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Cepaea nemoralis

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594: 779: 802:, and this species must mate to produce fertile eggs. Mating tends to be concentrated in late spring and early summer, though it can continue through the autumn. The snails may store the sperm they receive from their partner for some time, and individual broods can have mixed paternity. In Britain it lays clutches of 30–50 (in France 40–80) oval eggs between June and August (in France May–October, in W France until November). The size of the egg is 3.1 × 2.6 mm or egg diameter can be 2.3–3.0 mm. Juveniles hatch after 15–20 days. 86: 630: 431: 415: 606: 35: 61: 642: 618: 661: 1991:
Giffard, B.; Winter, S.; Guidoni, S.; Nicolai, A.; Castaldini, M.; Cluzeau, D.; Coll, P.; Cortet, J.; Le Cadre, E.; d’Errico, G.; Forneck, A.; Gagnarli, E.; Griesser, M.; Guernion, M.; Lagomarsino, A.; Landi, S.; Le Bissonnais, Y.; Mania, E.; Mocali, S.; Preda, C.; Priori, S.; Reineke, A; Rusch, A.;
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Silvertown, J.; Cook, L.; Cameron, R.; Dodd, M.; McConway, K.; Worthington, J.; Skelton, P.; Anton, C.; Bossdorf, O.; Baur, B.; Schilthuizen, M.; Fontaine, B.; Sattmann, H.; Bertorelle, G.; Correia, M.; Oliveira, C.; Pokryszko, B.; OĆŒgo, M.; StalaĆŸs, A.; Gill, E.; Rammul, Ü.; SĂłlymos, P.; FĂ©her, Z.;
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OĆŒgo, M.; Cameron, R.A.D.; HorsĂĄk, M.; Pokryszko, B.; Chudaƛ, M.; Cichy, A.; Kaczmarek, S.; Kobak, J.; Marzec, M.; Mierzwa-Szymkowiak, D.; Parzonko, D.; Pyka, G.; Rosin, Z.; Skawina, A.; Soroka, M.; Sulikowska-Drozd, A.; Surowiec, T.; Szymanek, M.; Templin, J.; UrbaƄska, M.; Zając, K.; Zielska, J.;
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is highly variable; it ranges from brown, through pink, to yellow or even whitish, with or without one to five dark-brown bands. Names for many colour variants were coined in the nineteenth century but this system has been replaced by an independent scoring of shell colour and the presence/absence
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Starting in 1857, there have been multiple introductions to North America, where it now occurs widely in Canada (from British Columbia to Newfoundland; in Ontario it is a pest of vineyards) and in the north-eastern part of the US, with further occurrences further south (e.g. California, Colorado,
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This snail is comparatively slow-growing, taking 1 to 3 years to develop from an egg to a breeding adult. The life-span for this species is up to seven or eight years, with the annual survival rate of adults about 50% (= 3% over five years, older adults suffer higher mortalities). In winter, the
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in shell colour and banding. The background colour of the shell varies along a continuum from brown through pink to yellow and sometimes almost white. Additionally the shells can be with or without dark bands. The bands vary in intensity of colour, in width and in number, from zero to five. The
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This is a very common and widespread species in Western Europe, occupying a wide range of habitats from coastal dunes to woodlands with full canopy cover, including gardens and abandoned land. In Eastern Europe, where it is a new arrival, it has turned up particularly in urban areas and other
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This species feeds mainly on dead or senescent plants. It prefers broad-leaved plants over grasses. Although mostly not a pest of crops, it can be a nuisance in vineyards because it is inadvertently picked with the grapes.
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disturbed habitats. It can be found up to an altitude of 1600 m in the Alps, 1800 m in the Pyrenees, 900 m in Wales, 600 m in Scotland. Density of adults is often of the order of 2 per square metre.
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or climatic selection: paler, more reflective colours in sunny environments reduce water loss and overheating. Climatic selection can also explain why yellow shells are more common in the south.
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is from Western and Northern Europe to Central Europe, but it has been spreading eastwards especially over the last few decades. Thus it is known from most of the Iberian Peninsula,
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develops a 'search image' for the commonest morph, so that the rarer morphs are less likely to be predated. Natural selection would then favour a diversity of
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per regna tria naturĂŠ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio decima, reformata.
