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2336:, which spiders lack, and to conceal the fact that they have eight legs rather than six; they develop large color patches round one pair of eyes to disguise the fact that they generally have eight simple eyes, while ants have two compound eyes; they cover their bodies with reflective bristles to resemble the shiny bodies of ants. In some spider species, males and females mimic different ant species, as female spiders are usually much larger than males. Ant-mimicking spiders also modify their behavior to resemble that of the target species of ant; for example, many adopt a zig-zag pattern of movement, ant-mimicking jumping spiders avoid jumping, and spiders of the genus
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spacing, larger sticky droplets and thicker lines would improve retention, but would make it easier for potential prey to see and avoid the web, at least during the day. However, there are no consistent differences between orb webs built for use during the day and those built for use at night. In fact, there is no simple relationship between orb web design features and the prey they capture, as each orb-weaving species takes a wide range of prey.
3286:
877:, and fold away behind the upper sections while not in use. The upper sections generally have thick "beards" that filter solid lumps out of their food, as spiders can take only liquid food. Scorpions' pedipalps generally form large claws for capturing prey, while those of spiders are fairly small appendages whose bases also act as an extension of the mouth; in addition, those of male spiders have enlarged last sections used for
3491:
2466:
2250:, and then swing the bolas at the moths. Although they miss on about 50% of strikes, they catch about the same weight of insects per night as web-weaving spiders of similar size. The spiders eat the bolas if they have not made a kill in about 30 minutes, rest for a while, and then make new bolas. Juveniles and adult males are much smaller and do not make bolas. Instead they release different pheromones that attract
1393:
1569:
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1428:(which may be thought of as a foot of sorts); the tarsus ends in a claw made up of either two or three points, depending on the family to which the spider belongs. Although all arthropods use muscles attached to the inside of the exoskeleton to flex their limbs, spiders and a few other groups still use hydraulic pressure to extend them, a system inherited from their pre-arthropod ancestors. The only
1533:, a comblike set of bristles on the jointed tip of the cribellum, and combined into a composite woolly thread that is very effective in snagging the bristles of insects. The earliest spiders had cribella, which produced the first silk capable of capturing insects, before spiders developed silk coated with sticky droplets. However, most modern groups of spiders have lost the cribellum.
131:
4068:. Most spend the majority of their time in burrows, and some run silk tripwires out from these, but a few build webs to capture prey. However, mygalomorphs cannot produce the piriform silk that the Araneomorphae use as an instant adhesive to glue silk to surfaces or to other strands of silk, and this makes web construction more difficult for mygalomorphs. Since mygalomorphs rarely "
2530:(bristles) with fragile bases and a row of barbs on the tip. The barbs cause intense irritation but there is no evidence that they carry any kind of venom. A few defend themselves against wasps by including networks of very robust threads in their webs, giving the spider time to flee while the wasps are struggling with the obstacles. The golden wheeling spider,
2108:. Laboratory studies show that they do so deliberately and over extended periods, and periodically clean themselves while feeding. These spiders also prefer sugar solutions to plain water, which indicates that they are seeking nutrients. Since many spiders are nocturnal, the extent of nectar consumption by spiders may have been underestimated. Nectar contains
993:, but the remaining members of this group have just the anterior pair of book lungs intact while the posterior pair of breathing organs are partly or fully modified into tracheae, through which oxygen is diffused into the haemolymph or directly to the tissue and organs. The tracheal system has most likely evolved in small ancestors to help resist
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4170:(family Atracidae) include fang display. Their venom, although they rarely inject much, has resulted in 13 attributed human deaths over 50 years. They have been deemed to be the world's most dangerous spiders on clinical and venom toxicity grounds, though this claim has also been attributed to the Brazilian wandering spider (genus
1679:
are two further openings leading directly to the spermathecae, creating a "flow through" system rather than a "first-in first-out" one. Eggs are as a general rule only fertilized during oviposition when the stored sperm is released from its chamber, rather than in the ovarian cavity. A few exceptions exist, such as
842:. In spiders, the cephalothorax and abdomen are connected by a small cylindrical section, the pedicel. The pattern of segment fusion that forms chelicerates' heads is unique among arthropods, and what would normally be the first head segment disappears at an early stage of development, so that chelicerates lack the
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2763:
Horizontal orb webs are fairly common, despite being less effective at intercepting and retaining prey and more vulnerable to damage by rain and falling debris. Various researchers have suggested that horizontal webs offer compensating advantages, such as reduced vulnerability to wind damage; reduced
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Female spiders' reproductive tracts are arranged in one of two ways. The ancestral arrangement ("haplogyne" or "non-entelegyne") consists of a single genital opening, leading to two seminal receptacles (spermathecae) in which females store sperm. In the more advanced arrangement ("entelegyne"), there
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cannot extend its legs, and the legs of dead spiders curl up. Spiders can generate pressures up to eight times their resting level to extend their legs, and jumping spiders can jump up to 50 times their own length by suddenly increasing the blood pressure in the third or fourth pair of legs. Although
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in web construction, in other words use of similar techniques by remotely related species, is rampant. Orb web designs and the spinning behaviors that produce them are the best understood. The basic radial-then-spiral sequence visible in orb webs and the sense of direction required to build them may
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period. In addition to preserving spiders' anatomy in very fine detail, pieces of amber show spiders mating, killing prey, producing silk and possibly caring for their young. In a few cases, amber has preserved spiders' egg sacs and webs, occasionally with prey attached; the oldest fossil web found
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kills and eats the male after it inserts its second palp into the female's genital opening; in fact, the males co-operate by trying to impale themselves on the females' fangs. Observation shows that most male redbacks never get an opportunity to mate, and the "lucky" ones increase the likely number
930:
flows. The heart is a tube in the upper part of the body, with a few ostia that act as non-return valves allowing blood to enter the heart from the hemocoel but prevent it from leaving before it reaches the front end. However, in spiders, it occupies only the upper part of the abdomen, and blood is
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There are several unusual variants of orb web, many of them convergently evolved, including: attachment of lines to the surface of water, possibly to trap insects in or on the surface; webs with twigs through their centers, possibly to hide the spiders from predators; "ladderlike" webs that appear
1332:
would block out information about the outside world, except that they are penetrated by many sensors or connections from sensors to the nervous system. In fact, spiders and other arthropods have modified their cuticles into elaborate arrays of sensors. Various touch sensors, mostly bristles called
2840:
Many experiments have been conducted to study the effect of zero gravity on the design of spider webs. In late 2020, reports of recent experiments were published that indicated that although web design was affected adversely in zero gravity conditions, having access to a light source could orient
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weave irregular, tangled, three-dimensional webs, popularly known as cobwebs. There seems to be an evolutionary trend towards a reduction in the amount of sticky silk used, leading to its total absence in some species. The construction of cobwebs is less stereotyped than that of orb-webs, and may
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build underwater "diving bell" webs that they fill with air and use for digesting prey and molting. Mating and raising the offspring happens in the female's bell. They live almost entirely within the bells, darting out to catch prey animals that touch the bell or the threads that anchor it. A few
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rituals to prevent the large females from eating the small males before fertilization, except where the male is so much smaller that he is not worth eating. In web-weaving species, precise patterns of vibrations in the web are a major part of the rituals, while patterns of touches on the female's
2178:
The best-known method of prey capture is by means of sticky webs. Varying placement of webs allows different species of spider to trap different insects in the same area, for example flat horizontal webs trap insects that fly up from vegetation underneath while flat vertical webs trap insects in
2014:
or larval
Lepidoptera, then the abdomen becomes green. Environmentally induced color changes may be morphological (occurring over several days) or physiological (occurring near instantly). Morphological changes require pigment synthesis and degradation. In contrast to this, physiological changes
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stings. Many alleged cases of spider bites may represent incorrect diagnoses, which would make it more difficult to check the effectiveness of treatments for genuine bites. A review published in 2016 agreed with this conclusion, showing that 78% of 134 published medical case studies of supposed
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Spiders often attach highly visible silk bands, called decorations or stabilimenta, to their webs. Field research suggests that webs with more decorative bands captured more prey per hour. However, a laboratory study showed that spiders reduce the building of these decorations if they sense the
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without breaking (stopping), and trapping the prey by entangling it or sticking to it (retention). No single design is best for all prey. For example: wider spacing of lines will increase the web's area and hence its ability to intercept prey, but reduce its stopping power and retention; closer
548:
rituals. Males of most species survive a few matings, limited mainly by their short life spans. Females weave silk egg cases, each of which may contain hundreds of eggs. Females of many species care for their young, for example by carrying them around or by sharing food with them. A minority of
2214:
has two different techniques for trapping prey: backwards strikes to catch flying insects, whose vibrations it detects; and forward strikes to catch ground-walking prey that it sees. These two techniques have also been observed in other deinopids. Walking insects form most of the prey of most
1715:, the remaining palp is then torn off by the female. The separated palp remains attached to the female's epigynum for about four hours and apparently continues to function independently. In the meantime, the female feeds on the palpless male. In over 60% of cases, the female of the Australian
8637:
Mammola, Stefano; Malumbres-Olarte, Jagoba; Arabesky, Valeria; Barrales-Alcalá, Diego
Alejandro; Barrion-Dupo, Aimee Lynn; Benamú, Marco Antonio; Bird, Tharina L.; Bogomolova, Maria; Cardoso, Pedro; Chatzaki, Maria; Cheng, Ren-Chung; Chu, Tien-Ai; Classen-Rodríguez, Leticia M.; Čupić, Iva;
1726:
1448:, consist of bristles whose ends are split into as many as 1,000 branches, and enable spiders with scopulae to walk up vertical glass and upside down on ceilings. It appears that scopulae get their grip from contact with extremely thin layers of water on surfaces. Spiders, like most other
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Information about spiders in the media is often emphasizing how dangerous and unpleasant they are. Among online newspaper articles on spider–human encounters and bites published from 2010 to 2020, a study found that 47% of articles contained errors and 43% were sensationalist.
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2759:
The hubs of orb webs, where the spiders lurk, are usually above the center, as the spiders can move downwards faster than upwards. If there is an obvious direction in which the spider can retreat to avoid its own predators, the hub is usually offset towards that direction.
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of mature males. When a male detects signs of a female nearby he checks whether she is of the same species and whether she is ready to mate; for example in species that produce webs or "safety ropes", the male can identify the species and sex of these objects by "smell".
516:, mostly preying on insects and other spiders, although a few large species also take birds and lizards. An estimated 25 million tons of spiders kill 400–800 million tons of prey every year. Spiders use numerous strategies to capture prey: trapping it in sticky webs,
1685:. In these species the female appears to be able to activate the dormant sperm before oviposition, allowing them to migrate to the ovarian cavity where fertilization occurs. The only known example of direct fertilization between male and female is an Israeli spider,
1188:
the ganglia of all segments behind the esophagus are fused, so that the cephalothorax is largely filled with nervous tissue and there are no ganglia in the abdomen; in the
Mesothelae, the ganglia of the abdomen and the rear part of the cephalothorax remain unfused.
2831:
generally make horizontal but uneven sheets, with tangles of stopping threads above. Insects that hit the stopping threads fall onto the sheet or are shaken onto it by the spider, and are held by sticky threads on the sheet until the spider can attack from below.
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weave even smaller webs, hold them outstretched between their first two pairs of legs, and lunge and push the webs as much as twice their own body length to trap prey, and this move may increase the webs' area by a factor of up to ten. Experiments have shown that
4245:—it is the abnormal fear of spiders or anything reminiscent of spiders, such as webs or spiderlike shapes. It is one of the most common specific phobias, and some statistics show that 50% of women and 10% of men show symptoms. It may be an exaggerated form of an
1279:
that point away from incoming light, just like in vertebrates, while the arrangement is the opposite in the former. The principal eyes are also the only ones with eye muscles, allowing them to move the retina. Having no muscles, the secondary eyes are immobile.
10063:
1295:. This acuity is achieved by a telephotographic series of lenses, a four-layer retina, and the ability to swivel the eyes and integrate images from different stages in the scan. The downside is that the scanning and integrating processes are relatively slow.
1255:
are only capable of detecting the direction from which light is coming, using the shadow cast by the walls of the cup. However, in spiders these eyes are capable of forming images. The other pairs, called secondary eyes, are thought to be derived from the
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that lurk in burrows, often closed by trapdoors and often surrounded by networks of silk threads that alert these spiders to the presence of prey. Other ambush predators do without such aids, including many crab spiders, and a few species that prey on
1574:
4151:
Although spiders are widely feared, only a few species are dangerous to people. Spiders will only bite humans in self-defense, and few produce worse effects than a mosquito bite or bee sting. Most of those with medically serious bites, such as
1995:
While in many spiders color is fixed throughout their lifespan, in some groups, color may be variable in response to environmental and internal conditions. Choice of prey may be able to alter the color of spiders. For example, the abdomen of
2726:. Now, however, it appears that non-orb spiders are a subgroup that evolved from orb-web spiders, and non-orb spiders have over 40% more species and are four times as abundant as orb-web spiders. Their greater success may be because sphecid
2722:
have been inherited from the common ancestors of most spider groups. However, the majority of spiders build non-orb webs. It used to be thought that the sticky orb web was an evolutionary innovation resulting in the diversification of the
2120:
and minerals in addition to sugars, and studies have shown that other spider species live longer when nectar is available. Feeding on nectar avoids the risks of struggles with prey, and the costs of producing venom and digestive enzymes.
5518:
Zhou, J., Lai, J., Menda, G., Stafstrom, J.A., Miles, C.I., Hoy, R.R. and Miles, R.N., 2022. Outsourced hearing in an orb-weaving spider that uses its web as an auditory sensor. Proceedings of the
National Academy of Sciences, 119(14),
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approach from which these spiders learn very quickly how to overcome new prey species. However, they seem to be relatively slow "thinkers", which is not surprising, as their brains are vastly smaller than those of mammalian predators.
400:. However, as there is currently neither paleontological nor embryological evidence that spiders ever had a separate thorax-like division, there exists an argument against the validity of the term cephalothorax, which means fused
2503:, several coloration schemes are present in a ratio that appears to remain constant, and this may make it more difficult for predators to recognize the species. Most spiders are insufficiently dangerous or unpleasant-tasting for
1501:
very similar to that used in insect silk. It is initially a liquid, and hardens not by exposure to air but as a result of being drawn out, which changes the internal structure of the protein. It is similar in tensile strength to
1381:, sensors that report the force exerted by muscles and the degree of bending in the body and joints, are well-understood. On the other hand, little is known about what other internal sensors spiders or other arthropods may have.
9278:
Probably the most recognized of the 10 most common phobias, arachnophobia is the fear of spiders. The statistics clearly show that more than 50% of women and 10% of men show signs of this leader on the 10 most common phobias
2348:. Ant mimicry in many spiders and other arthropods may be for protection from predators that hunt by sight, including birds, lizards and spiders. However, several ant-mimicking spiders prey either on ants or on the ants' "
1695:. In this species the male will penetrate its pedipalps through the female's body wall and inject his sperm directly into her ovaries, where the embryos inside the fertilized eggs will start to develop before being laid.
3124:
feeding system. Instead, they have mouths that point forwards and downwards, and all have some means of breathing air. Spiders (Araneae) are distinguished from other arachnid groups by several characteristics, including
1082:
from the midgut into the prey and then suck the liquified tissues of the prey into the gut, eventually leaving behind the empty husk of the prey. Others grind the prey to pulp using the chelicerae and the bases of the
2871:
Although the fossil record of spiders is considered poor, almost 1000 species have been described from fossils. Because spiders' bodies are quite soft, the vast majority of fossil spiders have been found preserved in
2945:
period, bears the earliest known silk-producing spigots, and was therefore hailed as a spider at the time of its discovery. However, these spigots may have been mounted on the underside of the abdomen rather than on
2128:
caught in webs may also be eaten, and studies have shown that young spiders have a better chance of survival if they have the opportunity to eat pollen. In captivity, several spider species are also known to feed on
438:
vary widely in size, shape and the amount of sticky thread used. It now appears that the spiral orb web may be one of the earliest forms, and spiders that produce tangled cobwebs are more abundant and diverse than
4416:
is a dance to rid the young woman of the lustful effects of a spider bite. Web-spinning also caused the association of the spider with creation myths, as they seem to have the ability to produce their own worlds.
2368:, but while hunting it imitates the behavior of a dying ant to attract worker ants. After a kill, some ant-mimicking spiders hold their victims between themselves and large groups of ants to avoid being attacked.
1246:
Spiders have primarily four pairs of eyes on the top-front area of the cephalothorax, arranged in patterns that vary from one family to another. The principal pair at the front are of the type called pigment-cup
1351:
may have up to 1,000 such chemosensitive setae, most on the tarsi of the first pair of legs. Males have more chemosensitive bristles on their pedipalps than females. They have been shown to be responsive to sex
8131:
Scholtz, Gerhard; Kamenz, Carsten (2006). "The book lungs of
Scorpiones and Tetrapulmonata (Chelicerata, Arachnida): evidence for homology and a single terrestrialisation event of a common arachnid ancestor".
4072:" by using air currents for transport, their populations often form clumps. In addition to arthropods, some mygalomorphs are known to prey on frogs, small mammals, lizards, snakes, snails, and small birds.
1814:'s brood clings to rough bristles on the mother's back, and females of some species respond to the "begging" behaviour of their young by giving them their prey, provided it is no longer struggling, or even
1720:
of offspring by ensuring that the females are well-fed. However, males of most species survive a few matings, limited mainly by their short life spans. Some even live for a while in their mates' webs.
1707:
amputate one of their palps before maturation and enter adult life with one palp only. The palps are 20% of the male's body mass in this species, and detaching one of the two improves mobility. In the
1370:
that detect force and vibrations. In web-building spiders, all these mechanical and chemical sensors are more important than the eyes, while the eyes are most important to spiders that hunt actively.
9576:
2386:
4386:
to a weaving contest. Arachne won, but Athena destroyed her tapestry out of jealousy, causing
Arachne to hang herself. In an act of mercy, Athena brought Arachne back to life as the first spider.
4412:
In some cultures, spiders have symbolized patience due to their hunting technique of setting webs and waiting for prey, as well as mischief and malice due to their venomous bites. The
Italian
3109:, consisting of a last common ancestor and all of its descendants). There has been debate about what their closest evolutionary relatives are, and how all of these evolved from the ancestral
1960:, guanine creates their silvery appearance. While guanine is originally an end-product of protein metabolism, its excretion can be blocked in spiders, leading to an increase in its storage.
1078:, spiders have a narrow gut that can only cope with liquid food and two sets of filters to keep solids out. They use one of two different systems of external digestion. Some pump digestive
533:
show signs of intelligence in their choice of tactics and ability to develop new ones. Spiders' guts are too narrow to take solids, so they liquefy their food by flooding it with digestive
2507:
to offer much benefit. However, a few species with powerful venom, large jaws or irritant bristles have patches of warning colors, and some actively display these colors when threatened.
1835:("skin") cannot stretch. In some species males mate with newly molted females, which are too weak to be dangerous to the males. Most spiders live for only one to two years, although some
9617:
4123:. They are distinguished by having fangs that oppose each other and cross in a pinching action, in contrast to the Mygalomorphae, which have fangs that are nearly parallel in alignment.
1756:
2021:
for instance can change its body color to match the substrate it lives on which makes it more difficult to be detected by prey. An example of physiological color change is observed in
1573:
1094:, compartments with no other exit, that extract nutrients from the food; most are in the abdomen, which is dominated by the digestive system, but a few are found in the cephalothorax.
1631:, which have excellent eyesight. If courtship is successful, the male injects his sperm from the palpal bulbs into the female via one or two openings on the underside of her abdomen.
408:. Similarly, arguments can be formed against the use of the term "abdomen", as the opisthosoma of all spiders contains a heart and respiratory organs, organs atypical of an abdomen.
9163:
Menassa, R.; Zhu, H.; Karatzas, C.N.; Lazaris, A.; Richman, A.; Brandle, J. (2004). "Spider dragline silk proteins in transgenic tobacco leaves: accumulation and field production".
1672:
Schematic diagrams showing sperm entering and being stored in the spermathecae; eggs leaving the ovaries and being fertilized; and finally a fertilized egg leaving the female's body
1366:, to attract the opposite sex. Because they are able to tell the sexes apart, it is assumed the blood scent is mixed with pheromones. Spiders also have in the joints of their limbs
10013:
8444:
Forster, R.R. & Platnick, N.I. (1984). "A review of the archaeid spiders and their relatives, with notes on the limits of the superfamily
Palpimanoidea (Arachnida, Araneae)".
2968:
may not have been true spiders, and probably used silk for lining nests or producing egg cases rather than for building webs. The largest known fossil spider as of 2011 is the
2429:
1795:
level. In some species, the females die afterwards, but females of other species protect the sacs by attaching them to their webs, hiding them in nests, carrying them in the
7526:"A review of the fossil record of spiders (Araneae) with special reference to Africa, and description of a new specimen from the Triassic Molteno Formation of South Africa"
1264:, but no longer have the separate facets typical of compound eyes. Unlike the principal eyes, in many spiders these secondary eyes detect light reflected from a reflective
1444:
Most spiders that hunt actively, rather than relying on webs, have dense tufts of fine bristles between the paired claws at the tips of their legs. These tufts, known as
524:, mimicking the prey to avoid detection, or running it down. Most detect prey mainly by sensing vibrations, but the active hunters have acute vision and hunters of the
1810:
stages inside the egg sac and emerge as spiderlings, very small and sexually immature but similar in shape to adults. Some spiders care for their young, for example a
9097:
7284:
Herberstein, M.E. (2000). "Foraging behaviour in orb-web spiders (Araneidae): Do web decorations increase prey capture success in
Argiope keyserlingi Karsch, 1878?".
2717:
may build very similar or significantly different webs. Nor is there much correspondence between spiders' classification and the chemical composition of their silks.
5853:"Structure and function of the female reproductive system in three species of goblin spiders (Arachnida: Araneae: Oonopidae): Morphology of oonopid spider genitalia"
2787:
for example, is known for its characteristic missing sector orb web. The missing sector contains a signal thread used to detect prey vibrations on the female's web.
2408:
5374:
549:
species are social, building communal webs that may house anywhere from a few to 50,000 individuals. Social behavior ranges from precarious toleration, as in the
5840:
Complex
Genital System of a Haplogyne Spider (Arachnida, Araneae, Tetrablemmidae) Indicates Internal Fertilization and Full Female Control Over Transferred Sperm
1087:, while flooding it with enzymes; in these species, the chelicerae and the bases of the pedipalps form a preoral cavity that holds the food they are processing.
8777:"An infestation of 2,055 brown recluse spiders (Araneae: Sicariidae) and no envenomations in a Kansas home: implications for bite diagnoses in nonendemic areas"
7001:
Avilés, L.; Maddison, W.P.; Agnarsson, I. (2006). "A New Independently Derived Social Spider with Explosive Colony Proliferation and a Female Size Dimorphism".
4194:, spider bites were not verified in over 90% of the reports. Even when verification had occurred, details of the treatment and its effects were often lacking.
8223:
4184:
spider bites did not meet the necessary criteria for a spider bite to be verified. In the case of the two genera with the highest reported number of bites,
895:, while the underside is covered by two rather flat plates. The abdomen is soft and egg-shaped. It shows no sign of segmentation, except that the primitive
6142:
11806:
9261:
374:. However, there has been debate among scientists about how families should be classified, with over 20 different classifications proposed since 1900.
6281:
Defrize, J.; Thery, M.; Casas, J. (1 May 2010). "Background colour matching by a crab spider in the field: a community sensory ecology perspective".
5414:"Descriptions of two new genera of the spider family Caponiidae (Arachnida, Araneae) and an update of Tisentnops and Taintnops from Brazil and Chile"
884:
In spiders, the cephalothorax and abdomen are joined by a small, cylindrical pedicel, which enables the abdomen to move independently when producing
2605:
build colonies that may contain several thousand individuals that co-operate in prey capture and share food. Other communal spiders include several
9228:
7571:
6512:
5218:
1172:
as local control centers in all segments; a brain formed by fusion of the ganglia for the head segments ahead of and behind the mouth, so that the
6421:"Fungus and fruit consumption by harvestmen and spiders (Opiliones, Araneae): the vegetarian side of two predominantly predaceous arachnid groups"
2751:
About half the potential prey that hit orb webs escape. A web has to perform three functions: intercepting the prey (intersection), absorbing its
544:
To avoid being eaten by the females, which are typically much larger, male spiders identify themselves as potential mates by a variety of complex
12217:
12065:
5937:
sp. nov (Araneae: Theridiidae) and its exceptional copulatory behaviour: emasculation, male palpal organ as a mating plug and sexual cannibalism"
564:
While the venom of a few species is dangerous to humans, scientists are now researching the use of spider venom in medicine and as non-polluting
1738:) (near the top of the leaf) is protected from the female by producing the right vibrations in the web, and may be too small to be worth eating.
