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Chlorophyll

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74: 1562: 1592:, are in cold polar waters or in places where ocean currents bring cold water to the surface, such as around the equator and along the shores of continents. It is not the cold water itself that stimulates the phytoplankton. Instead, the cool temperatures are often a sign that the water has welled up to the surface from deeper in the ocean, carrying nutrients that have built up over time. In polar waters, nutrients accumulate in surface waters during the dark winter months when plants cannot grow. When sunlight returns in the spring and summer, the plants flourish in high concentrations. 60: 92: 106: 311: 1149: 1191: 4585: 498: 1116: 1098: 1134: 1083: 1068: 1588:, are blue. Places where chlorophyll concentrations were high, meaning many phytoplankton were growing, are yellow. The observations come from the Moderate Resolution Imaging Spectroradiometer (MODIS) on NASA's Aqua satellite. Land is dark gray, and places where MODIS could not collect data because of sea ice, polar darkness, or clouds are light gray. The highest chlorophyll concentrations, where tiny surface-dwelling ocean plants are 436:
The function of the reaction center of chlorophyll is to absorb light energy and transfer it to other parts of the photosystem. The absorbed energy of the photon is transferred to an electron in a process called charge separation. The removal of the electron from the chlorophyll is an oxidation
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in Earth's atmosphere. Photosystem I typically works in series with Photosystem II; thus the P700 of Photosystem I is usually reduced as it accepts the electron, via many intermediates in the thylakoid membrane, by electrons coming, ultimately, from Photosystem II. Electron
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Reaction center chlorophyll–protein complexes are capable of directly absorbing light and performing charge separation events without the assistance of other chlorophyll pigments, but the probability of that happening under a given light intensity is small. Thus, the other chlorophylls in the
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Porra RJ, Thompson WA, Kriedemann PE (1989). "Determination of accurate extinction coefficients and simultaneous equations for assaying chlorophylls a and b extracted with four different solvents: verification of the concentration of chlorophyll standards by atomic absorption spectroscopy".
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as well as the red portion. Conversely, it is a poor absorber of green and near-green portions of the spectrum. Hence chlorophyll-containing tissues appear green because green light, diffusively reflected by structures like cell walls, is less absorbed. Two types of chlorophyll exist in the
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0.96 compared with chemical testing in the range from 41 mg m up to 675 mg m. Gitelson also developed a formula for direct readout of chlorophyll content in mg m. The formula provided a reliable method of measuring chlorophyll content from 41 mg m up to
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is reserved for a pigment that has been extracted from algae in 1966 but not chemically described. Besides the lettered chlorophylls, a wide variety of sidechain modifications to the chlorophyll structures are known in the wild. For example,
1982:"Das Hauptproduct der alkalischen Hydrolyse bilden tiefgrüne Alkalisalze. In ihnen liegen complexe Magnesiumverbindungen vor, die das Metall in einer gegen Alkali auch bei hoher Temperatur merkwürdig widerstandsfähigen Bindung enthalten." 1984:(Deep green alkali salts form the main product of alkali hydrolysis. In them, complex magnesium compounds are present, which contain the metal in a bond that is extraordinarily resistant to alkali even at high temperature.) 1918:"Nous n'avons aucun droit pour nommer une substance connue depuis long-temps, et à l'histoire de laquelle nous n'avons ajouté que quelques faits ; cependant nous proposerons, sans y mettre aucune importance, le nom de 441:. The charged reaction center of chlorophyll (P680) is then reduced back to its ground state by accepting an electron stripped from water. The electron that reduces P680 ultimately comes from the oxidation of water into O 433:) of their red-peak absorption maximum. The identity, function and spectral properties of the types of chlorophyll in each photosystem are distinct and determined by each other and the protein structure surrounding them. 1612:
gains its green color naturally from the chlorophyll introduced through the large variety of herbs used in its production. Chlorophyll is not soluble in water, and it is first mixed with a small quantity of
1482:(ALA), one of the intermediate compounds in the biosynthesis pathway. Plants that are fed by ALA accumulate high and toxic levels of protochlorophyllide; so do the mutants with a damaged regulatory system. 1328:
is an optical sensor used in plant science and agriculture for the assessment of chlorophyll contents in leaves. This device allows researchers to perform real-time and non-destructive measurements.
1787:"Influence of green, red and blue light emitting diodes on multiprotein complex proteins and photosynthetic activity under different light intensities in lettuce leaves (Lactuca sativa L.)" 1934:(We have no right to name a substance known for a long time, and to whose story we have added only a few facts ; however, we will propose, without giving it any importance, the name 1198:
Chlorophylls can be extracted from the protein into organic solvents. In this way, the concentration of chlorophyll within a leaf can be estimated. Methods also exist to separate
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This specific pair performs the final function of chlorophylls: Charge separation, which produces the unbound protons (H) and electrons (e) that separately propel biosynthesis.
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Muraki N, Nomata J, Ebata K, Mizoguchi T, Shiba T, Tamiaki H, Kurisu G, Fujita Y (May 2010). "X-ray crystal structure of the light-independent protochlorophyllide reductase".
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Plants are perceived as green because chlorophyll absorbs mainly the blue and red wavelengths but green light, reflected by plant structures like cell walls, is less absorbed.
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The chlorophyll maps show milligrams of chlorophyll per cubic meter of seawater each month. Places where chlorophyll amounts were very low, indicating very low numbers of
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Akhtar M (2007). "The Modification of Acetate and Propionate Side Chains During the Biosynthesis of Haem and Chlorophylls: Mechanistic and Stereochemical Studies".
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exceeds 10 M cm, which is among the highest for small-molecule organic compounds. In 90% acetone-water, the peak absorption wavelengths of chlorophyll
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Chen M (2019). "Chlorophylls d and f: Synthesis, occurrence, light-harvesting, and pigment organization in chlorophyll-binding protein complexes".
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group. This difference causes a considerable difference in the absorption spectrum, allowing plants to absorb a greater portion of visible light.
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Zolla L, Rinalducci S (December 2002). "Involvement of active oxygen species in degradation of light-harvesting proteins under light stresses".
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Taniguchi M, Lindsey JS (January 2017). "Synthetic Chlorins, Possible Surrogates for Chlorophylls, Prepared by Derivatization of Porphyrins".
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photosystem and antenna pigment proteins all cooperatively absorb and funnel light energy to the reaction center. Besides chlorophyll 
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The electron flow produced by the reaction center chlorophyll pigments is used to pump H ions across the thylakoid membrane, setting up a
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attached to the Mg center are often omitted for clarity. Appended to the chlorin ring are various side chains, usually including a long
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Woodward RB, Ayer WA, Beaton JM, Bickelhaupt F, Bonnett R, Buchschacher P, et al. (1990). "The total synthesis of chlorophyll
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Scheer H (2006). "An Overview of Chlorophylls and Bacteriochlorophylls: Biochemistry, Biophysics, Functions and Applications".