1469:Ć»bikowska, E.; Ć»oƂądek, J. (17 April 2019). "Cepaea nemoralis (Gastropoda: Pulmonata) in Poland: patterns of variation in a range-expanding species". 713:, Hungary, Bosnia, Croatia, and the northern half of Italy. In Central and Eastern Europe it has spread particularly along the Baltic coast (e.g. in 593: 2339: 710: 2154: 694: 2404: 564:
pressures in different areas with mixing by migration, and balanced polymorphism. Balanced polymorphism could arise when a predator like the
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Another question is why the variation persists, usually even within a locality. Researchers have variously argued that the cause is random
702: 1440: 2313: 2365: 1732:Čejka, T. (3 August 2020). "The terrestrial molluscan fauna in the Slovak part of the Danubian Lowland: an annotated checklist". 1255:
European non-marine molluscs, a guide for species identification = Bestimmungsbuch fĂŒr europĂ€ische Land- und SĂŒsswassermollusken
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is among the largest and, because of its bright colouration, one of the best-known snails in Western Europe. The colour of the
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Chang, H.-W.; Emlen, John M. (1 April 1993). "Seasonal variation of microhabitat distribution of the polymorphic land snail
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The thickened and slightly out-turned apertural lip of adults is usually dark brown, but can be white in some regions. The
2115: 1117: 545:. For instance, in stable habitats shells tend to be darker in woodland than in open habitats. The explanation might be 360:
is closed in adults but narrowly open in juveniles. The shell surface is semi-glossy. An adult shell consists of 4œ–5Âœ
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Schroers, H.-J.; Simoni, S.; Steiner, M.; Temneanu, E.; Bacher, S.; Costantini, E.A.C.; Zaller, J.; Leyer, I. (2022).
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2003 BCPC Symposiumproceedings No. 80: Slugs & snails: agricultural, veterinary & environmental perspectives
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Juƙičková, L.; Horsák, M.; Beran, L. (2001). "Check-list of the molluscs (Mollusca) of the Czech Republic".
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Turner, H.; Kuiper, J.G.J.; Thew, N.; Bernasconi, R.; RĂŒetschi, J.; WĂŒthrich, M.; Gosteli, M. (1998).
381:. They share much the same habitat and exhibit a similar range of shell colours and banding patterns. 2435: 1905:
Williamson, P.; Cameron, R.A.D.; Carter, M.A. (February 1977). "Population dynamics of the landsnail
497: 485: 481: 386: 1701:(Gastropoda: Helicidae) in Eastern Europe: spreading history and the shell colouration variability" 1140:"The scoring of polymorphic colour and pattern variation and its genetic basis in molluscan shells" 85: 2396: 842:
This article includes public domain text from the reference and CC-BY-2.5 text from the reference
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is rare and scattered in northern Scotland, where it has been introduced. It is not found in the
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tends to grow larger, but usually the species can most easily be recognised by the colour of the
939: 879: 721:, Estonia, southern Finland, the east coast of Sweden) but also now elsewhere in Poland, and in 2344: 2165: 725:, Belarus and Russia. In Slovakia it was still known only from a single garden centre by 2020. 617: 526: 991:"Citizen science reveals unexpected continental-scale evolutionary change in a model organism" 853:
2013 IUCN Red List of Threatened Species. <www.iucnredlist.org>. Cited 28 December 2013.
778: 1994:"Vineyard management and its impacts on soil biodiversity, functions, and ecosystem services" 1695:
Gural-Sverlova, N.; Egorov, R.; Kruglova, O.; Kovalevich, N.; Gural, R. (27 September 2021).
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Both in America and Eastern Europe it is known that some introductions have been deliberate.
641: 605: 180: 2383: 1583:"Deep structure, long‐distance migration and admixture in the colour polymorphic land snail 2261: 2212: 2133: 1550:(Gastropoda: Pulmonata) in the Czech Republic with comments on other land snail immigrants" 267: 8: 2217: 476: 357: 330: 50: 2030: 1741: 1405:
Philosophical Transactions of the Royal Society of London. Series B: Biological Sciences
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In areas where lip colour is variable, dissection is necessary. A cross-section of the
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Layton, K.K.S.; Warne, C.P.K.; Nicolai, A.; Ansart, A.; deWaard, J.R. (April 2019).
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Kerstes, N.A.G.; Breeschoten, T.; Kalkman, V.J.; Schilthuizen, M. (19 July 2019).
517: 2417: 2149: 1042:"Snail shell colour evolution in urban heat islands detected via citizen science" 1007: 864: 557: 361: 349: 334: 137: 1482: 530:
genetics underlying this variation is extensively understood and is shared with
2300: 2287: 326: 1566: 1541: 1325: 1305:"Discrete or indiscrete? Redefining the colour polymorphism of the land snail 1201:"Shooting darts: co-evolution and counter-adaptation in hermaphroditic snails" 1057: 2452: 2378: 2188: 2010: 1993: 799: 745: 553: 287: 197: 70: 65: 2357: 2073: 1677: 1660: 1380: 1359: 1969: 1620: 1416: 1344: 1236: 1217: 1075: 1026: 2124:
taxonomy,short description, distribution, biology,status (threats), images
2102:. International Veterinary Information Service. Accessed 24 November 2009. 1787: 1765:"Molecular evidence for multiple introductions of the banded grove snail ( 1717: 1696: 1634:
Koivunen, A.; Malinen, P.; Ormio, H.; Terhivuo, J.; Valovirta, I. (2014).
