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Sepal

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135: 88: 127: 2248: 110: 96: 38: 511:. In some species, the calyx not only persists after flowering but instead of withering, begins to grow until it forms a bladder-like enclosure around the fruit. This is an effective protection against some kinds of birds and insects, for example in 330:
are difficult to distinguish, e.g. the petals and sepals share the same color or the petals are absent and the sepals are colorful. When the undifferentiated tepals resemble petals, they are referred to as "petaloid", as in
485:) the calyx grows along with the fruit, possibly to protect the attachment point. Some plants retain a thorny calyx, either dried or live, as protection for the fruit or seeds. Examples include species of 426:
Mechanical cues may be responsible for sepal growth and there is a strong evidence suggesting that microtubules are present and determine the tensile strength and direction of growth at a molecular level.
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Hervieux, Nathan; Dumond, Mathilde; Sapala, Aleksandra; Routier-Kierzkowska, Anne-Lise; Kierzkowski, Daniel; Roeder, Adrienne H.K.; Smith, Richard S.; Boudaoud, Arezki; Hamant, Olivier (April 2016).
438:, both sepals and petals are modified leaves. The calyx (the sepals) and the corolla (the petals) are the outer sterile whorls of the flower, which together form what is known as the 2078: 2083: 2216: 720:
Jackson, Benjamin, Daydon; A Glossary of Botanic Terms with their Derivation and Accent; Published by Gerald Duckworth & Co. London, 4th ed 1928
450:, the calyx is the primary whorl, forming a flower up to 20 inches (51 cm) wide, with one sepal growing to a length of 13 feet (4.0 m) – 56: 2134: 1950: 91:
Diagram showing the parts of a mature flower. In this example, the perianth is separated into a calyx (sepals) and corolla (petals)
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After flowering, most plants have no more use for the calyx which withers or becomes vestigial, although in a few plants such as
202:(flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the 648: 339:, an alternative name is lilioid monocots. Examples of plants in which the term tepal is appropriate include genera such as 389:. They may be free (polysepalous) or fused together (gamosepalous). Often, the sepals are much reduced, appearing somewhat 900:
The identification of flowering plant families, including a key to those native and cultivated in north temperate regions
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John Entick, William Crakelt, Tyronis thesaurus, or, Entick's new Latin English dictionary. Publisher: E.J. Coale, 1822
705: 979:"Photosynthetic performance of vegetative and reproductive structures of green hellebore (Helleborus viridis L. agg.)" 2140: 907: 783: 669: 74: 393:-like, or as scales, teeth, or ridges. Most often such structures protrude until the fruit is mature and falls off. 1035: 1474: 466:. However, photosynthesis in sepals occurs at a slower rate than in ordinary leaves due to sepals having a lower 219: 1955: 17: 1405: 303: 276: 230: 2124: 1711: 1226: 2206: 1960: 1940: 640: 2272: 2090: 2228: 2073: 1123: 451: 2058: 1982: 223: 1787: 2223: 616: 611: 1945: 1842: 1797: 1469: 1028: 852: 435: 1445: 875: 8: 2115: 1660: 115: 101: 856: 29:
Any of the separate parts of the calyx of a flower (excluding the bracts), usually green
2053: 1918: 1696: 1388: 1174: 1070: 1000: 959: 817: 2211: 1060: 1055: 903: 880: 779: 701: 665: 644: 379: 332: 1004: 1997: 1903: 1898: 1852: 1745: 1730: 1691: 1616: 1457: 1381: 1278: 1220: 990: 951: 870: 860: 685: 681: 534: 513: 153: 1792: 2251: 2233: 2095: 1924: 1874: 1862: 1824: 1760: 1587: 1555: 1516: 1489: 1318: 1313: 1204: 1140: 1021: 522: 386: 120: 939: 370:. Flower merosity is indicative of a plant's classification. The merosity of a 2145: 2110: 2063: 2045: 2009: 1765: 1592: 1570: 1536: 1511: 1494: 1484: 1479: 1361: 1179: 1164: 1118: 1113: 463: 251: 134: 52: 995: 978: 865: 840: 2266: 2181: 1929: 1847: 1819: 1782: 1750: 1686: 1655: 1606: 1506: 1308: 1291: 1214: 1194: 1135: 1130: 1078: 621: 504: 375: 312:'cup, goblet'; both words have been used interchangeably in botanical Latin. 266: 260: 2176: 2166: 2068: 1777: 1286: 1253: 884: 446: 390: 1992: 1977: 1972: 1908: 1770: 1550: 1521: 1420: 1415: 1263: 1103: 1098: 1083: 199: 126: 87: 2021: 2014: 2004: 1987: 1670: 1665: 1601: 1393: 1240: 1199: 1093: 963: 841:"A Mechanical Feedback Restricts Sepal Growth and Shape in Arabidopsis" 838: 492: 419:
includes the bases of sepals, petals, and the attachment points of the
416: 408: 335:, orders of monocots with brightly colored tepals. Since they include 2129: 2026: 1755: 1560: 1541: 1452: 1376: 1296: 1248: 1189: 1169: 1108: 557: 496: 359: 307: 280: 236: 109: 955: 940:"Revision of the North and Central American Species of Aristolochia" 396:
Examples of flowers with much-reduced perianths are found among the
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In some flowers, the sepals are fused towards the base, forming a
1356: 1351: 1268: 1258: 397: 1809: 1701: 1621: 1575: 1565: 1526: 1464: 1440: 1435: 1430: 1425: 1044: 700:. United Kingdom: Oxford University Press. 2007. p. 3804. 487: 462:
Similarly to ordinary leaves, sepals are capable of performing
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The development and form of the sepals vary considerably among
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Aschan, G.; Pfanz, H.; Vodnik, D.; Batič, F. (1 March 2005).
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Stearn, William T. (2000). Botanical Latin, 4th ed.: 38-39.
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In many Fabaceae flowers, a calyx tube surrounds the petals.
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Stearn, William T. (2000). Botanical Latin, 4th ed.: 38
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flower is typically four or five. The merosity of a
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Organogenesis of Flowers. A Photographic Text-Atlas
594:"Oxford Languages | The Home of Language Data" 415:). In other flowers (e.g., Rosaceae, Myrtaceae), a 174: 156: 47:
may be too technical for most readers to understand