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Marker AF (1972). "The use of acetone and methanol in the estimation of chlorophyll in the presence of phaeophytin in plant".
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transfer reactions in the thylakoid membranes are complex, however, and the source of electrons used to reduce P700 can vary.
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by the presence of a fifth, ketone-containing ring beyond the four pyrrole-like rings. Most chlorophylls are classified as
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The function of the vast majority of chlorophyll (up to several hundred molecules per photosystem) is to absorb light.
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published a total synthesis of the molecule. In 1967, the last remaining stereochemical elucidation was completed by
2840:"Pre-loading of chlorophyll synthase with tetraprenyl diphosphate is an obligatory step in chlorophyll biosynthesis" 4861: 3372: 3811: 4292: 1470:, one of the biosynthetic intermediates, occurs mostly in the free form and, under light conditions, acts as a 1765: 1552:, completing the cycle. In later stages of senescence, chlorophyllides are converted to a group of colourless 3742: 3159: 2697:"New proximal sensors of vegetation: towards a non destructive quantitative estimation of plant constituents" 1873: 437:
reaction. The chlorophyll donates the high energy electron to a series of molecular intermediates called an
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and H through several intermediates. This reaction is how photosynthetic organisms such as plants produce O
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Chen M, Schliep M, Willows RD, Cai ZL, Neilan BA, Scheer H (September 2010). "A red-shifted chlorophyll".
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the phytyl sidechain to reverse the reaction in which chlorophylls are biosynthesised from chlorophyllide
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can provide a linear relationship of chlorophyll content when compared with chemical testing. The ratio
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in chlorophyll was discovered in 1906, and was the first detection of that element in living tissue.
3220:"Colorless tetrapyrrolic chlorophyll catabolites found in ripening fruit are effective antioxidants" 2294: 2024:
Woodward RB, Ayer WA, Beaton JM, Bickelhaupt F, Bonnett R, Buchschacher P, et al. (July 1960).
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Motilva MJ (2008). "Chlorophylls – from functionality in food to health relevance".
1608:. Chefs use chlorophyll to color a variety of foods and beverages green, such as pasta and spirits. 4565: 4410: 3460: 2252: 1675: 1670: 1499: 438: 195: 4607: 2506: 2025: 1681: 1310: 393:
Having done so, these same centers execute their second function: The transfer of that energy by
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Ratio fluorescence emission can be used to measure chlorophyll content. By exciting chlorophyll
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plants, the later steps in the biosynthetic pathway are light-dependent. Such plants are pale (
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Seen through a microscope, chlorophyll is concentrated within organisms in structures called
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has approximate maxima of 453 nm and 642 nm. The absorption peaks of chlorophyll
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These compounds have also been identified in ripening fruits and they give characteristic
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Gitelson AA, Buschmann C, Lichtenthaler HK (1999). "The Chlorophyll Fluorescence Ratio
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Meskauskiene R, Nater M, Goslings D, Kessler F, op den Camp R, Apel K (October 2001).
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fluorescence at a lower wavelength, the ratio of chlorophyll fluorescence emission at
1133: 1082: 1067: 525:). They share a common biosynthetic pathway with porphyrins, including the precursor 4548: 3397: 3392: 3336: 3299: 3249: 3200: 3141: 3136: 3093: 3066: 3023: 2952: 2909: 2862: 2820: 2785: 2775: 2744: 2626: 2573: 2550: 2502: 2415: 2401: 2366: 2358: 2321: 2233: 2221: 2134: 2004: 1818: 1634: 1569: 640: 486: 2964: 2921: 1556:
known as nonfluorescent chlorophyll catabolites (NCC's) with the general structure:
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has approximate absorbance maxima of 430 nm and 662 nm, while chlorophyll
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A 2002 study found that "leaves exposed to strong light contained degraded major
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Proceedings of the National Academy of Sciences of the United States of America
3094:"FLU: a negative regulator of chlorophyll biosynthesis in Arabidopsis thaliana" 2978:
Tsuzuki Y, Tsukatani Y, Yamakawa H, Itoh S, Fujita Y, Yamamoto H (March 2022).
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Willows RD (June 2003). "Biosynthesis of chlorophylls from protoporphyrin IX".
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Ciba Foundation Symposium 180 - the Biosynthesis of the Tetrapyrrole Pigments
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Several chlorophylls are known. All are defined as derivatives of the parent
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Bollivar DW (November 2006). "Recent advances in chlorophyll biosynthesis".
2890:"Recent advances in chlorophyll biosynthesis and breakdown in higher plants" 2317: 2217: 310: 4756: 4652: 4513: 4503: 4405: 4114: 3623: 3590: 3340: 3253: 3235: 3204: 3145: 3118: 3070: 3027: 3008: 2956: 2913: 2866: 2824: 2748: 2554: 2370: 2312:. Advances in Photosynthesis and Respiration. Vol. 25. pp. 1–26. 2225: 1962:[Contribution to the knowledge of the composition of chlorophyll]. 1822: 1715:"Chlorophyll does not reflect green light – how to correct a misconception" 1553: 1235: 764: 337:) in a solvent. The spectra of chlorophyll molecules are slightly modified 244: 4599: 2789: 1946:, leaf : this name would indicate the role that it plays in nature.) 105: 4815: 4794: 4746: 4705: 4329: 4314: 4309: 4245: 3887: 3858: 3757: 3752: 3600: 3440: 3435: 3420: 1932:, feuille : ce nom indiquerait le rôle qu'elle joue dans la nature." 1803: 1600:
Synthetic chlorophyll is registered as a food additive colorant, and its
1190: 884: 804: 553:). The most widely distributed form in terrestrial plants is chlorophyll 383: 375: 140: 80: 3062: 2858: 2046: 1916:
On p. 490, the authors propose a new name for chlorophyll. From p. 490:
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Chlorophyll is responsible for the green color of many plants and algae.
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Schmid HC, Rassadina V, Oster U, Schoch S, Rüdiger W (November 2002).
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Hörtensteiner S (2006). "Chlorophyll degradation during senescence".
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Battersby AR (December 2000). "Tetrapyrroles: the pigments of life".
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spanning from 1905 to 1915, the general structure of chlorophyll
132: 112: 3091: 2529:"Chlorophyll content monitoring in sugar maple (Acer saccharum)" 1712: 1494:
involves the degradation of chlorophyll: for example the enzyme
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This conversion forms an ester of the carboxylic acid group in
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is the enzyme that completes the biosynthesis of chlorophyll
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Chlorophyll forms deep green solutions in organic solvents.