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Davison, A.; Jackson, H.J.; Murphy, E.W.; Reader, T. (August 2019).
2279: 2182: 2054:"Variation in growth rate as an ecological factor in the landsnail 741: 733: 457:
has bifurcated blades. The mucous gland has 3 or fewer branches in
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Peltanová, Alena; Dvoƙák, Libor; Juƙičková, Lucie (1 April 2012).
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OĆŒgo, M.; Örstan, A.; Kirschenstein, M.; Cameron, R. (May 2016).
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A field guide to the land snails of Britain and north-west Europe
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Jones, J.S.; Leith, B.H.; Rawlings, P. (1977). "Polymorphism in
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Suomen kotilot ja etanat : opas maanilviĂ€isten maailmaan
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Ramos‐Gonzalez, D.; Saenko, S.V.; Davison, A. (August 2022).
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It is one of the most common large species of land snail in
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Life & Environment, University of the West of Scotland
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The polymorphism has also been intensely studied for its
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10.3101/1098-7096(2006)66[82:CNGHTI]2.0.CO;2
1762: 1539: 1904: 1884:. NeuchĂątel: CentreSuisse de cartographie de la faune. 576:. Most probably, the polymorphism has multiple causes. 393:
consistently has a dark-brown lip to its shell, whilst
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Caracoles y babosas de la Península Ibérica y Baleares
1248: 1246: 1690: 1688: 1128:, last modified 6 February 2011, accessed 1 May 2011. 1859:. Corvallis, Oregon: Oregon State University Press. 1520: 1364:(L.): interrogating the Evolution Megalab database" 1243: 1882:Atlas der Mollusken der Schweiz und Liechtensteins 1685: 1167: 810:, but can become active again during warm spells. 581:Different coloration and banding of the shells of 453:shows a cross with simple blades, whereas that of 389:of adult shells. In a high proportion of regions, 496:is distinct in having a small blunt tooth in its 2450: 1439:Dvoƙák L., Honěk A. & MartĂ­nkovĂĄ Z. (2003). 1198: 983: 981: 979: 977: 975: 973: 971: 969: 967: 965: 504:lies noticeably closer to the columella than in 353:and fusion of individual bands numbered 1 to 5. 1252: 500:, whilst the lowest brown band on the shell of 1898: 1857:Land snails and slugs of the Pacific Northwest 1807: 1805: 1495: 1199:Koene, J.M.; Schulenburg, H. (30 March 2005). 770:but extends further north and less far south. 2096:"Canine Angiostrongylosis (French Heartworm)" 1818:(Gastropoda, Helicidae): The invited invader" 1277: 962: 1163: 1161: 1159: 1157: 1802: 1463: 1461: 1459: 1457: 1357: 1351: 1296: 2139:Evolution MegaLab was an online survey of 2051: 1943: 1822:Journal of the Kentucky Academy of Science 1271: 59: 33: 2009: 1716: 1676: 1610: 1565: 1471:Biological Journal of the Linnean Society 1424: 1379: 1334: 1324: 1226: 1216: 1154: 1065: 1016: 1006: 816:is known experimentally to be a host for 2100:Companion and Exotic Animal Parasitology 1661:"Dispersal of land snails by sea storms" 1454: 1388: 1358:Cameron, R.A.D.; Cook, L.M. (May 2012). 