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Barren-ground caribou

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air before it enters the throat and lungs, bez tines set close to the brow tines, distinctive coat patterns, short legs and other adaptations for running long distances, and multiple behaviors suited to tundra, but not to forest (such as synchronized calving and aggregation during rutting and post-calving). As well, many genes, including those for vitamin D metabolism, fat metabolism, retinal development, circadian rhythm, and tolerance to cold temperatures, are found in tundra caribou that are lacking or rudimentary in forest types. For this reason, forest-adapted reindeer and caribou could not survive in tundra or polar deserts, nor could barren-ground or Peary caribou survive (or at least successfully reproduce) in boreal forest, and this explains the rarity of introgression of barren-ground caribou into woodland caribou, and almost none the other way.
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which may still be covered in snow or ice, thereby reducing access to food. Caribou give birth in early spring when insect populations are low, to enable sufficient rearing of healthy and strong calves. Early onset of spring temperatures in the Arctic further effect the phenology of the pregnancy time periods of barren-ground caribou. Insect avoidance forces caribou to expend large amounts of energy through migrational avoidance of insects. Changes in the climate can increase parasitic occurrences, thereby providing an additional threat to the subspecies.
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Beverly and Qamanirjuaq Caribou Management Board. The range of the Beverly herd spans the tundra from northern Manitoba and Saskatchewan and well into the Northwest Territories and Nunavut. In 1994 survey there were 276,000 caribou, an all-time record. According to a 2011 survey based on data collected using cutting-edge digital tools and fly-over visual surveillance, there were approximately 124,000 caribou in the Beverly herd and 83,300 in the Ahiak herd. The calving grounds of the Beverly herd are located around
234: 92: 42: 592:, the caribou's main food source. Frozen feeding grounds during winter months results in greater energy expenditure as the caribou attempt to access the lichen locked beneath the ice. This can result in malnutrition, starvation and death. Research has shown that changes in climate can alter the quality of lichen in the Arctic, making it less nutritious. A changing climate also introduces the threat of foreign plant species to the region, creating competition. 69: 1035:"Ahiak Herd, Baffin Island herds, Bathurst Herd, Beverly Herd, Bluenose-East Herd, Bluenose-West Herd, Cape Bathurst Herd, Dolphin and Union Herd, George River Herd, Leaf River Herd, Lorillard and Wager Bay herds, Peary caribou, Pen Islands and Cape Churchill herds, Porcupine Herd, Qamanirjuaq Herd, Southampton Island Herd, Tuktoyaktuk Peninsula Herd" 380:
order, from Alaska to Hudson Bay, these are the Porcupine herd, Cape Bathurst herd, Bluenose West herd, Bluenose East herd, Bathurst herd, Ahiak herd, Beverly herd, and Qamanirjuaq herd. About 120 000 other barren-ground caribou live in smaller herds that spend the entire year on the tundra. Half of these are confined to Baffin Island.
448:, but the herd shifted its traditional birthing area. Ross Thompson, executive director of the Beverly and Qamanirjuaq Caribou Management Board, explains the low calving rate mainly on habitat deterioration and disturbance with other factors contributing to the low growth rate – parasites, predation and poor weather. 626:
An additional stressor on barren-ground caribou is the irritation from insect behavior, which can dictate the movement and health of caribou during the summer months. Increased warming temperatures and early springs result in greater insect numbers. Insect harassment force caribou to migrate to areas
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Conversely, Molecular data also revealed that the four western Canadian montane ecotypes are not woodland caribou as currently classified (in Canada): they share a common ancestor with modern barren-ground/tundra reindeer and caribou, but distantly, having diverged > 60,000 years ago — before the
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Archaeologists distinguish “modern” tundra reindeer and barren-ground caribou from primitive forms — living and extinct — that did not have adaptations to extreme cold and to long-distance migration. They include a broad, high muzzle to increase the volume of the nasal cavity to warm and moisten the
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The Beverly herd (located primarily in Saskatchewan and the Northwest Territories, with portions in Nunavut, Manitoba and Alberta) and the Qamanirjuaq herd (located primarily in Manitoba and Nunavut, with portions in the southeastern NWT and northeastern Saskatchewan) fall under the auspices of the
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The timing of migration periods is closely linked to seasonal changes and as unpredictable climate conditions increase, barren-ground caribou must migrate over larger distances. Migration is dictated by the access to easily available lichen. An increased distance of migration places further stress
339:, with the females weighing around 90 kg (200 lb) and the males around 150 kg (330 lb). However, on some of the smaller islands, the average weight may be less. The large migratory herds of barren-ground caribou take their names from the traditional calving grounds, such as the 2149:
Yannic G, Pellissier L, Ortego J, Lecomte N, Couturier S, Cuyler C, Dussault C, Hundertmark KJ, Irvine RJ, Jenkins DA, Kolpashikov L, Mager K, Musiani M, Parker KL, Røed KH, Sipko T, Þórisson SG, V.Weckworth B, Guisan A, Bernatchez L, Côté SD (2013) Genetic diversity in caribou linked to past and
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having a negative impact on the population. Changing climate conditions in the Arctic are predicted to threaten barren-ground caribou in the immediate future. The risks associated with climate change can impact feeding habits, access to food and quality of food, birthing rates and calf rearing,
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John Nagy, University of Alberta's wildlife biologist and researcher, argued that the Beverly herd was robust, not declining. He claimed the herd had moved their calving grounds "near the western Queen Maud Gulf coast to the north of the herd's "traditional" calving ground in the Gary Lakes area
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Barren-ground caribou have evolved to match their calving period with the period in which lichen has traditionally bloomed. The phenological process and timing between birthing and easily accessible lichen is critical to the survival rate of the subspecies. The trophic mismatch, due to abnormal