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Hideous absinthe : a history of the devil in a bottle
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as an Accurate Measure of Chlorophyll Content in Plants".
1871:(September 1951). "Joseph Pelletier and Joseph Caventou". 489:, which occur in these pigment–protein antenna complexes. 461:
a chemiosmotic potential used mainly in the production of
386:. In these complexes, chlorophyll serves three functions: 111:
There are several types of chlorophyll, but all share the
4141: 3703: 3350: 2499:"Methods for analysis of benthic photosynthetic pigment" 2888:
Eckhardt U, Grimm B, Hörtensteiner S (September 2004).
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are 444 nm and 630 nm; peaks for chlorophyll
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are 442 nm and 630 nm; peaks for chlorophyll
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are 460 nm and 647 nm; peaks for chlorophyll
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are 430 nm and 664 nm; peaks for chlorophyll
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which have their own distinct reaction centres, named
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5th Pigments in Food congress- for quality and health
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and green algae have an additional light-independent
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into sugars as well as other biosynthetic reactions.
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were deduced based on NMR, optical and mass spectra.
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Pigments in vegetables: chlorophylls and carotenoids
1960:"Zur Kenntniss der Zusammensetzung des Chlorophylls" 576:
The structures of chlorophylls are summarized below:
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depending on specific pigment-protein interactions.
30:"Leaf green" redirects here. For the RAL color, see 3218:Müller T, Ulrich M, Ongania KH, Kräutler B (2007). 2596:
Biochimica et Biophysica Acta (BBA) - Bioenergetics
1713:Virtanen O, Constantinidou E, Tyystjärvi E (2020). 1637:". This appeared to the authors as support for the 1525:and the latter can be re-esterified to chlorophyll 1238:at 673 nm (maximum) and 726 nm. The peak 2620: 1906:[Notice on the green material in leaves]. 1386: 27:Green pigments found in plants, algae and bacteria 1901: 408:The two currently accepted photosystem units are 374:Chlorophyll molecules are arranged in and around 4848: 3298:. United Kingdom: I.B.Tauris, 2004. p. 22. 2340: 1785:Muneer S, Kim EJ, Park JS, Lee JH (March 2014). 1784: 1231:are at 465 nm and 665 nm. Chlorophyll 449:gas, and is the source for practically all the O 4714:1,7-Bis(4-hydroxyphenyl)-1,4,6-heptatrien-3-one 3318: 3178: 3176: 1846:University of California Museum of Paleontology 190:Chlorophylls absorb light most strongly in the 2455: 2168:"Infrared chlorophyll could boost solar cells" 1272:are 401 nm, 455 nm and 696 nm. 465:(stored chemical energy) or to reduce NADP to 183:("leaf"). Chlorophyll allows plants to absorb 83:– shown here grouped inside plant cells. 4615: 3366: 3182: 2621:Larkum AW, Douglas S, Raven JA, eds. (2003). 1692:of red-blue spectral affinity of chlorophyll. 4472:International Association for Plant Taxonomy 3173: 2980:"Effects of Light and Oxygen on Chlorophyll 1667:, a semi-synthetic derivative of chlorophyll 1661:, related compounds in phototrophic bacteria 1342:In some plants, chlorophyll is derived from 501:Space-filling model of the chlorophyll  367:, which allows plants to absorb energy from 274:and other oxygenic microorganisms that form 220:Chlorophyll was first isolated and named by 4629: 2526: 2449: 1957: 1791:International Journal of Molecular Sciences 1758: 121:which forms the right side of this diagram. 4622: 4608: 3373: 3359: 2422: 1904:"Notice sur la matière verte des feuilles" 557:. The only difference between chlorophyll 235:After initial work done by German chemist 3285: 3243: 3135: 3117: 3017: 3007: 2726: 2544: 1994: 1992: 1812: 1802: 1730: 1649:" in the short-term behaviour of plants. 4416:International Code of Nomenclature (ICN) 3197:10.1146/annurev.arplant.57.032905.105212 2984:Biosynthesis in a Marine Cyanobacterium 2934: 2755: 2034:Journal of the American Chemical Society 1867: 1189: 496: 309: 3312: 2971: 2802: 2694: 2191: 2189: 2066: 1998: 14: 4849: 2796: 2761: 2496: 2428: 2377: 2307: 2246: 1989: 1317:675 mg m with a correlation 397:to a specific chlorophyll pair in the 4603: 3354: 3291: 2567: 2456:Jeffrey SW, Shibata (February 1969). 2310:Chlorophylls and Bacteriochlorophylls 2271:"Unit 1.3. Photosynthetic cells" 1835: 1708: 1706: 1421:is made by the same enzyme acting on 1174:, a cyanobacterium, uses 8-vinyl Chl 492: 2527:Cate TM, Perkins TD (October 2003). 