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Structures built by animals

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other debris slow the flow of the water. The beavers then form a small platform of sticks stretching across the water source. More sticks and branches are added to build the dam up over time. The structure in the center of the dam, known as the lodge, serves as a home for the beavers and protects them from predators. The primary reason behind the construction of beaver dams is to surround the lodge with deep water, which protects the beaver from land-dwelling predators. The entrance of the dam is underwater to prevent predators such as bears and wolves from entering, and the sticks at the top of the lodge are not packed tightly, which allows air into the structure.
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to predators who might rip open the nest to eat the larva. One method the queens use to prevent this is covering the developing pupae in pulp, which acts as a reinforcer and makes it more difficult from predators to break open the pupae. This pulp is a mixture of plant matter and liquids from the mouth of the queen wasp. While there are costs associated with using pulp, such as requiring time and energy to collect materials and hindering the emergence of the worker wasps from the cocoon, it does lower the risk of predation. Nests in areas with higher predation rates have been found to contain more pulp on these cocoons than nests in low predation areas.
742: 489:) form "living bridges" to assist in transportation. Army ant colonies may move locations each day in search of food. These bridges provide a path over obstacles and allow for the ants to search for food at an increased speed. The bridges are constructed when the ants join their bodies together, and can vary in size and shape depending on the situation the ants face. Ants are confined to their position when they are forming these bridges, preventing them from moving. The bridges are broken apart when they are no longer needed. 448: 252: 363: 30: 264: 2267: 439:) larva. These larva prey on small arthropods, such as ants. The larva dig pits into fine-particle soil to capture their prey, which fall into the holes and are often unable to climb out. The antlions may alter these pits based on prey availability. In areas with less available prey, antlions will make wider holes to increase the chance of catching an insect. If prey are able to climb out of the hole, antlions will increase the depth of the hole. 753: 159: 473: 677: 435:
size of their webs when prey are scarce, and can add extra ornamental pieces to their web in order to attract more prey. Traps can allow organisms to capture larger prey, provide protection from predators, or serve as an area for mating, as seen with spiders. Another method of trap creation is used by the antlion (
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Structures formed from plant material include beaver dams, which are constructed by foraged branches and sticks. The dam is a wall of sticks constructed on a moving water source, which forces the water to collect in one area and to stop flowing. Beavers begin to build a dam in an area where rocks and
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The nests of these wasps contain “defensive structures”, which are formations built onto or inside of the nest to prevent predation. New nests are formed in the spring by young queens, as worker wasps have not hatched at this time. While these worker wasps are growing in the nest, they are vulnerable
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Chaboo, C.S., S. Adam, K. Nishida, L. Schletzbaum. 2023. Architecture, construction, retention, and repair of fecal shields in three tribes of tortoise beetles (Insecta: Coleoptera: Chrysomelidae Cassidinae: Cassidini, Mesomphaliini, Spilophorini). ZooKeys Special Issue, Research on Chrysomelidae 9.
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Cocoons are another type of structure formed to protect the organism from predation. In order to transform from a larva into a butterfly or moth, a caterpillar must undergo drastic changes in its body. These changes require significant amounts of energy and occur over long periods of time, making a
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caterpillar uses the hairs that cover its body as a defensive mechanism against predators. When it is time to form a cocoon, the caterpillar rips the hairs off of its body and places them around the pupating site. This creates a series of defensive walls to protect the vulnerable caterpillar while
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Flowering plants provide a variety of resources – twigs, leaves, petioles, roots, flowers and seeds. Basal plants, such as lichens, mosses and ferns also find use in structures built by animals. The leaves of grasses and palms being elongate and parallel-veined are very commonly used for building.