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They spend much or all of the year on the tundra from Alaska to Baffin Island. Most, or about 1.2 million, of the barren-ground caribou in Canada live in eight large migratory herds, which migrate seasonally from the tundra to the taiga, sparsely treed coniferous forests south of the tundra. In
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during the summer, migrate to the mainland in the fall after the sea ice has formed. At this time, the smaller groups may form into a larger herd and several hundred animals may be seen. Mainland barren-ground caribou herds move to coastal areas for part of each year, with the exception of the
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Bondarev AA, Tesakov AS, Simakova AN, Dorogov AL (2017) Reindeer (Rangifer) from Early Pleistocene of the south of Western Siberia (in Russian). In: Bogdanov AA, others (Eds) Integrative palaeontology: development prospects for geological objectives, material of the LXIII session of the
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The barren-ground caribou usually breeds in the fall and calves in June but may not drop their single calf until July. Usually the female gives birth away from the herd and if possible on a patch of snow. After birth, the female licks the calf clean and eats the tissues and the
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On the mainland of Canada, the animals may travel in herds of several thousand, but they move in smaller groups (no more than 50) on the islands. They are migratory animals and may travel 1,200 km (750 mi) in a season. Some groups, such as those living on
685:: Evolution). Before Banfield (1961), taxonomists using cranial, dental and skeletal measurements had unequivocally allied these western montane ecotypes with barren-ground caribou, naming them (as in Osgood 1909 Murie, 1935 and Anderson 1946, among others) 677:, had features (robust and short pedicles, smooth antler surface, and high position of second tine) that relate it to forest caribou. Because woodland and barren-ground caribou likely do not share a direct common ancestor, they cannot be conspecific. 2112:
Cavedon M, Gubili C, Heppenheimer E, vonHoldt B, Mariani S, Hebblewhite M, Hegel T, Hervieux D, Serrouya R, Steenweg R (2019) Genomics, environment and balancing selection in behaviourally bimodal populations: The caribou case. Molecular Ecology 28:
745:(Millais 1915), after the Ogilvie Mountains that form part of its winter range. Youngman (1975) realized Banfield's mistake and referred the barren-ground caribou of the Porcupine herd (and by implication, to other Alaskan barren-ground herds) to 452:
Most of the caribou populations in the north are cycling down. It's causing a lot of anxiety for a lot of hunters. We want to...give everybody time to work together to come up with solutions for the short term and until the caribou populations
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Prunier J, Carrier A, Gilbert I, Poisson W, Albert V, Taillon J, Bourret V, CĂ´tĂŠ SD, Droit A, Robert C (2021) CNVs with adaptive potential in Rangifer tarandus: genome architecture and new annotated assembly. Life Science Alliance 5: 1-16.
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from their summer grazing on Victoria Island to their winter grazing area on the Nunavut-NWT mainland in Canada. It is unusual for North American caribou to cross sea ice seasonally and the only other caribou subspecies to do so is the
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Climate change negatively impacts barren-ground caribou's access to food. Extreme weather conditions can cause increased amounts of rain and freezing rain during winter months. This results in an ice layer which blocks access to
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In consideration of the voluminous genetic and other data revealing diversity at the species and subspecies levels that is not reflected in the current taxonomy, a recent revision resurrects several species (Greenland caribou,
400:. In general, during the summer, the coat of the caribou is brown, and much lighter in the winter. The neck and rump tend towards a creamy-white colour. However, the general coloration may differ depending on the region. 2168:
Horn R, Marques AJD, Manseau M, Golding B, KlĂźtsch CFC, Abraham K, Wilson PJ (2018) Parallel evolution of site-specific changes in divergent caribou lineages. Ecology and Evolution 8: 6053-6064. doi:10.1002/ece3.4154.
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Cavedon M, vonHoldt B, Hebblewhite M, Hegel T, Heppenheimer E, Hervieux D, Mariani S, Schwantje H, Steenweg R, Theoret J (2022) Genomic legacy of migration in endangered caribou. PLoS genetics 18: e1009974.
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Millais JG (1915) The caribou. In: Carruthers D, Millais JG, Byl PBVD, Wallace HF, Kennion L-CRL, Barklay FG (Eds) The Gun at Home and Abroad. London & Counties Press Association Ltd., London, U.K.,
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From 1898 to 1937, mammalogists named 11 new species (other than Greenland caribou, barren-ground caribou and woodland caribou, which had been named earlier) in Canada and Alaska (see synonymies in
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Bastille-Rousseau, Guillaume; Schaefer, James A.; Lewis, Keith P.; Mumma, Matthew A.; Ellington, E. Hance; Rayl, Nathaniel D.; Mahoney, Shane P.; Pouliot, Darren; Murray, Dennis L. (2016-03-01).
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north of Baker Lake." He based his findings on data collected from 510 barren-ground caribou tracked with satellite collars in the Northwest Territories and Nunavut from 1993 to 2009.
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and energy expense on the caribou. Warming weather conditions reduce ice thickness over rivers and lakes, making it difficult for caribou to cross. The reduced ice cover creates a
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Cronin MA, MacNeil MD, Patton JC (2005) Variation in mitochondrial DNA and microsatellite DNA in caribou (Rangifer tarandus) in North America. Journal of Mammalogy 86: 495-505.
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Croitor R (2018) Plio-Pleistocene deer of Western Palearctic: taxonomy, systematics, phylogeny. Institute of Zoology of the Academy of Sciences of Moldova, Chișinău, 142 pp.
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Mallory, Conor D.; Campbell, Mitch W.; Boyce, Mark S. (2018-01-03). "Climate influences body condition and synchrony of barren-ground caribou abundance in Northern Canada".
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Allen 1902) of the west end of the Alaska Peninsula and nearby islands to all of Alaska and part of Yukon, including the Porcupine herd, which was originally described as
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temperature variations linked to climate change, have resulted in malnutrition in their young, as well as reduced reproductive rates contributing to population decline.
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Banfield AWF (1961) A revision of the reindeer and caribou, genus Rangifer. National Museum of Canada Bulletin 177, Biological Series No. 66, Ottawa, Ontario, 137 pp.