1174:Annual Review of Ecology and Systematics 777: 659: 1811: 1445:(L.) populations in the Czech Republic" 1360:"Habitat and the shell polymorphism of 1172:: a problem with too many solutions?". 701:, Denmark, southern Sweden and Norway, 480:both have a lip that is brown near the 2451: 2018: 1278:Kerney, M.P.; Cameron, R.A.D. (1979). 773: 511: 2164: 2163: 1854: 1731: 1257:. Göttingen: Planet Poster Editions. 1113: 1111: 1109: 1107: 1105: 2306:cba91759-90d4-4c12-8453-4d03b11fc37a 2052:Oosterhoff, L. M. (1 January 1976). 1394: 1137: 1103: 1101: 1099: 1097: 1095: 1093: 1091: 1089: 1087: 1085: 744:. It seems to have been affected by 2459:IUCN Red List least concern species 2031:The biology of terrestrial molluscs 1186:10.1146/annurev.es.08.110177.000545 13: 1998:Frontiers in Ecology and Evolution 728:At the northern edge of its range, 468:Two superficially similar species 14: 2490: 2109: 1742:10.22541/au.159620996.68925423/v2 1496:Cadevall, J.; Orozco, A. (2016). 1082: 367: 321:is the type species of the genus 869:pp. , 1-824. Holmiae. (Salvius). 640: 628: 616: 604: 592: 518:Cepaea § Shell polymorphism 429: 413: 84: 2143:polymorphism (no longer online) 2088: 2045: 1984: 1937: 1873: 1848: 1756: 1725: 1652: 1627: 1591:Journal of Evolutionary Biology 1574: 1533: 1514: 1489: 1433: 1192: 655: 2062:Netherlands Journal of Zoology 1812:Whitson, M. (September 2005). 1395:Cook, L.M. (29 October 1998). 1253:Welter-Schultes, F.W. (2012). 1131: 1033: 932: 902: 872: 856: 847: 340: 1: 2034:. CABI Publishing, Oxon, UK, 1911:The Journal of Animal Ecology 837: 286:, and has been introduced to 2469:Gastropods described in 1758 1665:Journal of Molluscan Studies 1368:Journal of Molluscan Studies 1008:10.1371/journal.pone.0018927 492:species it is smooth. Also, 7: 2479:Taxa named by Carl Linnaeus 2094:Conboy G.A. (30 May 2000). 1776:Canadian Journal of Zoology 950:. Flanders Marine Institute 920:. Flanders Marine Institute 890:. Flanders Marine Institute 756:Texas and South Carolina). 677:The native distribution of 471:Caucasotachea vindobonensis 10: 2495: 2028:. In: Barker G. M. (ed.). 1705:Malacologica Bohemoslovaca 1542:"The spread of non-native 989:Juan, X. (27 April 2011). 752:in some parts of England. 515: 484:becoming pale towards the 310:Cepaea nemoralis nemoralis 2172: 2098:. In: Bowman D. D. (Ed.) 2042:. 1-146, cited page: 428. 1567:10.2478/s11756-012-0020-2 1483:10.1093/biolinnean/blz029 1326:10.1038/s41437-019-0189-z 1058:10.1038/s42003-019-0511-6 623:Yellow with several bands 220:Cepaea (Cepaea) nemoralis 216: 209: 186: 179: 81:Scientific classification 79: 57: 48: 41: 32: 23: 2011:10.3389/fevo.2022.850272 1205:BMC Evolutionary Biology 912:Cepaea nemoralis etrusca 766:mostly overlaps that of 673:Snail traversing a leaf. 407:, cross sections on left 302:Cepaea nemoralis etrusca 2074:10.1163/002829677X00072 2026:Life History Strategies 1909:L.: a six-year study". 1697:"Introduced land snail 1397:"A two–stage model for 819:Angiostrongylus vasorum 574:antipredator adaptation 1417:10.1098/rstb.1998.0311 1218:10.1186/1471-2148-5-25 1046:Communications Biology 783: 674: 647:Reddish-brown unbanded 375:is closely related to 1788:10.1139/cjz-2018-0084 1718:10.5817/MaB2021-20-75 1678:10.1093/mollus/eyv060 1638:. Helsinki: Tibiale. 1523:Acta Soc. Zool. Bohem 1381:10.1093/mollus/eyr052 1118:"Species summary for 798:land snails, it is a 781: 672: 2474:Gastropods of Europe 2301:Fauna Europaea (new) 2134:Encyclopedia of Life 1855:Burke, T.E. (2013). 