470:density which limits the spaces for gas exchange. 2264: 682:Corollarium ad Philosophiam botanicam Linnaei 18 756: 628: 1029: 738: 521:. In other species, the calyx grows into an 2135:International Association for Plant Taxonomy 732: 363:have well-distinguished sepals and petals. 142:fruit is the source of its vulgar nicknames. 897: 296: 1036: 1022: 741:A Concise Etymological Dictionary of Latin 994: 891: 874: 864: 698:Shorter Oxford English dictionary, 6th ed 382:flower is three, or a multiple of three. 326:is usually applied when the parts of the 75:Learn how and when to remove this message 59:, without removing the technical details. 2079:International Code of Nomenclature (ICN) 366:The number of sepals in a flower is its 246:Collectively, the sepals are called the 133: 125: 108: 94: 86: 944:Annals of the Missouri Botanical Garden 937: 795: 762: 714: 634: 14: 2265: 922: 816:. University of Hawaii. Archived from 254:of parts that form a flower. The word 1017: 743:. Halle (Saale): Max Niemeyer Verlag. 57:make it understandable to non-experts 286: 31: 690: 24: 927:. New York: Sterling. p. 208. 25: 2284: 2084:ICN for Cultivated Plants (ICNCP) 250:(plural: calyces), the outermost 2247: 2246: 898:Davis, P.H.; Cullen, J. (1979). 807: 152: 36: 970: 931: 916: 832: 801: 778:. University of Toronto Press. 768: 747: 876:11858/00-001M-0000-002B-1620-2 723: 674: 658: 604: 586: 579:'petal', influenced by French 546: 454:, the largest of all calyces. 351:. In contrast, genera such as 315: 285:'bud, calyx, husk, wrapping' ( 222:in 1790, and derived from 113:After blooming, the sepals of 13: 1: 938:Pfeifer, HW (November 1966). 540: 430: 2125:History of plant systematics 1712:Thorns, spines, and prickles 925:Tropical Plants of the World 308: 281: 220:Noël Martin Joseph de Necker 209: 7: 528: 457: 291: 258:was adopted from the Latin 10: 2289: 1941:Alternation of generations 1043: 641:Royal Botanic Gardens, Kew 444:. In some plants, such as 302: 275: 264:, not to be confused with 229: 105:showing petals and sepals. 2242: 2190: 2154: 2091:Cultivated plant taxonomy 2054:Biological classification 2044: 1917: 1833: 1729: 1679: 1404: 1334: 1277: 1239: 1213: 1149: 1069: 1051: 996:10.1007/s11099-005-5064-x 866:10.1016/j.cub.2016.03.004 270:'cup, goblet'. The Latin 1951:Evolutionary development 923:Rohwer, Jens G. (2002). 680:Necker, N.J. de (1790). 452:Aristolochia grandiflora 1602:Hypanthium (Floral cup) 798:, pp. 51 & 91. 138:The large calyx of the 2217:by author abbreviation 2141:Plant taxonomy systems 2059:Botanical nomenclature 739:Tucker, T. G. (1931). 637:The Kew Plant Glossary 635:Beentje, Henk (2010). 501:Physalis philadelphica 301:is derived from Greek 297: 274:is derived from Greek 265: 259: 143: 131: 123: 119:expand into an edible 106: 99:Tetramerous flower of 92: 2224:Botanical expeditions 617:CollinsDictionary.com 137: 129: 112: 98: 90: 1956:Evolutionary history 1946:Double fertilization 1798:Cellular respiration 639:. Richmond, Surrey: 1175:Non-vascular plants 857:2016CBio...26.1019H 194:) is a part of the 116:Hibiscus sabdariffa 102:Ludwigia octovalvis 1680:Surface structures 1475:Flower development 820:on 5 December 2008 774:Sattler, R. 1973. 407:(as in the family 243: 'covering'. 