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Harding LE (2022) Available names for Rangifer (Mammalia, Artiodactyla, Cervidae) species and subspecies. ZooKeys 1119: 117-151. doi:10.3897/zookeys.1119.80233.
987: 673:: Evolution). Woodland caribou are likely more related to extinct forest caribou subspecies than to barren-ground caribou. For example, the extinct caribou 642:
originated Late Pliocene and diversified in the Early Pleistocene, a 2+ million-year period of multiple glacier advances and retreats. The oldest undoubted
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Osgood WH (1909) Biological investigations in Alaska and Yukon Territory. US Department of Agriculture Biological survey of North American fauna 1: 1-285.
1383: 1182:"Beverly caribou have been on the move, wildlife biologist says: The herd, now calving on the Queen Maud Gulf coast, is "robust" and not near extinction" 2207:
Anderson RM (1946) Catalogue of Canadian Recent mammals. National Museum of Canada Bulletin No. 102, Biological Series 31, Ottawa, Ontario, 238 pp.
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Zamin, Tara J.; CĂ´tĂŠ, Steeve D.; Tremblay, Jean-Pierre; Grogan, Paul (2017-10-01). "Experimental warming alters migratory caribou forage quality".
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Flagstad O, Røed KH (2003) Refugial origins of reindeer (Rangifer tarandus L.) inferred from mitochondrial DNA sequences. Evolution 57: 658-670.
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Murie OJ (1935) Alaska-Yukon caribou. Vol. 54, United States Department of Agriculture Bureau of Biological Survey, Washington D.C., 93 pp.
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modern barren-ground ecotype had evolved its cold- and darkness-adapted physiologies and mass-migration and aggregation behaviors, (see
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Geist V (1998) Deer of the world: their evolution, behavior, and ecology. Stackpole Books, Mechanicsburg, Pennsylvania, 421 pp.
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Allen JA (1902) A new caribou from the Alaska Peninsula. Bulletin of the American Museum of Natural History XVI: 119-127.
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Approximate range of barren-ground caribou. Overlap with other subspecies of caribou is possible for contiguous range. 1.
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Asplund, Johan; Wardle, David A. (2017-08-01). "How lichens impact on terrestrial community and ecosystem properties".
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Youngman PM (1975) Mammals of the Yukon Territory. National Museum of Canada, Publications in Zoology No. 10, 192 pp.
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Røed KH (2005) Refugial origin and postglacial colonization of holarctic reindeer and caribou. Rangifer 25: 19-30.
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Nicholson, Kerry L.; Arthur, Stephen M.; Horne, Jon S.; Garton, Edward O.; Vecchio, Patricia A. Del (2016-04-05).
984: 574:(SARA) registry placed barren-ground caribou under the status of "special concern". Their status was a result of 254: 76: 2516: 1641:"Phase-dependent climate–predator interactions explain three decades of variation in neonatal caribou survival" 2311: 1391: 2536: 2083:
Harington CR (2011) Pleistocene vertebrates of the Yukon Territory. Quaternary Science Reviews 30: 2341-2354.
842: 2521: 2452: 1344:"Sea Ice and Migration of the Dolphin and Union Caribou Herd in the Canadian Arctic: An Uncertain Future" 1156:"Researchers investigate decline of Beverly caribou herd: Population dropped by half over 20-year period" 541: 1054: 2526: 250: 930:
Variation in Mitochondrial DNA and Microsatellite DNA in caribou (Rangifer tarandus) in North America.
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Sara Minogue. Mining exploration on Nunavut's Bluenose East caribou calving grounds allowed by board
545: 537: 436: 2470: 2306: 887: 710: 336: 246: 2340: 1711:"Weather conditions and variation in timing of spring and fall migrations of migratory caribou" 352: 210: 1988:"Modeling Caribou Movements: Seasonal Ranges and Migration Routes of the Central Arctic Herd" 523: 320: 335:. The barren-ground caribou is a medium-sized caribou, smaller and lighter-colored than the 2400: 2290: 1999: 1866: 1782: 1652: 1588: 1524: 713:, 1946). Banfield (1961) synonymized them down to one species with four living subspecies ( 8: 1343: 484: 58: 2003: 1870: 1786: 1656: 1592: 1528: 1262:, Government of Northwest Territories, Department of Environment and Natural Resources, 2030: 1987: 1963: 1897: 1854: 1813: 1770: 1612: 1006:: Nunavut, N.W.T. governments opposed project, but it went ahead. CBC News Mar 04, 2016 86: 579:
greater distance of migration, thinning ice during migration and insect disturbances.
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future climate change. Nature Climate Change 4: 132-137. doi:10.1038/NCLIMATE2074.
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fossil is from Omsk, Russia) dated to 2.1-1.8 Ma.  The oldest North American
2273: 2012: 1879: 1795: 1453:"Observed and predicted effects of climate change on Arctic caribou and reindeer" 991: 638:“evolved as a mountain deer, ...exploiting the subalpine and alpine meadows...”. 611: 571: 496: 445: 360: 340: 1018:
Technical Thematic Report No. 10. - Northern caribou population trends in Canada
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fossil is from the Yukon, 1.6 million years before present (BP). Several named
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and the nearby mainland. They are endemic to Canada. They migrate across the
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In Canada about fifty percent of all caribou are barren-ground caribou.
2418: 721:), but most specialists did not accept this and continued to recognize 416: 300: 2431: 1933: 1479: 701:, respectively, and this phylogeny was confirmed by genetic analysis. 2159:
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Palaeontological Society, April, 3-7, 2017. Sankt-Peterburg, 173-175.
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Le Corre, Mael; Dussault, Christian; CĂ´tĂŠ, Steeve D. (2017-02-08).
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and North America predate the evolution of modern tundra reindeer.
420: 405: 304: 187: 173: 123: 1298:"COSEWIC Assessment and Update Status Report on the Peary Caribou 752:
Current taxonomy recognizes just one species of reindeer/caribou:
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communities who depend on caribou as a source of income and food.