7215:
2904:
1229:(center pair) are very acute. The outer pair are "secondary eyes" and there are other pairs of secondary eyes on the sides and top of its head.
6055:"Prolonged milk provisioning and extended maternal care in the milking spider Toxeus magnus: biological implications and questions unresolved"
12117:
10034:
3614:
6597:
5259:
12222:
8489:
3117:
3009:
1432:
muscles in spider legs are located in the three hip joints (bordering the coxa and the trochanter). As a result, a spider with a punctured
1400:
Each of the eight legs of a spider consists of seven distinct parts. The part closest to and attaching the leg to the cephalothorax is the
580:
to see if these can be used as silk factories. As a result of their wide range of behaviors, spiders have become common symbols in art and
6958:
Agnarsson, I. & Kuntner, M. (2005). "Madagascar: an unexpected hotspot of social Anelosimus spider diversity (Araneae: Theridiidae)".
6246:
Gillespie, Rosemary G. (1989). "Diet-Induced Color Change in the Hawaiian Happy-Face Spider Theridion grallator, (Araneae, Theridiidae)".
1117:. Production of uric acid and its removal via Malphigian tubules are a water-conserving feature that has evolved independently in several
2493:, both of which have good color vision. Many spider species are colored so as to merge with their most common backgrounds, and some have
2310:
hunt other spiders in ways that seem intelligent, outflanking their victims or luring them from their webs. Laboratory studies show that
1933:
1627:
body are important in many spiders that hunt actively, and may "hypnotize" the female. Gestures and dances by the male are important for
8817:
1773:
7989:
Lozano-Fernandez, Jesus; Tanner, Alastair R.; Puttick, Mark N.; Vinther, Jakob; Edgecombe, Gregory D.; Pisani, Davide (11 March 2020).
7730:
7720:
2985:
8390:"New and Remarkable Spiders of the Families Liphistiidae, Argyronetidae, Pisauridae, Theridiidae and Araneidae (Arachnida) from Japan"
2189:
spiders use the surfaces of lakes and ponds as "webs", detecting trapped insects by the vibrations that these cause while struggling.
1090:
The stomach in the cephalothorax acts as a pump that sends the food deeper into the digestive system. The midgut bears many digestive
1529:, a modified spinneret with up to 40,000 spigots, each of which produces a single very fine fiber. The fibers are pulled out by the
1001:, but in the majority of spiders this pair of spiracles has fused into a single one in the middle, and moved backwards close to the
12039:
8338:
8275:
2332:
to mimic the three distinct regions (tagmata) of an ant's body; they wave the first pair of legs in front of their heads to mimic
12091:
9923:
How to Know the Spiders: Pictured-Keys for Determining the More Common Spiders, with Suggestions for Collecting and Studying Them
1272:
can be spotted by torchlight reflected from the tapeta. On the other hand, the secondary eyes of jumping spiders have no tapeta.
9053:
Stuber, Marielle & Nentwig, Wolfgang (2016). "How informative are case studies of spider bites in the medical literature?".
6370:
4249:
response that helped early humans to survive, or a cultural phenomenon that is most common in predominantly European societies.
2798:
to test their web-spinning capabilities in zero gravity. At first, both produced rather sloppy webs, but they adapted quickly.
2768:
to catch insects in slow horizontal flight. However, there is no single explanation for the common use of horizontal orb webs.
2713:
There is no consistent relationship between the classification of spiders and the types of web they build: species in the same
1474:
The abdomen has no appendages except those that have been modified to form one to four (usually three) pairs of short, movable
1337:, respond to different levels of force, from strong contact to very weak air currents. Chemical sensors provide equivalents of
2015:
occur by changing the position of pigment-containing cells. An example of morphological color changes is background matching.
9976:
9949:
9898:
9852:
9806:
9722:
9695:
9666:
9551:
9524:
9486:
9461:
9434:
9407:
9308:
8939:
8621:
8562:
8537:
8428:
8372:
8242:
8195:
6867:
Blackledge, T.A. & Wenzel, J.W. (2001). "Silk Mediated Defense by an Orb Web Spider against Predatory Mud-dauber Wasps".
6772:
6126:
6037:
5777:
5542:
5492:
5275:
4765:
4740:
4642:
4549:
2558:
A few spider species that build webs live together in large colonies and show social behavior, although not as complex as in
1964:
occur in some species, which are the result of the diffraction, scattering or interference of light, for example by modified
922:
in which the coelom is reduced to small areas around the reproductive and excretory systems. Its place is largely taken by a
9198:
7394:
869:. Scorpions' chelicerae have three sections and are used in feeding. Spiders' chelicerae have two sections and terminate in
12096:
11156:
10018:
9138:
1744:
11454:
9324:
Davey, G.C.L. (1994). "The "Disgusting" Spider: The Role of Disease and Illness in the Perpetuation of Fear of Spiders".
7506:
6923:
Vollrath, F. (1986). "Eusociality and extraordinary sex ratios in the spider Anelosimus eximius (Araneae: Theridiidae)".
5884:"The spider Harpactea sadistica: co-evolution of traumatic insemination and complex female genital morphology in spiders"
3647:
2202:
stretch their webs and then release them when prey strike them, but do not actively move their webs. Those of the family
1384:
Some spiders use their webs for hearing, where the giant webs function as extended and reconfigurable auditory sensors.
1029:
9785:
8578:
8319:
7578:
4102:
In addition to accounting for over 90% of spider species, the Araneomorphae, also known as the "true spiders", include
1872:
from Colombia, are less than 0.37 mm (0.015 in) in body length. The largest and heaviest spiders occur among
9656:
4166:), would rather flee and bite only when trapped, although this can easily arise by accident. The defensive tactics of
2153:
caught on the webs of orb-weavers may be ingested along with the old web before construction of a new web. The enzyme
10107:
9541:
5384:
4464:
584:, symbolizing various combinations of patience, cruelty and creative powers. An irrational fear of spiders is called
7679:
5150:
2654:), which are notoriously cannibalistic, have formed small colonies in captivity, sharing webs and feeding together.
1598:
is not inserted into the female's body by the male's genitals but by an intermediate stage. Unlike many land-living
12212:
12179:
12018:
11258:
8248:
5507:
1184:
are the sole surviving family, spiders have the much more centralized nervous system that is typical of arachnids:
5219:""Eight-legged cats" and how they see – a review of recent research on jumping spiders (Araneae: Salticidae)"
4179:
There were about 100 reliably reported deaths from spider bites in the 20th century, compared to about 1,500 from
854:, and they lack anything that would function directly as "jaws". The first appendages behind the mouth are called
12122:
12031:
8234:
5658:"Spitting Performance Parameters and Their Biomechanical Implications in the Spitting Spider, Scytodes thoracica"
2242:
made of a single thread, tipped with a large ball of very wet sticky silk. They emit chemicals that resemble the
6513:"Adaptations to an aquatic life may be responsible for the reversed sexual size dimorphism in the water spider,
2160:
Spiders have been observed to consume plant material belonging to a large variety of taxa and type. Conversely,
1606:(packages of sperm), but spin small sperm webs onto which they ejaculate and then transfer the sperm to special
11372:
10053:
9265:
6176:
5111:"The effects of temperature on the web-building behaviour of the common house spider, Achaearanea tepidariorum"
4304:
4065:
3078:
account for the great majority of modern spiders, including those that weave the familiar orb-shaped webs. The
2497:, stripes and blotches that break up their outlines. In a few species, such as the Hawaiian happy-face spider,
2124:
Various species are known to feed on dead arthropods (scavenging), web silk, and their own shed exoskeletons.
12104:
4333:
of weaving, was envisioned as a spider spinning her web. According to her main myth, she resisted her father
9353:"Edible arachnids and myriapods worldwide – updated list, nutritional profile and food hygiene implications"
2586:
are at least somewhat social. Members of other species in the same family but several different genera have
1926:, resulting in its green color. Guanine is responsible for the white markings of the European garden spider
1876:, which can have body lengths up to 90 mm (3.5 in) and leg spans up to 250 mm (9.8 in).
1437:
larger spiders use hydraulics to straighten their legs, unlike smaller jumping spiders they depend on their
10246:
8082:"Exploring the evolution and terrestrialization of scorpions (Arachnida: Scorpiones) with rocks and clocks"
8080:
Howard, Richard J.; Edgecombe, Gregory D.; Legg, David A.; Pisani, Davide; Lozano-Fernandez, Jesus (2019).
5974:
4167:
2446:
1986:
both have areas of modified cuticle that act as light reflectors. The peacock spiders of Australia (genus
745:
434:
Their abdomens bear appendages, modified into spinnerets that extrude silk from up to six types of glands.
401:
397:
7045:
6557:
12078:
11948:
11935:
11253:
11216:
9514:
8718:
8464:
8273:
Coddington, Jonathan A.; Levi, Herbert W. (1991). "Systematics and Evolution of Spiders (Araneae) STOR".
5299:
1900:) have been identified in spiders, although other pigments have been detected but not yet characterized.
1559:
1396:
Image of a spider leg: 1–coxa; 2–trochanter; 3–femur; 4–patella; 5–tibia; 6–metatarsus; 7–tarsus; 8–claws
9236:
8516:
Assembling the Tree of Life – Phylogeny of Spiders: a review of the strictly fossil spider families
7702:
7568:
7109:
Bertani, R.; Fukushima, C. S.; Martins, R. (2008). "Sociable widow spiders? Evidence of subsociality in
6795:
Mclver, J.D. & Stonedahl, G. (1993). "Myrmecomorphy: Morphological and Behavioral Mimicry of Ants".
6531:
5233:
2730:, which are often the dominant predators of spiders, much prefer to attack spiders that have flat webs.
1362:
uses the scent of blood from mammals and other vertebrates, which is obtained by capturing blood-filled
935:
that pass through the pedicle and open into several parts of the cephalothorax. Hence spiders have open
10075:
5852:
4469:
12171:
11953:
4781:
Meehan, Christopher J.; Olson, Eric J.; Reudink, Matthew W.; Kyser, T. Kurt; Curry, Robert L. (2009).
3113:, which were marine animals. This 2019 cladogram illustrates the spiders' phylogenetic relationships.
1824:
produce a nutritious milk-like substance for their offspring, and fed until they are sexually mature.
931:
discharged into the hemocoel by one artery that opens at the rear end of the abdomen and by branching
12184:
12132:
9826:
7347:
6822:
3342:
3097:
fossils, spiders most likely diverged from other chelicerates between 375 and 328 million years ago.
3086:
periods provide a large number of fossil spiders, including representatives of many modern families.
2374:
2192:
Net-casting spiders weave only small webs, but then manipulate them to trap prey. Those of the genus
2179:
horizontal flight. Web-building spiders have poor vision, but are extremely sensitive to vibrations.
1815:
1681:
1519:
1034:
7883:"A giant spider from the Jurassic of China reveals greater diversity of the orbicularian stem group"
6881:
6753:
6698:
Yeargan, K.V. & Quate, L.W. (1997). "Adult male bolas spiders retain juvenile hunting tactics".
4295: – provided the highly irritant bristles, the spiders' main defense system, are removed first.
2238:
build "webs" that consist of only a single "trapeze line", which they patrol. They also construct a
160:
11206:
11149:
9117:
7635:
Penney, D. & Selden, P.A. (2007). "Spinning with the dinosaurs: the fossil record of spiders".
5839:
5794:
4459:
3189:
3071:
2470:
2328:
spiders face several challenges: they generally develop slimmer abdomens and false "waists" in the
1299:
10027:
7253:"Morphological phylogeny of cobweb spiders and their relatives (Araneae, Araneoidea, Theridiidae)"
806:
and proteins; heads that are composed of several segments that fuse during the development of the
11447:
5484:
3257:
601:
46:
31:
17:
9300:
8613:
8500:
6369:
Jackson, R.R.; Pollard, Simon D.; Nelson, Ximena J.; Edwards, G.B.; Barrion, Alberto T. (2001).
6340:
Exploitation of the Pseudomyrmex–Acacia mutualism by a predominantly vegetarian jumping spider (
6029:
6021:
4337:'s sexual advances by ensconcing herself in her web, but let him in after he promised her fresh
3158:
3074:, and their spinnerets appear adapted for building funnel-shaped webs to catch jumping insects.
12158:
11875:
9352:
9112:
8889:"The Global Epidemiology, Syndromic Classification, Management, and Prevention of Spider Bites"
6876:
6844:
Cooke, J.A.L.; Roth, V.D.; Miller, F.H. (1972). "The urticating hairs of theraphosid spiders".
6569:
5083:
3972:
build camouflaged silk tubes with a second trapdoor as an emergency exit. Members of the genus
2582:
has a strong tendency towards sociality: all known American species are social, and species in
2537:
1734:
1692:
1591:
1161:
381:
9712:
9451:
8420:
8412:
6118:
6112:
5530:
4840:"An Estimated 400–800 Million Tons of Prey Are Annually Killed by the Global Spider Community"
4732:
2648:
lives in small colonies which help to protect eggs and spiderlings. Even widow spiders (genus
997:. The trachea were originally connected to the surroundings through a pair of openings called
12153:
10994:
10278:
10100:
8929:
8821:
7818:
7530:
7416:
5268:
Animal cognition in nature: the convergence of psychology and biology in laboratory and field
3980:" outwards from their tunnels to help them detect approaching prey, while those of the genus
2494:
1961:
1763:
998:
553:, to cooperative hunting and food-sharing. Although most spiders live for at most two years,
240:
12145:
12070:
8605:
7834:"A golden orb-weaver spider(Araneae: Nephilidae: Nephila) from the Middle Jurassic of China"
7799:
7348:"Presence of potential prey affects web-building in an orb-weaving spider Zygiella x-notata"
5476:
5081:
Coddington, J.A. & Levi, H.W. (1991). "Systematics and Evolution of Spiders (Araneae)".
4895:
4429:
worshipped nature. They placed emphasis on animals and often depicted spiders in their art.
1658:
1536:
Even species that do not build webs to catch prey use silk in several ways: as wrappers for
1109:("little tubes") extract these wastes from the blood in the hemocoel and dump them into the
663:
11987:
11922:
11196:
10545:
9062:
8651:
8183:
8141:
7955:
7894:
7753:
7644:
7539:
7458:
7161:
7010:
6967:
6707:
6612:
5806:
5714:
5421:
5122:
4851:
4794:
4581:
4342:
3120:("jaw bases") at the bases of their legs; both of these features are part of the ancestral
3116:
Arachnids lack some features of other chelicerates, including backward-pointing mouths and
2718:
2665:
2587:
2394:
2282:, can adjust their ultraviolet reflectance to match the flowers in which they are lurking.
2023:
1922:
1893:
1644:
1309:
569:
8865:
8731:
7150:"Know your foe: synanthropic spiders are deterred by semiochemicals of European fire ants"
4325:
Spiders have been the focus of stories and mythologies of various cultures for centuries.
8:
11181:
11142:
10853:
9215:
There are many common phobias, but surprisingly, the most common phobia is arachnophobia.
4221:
4069:
4033:
2526:
on their abdomens and use their legs to flick them at attackers. These bristles are fine
2499:
2234:
2074:
1998:
1843:
was documented to have lived in the wild for 43 years, dying of a parasitic wasp attack.
1687:
1317:
944:
611:
9066:
8655:
8288:
8187:
8145:
7959:
7898:
7757:
7648:
7543:
7462:
7311:
Li, D. & Lee, W.S. (2004). "Predator-induced plasticity in web-building behaviour".
7234:
7165:
7014:
6971:
6808:
6711:
6616:
6338:
5810:
5718:
5425:
5126:
5095:
4855:
4798:
4585:
2685:
they release for communication have a notable deterrent effect on these spider species.
1421:
1405:
1070:
spiders, have lost their venom glands, and kill their prey with silk instead. Like most
11440:
11191:
10918:
10762:
10254:
9921:
9637:
9424:
9380:
9293:
9030:
9005:
8981:
8956:
8695:
8606:
8300:
8017:
7990:
7971:
7923:
7882:
7858:
7833:
7771:
7660:
7614:
7479:
7446:
7421:(Araneae: Linyphiidae): Web reduction – ethological and morphological adaptations"
7375:
7328:
7182:
7149:
7130:
7091:
7026:
6983:
6940:
6731:
6636:
6314:
6263:
6228:
6079:
6070:
6054:
5941:
5908:
5883:
5738:
5682:
5657:
5477:
5452:
5413:
5317:
4872:
4839:
4820:
4725:
4698:
4678:
4604:
4569:
3037:
2892:, places where conditions were exceptionally suited to preserving fairly soft tissues.
2705:
2617:
2567:
2184:
1928:
1840:
1563:
1490:. There are at least six types of silk gland, each producing a different type of silk.
1358:
1239:
1165:
1106:
936:
371:
155:
9126:
8753:
5705:
Vollrath, F. & Knight, D.P. (2001). "Liquid crystalline spinning of spider silk".
3219:
2764:
visibility to prey flying upwards, because of the backlighting from the sky; enabling
1198:) exhibit complex behaviour, including the ability to use a trial-and-error approach.
427:
are fused into one mass in the cephalothorax. Unlike most arthropods, spiders have no
380:, spiders (as with all arachnids) differ from other arthropods in that the usual body
12140:
11974:
11913:
11766:
10976:
10770:
10655:
10476:
9982:
9972:
9955:
9945:
9928:
9904:
9894:
9875:
9858:
9848:
9802:
9781:
9758:
9718:
9691:
9662:
9547:
9520:
9482:
9457:
9430:
9403:
9384:
9372:
9304:
9180:
9176:
9130:
9078:
9035:
8986:
8935:
8910:
8888:
8798:
8735:
8716:
Vetter, Richard S.; Isbister, Geoffrey K. (2008). "Medical Aspects of Spider Bites".
8699:
8687:
8679:
8617:
8558:
8533:
8424:
8368:
8292:
8238:
8191:
8175:
8157:
8113:
8065:
8048:
8022:
7928:
7910:
7863:
7664:
7656:
7484:
7367:
7269:
7252:
7246:
7244:
7187:
7022:
6979:
6768:
6723:
6628:
6486:
6440:
6394:
6306:
6298:
6255:
6220:
6172:
6122:
6084:
6033:
5913:
5868:
5822:
5773:
5742:
5730:
5687:
5611:
5592:
5584:
5538:
5488:
5457:
5439:
5380:
5321:
5271:
5135:
5110:
4877:
4812:
4761:
4736:
4702:
4690:
4682:
4662:
4638:
4609:
4545:
4446:
4330:
4119:
3930:
2782:
2644:
2642:("thieves"), and larger colonies are more successful in this. The herbivorous spider
2333:
2199:
2164:
spiders comprise the vast majority (over 80%) of reported incidents of plant-eating.
2061:
1917:
1855:
1623:
1583:
1466:
1413:
1329:
1304:
843:
545:
504:
440:
416:
9684:
9641:
8905:
8843:
7975:
7775:
7618:
7379:
7332:
7134:
7095:
7030:
6987:
6944:
6640:
6419:
Nyffeler, Martin; Machado, Glauco; Hartmann, Arik; Keiser, Carl N. (14 March 2023).
6318:
6203:
Oxford, G.S.; Gillespie, R.G. (1998). "Evolution and Ecology of Spider Coloration".
4824:
3320:
2780:. However, the significance of many variations is unclear. The orb-weaving species,
1425:
1275:
Other differences between the principal and secondary eyes are that the latter have
423:, spiders have the most centralized nervous systems of all arthropods, as all their
12207:
11849:
11421:
10695:
10669:
10463:
10330:
10093:
9629:
9593:
9364:
9333:
9172:
9122:
9070:
9025:
9017:
8976:
8968:
8927:
8900:
8788:
8727:
8669:
8659:
8453:
8284:
8149:
8103:
8093:
8060:
8012:
8002:
7963:
7918:
7902:
7853:
7845:
7814:
7761:
7652:
7606:
7557:
7547:
7474:
7466:
7359:
7320:
7293:
7264:
7230:
7177:
7169:
7122:
7083:
7018:
6975:
6932:
6886:
6849:
6804:
6715:
6678:
6620:
6478:
6432:
6386:
6290:
6232:
6212:
6074:
6066:
5993:
5983:
5950:
5903:
5895:
5864:
5814:
5722:
5677:
5669:
5626:
5574:
5447:
5429:
5307:
5263:
5130:
5091:
4867:
4859:
4802:
4674:
4634:
4599:
4589:
4541:
4537:
4213:
4091:
3935:
3780:
2850:
2841:
spiders and enable them to build their normally shaped webs under such conditions.
2572:
2217:
1970:
1417:
1409:
1233:
459:
367:
300:
9074:
8049:"A phylogenetic analysis of the arachnid orders based on morphological characters"
7594:
7324:
7241:
6735:
6624:
6216:
5818:
5795:"Fertilization occurs internally in the spider Achaearanea tepidariorum (C. Koch)"
3986:
do not and instead rely on their built-in vibration sensors. Spiders of the genus
1401:
798:. As arthropods, they have: segmented bodies with jointed limbs, all covered in a
469:, but these animals apparently lacked spinnerets. True spiders have been found in
11382:
11351:
11225:
11201:
11186:
11117:
11050:
10962:
10747:
10537:
10481:
10322:
10299:
10057:
10038:
10008:
9888:
9796:
9202:
8437:
8360:
7793:
7791:
7789:
7787:
7785:
7592:
7575:
5109:
Barghusen, L.E.; Claussen, D.L.; Anderson, M.S.; Bailer, A.J. (1 February 1997).
5108:
4628:
4474:
4242:
4006:
3994:
3990:
have no venom glands, although they do have venom gland outlets on the fang tip.
3723:
3565:
3090:
2743:
2659:
2630:
2627:
2523:
2436:
2421:
2318:
2306:
2270:
1491:
1265:
1194:
1130:
811:
529:
481:
385:
347:
312:
283:
12109:
11979:
11927:
9778:
Forest Spiders of South East Asia: With a Revision of the Sac and Ground Spiders
9453:
Tales, Rumors, and Gossip: Exploring Contemporary Folk Literature in Grades 7–12
6761:
The Cognitive Animal: Empirical and Theoretical Perspectives on Animal Cognition
6558:"Web manipulation and two stereotyped attack behaviors in the ogre-faced spider
1021:
623:(a reference to how spiders make their webs), from the Proto-Indo-European root
12026:
12013:
11940:
11854:
11758:
11748:
11743:
11603:
11578:
11513:
11403:
11268:
10939:
10898:
10827:
10574:
10442:
10418:
10408:
10351:
10238:
9840:
9831:
8921:
8793:
8776:
8153:
8098:
8081:
7502:
7470:
7209:
7207:
7205:
7203:
7201:
6378:
4532:
Cushing, P.E. (2008). "Spiders (Arachnida: Araneae)". In Capinera, J.L. (ed.).
4398:
4263:
4153:
4114:
4103:
4096:
3951:
3593:
3507:
3337:
3298:
2989:
2639:
2536:, of the Namibian desert escapes parasitic wasps by flipping onto its side and
2490:
2291:
2287:
2057:
2045:
2017:
1716:
1628:
1541:
1522:. In other words, it can stretch much further before breaking or losing shape.
1222:
1006:
648:
572:
superior to synthetic materials, and spider silk genes have been inserted into
225:
10069:
9633:
9597:
9249:
Arachnophobia, or fear of spiders, is one of the most common specific phobias.
8664:
8639:
8636:
7906:
7782:
7126:
6890:
6390:
6332:
6330:
6328:
5954:
5312:
4863:
4807:
4782:
4365:
and planting it in the ground to produce eight previously nonexistent plants.
4086:
4027:
2889:
2738:
1148:"), which use large amounts of water to excrete nitrogenous waste products as
814:, sets of segments that serve similar functions: the foremost one, called the
12201:
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11243:
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10903:
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10753:
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10703:
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so far is 100 million years old. Earlier spider fossils come from a few
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2553:
2519:
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2040:
1982:
1820:
1603:
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9546:. Newcastle upon Tyne, England: Cambridge Scholars Publishing. p. 166.