2383: 2186: 2147: 2026:"The total synthesis of chlorophyll" 1486:Senescence and the chlorophyll cycle 1428:. The same is known for chlorophyll 1346:and is synthesised along a branched 1521:can be converted to chlorophyllide 1387:{\displaystyle \rightleftharpoons } 485:, there are other pigments, called 247:in 1940. By 1960, when most of the 24: 2695:Cerovic, Zoran G. (October 2012). 2443:10.1111/j.1365-2427.1972.tb00377.x 1902:Pelletier PJ, Caventou JB (1817). 1703: 1186:Measurement of chlorophyll content 25: 4878: 4421:ICN for Cultivated Plants (ICNCP) 2003:(Print). University of Helsinki. 1624: 1459:and grow green even in darkness. 517:, which are reduced relatives of 317:spectra of free chlorophyll  305: 278:in 2010; a molecular formula of C 4584: 4583: 3268:"Chlorophyll : Global Maps" 2056:from the original on 2011-04-10. 1560: 1147: 1132: 1114: 1096: 1081: 1066: 131:is any of several related green 104: 90: 72: 58: 3260: 3211: 3152: 3085: 3034: 2928: 2881: 2831: 2720: 2688: 2639: 2614: 2586: 2561: 2520: 2490: 2334: 2301: 2263: 2240: 2160: 2141: 2108: 2060: 1745: 1719:Journal of Biological Education 1595: 1579: 1331: 270:was announced to be present in 151:. Its name is derived from the 52:Chlorophyll at different scales 3185:Annual Review of Plant Biology 2386:Advances in Botanical Research 2017: 1951: 1858: 1829: 1778: 1739: 1381: 199:photosystems of green plants: 13: 1: 2682:10.1016/S0034-4257(99)00023-1 2608:10.1016/S0005-2728(89)80347-0 1874:Journal of Chemical Education 1732:10.1080/00219266.2020.1858930 1696: 1541:can be directly reduced (via 569:group where the latter has a 4462:History of plant systematics 4049:Thorns, spines, and prickles 2150:"A New Form of Chlorophyll?" 2135:10.1016/0040-4020(90)80003-Z 1240:molar absorption coefficient 179: 165: 7: 2702:. Ebernburg. Archived from 2546:10.1093/treephys/23.15.1077 2501:. School of Life Sciences, 2355:10.1021/acs.chemrev.5b00696 1771:Online Etymology Dictionary 1652: 10: 4883: 4278:Alternation of generations 3380: 2497:Gilpin L (21 March 2001). 2398:10.1016/bs.abr.2019.03.006 2275:Essentials of Cell Biology 1335: 1062:Structures of chlorophylls 215: 171: 157: 29: 4803: 4775: 4737: 4704: 4661: 4638: 4579: 4527: 4491: 4428:Cultivated plant taxonomy 4391:Biological classification 4381: 4254: 4170: 4066: 4016: 3741: 3671: 3614: 3576: 3550: 3486: 3406: 3388: 3272:Earthobservatory.nasa.gov 2949:10.1007/s11120-006-9076-6 2906:10.1007/s11103-004-2331-3 2772:10.1002/9780470514535.ch8 2572:. Van Nostrand Reinhold. 1838:"Photosynthetic Pigments" 1678:, to measure plant stress 565:is that the former has a 395:resonance energy transfer 378:that are embedded in the 363:Chlorophyll is vital for 294:Mg and a structure of (2- 228:in 1817. The presence of 4288:Evolutionary development 2253:University of Cincinnati 1976:10.1002/jlac.19063500103 1676:Chlorophyll fluorescence 1671:Deep chlorophyll maximum 1537:. Moreover, chlorophyll 1462:Chlorophyll is bound to 1451:) if grown in darkness. 439:electron transport chain 222:Joseph Bienaimé Caventou 196:electromagnetic spectrum 4862:Photosynthetic pigments 3939:Hypanthium (Floral cup) 2937:Photosynthesis Research 2894:Plant Molecular Biology 2805:Natural Product Reports 2729:Natural Product Reports 2623:Photosynthesis in algae 2318:10.1007/1-4020-4516-6_1 2277:. Nature. July 5, 2013. 2218:10.1126/science.1191127 1682:Purple Earth hypothesis 1517:. Since chlorophyllide 469:. NADPH is a universal 355: Chlorophyll  346: Chlorophyll  226:Pierre Joseph Pelletier 4554:by author abbreviation 4478:Plant taxonomy systems 4396:Botanical nomenclature 3236:10.1002/anie.200703587 3119:10.1073/pnas.221252798 3009:10.3390/plants11070915 2148:Jabr F (August 2010). 1958:Willstätter R (1906). 1842:UCMP Glossary (online) 1617:to obtain the desired 1548:) back to chlorophyll 1388: 1195: 506: 360: 4561:Botanical expeditions 1752:University of Bristol 1686:scientific hypothesis 1543:7-hydroxychlorophyll 1389: 1374:+ phytyl diphosphate 1311:correlation value of 1193: 500: 313: 257:Robert Burns Woodward 4719:Bisdemethoxycurcumin 4293:Evolutionary history 4283:Double fertilization 4135:Cellular respiration 2847:Biological Chemistry 2662:Remote Sens. Environ 1908:Journal de Pharmacie 1804:10.3390/ijms15034657 1645:species play a role 1378: 1360:Chlorophyll synthase 1350:that is shared with 1348:biosynthetic pathway 527:uroporphyrinogen III 401:of the photosystems. 3512:Non-vascular plants 3160:"Chlorophyll Cycle" 3110:2001PNAS...9812826M 3063:10.1038/nature08950 3055:2010Natur.465..110M 2859:10.1515/BC.2002.198 2674:1999RSEnv..69..296G 2463:Biological Bulletin 2210:2010Sci...329.1318C 2154:Scientific American 2086:1967Natur.216..151F 2047:10.1021/ja01499a093 1887:1951JChEd..28..454D 1659:Bacteriochlorophyll 1480:aminolevulinic acid 1468:Protochlorophyllide 1453:Non-vascular plants 1409:with the 20-carbon 459:proton-motive force 237:Richard Willstätter 169:("pale green") and 32:RAL 6002 Leaf green 4867:E-number additives 4826:Phycoerythrocyanin 4724:Desmethoxycurcumin 4017:Surface structures 3812:Flower development 2625:. London: Kluwer. 