751:
2749:
2747:
2742:
2740:
2738:
2733:
2732:
2730:
2728:
2727:
2722:
2716:
2714:
2709:
2708:
2706:
2704:
2703:
2698:
2695:
2685:
2681:
2671:
2669:
2664:
2663:
2661:
2659:
2658:
2653:
2647:
2645:
2640:
2638:
2636:
2635:T. johnstonii
2631:
2629:
2627:
2622:
2620:
2618:
2613:
2611:
2609:
2604:
2603:
2601:
2591:
2590:
2585:
2579:
2577:
2576:S. pelegensis
2572:
2570:
2568:
2567:S. celebensis
2563:
2562:
2560:
2558:
2557:
2552:
2546:
2544:
2539:
2537:
2535:
2530:
2528:
2526:
2521:
2519:
2517:
2512:
2510:
2508:
2503:
2502:
2500:
2498:
2497:
2492:
2486:
2484:
2479:
2477:
2475:
2470:
2468:
2466:
2461:
2459:
2457:
2452:
2450:
2448:
2447:P. orientalis
2443:
2441:
2439:
2434:
2432:
2430:
2425:
2423:
2421:
2416:
2414:
2412:
2407:
2405:
2403:
2398:
2396:
2394:
2389:
2387:
2385:
2384:P. carmelitae
2380:
2378:
2376:
2375:P. alexandrae
2373:Gebe cuscus (
2371:
2370:
2368:
2366:
2365:
2360:
2354:
2352:
2347:
2345:
2343:
2338:
2337:
2335:
2325:
2324:
2319:
2316:
2310:
2302:
2301:Phalangeridae
2298:
2294:
2290:
2281:
2277:
2259:
2257:
2252:
2250:
2248:
2243:
2242:
2240:
2238:
2237:
2232:
2226:
2224:
2219:
2218:
2216:
2214:
2213:
2208:
2205:
2195:
2191:
2181:
2179:
2174:
2173:
2171:
2169:
2168:
2167:Phascolarctos
2163:
2160:
2158:
2154:
2150:
2146:
2145:Vombatiformes
2140:
2136:
2130:
2126:
2124:
2120:
2118:
2114:
2112:
2108:
2107:
2104:
2099:
2098:Diprotodontia
2091:
2086:
2084:
2079:
2077:
2072:
2071:
2068:
2044:
2038:
2036:
2030:
2028:
2022:
2020:
2014:
2012:
2006:
2005:
2003:
2001:
1997:
1991:
1987:Artiodactyla
1985:
1983:
1977:
1975:
1969:
1967:
1961:
1959:
1953:
1951:
1947:Eulipotyphla
1945:
1944:
1942:
1940:
1936:
1933:
1931:
1930:Boreoeutheria
1927:
1921:
1915:
1913:
1907:
1905:
1899:
1897:
1891:
1889:
1883:
1881:
1877:Afrosoricida
1875:
1874:
1872:
1870:
1866:
1860:
1854:
1852:
1846:
1845:
1843:
1841:
1837:
1834:
1828:
1820:
1816:
1806:
1800:
1798:
1792:
1790:
1784:
1782:
1776:
1774:
1768:
1767:
1765:
1763:
1759:
1753:
1747:
1745:
1739:
1738:
1736:
1734:
1730:
1727:
1721:
1713:
1709:
1706:
1704:
1700:
1690:
1684:
1683:
1681:
1679:
1675:
1672:
1670:
1666:
1660:
1656:
1654:
1650:
1648:
1644:
1642:
1638:
1637:
1634:
1629:
1621:
1616:
1614:
1609:
1607:
1602:
1601:
1598:
1580:
1576:
1572:
1568:
1564:
1560:
1556:
1549:
1541:
1537:
1533:
1529:
1525:
1521:
1517:
1510:
1501:
1496:
1492:
1488:
1484:
1480:
1473:
1464:
1459:
1455:
1451:
1447:
1443:
1436:
1427:
1422:
1418:
1414:
1407:
1400:
1392:
1388:
1383:
1378:
1373:
1368:
1364:
1360:
1356:
1349:
1343:
1338:
1332:
1327:
1312:
1308:
1304:
1300:
1296:
1292:
1285:
1278:
1270:
1266:
1262:
1260:0-801-88221-4
1256:
1252:
1251:
1246:
1245:Wilson, D. E.