9337:
8582:
7766:
7737:
7198:
5988:
5969:
5563:"Hydraulic Leg Extension Is Not Necessarily the Main Drive in Large Spiders"
4594:
1920:
process, resulting in a brown coloration. Bilins are found, for example, in
1912:, very common in other animals, are apparently absent. In some species, the
541:, as arachnids do not have the mandibles that crustaceans and insects have.
138:
Representatives of the three major extant spider groups (counterclockwise):
11784:
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10908:
10778:
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10509:
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9879:
9753:
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9134:
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9039:
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8691:
8161:
8026:
7932:
7867:
7849:
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7363:
7191:
7074:
Cangialosi, K.R. (1990). "Social spider defense against kleptoparasitism".
6747:
6745:
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6632:
6325:
6310:
6224:
6088:
5917:
5899:
5734:
5691:
5596:
5528:
5461:
4896:"Spider | Origin and meaning of spider by Online Etymology Dictionary"
4881:
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present in their digestive fluid allows for the digestion of these spores.
2085:
2066:
1956:
1374:
1257:
1181:
1067:
990:
900:
293:
10085:
10044:
9479:
Gods, Demons and Symbols of Ancient Mesopotamia: An Illustrated Dictionary
9368:
9021:
7593:
Dunlop, Jason A.; David Penney; O. Erik Tetlie; Lyall I. Anderson (2008).
7552:
7525:
6719:
5637:
5560:
4382:(Ancient Greek for "spider") was a Lydian girl who challenged the goddess
2806:
12083:
12052:
11907:
11898:
11771:
11735:
11488:
11377:
11341:
11331:
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11173:
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10984:
10948:
10877:
10861:
10810:
10801:
10792:
10633:
10603:
10499:
10359:
10142:
10081:
Record breaking achievements by spiders and the scientists who study them
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5932:
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5630:
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4454:
4402:
4209:
4203:
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4146:
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3788:
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3213:
3171:
3153:
3140:
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in China. Its body length is almost 25 mm, (i.e., almost one inch).
2964:
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2815:
2765:
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2650:
2575:
2559:
2504:
2453:
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2007:
1938:
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1811:
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1611:
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1377:
sensors and rely on their eyes to tell them which way is up. Arthropods'
1269:
1261:
1217:
1176:
is encircled by this conglomeration of ganglia. Except for the primitive
1043:
994:
986:
966:
957:
Spiders have developed several different respiratory anatomies, based on
858:, and serve different functions within different groups of chelicerates.
839:
791:
550:
475:
393:
339:
202:
63:
8931:
Venomous and Poisonous Marine Animals: A Medical and Biological Handbook
8674:
8389:
8108:
7630:
7628:
7523:
7173:
6742:
6465:
Nyffeler, Martin; Olson, Eric J.; Symondson, William O.C. (April 2016).
5561:
Weihmann, Tom; Günther, Michael; Blickhan, Reinhard (15 February 2012).
1298:
There are spiders with a reduced number of eyes, the most common having
1192:
Despite the relatively small central nervous system, some spiders (like
889:
11844:
11662:
11521:
11398:
11301:
11296:
11238:
11088:
11079:
11060:
11021:
11012:
10933:
10885:
10869:
10845:
10840:
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10783:
10734:
10718:
10687:
10679:
10616:
10553:
10504:
10434:
10426:
10413:
10400:
10343:
10317:
10286:
10270:
10164:
9971:. Cambridge Studies in Ecology. Cambridge: Cambridge University Press.
8715:
8304:
7087:
6936:
6420:
6294:
6267:
6143:"World's oldest known spider dies at 43 after a quiet life underground"
5579:
5562:
4413:
4318:
4309:
4172:
4061:
4045:
3982:
3968:
3925:
3841:
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3000:
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2101:
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1913:
1905:
1889:
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1313:
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1133:
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lineages that can live far away from water, for example the tubules of
1059:
1047:
970:
947:
896:
851:
452:
435:
420:
355:
335:
323:
256:
139:
108:
73:
11966:
7610:
7562:
7108:
5529:
Barnes, R.S.K.; Calow, P.; Olive, P.; Golding, D.; Spicer, J. (2001).
4783:"Herbivory in a Spider Through Exploitation of an Ant–Plant Mutualism"
4440:
3285:
2860:
2298:
capture prey by chasing it, and rely mainly on vision to locate prey.
2027:, which can change its body color from white to brown near instantly.
926:, a cavity that runs most of the length of the body and through which
431:
muscles in their limbs and instead extend them by hydraulic pressure.
12057:
12000:
11703:
11640:
11586:
11551:
11534:
11482:
11346:
11306:
11281:
11035:
10989:
10953:
10708:
10566:
10523:
10517:
10490:
10364:
10312:
10136:
9661:. Santa Barbara, California: Greenwood Publishing Group. p. 31.
9618:"Ekphrasis and the Theme of Artistic Failure in Ovid's Metamorphoses"
7967:
7625:
7000:
6853:
6764:
6482:
5998:
5726:
4909:
4907:
4905:
4506:
4406:
4390:
4354:
4292:
4280:
4268:
4180:
4107:
4057:
3485:
3417:
3233:
3184:
3126:
3121:
3061:
3020:
2979:
2951:
2947:
2936:
2911:
2878:
2592:
2515:
2349:
2266:
2251:
2243:
2222:
2194:
2161:
2154:
2134:
2011:
1950:
1873:
1836:
1828:
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1599:
1545:
1526:
1515:
1475:
1353:
1342:
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1141:
1118:
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974:
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795:
581:
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554:
509:
494:
464:
319:
192:
172:
113:
57:
12044:
11869:
10049:
8457:
8217:
8215:
8213:
8211:
8209:
8207:
7988:
7346:
Pasquet, Alain; Ridwan, Ahmad; Leborgne, Raymond (1 February 1994).
7297:
5212:
5210:
5208:
5206:
4044:
period, are generally heavily built and ″hairy″, with large, robust
2876:. The oldest known amber that contains fossil arthropods dates from
1356:
produced by females, both contact and air-borne. The jumping spider
1320:
species have no eyes, or possess vestigial eyes incapable of sight.
1050:
are fangs, and the great majority of spiders can use them to inject
12005:
11961:
11892:
11628:
11559:
11463:
11432:
11336:
11326:
11055:
10913:
10664:
10641:
10458:
10148:
10080:
8488:
Penney, D. & Selden, P.A. Deltshev, C. & Stoev, P. (eds.).
7800:"The Role of Behavior in the Evolution of Spiders, Silks, and Webs"
6843:
6466:
5032:
5030:
5028:
5026:
5024:
5022:
5020:
4284:
4272:
4246:
4041:
3977:
3963:
3946:
The only living members of the primitive Mesothelae are the family
3861:
Four pairs, in some species one pair fused, under middle of abdomen
3490:
3278:
3130:
3079:
3056:
2942:
2925:
2917:
2896:
2791:
2752:
2358:
2209:
2142:
1944:
1792:
1703:
1615:
1511:
1452:, keep at least four legs on the surface while walking or running.
1449:
1445:
1429:
1363:
1169:
1098:
1084:
1075:
1071:
923:
892:
866:
862:
855:
538:
513:
456:
444:
428:
424:
351:
343:
212:
103:
98:
83:
78:
68:
50:
9095:
8928:
Williamson, J.A.; Fenner, P.J.; Burnett, J.W.; Rifkin, J. (1996).
7507:"Space Station Spiders Found a Hack to Build Webs Without Gravity"
7445:
Zschokke, Samuel; Countryman, Stefanie; Cushing, Paula E. (2021).
6336:
4902:
3962:. Most of the Liphistiidae construct silk-lined burrows with thin
3779:
of spiders (order Araneae) have been identified, grouped into 132
11670:
11316:
11311:
10926:
10726:
10335:
9890:
The Book of the Spider: From Arachnophobia to the Love of Spiders
8204:
7148:
Fischer, Andreas; Lee, Yerin; Dong, T'ea; Gries, Gerhard (2021).
6368:
5412:
Brescovit, Antonio D.; Sánchez-Ruiz, Alexander (6 October 2016).
5203:
4379:
4338:
4002:
3827:
3776:
3026:
2959:
2465:
2146:
2117:
2049:
1988:
1901:
1897:
1885:
1832:
1607:
1498:
1347:
1149:
978:
962:
888:. The upper surface of the cephalothorax is covered by a single,
835:
819:
799:
377:
363:
359:
118:
93:
10050:
Online Videos of Jumping Spiders (Salticids) and other arachnids
7703:"A trigonotarbid arachnid from the Upper Silurian of Shropshire"
6015:
6013:
6011:
6009:
5344:
5342:
5340:
5338:
5193:
5191:
5178:
5176:
5047:
5045:
5017:
3066:. Some Triassic mygalomorphs appear to be members of the family
2402:
serves both as a trap and as a way of making its home in a leaf.
1992:) are notable for their bright structural colours in the males.
1932:. It is in many species accumulated in specialized cells called
568:. Spider silk provides a combination of lightness, strength and
11476:
10130:
10045:
New Mexico State University "The Spiders of the Arid Southwest"
10022:
5007:
5005:
5003:
5001:
4999:
4997:
4995:
4993:
4991:
4989:
4987:
4985:
4983:
4981:
4979:
4977:
4975:
4973:
4971:
4969:
4967:
4965:
4963:
4961:
4959:
4957:
4394:
4383:
4362:
4288:
3511:
3041:
2954:
and whose mobility is important in the building of webs. Hence
2440:
uses both webs and cunning, versatile tactics to overcome prey.
2130:
2125:
2105:
2070:
1976:
1909:
1791:
in one or more silk egg sacs, which maintain a fairly constant
1507:
1438:
1292:
1248:
1226:
1145:
1126:
1122:
1110:
1079:
982:
951:
932:
919:
831:
823:
807:
803:
573:
534:
448:
412:
405:
182:
9714:
American Anthropology: Papers from the American Anthropologist
9519:. New Haven, Connecticut: Yale University Press. p. 184.
8581:. Agricultural Research Council of New Zealand. Archived from
8042:
8040:
8038:
8036:
5970:"Risky mate search and male self-sacrifice in redback spiders"
4955:
4953:
4951:
4949:
4947:
4945:
4943:
4941:
4939:
4937:
2638:). Social predatory spiders need to defend their prey against
1392:
12166:
11716:
8354:
8352:
6418:
6095:
6006:
5749:
5335:
5188:
5173:
5057:
5042:
4358:
4053:
4049:
3959:
3955:
3784:
3748:
3739:
3729:
3248:
3106:
2873:
2865:
2795:
2714:
2598:
2353:
2302:
2239:
2150:
2113:
1818:
food. In one exceptional case, females of the jumping spider
1807:
1708:
1699:
1595:
1537:
1503:
1487:
1338:
1125:
and arachnids develop from completely different parts of the
1091:
1055:
1051:
927:
878:
874:
577:
525:
521:
517:
331:
11992:
9290:
8612:(2nd ed.). New York: Oxford University Press. pp.
8394:
Bulletin of the National Science Museum (Of Japan), Series A
8229:. In Ubick, D.; Paquin, P.; Cushing, P.E.; Roth, V. (eds.).
7447:"Spiders in space—orb-web-related behaviour in zero gravity"
6866:
5531:"Invertebrates with Legs: the Arthropods and Similar Groups"
2002:
will become orange if the spider ingests certain species of
783:
Legs are labelled I, II, III, IV from anterior to posterior.
11776:
8443:
8180:
Wonderful Life: The Burgess Shale and the Nature of History
8033:
7831:
6555:
6371:"Jumping spiders (Araneae: Salticidae) that feed on nectar"
6053:
Dong, Bing; Quan, Rui-Chang; Chen, Zhan-Qi (18 July 2019).
4934:
4924:
4922:
4426:
4369:
4350:
4346:
4334:
4326:
4217:
2924:
and segmented abdomen, as well as eight legs and a pair of
2777:
2727:
2578:) can form colonies of up to 50,000 individuals. The genus
2541:
2527:
2486:
2247:
2138:
1965:
1334:
1287:
of some jumping spiders exceeds by a factor of ten that of
1114:
885:
870:
827:
327:
130:
11134:
9481:. London, England: The British Museum Press. p. 182.
8349:
5930:
5508:'Vampire' spiders use blood as perfume | CBC News – CBC.ca
5216:
5080:
4013:
rocks, have been classified as members of the Mesothelae.
2056:
Although spiders are generally regarded as predatory, the
1441:
muscles to generate the propulsive force for their jumps.
1408:
that works as a hinge for the following long segment, the
8820:. Department of Entomology, Virginia Tech. Archived from
8266:
8079:
7444:
6794:
6759:. In Bekoff, M.; Allen, C. & Burghardt, G.M. (eds.).
6337:
Meehan, C.J.; Olson, E.J.; Curry, R.L. (21 August 2008).
2275:
2078:
2003:
1788:
754:
female, ventral aspect. Nos 1 to 14 as for dorsal aspect.
624:
7797:
6957:
6697:
5704:
5609:
4919:
3420:
shows the relation among spider suborders and families:
846:
typical of most arthropods. In fact, chelicerates' only
354:. Spiders are found worldwide on every continent except
9658:
The Greenwood Encyclopedia of Folktales and Fairy Tales
9543:
Aspects of Ecphrastic Technique in Ovid's Metamorphoses
9295:
Cognitive Science: An Introduction to the Study of Mind
7991:"A Cambrian–Ordovician Terrestrialization of Arachnids"
7392:
6663:"The Natural History and Behavior of the Bolas Spider,
5967:
4780:
4567:
1866:
Spiders occur in a large range of sizes. The smallest,
1013:, so environmental temperatures affect their activity.
9351:
Costa-Neto, E.M.; Grabowski, N.T. (27 November 2020).
8465:"A review of the archaeid spiders and their relatives"
7634:
7046:"Cooperative prey capture in the communal web spider,
6510:
6464:
6198:
6196:
6194:
6192:
6190:
6188:
6186:
6184:
5483:(3rd p/b ed.). Oxford University Press. pp.
4661:
Dimitrov, Dimitar; Hormiga, Gustavo (7 January 2021).
4357:
heard Uttu's screams and rescued her, removing Enki's
3044:, there are very few fossil spiders from this period.
981:
surface of the abdomen allow air to enter and diffuse
346:
and rank seventh in total species diversity among all
8073:
7524:
Selden, P.A.; Anderson, H.M.; Anderson, J.M. (2009).
7345:
6751:
6028:(2nd ed.). Oxford University Press US. pp.
5411:
5260:"Cognitive Abilities of Araneophagic Jumping Spiders"
5076:
5074:
5072:
4570:"Fossil evidence for the origin of spider spinnerets"
2481:
There is strong evidence that spiders' coloration is
2342:
walk on the outer edges of leaves in the same way as
2317:
s instinctive tactics are only starting points for a
484:, the Mesothelae. The main groups of modern spiders,
480:
and are very similar to the most primitive surviving
7807:
Annual Review of Ecology, Evolution, and Systematics
6906:"Fog, wind and heat – life in the Namib desert"
4838:
Nyffeler, Martin; Birkhofer, Klaus (14 March 2017).
4563:
4561:
4436:
4317:
ceramic depicts a spider, and dates from around 300
4131:
604:
9927:(1st ed.). Dubuque, Iowa: W.C. Brown Company.
9516:
The Harps that Once: Sumerian Poetry in Translation
9350:
7881:Selden, Paul A.; Shih, ChungKun; Ren, Dong (2013).
7738:"A replacement name for the trigonotarbid arachnid
7147:
6181:
1016:
1005:. Spiders that have tracheae generally have higher
561:spiders can live up to 25 years in captivity.
492:, first appeared in the Triassic period, more than
358:, and have become established in nearly every land
9920:
9683:
9575:
9292:
8954:
8815:
8487:
8446:Bulletin of the American Museum of Natural History
8231:Spiders of North America: an identification manual
5069:
4724:
1312:. Other species have four eyes and members of the
1113:chamber, from which they are expelled through the
822:, is a complete fusion of the segments that in an
810:. Being chelicerates, their bodies consist of two
9775:
9344:
9262:"The 10 Most Common Phobias – Did You Know?"
8893:American Journal of Tropical Medicine and Hygiene
8358:
7946:Selden, P.A. (1996). "Fossil mesothele spiders".
7595:"How many species of fossil arachnids are there?"
7283:
6825:. University of California Museum of Paleontology
6280:
6019:
5399:
4837:
4558:
4048:and fangs (technically, spiders do not have true
2590:developed social behavior. For example, although
2254:, and catch them with their front pairs of legs.
2225:flies that they catch with the backwards strike.
396:, or abdomen, and joined by a small, cylindrical
12199:
9794:
9508:
9506:
9504:
9502:
9500:
9498:
8640:"The global spread of misinformation on spiders"
8221:
7982:
7073:
5755:
5643:
5360:
5348:
5197:
5182:
5063:
5051:
5036:
5011:
4928:
4913:
4568:Selden, P.A. & Shear, W.A. (December 2008).
4220:' milk and in the leaves of plants, by means of
4117:, as well as the only known herbivorous spider,
3767:, of which the latter contains two infraorders,
3105:The spiders (Araneae) are monophyletic (i.e., a
3003:, a primitive group now represented only by the
2485:that helps them to evade their major predators,
1839:can live in captivity for over 20 years, and an
1590:Spiders reproduce sexually and fertilization is
1470:An orb weaver producing silk from its spinnerets
10307:(funnel-webs or venomous funnel-web tarantulas)
9795:Ruppert, E.E.; Fox, R.S.; Barnes, R.D. (2004).
9768:
9752:The Spirit of Ancient Peru: Treasures from the
9096:Hinman, M.B.; Jones J.A.; Lewis, R. W. (2000).
7310:
6598:"Aggressive Chemical Mimicry by a Bolas Spider"
6202:
5556:
5554:
5258:Wilcox, R. Stimson; Jackson, Robert R. (1998).
4663:"Spider Diversification Through Space and Time"
4660:
4040:The Mygalomorphae, which first appeared in the
3133:that are specially adapted for sperm transfer.
2681:ant colonies. These ants are predators and the
1373:Like most arthropods, spiders lack balance and
537:. They also grind food with the bases of their
10066:, from the International Field Guides database
9003:
8553:Schultz, Stanley; Schultz, Margeurite (2009).
8552:
8528:Schultz, Stanley; Schultz, Marguerite (2009).
8527:
8272:
7832:Selden, P.A.; ChungKun Shih; Dong Ren (2011).
7213:
6660:
6595:
6364:
6362:
3893:One, two or three pairs under rear of abdomen
2356:. When at rest, the ant-mimicking crab spider
1551:
977:filled with haemolymph, where openings on the
906:
903:, have segmented plates on the upper surface.
419:. In all except the most primitive group, the
11448:
11150:
10101:
9717:. University of Nebraska Press. p. 455.
9577:"The spinner and the poet: Arachne in Ovid's
9495:
9323:
9052:
8557:. Hauppauge, New York: Barron's. p. 88.
8532:. Hauppauge, New York: Barron's. p. 28.
8337:Liddell, Henry George; Scott, Robert (1940).
8130:
7880:
7250:
7043:
6903:
5772:(3rd p/b ed.). Oxford University Press.
5655:
5257:
4760:. Hauppauge, New York: Barron's. p. 23.
4731:. New York: Oxford University Press. p.
4626:
3070:, whose modern members include the notorious
2835:
1058:in the roots of the chelicerae. The families
1025:A syrphid fly captured in the web of a spider
10698:(wandering spiders or tropical wolf spiders)
9915:
9291:Friedenberg, J. & Silverman, G. (2005).
8774:
8417:Spiders – webs, behavior, and evolution
8336:
8046:
7945:
7735:
7724:but was renamed when it was realized that a
7700:
6922:
6052:
5551:
4756:Shultz, Stanley; Shultz, Marguerite (2009).
4755:
4341:as a marriage gift, thereby allowing him to
1974:results from bristles reflecting the light,
1494:also produce silk in modified venom glands.
910:
10115:
9569:
9567:
9565:
9563:
9402:. Lonely Planet Publications. p. 308.
9046:
8957:"Misdiagnosis of brown recluse spider bite"
8410:
8318:Lewis, Charlton T.; Short, Charles (1879).
8174:
6556:Coddington, J. & Sobrevila, C. (1987).
6359:
6167:Levi, Herbert W. and Levi, Lorna R. (2001)
5961:
5799:Invertebrate Reproduction & Development
5612:"The Hydraulic Mechanism of the Spider Leg"
4716:
4507:"Currently valid spider genera and species"
4501:
4499:
4497:
4495:
1614:or palpal organs, borne on the tips of the
1105:, which can be excreted as a dry material.
1009:and better water conservation. Spiders are
362:. As of September 2024, 52,309 spider
11455:
11441:
11157:
11143:
10204:
10108:
10094:
9476:
8886:
8711:
8709:
8317:
7414:
7113:Walckenaer, 1805 (Araneae, Theridiidae)".
5931:Knoflach, B. & van Harten, A. (2001).
5468:
5217:Harland, D.P. & Jackson, R.R. (2000).
4656:
4654:
4627:Sebastin, P.A.; Peter, K.V., eds. (2009).
2084:Juveniles of some spiders in the families
1750:Orange spider egg sac hanging from ceiling
1424:is next, and it connects the tibia to the
1291:, which have by far the best vision among
865:are members of one chelicerate group, the
129:
10257:(atypical tarantulas or purseweb spiders)
9710:
9611:
9609:
9607:
9226:
9116:
9029:
8980:
8904:
8792:
8673:
8663:
8499:. Supplement No. 1: 25–39. Archived from
8224:"Phylogeny and Classification of Spiders"
8107:
8097:
8064:
8053:Zoological Journal of the Linnean Society
8016:
8006:
7922:
7857:
7765:
7588:
7586:
7561:
7551:
7478:
7268:
7257:Zoological Journal of the Linnean Society
7181:
6880:
6682:
6245:
6078:
5997:
5987:
5907:
5681:
5578:
5451:
5433:
5311:
5300:"Spiders are much smarter than you think"
5134:
4871:
4806:
4603:
4593:
3411:
3100:
2895:The oldest known exclusively terrestrial
1916:of the legs and prosoma is modified by a
1602:, male spiders do not produce ready-made
1486:, each of which is connected to one silk
1155:
642:
512:in 2008, but all other known species are
11125:are families with more than 1000 species
10012:) is being considered for deletion. See
9886:
9869:
9839:
9675:
9560:
9512:
9422:
9264:. 10 Most Common Phobias. Archived from
9098:"Synthetic spider silk: a modular fiber"
8638:Dhiya’ulhaq, Naufal Urfi (August 2022).
8276:Annual Review of Ecology and Systematics
7819:10.1146/annurev.ecolsys.37.091305.110221
7798:Vollrath, F. & Selden, P.A. (2007).
7223:Annual Review of Ecology and Systematics
5610:Parry, D.A. & Brown, R.H.J. (1959).
4492:
4308:
4262:
4085:
4026:
3929:
3759:Spiders are divided into two suborders,
2859:
2805:
2737:
2698:
2464:
2039:
2035:
1850:
1831:, spiders have to molt to grow as their
1732:The tiny male of the golden orb weaver (
1567:
1465:
1391:
1232:
1216:
1129:. However, a few primitive spiders, the
1028:
1020:
10162:
10028:Picture story about the jumping spider
9754:Museo Arqueológico Rafael Larco Herrera
9573:
9397:
8706:
8387:
8330:
8182:. Hutchinson Radius. pp. 102–06 .
7825:
7501:
7216:"Function and Phylogeny of Spider Webs"
5924:
5881:
4774:
4651:
4620:
4531:
2261:, the "trapdoor spiders" of the family
2030:
1767:spiderlings next to their eggs' capsule
985:. This is also the case for some basal
14:
12218:Extant Pennsylvanian first appearances
12200:
9824:
9776:Deeleman-Reinhold, Christa L. (2001).
9750:Berrin, Katherine & Larco Museum.
9616:Leach, Eleanor Winsor (January 1974).
9604:
9539:
9477:Black, Jeremy; Green, Anthony (1992).
9449:
8756:. Illinois Department of Public Health
8603:
8419:. Stanford University Press. pp.