1500:. Barcelona: Omega. 914:(RossmĂ€ssler, 1835)" 862:Linnaeus C. (1758). 599:A diversity of forms 570:colours and patterns 24:Lemon or Grove snail 16:Species of gastropod 1769:) in North America" 1411:(1375): 1577–1593. 1282:. London: Collins. 1138:Cain, A.J. (1988). 774:Biology and ecology 512:Colour polymorphism 477:Macularia sylvatica 461:, but 4 or more in 331:ecological genetics 305:(RossmĂ€ssler, 1835) 51:Conservation status 2148:2019-07-06 at the 2024:Heller J. (2001). 1962:10.1007/BF00328957 1548:Monacha cartusiana 784: 750:soil acidification 675: 325:. It is used as a 246:brown-lipped snail 2446: 2445: 2166:Taxon identifiers 1866:978-0-87071-685-0 1603:10.1111/jeb.14060 884:(Linnaeus, 1758)" 832:Turdus philomelos 670: 635:Yellow mid-banded 397:has a white lip. 314: 306: 262:of air-breathing 238: 237: 232: 224: 172:C. nemoralis 74: 2486: 2439: 2438: 2426: 2425: 2413: 2412: 2400: 2399: 2387: 2386: 2374: 2373: 2361: 2360: 2348: 2347: 2335: 2334: 2322: 2321: 2309: 2308: 2296: 2295: 2283: 2282: 2270: 2269: 2257: 2256: 2244: 2243: 2231: 2230: 2221: 2220: 2218:Cepaea_nemoralis 2208: 2207: 2206: 2204:Cepaea nemoralis 2193: 2192: 2191: 2174:Cepaea nemoralis 2161: 2160: 2129:Cepaea nemoralis 2117:Cepaea nemoralis 2103: 2092: 2086: 2085: 2056:Cepaea nemoralis 2049: 2043: 2022: 2016: 2015: 2013: 1988: 1982: 1981: 1946:Cepaea nemoralis 1941: 1935: 1934: 1907:Cepaea nemoralis 1902: 1896: 1895: 1877: 1871: 1870: 1852: 1846: 1845: 1816:Cepaea nemoralis 1809: 1800: 1799: 1773: 1767:Cepaea nemoralis 1760: 1754: 1753: 1729: 1723: 1722: 1720: 1699:Cepaea nemoralis 1692: 1683: 1682: 1680: 1656: 1650: 1649: 1631: 1625: 1624: 1614: 1597:(8): 1110–1125. 1585:Cepaea nemoralis 1578: 1572: 1571: 1569: 1544:Cepaea nemoralis 1537: 1531: 1530: 1518: 1512: 1511: 1493: 1487: 1486: 1465: 1452: 1443:Cepaea nemoralis 1437: 1431: 1430: 1428: 1392: 1386: 1385: 1383: 1362:Cepaea nemoralis 1355: 1349: 1348: 1338: 1328: 1307:Cepaea nemoralis 1300: 1294: 1293: 1275: 1269: 1268: 1250: 1241: 1240: 1230: 1220: 1196: 1190: 1189: 1165: 1152: 1151: 1135: 1129: 1120:Cepaea nemoralis 1115: 1080: 1079: 1069: 1037: 1031: 1030: 1020: 1010: 985: 960: 959: 957: 955: 936: 930: 929: 927: 925: 906: 900: 899: 897: 895: 882:Cepaea nemoralis 876: 870: 860: 854: 851: 824:Cepaea nemoralis 814:Cepaea nemoralis 671: 644: 632: 620: 608: 596: 583:Cepaea nemoralis 523:Cepaea nemoralis 502:C. vindobonensis 433: 417: 383:Cepaea nemoralis 378:Cepaea hortensis 373:Cepaea nemoralis 346:Cepaea nemoralis 319:Cepaea nemoralis 313:(Linnaeus, 1758) 312: 304: 255:Cepaea nemoralis 230: 223:(Linnaeus, 1758) 222: 192: 190:Cepaea nemoralis 89: 88: 68: 63: 62: 43:Cepaea nemoralis 37: 27:Cepaea nemoralis 21: 20: 2494: 2493: 2489: 2488: 2487: 2485: 2484: 2483: 2449: 2448: 2447: 2442: 2434: 2429: 2421: 2418:Observation.org 2416: 2408: 2403: 2395: 2390: 2382: 2377: 2369: 2364: 2356: 2351: 2343: 2338: 2330: 2325: 2317: 2312: 2304: 2299: 2291: 2286: 2278: 2273: 2265: 2260: 2252: 2247: 2239: 2234: 2226: 2224: 2216: 2211: 2202: 2201: 2196: 2187: 2186: 2181: 2168: 2150:Wayback Machine 2112: 2107: 2106: 2093: 2089: 2050: 2046: 2023: 2019: 1989: 1985: 1942: 1938: 1903: 1899: 1892: 1878: 1874: 1867: 1853: 1849: 1810: 1803: 1771: 1761: 1757: 1730: 1726: 1693: 1686: 1657: 1653: 1646: 1632: 1628: 1579: 1575: 1538: 1534: 1519: 1515: 1508: 1494: 