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Mammal Species of the World: A Taxonomic and Geographic Reference
655: 424: 316: 2405: 1384:"HTML Version of the Document - Species at Risk Public Registry" 1110:
Beverly caribou decline not as drastic as once feared: new study
900:(3rd ed.). Baltimore, MD: Johns Hopkins University Press. 589: 412: 397: 332: 312: 133: 113: 1048:"The Beverly Caribou Herd: Continental Wilderness Travellers" 488: 1852: 1769:
Joly, Kyle; Wasser, Samuel K.; Booth, Rebecca (2015-06-10).
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The barren-ground caribou, one of several subspecies called
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Harper, Francis (21 October 1955), Hall, E. Raymond (ed.),
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Beverly and Qamanirjuaq Caribou Management Board website
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Terrestrial Mammals of Nunavut by Ingrid Anand-Wheeler.
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Canada, Government of Canada, Environment (2011-04-27).
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Terrestrial Mammals of Nunavut by Ingrid Anand-Wheeler.
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Cronin, M. A., M. D. Macneil, and J. C. Patton (2005).
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The Beverly Caribou Herd’s headlong rush to extinction
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The Dolphin-Union caribou herd, locally known as the
1708: 1578: 1341: 506:The major predator of barren-ground caribou is the 1222: 1256:Species at Risk in the Northwest Territories 2012 2498: 2322:Map of the Beverly and Qamanirjuaq caribou range 1451:Mallory, Conor D.; Boyce, Mark S. (2017-06-28). 1768: 733:). Banfield (1961) had extended the tiny, pale 1131:"Another caribou herd in steep decline: study" 1919: 557:), which are smaller in size and population. 517: 1450: 1411:: CS1 maint: multiple names: authors list ( 1102: 1100: 940: 938: 828:) in North America) and several subspecies. 1021:, Biodiversity Canada, 2010, archived from 725:(Labrador caribou), Newfoundland caribou ( 665:DNA analysis shows that woodland caribou ( 419:(sedges) and grasses along with twigs and 323:, in northern Alaska and in south-western 232: 67: 40: 2029: 2011: 1941: 1896: 1878: 1812: 1794: 1726: 1664: 1544: 1478: 1468: 1097: 935: 2330:, a documentary on Barren-ground caribou 944: 560: 1446: 1444: 1442: 1306:Dolphin and Union population in Canada" 1290: 838:Caribou herds and populations in Canada 604: 458:2013, Regional biologist Mitch Campbell 370: 14: 2499: 2288: 1981: 1979: 1977: 1848: 1846: 1844: 1842: 1840: 1838: 1836: 1834: 1832: 1764: 1762: 1760: 1758: 1756: 1754: 1752: 1750: 1748: 1746: 1704: 1702: 1700: 1698: 1696: 1694: 1692: 1634: 1632: 1630: 1628: 1626: 1510: 1508: 1506: 1504: 1502: 1500: 1498: 1440: 1438: 1436: 1434: 1432: 1430: 1428: 1426: 1424: 1422: 1381: 1248: 1246: 1179: 1173: 2507:NatureServe apparently secure species 2339: 2338: 2176: 2174: 2060: 2058: 1574: 1572: 1220: 1106: 922: 886: 245:, which is subdivided into ecotypes: 1335: 1082:. arctic-caribou.com. Archived from 932:Journal of Mammalogy 86(3): 495–505. 621: 49:Barren-ground caribou herd near the 1974: 1829: 1743: 1689: 1623: 1495: 1419: 1243: 880: 756:. Subspecies in North America are, 24: 2171: 2055: 1569: 1388:www.registrelep-sararegistry.gc.ca 570:In 2004 the Canadian Government's 396:, both the males and females have 25: 2548: 2300: 1180:George, Jane (30 November 2011), 595: 565: 487:, is a major food source for the 1252: 945:Kavanagh, Maureen, ed. (2005) , 90: 2379:Rangifer tarandus groenlandicus 2349:Rangifer tarandus groenlandicus 2247: 2238: 2229: 2219: 2210: 2201: 2192: 2183: 2162: 2153: 2143: 2134: 2125: 2116: 2106: 2096: 2086: 2077: 2067: 2046: 1913: 1375: 1304:Rangifer tarandus groenlandicus 1214: 1198: 1148: 1123: 582: 534:Rangifer tarandus groenlandicus 359:), the Bluenose West herd, the 293:Rangifer tarandus groenlandicus 220:Rangifer tarandus groenlandicus 2093:doi:10.1101/2021.07.22.453386. 