2431:Freshwater Biology 2247:Carter JS (1996). 1964:Annalen der Chemie 1688:that explains the 1384: 1196: 507: 493:Chemical structure 487:accessory pigments 361: 243:was elucidated by 4844: 4843: 4767:Condensed tannins 4597: 4596: 4236:Herbaceous plants 4062: 4061: 3333:10.1021/bi0265776 3327:(48): 14391–402. 3224:Angewandte Chemie 2632:978-0-7923-6333-0 2509:on April 14, 2008 2503:Napier University 2327:978-1-4020-4515-8 2129:(22): 7599–7659. 2080:(5111): 151–152. 2041:(14): 3800–3802. 2010:978-952-10-4846-3 1895:10.1021/ed028p454 1836:Speer BR (1997). 1635:isolated proteins 1058: 1057: 641:Molecular formula 473:used to reduce CO 43:Pokémon LeafGreen 16:(Redirected from 4874: 4624: 4617: 4610: 4601: 4600: 4587: 4586: 4566:Individual trees 4241:Secondary growth 4212:Succulent plants 4200:Prostrate shrubs 4083:Apical dominance 4068:Plant physiology 4029:Epicuticular wax 3574: 3573: 3567: 3558:Plant morphology 3375: 3368: 3361: 3352: 3351: 3345: 3344: 3316: 3310: 3309: 3292:Adams J (2004). 3289: 3283: 3282: 3280: 3278: 3264: 3258: 3257: 3247: 3230:(45): 8699–702. 3215: 3209: 3208: 3180: 3171: 3170: 3168: 3167: 3156: 3150: 3149: 3139: 3121: 3104:(22): 12826–31. 3089: 3083: 3082: 3038: 3032: 3031: 3021: 3011: 2975: 2969: 2968: 2932: 2926: 2925: 2885: 2879: 2878: 2844: 2835: 2829: 2828: 2817:10.1039/B110549N 2800: 2794: 2793: 2759: 2753: 2752: 2741:10.1039/B002635M 2724: 2718: 2717: 2715: 2714: 2708: 2701: 2692: 2686: 2685: 2643: 2637: 2636: 2618: 2612: 2611: 2590: 2584: 2583: 2568:Gross J (1991). 2565: 2559: 2558: 2548: 2524: 2518: 2517: 2515: 2514: 2505:. Archived from 2494: 2488: 2487: 2453: 2447: 2446: 2426: 2420: 2419: 2381: 2375: 2374: 2343:Chemical Reviews 2338: 2332: 2331: 2305: 2299: 2298: 2292: 2288: 2286: 2278: 2267: 2261: 2260: 2255:. Archived from 2249:"Photosynthesis" 2244: 2238: 2237: 2204:(5997): 1318–9. 2193: 2184: 2183: 2181: 2179: 2174:. 19 August 2010 2164: 2158: 2157: 2145: 2139: 2138: 2112: 2106: 2105: 2094:10.1038/216151a0 2064: 2058: 2057: 2055: 2030: 2021: 2015: 2014: 1996: 1987: 1986: 1955: 1949: 1945: 1930: 1915: 1898: 1862: 1856: 1855: 1853: 1852: 1833: 1827: 1826: 1816: 1806: 1782: 1776: 1775: 1762: 1756: 1755: 1743: 1737: 1736: 1734: 1710: 1631:antenna proteins 1564: 1547: 1492:plant senescence 1393: 1391: 1390: 1385: 1294: 1292: 1286: 1284: 1151: 1136: 1118: 1100: 1085: 1070: 579: 578: 561:and chlorophyll 552: 420: 413: 354: 345: 335: 325: 182: 176: 175: 174: 168: 162: 161: 160: 108: 94: 76: 62: 40:video game, see 21: 4882: 4881: 4877: 4876: 4875: 4873: 4872: 4871: 4847: 4846: 4845: 4840: 4811:Allophycocyanin 4799: 4771: 4733: 4700: 4657: 4634: 4628: 4598: 4593: 4575: 4544:Botanical terms 4537: 4523: 4487: 4433:Citrus taxonomy 4411:Author citation 4377: 4271: 4250: 4172: 4166: 4162:Turgor pressure 4070: 4058: 4012: 3827:Floral symmetry 3745: 3737: 3667: 3656:Vascular bundle 3651:Vascular tissue 3610: 3570: 3561: 3560: 3546: 3517:Vascular plants 3482: 3478:Plant pathology 3402: 3384: 3379: 3349: 3348: 3317: 3313: 3306: 3290: 3286: 3276: 3274: 3266: 3265: 3261: 3216: 3212: 3181: 3174: 3165: 3163: 3158: 3157: 3153: 3090: 3086: 3049:(7294): 110–4. 3039: 3035: 2976: 2972: 2933: 2929: 2886: 2882: 2853:(11): 1769–78. 2842: 2836: 2832: 2801: 2797: 2782: 2760: 2756: 2725: 2721: 2712: 2710: 2706: 2699: 2693: 2689: 2659: 2652: 2644: 2640: 2633: 2619: 2615: 2591: 2587: 2580: 2566: 2562: 2533:Tree Physiology 2525: 2521: 2512: 2510: 2495: 2491: 2476:10.2307/1539668 2454: 2450: 2427: 2423: 2408: 2382: 2378: 2339: 2335: 2328: 2306: 2302: 2290: 2289: 2280: 2279: 2269: 2268: 2264: 2245: 2241: 2194: 2187: 2177: 2175: 2166: 2165: 2161: 2146: 2142: 2113: 2109: 2065: 2061: 2053: 2028: 2022: 2018: 2011: 1997: 1990: 1956: 1952: 1943: 1928: 1863: 1859: 1850: 1848: 1834: 1830: 1783: 1779: 1764: 1763: 1759: 1744: 1740: 1711: 1704: 1699: 1655: 1627: 1598: 1582: 1542: 1490:The process of 1488: 1472:photosensitizer 1423:chlorophyllide 1404:chlorophyllide 1379: 1376: 1375: 1370:chlorophyllide 1340: 1334: 1321:value of 0.95. 