7583:
6511:Schütz, D. & Taborsky, M. (2003).
6110:
6104:
5850:
5792:
5767:
5761:
5656:Suter, R. B.; Stratton, G. E. (2009).
5474:
4722:
939:. The blood of many spiders that have
11874:
11873:
11833:
11501:
11436:
11138:
10203:
10161:
10089:
9681:
9654:
9615:
8732:10.1146/annurev.ento.53.103106.093503
7677:
6752:Wilcox, S. & Jackson, R. (2002).
5537:. Blackwell Publishing. p. 168.
5376:A Spider's World: Senses and Behavior
5372:
5297:
5291:
4298:
3966:, although some species of the genus
2069:, a solid plant material produced by
826:would form two separate tagmata, the
12172:E6743423-ECE0-112E-7821-15817558C6C1
12133:8fe5dc6f-2d05-45f2-98ee-598d13b7fe9d
12032:c0be2190-1e35-453a-a53f-22a44bf8c472
11462:
9966:
9939:
9456:. Libraries Unlimited. p. 186.
7393:Thomson, Peggy & Park, Edwards.
6117:. Oxford University Press. pp.
4126:
3728:The order name Araneae derives from
3025:, had five spinnerets. Although the
2657:In experiments, spider species like
1345:, often by means of setae. An adult
1308:) with a pair of eyes absent on the
12223:Taxa named by Carl Alexander Clerck
9357:Journal of Insects as Food and Feed
9006:"Spider mythology across the world"
8289:10.1146/annurev.es.22.110191.003025
8086:Organisms Diversity & Evolution
7235:10.1146/annurev.es.21.110190.002013
7076:Behavioral Ecology and Sociobiology
6925:Behavioral Ecology and Sociobiology
6809:10.1146/annurev.en.38.010193.002031
5567:The Journal of Experimental Biology
5096:10.1146/annurev.es.22.110191.003025
3913:From sides to center, like pincers
3047:The main groups of modern spiders,
1328:As with other arthropods, spiders'
1168:running below the gut, with paired
918:Like other arthropods, spiders are
682:carapace of prosoma (cephalothorax)
24:
10556:(violin spiders, assassin spiders)
9817:
9690:. New York: Simon & Schuster.
9201:. phobias-help.com. Archived from
8866:"Pub chef bitten by deadly spider"
8579:"Natural history of Mygalomorphae"
8413:"The Role of Silk in Prey Capture"
7680:"Oldest spider web found in amber"
6823:"Different smiles, single species"
6071:10.24272/j.issn.2095-8137.2019.041
4679:10.1146/annurev-ento-061520-083414
4421:are depictions of spiderwebs. The
4368:In a story told by the Roman poet
3824:Segmented plates on top of abdomen
3089:According to a 2020 study using a
1455:
1416:, which acts as the hinge for the
25:
12234:
10016:to help reach a consensus. ›
9995:
9825:Bilger, Burkhard (5 March 2007).
8359:Leroy, J & Leroy, A. (2003).
5151:"Spiders-Arañas – Dr. Sam Thelin"
4465:List of animals that produce silk
4279:Spiders are used as food. Cooked
4132:Media coverage and misconceptions
2215:deinopids, but one population of
2167:
1841:Australian female trapdoor spider
1782:carrying its young on its abdomen
1594:but indirect, in other words the
1506:and biological materials such as
1412:; next is the spider's knee, the
1140:, retain the ancestral arthropod
1046:, the final sections of spiders'
850:ahead of the mouth are a pair of
11814:are possibly paraphyletic groups
11417:
11416:
11215:
10795:(sheet weavers or money spiders)
9847:. Taplinger Publishing Company.
9744:
9741:. New York: Praeger Press. 1972.
9731:
9704:
9648:
9533:
9470:
9443:
9416:
9391:
9317:
9284:
9254:
9220:
9191:
9177:10.1111/j.1467-7652.2004.00087.x
9156:
9089:
8997:
8948:
8880:
8858:
8846:. Australian Venom Research Unit
8836:
8809:
8768:
8746:
8630:
8597:
8571:
8546:
8521:
8481:
8404:
8381:
8311:
8168:
8066:10.1111/j.1096-3642.2007.00284.x
7728:arachnid had already been named
7718:The fossil was originally named
7657:10.1111/j.1365-2451.2007.00641.x
7395:"Odd Tales from the Smithsonian"
7270:10.1111/j.1096-3642.2004.00120.x
7023:10.1111/j.1744-7429.2006.00202.x
6980:10.1111/j.1365-3113.2005.00289.x
5968:Andrade, Maydianne C.B. (2003).
5869:10.1111/j.1744-7410.2011.00227.x
5298:Mason, Betsy (28 October 2021).
5136:10.1046/j.1365-2435.1997.00040.x
4439:
4227:
4216:are being made to produce it in
4075:
4016:
3646:
3510:
3489:
3341:
3319:
3284:
3256:
3218:
3188:
3157:
2855:
2547:
2445:
2428:
2407:
2385:
2373:
1968:or scales. The white prosoma of
1772:
1755:
1743:
1725:
1657:
1643:
1622:Spiders generally use elaborate
1548:" by the young of some species.
1404:; the next segment is the short
1017:Feeding, digestion and excretion
744:
662:
342:. They are the largest order of
159:
61:
10249:(Australian funnel-web spiders)
10064:List of field guides to spiders
9586:The Cambridge Classical Journal
9229:"Spider Fears or Arachnophobia"
9227:Fritscher, Lisa (3 June 2009).
8906:10.4269/ajtmh.2004.71.2.0700239
8235:American Arachnological Society
8124:
7939:
7874:
7694:
7671:
7517:
7495:
7438:
7408:
7386:
7339:
7304:
7277:
7141:
7102:
7067:
7037:
6994:
6951:
6916:
6897:
6860:
6837:
6815:
6788:
6691:
6671:Psyche: A Journal of Entomology
6654:
6589:
6549:
6504:
6458:
6412:
6283:Journal of Experimental Biology
6274:
6239:
6161:
6135:
6046:
5875:
5844:
5833:
5786:
5698:
5649:
5619:Journal of Experimental Biology
5603:
5522:
5512:
5501:
5405:
5393:
5366:
5354:
5251:
5143:
5102:
4888:
4212:is both light and very strong,
4066:Australasian funnel-web spiders
4056:). Well-known examples include
2414:A trapdoor spider in the genus
2065:gets over 90% of its food from
1540:and for fertilized eggs; as a "
1323:
1316:family can have as few as two.
1251:("little eyes"), which in most
1201:
899:, whose living members are the
834:; the rear tagma is called the
301:134 families, c. 52,000 species
9739:The Mochica: A Culture of Peru
8934:. UNSW Press. pp. 65–68.
8816:Hannum, C. & Miller, D.M.
7428:European Journal of Entomology
6904:Armstrong, S. (14 July 1990).
5756:Ruppert, Fox & Barnes 2004
5644:Ruppert, Fox & Barnes 2004
5535:The Invertebrates: A Synthesis
5361:Ruppert, Fox & Barnes 2004
5349:Ruppert, Fox & Barnes 2004
5198:Ruppert, Fox & Barnes 2004
5183:Ruppert, Fox & Barnes 2004
5064:Ruppert, Fox & Barnes 2004
5052:Ruppert, Fox & Barnes 2004
5037:Ruppert, Fox & Barnes 2004
5012:Ruppert, Fox & Barnes 2004
4929:Ruppert, Fox & Barnes 2004
4914:Ruppert, Fox & Barnes 2004
4831:
4749:
4542:10.1007/978-1-4020-6359-6_4320
4525:
4305:Cultural depictions of spiders
4031:A Mexican red-kneed tarantula
2794:took two orb-web spiders into
2601:with no other social species,
1544:"; for nest-building; and as "
13:
1:
10756:(flat-bellied ground spiders)
10233:(mouse spiders and relatives)
10001:
9801:(7th ed.). Brooks/Cole.
9711:De Laguna, Frederica (2002).
9127:10.1016/S0167-7799(00)01481-5
9075:10.1016/j.toxicon.2016.02.023
8781:Journal of Medical Entomology
8775:Vetter RS, Barger DK (2002).
7325:10.1016/j.anbehav.2003.06.011
7286:Australian Journal of Zoology
6625:10.1126/science.198.4322.1173
6524:Evolutionary Ecology Research
6217:10.1146/annurev.ento.43.1.619
5819:10.1080/07924259.1995.9672484
4513:. Natural History Museum Bern
4283:are considered a delicacy in
4271:are considered a delicacy in
4168:Australian funnel-web spiders
3919:
2920:period, and had a triangular
1879:
1862:), the largest spider by mass
1497:Silk is mainly composed of a
1387:
761:pedicel (also called pedicle)
11805:Classification is based on
10619:(araneomorph funnel weavers)
9919:; Kaston, Elizabeth (1953).
9769:General and cited references
8887:Diaz, J.H. (1 August 2004).
8555:The Tarantula Keeper's Guide
8530:The Tarantula Keeper's Guide
6171:, Golden Press, pp. 20, 44.
5373:Barth, Friedrich G. (2013).
4758:The Tarantula Keeper's Guide
4485:
3836:Striking direction of fangs
2999:spiders were members of the
2844:
2688:
2454:ant-mimicking jumping spider
2301:Some jumping spiders of the
1806:Baby spiders pass all their
1637:Spider fertilization systems
1237:Eyes of the jumping spider,
653:
591:
7:
11834:
11665:(shorttailed whipscorpions)
11254:List of families of spiders
11164:
10273:(cork-lid trapdoor spiders)
9940:Main, Barbara York (1975).
9513:Jacobsen, Thorkild (1987).
9235:. About.com. Archived from
9165:Plant Biotechnology Journal
9010:Western Journal of Medicine
8961:Western Journal of Medicine
8955:Nishioka, S. de A. (2001).
8719:Annual Review of Entomology
8490:"European Arachnology 2005"
6797:Annual Review of Entomology
6754:"Jumping Spider Tricksters"
6562:Marx (Araneae, Deinopidae)"
6530:(1): 105–17. Archived from
6205:Annual Review of Entomology
4667:Annual Review of Entomology
4432:
4160:) and widow spiders (genus
3717:
2776:most effective in catching
1651:Haplogyne or non-entelegyne
1560:Sexual selection in spiders
1552:Reproduction and life cycle
907:Circulation and respiration
10:
12239:
11502:
10569:(long-legged cave spiders)
10265:(brushed trapdoor spiders)
10241:(folding trapdoor spiders)
9969:Spiders in Ecological Webs
9893:. New York: Random House.
9540:Norton, Elizabeth (2013).
8794:10.1603/0022-2585-39.6.948
8604:Foelix, Rainer F. (2011).
8367:. Struik. pp. 15–21.
8365:Spiders of Southern Africa
8345:. Perseus Digital Library.
8326:. Perseus Digital Library.
8154:10.1016/j.zool.2005.06.003
8099:10.1007/s13127-019-00390-7
7599:The Journal of Arachnology
7471:10.1007/s00114-020-01708-8
7154:Royal Society Open Science
6425:The Journal of Arachnology
6248:The Journal of Arachnology
6111:Foelix, Rainer F. (1996).
6020:Foelix, Rainer F. (1996).
5768:Foelix, Rainer F. (2011).
5475:Foelix, Rainer F. (2011).
4723:Foelix, Rainer F. (1996).
4536:. Springer. p. 3496.
4534:Encyclopedia of Entomology
4470:List of endangered spiders
4302:
4256:
4252:
4231:
4201:
4144:
4079:
4020:
3923:
3749:
3740:
3721:
2848:
2836:Web design in zero gravity
2692:
2551:
2460:
2362:does not closely resemble
2221:appears to live mainly on
2171:
1557:
1482:. Each spinneret has many
1459:
1210:
973:spiders have two pairs of
954:transport more efficient.
646:
29:
11882:
11840:
11829:
11801:
11757:
11734:
11715:
11699:
11690:
11652:
11643:(tailless whip scorpions)
11636:
11627:
11611:
11602:
11571:
11544:
11533:
11512:
11508:
11497:
11471:
11412:
11391:
11360:
11267:
11224:
11213:
11172:
11113:
10611:
10602:
10587:Trogloraptor marchingtoni
10396:
10389:
10380:
10226:
10217:
10213:
10199:
10171:
10157:
10125:
9634:10.1017/S0048671X00004549
9598:10.1017/S006867350000523X
8665:10.1016/j.cub.2022.07.026
8222:Coddington, J.A. (2005).
7907:10.1007/s00114-013-1121-7
7127:10.1007/s10164-007-0082-8
6891:10.1163/15685390151074357
6846:American Museum Novitates
6467:"Plant-eating by spiders"
6391:10.1017/S095283690100108X
6347:. 93rd ESA Annual Meeting
5955:10.1017/S0952836901000954
5851:Burger, Matthias (2011).
5793:Suzuki, Hirohumi (1995).
5662:Journal of Insect Science
5313:10.1146/knowable-102821-1
5266:; Kamil, Alan C. (eds.).
4864:10.1007/s00114-017-1440-1
4808:10.1016/j.cub.2009.08.049
4062:ctenizid trapdoor spiders
3892:
3889:
3886:
3863:
3806:
3803:(numbers are approximate)
3799:
3640:
3625:
3618:
3591:
3576:
3569:
3559:
3544:
3537:
3505:
3483:
3476:
3466:
3459:
3441:
3431:
3334:
3309:
3302:
3282:
3272:
3245:
3237:
3210:
3203:
3182:
3175:
3151:
3144:
2801:
2709:(European garden spider).
2398:or leaf-curling spider's
1803:and dragging them along.
1799:or attaching them to the
1682:Parasteatoda tepidariorum
1035:Cheiracanthium punctorium
989:spiders, like the family
330:generally able to inject
299:
292:
253:
248:
156:Scientific classification
154:
137:
128:
41:
11063:(cribellate orb weavers)
10997:(long jawed orb-weavers)
10649:(anyphaenid sac spiders)
10445:(crevice weaver spiders)
10281:(wafer trapdoor spiders)
10014:templates for discussion
8497:Acta Zoologica Bulgarica
8415:. In Shear, W.A. (ed.).
8008:10.3389/fgene.2020.00182
5435:10.3897/zookeys.622.8682
5262:. In Balda, Russell P.;
5232:: 231–40. Archived from
4460:Glossary of spider terms
4331:ancient Sumerian goddess
4140:
3072:Sydney funnel-web spider
2822:
2471:Sydney funnel-web spider
2075:mutualistic relationship
1787:Females lay up to 3,000
756:
12213:Carboniferous arachnids
11259:Lists of spider species
10864:(red and black spiders)
10804:(liocranid sac spiders)
10548:(tube-dwelling spiders)
10484:(midget ground weavers)
10346:(funnel-web tarantulas)
10338:(tree trapdoor spiders)
10289:(funnel-web tarantulas)
10030:Aelurillus v-insignitus
9967:Wise, David A. (1993).
9887:Hillyard, Paul (1994).
9870:Crompton, John (1950).
9574:Harries, Byron (1990).
9338:10.1163/156853094X00045
9105:Trends in Biotechnology
9004:Isbister, G.K. (2001).
8343:A Greek-English Lexicon
7887:Die Naturwissenschaften
7767:10.1111/1475-4983.00068
7214:Eberhard, W.G. (1990).
6661:Eberhard, W.G. (1980).
6596:Eberhard, W.G. (1977).
5882:Rezác M (August 2009).
4595:10.1073/pnas.0809174106
4197:
4009:, so far found only in
3864:Downwards and forwards
3064: million years ago
3023: million years ago
2982: million years ago
2974:Mongolarachne jurassica
2939: million years ago
2931:Attercopus fimbriunguis
2914: million years ago
2881: million years ago
2772:presence of predators.
1846:
1206:
710:
626:
606:
497: million years ago
467: million years ago
322:that have eight limbs,
32:Spider (disambiguation)
10893:(running crab spiders)
10786:(white-tailed spiders)
10628:(tangled nest spiders)
10453:(large-clawed spiders)
10429:(false violin spiders)
9872:The Life of the Spider
9655:Haase, Donald (2008).
9400:Lonely Planet Cambodia
8973:10.1136/ewjm.174.4.240
8361:"How spiders function"
7850:10.1098/rsbl.2011.0228
7364:10.1006/anbe.1994.1066
7251:Agnarsson, I. (2004).
7057:Journal of Arachnology
7050:(Araneae, Uloboridae)"
7044:Matsumoto, T. (1998).
6570:Journal of Arachnology
6471:Journal of Arachnology
5900:10.1098/rspb.2009.0104
5400:Deeleman-Reinhold 2001
5084:Annu. Rev. Ecol. Syst.
4322:
4276:
4099:
4037:
3943:
3412:Internal relationships
3101:External relationships
3055:, first appear in the
2868:
2814:Members of the family
2811:
2748:
2733:
2710:
2533:Carparachne aureoflava
2478:
2053:
1884:Only three classes of
1863:
1735:Trichonephila clavipes
1693:traumatic insemination
1587:
1471:
1397:
1243:
1230:
1164:consists of a pair of
1162:central nervous system
1156:Central nervous system
1039:
1026:
915:
698:PL (posterior lateral)
672:female, dorsal aspect
643:Anatomy and physiology
634:to draw, stretch, spin
415:, spiders do not have
370:have been recorded by
12154:Paleobiology Database
11524:(microwhip scorpions)
10921:(nursery web spiders)
10856:(cave cobweb spiders)
10848:(spurred orb-weavers)
10765:(dwarf sheet spiders)
10721:(net-casting spiders)
10512:(palp-footed spiders)
10354:(bald-legged spiders)
9450:De Vos, Gail (1996).
9369:10.3920/JIFF2020.0046
9022:10.1136/ewjm.175.2.86
8047:Shultz, J.W. (2007).
7995:Frontiers in Genetics
7736:Dunlop, J.A. (1999).
7701:Dunlop, J.A. (1996).
7553:10.5733/afin.050.0103
7531:African Invertebrates
7451:The Science of Nature
6960:Systematic Entomology
6720:10.1007/s004420050347
6665:Mastophora dizzydeani
6169:Spiders and their Kin
5989:10.1093/beheco/arg015
4844:The Science of Nature
4633:. Universities Press/
4397:are prominent in the
4312:
4266:
4089:
4030:
3993:The extinct families
3933:
3775:. Some 50,356 living
3243:Non‑pulmonates
2950:, which are modified
2863:
2809:
2747:, a golden orb weaver
2741:
2703:The large orb web of
2702:
2495:disruptive coloration
2468:
2380:Crab spider with prey
2043:
2036:Non-predatory feeding
2010:, but if it consumes
1854:
1764:Gasteracantha mammosa
1580:
1558:Further information:
1469:
1395:
1236:
1220:
1097:Most spiders convert
1054:into prey from venom
1032:
1024:
914:
701:PM (posterior median)
692:AL (anterior lateral)
685:opisthosoma (abdomen)
12027:Fauna Europaea (new)
11706:(hooded tickspiders)
11006:(tangle-web spiders)
10773:(tree trunk spiders)
10729:(intertidal spiders)
10658:(orb-weaver spiders)
10037:28 July 2013 at the
9874:. New York: Mentor.
9845:The World of Spiders
9798:Invertebrate Zoology
9780:. Brill Publishers.
9682:Garai, Jana (1973).
8868:. BBC. 27 April 2005
8844:"Funnel web spiders"
8411:Coyle, F.A. (1986).
7505:(10 December 2020).
7048:Philoponella raffray
6437:10.1636/JoA-S-22-015
5857:Invertebrate Biology
5674:10.1673/031.009.6201
5631:10.1242/jeb.36.2.423
5264:Pepperberg, Irene M.
5239:on 28 September 2006
4511:World Spider Catalog
4291:Indians of southern
3887:Only in some fossils
3032:299 to 251
2864:Spider preserved in
2719:Convergent evolution
2666:Latrodectus hesperus
2469:Threat display by a
2395:Phonognatha graeffei
2031:Ecology and behavior
2024:Cyrtophora cicatrosa
1936:. In genera such as
1923:Micrommata virescens
1691:, which has evolved
1610:-styled structures,
1581:Mating behaviour of
1525:Some spiders have a
1310:anterior median line
1160:The basic arthropod
1101:waste products into
695:AM (anterior median)
476:318 to 299
447:with silk-producing
392:or prosoma, and the
318:) are air-breathing
30:For other uses, see
11373:Cultural depictions
10979:(dwarf orb-weavers)
10466:(lampshade spiders)
10437:(woodlouse hunters)
10186:(segmented spiders)
10178:(segmented spiders)
10056:12 May 2011 at the
9944:. Sydney: Collins.
9737:Benson, Elizabeth.
9686:The Book of Symbols
9326:Society and Animals
9268:on 3 September 2009
9144:on 16 December 2008
9067:2016Txcn..114...40S
8656:2022CBio...32.R871M
8585:on 26 December 2008
8506:on 16 December 2008
8188:1989wlbs.book.....G
8146:2006Zool..109....2S
7960:1996Natur.379..498S
7899:2013NW....100.1171S
7758:1999Palgy..42..191D
7649:2007GeolT..23..231P
7544:2009AfrIn..50..105S
7463:2021SciNa.108....1Z
7419:Drapetisca socialis
7415:Schütt, K. (1995).
7174:10.1098/rsos.210279
7166:2021RSOS....810279F
7115:Journal of Ethology
7015:2006Biotr..38..743A
6972:2005SysEn..30..575A
6712:1997Oecol.112..572Y
6617:1977Sci...198.1173E
6537:on 16 December 2008
6515:Argyroneta aquatica
6059:Zoological Research
5811:1995InvRD..28..211S
5719:2001Natur.410..541V
5426:2016ZooK..622...47B
5161:on 21 February 2017
5127:1997FuEco..11....4B
4856:2017SciNa.104...30N
4799:2009CBio...19.R892M
4586:2008PNAS..10520781S
4222:genetic engineering
4034:Brachypelma hamorii
3812:Suborder/Infraorder
3093:calibrated with 27
2819:take several days.
2510:Many of the family
2500:Theridion grallator
2185:Argyroneta aquatica
1999:Theridion grallator
1688:Harpactea sadistica
1518:, but is much more
945:respiratory pigment
937:circulatory systems
873:that are generally
779:posterior spinneret
384:are fused into two
11579:Pseudoscorpionida
11013:Theridiosomatidae
10977:Symphytognathidae
10965:(huntsman spiders)
10872:(disc web spiders)
10690:(dark sac spiders)
10540:(spitting spiders)
10325:(dwarf tarantulas)
8824:on 18 October 2008
8608:Biology of Spiders
8324:A Latin Dictionary
8254:on 9 December 2008
8237:. pp. 18–24.
7574:2011-08-10 at the
7088:10.1007/BF00183313
6937:10.1007/BF00300005
6767:. pp. 27–34.
6684:10.1155/1980/81062
6295:10.1242/jeb.039743
6114:Biology of Spiders
6026:Biology of Spiders
5975:Behavioral Ecology
5942:Journal of Zoology
5894:(1668): 2697–701.
5770:Biology of Spiders
5580:10.1242/jeb.054585
5479:Biology of Spiders
5270:. Academic Press.