1107:Varga, Peter (14 March 2013), 1072: 1040: 1009: 997: 978: 855: 715:caribou, pearyi, groenlandicus 13: 1: 2260: 843:Reindeer hunting in Greenland 2013:10.1371/journal.pone.0150333 1880:10.1371/journal.pone.0172669 1796:10.1371/journal.pone.0127586 951:Canadian Wildlife Service/EC 848: 630: 7: 1053:. taiga.net. Archived from 994:February 22, 2016 Don Jaque 831: 704: 10: 2553: 2328:Caribou of Northern Canada 1302:and Barren-ground Caribou 863:"NatureServe Explorer 2.0" 521: 518:Dolphin-Union caribou herd 503:) of present-day Nunavut. 253:and mountain woodland; 2. 2347: 1645:Journal of Animal Ecology 1601:10.1007/s00300-017-2248-3 297:recent taxonomic revision 243:Rangifer tarandus caribou 240: 231: 216: 209: 87:Scientific classification 85: 65: 56: 48: 39: 34: 1300:Rangifer tarandus pearyi 894:; Reeder, D. M. (eds.). 867:explorer.natureserve.org 546:Dolphin and Union Strait 415:, but they also feed on 411:The main food source is 202:R. t. groenlandicus 2532:Endemic fauna of Canada 1666:10.1111/1365-2656.12466 1517:Ecological Applications 711:Rudolph Martin Anderson 675:Torontoceros hypogaeus 337:boreal woodland caribou 311:) that is found in the 2103:doi:10.1111/mec.15039. 1728:10.1093/jmammal/gyw177 1221:Mowat, Farley (2000). 947:"Hinterland Who's Who" 824:; and Arctic caribou, 729:and Osborn's caribou ( 461: 390: 35:Barren-ground caribou 2517:Mammals of the Arctic 2307:Barren-ground Caribou 1457:Environmental Reviews 561:Environmental threats 524:Dolphin-Union caribou 450: 377: 321:Northwest Territories 289:barren-ground caribou 18:Barren-ground Caribou 2537:Mammals of Greenland 2312:Hinterland Who's Who 1715:Journal of Mammalogy 1470:10.1139/er-2017-0032 1229:. Steerforth Press. 820:; woodland caribou, 605:Effects on migration 495:bands living in the 386:Hinterland Who's Who 371:Range and population 2522:Arctic land animals 2292:Caribou of Keewatin 2004:2016PLoSO..1150333N 1871:2017PLoSO..1272669R 1787:2015PLoSO..1027586J 1657:2016JAnEc..85..445B 1593:2018PoBio..41..855M 1529:2017EcoAp..27.2061Z 1363:10.14430/arctic3331 1225:Walking on the Land 1080:"The caribou herds" 967:on 24 December 2013 810:R. t. platyrhynchus 770:R. t. groenlandicus 747:R. t. groenlandicus 485:Inuktitut syllabics 345:Baffin Island herds 307:(or the caribou in 295:; but subject to a 274:R. t. groenlandicus 269:R. t. groenlandicus 259:(extinct 1908); 3. 59:Conservation status 2272:2007-09-29 at the 1922:Biological Reviews 1546:20.500.11794/17064 990:2020-12-19 at the 784:; and in Eurasia, 512:Canis lupus arctos 327:. It includes the 251:migratory woodland 73:Apparently Secure 27:Subspecies of deer 2527:Mammals of Canada 2494: 2493: 2341:Taxon identifiers 1934:10.1111/brv.12305 1322:978-0-662-37375-9 1269:978-0-7708-0196-0 1137:, 1 December 2008 1025:on August 7, 2018 960:978-0-662-39659-8 754:Rangifer tarandus 727:R. t. terranovae) 622:Insect harassment 491:, especially the 479:, and written as 329:Porcupine caribou 285: 284: 80: 16:(Redirected from 2544: 2487: 2486: 2474: 2473: 2461: 2460: 2448: 2447: 2435: 2434: 2422: 2421: 2409: 2408: 2396: 2395: 2383: 2382: 2381: 2368: 2367: 2366: 2336: 2335: 2296: 2254: 2251: 2245: 2242: 2236: 2233: 2227: 2223: 2217: 2214: 2208: 2205: 2199: 2196: 2190: 2187: 2181: 2178: 2169: 2166: 2160: 2157: 2151: 2147: 2141: 2138: 2132: 2129: 2123: 2120: 2114: 2110: 2104: 2100: 2094: 2090: 2084: 2081: 2075: 2071: 2065: 2062: 2053: 2050: 2044: 2043: 2033: 2015: 1983: 1972: 1971: 1945: 1928:(3): 1720–1738. 1917: 1911: 1910: 1900: 1882: 1850: 1827: 1826: 1816: 1798: 1766: 1741: 1740: 1730: 1706: 1687: 1686: 1668: 1636: 1621: 1620: 1576: 1567: 1566: 1548: 1537:10.1002/eap.1590 1523:(7): 2061–2073. 