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Animals primarily build habitat for protection from extreme temperatures and from predation. Constructed structures raise physical problems which need to be resolved, such as humidity control or ventilation, which increases the complexity of the structure. Over time, through evolution, animals use
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Trap-building is a method used to catch prey instead of active hunting. Animals that snare prey will construct a trap and then wait nearby until an organism is caught. This is observed in web-building spiders, who weave elaborate webs of sticky spider silk that entangle prey. Spiders increase the
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to form fine sticky strands used for capturing prey. In Chrysomelidae (leaf beetles), larvae in a few subfamilies retain their feces as shield or body armor that may be thermoregulatory, offensive, or defensive In other cases, the scale wax, produced on the bodies of honey bees, is gathered and
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Predators are attracted to animal-built structures either by the prey or its offspring, or the stored caches of food. Structures built by animals may provide protection from predators through avoiding detection, by means such as camouflage and concealment, or through prevention of invasion, once
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The majority of self-secreted materials are produced by insects and selection acts on this characteristic of production of self-secreting materials and increases the fitness of the animal. In some cases, the self-secreted material is directly applied, as in the case of ecribellate silk, spun by
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spiders, to form sticky traps for prey, or it may be processed, as in the case of salivary excretion used for creation of paper by paper wasps, by blending it directly with wood pulp. Self-secreted materials may be processed in some cases. In cribellate spiders, silk produced by the spider are
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Materials used by animals in building structures need to not only be suitable for the kind of structure to be built but also to be manipulable by the animals. These materials may be organic in nature or mineral. They may also be categorised as "collected material" and "self-secreted material".
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also create display structures to attract mates. During the mating season, male Bowerbirds will collect twigs and colourful objects to create structures known as "bowers", which attract the attention of females. Bowers that are more colourful and well constructed are more attractive to female
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These, along with palm fibers and horse-hair fern are used to build hanging baskets. Wooden twigs form the greater proportion of materials used in the nests of large birds. Plants and trees not only provide resources but also sites. Branches provide support in the form of
401:. The primary sources of energy for an animal are the sun and its metabolism. The dynamics of heat in animal shelters is influenced by the construction material which may act as a barrier, as a heat sink or to dissipate heat. The cocoons of insect are a case in point. 422:
when the temperature falls resulting in heating. Cooling is aided by evaporation of excess water from the pupal cells. When the ambient temperature drops, the silk absorbs moisture and restores the moisture content by spreading water through all parts of its cocoon.
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Ground-nesting birds which rely on crypsis for concealment have nests made from local materials which blend in with the background, the eggs and young too are cryptic; whereas birds which do not use crypsis for hiding their nests may not have cryptic eggs or young.
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will build tall, thin pillars out of mud. These structures attract female crabs to male crab burrows and provide a hiding place for both males and females when predators are nearby. Beavers will often seal their dams and lodges with mud for extra support.
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caterpillar very vulnerable to predation. To overcome this, caterpillars will produce silk to form a cocoon or pupa, a structure in which the caterpillar will reside while pupating to lower its risk of predation. Some species of caterpillar, such as the
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larvae use stone pieces and also cut sections from green leaves for use in construction. The stone pieces are selected as per their size and shape from a large variety. In the case of leaf sections, these are cut and shaped to required size. Similarly
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blended with saliva, to form comb wax, the building material. Not all self-secreted materials are developed specifically for that purpose. For example, bird feathers are used for lining and insulation, a typical example being that of the female
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In ectothermic animals, moderation of temperature, along with architectural modifications to absorb, trap or dissipate energy, maximises the rate of development, as in the case of the communal silk nests of the small eggar moth
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bush. The structural stability of the nest is provided by a mesh of moss and spider silk. The tiny leaves of the moss act as hooks and the spider silk of egg cocoons provides the loops; thus forming a natural form of
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places bits of grass and other material loosely streaming both above and below the nest chamber to break the shape of the nest and to cause it to resemble random debris without any underlying structure.
339:, M. Hansell, where the lichen flakes are thought to resemble small patches of light seen through as in the case of insubstantial objects of insufficient importance to receive a predator's interest. 668:) of Asian tropical forests uses spider silk differently. It constructs a nest of plant strips which it suspends below a large leaf using spider silk for about a 150 or so of "pop-rivets". 634:"...the most amazing thing about it (the building behaviour) is, in my opinion that so few, so simple and so rigid movements together lead to the construction of so superb a result." 622:. The tit lines the outside with hundreds of flakes of pale lichens – this provides camouflage. Inside, it lines the nest with more than 2000 feathers to insulate the nest. 311:. Some species of birds will use materials foraged from nature to camouflage their nests and prevent their offspring from being hunted. Blue–gray gnatcatchers ( 794:) are able to spin multiple cocoons in the event that one gets destroyed. Other caterpillars will even form defensive structures to accompany their pupas. The 545:
Some materials in nature act as ready made "building blocks" to the animals in question, such as feathers and leaf petioles for some birds and animal
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birds have species which utilise spider silk. Most birds use spider silk as in the case of the long-tailed tit, previously discussed; however the
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Animal structures can serve as a means of communication with other organisms. Animals may construct to attract mates, as seen in species of male
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Internal architectural devices, such as walls may block convection or the construction of air flow systems may cool the nest or habitat.
