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Anus

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537: 82: 522: 805: 67: 503:. In deuterostomes, the original dent becomes the anus while the gut eventually tunnels through to make another opening, which forms the mouth. The protostomes were so named because it was thought that in their embryos the dent formed the mouth first ( 509:
meaning "first") and the anus was formed later at the opening made by the other end of the gut. Research from 2001 shows the edges of the dent close up in the middles of protosomes, leaving openings at the ends which become the mouths and anuses.
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The development of the anus was an important stage in the evolution of multicellular animals. It appears to have happened at least twice, following different paths in
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Arendt, D.; Technau, U. & Wittbrodt, J. (4 January 2001). "Evolution of the bilaterian larval foregut".
450:, in which the body was built of repeated "modules" which could later specialize, such as the heads of most 527: 830: 608: 17: 735: 583: 616: 610: 210: 198: 186: 150: 126: 114: 809: 715: 49: 44:
This article is about the anuses of animals in general. For details specific to humans, see
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mammals also have a cloaca, which is thought to be a feature inherited from the earliest
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have a single orifice for excreting both solids and liquids and, in females, a separate
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Langstroth, Lovell; Libby Langstroth; Todd Newberry; Monterey Bay Aquarium (2000).
561: 461:, there are species with one and sometimes two permanent anuses, species like the 438:. This accompanied or facilitated other important evolutionary developments: the 260: 582:
Helms, Doris R.; Helms, Carl W.; Kosinski, Robert J.; Cummings, John C. (1997).
427:, although the channels flowing to that orifice are almost completely separate. 290:, which (depending on the type of animal) include: indigestible matter such as 328:; ingested matter which would be toxic if it remained in the digestive tract; 825: 819: 500: 203: 778: 478: 435: 355: 351: 302: 252: 109: 541: 496: 536: 505: 711: 549: 492: 474: 462: 458: 451: 439: 431: 416: 392: 332: 309: 272: 102: 97: 81: 45: 770: 484: 420: 412: 404: 396: 372: 362: 347: 336: 329: 321: 276: 66: 703: 641: 359: 317: 746: 667: 465:
grows an anus, which then disappear when it is no longer needed.
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Bowel contents that pass through the anus include the gaseous
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What is a warty comb jelly? | BBC Science Focus Magazine
639: 558: – Medical functional test of the anus and rectum 454:, which are composed of fused, specialized segments. 640:
Chin, K.; Erickson, G.M.; et al. (1998-06-18).
517: 365:, and failure to stop unwanted passages results in 30:"Butthole" redirects here. For the slang word, see 612:A living bay: the underwater world of Monterey Bay 585:Biology in the Laboratory With BioBytes 3.1 CD-ROM 552: – Functional segment of the large intestine 817: 681: 415:mammals have completely separate orifices for 387:) for excreting liquid and solid wastes, for 86:Formation of anus in proto- and deuterostomes 425:another for both urination and reproduction 615:. University of California Press. p.  80: 65: 698:(25). Society for Science &#38: 391. 564: – Birth defect of malformed rectum 499:, the first phase in the growth of the 27:Digestive tract waste expulsion opening 14: 818: 251:, 'ring' or 'circle') is the external 483:In animals at least as complex as an 684:"Getting the scoop from the poop of 383:use a similar orifice (known as the 271:. Its function is to facilitate the 236: 24: 267:), i.e. the opposite end from the 25: 842: 797: 642:"A king-sized theropod coprolite" 320:have been extracted, for example 803: 588:. W. H. Freeman. p. 36-12. 535: 520: 163:Superficial inguinal lymph nodes 740: 729: 682:Monastersky, R. (1998-06-20). 