144: 132: 124: 107: 93: 2260: 2259: 1899:Herbaceous plants 1725: 1724: 650:978-1-84246-422-9 598:languages.oup.com 495:(for example the 483:Solanum melongena 333:petaloid monocots 85: 84: 77: 16:(Redirected from 2280: 2273:Plant morphology 2250: 2249: 2229:Individual trees 1904:Secondary growth 1875:Succulent plants 1863:Prostrate shrubs 1746:Apical dominance 1731:Plant physiology 1692:Epicuticular wax 1237: 1236: 1230: 1221:Plant morphology 1038: 1031: 1024: 1015: 1014: 1009: 1008: 998: 974: 968: 967: 935: 929: 928: 920: 914: 913: 895: 889: 888: 878: 868: 851:(8): 1019–1028. 836: 830: 829: 827: 825: 805: 799: 793: 787: 772: 766: 760: 754: 751: 745: 744: 736: 730: 727: 721: 718: 712: 711: 694: 688: 678: 672: 662: 656: 654: 632: 626: 625: 608: 602: 601: 590: 584: 550: 535:Plant morphology 514:Hibiscus trionum 387:flowering plants 311: 306: 300: 294: 288: 284: 279: 240: 233: 193: 192: 189: 188: 185: 182: 179: 176: 171: 170: 167: 164: 161: 158: 80: 73: 69: 66: 60: 40: 39: 32: 21: 2288: 2287: 2283: 2282: 2281: 2279: 2278: 2277: 2263: 2262: 2261: 2256: 2238: 2207:Botanical terms 2200: 2186: 2150: 2096:Citrus taxonomy 2074:Author citation 2040: 1934: 1913: 1835: 1829: 1825:Turgor pressure 1733: 1721: 1675: 1490:Floral symmetry 1408: 1400: 1330: 1319:Vascular bundle 1314:Vascular tissue 1273: 1233: 1224: 1223: 1209: 1180:Vascular plants 1145: 1141:Plant pathology 1065: 1047: 1042: 1012: 983:Photosynthetica 975: 971: 956:10.2307/2394940 936: 932: 921: 917: 910: 896: 892: 845:Current Biology 837: 833: 823: 821: 806: 802: 794: 790: 773: 769: 761: 757: 752: 748: 737: 733: 728: 724: 719: 715: 708: 696: 695: 691: 679: 675: 663: 659: 651: 633: 629: 610: 609: 605: 592: 591: 587: 575:from Neo-Latin 571:'covering' and 551: 547: 543: 531: 523:accessory fruit 519:Cape gooseberry 460: 436:Morphologically 433: 318: 212: 206:when in bloom. 173: 155: 151: 121:accessory fruit 81: 70: 64: 61: 53:help improve it 50: 41: 37: 30: 23: 22: 15: 12: 11: 5: 2286: 2276: 2275: 2258: 2257: 2255: 2254: 2243: 2240: 2239: 2237: 2236: 2231: 2226: 2221: 2220: 2219: 2209: 2203: 2201: 2199: 2198: 2197:Related topics 2195: 2191: 2188: 2187: 2185: 2184: 2179: 2174: 2169: 2164: 2158: 2156: 2152: 2151: 2149: 2148: 2146:Taxonomic rank 2143: 2138: 2132: 2127: 2122: 2121: 2120: 2119: 2118: 2113: 2108: 2098: 2088: 2087: 2086: 2081: 2076: 2071: 2066: 2064:Botanical name 2056: 2050: 2048: 2046:Plant taxonomy 2042: 2041: 2039: 2038: 2037: 2036: 2031: 2030: 2029: 2022:Megasporangium 2019: 2018: 2017: 2010:Microsporangia 2002: 2001: 2000: 1995: 1990: 1985: 1975: 1970: 1965: 1964: 1963: 1953: 1948: 1943: 1937: 1935: 1933: 1932: 1927: 1921: 1915: 1914: 1912: 1911: 1906: 1901: 1896: 1895: 1894: 1893: 1892: 1882: 1877: 1872: 1871: 1870: 1865: 1855: 1850: 1848:Cushion plants 1839: 1837: 1831: 1830: 1828: 1827: 1822: 1817: 1812: 1807: 1802: 1801: 1800: 1795: 1785: 1783:Plant hormones 1780: 1775: 1774: 1773: 1766:Photosynthesis 1763: 1758: 1753: 1748: 1743: 1737: 1735: 1727: 1726: 1723: 1722: 1720: 1719: 1714: 1709: 1704: 1699: 1694: 1689: 1683: 1681: 1677: 1676: 1674: 1673: 1668: 1663: 1658: 1653: 1652: 1651: 1646: 1641: 1631: 1630: 1629: 1624: 1619: 1614: 1604: 1599: 1598: 1597: 1596: 1595: 1590: 1585: 1584: 1583: 1578: 1558: 1553: 1548: 1547: 1546: 1545: 1544: 1539: 1529: 1524: 1519: 1514: 1509: 1499: 1498: 