1432:"Bower quality, number of decorations and mating success of male satin bowerbirds (Ptilonorhynchus violaceus): an experimental analysis" 1336:"Plasticity in extended phenotypes: how the antlion Myrmeleon crudelis adjusts the pit traps depending on biotic and abiotic conditions" 2296: 2206: 724:
and compressive strength. Males in some species of crab will construct structures out of mud to attract mates and avoid predators.
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predators have located the hideout or prey, or a combination of both. As a last resort, structures may provide means of escape.
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An animal builder may collect a variety of materials and use them in complex ways to form useful habitat. The nest of the
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places bits of grass and other material loosely streaming around its nest to break the shape and to masquerade as debris.
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Birds form the majority of the group of animals which collect building material of animal origin. They collect animal
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Some animals collect materials with plastic properties which are used to construct and shape the nest. These include
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Götmark, F. (1993). "Conspicuous nests may select for non-cryptic eggs: a comparative study of avian families".
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use materials such as spider webbing, silk, and lichen, while other species such as great crested flycatchers (
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with different forms carrying out specialized roles. These constructions may arise from complex building
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An interesting example is the case of silk caps which cover the pupal cells of the Oriental hornet
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for communication between members of the species (intra-specific communication), i.e. display.
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Mud is used by a few species of a wide variety of families including wasps and birds. Mud is
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bowerbirds, as the quality of the constructed bowers reflects the quality of the male bird.
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and feathers of other species of birds to line their nests. Almost 56% of all families of
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Graham, Jason M.; Kao, Albert B.; Wilhelm, Dylana A.; Garnier, Simon (21 December 2017).
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Recently, some researchers have argued that the structures built by animals affect the
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wasps collect mud and blend them with water to construct free standing nests of mud.
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when dried. Amongst birds, 5% of all birds use mud and stones in their nest for
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Vaughan, Terry A.; Ryan, James M.; Czaplewski, Nicholas J. (15 February 2010).
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Among the structures created by animals to prevent predation are those of the
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Christy, John H.; Backwell, Patricia R.; Schober, Ursula (1 January 2003).
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Christy, John H.; Backwell, Patricia R.; Schober, Ursula (January 2003).
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Farji-Brener, Alejandro G.; Amador-Vargas, Sabrina (19 December 2020).
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shelters for other purposes such as reproduction, food storage, etc.
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while leaves and green twigs provide flexible but strong supports.
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Temperature extremes harm animals irrespective of whether they are
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Often, these structures incorporate sophisticated features such as
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properties, the silk stores excess daytime heat in the form of
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Animal architects: building and the evolution of intelligence
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Built by animals: the natural history of animal architecture
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Built by animals: the natural history of animal architecture
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Frith, Clifford B.; Frith, Dawn W.; Barnes, Eustace (2004).
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which depends upon crypsis to avoid detection of its nest
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ZooKeys 1177: 87–146. DOI: 10.3897/zookeys.1177.102600.
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cut and shape thorns or twigs to form their case. Some
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Animals create structures primarily for three reasons:
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may also be involved in building structures by animals.
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The ethology of domestic animals: an introductory text
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are the largest structures built by non-human animals.
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Gould, James L.; Gould, Carol Grant (12 March 2007).
