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Archibald Hill

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1358: 2547: 1931: 1396:, he met during the course of his long and extraordinary life. After-dinner conversations in the drawing room would inevitably involve passionate debates about science or politics. "Every Sunday we would have to attend a tea party at grandpa’s house and apart from entertaining some extraordinary guests, he would devise some great games for us, such as frog racing in the garden or looking through the lens of a (dissected) sheep’s eye". Sir 1645: 44: 421:. Someone dubbed his motley group "Hill's Brigands", which they proudly adopted. Later in the war they also worked on locating enemy planes from their sound. He sped between their working sites on his beloved motorcycle. At the end of the war, Major Hill issued certificates to more than one hundred Brigands. He was appointed an 485:
from 1940 to 1945. In 1940, he was posted to the British Embassy in Washington to promote war research in the still neutral United States. He was authorized to swap secrets with the Americans, but this could not work: how do you place a value on another's secret? Hill saw the answer and persuaded the
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who proposed the Blue plaque, said: "The Nobel Prize winner A. V. Hill contributed vastly to our understanding of muscle physiology. His work has resulted in wide-ranging application in sports medicine. As an outstanding Humanitarian and Parliamentarian, he was uncompromising in his condemnation of
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persecution. He prominently displayed in his laboratory a toy figure of Adolf Hitler with saluting arm upraised, which he explained was in gratitude for all the scientists Germany had expelled, some of whom were now working with him. Hill believed that "Laughter is the best detergent for nonsense".
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Hill made many exacting measurements of the heat released when skeletal muscles contract and relax. A key finding was that heat is produced during contraction, which requires investment of chemical energy, but not during relaxation, which is passive. His earliest measurements used equipment left
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to come for a day to advise them on how to train anti-aircraft gunners. On site, Hill immediately proposed a simple two mirror method to determine airplanes' heights. Transferred to Munitions, he realized that the mirrors could measure where smoke shells burst and if he fitted this data with the
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was president. Both had been frustrated by the delay in putting scientists to work in the previous war. The Royal Society collated a list of scientists and Hill represented the Society at the Ministry of Labour. When the war came Hill led a campaign to liberate refugee scientists who had been
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and he went to Germany to learn more about their work. He continually improved his apparatus to make it more sensitive and to reduce the time lag between the heat released by the preparation and its recording by his
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from 1955 to 1960. In 1967, he retired to Cambridge where he gradually lost the use of his legs. He died "held in the greatest affection by more than a hundred scientific descendants all over the world".
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After the war he rebuilt his laboratory at University College and vigorously carried on research. In 1951, his advocacy was rewarded by the establishment of a Biophysics Department under his leadership.
358:. In this paper, Hill's first publication, he derived both the equilibrium form of the Langmuir equation, and also the exponential approach to equilibrium. The paper, written under the supervision of 606: 1206:, Hill elucidated the processes whereby mechanical work is produced in muscles. The two shared the 1922 Nobel Prize in Physiology and Medicine for this work. Hill introduced the concepts of 1073: 4901: 3097: 954: 1404:
the Nazi regime for its persecution of scientists and others. A. V. Hill played a crucial role in assisting and rescuing many refugees to continue their work in this country".
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British to show the Americans everything they were working on (except for the atomic bomb). The mobilization of Allied scientists was one of the major successes in the war.
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Although Hill's work in muscle physiology is probably the most important, and certainly responsible for his Nobel Prize, he is also very well known in biochemistry for the
4856: 698: 4916: 2648: 1198:. Using himself as the subject — he ran every morning from 7:15 to 10:30 — he showed that running a dash relies on energy stores which afterwards are replenished by 857: 801: 747: 1128: 903: 827: 2370: 1148: 974: 923: 877: 767: 718: 669: 649: 629: 4981: 4926: 1774: 1670: 489:
He visited India between November 1943 and April 1944 to survey scientific and technological research. His suggestions influenced the establishment of the
3090: 509: 4921: 4640: 1380:, where Hill was a Governor from 1929 to 1960. It has now been sold, redeveloped and renamed as Hurstbourne. In Hill's time, according to his grandson 501: 3106: 2577: 482: 4936: 1167:, Hill measured a temperature rise of only 0.003 °C. After publication he learned that German physiologists had already reported on heat and 4861: 2270:. Washington, DC: American Academy of Physical Education and the American Alliance for Health, Physical Education and Recreation. pp. 63–65. 1332: 2061:
Hale, Tudor (15 February 2008). "History of developments in sport and exercise physiology: A. V. Hill, maximal oxygen uptake, and oxygen debt".