1512: 1493: 1492: 1482: 1472: 1448: 1417: 1416: 1410: 1402: 1400: 1399: 1390:. Archived from 1379: 1373: 1372: 1371: 1369: 1348: 1339: 1333: 1332: 1331: 1329: 1310: 1294: 1288: 1287: 1286: 1284: 1278: 1272:, archived from 1261: 1250: 1241: 1240: 1228: 1218: 1212: 1202: 1196: 1195: 1194: 1192: 1177: 1171: 1170: 1169: 1167: 1152: 1146: 1145: 1144: 1142: 1127: 1121: 1120: 1119: 1117: 1104: 1095: 1094: 1092: 1091: 1076: 1070: 1069: 1067: 1065: 1059: 1052: 1044: 1038: 1037: 1032: 1030: 1013: 1007: 1001: 995: 982: 976: 975: 974: 972: 963:, archived from 942: 933: 926: 920: 919: 884: 878: 877: 875: 873: 859: 818:R. groenlandicus 798:R. t. phylarchus 794:R. t. valentinae 782:R. t. terranovae 459: 388: 365:Qamanirjuaq herd 236: 226:(Borowski, 1780) 222: 188:R. tarandus 95: 94: 74: 71: 70: 44: 32: 31: 21: 2552: 2551: 2547: 2546: 2545: 2543: 2542: 2541: 2497: 2496: 2495: 2490: 2482: 2477: 2469: 2464: 2456: 2451: 2443: 2438: 2430: 2425: 2417: 2412: 2404: 2399: 2391: 2386: 2377: 2376: 2371: 2362: 2361: 2356: 2343: 2303: 2274:Wayback Machine 2263: 2258: 2257: 2252: 2248: 2243: 2239: 2234: 2230: 2224: 2220: 2215: 2211: 2206: 2202: 2197: 2193: 2188: 2184: 2179: 2172: 2167: 2163: 2158: 2154: 2148: 2144: 2139: 2135: 2130: 2126: 2121: 2117: 2111: 2107: 2101: 2097: 2091: 2087: 2082: 2078: 2072: 2068: 2063: 2056: 2051: 2047: 1998:(4): e0150333. 1984: 1975: 1918: 1914: 1865:(3): e0172669. 1851: 1830: 1781:(6): e0127586. 1767: 1744: 1707: 1690: 1637: 1624: 1577: 1570: 1513: 1496: 1449: 1420: 1404: 1403: 1397: 1395: 1380: 1376: 1367: 1365: 1346: 1340: 1336: 1327: 1325: 1323: 1308: 1296: 1295: 1291: 1282: 1280: 1276: 1270: 1259: 1251: 1244: 1237: 1219: 1215: 1203: 1199: 1190: 1188: 1178: 1174: 1165: 1163: 1154: 1153: 1149: 1140: 1138: 1129: 1128: 1124: 1115: 1113: 1105: 1098: 1089: 1087: 1078: 1077: 1073: 1063: 1061: 1057: 1050: 1046: 1045: 1041: 1028: 1026: 1015: 1014: 1010: 1002: 998: 992:Wayback Machine 983: 979: 970: 968: 961: 943: 936: 927: 923: 908: 885: 881: 871: 869: 861: 860: 856: 851: 834: 806:R. t. sibiricus 790:R. t. buskensis 739:Rangifer granti 707: 695:R. a. fortidens 633: 624: 612:natural barrier 607: 598: 585: 572:Species at Risk 568: 563: 538:Victoria Island 526: 520: 497:Kivalliq Region 481:ᓇᐹᕐᑐᕐᑲᓐᖏᑦᑐᒥ ᑐᒃᑐ 460: 457: 446:Queen Maud Gulf 437:Victoria Island 431:and bird eggs. 427:, fish such as 389: 384: 373: 315:territories of 247:boreal woodland 227: 224: 218: 205: 191: 89: 81: 72: 68: 61: 28: 23: 22: 15: 12: 11: 5: 2550: 2540: 2539: 2534: 2529: 2524: 2519: 2514: 2509: 2492: 2491: 2489: 2488: 2475: 2462: 2449: 2436: 2423: 2410: 2397: 2384: 2369: 2353: 2351: 2345: 2344: 2333: 2332: 2324: 2319: 2314: 2309: 2302: 2301:External links 2299: 2298: 2297: 2286: 2276: 2262: 2259: 2256: 2255: 2246: 2237: 2228: 2218: 2209: 2200: 2191: 2182: 2170: 2161: 2152: 2142: 2133: 2124: 2115: 2105: 2095: 2085: 2076: 2066: 2054: 2045: 1973: 1912: 1828: 1742: 1688: 1651:(2): 445–456. 1622: 1587:(5): 855–864. 1568: 1494: 1418: 1374: 1357:(4): 414–428, 1334: 1321: 1289: 1268: 1242: 1236:978-1586420246 1235: 1213: 1197: 1186:Nunatsiaq News 1172: 1162:, 20 June 2013 1147: 1122: 1096: 1071: 1060:on 9 June 2013 1039: 1008: 996: 977: 959: 934: 921: 906: 879: 853: 852: 850: 847: 846: 845: 840: 833: 830: 802:R. t. pearsoni 786:R. t. tarandus 743:R. ogilviensis 706: 703: 691:R. a. montanus 632: 629: 623: 620: 606: 603: 597: 596:Calving season 594: 584: 581: 576:climate change 567: 566:Climate change 564: 562: 559: 536:) that occupy 530:island caribou 522:Main article: 519: 516: 455: 440:Beverly herd. 382: 372: 369: 361:Porcupine herd 283: 282: 262:R. t. arcticus 238: 237: 229: 228: 225: 214: 213: 211:Trinomial name 207: 206: 199: 197: 193: 192: 185: 183: 179: 178: 171: 167: 166: 161: 157: 156: 151: 147: 146: 141: 137: 136: 131: 127: 126: 121: 117: 116: 111: 107: 106: 101: 97: 96: 83: 82: 66: 63: 62: 57: 54: 53: 46: 45: 37: 36: 26: 9: 6: 4: 3: 2: 2549: 2538: 2535: 2533: 2530: 2528: 2525: 2523: 2520: 2518: 2515: 2513: 2510: 2508: 2505: 2504: 2502: 2485: 2480: 2476: 2472: 2467: 2463: 2459: 2454: 2450: 2446: 2441: 2437: 2433: 2428: 2424: 2420: 2415: 2411: 2407: 2402: 2398: 2394: 2389: 2385: 2380: 2374: 2370: 2365: 2359: 2355: 2354: 2352: 2350: 2346: 2342: 2337: 2331: 2329: 2325: 2323: 2320: 2318: 2315: 2313: 2310: 2308: 2305: 2304: 2294: 2293: 2287: 2285: 2284:1-55325-035-4 2281: 2277: 2275: 2271: 2268: 2265: 2264: 2250: 2241: 2232: 2222: 2213: 2204: 2195: 2186: 2177: 2175: 2165: 2156: 2146: 2137: 2128: 2119: 2109: 2099: 2089: 2080: 2070: 2061: 2059: 2049: 2041: 2037: 2032: 2027: 2023: 2019: 2014: 2009: 2005: 2001: 1997: 1993: 1989: 1982: 1980: 1978: 1969: 1965: 1961: 1957: 1953: 1949: 1944: 1943:11250/2578209 1939: 1935: 1931: 1927: 1923: 1916: 1908: 1904: 1899: 1894: 1890: 1886: 1881: 1876: 1872: 1868: 1864: 1860: 1856: 1849: 1847: 1845: 1843: 1841: 1839: 1837: 1835: 1833: 1824: 1820: 1815: 1810: 1806: 1802: 1797: 1792: 1788: 1784: 1780: 1776: 1772: 1765: 1763: 1761: 1759: 1757: 1755: 1753: 1751: 1749: 1747: 1738: 1734: 1729: 1724: 1721:(1): gyw177. 