782:), which produces down feathers for lining its nest. 1563:Bohart, Richard Mitchell; Menke, Arnold S. (1976). 1231:. Jones & Bartlett Learning. pp. 440–441. 876: 1641: 1186: 952: 1490:"Optimal construction of army ant living bridges" 1267: 2283: 1594: 1268:Scharf, Inon; Lubin, Yael; Ovadia, Ofer (2011). 990: 946: 944: 1785: 1783: 1556: 1549: 1547: 1545: 1543: 1541: 1539: 1218: 172:Building behaviour is common in many non-human 1635: 1633: 1631: 1566:Sphecid wasps of the world: a generic revision 1119:Furuichi, Sho; Kasuya, Eiiti (February 2014). 1118: 2202:Association for the Study of Animal Behaviour 1846: 1033: 1031: 1029: 1027: 1025: 1023: 1021: 984: 941: 1780: 1676: 1600: 1536: 802: 647: 366:Communal silk nests of the small eggar moth 135:responses and interactions, as exhibited by 111:such as in the case of night-time nests for 33:A so-called "cathedral" mound produced by a 1628: 1562: 1178: 1159: 1065: 235:to catch prey and for foraging, i.e. traps. 2207:International Society for Applied Ethology 1853: 1839: 1569:. University of California Press. p.  1018: 950: 811:of the constructor, a phenomenon known as 736: 553:. Other materials need to be "processed". 1078:. Oxford University Press. pp. 1–2. 1044:. Oxford University Press. pp. 1–2. 910: 842: 451:Bowerbird in front of a constructed bower 232:to create protected habitats, i.e. homes. 1789: 1601:Burton, Maurice; Burton, Robert (1970). 882: 751: 740: 675: 505: 471: 446: 361: 346:In a case apparently of masquerade, the 262: 250: 157: 28: 1790:Capinera, John L. (17 September 2008). 1639: 1604:The international wildlife encyclopedia 1340:Israel Journal of Ecology and Evolution 1184: 1165: 997:. Oxford University Press. p. 12. 593:, is constructed from four materials – 14: 2284: 1718: 1716: 1714: 1712: 1710: 1708: 1429: 1193:. Cambridge University Press. p.  855:. PHI Learning Pvt. Ltd. p. 100. 849:Mandal, Fatik Baran (1 January 2010). 848: 625:About the construction of nest by the 501: 188:. 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In Jensen, Per (ed.). 666:Arachnothera longirostra 387:, as in the case of the 315:) and long-tailed tits ( 2116:Irenäus Eibl-Eibesfeldt 1896:Animal sexual behaviour 799:resides in its cocoon. 737:Self-secreted materials 629:, it has been written: 389:Arctic ground squirrels 168:) starting a new colony 2055:Tool use by non-humans 2008:Philosophical ethology 1953:Comparative psychology 1901:Animal welfare science 760: 749: 716:when wet and provides 689: 636: 515: 482: 452: 372: 272: 260: 169: 89:temperature regulation 38: 755: 744: 679: 632: 509: 475: 450: 365: 351:Cranioleuca erythrops 333:Terpsiphone paradisei 323:and common waxbills ( 266: 254: 161: 32: 2161:William Homan Thorpe 1926:Behavioural genetics 1886:Animal consciousness 1881:Animal communication 927:. CABI. p. 34. 