1242:
1241:Groves, C. P.
1236:
1232:
1222:
1219:
1217:
1214:
1212:
1209:
1208:
1202:
1194:
1190:
1186:
1183:
1182:rat-kangaroos
1179:
1175:
1171:
1167:
1164:
1160:
1156:
1152:
1149:
1145:
1144:
1143:
1142:
1137:
1130:
1126:
1122:
1118:
1115:
1111:
1107:
1104:
1100:
1096:
1092:
1088:
1084:
1080:
1076:
1073:
1069:
1065:
1064:
1063:
1059:
1054:
1050:
1046:
1043:
1042:pygmy possums
1039:
1035:
1032:
1028:
1024:
1023:Phalangeridae
1020:
1019:
1018:
1014:
1013:
1012:
1011:
1006:
1002:
998:
995:
991:
987:
984:
980:
976:
973:
972:giant wombats
969:
965:
963:
959:
957:
953:
951:(one species)
950:
946:
942:
939:
935:
931:
930:
929:
928:
927:Vombatiformes
923:
922:
921:
920:
916:
914:
910:
906:
902:
898:
894:
893:Vombatiformes
890:
876:
875:
867:
866:
858:
857:
849:
848:
840:
839:
831:
830:
827:
826:
820:
819:
816:
815:
812:
811:
805:
804:
798:
797:
794:
793:
790:
789:
783:
782:
775:
774:
771:
770:
762:
761:
753:
752:
744:
743:
740:
739:
733:
732:
729:
728:
725:
724:
718:
717:
711:
710:
707:
706:
703:
702:
696:
695:
689:
688:
685:
684:
681:
680:
674:
673:
666:
665:
658:
655:
654:
651:
650:
642:
641:
638:
633:
632:
629:
628:
625:
620:
619:
612:
609:
608:
602:
599:
598:
595:
594:
586:
585:
582:
581:
574:
573:
570:
569:
566:
565:
558:
557:
551:
550:Vombatiformes
548:
547:
540:
539:
535:
534:
530:
529:
525:
515:
513:
512:Early Miocene
509:
508:
503:
499:
492:Fossil record
489:
486:
481:
476:
474:
470:
466:
460:
458:
453:
451:
447:
443:
438:
436:
432:
428:
424:
420:
416:
415:insectivorous
412:
408:
400:
396:
392:
387:
378:
376:
375:
370:
369:
364:
360:
356:
352:
348:
344:
340:
336:
332:
326:
272:
271:Diprotodontia
264:
263:
259:
255:
254:Vombatiformes
250:
245:
240:
235:
234:Diprotodontia
232:
229:
228:
225:
224:Eomarsupialia
222:
219:
216:
215:
212:
209:
206:
205:
202:
199:
196:
195:
192:
189:
186:
185:
182:
179:
176:
175:
172:
169:
166:
165:
160:
155:
151:
147:
143:
139:
135:
131:
127:
123:
119:
114:
110:
105:
100:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
33:
24:Diprotodontia
22:
19:
4686:Diprotodonts
4400:
4362:H. moschatus
4361:
4349:
4325:P. gilbertii
4324:
4315:
4306:
4286:
4274:
4265:
4256:
4247:
4227:
4216:A. rufescens
4215:
4203:
4178:
4166:
4154:
4145:
4136:
4127:
4118:
4109:
4100:
4089:(pademelons)
4080:
4068:
4056:
4044:
4035:
4026:
4017:
4009:P. lateralis
4008:
3999:
3990:
3981:
3973:P. coenensis
3972:
3964:P. assimilis
3963:
3955:
3954:
3946:P. xanthopus
3945:
3936:
3927:
3920:P. xanthopus
3919:
3918:
3909:
3900:
3892:P. burbidgei
3891:
3882:
3874:
3873:
3855:
3843:
3834:
3826:O. bernardus
3825:
3816:
3804:
3793:O. unguifera
3792:
3783:
3763:
3751:
3742:
3733:
3724:
3715:
3706:
3697:
3687:Notamacropus
3685:
3674:M. giganteus
3673:
3664:
3652:
3640:
3631:
3613:Lagorchestes
3611:
3600:D. vanheurni
3599:
3590:
3578:
3566:
3557:
3548:
3539:
3527:
3515:
3507:D. stellarum
3506:
3497:
3488:
3479:
3470:
3461:
3453:D. matschiei
3452:
3444:D. lumholtzi
3443:
3434:
3425:
3416:
3407:
3387:
3376:L. fasciatus
3375:
3365:Lagostrophus
3363:
3343:Macropodidae
3300:
3290:Distoechurus
3288:
3276:
3264:
3239:
3231:P. coronatus
3230:
3221:
3212:
3203:
3191:
3180:P. schlegeli
3179:
3170:
3161:
3152:
3143:
3134:
3125:
3117:P. canescens
3116:
3104:
3092:
3080:
3068:
3058:Petropseudes
3056:
3044:
3035:
3026:
3006:
2994:
2984:Hemibelideus
2982:
2957:
2948:
2940:P. breviceps
2939:
2931:P. biacensis
2930:
2922:P. australis
2921:
2912:
2900:
2888:
2876:
2864:
2855:
2846:
2837:
2827:Dactylopsila
2825:
2801:T. rostratus
2800:
2788:
2780:Tarsipedidae
2763:
2754:
2746:C. concinnus
2745:
2736:
2724:
2712:
2700:
2667:
2655:
2644:T. vulpecula
2643:
2634:
2625:
2616:
2607:
2587:
2575:
2566:
2556:Strigocuscus
2554:
2542:
2534:S. rufoniger
2533:
2525:S. papuensis
2524:
2516:S. maculatus
2515:
2506:
2494:
2482:
2473:
2464:
2455:
2446:
2437:
2428:
2419:
2410:
2401:
2392:
2383:
2374:
2362:
2350:
2342:A. melanotis
2341:
2321:
2255:
2247:L. latifrons
2246:
2234:
2222:
2210:
2177:
2165:
2097:
2026:(Treeshrews)
1909:Proboscidea
1850:(Armadillos)
1801:
1733:Ameridelphia
1686:Monotremata
1582:. Retrieved
1562:
1558:
1548:
1523:
1519:
1509:
1482:
1472:
1445:
1441:
1435:
1416:
1413:PLOS Biology
1412:
1399:
1362:
1358:
1348:
1337:
1326:
1314:. Retrieved
1294:
1290:
1277:
1249:
1235:
1200:
1155:Macropodidae
1138:
1095:sugar glider
1072:honey possum
1068:Tarsipedidae
1060:Superfamily
1015:Superfamily
1007:
1001:Wynyardiidae
924:
918:
917:
905:Phalangerida
886:
825:Macropodidae
823:
808:
786:
736:
721:
701:Tarsipedidae
699:
677:
667:Petauroidea
636:
624:Burramyidae
623:
601:Phalangerida
578:
562:
505:
495:
477:
461:
454:
439:
404:
398:
372:
366:
363:hippopotamus
270:
269:
252:
233:
217:
207:Infraclass:
145:
137:
129:
121:
18:
4567:iNaturalist
4425:Wikispecies
4307:P. longipes
4248:B. gaimardi
4205:Aepyprymnus
4045:P. sharmani
4000:P. inornata
3991:P. herberti
3910:P. wilkinsi
3901:P. concinna
3835:O. robustus
3784:O. fraenata
3765:Onychogalea
3707:N. dorsalis
3641:L. hirsutus
3591:D. macleayi
3580:Dorcopsulus
3567:D. muelleri
3558:D. luctuosa
3417:D. dorianus
3389:Dendrolagus
3301:D. pennatus
3277:A. pygmaeus
3256:Acrobatidae
3222:P. corinnae
3162:P. larvatus
3135:P. cinereus
3008:Petauroides
2949:P. gracilis
2847:D. palpator
2838:D. megalura
2737:C. caudatus
2726:Cercartetus
2684:Burramyidae
2589:Trichosurus
2507:S. kraemeri
2496:Spilocuscus
2483:P. vestitus
2474:P. sericeus
2420:P. matabiru
2411:P. lullulae
2393:P. gymnotis
2307:(including
2256:L. krefftii
2236:Lasiorhinus
2178:P. cinereus
2129:Marsupialia
2127:Infraclass
2032:Dermoptera
2024:Scandentia
2016:Lagomorpha
1965:(Pangolins)
1955:Chiroptera
1911:(Elephants)
1901:Hyracoidea
1657:(unranked)
1165:and allies)
1121:Acrobatidae
1116:and allies)
1062:Petauroidea
1038:Burramyidae
679:Acrobatidae
659: s.l.