1308: 1301: 1290: 1288: 1282: 1280: 1266: 1259: 1242:of chlorophyll 1188: 1171:Prochlorococcus 1161: 1158: 1152: 1143: 1137: 1128: 1126: 1119: 1110: 1108: 1101: 1092: 1086: 1077: 1071: 1030: 1025: 1020: 1015: 1010: 1005: 995: 991: 984: 980: 965: 961: 954: 950: 939: 935: 929: 925: 919: 913: 909: 903: 899: 892: 888: 876: 870: 864: 858: 849: 838: 827: 821: 815: 808: 793: 787: 781: 775: 768: 755: 751: 747: 743: 736: 732: 728: 724: 717: 713: 709: 705: 698: 694: 690: 686: 679: 675: 671: 667: 660: 656: 652: 648: 617: 606: 550: 546: 542: 495: 476: 452: 448: 444: 415: 409: 399:reaction center 359: 352: 350: 343: 333: 323: 308: 293: 289: 285: 281: 251:of chlorophyll 249:stereochemistry 218: 172: 158: 126: 125: 124: 123: 122: 109: 100: 99: 98: 95: 86: 85: 84: 77: 68: 67: 66: 63: 54: 53: 47: 34:. For the 2004 28: 23: 22: 15: 12: 11: 5: 4880: 4870: 4869: 4864: 4859: 4842: 4841: 4839: 4838: 4833: 4828: 4823: 4818: 4813: 4807: 4805: 4801: 4800: 4798: 4797: 4792: 4787: 4781: 4779: 4773: 4772: 4770: 4769: 4764: 4759: 4754: 4752:Anthocyanidins 4749: 4743: 4741: 4735: 4734: 4732: 4731: 4726: 4721: 4716: 4710: 4708: 4702: 4701: 4699: 4698: 4693: 4688: 4683: 4678: 4673: 4667: 4665: 4659: 4658: 4656: 4655: 4650: 4644: 4642: 4636: 4635: 4632:plant pigments 4627: 4626: 4619: 4612: 4604: 4595: 4594: 4592: 4591: 4580: 4577: 4576: 4574: 4573: 4568: 4563: 4558: 4557: 4556: 4546: 4540: 4538: 4536: 4535: 4534:Related topics 4532: 4528: 4525: 4524: 4522: 4521: 4516: 4511: 4506: 4501: 4495: 4493: 4489: 4488: 4486: 4485: 4483:Taxonomic rank 4480: 4475: 4469: 4464: 4459: 4458: 4457: 4456: 4455: 4450: 4445: 4435: 4425: 4424: 4423: 4418: 4413: 4408: 4403: 4401:Botanical name 4393: 4387: 4385: 4383:Plant taxonomy 4379: 4378: 4376: 4375: 4374: 4373: 4368: 4367: 4366: 4359:Megasporangium 4356: 4355: 4354: 4347:Microsporangia 4339: 4338: 4337: 4332: 4327: 4322: 4312: 4307: 4302: 4301: 4300: 4290: 4285: 4280: 4274: 4272: 4270: 4269: 4264: 4258: 4252: 4251: 4249: 4248: 4243: 4238: 4233: 4232: 4231: 4230: 4229: 4219: 4214: 4209: 4208: 4207: 4202: 4192: 4187: 4185:Cushion plants 4176: 4174: 4168: 4167: 4165: 4164: 4159: 4154: 4149: 4144: 4139: 4138: 4137: 4132: 4122: 4120:Plant hormones 4117: 4112: 4111: 4110: 4103:Photosynthesis 4100: 4095: 4090: 4085: 4080: 4074: 4072: 4064: 4063: 4060: 4059: 4057: 4056: 4051: 4046: 4041: 4036: 4031: 4026: 4020: 4018: 4014: 4013: 4011: 4010: 4005: 4000: 3995: 3990: 3989: 3988: 3983: 3978: 3968: 3967: 3966: 3961: 3956: 3951: 3941: 3936: 3935: 3934: 3933: 3932: 3927: 3922: 3921: 3920: 3915: 3895: 3890: 3885: 3884: 3883: 3882: 3881: 3876: 3866: 3861: 3856: 3851: 3846: 3836: 3835: 3834: 3829: 3824: 3822:Floral formula 3819: 3817:Floral diagram 3814: 3809: 3799: 3798: 3797: 3792: 3787: 3786: 3785: 3780: 3770: 3760: 3755: 3749: 3747: 3746:(incl. Flower) 3739: 3738: 3736: 3735: 3734: 3733: 3728: 3723: 3722: 3721: 3716: 3706: 3696: 3691: 3686: 3681: 3675: 3673: 3669: 3668: 3666: 3665: 3660: 3659: 3658: 3648: 3646:Storage organs 3643: 3638: 3637: 3636: 3626: 3620: 3618: 3612: 3611: 3609: 3608: 3603: 3598: 3593: 3588: 3582: 3580: 3571: 3569: 3568: 3554: 3548: 3547: 3545: 3544: 3539: 3534: 3532:Spermatophytes 3529: 3524: 3519: 3514: 3509: 3504: 3502:Archaeplastida 3499: 3493: 3491: 3484: 3483: 3481: 3480: 3475: 3470: 3465: 3464: 3463: 3456:Phytogeography 3453: 3451:Phytochemistry 3448: 3443: 3438: 3433: 3428: 3423: 3418: 3412: 3410: 3408:Subdisciplines 3404: 3403: 3401: 3400: 3395: 3389: 3386: 3385: 3378: 3377: 3370: 3363: 3355: 3347: 3346: 3311: 3305:978-1860649202 3304: 3284: 3259: 3210: 3172: 3151: 3084: 3033: 2970: 2927: 2880: 2830: 2795: 2780: 2754: 2719: 2687: 2668:(3): 296–302. 2657: 2650: 2638: 2631: 2613: 2602:(3): 384–394. 2585: 2579:978-0442006570 2578: 2560: 2539:(15): 1077–9. 2519: 2489: 2448: 2437:(4): 361–385. 2421: 2406: 2376: 2349:(2): 344–535. 2333: 2326: 2300: 2291:|website= 2262: 2259:on 2013-06-29. 2239: 2185: 2159: 2140: 2107: 2059: 2016: 2009: 1988: 1970:(1–2): 48–82. 1950: 1948: 1947: 1926:, couleur, et 1899: 1857: 1828: 1797:(3): 4657–70. 1777: 1757: 1738: 1701: 1700: 1698: 1695: 1694: 1693: 1679: 1673: 1668: 1662: 1654: 1651: 1626: 1625:Biological use 1623: 1597: 1594: 1581: 1578: 1570:autumn colours 1566: 1565: 1496:chlorophyllase 1487: 1484: 1417:. Chlorophyll 1400: 1399: 1383: 1338:Chlorophyllide 1336:Main article: 1333: 1330: 1306: 1299: 1264: 1257: 1219:, chlorophyll 1187: 1184: 1160: 1159: 1153: 1146: 1144: 1138: 1131: 1129: 1124: 1120: 1113: 1111: 1106: 1102: 1095: 1093: 1087: 1080: 1078: 1072: 1065: 1063: 1059: 1056: 1055: 1054:Cyanobacteria 1052: 1051:Cyanobacteria 1049: 1048:Various algae 1046: 1045:Various algae 1043: 1042:Mostly plants 1040: 1037: 1033: 1032: 1027: 1022: 1017: 1012: 1007: 1002: 998: 997: 993: 989: 986: 982: 978: 975: 972: 967: 963: 959: 956: 952: 948: 945: 941: 940: 937: 933: 930: 927: 923: 920: 917: 914: 911: 907: 904: 901: 897: 894: 890: 886: 882: 878: 877: 874: 871: 868: 865: 862: 859: 856: 853: 850: 847: 844: 840: 839: 836: 833: 828: 825: 822: 819: 816: 813: 810: 806: 802: 798: 797: 794: 791: 788: 785: 782: 779: 776: 773: 770: 766: 762: 758: 757: 753: 749: 745: 741: 738: 734: 730: 726: 722: 719: 715: 711: 707: 703: 700: 696: 692: 688: 684: 681: 677: 673: 669: 665: 662: 658: 654: 650: 646: 643: 637: 636: 628: 620: 615: 609: 604: 598: 590: 582: 548: 544: 494: 491: 474: 450: 446: 442: 417:photosystem II 406: 