1497: 1492: 1487: 1485:Floral formula 1482: 1480:Floral diagram 1477: 1472: 1462: 1461: 1460: 1455: 1450: 1449: 1448: 1443: 1433: 1423: 1418: 1412: 1410: 1409:(incl. Flower) 1402: 1401: 1399: 1398: 1397: 1396: 1391: 1386: 1385: 1384: 1379: 1369: 1359: 1354: 1349: 1344: 1338: 1336: 1332: 1331: 1329: 1328: 1323: 1322: 1321: 1311: 1309:Storage organs 1306: 1301: 1300: 1299: 1289: 1283: 1281: 1275: 1274: 1272: 1271: 1266: 1261: 1256: 1251: 1245: 1243: 1234: 1232: 1231: 1217: 1211: 1210: 1208: 1207: 1202: 1197: 1195:Spermatophytes 1192: 1187: 1182: 1177: 1172: 1167: 1165:Archaeplastida 1162: 1156: 1154: 1147: 1146: 1144: 1143: 1138: 1133: 1128: 1127: 1126: 1119:Phytogeography 1116: 1114:Phytochemistry 1111: 1106: 1101: 1096: 1091: 1086: 1081: 1075: 1073: 1071:Subdisciplines 1067: 1066: 1064: 1063: 1058: 1052: 1049: 1048: 1041: 1040: 1033: 1026: 1018: 1011: 1010: 969: 930: 915: 908: 890: 831: 808:Carr, Gerald. 800: 788: 767: 755: 746: 731: 722: 713: 707:978-0199206872 706: 689: 673: 657: 649: 627: 603: 585: 544: 542: 539: 538: 537: 530: 527: 491:, some of the 464:photosynthesis 459: 456: 432: 429: 317: 314: 295:'bud'), while 218:was coined by 211: 208: 83: 82: 44: 42: 35: 28: 18:Calyx (botany) 9: 6: 4: 3: 2: 2285: 2274: 2271: 2270: 2268: 2253: 2245: 2244: 2241: 2235: 2232: 2230: 2227: 2225: 2222: 2218: 2215: 2214: 2213: 2210: 2208: 2205: 2204: 2202: 2196: 2193: 2192: 2189: 2183: 2182:Phytochemical 2180: 2178: 2175: 2173: 2170: 2168: 2165: 2163: 2160: 2159: 2157: 2153: 2147: 2144: 2142: 2139: 2136: 2133: 2131: 2128: 2126: 2123: 2117: 2114: 2112: 2109: 2107: 2104: 2103: 2102: 2099: 2097: 2094: 2093: 2092: 2089: 2085: 2082: 2080: 2077: 2075: 2072: 2070: 2067: 2065: 2062: 2061: 2060: 2057: 2055: 2052: 2051: 2049: 2047: 2043: 2035: 2032: 2028: 2025: 2024: 2023: 2020: 2016: 2013: 2012: 2011: 2008: 2007: 2006: 2003: 1999: 1996: 1994: 1991: 1989: 1986: 1984: 1981: 1980: 1979: 1976: 1974: 1971: 1969: 1966: 1962: 1959: 1958: 1957: 1954: 1952: 1949: 1947: 1944: 1942: 1939: 1938: 1936: 1931: 1928: 1926: 1923: 1922: 1920: 1916: 1910: 1907: 1905: 1902: 1900: 1897: 1891: 1888: 1887: 1886: 1883: 1881: 1878: 1876: 1873: 1869: 1866: 1864: 1861: 1860: 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Retrieved 818:the original 811: 803: 796:Beentje 2010 791: 775: 770: 763:Beentje 2010 758: 749: 740: 734: 725: 716: 697: 692: 676: 660: 636: 630: 615: 606: 597: 588: 580: 576: 572: 568: 564: 560: 553: 552:From French 548: 512: 509:Trapa natans 508: 500: 486: 482: 472: 461: 447:Aristolochia 445: 439: 434: 425: 404: 402: 395: 384: 365: 358: 352: 346: 340: 321: 319: 271: 255: 247: 245: 237: 234: 227: 215: 213: 147: 145: 114: 100: 71: 62: 46: 26: 1993:Pollen tube 1988:Pollinators 1978:Pollination 1973:Germination 1788:Respiration 1771:Chlorophyll 1617:Pedicellate 1551:Gametophyte 1470:Aestivation 1421:Antheridium 1416:Archegonium 1264:Plasmodesma 1241:Plant cells 1104:Paleobotany 1099:Ethnobotany 1084:Astrobotany 824:20 December 567:from Greek 503:), and the 380:palaeodicot 316:Description 200:angiosperms 2015:Microspore 2005:Sporangium 1983:Artificial 1671:Sporophyte 1666:Sporophyll 1661:Receptacle 1556:Gynandrium 1426:Androecium 1335:Vegetative 1205:Angiosperm 1200:Gymnosperm 1094:Dendrology 950:(2): 164. 