613: s.s.
442:Australasia
427:fungivorous
211:Marsupialia
4680:Categories
4295:(potoroos)
4275:B. tropica
4257:B. lesueur
4236:(bettongs)
4195:Potoroidae
4179:W. bicolor
4137:T. lanatus
4128:T. calabyi
4018:P. mareeba
3982:P. godmani
3899:Nabarlek (
3806:Osphranter
3716:N. eugenii
3516:D. ursinus
3498:D. scottae
3471:D. notatus
3435:D. inustus
3349:(includes
3240:P. cupreus
3213:P. archeri
3144:P. forbesi
2817:Petauridae
2755:C. lepidus
2617:T. caninus
2543:S. wilsoni
2456:P. ornatus
2438:P. mimicus
2429:P. matanim
2351:A. ursinus
2223:V. ursinus
2194:Vombatidae
1971:Carnivora
1963:Pholidota
1895:(Aardvark)
1869:Afrotheria
1848:Cingulata
1829:inclusive)
1751:(Opossums)
1722:inclusive)
1712:Metatheria
1669:Yinotheria
1653:Vertebrata
1651:Subphylum
1584:24 October
1227:References
1170:Potoroidae
1148:Balbaridae
1079:Petauridae
934:Vombatidae
810:Potoroidae
723:Petauridae
580:Vombatidae
473:syndactyly
450:New Guinea
423:Potoroidae
419:omnivorous
407:herbivores
374:Thylacoleo
368:Diprotodon
339:marsupials
247:Suborders
30:28–0
4229:Bettongia
4155:T. thetis
4119:T. brunii
4110:T. browni
4082:Thylogale
3857:Petrogale
3743:N. parryi
3698:N. agilis
3549:D. hageni
3540:D. atrata
3529:Dorcopsis
3496:Tenkile (
3489:D. spadix
3462:D. mbaiso
3460:Dingiso (
3351:wallabies
3329:Suborder
3266:Acrobates
3171:P. mayeri
3126:P. caroli
3045:P. volans
2713:B. parvus
2364:Phalanger
2288:(possums)
2284:Suborder
2200:(wombats)
2143:Suborder
2040:Primates
2034:(Colugos)
2008:Rodentia
1903:(Hyraxes)
1840:Xenarthra
1827:Placental
1720:Marsupial
1540:0022-2372
1359:PLOS Biol
1163:wallabies
1159:kangaroos
1139:Suborder
1008:Suborder
962:Maradidae
956:Ilariidae
925:Suborder
909:suborders
889:suborders
510:from the
446:Oligocene
435:carnivory
347:wallabies
343:kangaroos
177:Kingdom:
171:Eukaryota
132:), young
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4465:BioLib:
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4168:Wallabia
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3844:O. rufus
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2913:P. abidi
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2790:Tarsipes
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2323:Ailurops
2309:cuscuses
2212:Vombatus
2123:Mammalia
2117:Chordata
2111:Animalia
2109:Kingdom
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1819:Eutheria
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954:Family †
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181:Animalia
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101:– Recent
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2115:Phylum
2100:species
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1021:Family
943:Family
938:wombats
932:Family
897:wombats
469:rodents
467:, like
465:wombats
411:extinct
365:-sized
359:wombats
351:possums
329:, from
230:Order:
197:Class:
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4598:552297
4520:1DIPRO
4416:Q26332
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1957:(Bats)
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1630:orders
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901:koala
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335:order
331:Greek
218:Clade
118:koala
4632:NCBI
4593:ITIS
4559:1452
4554:GBIF
4515:EPPO
4507:1694
4476:BOLD
1586:2015
1575:PMID
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