5115:Functional Ecology
4727:Biology of Spiders
4425:people of ancient
4389:Stories about the
4323:
4299:Spiders in culture
4277:
4100:
4038:
3944:
3939:, a member of the
3187:(horseshoe crabs)
2869:
2812:
2749:
2711:
2706:Araneus diadematus
2672:Eratigena agrestis
2618:Agelena consociata
2505:warning coloration
2479:
2077:with a species of
2054:
1929:Araneus diadematus
1864:
1588:
1564:Spider cannibalism
1472:
1398:
1359:Evarcha culicivora
1244:
1240:Plexippus paykulli
1231:
1107:Malphigian tubules
1040:
1038:, displaying fangs
1027:
916:
776:anterior spinneret
758:sternum of prosoma
752:Palystes castaneus
670:Palystes castaneus
455:) appeared in the
441:orb-weaver spiders
27:Order of arachnids
12195:
12194:
12141:Open Tree of Life
11876:Taxon identifiers
11867:
11866:
11863:
11862:
11825:
11824:
11821:
11820:
11797:
11796:
11793:
11792:
11780:
11767:Opilioacariformes
11727:
11707:
11686:
11685:
11682:
11681:
11674:
11666:
11644:
11619:
11598:
11597:
11590:
11582:
11581:(pseudoscorpions)
11563:
11555:
11525:
11430:
11429:
11361:Human interaction
11132:
11131:
11109:
11108:
11105:
11104:
11101:
11100:
11097:
11096:
11083:
11064:
11025:
11016:
11007:
10998:
10980:
10966:
10957:
10943:
10942:(jumping spiders)
10930:
10922:
10894:
10881:
10873:
10865:
10857:
10849:
10836:
10823:
10814:
10805:
10796:
10787:
10774:
10766:
10757:
10743:
10730:
10722:
10699:
10691:
10683:
10659:
10650:
10637:
10636:(termite hunters)
10629:
10620:
10598:
10597:
10590:
10583:Trogloraptoridae
10578:
10577:(armored spiders)
10570:
10557:
10549:
10541:
10527:
10513:
10494:
10485:
10477:Mecysmaucheniidae
10467:
10454:
10446:
10438:
10430:
10422:
10421:(coneweb spiders)
10404:
10403:(pelican spiders)
10376:
10375:
10368:
10367:(true tarantulas)
10355:
10347:
10339:
10326:
10323:Mecicobothriidae
10308:
10290:
10282:
10274:
10266:
10258:
10250:
10242:
10234:
10195:
10194:
10187:
10179:
9978:978-0-521-32547-9
9951:978-0-00-211443-1
9900:978-0-679-40881-9
9854:978-0-8008-8598-4
9835:. pp. 66–73.
9808:978-0-03-025982-1
9759:Thames and Hudson
9724:978-0-8032-8280-3
9697:978-0-671-21773-0
9668:978-0-313-33441-2
9553:978-1-4438-4271-6
9526:978-0-300-07278-5
9488:978-0-7141-1705-8
9463:978-1-56308-190-3
9436:978-0-452-28700-6
9423:Weil, C. (2006).
9409:978-1-74059-111-9
9310:978-1-4129-2568-6
9299:. Sage. pp.
9199:"A Common Phobia"
8941:978-0-86840-279-6
8650:(16): R871–R873.
8623:978-0-19-973482-5
8564:978-0-7641-3885-0
8539:978-0-7641-3885-0
8430:978-0-8047-1203-3
8374:978-1-86872-944-9
8244:978-0-9771439-0-0
8197:978-0-09-174271-3
7893:(12): 1171–1181.
7611:10.1636/CH07-89.1
6774:978-0-262-52322-6
6667:sp. n. (Araneae)"
6611:(4322): 1173–75.
6560:Deinopis spinosus
6342:Bagheera kiplingi
6128:978-0-674-07431-6
6039:978-0-19-509594-4
5779:978-0-19-973482-5
5758:, pp. 537–39
5646:, pp. 325–49
5544:978-0-632-04761-1
5494:978-0-19-973482-5
5363:, pp. 578–80
5351:, pp. 532–37
5304:Knowable Magazine
5277:978-0-12-077030-4
5200:, pp. 531–32
5185:, pp. 529–30
5066:, pp. 527–28
5054:, pp. 565–69
5039:, pp. 559–64
5014:, pp. 571–84
4931:, pp. 518–22
4916:, pp. 554–55
4767:978-0-7641-3885-0
4742:978-0-19-509593-7
4644:978-81-7371-641-6
4551:978-1-4020-6242-1
4447:Arthropods portal
4353:her. Enki's wife
4127:Human interaction
4120:Bagheera kiplingi
3917:
3916:
3804:
3714:
3713:
3705:
3704:
3696:
3695:
3687:
3686:
3678:
3677:
3669:
3668:
3660:
3659:
3604:
3603:
3523:
3522:
3408:
3407:
3399:
3398:
3390:
3389:
3381:
3380:
3372:
3371:
3363:
3362:
3354:
3353:
3034:million years ago
3007:. The mesothelid
2783:Zygiella x-notata
2675:stayed away from
2645:Bagheera kiplingi
2603:T. nigroannulatum
2514:, which includes
2200:Theridiosomatidae
2182:The water spider
2062:Bagheera kiplingi
1962:Structural colors
1860:Theraphosa blondi
1856:Goliath birdeater
1578:
1305:Periegops suterii
1260:of the ancestral
965:system, or both.
508:was described as
505:Bagheera kiplingi
478:million years ago
306:
305:
244:
16:(Redirected from
12230:
12188:
12187:
12175:
12174:
12162:
12161:
12149:
12148:
12136:
12135:
12126:
12125:
12113:
12112:
12110:NHMSYS0000841308
12100:
12099:
12087:
12086:
12074:
12073:
12061:
12060:
12048:
12047:
12035:
12034:
12022:
12021:
12009:
12008:
11996:
11995:
11983:
11982:
11970:
11969:
11957:
11956:
11944:
11943:
11931:
11930:
11918:
11917:
11916:
11903:
11902:
11901:
11871:
11870:
11850:Phalangiotarbida
11831:
11830:
11813:
11778:
11732:
11731:
11726:
11725:
11721:
11719:
11705:
11697:
11696:
11693:
11672:
11664:
11642:
11634:
11633:
11617:
11609:
11608:
11588:
11580:
11561:
11553:
11542:
11541:
11537:
11523:
11510:
11509:
11499:
11498:
11457:
11450:
11443:
11434:
11433:
11420:
11419:
11219:
11159:
11152:
11145:
11136:
11135:
11081:
11062:
11023:
11014:
11005:
10996:
10978:
10964:
10955:
10941:
10924:
10920:
10892:
10879:
10871:
10863:
10855:
10847:
10835:(pirate spiders)
10834:
10822:(shield spiders)
10821:
10812:
10803:
10794:
10785:
10772:
10764:
10755:
10742:(velvet spiders)
10741:
10728:
10720:
10697:
10689:
10681:
10670:Cheiracanthiidae
10657:
10648:
10635:
10627:
10618:
10609:
10608:
10584:
10576:
10568:
10555:
10547:
10539:
10526:(cellar spiders)
10525:
10511:
10493:(goblin spiders)
10492:
10483:
10482:Ochyroceratidae
10465:
10452:
10444:
10436:
10428:
10420:
10402:
10394:
10393:
10387:
10386:
10366:
10353:
10345:
10337:
10331:Microstigmatidae
10324:
10306:
10288:
10280:
10279:Cyrtaucheniidae
10272:
10264:
10256:
10248:
10240:
10232:
10224:
10223:
10215:
10214:
10201:
10200:
10185:
10177:
10159:
10158:
10110:
10103:
10096:
10087:
10086:
10072:
9990:
9963:
9936:
9926:
9912:
9883:
9866:
9836:
9812:
9791:
9762:
9748:
9742:
9735:
9729:
9728:
9708:
9702:
9701:
9689:
9679:
9673:
9672:
9652:
9646:
9645:
9613:
9602:
9601:
9583:
9571:
9558:
9557:
9537:
9531:
9530:
9510:
9493:
9492:
9474:
9468:
9467:
9447:
9441:
9440:
9420:
9414:
9413:
9398:Ray, N. (2002).
9395:
9389:
9388:
9348:
9342:
9341:
9321:
9315:
9314:
9298:
9288:
9282:
9281:
9275:
9273:
9258:
9252:
9251:
9246:
9244:
9224:
9218:
9217:
9212:
9210:
9195:
9189:
9188:
9160:
9154:
9153:
9151:
9149:
9143:
9137:. Archived from
9120:
9102:
9093:
9087:
9086:
9050:
9044:
9043:
9033:
9001:
8995:
8994:
8984:
8952:
8946:
8945:
8925:
8919:
8918:
8908:
8884:
8878:
8877:
8875:
8873:
8862:
8856:
8855:
8853:
8851:
8840:
8834:
8833:
8831:
8829:
8813:
8807:
8806:
8796:
8772:
8766:
8765:
8763:
8761:
8750:
8744:
8743:
8713:
8704:
8703:
8677:
8667:
8634:
8628:
8627:
8611:
8601:
8595:
8594:
8592:
8590:
8575:
8569:
8568:
8550:
8544:
8543:
8525:
8519:
8518:
8513:
8511:
8505:
8494:
8485:
8479:
8478:
8476:
8474:
8469:
8461:
8441:
8435:
8434:
8408:
8402:
8401:
8388:Ono, H. (2002).
8385:
8379:
8378:
8356:
8347:
8346:
8334:
8328:
8327:
8315:
8309:
8308:
8270:
8264:
8263:
8261:
8259:
8253:
8247:. Archived from
8228:
8219:
8202:
8201:
8172:
8166:
8165:
8128:
8122:
8121:
8111:
8101:
8077:
8071:
8070:
8068:
8044:
8031:
8030:
8020:
8010:
7986:
7980:
7979:
7968:10.1038/379498b0
7954:(6565): 498–99.
7943:
7937:
7936:
7926:
7878:
7872:
7871:
7861:
7829:
7823:
7822:
7804:
7795:
7780:
7779:
7769:
7717:
7707:
7698:
7692:
7691:
7689:
7687:
7675:
7669:
7668:
7632:
7623:
7622:
7590:
7581:
7567:
7565:
7555:
7521:
7515:
7514:
7499:
7493:
7492:
7482:
7442:
7436:
7435:
7425:
7412:
7406:
7405:
7403:
7401:
7390:
7384:
7383:
7352:Animal Behaviour
7343:
7337:
7336:
7313:Animal Behaviour
7308:
7302:
7301:
7281:
7275:
7274:
7272:
7248:
7239:
7238:
7220:
7211:
7196:
7195:
7185:
7145:
7139:
7138:
7106:
7100:
7099:
7071:
7065:
7064:
7054:
7041:
7035:
7034:
6998:
6992:
6991:
6955:
6949:
6948:
6920:
6914:
6913:
6901:
6895:
6894:
6884:
6864:
6858:
6857:
6841:
6835:
6834:
6832:
6830:
6819:
6813:
6812:
6792:
6786:
6785:
6783:
6781:
6758:
6749:
6740:
6739:
6695:
6689:
6688:
6686:
6658:
6652:
6651:
6649:
6647:
6602:
6593:
6587:
6586:
6584:
6582:
6566:
6553:
6547:
6546:
6544:
6542:
6536:
6521:
6508:
6502:
6501:
6499:
6497:
6483:10.1636/P15-45.1
6462:
6456:
6455:
6453:
6451:
6416:
6410:
6409:
6407:
6405:
6400:on 18 March 2009
6399:
6393:. Archived from
6375:
6366:
6357:
6356:
6354:
6352:
6334:
6323:
6322:
6278:
6272:
6271:
6243:
6237:
6236:
6200:
6179:
6165:
6159:
6158:
6156:
6154:
6139:
6133:
6132:
6108:
6102:
6099:
6093:
6092:
6082:
6050:
6044:
6043:
6017:
6004:
6003:
6001:
5991:
5965:
5959:
5958:
5928:
5922:
5921:
5911:
5879:
5873:
5872:
5848:
5842:
5837:
5831:
5830:
5790:
5784:
5783:
5765:
5759:
5753:
5747:
5746:
5727:10.1038/35069000
5713:(6828): 541–48.
5702:
5696:
5695:
5685:
5653:
5647:
5641:
5635:
5634:
5616:
5607:
5601:
5600:
5582:
5558:
5549:
5548:
5526:
5520:
5516:
5510:
5505:
5499:
5498:
5482:
5472:
5466:
5465:
5455:
5437:
5409:
5403:
5397:
5391:
5390:
5370:
5364:
5358:
5352:
5346:
5333:
5332:
5330:
5328:
5315:
5295:
5289:
5288:
5286:
5284:
5255:
5249:
5248:
5246:
5244:
5238:
5223:
5214:
5201:
5195:
5186:
5180:
5171:
5170:
5168:
5166:
5157:. Archived from
5155:Drsamchapala.com
5147:
5141:
5140:
5138:
5106:
5100:
5099:
5078:
5067:
5061:
5055:
5049:
5040:
5034:
5015:
5009:
4932:
4926:
4917:
4911:
4900:
4899:
4892:
4886:
4885:
4875:
4835:
4829:
4828:
4810:
4778:
4772:
4771:
4753:
4747:
4746:
4730:
4720:
4714:
4713:
4711:
4709:
4658:
4649:
4648:
4635:Orient Blackswan
4630:Spiders of India
4624:
4618:
4617:
4607:
4597:
4580:(52): 20781–85.
4565:
4556:
4555:
4529:
4523:
4522:
4520:
4518:
4503:
4449:
4444:
4443:
4092:Leucauge venusta
3950:, found only in
3936:Ryuthela sasakii
3802:
3800:Spider diversity
3794:
3793:
3752:
3751:
3743:
3742:
3651:
3650:
3621:
3620:
3572:
3571:
3562:
3561:
3540:
3539:
3514:
3494:
3493:
3479:
3478:
3469:
3468:
3462:
3461:
3434:
3433:
3424:
3423:
3345:
3323:
3305:
3304:
3288:
3275:
3274:
3260:
3240:
3239:
3223:
3222:
3216:(sea scorpions)
3206:
3205:
3192:
3178:
3177:
3161:
3147:
3146:
3137:
3136:
3065:
3035:
3024:
3014:, from the Late
2984:, recorded from
2983:
2958:and the similar
2940:
2915:
2882:
2851:Spider evolution
2611:species (family
2524:urticating hairs
2449:
2432:
2411:
2389:
2377:
2316:
2271:ambush predators
2218:Deinopis subrufa
1776:
1759:
1747:
1729:
1661:
1647:
1579:
1492:Spitting spiders
767:book lung stigma
748:
666:
638:
635:
632:
629:
622:
619:
616:
609:
498:
479:
468:
239:
164:
163:
133:
123:
60:
45:Temporal range:
39:
38:
21:
12238:
12237:
12233:
12232:
12231:
12229:
12228:
12227:
12198:
12197:
12196:
12191:
12183:
12178:
12170:
12165:
12157:
12152:
12144:
12139:
12131:
12129:
12121:
12116:
12108:
12103:
12095:
12090:
12082:
12077:
12069:
12064:
12056:
12051:
12043:
12038:
12030:
12025:
12017:
12012:
12004:
11999:
11991:
11986:
11978:
11973:
11965:
11960:
11952:
11947:
11939:
11934:
11926:
11921:
11912:
11911:
11906:
11897:
11896:
11891:
11878:
11868:
11859:
11836:
11817:
11811:
11789:
11753:
11723:
11722:
11720:
11717:
11711:
11691:
11678:
11648:
11623:
11594:
11589:(camel spiders)
11567:
11535:
11529:
11504:
11493:
11467:
11461:
11431:
11426:
11408:
11404:Web decorations
11387:
11383:Spider fighting
11356:
11352:Urticating hair
11302:Cheliceral fang
11263:
11220:
11211:
11168:
11163:
11133:
11128:
11118:Spider taxonomy
11093:
11051:Trochanteriidae
10995:Tetragnathidae
10748:Gallieniellidae
10594:
10575:Tetrablemmidae
10390:Non-entelegynes
10372:
10352:Paratropididae
10300:Halonoproctidae
10239:Antrodiaetidae
10209:
10191:
10167:
10153:
10121:
10114:
10070:
10058:Wayback Machine
10039:Wayback Machine
10017:
9998:
9993:
9979:
9952:
9901:
9855:
9841:Bristowe, W. S.
9820:
9818:Further reading
9815:
9809:
9788:
9771:
9766:
9765:
9749:
9745:
9736:
9732:
9725:
9709:
9705:
9698:
9680:
9676:
9669:
9653:
9649:
9614:
9605:
9572:
9561:
9554:
9538:
9534:
9527:
9511:
9496:
9489:
9475:
9471:
9464:
9448:
9444:
9437:
9421:
9417:
9410:
9396:
9392:
9349:
9345:
9322:
9318:
9311:
9289:
9285:
9271:
9269:
9260:
9259:
9255:
9242:
9240:
9239:on 19 June 2009
9225:
9221:
9208:
9206:
9205:on 25 June 2009
9197:
9196:
9192:
9161:
9157:
9147:
9145:
9141:
9100:
9094:
9090:
9051:
9047:
9002:
8998:
8953:
8949:
8942:
8926:
8922:
8885:
8881:
8871:
8869:
8864:
8863:
8859:
8849:
8847:
8842:
8841:
8837:
8827:
8825:
8818:"Widow Spiders"
8814:
8810:
8773:
8769:
8759:
8757:
8752:
8751:
8747:
8714:
8707:
8644:Current Biology
8635:
8631:
8624:
8602:
8598:
8588:
8586:
8577:
8576:
8572:
8565:
8551:
8547:
8540:
8526:
8522:
8509:
8507:
8503:
8492:
8486:
8482:
8472:
8470:
8467:
8463:
8442:
8438:
8431:
8409:
8405:
8386:
8382:
8375:
8357:
8350:
8335:
8331:
8316:
8312:
8271:
8267:
8257:
8255:
8251:
8245:
8226:
8220:
8205:
8198:
8173:
8169:
8129:
8125:
8078:
8074:
8045:
8034:
7987:
7983:
7944:
7940:
7879:
7875:
7838:Biology Letters
7830:
7826:
7802:
7796:
7783:
7705:
7699:
7695:
7685:
7683:
7682:. New Scientist
7676:
7672:
7633:
7626:
7591:
7584:
7576:Wayback Machine
7522:
7518:
7503:Dvorsky, George
7500:
7496:
7443:
7439:
7423:
7413:
7409:
7399:
7397:
7391:
7387:
7344:
7340:
7309:
7305:
7298:10.1071/ZO00007
7282:
7278:
7249:
7242:
7218:
7212:
7199:
7146:
7142:
7107:
7103:
7072:
7068:
7052:
7042:
7038:
6999:
6995:
6956:
6952:
6921:
6917:
6902:
6898:
6882:10.1.1.512.3868
6865:
6861:
6842:
6838:
6828:
6826:
6821:
6820:
6816:
6793:
6789:
6779:
6777:
6775:
6756:
6750:
6743:
6696:
6692:
6677:(3–4): 143–70.
6659:
6655:
6645:
6643:
6600:
6594:
6590:
6580:
6578:
6564:
6554:
6550:
6540:
6538:
6534:
6519:
6509:
6505:
6495:
6493:
6463:
6459:
6449:
6447:
6417:
6413:
6403:
6401:
6397:
6373:
6367:
6360:
6350:
6348:
6335:
6326:
6279:
6275:
6244:
6240:
6201:
6182:
6166:
6162:
6152:
6150:
6149:. 30 April 2018
6141:
6140:
6136:
6129:
6109:
6105:
6101:Ruppert, 523–24
6100:
6096:
6051:
6047:
6040:
6018:
6007:
5966:
5962:
5929:
5925:
5888:Proc. Biol. Sci
5880:
5876:
5849:
5845:
5838:
5834:
5791:
5787:
5780:
5766:
5762:
5754:
5750:
5703:
5699:
5654:
5650:
5642:
5638:
5614:
5608:
5604:
5559:
5552:
5545:
5527:
5523:
5517:
5513:
5506:
5502:
5495:
5473:
5469:
5410:
5406:
5398:
5394:
5387:
5371:
5367:
5359:
5355:
5347:
5336:
5326:
5324:
5296:
5292:
5282:
5280:
5278:
5256:
5252:
5242:
5240:
5236:
5221:
5215:
5204:
5196:
5189:
5181:
5174:
5164:
5162:
5149:
5148:
5144:
5107:
5103:
5079:
5070:
5062:
5058:
5050:
5043:
5035:
5018:
5010:
4935:
4927:
4920:
4912:
4903:
4894:
4893:
4889:
4836:
4832:
4793:(19): R892–93.
4787:Current Biology
4779:
4775:
4768:
4754:
4750:
4743:
4721:
4717:
4707:
4705:
4659:
4652:
4645:
4625:
4621:
4566:
4559:
4552:
4530:
4526:
4516:
4514:
4505:
4504:
4493:
4488:
4475:Spider taxonomy
4445:
4438:
4435:
4307:
4301:
4261:
4255:
4243:specific phobia
4236:
4230:
4206:
4200:
4154:recluse spiders
4149:
4143:
4134:
4129:
4115:jumping spiders
4104:orb-web spiders
4084:
4078:
4025:
4019:
4007:Arthromygalidae
3995:Arthrolycosidae
3928:
3922:
3898:Opisthothelae:
3801:
3726:
3724:Spider taxonomy
3720:
3715:
3706:
3697:
3688:
3679:
3670:
3661:
3645:
3605:
3566:Austrochiloidea
3524:
3488:
3414:
3409:
3400:
3391:
3382:
3373:
3364:
3355:
3217:
3129:and, in males,
3103:
3091:molecular clock
3060:
3038:diversification
3030:
3019:
2978:
2935:
2910:
2877:
2858:
2853:
2847:
2838:
2825:
2804:
2744:Nephila clavata
2736:
2697:
2691:
2660:Steatoda grossa
2640:kleptoparasites
2556:
2550:
2491:parasitic wasps
2463:
2456:
2450:
2441:
2433:
2424:
2422:ambush predator
2412:
2403:
2390:
2381:
2378:
2319:trial-and-error
2314:
2292:fishing spiders
2288:jumping spiders
2198:and the family
2176:
2170:
2038:
2033:
1882:
1849:
1783:
1777:
1768:
1760:
1751:
1748:
1739:
1730:
1676:
1675:
1674:
1673:
1669:
1668:
1667:
1662:
1654:
1653:
1648:
1639:
1638:
1629:jumping spiders
1584:Neriene radiata
1568:
1566:
1554:
1464:
1458:
1456:Silk production
1390:
1326:
1266:tapetum lucidum
1215:
1209:
1204:
1180:, of which the
1158:
1136:and infraorder
1042:Uniquely among
1019:
1007:metabolic rates
909:
794:and therefore,
788:
787:
786:
785:
784:
782:
770:epigastric fold
749:
741:
740:
739:
707:
704:
667:
656:
651:
645:
636:
633:
630:
620:
617:
614:
594:
520:it with sticky
493:
474:
463:
404:(head) and the
284:Spider taxonomy
259:
238:
158:
124:
122:
121:
116:
111:
106:
101:
96:
91:
86:
81:
76:
71:
66:
55:
54:
43:
35:
28:
23:
22:
15:
12:
11:
5:
12236:
12226:
12225:
12220:
12215:
12210:
12193:
12192:
12190:
12189:
12176:
12163:
12150:
12137:
12127:
12114:
12101:
12088:
12075:
12062:
12049:
12036:
12023:
12014:Fauna Europaea
12010:
11997:
11984:
11971:
11958:
11945:
11932:
11919:
11904:
11888:
11886:
11880:
11879:
11865:
11864:
11861:
11860:
11858:
11857:
11855:Trigonotarbida
11852:
11847:
11841:
11838:
11837:
11827:
11826:
11823:
11822:
11819:
11818:
11816:
11815:
11812:Items in green
11809:
11802:
11799:
11798:
11795:
11794:
11791:
11790:
11788:
11787:
11782:
11774:
11769:
11763:
11761:
11759:Parasitiformes
11755:
11754:
11752:
11751:
11749:Trombidiformes
11746:
11744:Sarcoptiformes
11740:
11738:
11729:
11724:(ticks, mites)
11713:
11712:
11710:
11709:
11700:
11694:
11688:
11687:
11684:
11683:
11680:
11679:
11677:
11676:
11668:
11659:
11657:
11650:
11649:
11647:
11646:
11637:
11631:
11625:
11624:
11622:
11621:
11612:
11606:
11604:Tetrapulmonata
11600:
11599:
11596:
11595:
11593:
11592:
11584:
11575:
11573:
11569:
11568:
11566:
11565:
11557:
11548:
11546:
11539:
11531:
11530:
11528:
11527:
11518:
11516:
11514:Incertae sedis
11506:
11505:
11495:
11494:
11492:
11491:
11485:
11479:
11472:
11469:
11468:
11460:
11459:
11452:
11445:
11437:
11428:
11427:
11425:
11424:
11413:
11410:
11409:
11407:
11406:
11401:
11395:
11393:
11389:
11388:
11386:
11385:
11380:
11375:
11370:
11364:
11362:
11358:
11357:
11355:
11354:
11349:
11344:
11339:
11334:
11329:
11324:
11319:
11314:
11309:
11304:
11299:
11294:
11289:
11284:
11279:
11273:
11271:
11265:
11264:
11262:
11261:
11256:
11251:
11246:
11241:
11236:
11230:
11228:
11222:
11221:
11214:
11212:
11210:
11209:
11204:
11202:Classification
11199:
11194:
11189:
11184:
11178:
11176:
11170:
11169:
11162:
11161:
11154:
11147:
11139:
11130:
11129:
11127:
11126:
11120:
11114:
11111:
11110:
11107:
11106:
11103:
11102:
11099:
11098:
11095:
11094:
11092:
11091:
11086:
11076:
11071:
11066:
11058:
11053:
11048:
11043:
11038:
11033:
11028:
11024:(crab spiders)
11018:
11010:
11000:
10992:
10987:
10982:
10974:
10969:
10959:
10956:(bark hunters)
10951:
10946:
10936:
10931:
10916:
10911:
10906:
10901:
10899:Phrurolithidae
10896:
10891:Philodromidae
10888:
10883:
10880:(lynx spiders)
10875:
10867:
10859:
10851:
10843:
10838:
10830:
10828:Megadictynidae
10825:
10817:
10813:(wolf spiders)
10807:
10799:
10789:
10781:
10779:Homalonychidae
10776:
10768:
10760:
10750:
10745:
10737:
10732:
10724:
10716:
10711:
10706:
10701:
10693:
10685:
10677:
10672:
10667:
10662:
10652:
10644:
10639:
10631:
10623:
10612:
10606:
10600:
10599:
10596:
10595:
10593:
10592:
10580:
10572:
10564:
10559:
10551:
10543:
10535:
10530:
10520:
10515:
10507:
10502:
10497:
10487:
10479:
10474:
10469:
10461:
10456:
10451:Gradungulidae
10448:
10440:
10432:
10424:
10416:
10411:
10409:Austrochilidae
10406:
10397:
10391:
10384:
10378:
10377:
10374:
10373:
10371:
10370:
10365:Theraphosidae
10362:
10360:Porrhothelidae
10357:
10349:
10341:
10333:
10328:
10320:
10315:
10310:
10302:
10297:
10292:
10284:
10276:
10268:
10260:
10252:
10244:
10236:
10231:Actinopodidae
10227:
10221:
10211:
10210:
10197:
10196:
10193:
10192:
10190:
10189:
10184:Heptathelidae
10181:
10172:
10169:
10168:
10155:
10154:
10152:
10151:
10145:
10139:
10133:
10126:
10123:
10122:
10113:
10112:
10105:
10098:
10090:
10084:
10083:
10078:
10067:
10061:
10047:
10042:
10025:
9997:
9996:External links
9994:
9992:
9991:
9977:
9964:
9950:
9937:
9913:
9899:
9884:
9867:
9853:
9837:
9832:The New Yorker
9827:"Spider Woman"
9821:
9819:
9816:
9814:
9813:
9807:
9792:
9787:978-9004119598
9786:
9772:
9770:
9767:
9764:
9763:
9743:
9730:
9723:
9703:
9696:
9674:
9667:
9647:
9603:
9559:
9552:
9532:
9525:
9494:
9487:
9469:
9462:
9442:
9435:
9415:
9408:
9390:
9363:(3): 261–279.