780:Somateria mollissima 718:compressive strength 481:sp. forming a bridge 398:Eriogaster lanestris 369:Eriogaster lanestris 317:Aegithalos caudatus) 109:behaviour of animals 1916:Behavioural ecology 1506:2017JThBi.435..184G 1075:Animal architecture 1041:Animal architecture 886:Chimpanzee cultures 662:little spiderhunter 591:Aegithalos caudatus 502:Collected materials 348:red-faced spinetail 329:paradise flycatcher 321:Myiarchus crinitus) 313:Polioptila caerulea 269:red-faced spinetail 257:red-wattled lapwing 129:emergent properties 46:animal architecture 18:Animal architecture 2245:Behavioral Ecology 2166:Nikolaas Tinbergen 1958:Emotion in animals 1936:Cognitive ethology 1274:Biological Reviews 813:niche construction 796:Aethria carnicauda 761: 750: 746:Western honey bees 690: 603:spider egg cocoons 570:queens build with 516: 493:Building materials 483: 453: 437:Myrmeleon crudelis 373: 309:avoiding detection 273: 261: 247:Protected habitats 170: 39: 2279: 2278: 2171:Jakob von UexkĂĽll 1941:Comfort behaviour 1803:978-1-4020-6242-1 1659:978-0-19-920556-1 1614:978-0-7614-7266-7 1580:978-0-520-02318-5 1238:978-0-7637-6299-5 1204:978-0-521-46038-5 1085:978-0-19-850752-9 1051:978-0-19-850752-9 1004:978-0-19-854844-7 970:978-0-465-02782-8 934:978-1-84593-536-8 896:978-0-674-11663-4 862:978-81-203-4035-0 165:Polistes dominula 16:(Redirected from 2304: 2269: 2268: 2231:Animal Cognition 2224:Animal Behaviour 2176:Wolfgang Wickler 1876:Animal cognition 1855: 1848: 1841: 1832: 1831: 1825: 1821: 1815: 1814: 1812: 1810: 1787: 1778: 1777: 1737: 1728: 1727: 1720: 1683: 1680: 1674: 1673: 1668: 1666: 1647: 1637: 1626: 1625: 1623: 1621: 1598: 1592: 1591: 1589: 1587: 1560: 1554: 1551: 1534: 1533: 1485: 1468: 1467: 1436:Animal Behaviour 1427: 1416: 1415: 1375: 1364: 1363: 1331: 1314: 1313: 1265: 1250: 1249: 1247: 1245: 1222: 1216: 1215: 1213: 1211: 1192: 1182: 1176: 1175: 1163: 1157: 1156: 1116: 1097: 1096: 1094: 1092: 1069: 1063: 1062: 1060: 1058: 1035: 1016: 1015: 1013: 1011: 988: 982: 981: 979: 977: 958: 948: 939: 938: 914: 908: 907: 905: 903: 880: 874: 873: 871: 869: 846: 769:reworked in the 686:Myodes glareolus 643: 420:electric current 407:Vespa orientalis 385:thermoregulation 358:Thermoregulation 325:Estrilda astrild 117:neural responses 21: 2312: 2311: 2307: 2306: 2305: 2303: 2302: 2301: 2282: 2281: 2280: 2275: 2257: 2211: 2190: 2186:Solly Zuckerman 2126:Karl von Frisch 2111:Richard Dawkins 2096:John B. Calhoun 2081:Patrick Bateson 2069: 2003:Pain in animals 1864: 1859: 1829: 1828: 1822: 1818: 1808: 1806: 1804: 1788: 1781: 1738: 1731: 1726:. 1 April 2008. 1722: 1721: 1686: 1681: 1677: 1664: 1662: 1660: 1638: 1629: 1619: 1617: 1615: 1599: 1595: 1585: 1583: 1581: 1561: 1557: 1552: 1537: 1486: 1471: 1428: 1419: 1376: 1367: 1332: 1317: 1266: 1253: 1243: 1241: 1239: 1223: 1219: 1209: 1207: 1205: 1183: 1179: 1164: 1160: 1117: 1100: 1090: 1088: 1086: 1070: 1066: 1056: 1054: 1052: 1036: 1019: 1009: 1007: 1005: 989: 985: 975: 973: 971: 949: 942: 935: 917:A.; L. (2009). 915: 911: 901: 899: 897: 881: 877: 867: 865: 863: 847: 843: 838: 821: 805: 739: 710: 674: 650: 645: 638: 627:long-tailed tit 587:long-tailed tit 512:long-tailed tit 504: 495: 470: 445: 432: 416:electric charge 360: 249: 226: 216:and some other 115:, from inbuilt 44:, often called 23: 22: 15: 12: 11: 5: 2310: 2300: 2299: 2294: 2277: 2276: 2274: 2273: 2262: 2259: 2258: 2256: 2255: 2248: 2241: 2238:Animal Welfare 2234: 2227: 2219: 2217: 2213: 2212: 2210: 2209: 2204: 2198: 2196: 2192: 2191: 