1490: 1466: 1455: 1441:"The spread of 1438: 1434: 1393: 1389: 1356: 1352: 1301: 1297: 1290: 1276: 1272: 1265: 1251: 1244: 1197: 1193: 1166: 1155: 1136: 1132: 1116: 1083: 1038: 1034: 986: 963: 953: 951: 938: 937: 933: 923: 921: 908: 907: 903: 893: 891: 878: 877: 873: 865:Systema naturae 861: 857: 852: 848: 840: 822:. Predators of 776: 695:the Netherlands 660: 658: 653: 652: 651: 648: 645: 636: 633: 624: 621: 612: 611:Yellow unbanded 609: 600: 597: 586: 585: 558:founder effects 520: 514: 443: 442: 441: 440: 439: 434: 425: 424: 423: 418: 409: 408: 370: 343: 335:citizen science 333:, including in 228:Helix nemoralis 205: 194: 188: 175: 138:Stylommatophora 83: 75: 64: 60: 53: 25: 17: 12: 11: 5: 2492: 2482: 2481: 2476: 2471: 2466: 2461: 2444: 2443: 2441: 2440: 2427: 2414: 2401: 2388: 2375: 2362: 2349: 2336: 2323: 2310: 2297: 2288:Fauna Europaea 2284: 2271: 2258: 2245: 2232: 2222: 2209: 2194: 2178: 2176: 2170: 2169: 2158: 2157: 2152: 2136: 2125: 2111: 2110:External links 2108: 2105: 2104: 2087: 2044: 2017: 1983: 1956:(4): 501–507. 1936: 1897: 1890: 1872: 1865: 1847: 1801: 1782:(4): 392–398. 1755: 1724: 1684: 1671:(2): 341–343. 1651: 1644: 1626: 1573: 1560:(2): 384–389. 1532: 1513: 1506: 1488: 1453: 1432: 1387: 1374:(2): 179–184. 1350: 1319:(2): 162–175. 1295: 1288: 1270: 1263: 1242: 1191: 1180:(1): 109–143. 1153: 1130: 1081: 1032: 961: 931: 901: 871: 855: 845: 844: 839: 836: 775: 772: 711:Czech Republic 657: 654: 650: 649: 646: 639: 637: 634: 627: 625: 622: 615: 613: 610: 603: 601: 598: 591: 588: 587: 580: 579: 578: 516:Main article: 513: 510: 435: 428: 427: 426: 419: 412: 411: 410: 403:Love darts of 402: 401: 400: 399: 369: 368:Identification 366: 342: 339: 327:model organism 316: 315: 307: 298: 297: 295: 236: 235: 234: 233: 231:Linnaeus, 1758 225: 214: 213: 207: 206: 195: 184: 183: 177: 176: 169: 167: 163: 162: 155: 151: 150: 145: 141: 140: 135: 131: 130: 125: 121: 120: 115: 111: 110: 105: 101: 100: 95: 91: 90: 77: 76: 58: 55: 54: 49: 46: 45: 39: 38: 30: 29: 15: 9: 6: 4: 3: 2: 2491: 2480: 2477: 2475: 2472: 2470: 2467: 2465: 2462: 2460: 2457: 2456: 2454: 2437: 2432: 2428: 2424: 2419: 2415: 2411: 2406: 2402: 2398: 2393: 2389: 2385: 2380: 2376: 2372: 2367: 2363: 2359: 2354: 2350: 2346: 2341: 2337: 2333: 2328: 2324: 2320: 2315: 2311: 2307: 2302: 2298: 2294: 2289: 2285: 2281: 2276: 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1507:9788428215992 1503: 1499: 1492: 1484: 1480: 1476: 1472: 1464: 1462: 1460: 1458: 1450: 1446: 1444: 1436: 1427: 1422: 1418: 1414: 1410: 1406: 1402: 1401:polymorphism" 1400: 1391: 1382: 1377: 1373: 1369: 1365: 1363: 1354: 1346: 1342: 1337: 1332: 1327: 1322: 1318: 1314: 1310: 1308: 1299: 1291: 1285: 1281: 1274: 1266: 1264:9783933922755 1260: 1256: 1249: 1247: 1238: 1234: 1229: 1224: 1219: 1214: 1210: 1206: 1202: 1195: 1187: 1183: 1179: 1175: 1171: 1164: 1162: 1160: 1158: 1149: 1145: 1141: 1134: 1127: 1123: 1121: 1114: 1112: 1110: 1108: 1106: 1104: 1102: 1100: 1098: 1096: 1094: 1092: 1090: 1088: 1086: 1077: 1073: 1068: 1063: 1059: 1055: 1051: 1047: 1043: 1036: 1028: 1024: 1019: 1014: 1009: 1004: 1001:(4): e18927. 