1720: 1716: 1712: 1705: 1703: 1701: 1699: 1697: 1695: 1693: 1684: 1680: 1676: 1672: 1667: 1662: 1658: 1654: 1650: 1646: 1642: 1635: 1633: 1631: 1629: 1627: 1618: 1614: 1610: 1606: 1602: 1598: 1594: 1590: 1586: 1582: 1581:Polar Biology 1575: 1573: 1564: 1560: 1556: 1552: 1547: 1542: 1538: 1534: 1530: 1526: 1522: 1518: 1511: 1509: 1507: 1505: 1503: 1501: 1499: 1490: 1486: 1481: 1476: 1471: 1466: 1462: 1458: 1454: 1447: 1445: 1443: 1441: 1439: 1437: 1435: 1433: 1431: 1429: 1427: 1425: 1423: 1414: 1408: 1394:on 2018-02-07 1393: 1389: 1385: 1378: 1364: 1360: 1356: 1352: 1345: 1338: 1324: 1318: 1314: 1307: 1305: 1301: 1293: 1279:on 2015-09-24 1275: 1271: 1265: 1258: 1257: 1249: 1247: 1238: 1232: 1227: 1226: 1217: 1211: 1210:1-55325-035-4 1207: 1201: 1187: 1183: 1176: 1161: 1157: 1151: 1136: 1132: 1126: 1112: 1111: 1103: 1101: 1086:on 2012-01-06 1085: 1081: 1075: 1056: 1049: 1043: 1036: 1024: 1020: 1019: 1012: 1005: 1000: 993: 989: 986: 981: 966: 962: 956: 952: 948: 941: 939: 931: 925: 917: 913: 909: 907:0-8018-8221-4 903: 899: 898: 893: 892:Wilson, D. E. 889: 883: 868: 864: 858: 854: 844: 841: 839: 836: 835: 829: 827: 823: 819: 813: 811: 807: 803: 799: 795: 791: 787: 783: 779: 775: 774:R. t. osborni 771: 767: 766:R. t. dawsoni 763: 762:R. t. caribou 759: 755: 750: 748: 744: 740: 736: 732: 731:R. t. osborni 728: 724: 720: 716: 712: 702: 700: 699:R. a. osborni 696: 692: 688: 684: 678: 676: 672: 668: 663: 659: 657: 653: 649: 645: 641: 637: 628: 619: 617: 613: 602: 593: 591: 580: 577: 573: 558: 556: 552: 551:Peary caribou 547: 543: 539: 535: 531: 525: 515: 513: 509: 504: 502: 498: 494: 493:Caribou Inuit 490: 486: 482: 478: 474: 470: 465: 454: 449: 447: 441: 438: 432: 430: 426: 422: 418: 414: 409: 407: 401: 399: 395: 394:Peary caribou 387: 381: 376: 368: 366: 362: 358: 354: 350: 349:Bathurst herd 346: 342: 338: 334: 331:of Yukon and 330: 326: 322: 318: 314: 310: 309:North America 306: 302: 298: 294: 290: 281: 280: 275: 271: 270: 264: 263: 258: 257: 256:R. t. dawsoni 252: 248: 244: 239: 235: 230: 223: 221: 215: 212: 208: 204: 203: 198: 195: 194: 190: 189: 184: 181: 180: 177: 176: 172: 169: 168: 165: 162: 159: 158: 155: 152: 149: 148: 145: 142: 139: 138: 135: 132: 129: 128: 125: 122: 119: 118: 115: 112: 109: 108: 105: 102: 99: 98: 93: 88: 84: 78: 64: 60: 55: 52: 47: 43: 38: 33: 30: 19: 2348: 2327: 2291: 2249: 2240: 2231: 2221: 2212: 2203: 2194: 2185: 2164: 2155: 2145: 2136: 2127: 2118: 2108: 2098: 2088: 2079: 2069: 2048: 1995: 1991: 1925: 1921: 1915: 1862: 1858: 1778: 1774: 1718: 1714: 1648: 1644: 1584: 1580: 1520: 1516: 1460: 1456: 1396:. Retrieved 1392:the original 1387: 1377: 1366:, retrieved 1354: 1350: 1337: 1326:, retrieved 1315:, May 2004, 1312: 1303: 1299: 1292: 1281:, retrieved 1274:the original 1255: 1224: 1216: 1200: 1189:, retrieved 1185: 1175: 1164:, retrieved 1159: 1150: 1139:, retrieved 1134: 1125: 1114:, retrieved 1109: 1088:. Retrieved 1084:the original 1074: 1062:. Retrieved 1055:the original 1042: 1034: 1027:, retrieved 1023:the original 1017: 1011: 999: 980: 969:, retrieved 965:the original 950: 929: 924: 896: 882: 870:. Retrieved 866: 857: 825: 821: 817: 814: 809: 805: 801: 797: 796:in Europe), 793: 789: 785: 781: 778:R. t. pearyi 777: 773: 769: 765: 761: 758:R. t. caboti 757: 753: 751: 746: 742: 738: 737:(originally 734: 730: 726: 723:R. t. caboti 722: 718: 714: 708: 698: 694: 690: 687:R. a. stonei 686: 679: 674: 666: 664: 660: 651: 647: 643: 639: 635: 634: 625: 608: 599: 586: 583:Food sources 569: 555:R. t. pearyi 554: 540:in Canada's 533: 529: 527: 511: 505: 501:Barren Lands 500: 480: 468: 466: 462: 451: 442: 433: 410: 402: 391: 385: 378: 374: 353:Beverly herd 292: 288: 286: 279:R. t. pearyi 278: 273: 268: 266: 261: 255: 242: 219: 217: 201: 200: 196:Subspecies: 186: 174: 144:Artiodactyla 51:Thelon River 29: 2466:NatureServe 2427:iNaturalist 2373:Wikispecies 2267:NatureServe 1191:27 December 1166:27 December 1141:27 December 1116:27 December 1029:October 30, 971:21 December 826:R. arcticus 654:fossils in 542:High Arctic 508:Arctic wolf 473:Inuinnaqtun 429:Arctic char 164:Capreolinae 160:Subfamily: 77:NatureServe 2501:Categories 2261:References 1480:1807/79796 1398:2018-02-09 1368:31 October 1328:1 November 1283:31 October 1090:2008-01-16 1064:16 January 822:R. caribou 667:R. caribou 616:Indigenous 417:Cyperaceae 341:Ahiak herd 301:subspecies 2113:1946-1963 2022:1932-6203 1952:1469-185X 1889:1932-6203 1805:1932-6203 1737:0022-2372 1675:1365-2656 1609:0722-4060 1555:1939-5582 1489:1181-8700 1463:: 13–25. 