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Trails and Trials in Physiology: A Bibliography, 1909–1964; with reviews of certain topics and methods and a reconnaissance for further research
4871: 3083: 4866: 2641: 436:, a few years later becoming a Royal Society Research professor there, where he remained until retirement in 1951. In 1933, he became with 2474: 4986: 4598: 4587: 4572: 4553: 1807:"The mode of action of nicotine and curari, determined by the form of the contraction curve and the method of temperature coefficients" 505: 422: 283: 4971: 4966: 401:
describing a shell's flight they could provide accurate range tables for anti-aircraft guns. To measure and compute he assembled the
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theory, because the context for the derivation was the binding of nicotine and curare to the "receptive substance" at the
2597: 2589: 2565: 402: 351:. While still an undergraduate at Trinity College, he derived in 1909 what came to be known as the Langmuir equation. 340: 2317: 4825: 4820: 4815: 4810: 4805: 4795: 4790: 4785: 4780: 4775: 4626: 2569: 1191: 4941: 4886: 4881: 4470: 2556: 461: 2427: 2047: 4486: 3751: 1219: 1195: 490: 140: 3663: 2452: 2343: 4891: 4876: 4682: 3178: 1309: 4202: 4961: 4561: 4413: 4389: 2854: 989: 4951: 4110: 2281: 1326: 523: 130: 2305: 3313: 3075: 2990: 2942: 2371:"A prize property: this £10 million Highgate mansion was once home to a Nazi-fighting Nobel Prize winner" 1345: 1271: 1250: 287: 261: 928: 4956: 4090: 3871: 2601: 336: 207: 103: 17: 1891:"Mirrors and smoke: A. V. Hill, his Brigands, and the science of anti-aircraft gunnery in World War I" 1376:, where he had lived from 1923 to 1967. Since then the house had been divided into flats and owned by 4678: 3831: 3647: 3399: 3375: 2998: 2878: 2870: 2621: 2239:
Cardinal, Bradley J. (2022). "The National Academy of Kinesiology: Its founding, focus, and future".
1626: 1241:(1914–2005), economist, married K.A.C. Humphreys, registrar of the West African Examinations Council. 770: 433: 355: 3170: 2798: 2617: 1930: 1384:, regular guests at the house included 18 exiled Nobel laureates, his brother-in-law, the economist 4659: 3954: 2886: 2453:"A. V. Hill, Nobel Prize Winner and Sports Medicine Pioneer, receives English Heritage Blue Plaque" 478: 448:(which in 1936 became the Society for the Protection of Science and Learning). By the start of the 203: 4769: 3799: 3627: 2400: 382: 374: 363: 4066: 3455: 1078:
This shows that when the Hill equation is accurately obeyed (which usually it is not) a plot of
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these symbols are misleading if taken to imply that it shows the number of binding sites on the
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Hill returned briefly to Cambridge in 1919 before taking the chair in physiology at the
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Presidential Address to the British Association Meeting, held at Belfast in 1952
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Members of the Parliament of the United Kingdom for the University of Cambridge
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is an integer, and in most experimental cases, apart from the trivial case of
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Hitler's Gift: The True Story of the Scientists Expelled by the Nazi Regime
2090: 1923: 1907: 1840: 1750: 1701: 1684: 1665: 1661: 1568: 1543: 1453: 1436: 1427: 1338: 1173: 465: 414: 296: 235: 166: 3105: 2541: 2147: 1437:"Muscular Exercise, Lactic Acid, and the Supply and Utilisation of Oxygen" 4650: 4442: 4357: 4353: 4266: 4242: 4150: 3815: 3611: 3528: 3158: 3150: 3134: 3038: 2846: 2790: 2782: 2758: 2252: 1369: 1319: 1164: 532: 227: 156: 1871: 43: 4729: 4293: 3552: 3500: 3447: 2966: 2894: 1397: 1184: 516: 398: 348: 300: 189: 185: 126: 1856:"The adsorption of gases on plane surface of glass, mica and platinum" 1693: 1414:
Gray, C. H. (1947). "The significance of the van den Bergh reaction".
4494: 4046: 2114:"Maximal oxygen uptake: "classical" versus "contemporary" viewpoints" 1725:"Scientific contributions of A. V. Hill: exercise physiology pioneer" 83: 1855: 1256:
Janet Hill (1918–2000) child psychiatrist, married the immunologist
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Hill, A. V. (April 1926). "The scientific study of athletics".