9343:
9316:
9309:
9283:
9253:
9219:
9190:
9155:
9118:10.1.1.682.313
9088:
9045:
8996:
8947:
8940:
8920:
8879:
8857:
8835:
8808:
8767:
8745:
8705:
8629:
8622:
8596:
8570:
8563:
8545:
8538:
8520:
8480:
8436:
8429:
8403:
8380:
8373:
8348:
8329:
8310:
8265:
8243:
8203:
8196:
8167:
8123:
8072:
8032:
7981:
7938:
7873:
7824:
7781:
7693:
7670:
7624:
7582:
7516:
7494:
7437:
7407:
7385:
7338:
7303:
7276:
7263:(4): 447–626.
7240:
7197:
7140:
7121:(2): 299–302.
7101:
7066:
7036:
6993:
6950:
6915:
6896:
6859:
6836:
6814:
6787:
6773:
6741:
6690:
6653:
6588:
6548:
6503:
6457:
6411:
6379:J. Zool. Lond.
6358:
6324:
6289:(9): 1425–35.
6273:
6238:
6180:
6160:
6134:
6127:
6103:
6094:
6065:(4): 241–243.
6045:
6038:
6022:"Reproduction"
6005:
5960:
5923:
5874:
5863:(2): 148–160.
5843:
5832:
5805:(3): 211–214.
5785:
5778:
5760:
5748:
5697:
5648:
5636:
5602:
5550:
5543:
5521:
5519:p.e2122789119.
5511:
5500:
5493:
5467:
5420:(622): 47–84.
5404:
5392:
5385:
5365:
5353:
5334:
5290:
5276:
5250:
5202:
5187:
5172:
5142:
5101:
5068:
5056:
5041:
5016:
4933:
4918:
4901:
4887:
4830:
4773:
4766:
4748:
4741:
4715:
4673:(1): 225–241.
4650:
4643:
4619:
4557:
4550:
4524:
4490:
4489:
4487:
4484:
4483:
4482:
4477:
4472:
4467:
4462:
4457:
4451:
4450:
4434:
4431:
4303:Main article:
4300:
4297:
4254:
4251:
4232:Main article:
4229:
4226:
4202:Main article:
4199:
4196:
4145:Main article:
4142:
4139:
4133:
4130:
4128:
4125:
4097:orb-web spider
4080:Main article:
4077:
4074:
4021:Main article:
4018:
4015:
3952:Southeast Asia
3924:Main article:
3921:
3918:
3915:
3914:
3911:
3908:
3905:
3902:
3895:
3894:
3891:
3888:
3885:
3882:
3879:
3876:
3866:
3865:
3862:
3859:
3856:
3853:
3850:
3847:
3844:
3838:
3837:
3834:
3831:
3825:
3822:
3819:
3816:
3813:
3809:
3808:
3805:
3798:
3789:arachnologists
3722:Main article:
3719:
3716:
3712:
3711:
3708:
3707:
3703:
3702:
3699:
3698:
3694:
3693:
3690:
3689:
3685:
3684:
3681:
3680:
3676:
3675:
3672:
3671:
3667:
3666:
3663:
3662:
3658:
3657:
3654:
3653:
3639:
3636:
3635:
3632:
3631:
3624:
3619:
3617:
3611:
3610:
3607:
3606:
3602:
3601:
3598:
3597:
3594:Austrochilidae
3590:
3587:
3586:
3583:
3582:
3575:
3570:
3568:
3560:
3558:
3555:
3554:
3551:
3550:
3543:
3538:
3536:
3530:
3529:
3526:
3525:
3521:
3520:
3517:
3516:
3508:Avicularioidea
3504:
3501:
3500:
3497:
3496:
3482:
3477:
3475:
3467:
3465:
3460:
3458:
3452:
3451:
3448:
3447:
3440:
3432:
3430:
3422:
3413:
3410:
3406:
3405:
3402:
3401:
3397:
3396:
3393:
3392:
3388:
3387:
3384:
3383:
3379:
3378:
3375:
3374:
3370:
3369:
3366:
3365:
3361:
3360:
3357:
3356:
3352:
3351:
3348:
3347:
3338:whip scorpions
3333:
3330:
3329:
3326:
3325:
3308:
3303:
3301:
3299:Tetrapulmonata
3295:
3294:
3291:
3290:
3281:
3273:
3271:
3267:
3266:
3263:
3262:
3244:
3238:
3236:
3230:
3229:
3226:
3225:
3209:
3204:
3202:
3199:
3198:
3195:
3194:
3181:
3176:
3174:
3168:
3167:
3164:
3163:
3156:(sea spiders)
3150:
3145:
3143:
3135:
3102:
3099:
2990:Inner Mongolia
2857:
2854:
2849:Main article:
2846:
2843:
2837:
2834:
2824:
2821:
2803:
2800:
2735:
2732:
2693:Main article:
2690:
2687:
2595:nigroannulatum
2560:social insects
2552:Main article:
2549:
2546:
2520:baboon spiders
2475:Atrax robustus
2462:
2459:
2458:
2457:
2451:
2444:
2442:
2434:
2427:
2425:
2413:
2406:
2404:
2391:
2384:
2382:
2379:
2372:
2257:The primitive
2228:Mature female
2172:Main article:
2169:
2168:Capturing prey
2166:
2104:feed on plant
2067:Beltian bodies
2058:jumping spider
2046:jumping spider
2037:
2034:
2032:
2029:
2018:Misumena vatia
1881:
1878:
1848:
1845:
1785:
1784:
1778:
1771:
1769:
1761:
1754:
1752:
1749:
1742:
1740:
1731:
1724:
1717:redback spider
1671:
1670:
1663:
1656:
1655:
1649:
1642:
1641:
1640:
1636:
1635:
1634:
1633:
1604:spermatophores
1553:
1550:
1460:Main article:
1457:
1454:
1389:
1386:
1379:proprioceptors
1368:slit sensillae
1325:
1322:
1223:jumping spider
1211:Main article:
1208:
1205:
1203:
1200:
1157:
1154:
1018:
1015:
908:
905:
781:
780:
777:
774:
771:
768:
765:
762:
759:
755:
750:
743:
742:
738:
737:
734:
731:
728:
725:
722:
719:
716:
713:
709:
706:
705:
703:
702:
699:
696:
693:
689:
686:
683:
680:
677:
673:
668:
661:
660:
659:
658:
657:
655:
652:
649:Spider anatomy
647:Main article:
644:
641:
602:Proto-Germanic
593:
590:
443:. Spider-like
304:
303:
297:
296:
290:
289:
278:
277:
276:
275:
270:
260:
251:
250:
246:
245:
233:
229:
228:
226:Tetrapulmonata
223:
216:
215:
210:
206:
205:
200:
196:
195:
190:
186:
185:
180:
176:
175:
170:
166:
165:
152:
151:
135:
134:
126:
125:
117:
112:
107:
102:
97:
92:
87:
82:
77:
72:
67:
62:
44:
26:
9:
6:
4:
3:
2:
12235:
12224:
12221:
12219:
12216:
12214:
12211:
12209:
12206:
12205:
12203:
12186:
12181:
12177:
12173:
12168:
12164:
12160:
12155:
12151:
12147:
12142:
12138:
12134:
12128:
12124:
12119:
12115:
12111:
12106:
12102:
12098:
12093:
12089:
12085:
12080:
12076:
12072:
12067:
12063:
12059:
12054:
12050:
12046:
12041:
12037:
12033:
12028:
12024:
12020:
12015:
12011:
12007:
12002:
11998:
11994:
11989:
11985:
11981:
11976:
11972:
11968:
11963:
11959:
11955:
11950:
11946:
11942:
11937:
11933:
11929:
11924:
11920:
11915:
11909:
11905:
11900:
11894:
11890:
11889:
11887:
11885:
11881:
11877:
11872:
11856:
11853:
11851:
11848:
11846:
11843:
11842:
11839:
11832:
11828:
11810:
11808:
11807:Shultz (2007)
11804:
11803:
11800:
11786:
11783:
11781:
11775:
11773:
11770:
11768:
11765:
11764:
11762:
11760:
11756:
11750:
11747:
11745:
11742:
11741:
11739:
11737:
11733:
11730:
11728:
11714:
11708:
11702:
11701:
11698:
11695:
11689:
11675:
11673:(vinegaroons)
11669:
11667:
11661:
11660:
11658:
11656:
11653:Thelyphonida
11651:
11645:
11639:
11638:
11635:
11632:
11630:
11626:
11620:
11614:
11613:
11610:
11607:
11605:
11601:
11591:
11585:
11583:
11577:
11576:
11574:
11570:
11564:
11558:
11556:
11550:
11549:
11547:
11543:
11540:
11538:
11532:
11526:
11520:
11519:
11517:
11515:
11511:
11507:
11500:
11496:
11490:
11486:
11484:
11480:
11478:
11474:
11473:
11470:
11465:
11458:
11453:
11451:
11446:
11444:
11439:
11438:
11435:
11423:
11415:
11414:
11411:
11405:
11402:
11400:
11397:
11396:
11394:
11390:
11384:
11381:
11379:
11376:
11374:
11371:
11369:
11368:Arachnophobia
11366:
11365:
11363:
11359:
11353:
11350:
11348:
11345:
11343:
11340:
11338:
11335:
11333:
11330:
11328:
11325:
11323:
11320:
11318:
11315:
11313:
11310:
11308:
11305:
11303:
11300:
11298:
11295:
11293:
11292:Cephalothorax
11290:
11288:
11285:
11283:
11280:
11278:
11277:Arthropod leg
11275:
11274:
11272:
11270:
11266:
11260:
11257:
11255:
11252:
11250:
11249:Opisthothelae
11247:
11245:
11244:Mygalomorphae
11242:
11240:
11237:
11235:
11234:Araneomorphae
11232:
11231:
11229:
11227:
11223:
11218:
11208:
11205:
11203:
11200:
11198:
11195:
11193:
11190:
11188:
11185:
11183:
11180:
11179:
11177:
11175:
11171:
11167:
11160:
11155:
11153:
11148:
11146:
11141:
11140:
11137:
11124:
11121:
11119:
11116:
11115:
11112:
11090:
11087:
11085:
11084:
11082:(ant spiders)
11077:
11075:
11072:
11070:
11067:
11065:
11059:
11057:
11054:
11052:
11049:
11047:
11044:
11042:
11039:
11037:
11034:
11032:
11029:
11027:
11026:
11019:
11017:
11015:(ray spiders)
11011:
11009:
11008:
11001:
10999:
10993:
10991:
10988:
10986:
10983:
10981:
10975:
10973:
10970:
10968:
10967:
10960:
10958:
10952:
10950:
10947:
10945:
10944:
10937:
10935:
10932:
10928:
10923:
10917:
10915:
10912:
10910:
10907:
10905:
10904:Physoglenidae
10902:
10900:
10897:
10895:
10889:
10887:
10884:
10882:
10876:
10874:
10868:
10866:
10860:
10858:
10852:
10850:
10844:
10842:
10839:
10837:
10831:
10829:
10826:
10824:
10818:
10816:
10815:
10808:
10806:
10800:
10798:
10797:
10790:
10788:
10782:
10780:
10777:
10775:
10769:
10767:
10761:
10759:
10758:
10751:
10749:
10746:
10744:
10738:
10736:
10733:
10731:
10725:
10723:
10717:
10715:
10714:Cycloctenidae
10712:
10710:
10707:
10705:
10704:Cyatholipidae
10702:
10700:
10694:
10692:
10686:
10684:
10682:(sac spiders)
10678:
10676:
10675:Cithaeronidae
10673:
10671:
10668:
10666:
10663:
10661:
10660:
10653:
10651:
10647:Anyphaenidae
10645:
10643:
10640:
10638:
10632:
10630:
10626:Amaurobiidae
10624:
10622:
10621:
10614:
10613:
10610:
10607:
10605:
10601:
10591:
10588:
10581:
10579:
10573:
10571:
10565:
10563:
10562:Stenochilidae
10560:
10558:
10552:
10550:
10546:Segestriidae
10544:
10542:
10536:
10534:
10533:Plectreuridae
10531:
10529:
10528:
10521:
10519:
10516:
10514:
10510:Palpimanidae
10508:
10506:
10503:
10501:
10498:
10496:
10495:
10488:
10486:
10480:
10478:
10475:
10473:
10470:
10468:
10464:Hypochilidae
10462:
10460:
10457:
10455:
10449:
10447:
10443:Filistatidae
10441:
10439:
10433:
10431:
10425:
10423:
10417:
10415:
10412:
10410:
10407:
10405:
10399:
10398:
10395:
10392:
10388:
10385:
10383:
10382:Araneomorphae
10379:
10369:
10363:
10361:
10358:
10356:
10350:
10348:
10342:
10340:
10334:
10332:
10329:
10327:
10321:
10319:
10318:Macrothelidae
10316:
10314:
10311:
10309:
10305:Hexathelidae
10303:
10301:
10298:
10296:
10293:
10291:
10285:
10283:
10277:
10275:
10269:
10267:
10263:Barychelidae
10261:
10259:
10253:
10251:
10245:
10243:
10237:
10235:
10229:
10228:
10225:
10222:
10220:
10219:Mygalomorphae
10216:
10212:
10208:
10207:Opisthothelae
10202:
10198:
10188:
10182:
10180:
10176:Liphistiidae
10174:
10173:
10170:
10166:
10160:
10156:
10150:
10146:
10144:
10140:
10138:
10134:
10132:
10128:
10127:
10124:
10119:
10111:
10106:
10104:
10099:
10097:
10092:
10091:
10088:
10082:
10079:
10077:
10073:
10068:
10065:
10062:
10059:
10055:
10051:
10048:
10046:
10043:
10040:
10036:
10032:
10031:
10026:
10024:
10020:
10015:
10011:
10010:
10005:
10000:
9999:
9988:
9984:
9980:
9974:
9970:
9965:
9961:
9957:
9953:
9947:
9943:
9938:
9934:
9930:
9925:
9924:
9918:
9917:Kaston, B. J.
9914:
9910:
9906:
9902:
9896:
9892:
9891:
9885:
9881:
9877:
9873:
9868:
9864:
9860:
9856:
9850:
9846:
9842:
9838:
9834:
9833:
9828:
9823:
9822:
9810:
9804:
9800:
9799:
9793:
9789:
9783:
9779:
9774:
9773:
9760:
9756:
9755:
9747:
9740:
9734:
9726:
9720:
9716:
9715:
9707:
9699:
9693:
9688:
9687:
9678:
9670:
9664:
9660:
9659:
9651:
9643:
9639:
9635:
9631:
9628:(2): 102–42.
9627:
9623:
9619:
9612:
9610:
9608:
9599:
9595:
9591:
9587:
9582:
9580:
9579:Metamorphoses
9570:
9568:
9566:
9564:
9555:
9549:
9545:
9544:
9536:
9528:
9522:
9518:
9517:
9509:
9507:
9505:
9503:
9501:
9499:
9490:
9484:
9480:
9473:
9465:
9459:
9455:
9454:
9446:
9438:
9432:
9428:
9427:
9419:
9411:
9405:
9401:
9394:
9386:
9382:
9378:
9374:
9370:
9366:
9362:
9358:
9354:
9347:
9339:
9335:
9331:
9327:
9320:
9312:
9306:
9302:
9297:
9296:
9287:
9280:
9267:
9263:
9257:
9250:
9238:
9234:
9230:
9223:
9216:
9204:
9200:
9194:
9186:
9182:
9178:
9174:
9171:(5): 431–38.
9170:
9166:
9159:
9140:
9136:
9132:
9128:
9124:
9119:
9114:
9111:(9): 374–79.
9110:
9106:
9099:
9092:
9084:
9080:
9076:
9072:
9068:
9064:
9060:
9056:
9049:
9041:
9037:
9032:
9027:
9023:
9019:
9015:
9011:
9007:
9000:
8992:
8988:
8983:
8978:
8974:
8970:
8966:
8962:
8958:
8951:
8943:
8937:
8933:
8932:
8924:
8916:
8912:
8907:
8902:
8899:(2): 239–50.
8898:
8894:
8890:
8883:
8867:
8861:
8845:
8839:
8823:
8819:
8812:
8804:
8800:
8795:
8790:
8787:(6): 948–51.
8786:
8782:
8778:
8771:
8755:
8749:
8741:
8737:
8733:
8729:
8725:
8721:
8720:
8712:
8710:
8701:
8697:
8693:
8689:
8685:
8681:
8676:
8671:
8666:
8661:
8657:
8653:
8649:
8645:
8641:
8633:
8625:
8619:
8615:
8610:
8609:
8600:
8584:
8580:
8574:
8566:
8560:
8556:
8549:
8541:
8535:
8531:
8524:
8517:
8502:
8498:
8491:
8484:
8466:
8462:Full text at
8459:
8455:
8451:
8447:
8440:
8432:
8426:
8422:
8418:
8414:
8407:
8399:
8395:
8391:
8384:
8376:
8370:
8366:
8362:
8355:
8353:
8344:
8340:
8333:
8325:
8321:
8314:
8306:
8302:
8298:
8294:
8290:
8286:
8282:
8278:
8277:
8269:
8250:
8246:
8240:
8236:
8232:
8225:
8218:
8216:
8214:
8212:
8210:
8208:
8199:
8193:
8189:
8185:
8181:
8177:
8171:
8163:
8159:
8155:
8151:
8147:
8143:
8139:
8135:
8127:
8119:
8115:
8110:
8105:
8100:
8095:
8091:
8087:
8083:
8076:
8067:
8062:
8059:(2): 221–65.
8058:
8054:
8050:
8043:
8041:
8039:
8037:
8028:
8024:
8019:
8014:
8009:
8004:
8000:
7996:
7992:
7985:
7977:
7973:
7969:
7965:
7961:
7957:
7953:
7949:
7942:
7934:
7930:
7925:
7920:
7916:
7912:
7908:
7904:
7900:
7896:
7892:
7888:
7884:
7877:
7869:
7865:
7860:
7855:
7851:
7847:
7844:(5): 775–78.
7843:
7839:
7835:
7828:
7820:
7816:
7812:
7808:
7801:
7794:
7792:
7790:
7788:
7786:
7777:
7773:
7768:
7763:
7759:
7755:
7751:
7747:
7746:Palaeontology
7743:
7741:
7733:
7732:
7727:
7726:Carboniferous
7723:
7722:
7715:
7711:
7710:Palaeontology
7704:
7697:
7681:
7674:
7666:
7662:
7658:
7654:
7650:
7646:
7643:(6): 231–37.
7642:
7638:
7637:Geology Today
7631:
7629:
7620:
7616:
7612:
7608:
7605:(2): 267–72.
7604:
7600:
7596:
7589:
7587:
7580:
7577:
7573:
7570:
7564:
7559:
7554:
7549:
7545:
7541:
7538:(1): 105–16.
7537:
7533:
7532:
7527:
7520:
7512:
7508:
7504:
7498:
7490:
7486:
7481:
7476:
7472:
7468:
7464:
7460:
7456:
7452:
7448:
7441:
7433:
7429:
7422:
7420:
7411:
7396:
7389:
7381:
7377:
7373:
7369:
7365:
7361:
7358:(2): 477–80.
7357:
7353:
7349:
7342:
7334:
7330:
7326:
7322:
7319:(2): 309–18.
7318:
7314:
7307:
7299:
7295:
7292:(2): 217–23.
7291:
7287:
7280:
7271:
7266:
7262:
7258:
7254:
7247:
7245:
7236:
7232:
7228:
7224:
7217:
7210:
7208:
7206:
7204:
7202:
7193:
7189:
7184:
7179:
7175:
7171:
7167:
7163:
7160:(5): 210279.
7159:
7155:
7151:
7144:
7136:
7132:
7128:
7124:
7120:
7116:
7112:
7105:
7097:
7093:
7089:
7085:
7081:
7077:
7070:
7062:
7058:
7051:
7049:
7040:
7032:
7028:
7024:
7020:
7016:
7012:
7009:(6): 743–53.
7008:
7004:
6997:
6989:
6985:
6981:
6977:
6973:
6969:
6966:(4): 575–92.
6965:
6961:
6954:
6946:
6942:
6938:
6934:
6931:(4): 283–87.
6930:
6926:
6919:
6911:
6910:New Scientist
6907:
6900:
6892:
6888:
6883:
6878:
6875:(2): 155–71.
6874:
6870:
6863:
6855:
6851:
6847:
6840:
6824:
6818:
6810:
6806:
6802:
6798:
6791:
6776:
6770:
6766:
6762:
6755:
6748:
6746:
6737:
6733:
6729:
6725:
6721:
6717:
6713:
6709:
6706:(4): 572–76.
6705:
6701:
6694:
6685:
6680:
6676:
6672:
6668:
6666:
6657:
6642:
6638:
6634:
6630:
6626:
6622:
6618:
6614:
6610:
6606:
6599:
6592:
6576:
6572:
6571:
6563:
6561:
6552:
6533:
6529:
6525:
6518:
6516:
6507:
6492:
6488:
6484:
6480:
6476:
6472:
6468:
6461:
6446:
6442:
6438:
6434:
6430:
6426:
6422:
6415:
6396:
6392:
6388:
6384:
6381:
6380:
6372:
6365:
6363:
6346:
6345:
6341:
6333:
6331:
6329:
6320:
6316:
6312:
6308:
6304:
6300:
6296:
6292:
6288:
6284:
6277:
6269:
6265:
6261:
6257:
6254:(2): 171–77.