1000: 996: 992: 984: 982: 980: 978: 976: 974: 972: 970: 968: 966: 949: 945: 943: 935: 919: 915: 913: 905: 889: 885: 883: 875: 868: 866: 859: 850: 846: 843: 835: 833: 829: 825: 821: 820: 815: 811: 809: 803: 801: 800:hermaphrodite 797: 792: 788: 780: 771: 769: 765: 762:The range of 760: 757: 753: 751: 747: 746:air pollution 743: 739: 735: 731: 726: 724: 720: 716: 712: 708: 704: 700: 696: 692: 688: 687:Great Britain 684: 680: 643: 638: 631: 626: 619: 614: 607: 602: 595: 590: 589: 584: 577: 575: 571: 567: 563: 559: 555: 554:genetic drift 550: 548: 544: 540: 535: 533: 528: 524: 519: 509: 507: 503: 499: 495: 491: 487: 483: 479: 478: 473: 472: 466: 464: 460: 456: 452: 448: 438: 432: 422: 416: 406: 398: 396: 392: 388: 384: 380: 379: 374: 365: 363: 359: 354: 351: 347: 338: 336: 332: 328: 324: 320: 311: 308: 303: 300: 299: 296: 293: 292: 291: 289: 288:North America 285: 280: 278: 275: 272: 269: 265: 261: 257: 256: 251: 247: 243: 229: 226: 221: 218: 217: 215: 212: 208: 203: 199: 193: 191: 185: 182: 181:Binomial name 178: 174: 173: 168: 165: 164: 161: 160: 156: 153: 152: 149: 146: 143: 142: 139: 136: 133: 132: 129: 126: 123: 122: 119: 116: 113: 112: 109: 106: 103: 102: 99: 96: 93: 92: 87: 82: 78: 72: 67: 66:Least Concern 56: 52: 47: 44: 40: 36: 31: 28: 22: 19: 2173: 2140: 2128: 2116: 2099: 2090: 2068:(1): 1–132. 2065: 2061: 2055: 2047: 2029: 2025: 2020: 2001: 1997: 1986: 1953: 1949: 1945: 1939: 1923:10.2307/3955 1914: 1910: 1906: 1900: 1881: 1875: 1856: 1850: 1828:(2): 82–88. 1825: 1821: 1815: 1779: 1775: 1766: 1758: 1733: 1727: 1708: 1704: 1698: 1668: 1664: 1654: 1635: 1629: 1594: 1590: 1584: 1576: 1557: 1553: 1547: 1543: 1535: 1526: 1522: 1516: 1497: 1491: 1474: 1470: 1448: 1442: 1435: 1408: 1404: 1398: 1390: 1371: 1367: 1361: 1353: 1316: 1312: 1306: 1298: 1279: 1273: 1254: 1208: 1204: 1194: 1177: 1173: 1169: 1147: 1143: 1133: 1119: 1049: 1045: 1035: 998: 994: 952:. Retrieved 948:MolluscaBase 947: 941: 934: 922:. Retrieved 918:MolluscaBase 917: 911: 904: 892:. Retrieved 888:MolluscaBase 887: 881: 874: 863: 858: 849: 841: 831: 826:include the 823: 817: 813: 812: 804: 793: 789: 785: 768:C. nemoralis 767: 764:C. hortensis 763: 761: 758: 754: 730:C. nemoralis 729: 727: 679:C. nemoralis 678: 676: 656:Distribution 582: 560:, different 551: 536: 532:C. hortensis 531: 522: 521: 505: 501: 494:M. sylvatica 493: 489: 475: 469: 467: 463:C. hortensis 462: 459:C. nemoralis 458: 455:C. hortensis 454: 451:C. nemoralis 450: 444: 437:C. hortensis 436: 421:C. nemoralis 420: 404: 395:C. hortensis 394: 391:C. nemoralis 390: 382: 377: 372: 371: 355: 345: 344: 322: 318: 317: 309: 301: 281: 254: 253: 249: 245: 241: 239: 227: 219: 189: 187: 171: 170: 158: 42: 26: 18: 2392:NatureServe 2327:iNaturalist 2198:Wikispecies 1477:(1): 1–11. 1144:Malacologia 944:Held, 1838" 828:song thrush 806:snails may 703:Switzerland 693:, Belgium, 566:song thrush 527:polymorphic 341:Description 268:terrestrial 250:lemon snail 242:grove snail 2453:Categories 2132:images at 2122:Animalbase 1917:(1): 181. 1891:2884140131 1451:: 99-102. 1289:000219676X 1126:AnimalBase 1052:(1): 264. 838:References 547:camouflage 525:is highly 337:projects. 294:Subspecies 264:land snail 128:Gastropoda 2464:Helicidae 1950:Oecologia 1750:242206099 1711:: 75–91. 1211:(1): 25. 808:hibernate 796:pulmonate 794:Like all 562:selection 539:evolution 482:columella 447:love dart 358:umbilicus 274:gastropod 271:pulmonate 166:Species: 148:Helicidae 104:Kingdom: 98:Eukaryota 2397:2.107314 2345:10235748 2225:BioLib: 2183:Wikidata 2146:Archived 2082:55537228 1978:22594544 1970:28313817 1842:86005518 1796:91897997 1734:Authorea 1621:35830483 1554:Biologia 1529:: 25–40. 