602: 575: 495:, which deepens to become the 491:forms a dent on one side, the 468: 316:material after the digestible 13: 1: 568: 540:The dictionary definition of 275:of wastes that remain after 7: 528:Evolutionary biology portal 513: 10: 847: 472: 43: 36: 29: 411:for reproduction. Female 209: 197: 185: 173: 168: 156: 144: 132: 120: 108: 96: 91: 79: 64: 59: 211:Anatomical terminology 151:Inferior rectal nerves 127:Inferior rectal artery 48:. For other uses, see 50:Anus (disambiguation) 812:at Wikimedia Commons 440:bilaterian body plan 139:Inferior rectal vein 763:2001Natur.409...81A 660:1998Natur.393..680C 556:Anorectal manometry 286:and the semi-solid 367:fecal incontinence 808:Media related to 626:978-0-520-22149-9 595:978-0-7167-3146-7 225: 224: 220: 16:(Redirected from 838: 831:Digestive system 807: 791: 790: 771:10.1038/35051075 744: 738: 733: 727: 726: 724: 723: 714:. Archived from 679: 637: 631: 630: 606: 600: 599: 579: 562:Imperforate anus 539: 530: 525: 524: 463:warty comb jelly 238: 217:edit on Wikidata 214: 84: 69: 57: 56: 21: 846: 845: 841: 840: 839: 837: 836: 835: 816: 815: 800: 795: 794: 757:(6816): 81–85. 745: 741: 734: 730: 721: 719: 704:10.2307/4010364 638: 634: 627: 607: 603: 596: 580: 576: 571: 526: 519: 516: 481: 473:Main articles: 471: 310:digestive rocks 261:digestive tract 221: 87: 75: 53: 42: 35: 28: 23: 22: 15: 12: 11: 5: 844: 834: 833: 828: 814: 813: 799: 798:External links 796: 793: 792: 739: 728: 632: 625: 601: 594: 573: 572: 570: 567: 566: 565: 559: 553: 547: 532: 531: 515: 512: 470: 467: 303:endozoochorous 223: 222: 213: 207: 206: 201: 195: 194: 189: 183: 182: 177: 171: 170: 166: 165: 160: 154: 153: 148: 142: 141: 136: 130: 129: 124: 118: 117: 112: 106: 105: 100: 94: 93: 89: 88: 85: 77: 76: 70: 62: 61: 26: 9: 6: 4: 3: 2: 843: 832: 829: 827: 824: 823: 821: 811: 806: 802: 801: 788: 784: 780: 776: 772: 768: 764: 760: 756: 752: 751: 743: 737: 732: 718:on 2013-05-11 717: 713: 709: 705: 701: 697: 693: 689: 687: 677: 673: 669: 668:10.1038/31461 665: 661: 657: 654:(6686): 680. 653: 649: 648: 643: 636: 628: 622: 618: 614: 613: 605: 597: 591: 587: 586: 578: 574: 563: 560: 557: 554: 551: 548: 546:at Wiktionary 545: 544: 538: 534: 533: 529: 523: 518: 511: 508: 507: 502: 498: 494: 490: 486: 480: 476: 466: 464: 460: 455: 453: 449: 445: 441: 437: 436:deuterostomes 433: 428: 426: 422: 418: 414: 410: 406: 402: 398: 394: 390: 386: 382: 378: 374: 370: 368: 364: 361: 357: 356:endosymbionts 353: 349: 346: 342: 338: 334: 331: 327: 323: 319: 315: 311: 307: 304: 300: 297: 293: 289: 285: 280: 278: 274: 270: 266: 262: 258: 254: 250: 246: 242: 234: 230: 218: 212: 208: 205: 202: 200: 196: 193: 190: 188: 184: 181: 178: 176: 172: 167: 164: 161: 159: 155: 152: 149: 147: 143: 140: 137: 135: 131: 128: 125: 123: 119: 116: 113: 111: 107: 104: 101: 99: 95: 90: 83: 78: 74: 68: 63: 58: 55: 51: 47: 40: 33: 19: 754: 748: 742: 731: 720:. Retrieved 716:the original 695: 692:Science News 691: 685: 651: 645: 635: 611: 604: 584: 577: 542: 504: 482: 479:Deuterostome 459:comb jellies 456: 429: 371: 352:gut bacteria 339:-containing 281: 256: 253:body orifice 244: 240: 232: 226: 192:A05.7.05.013 179: 54: 680:Summary at 497:archenteron 469:Development 432:protostomes 343:; and dead 333:metabolites 312:; residual 259:end of the 169:Identifiers 820:Categories 722:2009-04-24 569:References 550:Anal canal 493:blastopore 475:Protostome 452:arthropods 448:metamerism 417:defecation 405:Marsupials 393:egg-laying 389:copulation 373:Amphibians 363:sphincters 354:and other 350:or excess 115:Alimentary 103:Proctodeum 71:Anus of a 46:Human anus 485:earthworm 421:urination 413:placental 397:Monotreme 348:epithelia 337:bilirubin 322:cellulose 318:nutrients 277:digestion 273:expulsion 98:Precursor 779:11343117 514:See also 401:amniotes 377:reptiles 360:muscular 330:excreted 227:In most 18:Butthole 787:4406268 759:Bibcode 712:4010364 676:4343329 656:Bibcode 345:mucosal 299:pellets 255:at the 247:; from 229:animals 92:Details 32:Asshole 785:  777:  750:Nature 710:  686:T. rex 674:  647:Nature 623:  592:  506:proto– 489:embryo 487:, the 446:, and 444:coelom 442:, the 409:vagina 385:cloaca 326:lignin 284:flatus 241:anuses 231:, the 122:Artery 110:System 39:Wooper 783:S2CID 708:JSTOR 672:S2CID 381:birds 335:like 306:seeds 292:bones 288:feces 269:mouth 265:bowel 249:Latin 215:[ 175:Latin 158:Lymph 146:Nerve 826:Anus 810:Anus 775:PMID 621:ISBN 590:ISBN 543:anus 477:and 434:and 391:and 379:and 341:bile 314:food 308:and 296:hair 257:exit 233:anus 204:3022 187:TA98 180:anus 134:Vein 60:Anus 767:doi 755:409 700:doi 696:153 664:doi 652:393 617:244 501:gut 457:In 324:or 245:ani 243:or 237:pl. 199:TA2 73:dog 822:: 781:. 773:. 765:. 753:. 706:. 694:. 690:. 670:. 662:. 650:. 644:. 619:. 419:, 403:. 395:. 375:, 369:. 301:, 294:, 279:. 239:: 789:. 769:: 761:: 725:. 702:: 688:" 678:. 666:: 658:: 629:. 598:. 263:( 235:( 219:] 52:. 41:. 34:. 20:)

Index

Butthole
Asshole
Wooper
Human anus
Anus (disambiguation)

dog

Precursor
Proctodeum
System
Alimentary
Artery
Inferior rectal artery
Vein
Inferior rectal vein
Nerve
Inferior rectal nerves
Lymph
Superficial inguinal lymph nodes
Latin
TA98
A05.7.05.013
TA2
3022
Anatomical terminology
edit on Wikidata
animals
Latin
body orifice

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