405: 402: 391: 365:photosynthesis 351: 342: 307: 306:Photosynthesis 304: 298:)-chlorophyll 291: 287: 283: 279: 217: 214: 110: 103: 102: 101: 96: 89: 88: 87: 78: 71: 70: 69: 64: 57: 56: 55: 51: 50: 49: 48: 26: 9: 6: 4: 3: 2: 4879: 4868: 4865: 4863: 4860: 4858: 4857:Tetrapyrroles 4855: 4854: 4852: 4837: 4834: 4832: 4829: 4827: 4824: 4822: 4821:Phycoerythrin 4819: 4817: 4814: 4812: 4809: 4808: 4806: 4802: 4796: 4793: 4791: 4788: 4786: 4783: 4782: 4780: 4778: 4774: 4768: 4765: 4763: 4760: 4758: 4757:Anthoxanthins 4755: 4753: 4750: 4748: 4745: 4744: 4742: 4740: 4736: 4730: 4727: 4725: 4722: 4720: 4717: 4715: 4712: 4711: 4709: 4707: 4703: 4697: 4694: 4692: 4691:Chlorophyll f 4689: 4687: 4686:Chlorophyll d 4684: 4682: 4681:Chlorophyll c 4679: 4677: 4676:Chlorophyll b 4674: 4672: 4671:Chlorophyll a 4669: 4668: 4666: 4664: 4660: 4654: 4651: 4649: 4646: 4645: 4643: 4641: 4637: 4633: 4625: 4620: 4618: 4613: 4611: 4606: 4605: 4602: 4590: 4582: 4581: 4578: 4572: 4569: 4567: 4564: 4562: 4559: 4555: 4552: 4551: 4550: 4547: 4545: 4542: 4541: 4539: 4533: 4530: 4529: 4526: 4520: 4519:Phytochemical 4517: 4515: 4512: 4510: 4507: 4505: 4502: 4500: 4497: 4496: 4494: 4490: 4484: 4481: 4479: 4476: 4473: 4470: 4468: 4465: 4463: 4460: 4454: 4451: 4449: 4446: 4444: 4441: 4440: 4439: 4436: 4434: 4431: 4430: 4429: 4426: 4422: 4419: 4417: 4414: 4412: 4409: 4407: 4404: 4402: 4399: 4398: 4397: 4394: 4392: 4389: 4388: 4386: 4384: 4380: 4372: 4369: 4365: 4362: 4361: 4360: 4357: 4353: 4350: 4349: 4348: 4345: 4344: 4343: 4340: 4336: 4333: 4331: 4328: 4326: 4323: 4321: 4318: 4317: 4316: 4313: 4311: 4308: 4306: 4303: 4299: 4296: 4295: 4294: 4291: 4289: 4286: 4284: 4281: 4279: 4276: 4275: 4273: 4268: 4265: 4263: 4260: 4259: 4257: 4253: 4247: 4244: 4242: 4239: 4237: 4234: 4228: 4225: 4224: 4223: 4220: 4218: 4215: 4213: 4210: 4206: 4203: 4201: 4198: 4197: 4196: 4193: 4191: 4188: 4186: 4183: 4182: 4181: 4178: 4177: 4175: 4169: 4163: 4160: 4158: 4157:Transpiration 4155: 4153: 4150: 4148: 4145: 4143: 4140: 4136: 4133: 4131: 4128: 4127: 4126: 4123: 4121: 4118: 4116: 4113: 4109: 4106: 4105: 4104: 4101: 4099: 4096: 4094: 4091: 4089: 4086: 4084: 4081: 4079: 4076: 4075: 4073: 4069: 4065: 4055: 4052: 4050: 4047: 4045: 4042: 4040: 4037: 4035: 4032: 4030: 4027: 4025: 4022: 4021: 4019: 4015: 4009: 4006: 4004: 4001: 3999: 3996: 3994: 3991: 3987: 3984: 3982: 3979: 3977: 3974: 3973: 3972: 3969: 3965: 3962: 3960: 3957: 3955: 3952: 3950: 3947: 3946: 3945: 3944:Inflorescence 3942: 3940: 3937: 3931: 3928: 3926: 3923: 3919: 3916: 3914: 3911: 3910: 3909: 3906: 3905: 3904: 3901: 3900: 3899: 3896: 3894: 3891: 3889: 3886: 3880: 3877: 3875: 3872: 3871: 3870: 3867: 3865: 3862: 3860: 3857: 3855: 3852: 3850: 3847: 3845: 3842: 3841: 3840: 3837: 3833: 3830: 3828: 3825: 3823: 3820: 3818: 3815: 3813: 3810: 3808: 3805: 3804: 3803: 3800: 3796: 3793: 3791: 3788: 3784: 3781: 3779: 3776: 3775: 3774: 3771: 3769: 3766: 3765: 3764: 3761: 3759: 3756: 3754: 3751: 3750: 3748: 3744: 3740: 3732: 3729: 3727: 3724: 3720: 3717: 3715: 3712: 3711: 3710: 3707: 3705: 3702: 3701: 3700: 3697: 3695: 3692: 3690: 3687: 3685: 3682: 3680: 3677: 3676: 3674: 3670: 3664: 3661: 3657: 3654: 3653: 3652: 3649: 3647: 3644: 3642: 3639: 3635: 3632: 3631: 3630: 3629:Ground tissue 3627: 3625: 3622: 3621: 3619: 3617: 3613: 3607: 3604: 3602: 3599: 3597: 3594: 3592: 3589: 3587: 3584: 3583: 3581: 3579: 3575: 3572: 3565: 3559: 3556: 3555: 3553: 3552:Plant anatomy 3549: 3543: 3540: 3538: 3535: 3533: 3530: 3528: 3525: 3523: 3520: 3518: 3515: 3513: 3510: 3508: 3505: 3503: 3500: 3498: 3495: 3494: 3492: 3489: 3485: 3479: 3476: 3474: 3473:Plant ecology 3471: 3469: 3468:Plant anatomy 3466: 3462: 3459: 3458: 3457: 3454: 3452: 3449: 3447: 3444: 3442: 3439: 3437: 3434: 3432: 3429: 3427: 3424: 3422: 3419: 3417: 3416:Archaeobotany 3414: 3413: 3411: 3409: 3405: 3399: 3396: 3394: 3391: 3390: 3387: 3383: 3376: 3371: 3369: 3364: 3362: 3357: 3356: 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The axial 508: 502: 482: 479: 456: 435: 407: 384:chloroplasts 376:photosystems 373: 362: 356: 347: 338: 332: 328: 322: 318: 299: 266: 265:Chlorophyll 252: 245:Hans Fischer 240: 234: 219: 208: 202: 201:chlorophyll 192:blue portion 189: 187:from light. 141:chloroplasts 128: 127: 81:chloroplasts 42: 35: 18:Chlorophylls 4836:Xanthonoids 4816:Phycocyanin 4777:Carotenoids 4663:Chlorophyll 4648:Betacyanins 4330:Pollen tube 4325:Pollinators 4315:Pollination 4310:Germination 4125:Respiration 4108:Chlorophyll 3954:Pedicellate 3888:Gametophyte 3807:Aestivation 3758:Antheridium 3753:Archegonium 3601:Plasmodesma 3578:Plant cells 3441:Paleobotany 3436:Ethnobotany 3421:Astrobotany 2900:(1): 1–14. 2392:: 121–139. 