6253:
6249:
6242:
6234:
6230:
6226:
6222:
6218:
6214:
6210:
6206:
6199:
6197:
6195:
6193:
6191:
6189:
6187:
6185:
6178:
6174:
6170:
6164:
6148:
6144:
6138:
6130:
6124:
6120:
6116:
6115:
6107:
6098:
6090:
6086:
6081:
6076:
6072:
6068:
6064:
6060:
6056:
6049:
6041:
6035:
6031:
6027:
6023:
6016:
6014:
6012:
6010:
6000:
5995:
5990:
5985:
5982:(4): 531–38.
5981:
5977:
5976:
5971:
5964:
5956:
5952:
5949:(4): 449–59.
5948:
5944:
5943:
5938:
5936:
5935:Tidarren argo
5927:
5919:
5915:
5910:
5905:
5901:
5897:
5893:
5889:
5885:
5878:
5870:
5866:
5862:
5858:
5854:
5847:
5841:
5836:
5828:
5824:
5820:
5816:
5812:
5808:
5804:
5800:
5796:
5789:
5781:
5775:
5771:
5764:
5757:
5752:
5744:
5740:
5736:
5732:
5728:
5724:
5720:
5716:
5712:
5708:
5701:
5693:
5689:
5684:
5679:
5675:
5671:
5667:
5663:
5659:
5652:
5645:
5640:
5632:
5628:
5625:(2): 423–33.
5624:
5620:
5613:
5606:
5598:
5594:
5590:
5586:
5581:
5576:
5573:(4): 578–83.
5572:
5568:
5564:
5557:
5555:
5546:
5540:
5536:
5532:
5525:
5515:
5509:
5504:
5496:
5490:
5486:
5481:
5480:
5471:
5463:
5459:
5454:
5449:
5445:
5441:
5436:
5431:
5427:
5423:
5419:
5415:
5408:
5401:
5396:
5388:
5386:9783662048993
5382:
5378:
5377:
5369:
5362:
5357:
5350:
5345:
5343:
5341:
5339:
5323:
5319:
5314:
5309:
5305:
5301:
5294:
5279:
5273:
5269:
5265:
5261:
5254:
5235:
5231:
5227:
5220:
5213:
5211:
5209:
5207:
5199:
5194:
5192:
5184:
5179:
5177:
5160:
5156:
5152:
5146:
5137:
5132:
5128:
5124:
5120:
5116:
5112:
5105:
5097:
5093:
5089:
5086:
5085:
5077:
5075:
5073:
5065:
5060:
5053:
5048:
5046:
5038:
5033:
5031:
5029:
5027:
5025:
5023:
5021:
5013:
5008:
5006:
5004:
5002:
5000:
4998:
4996:
4994:
4992:
4990:
4988:
4986:
4984:
4982:
4980:
4978:
4976:
4974:
4972:
4970:
4968:
4966:
4964:
4962:
4960:
4958:
4956:
4954:
4952:
4950:
4948:
4946:
4944:
4942:
4940:
4938:
4930:
4925:
4923:
4915:
4910:
4908:
4906:
4897:
4891:
4883:
4879:
4874:
4869:
4865:
4861:
4857:
4853:
4849:
4845:
4841:
4834:
4826:
4822:
4818:
4814:
4809:
4804:
4800:
4796:
4792:
4788:
4784:
4777:
4769:
4763:
4759:
4752:
4744:
4738:
4734:
4729:
4728:
4719:
4704:
4700:
4696:
4692:
4688:
4684:
4680:
4676:
4672:
4668:
4664:
4657:
4655:
4646:
4640:
4636:
4632:
4631:
4623:
4615:
4611:
4606:
4601:
4596:
4591:
4587:
4583:
4579:
4575:
4571:
4564:
4562:
4553:
4547:
4543:
4539:
4535:
4528:
4512:
4508:
4502:
4500:
4498:
4496:
4491:
4481:
4478:
4476:
4473:
4471:
4468:
4466:
4463:
4461:
4458:
4456:
4453:
4452:
4448:
4442:
4437:
4430:
4428:
4424:
4420:
4419:Dreamcatchers
4415:
4410:
4408:
4404:
4400:
4396:
4392:
4387:
4385:
4381:
4377:
4376:
4375:Metamorphoses
4371:
4366:
4364:
4360:
4356:
4352:
4348:
4344:
4340:
4336:
4332:
4328:
4320:
4316:
4311:
4306:
4296:
4294:
4290:
4287:, and by the
4286:
4282:
4274:
4270:
4265:
4260:
4250:
4248:
4244:
4240:
4239:Arachnophobia
4235:
4234:Arachnophobia
4228:Arachnophobia
4225:
4223:
4219:
4215:
4211:
4205:
4195:
4193:
4192:
4187:
4182:
4177:
4175:
4174:
4169:
4165:
4164:
4159:
4155:
4148:
4138:
4124:
4122:
4121:
4116:
4112:
4109:
4105:
4098:
4094:
4093:
4088:
4083:
4082:Araneomorphae
4076:Araneomorphae
4073:
4071:
4067:
4063:
4059:
4055:
4052:, but rather
4051:
4047:
4043:
4036:
4035:
4029:
4024:
4023:Mygalomorphae
4017:Mygalomorphae
4014:
4012:
4011:Carboniferous
4008:
4004:
4000:
3999:Carboniferous
3996:
3991:
3989:
3985:
3984:
3979:
3975:
3971:
3970:
3965:
3961:
3957:
3953:
3949:
3942:
3938:
3937:
3932:
3927:
3912:
3909:
3906:
3903:
3901:
3900:Araneomorphae
3897:
3896:
3883:
3880:
3877:
3875:
3874:Mygalomorphae
3871:
3870:Opisthothelae
3868:
3867:
3860:
3857:
3854:
3851:
3848:
3845:
3843:
3840:
3839:
3835:
3832:
3829:
3826:
3823:
3820:
3817:
3814:
3811:
3810:
3796:
3795:
3792:
3790:
3786:
3782:
3778:
3774:
3773:Araneomorphae
3770:
3769:Mygalomorphae
3766:
3765:Opisthothelae
3762:
3757:
3755:
3746:
3738:
3737:Ancient Greek
3734:
3731:
3725:
3710:
3709:
3701:
3700:
3692:
3691:
3683:
3682:
3674:
3673:
3665:
3664:
3656:
3655:
3652:
3649:
3644:
3638:
3637:
3634:
3633:
3630:
3629:
3623:
3622:
3616:
3613:
3612:
3609:
3608:
3600:
3599:
3596:
3595:
3589:
3588:
3585:
3584:
3581:
3580:
3579:Gradungulidae
3574:
3573:
3567:
3564:
3563:
3557:
3556:
3553:
3552:
3549:
3548:
3542:
3541:
3535:
3534:Araneomorphae
3532:
3531:
3528:
3527:
3519:
3518:
3515:
3513:
3509:
3503:
3502:
3499:
3498:
3495:
3492:
3487:
3481:
3480:
3474:
3473:Mygalomorphae
3471:
3470:
3464:
3463:
3457:
3456:Opisthothelae
3454:
3453:
3450:
3449:
3446:
3445:
3439:
3436:
3435:
3429:
3426:
3425:
3421:
3419:
3404:
3403:
3395:
3394:
3386:
3385:
3377:
3376:
3368:
3367:
3359:
3358:
3350:
3349:
3346:
3344:
3339:
3332:
3331:
3328:
3327:
3324:
3322:
3317:
3313:
3307:
3306:
3300:
3297:
3296:
3293:
3292:
3289:
3287:
3280:
3277:
3276:
3269:
3268:
3265:
3264:
3261:
3259:
3254:
3250:
3242:
3241:
3235:
3232:
3231:
3228:
3227:
3224:
3221:
3215:
3208:
3207:
3201:
3200:
3197:
3196:
3193:
3191:
3186:
3180:
3179:
3173:
3170:
3169:
3166:
3165:
3162:
3160:
3155:
3149:
3148:
3142:
3139:
3138:
3134:
3132:
3128:
3123:
3119:
3114:
3112:
3108:
3098:
3096:
3092:
3087:
3085:
3081:
3077:
3076:Araneomorphae
3073:
3069:
3063:
3058:
3054:
3053:Araneomorphae
3050:
3049:Mygalomorphae
3045:
3043:
3039:
3033:
3028:
3022:
3017:
3016:Carboniferous
3013:
3012:montceauensis
3011:
3006:
3002:
2998:
2997:Carboniferous
2993:
2991:
2987:
2981:
2977:, from about
2976:
2975:
2971:
2967:
2966:
2961:
2957:
2953:
2949:
2944:
2938:
2933:
2932:
2927:
2923:
2922:cephalothorax
2919:
2913:
2909:, from about
2908:
2906:
2902:
2901:trigonotarbid
2898:
2893:
2891:
2886:
2883:in the Early
2880:
2875:
2867:
2862:
2856:Fossil record
2852:
2842:
2833:
2830:
2820:
2817:
2810:A funnel web.
2808:
2799:
2797:
2793:
2788:
2786:
2784:
2779:
2773:
2769:
2767:
2761:
2757:
2754:
2746:
2745:
2740:
2731:
2729:
2725:
2720:
2716:
2708:
2707:
2701:
2696:
2686:
2684:
2680:
2679:
2678:Myrmica rubra
2674:
2673:
2668:
2667:
2662:
2661:
2655:
2653:
2652:
2647:
2646:
2641:
2637:
2633:
2632:
2629:
2624:
2620:
2619:
2614:
2610:
2609:
2604:
2600:
2597:belongs to a
2596:
2594:
2589:
2588:independently
2585:
2581:
2577:
2574:
2570:
2569:
2566:
2561:
2555:
2554:Social spider
2548:Socialization
2545:
2543:
2539:
2535:
2534:
2529:
2525:
2521:
2517:
2513:
2512:Theraphosidae
2508:
2506:
2502:
2501:
2496:
2492:
2488:
2484:
2476:
2472:
2467:
2455:
2448:
2443:
2439:
2438:
2431:
2426:
2423:
2419:
2418:
2410:
2405:
2401:
2397:
2396:
2388:
2383:
2376:
2371:
2370:
2369:
2367:
2366:
2361:
2360:
2355:
2351:
2347:
2346:
2341:
2340:
2335:
2331:
2330:cephalothorax
2327:
2326:Ant-mimicking
2323:
2320:
2313:
2309:
2308:
2304:
2299:
2297:
2293:
2289:
2285:
2281:
2277:
2272:
2268:
2264:
2260:
2255:
2253:
2249:
2245:
2241:
2237:
2236:
2232:of the genus
2231:
2230:bolas spiders
2226:
2224:
2220:
2219:
2213:
2211:
2205:
2201:
2197:
2196:
2190:
2187:
2186:
2180:
2175:
2165:
2163:
2158:
2156:
2152:
2151:fungal spores
2148:
2144:
2140:
2136:
2132:
2127:
2122:
2119:
2115:
2111:
2107:
2103:
2099:
2095:
2091:
2087:
2082:
2080:
2076:
2073:as part of a
2072:
2068:
2064:
2063:
2059:
2051:
2047:
2042:
2028:
2026:
2025:
2020:
2019:
2013:
2009:
2005:
2001:
2000:
1993:
1991:
1990:
1985:
1984:
1979:
1978:
1973:
1972:
1967:
1963:
1959:
1958:
1953:
1952:
1947:
1946:
1941:
1940:
1935:
1931:
1930:
1925:
1924:
1919:
1915:
1911:
1907:
1903:
1899:
1895:
1891:
1887:
1877:
1875:
1871:
1870:
1861:
1857:
1853:
1844:
1842:
1838:
1834:
1830:
1825:
1823:
1822:
1821:Toxeus magnus
1817:
1813:
1809:
1804:
1802:
1798:
1794:
1790:
1781:
1775:
1770:
1766:
1765:
1758:
1753:
1746:
1741:
1737:
1736:
1728:
1723:
1722:
1721:
1718:
1714:
1713:Tidarren argo
1710:
1706:
1705:
1701:
1698:Males of the
1696:
1694:
1690:
1689:
1684:
1683:
1666:
1660:
1652:
1646:
1632:
1630:
1625:
1620:
1617:
1613:
1609:
1605:
1601:
1597:
1593:
1586:
1585:
1565:
1561:
1556:
1549:
1547:
1543:
1539:
1534:
1532:
1528:
1523:
1521:
1517:
1513:
1509:
1505:
1500:
1495:
1493:
1489:
1485:
1481:
1478:, which emit
1477:
1468:
1463:
1453:
1451:
1447:
1442:
1440:
1435:
1434:cephalothorax
1431:
1427:
1423:
1419:
1415:
1411:
1407:
1403:
1394:
1385:
1382:
1380:
1376:
1371:
1369:
1365:
1361:
1360:
1355:
1350:
1349:
1344:
1340:
1336:
1331:
1321:
1319:
1318:Cave dwelling
1315:
1311:
1307:
1306:
1301:
1296:
1294:
1290:
1286:
1285:visual acuity
1281:
1278:
1273:
1271:
1267:
1263:
1259:
1258:compound eyes
1254:
1250:
1242:
1241:
1235:
1228:
1224:
1219:
1214:
1213:Spider vision
1199:
1197:
1196:
1190:
1187:
1183:
1179:
1175:
1171:
1167:
1163:
1153:
1151:
1147:
1143:
1139:
1138:Mygalomorphae
1135:
1132:
1128:
1124:
1120:
1116:
1112:
1108:
1104:
1100:
1095:
1093:
1088:
1086:
1081:
1077:
1073:
1069:
1065:
1064:Holarchaeidae
1061:
1057:
1053:
1049:
1045:
1037:
1036:
1031:
1023:
1014:
1012:
1008:
1004:
1000:
996:
992:
988:
984:
980:
976:
972:
968:
964:
960:
955:
953:
949:
946:
943:contains the
942:
938:
934:
929:
925:
921:
913:
904:
902:
898:
894:
891:
887:
882:
880:
876:
872:
868:
864:
859:
857:
853:
849:
845:
841:
837:
833:
829:
825:
821:
817:
816:cephalothorax
813:
809:
805:
801:
797:
793:
778:
775:
772:
769:
766:
764:book lung sac
763:
760:
757:
753:
747:
735:
732:
729:
726:
723:
720:
717:
714:
711:
708:Leg segments:
700:
697:
694:
691:
690:
687:
684:
681:
679:trichobothria
678:
675:
674:
671:
665:
650:
640:
628:
613:
608:
603:
600:derives from
599:
589:
587:
586:arachnophobia
583:
579:
575:
571:
567:
562:
560:
556:
552:
551:widow spiders
547:
542:
540:
536:
532:
531:
527:
523:
519:
515:
511:
507:
506:
500:
496:
491:
490:Araneomorphae
487:
486:Mygalomorphae
483:
477:
472:
471:Carboniferous
466:
461:
458:
454:
450:
446:
442:
437:
432:
430:
426:
422:
418:
414:
409:
407:
403:
399:
395:
391:
390:cephalothorax
387:
383:
379:
375:
373:
369:
365:
361:
357:
353:
349:
345:
341:
338:that extrude
337:
333:
329:
325:
321:
317:
314:
310:
302:
298:
295:
291:
288:
286:
285:
274:
273:Araneomorphae
271:
269:
268:Mygalomorphae
266:
265:
264:
263:Opisthothelae
261:
258:
255:
254:
252:
247:
242:
237:
234:
231:
230:
227:
224:
221:
218:
217:
214:
211:
208:
207:
204:
201:
198:
197:
194:
191:
188:
187:
184:
181:
178:
177:
174:
171:
168:
167:
162:
157:
153:
149:
148:Mygalomorphae
145:
144:Araneomorphae
141:
136:
132:
127:
120:
115:
110:
105:
100:
95:
90:
85:
80:
75:
70:
65:
59:
52:
48:
47:Pennsylvanian
40:
37:
33:
19:
11883:
11785:Mesostigmata
11654:
11615:
11572:Haplocnemata
11554:(harvestmen)
11545:Stomothecata
11165:
11122:
11078:
11074:Xenoctenidae
11069:Viridasiidae
11046:Trechaleidae
11031:Titanoecidae
11020:
11004:Theridiidae
11002:
10972:Stiphidiidae
10963:Sparassidae
10961:
10954:Senoculidae
10938:
10909:Phyxelididae
10886:Penestomidae
10862:Nicodamidae
10809:
10802:Liocranidae
10793:Linyphiidae
10791:
10771:Hersiliidae
10754:Gnaphosidae
10752:
10680:Clubionidae
10654:
10634:Ammoxenidae
10615:
10586:
10522:
10518:Periegopidae
10489:
10472:Leptonetidae
10295:Euctenizidae
10117:
10071:Spider hunts
10029:
10007:
9968:
9941:
9922:
9889:
9871:
9844:
9830:
9797:
9777:
9757:. New York:
9751:
9746:
9738:
9733:
9713:
9706:
9685:
9677:
9657:
9650:
9625:
9621:
9589:
9585:
9578:
9542:
9535:
9515:
9478:
9472:
9452:
9445:
9425:
9418:
9399:
9393:
9360:
9356:
9346:
9332:(1): 17–25.
9329:
9325:
9319:
9294:
9286:
9277:
9270:. Retrieved
9266:the original
9256:
9248:
9241:. Retrieved
9237:the original
9232:
9222:
9214:
9207:. Retrieved
9203:the original
9193:
9168:
9164:
9158:
9146:. Retrieved
9139:the original
9108:
9104:
9091:
9058:
9054:
9048:
9016:(4): 86–87.
9013:
9009:
8999:
8964:
8960:
8950:
8930:
8923:
8896:
8892:
8882:
8870:. Retrieved
8860:
8848:. Retrieved
8838:
8826:. Retrieved
8822:the original
8811:
8784:
8780:
8770:
8758:. Retrieved
8748:
8723:
8717:
8675:10400.3/6470
8647:
8643:
8632:
8607:
8599:
8587:. Retrieved
8583:the original
8573:
8554:
8548:
8529:
8523:
8515:
8508:. Retrieved
8501:the original
8496:
8483:
8471:. Retrieved
8449:
8445:
8439:
8416:
8406:
8397:
8393:
8383:
8364:
8342:
8332:
8323:
8313:
8280:
8274:
8268:
8256:. Retrieved
8249:the original
8230:
8179:
8170:
8137:
8133:
8126:
8109:10261/217081
8092:(1): 71–86.
8089:
8085:
8075:
8056:
8052:
7998:
7994:
7984:
7951:
7947:
7941:
7890:
7886:
7876:
7841:
7837:
7827:
7810:
7806:
7749:
7745:
7739:
7729:
7719:
7716:(3): 605–14.
7713:
7709:
7696:
7684:. Retrieved
7673:
7640:
7636:
7602:
7598:
7535:
7529:
7519:
7510:
7497:
7454:
7450:
7440:
7431:
7427:
7418:
7410:
7398:. Retrieved
7388:
7355:
7351:
7341:
7316:
7312:
7306:
7289:
7285:
7279:
7260:
7256:
7226:
7222:
7157:
7153:
7143:
7118:
7114:
7110:
7104:
7079:
7075:
7069:
7060:
7056:
7047:
7039:
7006:
7002:
6996:
6963:
6959:
6953:
6928:
6924:
6918:
6909:
6899:
6872:
6868:
6862:
6845:
6839:
6827:. Retrieved
6817:
6800:
6796:
6790:
6778:. Retrieved
6760:
6703:
6699:
6693:
6674:
6670:
6664:
6656:
6644:. Retrieved
6608:
6604:
6591:
6579:. Retrieved
6574:
6568:
6559:
6551:
6539:. Retrieved
6532:the original
6527:
6523:
6514:
6506:
6494:. Retrieved
6477:(1): 15–27.
6474:
6470:
6460:
6448:. Retrieved
6428:
6424:
6414:
6402:. Retrieved
6395:the original
6382:
6377:
6349:. Retrieved
6343:
6339:
6286:
6282:
6276:
6251:
6247:
6241:
6208:
6204:
6168:
6163:
6151:. Retrieved
6147:The Guardian
6146:
6137:
6113:
6106:
6097:
6062:
6058:
6048:
6025:
5979:
5973:
5963:
5946:
5940:
5934:
5926:
5891:
5887:
5877:
5860:
5856:
5846:
5835:
5802:
5798:
5788:
5769:
5763:
5751:
5710:
5706:
5700:
5668:(62): 1–15.
5665:
5661:
5651:
5639:
5622:
5618:
5605:
5570:
5566:
5534:
5524:
5514:
5503:
5478:
5470:
5417:
5407:
5402:, p. 27
5395:
5379:. Springer.
5375:
5368:
5356:
5325:. Retrieved
5303:
5293:
5281:. Retrieved
5267:
5253:
5241:. Retrieved
5234:the original
5229:
5225:
5163:. Retrieved
5159:the original
5154:
5145:
5118:
5114:
5104:
5087:
5082:
5059:
4890:
4847:
4843:
4833:
4790:
4786:
4776:
4757:
4751:
4726:
4718:
4706:. Retrieved
4670:
4666:
4629:
4622:
4577:
4573:
4533:
4527:
4515:. Retrieved
4510:
4411:
4388:
4373:
4367:
4324:
4278:
4259:Fried spider
4237:
4207:
4189:
4185:
4178:
4171:
4161:
4157:
4150:
4135:
4118:
4111:wolf spiders
4101:
4090:
4039:
4032:
3992:
3987:
3981:
3973:
3967:
3948:Liphistiidae
3945:
3941:Liphistiidae
3934:
3758:
3753:
3744:
3732:
3727:
3641:
3626:
3592:
3577:
3547:Hypochilidae
3545:
3506:
3484:
3444:Liphistiidae
3442:
3427:
3415:
3335:
3315:
3311:
3310:
3283:
3246:
3211:
3183:
3152:
3115:
3111:chelicerates
3104:
3088:
3068:Hexathelidae
3059:well before
3046:
3008:
3005:Liphistiidae
2994:
2972:
2963:
2955:
2929:
2905:Palaeotarbus
2903:
2894:
2890:lagerstätten
2870:
2839:
2826:
2813:
2789:
2781:
2774:
2770:
2766:oscillations
2762:
2758:
2750:
2742:
2724:Orbiculariae
2712:
2704:
2676:
2670:
2664:
2658:
2656:
2649:
2643:
2626:
2616:
2608:Philoponella
2606:
2602:
2591:
2579:
2563:
2557:
2538:cartwheeling
2531:
2509:
2498:
2480:
2474:
2435:
2415:
2393:
2363:
2357:
2345:Pseudomyrmex
2343:
2337:
2324:
2311:
2305:
2300:
2296:crab spiders
2284:Wolf spiders
2278:, which see
2259:Liphistiidae
2256:
2233:
2227:
2216:
2208:
2193:
2191:
2183:
2181:
2177:
2159:
2123:
2086:Anyphaenidae
2083:
2060:
2055:
2022:
2016:
1997:
1994:
1987:
1981:
1975:
1969:
1957:Theridiosoma
1955:
1949:
1943:
1937:
1927:
1921:
1883:
1867:
1865:
1859:
1826:
1819:
1805:
1786:
1762:
1733:
1712:
1702:
1697:
1686:
1680:
1677:
1664:
1650:
1621:
1612:palpal bulbs
1589:
1582:
1555:
1535:
1524:
1496:
1473:
1443:
1399:
1383:
1375:acceleration
1372:
1357:
1346:
1327:
1324:Other senses
1303:
1297:
1282:
1274:
1270:wolf spiders
1262:chelicerates
1245:
1238:
1202:Sense organs
1193:
1191:
1185:
1182:Liphistiidae
1159:
1096:
1089:
1074:, including
1068:Liphistiidae
1044:chelicerates
1041:
1033:
991:Hypochilidae
956:
917:
901:Liphistiidae
883:
861:Spiders and
860:
792:chelicerates
790:Spiders are
789:
751:
669:
597:
595:
563:
543:
528:
503:
502:The species
501:
433:
410:
378:Anatomically
376:
315:
308:
307:
281:
279:
235:
219:
36:
12053:iNaturalist
11908:Wikispecies
11772:Holothyrida
11736:Acariformes
11692:Acaromorpha
11663:Schizomida
11562:(scorpions)
11560:Scorpiones
11522:Palpigradi
11489:Chelicerata
11487:Subphylum:
11378:Spider bite
11332:Palpal bulb
11322:Opisthosoma
11287:Calamistrum
11192:Cannibalism
11174:Arachnology
11080:Zodariidae
11061:Uloboridae
11041:Trachelidae
11022:Thomisidae
10990:Synotaxidae
10985:Synaphridae
10949:Selenopidae
10940:Salticidae
10925:(including
10919:Pisauridae
10870:Oecobiidae
10854:Nesticidae
10846:Mysmenidae
10820:Malkaridae
10784:Lamponidae
10719:Deinopidae
10688:Corinnidae
10617:Agelenidae
10604:Entelegynae
10554:Sicariidae
10538:Scytodidae
10500:Orsolobidae
10459:Huttoniidae
10435:Dysderidae
10427:Drymusidae
10419:Diguetidae
10401:Archaeidae
10344:Nemesiidae
10287:Dipluridae
10271:Ctenizidae
10143:Chelicerata
10141:Subphylum:
10002:‹ The
9426:Fierce Food
8400:(1): 51–60.