888:Grubb, P. 849:Footnotes 631:Evolution 477:Inuktitut 421:mushrooms 392:Like the 357:Kugluktuk 325:Greenland 182:Species: 110:Kingdom: 104:Eukaryota 2512:Reindeer 2471:2.737205 2458:14200335 2364:Q1757636 2358:Wikidata 2270:Archived 2226:255-280. 2040:27045587 1992:PLOS ONE 1968:25453156 1960:27730713 1907:28328953 1859:PLOS ONE 1823:26061003 1775:PLOS ONE 1683:26529139 1617:13593141 1563:28653471 1407:cite web 1160:CBC News 988:Archived 916:62265494 890:(2005). 872:30 March 832:See also 792:(called 705:Taxonomy 683:Reindeer 671:Reindeer 652:Rangifer 648:Rangifer 644:Rangifer 640:Rangifer 636:Rangifer 456:—  453:recover. 425:lemmings 406:placenta 383:—  363:and the 319:and the 313:Canadian 305:reindeer 175:Rangifer 154:Cervidae 150:Family: 134:Mammalia 124:Chordata 120:Phylum: 114:Animalia 100:Domain: 2419:5220116 2406:1268552 2031:4821536 2000:Bibcode 1898:5362199 1867:Bibcode 1814:4464525 1783:Bibcode 1653:Bibcode 1589:Bibcode 1525:Bibcode 1313:COSEWIC 656:Eurasia 398:antlers 317:Nunavut 303:of the 299:) is a 170:Genus: 140:Order: 130:Class: 75: ( 2445:898935 2432:238268 2282:  2038:  2028:  2020:  1966:  1958:  1950:  1905:  1895:  1887:  1821:  1811:  1803:  1735:  1681:  1673:  1615:  1607:  1561:  1553:  1487:  1351:Arctic 1319:  1266:  1253:GNWT, 1233:  1208:  957:  914:  904:  780:, and 735:granti 719:granti 590:lichen 413:lichen 351:, the 347:, the 343:, the 333:Alaska 2484:86328 2393:7L2T7 1964:S2CID 1613:S2CID 1347:(PDF) 1309:(PDF) 1277:(PDF) 1260:(PDF) 1058:(PDF) 1051:(PDF) 489:Inuit 469:tuktu 276:; 6. 272:; 5. 2479:NCBI 2440:ITIS 2414:GBIF 2280:ISBN 2036:PMID 2018:ISSN 1956:PMID 1948:ISSN 1903:PMID 1885:ISSN 1819:PMID 1801:ISSN 1733:ISSN 1679:PMID 1671:ISSN 1605:ISSN 1559:PMID 1551:ISSN 1485:ISSN 1413:link 1370:2014 1330:2014 1317:ISBN 1285:2014 1264:ISBN 1231:ISBN 1206:ISBN 1193:2013 1168:2013 1143:2013 1118:2013 1066:2008 1031:2017 973:2013 955:ISBN 912:OCLC 902:ISBN 874:2022 808:and 717:and 697:and 287:The 2453:MSW 2401:EoL 2388:CoL 2026:PMC 2008:doi 1938:hdl 1930:doi 1893:PMC 1875:doi 1809:PMC 1791:doi 1723:doi 1661:doi 1597:doi 1541:hdl 1533:doi 1475:hdl 1465:doi 1359:doi 1135:CBC 483:in 471:in 267:4. 2503:: 2481:: 2468:: 2455:: 2442:: 2429:: 2416:: 2403:: 2390:: 2375:: 2360:: 2173:^ 2057:^ 2034:. 2024:. 2016:. 2006:. 1996:11 1994:. 1990:. 1976:^ 1962:. 1954:. 1946:. 1936:. 1926:92 1924:. 1901:. 1891:. 1883:. 1873:. 1863:12 1861:. 1857:. 1831:^ 1817:. 1807:. 1799:. 1789:. 1779:10 1777:. 1773:. 1745:^ 1731:. 1719:98 1717:. 1713:. 1691:^ 1677:. 1669:. 1659:. 1649:85 1647:. 1643:. 1625:^ 1611:. 1603:. 1595:. 1585:41 1583:. 1571:^ 1557:. 1549:. 1539:. 1531:. 1521:27 1519:. 1497:^ 1483:. 1473:. 1461:26 1459:. 1455:. 1421:^ 1409:}} 1405:{{ 1386:. 1355:63 1353:, 1349:, 1311:, 1245:^ 1184:, 1158:, 1133:, 1099:^ 1033:, 953:, 949:, 937:^ 910:. 865:. 812:. 804:, 800:, 788:, 776:, 772:, 768:, 764:, 760:, 749:. 693:, 689:, 367:. 265:; 249:, 2042:. 2010:: 2002:: 1970:. 1940:: 1932:: 1909:. 1877:: 1869:: 1825:. 1793:: 1785:: 1739:. 1725:: 1685:. 1663:: 1655:: 1619:. 1599:: 1591:: 1565:. 1543:: 1535:: 1527:: 1491:. 1477:: 1467:: 1415:) 1401:. 1361:: 1239:. 1093:. 1068:. 918:. 876:. 553:( 510:( 499:( 475:/ 291:( 79:) 20:)

Index

Barren-ground Caribou

Thelon River
Conservation status
NatureServe
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Mammalia
Artiodactyla
Cervidae
Capreolinae
Rangifer
R. tarandus
Trinomial name

boreal woodland
migratory woodland
R. t. dawsoni
R. t. arcticus
R. t. pearyi
recent taxonomic revision
subspecies
reindeer
North America
Canadian
Nunavut
Northwest Territories

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