2122:Tetrahedron 1936:chlorophyll 1920:chlorophyle 1309:provided a 1036:Occurrence 996:COO−Phytyl 985:COO−Phytyl 974:−CH=CHCOOH 966:COO−Phytyl 955:COO−Phytyl 261:Ian Fleming 255:was known, 139:and in the 129:Chlorophyll 4851:Categories 4739:Flavonoids 4352:Microspore 4342:Sporangium 4320:Artificial 4008:Sporophyte 4003:Sporophyll 3998:Receptacle 3893:Gynandrium 3763:Androecium 3672:Vegetative 3542:Angiosperm 3537:Gymnosperm 3431:Dendrology 3277:2 February 3166:2020-06-04 3002:(7): 915. 2713:2024-07-12 2513:2010-07-17 1914:: 486–491. 1881:(9): 454. 1869:Delépine M 1851:2010-07-17 1725:(5): 1–8. 1697:References 1639:hypothesis 1507:hydrolyses 1474:, forming 1445:Angiosperm 1293:10 nm 1285:10 nm 1236:fluoresces 1039:Universal 1031:(chlorin) 1026:(chlorin) 1011:(chlorin) 1006:(chlorin) 970:−CH=CHCOOH 944:C17 group 523:hemoglobin 521:(found in 519:porphyrins 431:nanometers 315:Absorbance 4790:Retinoids 4785:Carotenes 4640:Betalains 4630:Types of 4549:Botanists 4467:Herbarium 4364:Megaspore 4262:Evolution 4205:Subshrubs 4173:and habit 4098:Nutrition 4093:Cellulose 4088:Bulk flow 4071:Materials 4034:Epidermis 3898:Gynoecium 3879:Endosperm 3874:Dispersal 3790:Staminode 3726:Sessility 3714:Cataphyll 3634:Mesophyll 3586:Cell wall 3527:Lycophyte 3507:Bryophyte 3461:Geobotany 3446:Phycology 3191:: 55–77. 2416:149632511 2293:ignored ( 2283:cite book 2234:206527174 2068:Fleming I 1690:evolution 1574:deciduous 1449:etiolated 1411:diterpene 1382:⇌ 1344:glutamate 881:C8 group 843:C7 group 801:C3 group 761:C2 group 531:magnesium 380:thylakoid 230:magnesium 135:found in 116:magnesium 4831:Quinones 4729:Curcumin 4589:Category 4509:Forestry 4499:Agronomy 4492:Practice 4443:Cultivar 4438:Cultigen 4298:timeline 4190:Rosettes 4078:Aleurone 4054:Trichome 3971:Perianth 3783:Filament 3641:Meristem 3564:glossary 3426:Bryology 3341:12450406 3254:17943948 3205:16669755 3146:11606728 3071:20400946 3028:35406896 2965:23808539 2957:17370354 2922:21174896 2914:15604725 2867:12530542 2825:12828371 2749:11152419 2555:12975132 2371:27498781 2226:20724585 2178:15 April 2051:Archived 1823:24642884 1653:See also 1619:solution 1610:Absinthe 1602:E number 1590:thriving 1576:plants. 1503:3.1.1.14 1464:proteins 1413:alcohol 1356:siroheme 515:chlorins 505:molecule 133:pigments 4762:Flavans 4696:Chlorin 4267:Ecology 4024:Cuticle 3854:Capsule 3844:Anatomy 3795:Tapetum 3719:Petiole 3694:Rhizome 3689:Rhizoid 3616:Tissues 3606:Vacuole 3596:Plastid 3398:Outline 3393:History 3245:2912502 3128:3056990 3106:Bibcode 3079:4427639 3051:Bibcode 3019:9003380 2875:3099209 2790:7842850 2670:Bibcode 2484:1539668 2363:1534468 2206:Bibcode 2198:Science 2102:4262313 2082:Bibcode 1940:chloros 1938:, from 1924:chloros 1883:Bibcode 1814:3975419 1746:May P. 1647:in vivo 541:chain ( 535:ligands 511:chlorin 339:in vivo 216:History 194:of the 180:phyllon 166:khloros 113:chlorin 37:Pokémon 4571:Plants 4474:(IAPT) 4227:Lianas 4195:Shrubs 4147:Starch 4039:Nectar 3959:Raceme 3925:Stigma 3913:Locule 3903:Carpel 3864:Pyrena 3802:Flower 3778:Anther 3773:Stamen 3768:Pollen 3490:groups 3382:Botany 3339:  3302:  3252:  3242:  3203:  3144:  3134:  3126:  3077:  3069:  3043:Nature 3026:  3016:  2996:Plants 2990:marina 2988:  2963:  2955:  2920:  2912:  2873:  2865:  2823:  2788:  2778:  2747:  2629:  2576:  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977:−CH 958:−CH 947:−CH 932:−CH 922:−CH 906:−CH 896:−CH 885:−CH 873:−CH 867:−CH 861:−CH 855:−CH 846:−CH 790:−CH 784:−CH 778:−CH 772:−CH 765:−CH 756:Mg 737:Mg 718:Mg 699:Mg 680:Mg 661:Mg 463:ATP 334:red 143:of 4853:: 3335:. 3325:41 3323:. 3270:. 3248:. 3238:. 3228:46 3226:. 3222:. 3199:. 3189:57 3187:. 3175:^ 3140:. 3130:. 3122:. 3112:. 3102:98 3100:. 3096:. 3073:. 3065:. 3057:. 3045:. 3022:. 3012:. 3000:11 2998:. 2994:. 2959:. 2951:. 2941:90 2939:. 2916:. 2908:. 2898:56 2896:. 2892:. 2869:. 2861:. 2849:. 2845:. 2819:. 2809:20 2807:. 2784:. 2774:. 2743:. 2733:17 2731:. 2676:. 2666:69 2664:. 2598:. 2549:. 2537:23 2535:. 2531:. 2478:. 2466:. 2460:. 2433:. 2410:. 2400:. 2390:90 2388:. 2365:. 2357:. 2345:. 2320:. 2287:: 2285:}} 2281:{{ 2273:. 2251:. 2228:. 2220:. 2212:. 2200:. 2188:^ 2170:. 2152:. 2127:46 2125:. 2096:. 2088:. 2076:. 2049:. 2039:82 2037:. 2031:. 1991:^ 1978:. 1889:. 1879:28 1877:. 1844:. 1840:. 1817:. 1807:. 1795:15 1793:. 1789:. 1768:. 1750:. 1723:56 1721:. 1717:. 1705:^ 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Index

Chlorophylls
RAL 6002 Leaf green
Pokémon
Pokémon LeafGreen
Lemon balm leaves
A microscope image of plant cells, with chloroplasts visible as small green balls
chloroplasts
A leaf absorbing blue and red light, but reflecting green light
The structure of chlorophyll d
chlorin
magnesium
ligand
pigments
cyanobacteria
chloroplasts
algae
plants
Greek
energy
blue portion
electromagnetic spectrum
chlorophyll a
b
Joseph Bienaimé Caventou
Pierre Joseph Pelletier
magnesium
Richard Willstätter
Hans Fischer
stereochemistry
Robert Burns Woodward

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