8176:Gould, S.J.
8140:(1): 2–13.
7111:Latrodectus
6496:1 September
6450:1 September
6431:(1): 1–18.
5327:10 December
5121:(1): 4–10.
4708:10 December
4517:24 November
4455:Arachnidism
4403:West Africa
4247:instinctive
4210:spider silk
4204:Spider silk
4191:Latrodectus
4163:Latrodectus
4147:Spider bite
4005:rocks, and
3997:, found in
3643:Entelegynae
3615:Araneoclada
3336:Pedipalpi (
3270:pulmonates
3214:Eurypterida
3172:Prosomapoda
3154:Pycnogonida
3141:Chelicerata
3118:gnathobases
3095:chelicerate
2970:araneomorph
2965:Permarachne
2829:Linyphiidae
2816:Theridiidae
2651:Latrodectus
2576:Theridiidae
2417:Cyclocosmia
2352:", such as
2280:ultraviolet
2149:. Airborne
2110:amino acids
2094:Clubionidae
2008:Lepidoptera
1939:Tetragnatha
1906:carotenoids
1890:ommochromes
1827:Like other
1816:regurgitate
1812:wolf spider
1780:Wolf spider
1542:safety rope
1531:calamistrum
1462:Spider silk
1289:dragonflies
1277:rhabdomeres
1166:nerve cords
1099:nitrogenous
1066:, and some
995:desiccation
987:araneomorph
967:Mygalomorph
840:opisthosoma
559:mygalomorph
510:herbivorous
473:rocks from
436:Spider webs
394:opisthosoma
372:taxonomists
203:Chelicerata
199:Subphylum:
56:319–0
12202:Categories
11704:Ricinulei
11641:Amblypygi
11587:Solifugae
11552:Opiliones
11483:Arthropoda
11399:Spider web
11297:Chelicerae
11239:Mesothelae
11182:Ballooning
11089:Zoropsidae
10934:Psechridae
10878:Oxyopidae
10841:Miturgidae
10833:Mimetidae
10811:Lycosidae
10763:Hahniidae
10735:Dictynidae
10656:Araneidae
10567:Telemidae
10524:Pholcidae
10505:Pacullidae
10491:Oonopidae
10414:Caponiidae
10247:Atracidae
10165:Mesothelae
10137:Arthropoda
9148:19 October
8967:(4): 240.
8872:11 October
8850:11 October
8828:11 October
8760:11 October
8726:: 409–29.
8589:13 October
8510:13 October
8473:13 October
8283:: 565–92.
8258:12 October
7813:: 819–46.
7752:(1): 191.
7686:15 October
7678:Hecht, H.
7563:1808/17085
7229:: 341–72.
7003:Biotropica
6829:10 October
6803:: 351–77.
6581:11 October
6541:11 October
6351:10 October
6211:: 619–43.
6177:1582381569
5243:11 October
5226:Cimbebasia
5165:31 October
5090:: 565–92.
4850:(30): 30.
4414:tarantella
4343:intoxicate
4281:tarantulas
4269:tarantulas
4257:See also:
4186:Loxosceles
4173:Phoneutria
4158:Loxosceles
4058:tarantulas
4046:chelicerae
3988:Heptathela
3983:Heptathela
3976:run silk "
3974:Liphistius
3969:Liphistius
3926:Mesothelae
3920:Mesothelae
3842:Mesothelae
3833:Spinnerets
3830:in abdomen
3783:and 4,280
3761:Mesothelae
3735:borrowing
3628:Haplogynae
3438:Mesothelae
3279:Scorpiones
3253:harvestmen
3127:spinnerets
3084:Cretaceous
3040:of flying
3036:saw rapid
3010:Paleothele
3001:Mesothelae
2956:Attercopus
2952:appendages
2948:spinnerets
2885:Cretaceous
2695:Spider web
2683:pheromones
2636:Dictynidae
2623:Agelenidae
2613:Uloboridae
2584:Madagascar
2580:Anelosimus
2565:Anelosimus
2540:down sand
2516:tarantulas
2483:camouflage
2365:Oecophylla
2339:Synemosyna
2267:tarantulas
2263:Ctenizidae
2252:moth flies
2244:pheromones
2235:Mastophora
2204:Deinopidae
2174:Spider web
2102:Salticidae
2098:Thomisidae
2090:Corinnidae
2006:and adult
1934:guanocytes
1914:exocuticle
1880:Coloration
1874:tarantulas
1869:Patu digua
1837:tarantulas
1829:arthropods
1801:spinnerets
1797:chelicerae
1665:Entelegyne
1600:arthropods
1546:parachutes
1476:spinnerets
1422:metatarsus
1406:trochanter
1388:Locomotion
1364:mosquitoes
1354:pheromones
1314:Caponiidae
1302:(example,
1253:arthropods
1178:Mesothelae
1134:Mesothelae
1060:Uloboridae
1048:chelicerae
1011:ectotherms
1003:spinnerets
975:book lungs
971:Mesothelae
959:book lungs
948:hemocyanin
941:book lungs
920:coelomates
897:Mesothelae
881:transfer.
852:chelicerae
848:appendages
796:arthropods
727:metatarsus
715:trochanter
607:spin-þron-
570:elasticity
566:pesticides
557:and other
555:tarantulas
453:Uraraneida
421:Mesothelae
356:Antarctica
336:spinnerets
324:chelicerae
320:arthropods
257:Mesothelae
249:Suborders
193:Arthropoda
140:Mesothelae
11845:Haptopoda
11629:Pedipalpi
11618:(spiders)
11536:Dromopoda
11464:Arachnida
11347:Spinneret
11307:Cribellum
11282:Book lung
11197:Evolution
11036:Toxopidae
10740:Eresidae
10709:Cybaeidae
10696:Ctenidae
10313:Idiopidae
10255:Atypidae
10205:Suborder
10163:Suborder
10149:Arachnida
10129:Kingdom:
9960:123151744
9933:628203833
9863:256272177
9592:: 64–82.
9429:. Plume.
9385:229391382
9377:2352-4588
9113:CiteSeerX
9061:: 40–44.
8754:"Spiders"
8700:251727654
8684:0960-9822
8452:: 1–106.
8297:0066-4162
8118:1439-6092
7915:0028-1042
7665:130267137
7434:: 553–63.
7372:0003-3472
7063:: 392–96.
6877:CiteSeerX
6869:Behaviour
6854:2246/2705
6765:MIT Press
6700:Oecologia
6646:31 August
6491:0161-8202
6445:0161-8202
6404:11 August
6385:: 25–29.
6303:0022-0949
6260:0161-8202
5999:1807/1012
5827:0792-4259
5743:205015549
5589:0022-0949
5444:1313-2989
5322:240206876
4703:221235817
4687:0066-4170
4486:Citations
4407:Caribbean
4399:folktales
4391:trickster
4361:from her
4355:Ninhursag
4345:her with
4293:Venezuela
4181:jellyfish
4108:cursorial
3978:tripwires
3964:trapdoors
3807:Features
3791:in 2022.
3486:Atypoidea
3418:cladogram
3234:Arachnida
3185:Xiphosura
3131:pedipalps
3122:arthropod
2962:arachnid
2926:pedipalps
2845:Evolution
2790:In 1973,
2689:Web types
2593:Theridion
2350:livestock
2294:and some
2265:and many
2195:Hyptiotes
2162:cursorial
2155:chitinase
2135:marmalade
2012:Homoptera
1951:Argyrodes
1624:courtship
1616:pedipalps
1527:cribellum
1516:cellulose
1450:arachnids
1174:esophagus
1144:("little
1142:nephridia
1119:arthropod
1103:uric acid
1085:pedipalps
1076:scorpions
1072:arachnids
999:spiracles
867:arachnids
863:scorpions
856:pedipalps
736:chelicera
654:Body plan
612:literally
596:The word
592:Etymology
582:mythology
546:courtship
539:pedipalps
514:predators
445:arachnids
352:organisms
344:arachnids
294:Diversity
213:Arachnida
179:Kingdom:
173:Eukaryota
11962:BugGuide
11893:Wikidata
11777:Ixodida
11671:Uropygi
11616:Araneae
11481:Phylum:
11477:Animalia
11475:Kingdom
11422:Category
11337:Scopulae
11327:Pedipalp
11226:Taxonomy
11207:Glossary
11187:Behavior
11056:Udubidae
10914:Pimoidae
10727:Desidae
10665:Arkyidae
10642:Anapidae
10336:Migidae
10135:Phylum:
10131:Animalia
10120:families
10054:Archived
10035:Archived
10004:template
9987:25833874
9909:35231232
9843:(1976).
9642:29668658
9272:2 August
9243:2 August
9209:2 August
9185:17168889
9135:10942961
9083:26923161
9040:11483545
8991:11290673
8915:15306718
8803:12495200
8740:17877450
8692:35998593
8458:2246/991
8339:"ἀράχνη"
8320:"ărānĕa"
8178:(1990).
8162:16386884
8027:32218802
7976:26323977
7933:24317464
7868:21508021
7776:83825904
7740:Eotarbus
7731:Eotarbus
7721:Eotarbus
7619:42371883
7572:Archived
7569:Abstract
7489:33270151
7457:(1): 1.
7380:53157471
7333:53166407
7192:34017604
7135:36475912
7096:38307165
7031:54023263
6988:13871079
6945:44727810
6848:(2498).
6780:25 March
6728:28307636
6641:35215325
6633:17818935
6577:: 213–25
6319:26035671
6311:20400626
6225:15012400
6153:30 April
6089:31161756
5918:19403531
5735:11279484
5692:20050781
5597:22279064
5462:27843380
4882:28289774
4825:27885893
4817:19825348
4695:32822555
4614:19104044
4433:See also
4405:and the
4393:-spider
4285:Cambodia
4273:Cambodia
4214:attempts
4208:Because
4064:and the
4042:Triassic
3815:Families
3781:families
3754:arákhnēs
3718:Taxonomy
3080:Jurassic
3057:Triassic
2995:Several
2986:Daohuogo
2943:Devonian
2918:Silurian
2897:arachnid
2792:Skylab 3
2753:momentum
2634:(family
2631:gregalis
2621:(family
2571:(in the
2359:Amyciaea
2334:antennae
2212:spinosus
2210:Deinopis
2147:sausages
2143:egg yolk
2118:vitamins
2048:seen in
1945:Leucauge
1902:Melanins
1793:humidity
1711:species
1704:Tidarren
1592:internal
1512:collagen
1446:scopulae
1430:extensor
1330:cuticles
1300:six eyes
1225:'s main
1131:suborder
963:tracheal
950:to make
933:arteries
924:hemocoel
893:carapace
875:venomous
844:antennae
802:made of
676:pedipalp
518:lassoing
482:suborder
462:, about
457:Devonian
429:extensor
417:antennae
402:cephalon
382:segments
368:families
189:Phylum:
183:Animalia
169:Domain:
51:Holocene
12208:Spiders
11941:Araneae
11928:Araneae
11914:Araneae
11884:Araneae
11835:Extinct
11779:(ticks)
11317:Exuviae
11312:Epigyne
11269:Anatomy
11166:Spiders
10927:Halidae
10147:Class:
10118:Araneae
10116:Extant
10076:YouTube
10019:Spiders
10006:below (
9942:Spiders
9880:1979220
9761:, 1997.
9233:Phobias
9063:Bibcode
9055:Toxicon
9031:1071491
8982:1071344
8652:Bibcode
8305:2097274
8184:Bibcode
8142:Bibcode
8134:Zoology
8018:7078165
8001:: 182.
7956:Bibcode
7924:3889289
7895:Bibcode
7859:3169061
7754:Bibcode
7742:Dunlop"
7645:Bibcode
7540:Bibcode
7511:Gizmodo
7480:7716925
7459:Bibcode
7400:21 July
7183:8131949
7162:Bibcode
7011:Bibcode
6968:Bibcode
6708:Bibcode
6613:Bibcode
6605:Science
6268:3705625
6233:6963733
6080:6680124
6030:176–212
5909:2839943
5807:Bibcode
5715:Bibcode
5683:3011943
5453:5096409
5422:Bibcode
5418:ZooKeys
5123:Bibcode
4873:5348567
4852:Bibcode
4795:Bibcode
4605:2634869
4582:Bibcode
4380:Arachne
4372:in his
4339:produce
4267:Cooked
4253:As food
4156:(genus
4070:balloon
4003:Permian
3828:Ganglia
3821:Species
3777:species
3750:ἀράχνης
3745:arákhnē
3428:Araneae
3340:, etc)
3316:spiders
3312:Araneae
3255:, etc)
3042:insects
3029:period
3027:Permian
2960:Permian
2941:in the
2934:, from
2916:in the
2899:is the
2568:eximius
2522:, have
2461:Defense
2223:tipulid
2131:bananas
2071:acacias
2050:Chennai
2004:Diptera
1989:Maratus
1971:Argiope
1918:tanning
1910:pterins
1898:guanine
1886:pigment
1833:cuticle
1608:syringe
1520:elastic
1499:protein
1484:spigots
1414:patella
1348:Araneus
1293:insects
1170:ganglia
1150:ammonia
1146:kidneys
1123:insects
1111:cloacal
1080:enzymes
979:ventral
836:abdomen
820:prosoma
812:tagmata
800:cuticle
773:epigyne
721:patella
627:(s)pen-
618:spinner
574:mammals
535:enzymes
449:spigots
425:ganglia
413:insects
411:Unlike
398:pedicel
386:tagmata
366:in 134
364:species
360:habitat
316:Araneae
309:Spiders
236:Araneae
232:Order:
209:Class:
18:Araneae
12185:150467
12146:614523
12130:NZOR:
12006:1ARANO
11503:Extant
11466:orders
10023:Curlie
10009:Curlie
9985:
9975:
9958:
9948:
9931:
9907:
9897:
9878:
9861:
9851:
9805:
9784:
9721:
9694:
9665:
9640:
9550:
9523:
9485:
9460:
9433:
9406:
9383:
9375:
9307:
9183:
9133:
9115:
9081:
9038:
9028:
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8979:
8938:
8913:
8801:
8738:
8698:
8690:
8682:
8620:
8561:
8536:
8427:
8421:272–73
8371:
8303:
8295:
8241:
8194:
8160:
8116:
8025:
8015:
7974:
7948:Nature
7931:
7921:
7913:
7866:
7856:
7774:
7663:
7617:
7487:
7477:
7378:
7370:
7331:
7190:
7180:
7133:
7094:
7029:
6986:
6943:
6879:
6771:
6736:424262
6734:
6726:
6639:
6631:
6489:
6443:
6317:
6309:
6301:
6266:
6258:
6231:
6223:
6175:
6125:
6119:232–33
6087:
6077:
6036:
5916:
5906:
5825:
5776:
5741:
5733:
5707:Nature
5690:
5680:
5595:
5587:
5541:
5491:
5460:
5450:
5442:
5383:
5320:
5274:
4880:
4870:
4823:
4815:
4764:
4739:
4701:
4693:
4685:
4641:
4612:
4602:
4548:
4480:Toxins
4395:Anansi
4384:Athena
4363:vagina
4329:, the
4289:Piaroa
4113:, and
4106:, the
3958:, and
3910:46,770
3818:Genera
3797:
3785:genera
3741:ἀράχνη
3733:aranea
2907:jerami
2802:Cobweb
2628:Mallos
2625:) and
2573:family
2437:Portia
2354:aphids
2312:Portia
2307:Portia
2126:Pollen
2114:lipids
2106:nectar
1977:Lycosa
1894:bilins
1808:larval
1709:Yemeni
1508:chitin
1439:flexor
1426:tarsus
1420:; the
1268:, and
1249:ocelli
1227:ocelli
1195:Portia
1127:embryo
1056:glands
983:oxygen
952:oxygen
890:convex
832:thorax
824:insect
808:embryo
804:chitin
730:tarsus
598:spider
578:plants
530:Portia
460:period
406:thorax
388:, the
348:orders
334:, and
280:
243:, 1757
241:Clerck
42:Spider
12180:WoRMS
12167:Plazi
12159:57473
12097:82732
12071:12065
12066:IRMNG
12058:47118
12019:10626
11899:Q1357
11718:Acari
9638:S2CID
9622:Ramus
9381:S2CID
9303:–45.
9279:list.
9142:(PDF)
9101:(PDF)
8696:S2CID
8504:(PDF)
8493:(PDF)
8468:(PDF)
8301:JSTOR
8252:(PDF)
8227:(PDF)
7972:S2CID
7803:(PDF)
7772:S2CID
7706:(PDF)
7661:S2CID
7615:S2CID
7424:(PDF)
7376:S2CID
7329:S2CID
7219:(PDF)
7131:S2CID
7092:S2CID
7082:(1).
7053:(PDF)
7027:S2CID
6984:S2CID
6941:S2CID
6757:(PDF)
6732:S2CID
6637:S2CID
6601:(PDF)
6565:(PDF)
6535:(PDF)
6520:(PDF)
6398:(PDF)
6374:(PDF)
6315:S2CID
6264:JSTOR
6229:S2CID
5739:S2CID
5615:(PDF)
5487:–01.
5318:S2CID
5283:8 May
5237:(PDF)
5222:(PDF)
4821:S2CID
4699:S2CID
4423:Moche
4359:semen
4315:Moche
4313:This
4241:is a
4218:goats
4141:Bites
4095:, an
4054:setae
4050:hairs
3960:Japan
3956:China
3907:3,899
3884:3,327
3747:from
3730:Latin
3249:ticks
3107:clade
3018:over
2874:amber
2866:amber
2823:Other
2796:space
2778:moths
2728:wasps
2715:genus
2599:genus
2542:dunes
2528:setae
2487:birds
2420:, an
2315:'
2303:genus
2248:moths
2240:bolas
1966:setae
1700:genus
1596:sperm
1538:sperm
1504:nylon
1488:gland
1418:tibia
1410:femur
1343:smell
1339:taste
1335:setae
1221:This
1052:venom
928:blood
879:sperm
871:fangs
724:tibia
718:femur
625:*
605:*
526:genus
522:bolas
332:venom
328:fangs
326:with
313:order
220:Clade
12123:6893
12118:NCBI
12092:ITIS
12045:1496
12040:GBIF
12001:EPPO
11967:1954
11949:BOLD
11655:s.l.
11392:Webs
11342:Silk
11123:Bold
9983:OCLC
9973:ISBN
9956:OCLC
9946:ISBN
9929:OCLC
9905:OCLC
9895:ISBN
9876:OCLC
9859:OCLC
9849:ISBN
9803:ISBN
9782:ISBN
9719:ISBN
9692:ISBN
9663:ISBN
9548:ISBN
9521:ISBN
9483:ISBN
9458:ISBN
9431:ISBN
9404:ISBN
9373:ISSN
9305:ISBN
9274:2009
9245:2009
9211:2009
9181:PMID
9150:2008
9131:PMID
9079:PMID
9036:PMID
8987:PMID
8936:ISBN
8911:PMID
8874:2008
8852:2008
8830:2008
8799:PMID
8762:2008
8736:PMID
8688:PMID
8680:ISSN
8618:ISBN
8616:–5.
8591:2008
8559:ISBN
8534:ISBN
8512:2008
8475:2008
8425:ISBN
8369:ISBN
8293:ISSN
8260:2008
8239:ISBN
8192:ISBN
8158:PMID
8114:ISSN
8023:PMID
7929:PMID
7911:ISSN
7864:PMID
7688:2008
7485:PMID
7402:2008
7368:ISSN
7188:PMID
6831:2008
6782:2011
6769:ISBN
6724:PMID
6648:2024
6629:PMID
6583:2008
6543:2008
6498:2024
6487:ISSN
6452:2024
6441:ISSN
6406:2007
6353:2008
6307:PMID
6299:ISSN
6256:ISSN
6221:PMID
6173:ISBN
6155:2018
6123:ISBN
6085:PMID
6034:ISBN
5914:PMID
5823:ISSN
5774:ISBN
5731:PMID
5688:PMID
5593:PMID
5585:ISSN
5539:ISBN
5489:ISBN
5458:PMID
5440:ISSN
5381:ISBN
5329:2021
5285:2016
5272:ISBN
5245:2008
5167:2017
4878:PMID
4813:PMID
4762:ISBN
4737:ISBN
4710:2021
4691:PMID
4683:ISSN
4639:ISBN
4610:PMID
4574:PNAS
4546:ISBN
4519:2023
4427:Peru
4370:Ovid
4351:rape
4349:and
4347:beer
4335:Enki
4327:Uttu
4198:Silk
4188:and
4001:and
3771:and
3763:and
3416:The
3082:and
3051:and
2827:The
2669:and
2518:and
2489:and
2392:The
2276:bees
2269:are
2145:and
2139:milk
2100:and
1983:Josa
1980:and
1908:and
1896:and
1847:Size
1789:eggs
1562:and
1514:and
1480:silk
1402:coxa
1341:and
1283:The
1207:Eyes
1115:anus
1092:ceca
1062:and
969:and
961:, a
886:silk
830:and
828:head
733:claw
712:coxa
688:eyes
576:and
488:and
340:silk
282:See
146:and
64:PreꞒ
12105:NBN
12084:118
12079:ISC
11993:166
11988:EoL
11975:CoL
11954:251
11936:AFD
11923:ADW
10074:on
10021:at
9630:doi
9594:doi
9365:doi
9334:doi
9301:244
9173:doi
9123:doi
9071:doi
9059:114
9026:PMC
9018:doi
9014:175
8977:PMC
8969:doi
8965:174
8901:doi
8789:doi
8728:doi
8670:hdl
8660:doi
8454:hdl
8450:178
8285:doi
8150:doi
8138:109
8104:hdl
8094:doi
8061:doi
8057:150
8013:PMC
8003:doi
7964:doi
7952:379
7919:PMC
7903:doi
7891:100
7854:PMC
7846:doi
7815:doi
7762:doi
7653:doi
7607:doi
7579:PDF
7558:hdl
7548:doi
7475:PMC
7467:doi
7455:108
7360:doi
7321:doi
7294:doi
7265:doi
7261:141
7231:doi
7178:PMC
7170:doi
7123:doi
7084:doi
7019:doi
6976:doi
6933:doi
6887:doi
6873:138
6850:hdl
6805:doi
6716:doi
6704:112
6679:doi
6621:doi
6609:198
6479:doi
6433:doi
6387:doi
6383:255
6291:doi
6287:213
6213:doi
6075:PMC
6067:doi
5994:hdl
5984:doi
5951:doi
5947:254
5904:PMC
5896:doi
5892:276
5865:doi
5861:130
5815:doi
5723:doi
5711:410
5678:PMC
5670:doi
5627:doi
5575:doi
5571:215
5485:100
5448:PMC
5430:doi
5308:doi
5131:doi
5092:doi
4868:PMC
4860:doi
4848:104
4803:doi
4675:doi
4600:PMC
4590:doi
4578:105
4538:doi
4401:of
4176:).
3881:368
3858:Yes
3855:Yes
3852:169
3787:by
3062:200
3021:299
2980:165
2937:386
2912:420
2879:130
2734:Orb
2615:),
2452:An
2400:web
2246:of
2079:ant
1954:or
1186:all
838:or
818:or
495:200
465:386
350:of
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12182::
12169::
12156::
12143::
12120::
12107::
12094::
12081::
12068::
12055::
12042::
12029::
12016::
12003::
11990::
11980:RN
11977::
11964::
11951::
11938::
11925::
11910::
11895::
9981:.
9954:.
9903:.
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9829:.
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