2189: 2188: 2183: 2178: 2173: 2168: 2163: 2158: 2153: 2151:Desmond Morris 2148: 2143: 2138: 2133: 2128: 2123: 2118: 2113: 2108: 2106:Marian Dawkins 2103: 2101:Charles Darwin 2098: 2093: 2088: 2083: 2077: 2075: 2071: 2070: 2068: 2067: 2062: 2057: 2052: 2047: 2046: 2045: 2040: 2035: 2030: 2020: 2015: 2010: 2005: 2000: 1995: 1990: 1985: 1983:Human ethology 1980: 1975: 1970: 1965: 1960: 1955: 1950: 1949: 1948: 1938: 1933: 1928: 1923: 1918: 1913: 1908: 1903: 1898: 1893: 1891:Animal culture 1888: 1883: 1878: 1872: 1870: 1866: 1865: 1858: 1857: 1850: 1843: 1835: 1827: 1826: 1816: 1802: 1779: 1729: 1684: 1675: 1658: 1627: 1613: 1593: 1579: 1555: 1535: 1469: 1442:(1): 266–271. 1417: 1365: 1346:(1–2): 41–47. 1315: 1280:(3): 626–639. 1251: 1237: 1217: 1203: 1177: 1158: 1131:(2): 215–221. 1098: 1084: 1064: 1050: 1017: 1003: 983: 969: 940: 933: 909: 895: 875: 861: 840: 839: 837: 834: 833: 832: 827: 820: 817: 804: 801: 748:on a wild nest 738: 735: 709: 708:Mud and stones 706: 673: 672:Plant material 670: 649: 646: 640:Niko Tinbergen 631: 524:stingless bees 503: 500: 494: 491: 487:Eciton hamatum 469: 468:Transportation 466: 444: 441: 431: 428: 412:thermoelectric 359: 356: 248: 245: 240: 239: 236: 233: 225: 222: 105:social animals 74:nests of birds 50:termite mounds 9: 6: 4: 3: 2: 2309: 2298: 2295: 2293: 2290: 2289: 2287: 2272: 2264: 2263: 2260: 2254: 2253: 2249: 2247: 2246: 2242: 2240: 2239: 2235: 2233: 2232: 2228: 2226: 2225: 2221: 2220: 2218: 2214: 2208: 2205: 2203: 2200: 2199: 2197: 2193: 2187: 2184: 2182: 2179: 2177: 2174: 2172: 2169: 2167: 2164: 2162: 2159: 2157: 2156:Thomas Sebeok 2154: 2152: 2149: 2147: 2146:Konrad Lorenz 2144: 2142: 2141:Julian Huxley 2139: 2137: 2136:Heini Hediger 2134: 2132: 2129: 2127: 2124: 2122: 2119: 2117: 2114: 2112: 2109: 2107: 2104: 2102: 2099: 2097: 2094: 2092: 2089: 2087: 2084: 2082: 2079: 2078: 2076: 2072: 2066: 2065:Zoomusicology 2063: 2061: 2058: 2056: 2053: 2051: 2048: 2044: 2041: 2039: 2036: 2034: 2031: 2029: 2026: 2025: 2024: 2021: 2019: 2016: 2014: 2011: 2009: 2006: 2004: 2001: 1999: 1998:Neuroethology 1996: 1994: 1991: 1989: 1986: 1984: 1981: 1979: 1976: 1974: 1971: 1969: 1966: 1964: 1961: 1959: 1956: 1954: 1951: 1947: 1944: 1943: 1942: 1939: 1937: 1934: 1932: 1929: 1927: 1924: 1922: 1919: 1917: 1914: 1912: 1909: 1907: 1906:Anthrozoology 1904: 1902: 1899: 1897: 1894: 1892: 1889: 1887: 1884: 1882: 1879: 1877: 1874: 1873: 1871: 1867: 1863: 1856: 1851: 1849: 1844: 1842: 1837: 1836: 1833: 1820: 1805: 1799: 1795: 1794: 1786: 1784: 1775: 1771: 1767: 1763: 1759: 1755: 1751: 1747: 1743: 1736: 1734: 1725: 1719: 1717: 1715: 1713: 1711: 1709: 1707: 1705: 1703: 1701: 1699: 1697: 1695: 1693: 1691: 1689: 1679: 1672: 1661: 1655: 1651: 1646: 1645: 1636: 1634: 1632: 1616: 1610: 1606: 1605: 1597: 1582: 1576: 1572: 1568: 1567: 1559: 1550: 1548: 1546: 1544: 1542: 1540: 1531: 1527: 1523: 1519: 1515: 1511: 1507: 1503: 1499: 1495: 1491: 1484: 1482: 1480: 1478: 1476: 1474: 1465: 1461: 1457: 1453: 1449: 1445: 1441: 1437: 1433: 1426: 1424: 1422: 1413: 1409: 1405: 1401: 1397: 1393: 1389: 1385: 1381: 1374: 1372: 1370: 1361: 1357: 1353: 1349: 1345: 1341: 1337: 1330: 1328: 1326: 1324: 1322: 1320: 