1315: 528: 385:. The British made no effort to make use of their scientists. 344: 316: 117: 409:(a wounded officer), and lads too young for service including 469: 2432:
Hektoen International – An online medical humanities journal
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is a parameter that expresses the degree of departure from
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Nobel Prize Winners in Medicine and Physiology: 1901–1950
472:. He was also biological secretary of the Royal Society; 2511: 2026:. Weinheim, Germany: Wiley-Blackwell. pp. 286–288. 1666:"Archibald Vivian Hill. 26 September 1886 – 3 June 1977" 1372:
was erected at Hill's former home, 16 Bishopswood Road,
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At the end of 1915, while home on leave he was asked by
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Proceedings of the Royal Society B: Biological Sciences
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Proceedings of the Royal Society B: Biological Sciences
1247:(1915–2002), physiologist, married Stella Mary Humphrey 377:; he was a crack shot. In 1914, at the outbreak of the 4902:
Foreign associates of the National Academy of Sciences
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Muscular Activity: Herter Lectures – Sixteenth Course
1136: 1084: 992: 962: 931: 911: 885: 865: 839: 809: 783: 755: 726: 706: 677: 657: 637: 617: 601:{\displaystyle v=V{\frac {a^{h}}{K_{0.5}^{h}+a^{h}}}} 544: 1883: 1881: 1671:
Biographical Memoirs of Fellows of the Royal Society
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Cooperativity of protein binding and enzyme kinetics
299:, one of the founders of the diverse disciplines of 4857:Academics of the Victoria University of Manchester 2428:"How Britain rescued scientists from Nazi tyranny" 2399: 2310: 2015: 1889: 1778:(online ed.). Oxford University Press. 2004. 1595:The Ethical Dilemma of Science, and Other Writings 1504:. Philadelphia: University of Pennsylvania Press. 1142: 1122: 1067: 968: 948: 917: 897: 871: 851: 821: 795: 761: 741: 712: 692: 663: 643: 623: 600: 502:British Association for the Advancement of Science 311:for his elucidation of the production of heat and 2656: 2021: 1937: 1878: 1581:. Philadelphia: University of Pennsylvania Press. 1253:(1919–1966), oceanographer, married Philippa Pass 4917:Members of the Order of the Companions of Honour 4838: 2475:"Famous homes in north London with many stories" 2393: 2391: 2048:"The Nobel Prize in Physiology or Medicine 1922" 1435:Hill, A. V.; Long, C. N. H.; Lupton, H. (1924). 1388:, and also friends as diverse and unexpected as 980:. Hill himself avoided any such interpretation. 2530: 2318:"On Archibald Vivien Hill and the honours list" 1961:Hitler's gift: Scientists who fled Nazi Germany 1772:"The Oxford Dictionary of National Biography". 1333:Member of the Order of the Companions of Honour 4648: 1434: 4982:Members of the American Philosophical Society 4634: 3091: 2642: 2388: 2111: 1943: 1887: 1618: 1361:Blue plaque at 16 Bishopswood Road, Highgate. 2362: 2214:The Journal of Health and Physical Education 1985:All Hell let loose: The World at War 1939-45 1523:. Baltimore: Williams & Wilkins Company. 4927:Officers of the Order of the British Empire 2118:Medicine and Science in Sports and Exercise 1847: 983:The equation can be rearranged as follows: 444:a founder member and vice-president of the 4641: 4627: 3098: 3084: 2649: 2635: 2545: 2516:(Paperback). New York: Arcade Publishing. 2024:Fundamentals of Enzyme Kinetics (4th edn.) 1656: 1654: 1325:International Member of the United States 506:International Council of Scientific Unions 423:Officer of the Order of the British Empire 405:, a team of men too old for conscription, 381:, Hill became the musketry officer of the 4922:Nobel laureates in Physiology or Medicine 2397: 2129: 1830: 1740: 1716: 1683: 1567: 1452: 859:. Note that there is no implication that 631:is the rate of reaction at concentration 2502: 2401:"Houses stamped with a mark of prestige" 2368: 2238: 1982: 1888:Van der Kloot, William (December 2011). 