1345:30804571 1313:Heredity 1237:15799778 1076:31341963 1027:21556137 995:PLOS ONE 954:23 April 924:23 April 894:23 April 742:Shetland 734:Hebrides 498:aperture 211:Synonyms 198:Linnaeus 144:Family: 118:Mollusca 114:Phylum: 108:Animalia 94:Domain: 71:IUCN 3.1 2319:2294280 2189:Q114893 1612:9541890 1426:1692378 1336:6629550 1228:1080126 1150:: 1–15. 1067:6642149 1018:3083392 723:Ukraine 707:Austria 699:Germany 691:Ireland 543:ecology 277:mollusc 260:species 258:) is a 154:Genus: 134:Order: 124:Class: 69: ( 2436:235792 2384:156896 2358:113097 2293:426306 2280:CEPANE 2267:449909 2141:Cepaea 2080:  2038:  1976:  1968:  1929:  1888:  1863:  1840:  1794:  1748:  1642:  1619:  1609:  1504:  1423:  1399:Cepaea 1343:  1333:  1286:  1261:  1235:  1225:  1170:Cepaea 1074:  1064:  1025:  1015:  942:Cepaea 782:Mating 738:Orkney 719:Latvia 715:Poland 709:, the 683:France 572:as an 506:Cepaea 490:Cepaea 486:suture 405:Cepaea 362:whorls 323:Cepaea 284:Europe 159:Cepaea 2431:WoRMS 2410:28835 2371:77910 2340:IRMNG 2332:48586 2241:28635 2078:S2CID 2058:(L.)" 1974:S2CID 1927:JSTOR 1838:S2CID 1792:S2CID 1772:(PDF) 1746:S2CID 350:shell 2423:7720 2405:NCBI 2379:IUCN 2366:ITIS 2314:GBIF 2275:EPPO 2254:SBY4 2236:BOLD 2228:2977 2036:ISBN 1966:PMID 1931:3955 1886:ISBN 1861:ISBN 1640:ISBN 1617:PMID 1546:and 1502:ISBN 1341:PMID 1284:ISBN 1259:ISBN 1233:PMID 1072:PMID 1023:PMID 956:2023 926:2023 896:2023 748:and 556:and 541:and 474:and 266:, a 240:The 202:1758 2353:ISC 2262:EoL 2249:CoL 2213:ADW 2120:at 2070:doi 2006:doi 1958:doi 1948:". 1919:doi 1830:doi 1784:doi 1738:doi 1713:doi 1673:doi 1607:PMC 1599:doi 1562:doi 1479:doi 1475:127 1421:PMC 1413:doi 1409:353 1376:doi 1331:PMC 1321:doi 1317:123 1223:PMC 1213:doi 1182:doi 1062:PMC 1054:doi 1013:PMC 1003:doi 834:). 740:or 449:of 387:lip 329:in 248:or 2455:: 2433:: 2420:: 2407:: 2394:: 2381:: 2368:: 2355:: 2342:: 2329:: 2316:: 2303:: 2290:: 2277:: 2264:: 2251:: 2238:: 2215:: 2200:: 2185:: 2076:. 2066:27 2064:. 2060:. 2004:. 2002:10 2000:. 1996:. 1972:. 1964:. 1954:93 1952:. 1925:. 1915:46 1913:. 1836:. 1826:66 1824:. 1820:. 1804:^ 1790:. 1780:97 1778:. 1774:. 1744:. 1736:. 1709:20 1707:. 1703:. 1687:^ 1669:82 1667:. 1663:. 1615:. 1605:. 1595:35 1593:. 1589:. 1558:67 1556:. 1552:. 1527:65 1525:. 1473:. 1456:^ 1447:. 1419:. 1407:. 1403:. 1372:78 1370:. 1366:. 1339:. 1329:. 1315:. 1311:. 1245:^ 1231:. 1221:. 1207:. 1203:. 1176:. 1156:^ 1148:28 1146:. 1142:. 1124:. 1084:^ 1070:. 1060:. 1048:. 1044:. 1021:. 1011:. 997:. 993:. 964:^ 946:. 916:. 886:. 736:, 717:, 705:, 697:, 689:, 685:, 534:. 508:. 465:. 290:. 279:. 244:, 200:, 2084:. 2072:: 2014:. 2008:: 1980:. 1960:: 1933:. 1921:: 1894:. 1869:. 1844:. 1832:: 1814:" 1798:. 1786:: 1752:. 1740:: 1721:. 1715:: 1681:. 1675:: 1648:. 1623:. 1601:: 1587:" 1570:. 1564:: 1510:. 1485:. 1481:: 1429:. 1415:: 1384:. 1378:: 1347:. 1323:: 1309:" 1292:. 1267:. 1239:. 1215:: 1209:5 1188:. 1184:: 1178:8 1122:" 1078:. 1056:: 1050:2 1029:. 1005:: 999:6 958:. 940:" 928:. 910:" 898:. 880:" 830:( 252:( 204:) 196:( 73:)

Index


Conservation status
Least Concern
IUCN 3.1
Scientific classification
Edit this classification
Eukaryota
Animalia
Mollusca
Gastropoda
Stylommatophora
Helicidae
Cepaea
Binomial name
Linnaeus
1758
Synonyms
species
land snail
terrestrial
pulmonate
gastropod
mollusc
Europe
North America
model organism
ecological genetics
citizen science
shell
umbilicus

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