1311: 1307: 1303: 1299: 1295: 1291: 1287: 1283: 1279: 1275: 1271: 1264: 1262: 1260: 1258: 1256: 1240: 1234: 1230: 1229: 1221: 1206: 1200: 1196: 1191: 1190: 1181: 1173: 1169: 1162: 1154: 1150: 1146: 1142: 1138: 1134: 1130: 1126: 1122: 1115: 1113: 1111: 1109: 1107: 1105: 1103: 1087: 1081: 1077: 1076: 1068: 1053: 1047: 1043: 1042: 1034: 1032: 1030: 1028: 1026: 1024: 1022: 1006: 1000: 996: 995: 987: 972: 966: 962: 957: 956: 947: 945: 936: 930: 926: 925: 920: 913: 898: 892: 888: 887: 879: 864: 858: 854: 853: 845: 841: 831: 828: 826: 823: 822: 816: 814: 810: 800: 797: 793: 789: 783: 781: 777: 772: 767: 758: 754: 747: 743: 734: 731: 727: 723: 719: 715: 705: 701: 699: 696: 687: 683: 678: 669: 667: 663: 659: 655: 644: 641: 635: 630: 628: 623: 621: 616: 612: 608: 604: 600: 596: 592: 588: 583: 581: 577: 573: 569: 565: 561: 556: 552: 548: 543: 541: 538:collected by 537: 533: 530:collected by 529: 525: 522:collected by 521: 513: 508: 499: 490: 488: 480: 479: 474: 465: 462: 458: 457:fiddler crabs 449: 440: 438: 430:Trap building 427: 424: 421: 417: 413: 409: 408: 402: 400: 399: 392: 390: 386: 382: 378: 371: 370: 364: 355: 352: 349: 344: 340: 338: 334: 330: 326: 322: 318: 314: 310: 306: 302: 298: 294: 289: 286: 282: 277: 270: 265: 258: 253: 244: 237: 234: 231: 230: 229: 221: 219: 215: 211: 207: 203: 199: 195: 191: 187: 183: 179: 175: 167: 166: 160: 156: 154: 150: 146: 145:communication 142: 138: 134: 130: 126: 122: 118: 114: 110: 106: 102: 98: 94: 90: 85: 83: 79: 75: 71: 67: 63: 59: 55: 51: 47: 43: 36: 31: 27: 19: 2250: 2243: 2236: 2229: 2222: 2181:E. O. Wilson 2131:Jane Goodall 2091:Donald Broom 2060:Zoosemiotics 2022: 2013:Sociobiology 1819: 1807:. Retrieved 1792: 1752:(2): 84–91. 1749: 1745: 1723: 1678: 1670: 1663:. Retrieved 1643: 1618:. Retrieved 1603: 1596: 1584:. Retrieved 1565: 1558: 1497: 1493: 1439: 1435: 1390:(2): 84–91. 1387: 1383: 1343: 1339: 1277: 1273: 1242:. Retrieved 1227: 1220: 1208:. Retrieved 1188: 1180: 1171: 1167: 1161: 1128: 1124: 1089:. Retrieved 1074: 1067: 1055:. Retrieved 1040: 1008:. Retrieved 993: 986: 974:. Retrieved 954: 923: 912: 900:. Retrieved 885: 878: 866:. Retrieved 851: 844: 806: 795: 791: 784: 779: 776:common eider 762: 729: 725: 711: 702: 695:cantilevered 691: 685: 665: 651: 637: 633: 624: 590: 584: 544: 540:hummingbirds 517: 496: 486: 484: 476: 460: 454: 436: 433: 425: 405: 403: 396: 393: 374: 367: 350: 345: 341: 332: 324: 320: 316: 312: 290: 284: 278: 274: 241: 227: 206:urochordates 171: 163: 131:from simple 86: 72:, elaborate 45: 41: 40: 26: 2121:Dian Fossey 2086:Marc Bekoff 2074:Ethologists 1500:: 184–198. 792:Bombyx mori 766:ecribellate 757:Beaver dams 730:Uca beebei, 485:Army ants ( 381:ectothermic 377:endothermic 301:concealment 281:paper wasps 210:crustaceans 133:instinctive 113:chimpanzees 101:ventilation 70:beaver dams 68:complexes, 2286:Categories 2023:Structures 2018:Stereotypy 1174:: 102–105. 