812:Lythraceae 541:References 493:Solanaceae 431:Morphology 417:hypanthium 409:Lythraceae 405:calyx tube 2212:Botanists 2130:Herbarium 2027:Megaspore 1925:Evolution 1868:Subshrubs 1836:and habit 1761:Nutrition 1756:Cellulose 1751:Bulk flow 1734:Materials 1697:Epidermis 1561:Gynoecium 1542:Endosperm 1537:Dispersal 1453:Staminode 1389:Sessility 1377:Cataphyll 1297:Mesophyll 1249:Cell wall 1190:Lycophyte 1170:Bryophyte 1124:Geobotany 1109:Phycology 558:Neo-Latin 497:Tomatillo 360:Phaseolus 320:The term 289:Sanskrit 214:The term 210:Etymology 2267:Category 2252:Category 2172:Forestry 2162:Agronomy 2155:Practice 2106:Cultivar 2101:Cultigen 1961:timeline 1853:Rosettes 1741:Aleurone 1717:Trichome 1634:Perianth 1446:Filament 1304:Meristem 1227:glossary 1089:Bryology 1005:24426595 885:27151660 655:, p. 106 583:'petal'. 529:See also 517:and the 479:eggplant 475:Lodoicea 468:stomatal 458:Function 441:perianth 413:Fabaceae 368:merosity 337:Liliales 328:perianth 1930:Ecology 1687:Cuticle 1517:Capsule 1507:Anatomy 1458:Tapetum 1382:Petiole 1357:Rhizome 1352:Rhizoid 1279:Tissues 1269:Vacuole 1259:Plastid 1061:Outline 1056:History 964:2394940 853:Bibcode 612:"sepal" 577:petalum 561:sepalum 556:, from 421:stamens 398:grasses 376:monocot 372:eudicot 216:sepalum 51:Please 2234:Plants 2137:(IAPT) 1890:Lianas 1858:Shrubs 1810:Starch 1702:Nectar 1622:Raceme 1588:Stigma 1576:Locule 1566:Carpel 1527:Pyrena 1465:Flower 1441:Anther 1436:Stamen 1431:Pollen 1153:groups 1045:Botany 1003:  962:  906:  883:  782:  704:  668:  647:  581:pétale 554:sépale 488:Acaena 411:, and 348:Tulipa 292:kalika 204:petals 196:flower 140:medlar 2194:Lists 2111:Group 2034:Spore 1968:Flora 1885:Vines 1880:Trees 1843:Habit 1815:Sugar 1707:Stoma 1649:Sepal 1644:Petal 1639:Tepal 1627:Umbel 1612:Bract 1593:Style 1581:Ovule 1571:Ovary 1512:Berry 1502:Fruit 1495:Whorl 1362:Shoot 1160:Algae 1151:Plant 1001:S2CID 960:JSTOR 573:-alum 569:skepē 323:tepal 309:kylix 304:κύλιξ 298:calix 282:kalyx 277:κάλυξ 272:calyx 267:calix 261:calyx 256:calyx 252:whorl 248:calyx 238:sképē 231:σκέπη 226: 148:sepal 2116:Grex 1998:Self 1532:Seed 1394:Stem 1372:Leaf 1347:Root 1342:Bulb 1326:Wood 1287:Cork 1185:Fern 904:ISBN 881:PMID 826:2008 780:ISBN 702:ISBN 666:ISBN 645:ISBN 565:sep- 477:and 357:and 354:Rosa 345:and 342:Aloe 1805:Sap 1522:Nut 1367:Bud 991:doi 952:doi 871:hdl 861:doi 391:awn 378:or 287:cf. 198:of 55:to 2269:: 999:. 987:43 985:. 981:. 958:. 948:53 946:. 942:. 879:. 869:. 859:. 849:26 847:. 843:. 686:31 684:, 643:. 620:. 614:. 596:. 525:. 507:, 499:, 423:. 400:. 187:əl 181:iː 169:əl 146:A 1229:) 1225:( 1037:e 1030:t 1023:v 1007:. 993:: 966:. 954:: 912:. 887:. 873:: 863:: 855:: 828:. 814:" 810:" 786:. 710:. 653:. 624:. 600:. 481:( 241:) 235:( 190:/ 184:p 178:s 175:ˈ 172:, 166:p 163:ɛ 160:s 157:ˈ 154:/ 150:( 78:) 72:( 67:) 63:( 49:. 20:)

Index

Calyx (botany)
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Ludwigia octovalvis

Hibiscus sabdariffa
accessory fruit


medlar
/ˈsɛpəl,ˈspəl/
flower
angiosperms
petals
Noël Martin Joseph de Necker
Ancient Greek
σκέπη
whorl
calyx
calix
κάλυξ
κύλιξ
tepal
perianth
petaloid monocots
Liliales
Aloe

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