1168:Ornis Fenn 836:References 726:Uca musica 682:bank voles 572:paper pulp 568:Paper wasp 337:ethologist 297:camouflage 218:arthropods 198:amphibians 149:aesthetics 2252:Behaviour 2195:Societies 2033:Honeycomb 1766:0340-5443 1522:0022-5193 1456:0003-3472 1404:1432-0762 1360:2224-4662 1310:205599463 1294:1469-185X 1228:Mammalogy 1210:10 August 1145:0340-5443 809:evolution 771:cribellum 722:toughness 658:passerine 555:Caddisfly 551:chaffinch 224:Functions 186:arachnids 54:ant hills 2292:Ethology 2271:Category 2216:Journals 2043:Instinct 1993:Learning 1988:Instinct 1963:Ethogram 1946:Grooming 1869:Branches 1862:Ethology 1774:34001395 1530:28939347 1464:53192968 1412:34001395 1302:21062400 1153:15263361 819:See also 788:silkworm 599:feathers 560:bagworms 549:for the 532:swallows 443:Displays 214:annelids 202:molluscs 194:reptiles 153:Tool use 141:learning 137:termites 127:, or as 62:beehives 1973:Feeding 1671:velcro. 1502:Bibcode 1244:10 July 1091:29 June 1057:29 June 868:29 June 714:plastic 642:, 1953. 615:bramble 564:sphecid 305:mimicry 293:crypsis 182:insects 174:mammals 121:hormone 82:spiders 35:termite 1809:2 July 1800:  1772:  1764:  1665:1 July 1656:  1652:, 77. 1620:1 July 1611:  1586:1 July 1577:  1528:  1520:  1462:  1454:  1410:  1402:  1358:  1308:  1300:  1292:  1235:  1201:  1151:  1143:  1082:  1048:  1010:1 July 1001:  976:1 July 967:  931:  902:2 July 893:  859:  778:duck ( 680:Young 620:velcro 595:lichen 580:saliva 478:Eciton 307:, for 303:, and 76:, and 66:burrow 37:colony 2050:Swarm 1978:Hover 1931:Breed 1770:S2CID 1460:S2CID 1408:S2CID 1306:S2CID 1149:S2CID 963:–18. 698:beams 611:gorse 520:resin 178:birds 93:traps 2038:Nest 2028:Hive 1811:2011 1798:ISBN 1762:ISSN 1667:2011 1654:ISBN 1622:2011 1609:ISBN 1588:2011 1575:ISBN 1526:PMID 1518:ISSN 1452:ISSN 1400:ISSN 1356:ISSN 1298:PMID 1290:ISSN 1246:2011 1233:ISBN 1212:2011 1199:ISBN 1141:ISSN 1093:2011 1080:ISBN 1059:2011 1046:ISBN 1012:2011 999:ISBN 978:2011 965:ISBN 929:ISBN 904:2011 891:ISBN 870:2011 857:ISBN 607:moss 605:and 576:wood 547:hair 536:silk 534:and 267:The 190:fish 184:and 143:and 125:sows 97:bait 78:webs 60:and 58:wasp 1754:doi 1510:doi 1498:435 1444:doi 1392:doi 1348:doi 1282:doi 1195:103 1133:doi 654:fur 613:or 528:mud 379:or 295:or 80:of 2288:: 1782:^ 1768:. 1760:. 1750:53 1748:. 1744:. 1732:^ 1687:^ 1669:. 1650:76 1630:^ 1573:. 1538:^ 1524:. 1516:. 1508:. 1496:. 1492:. 1472:^ 1458:. 1450:. 1440:33 1438:. 1434:. 1420:^ 1406:. 1398:. 1388:53 1386:. 1382:. 1368:^ 1354:. 1344:66 1342:. 1338:. 1318:^ 1304:. 1296:. 1288:. 1278:86 1276:. 1272:. 1254:^ 1197:. 1172:70 1170:. 1147:. 1139:. 1129:68 1127:. 1123:. 1101:^ 1020:^ 943:^ 815:. 601:, 597:, 589:, 542:. 526:, 510:A 391:. 299:, 283:, 212:, 208:, 204:, 200:, 196:, 192:, 180:, 176:, 151:. 99:, 95:, 91:, 84:. 64:, 56:, 52:, 1854:e 1847:t 1840:v 1813:. 1776:. 1756:: 1624:. 1590:. 1571:2 1532:. 1512:: 1504:: 1466:. 1446:: 1414:. 1394:: 1362:. 1350:: 1312:. 1284:: 1248:. 1214:. 1155:. 1135:: 1095:. 1061:. 1014:. 980:. 961:1 937:. 906:. 872:. 790:( 684:( 664:( 331:( 20:)

Index

Animal architecture

termite
termite mounds
ant hills
wasp
beehives
burrow
beaver dams
nests of birds
webs
spiders
temperature regulation
traps
bait
ventilation
social animals
behaviour of animals
chimpanzees
neural responses
hormone
sows
emergent properties
instinctive
termites
learning
communication
aesthetics
Tool use

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