1860:Journal of the American Chemical Society 1853: 1597:. New York: Rockefeller Institute Press. 1356: 1296:Nerves and Muscles: How We Feel and Move 2419: 2166:American Academy of Arts & Sciences 1798: 1775:Oxford Dictionary of National Biography 1722: 1651: 1625:Nobel Prize Outreach AB (3 June 1977). 1407: 1290:In 1926, he was invited to deliver the 1234:. They had two sons and two daughters: 1222:(1885-1974), daughter of the economist 1068:{\displaystyle \ln=h\ln a-h\ln K_{0.5}} 905:, it is not. Although many authors use 527:, which is used to quantify binding of 373:While a student he had enrolled in the 14: 4862:Academics of University College London 4839: 2507:. Philadelphia: W.B. Saunders Company. 2425: 2344:"Blue Plaque at AV Hill's former home" 2112:Bassett, D. R.; Howley, E. T. (1997). 1973:Van der Kloot, William (2014). p. 202. 1896:Notes and Records of the Royal Society 535:, written here as a kinetic equation: 4872:British Army personnel of World War I 4622: 3111:Nobel Prize in Physiology or Medicine 3079: 2630: 2265: 1948:. Stroud: Fonthill. pp. 191–214. 1518: 1499: 1303:American Academy of Arts and Sciences 1301:International Honorary Member of the 1285:Nobel Prize in Physiology or Medicine 309:Nobel Prize in Physiology or Medicine 152:Nobel Prize in Physiology or Medicine 4937:People educated at Blundell's School 4867:Alumni of Trinity College, Cambridge 2398:Rowlinson, Liz (18 September 2015). 2369:Crossley, Frankie (17 August 2017). 2060: 1997: 1804: 1660: 1601: 1592: 1576: 1541: 1468: 1413: 1264: 1157: 468:on the committee that gave birth to 2512:Medawar, Jean; Pyke, David (2012). 1946:Great Scientists wage the Great War 1765: 1587:Chemical Wave Transmission in Nerve 1292:Royal Institution Christmas Lecture 1163:behind by the Swedish physiologist 24: 2472: 1493:10.1038/scientificamerican0426-224 949:{\displaystyle {n_{\mathrm {H} }}} 939: 403:Anti-Aircraft Experimental Section 362:, is a landmark in the history of 25: 4998: 4987:Recipients of the Cothenius Medal 2610: 1959:Jean Medawar; David Pyke (2001). 1463:Textbook of Anti-Aircraft Gunnery 1416:The Quarterly Journal of Medicine 1202:. Paralleling the work of German 1192:Victoria University of Manchester 4972:Jodrell Professors of Physiology 4967:Cambridgeshire Regiment officers 2557:Parliament of the United Kingdom 2505:Lectures on nutrition: 1924–1925 2131:10.1097/00005768-199705000-00002 1929: 1643: 1213: 493:(IITs) in the following decade. 42: 27:British physiologist (1886–1977) 4977:Military personnel from Bristol 2466: 2426:Pearce, J.M.S. (2 April 2021). 2299: 2274: 2259: 2232: 2212:"It may interest you to know". 2205: 2180: 2154: 2105: 2054: 2040: 2006: 2000:Trails and Trials in Physiology 1991: 1976: 1967: 1963:. London: Piatkus. p. 122. 1944:Van der Kloot, William (2014). 1742:10.1152/japplphysiol.01246.2001 1130:gives a straight line of slope 500:In 1952, he became head of the 491:Indian Institutes of Technology 4947:Recipients of the Copley Medal 4932:British operations researchers 2226:10.1080/23267240.1938.10619861 1952: 1854:Langmuir, Irving (June 1918). 1823:10.1113/jphysiol.1909.sp001344 1352: 1310:American Philosophical Society 1226:, and sister of the economist 1117: 1114: 1102: 1091: 1025: 1022: 1010: 999: 481:Member of Parliament (MP) for 432:as professor of physiology at 13: 1: 2855:Edward Albert Sharpey-Schafer 2012:Katz, Bernard (1978). p. 133. 1987:. London: Harper. p. 81. 1729:Journal of Applied Physiology 1611: 1179:Hill is regarded, along with 510:Marine Biological Association 504:and Secretary General of the 4897:Fellows of the Royal Society 1792:UK public library membership 1327:National Academy of Sciences 1308:International Member of the 1200:increased oxygen consumption 1183:, as one of the founders of 671:is the rate at saturation, 322: 252:He is notably the father of 131:Hill equation (biochemistry) 7: 4203:Christiane Nüsslein-Volhard 2943:Charles Thomson Rees Wilson 1341:of the Royal Society (1948) 1318:in the National Academy of 1272:Order of the British Empire 446:Academic Assistance Council 354:This is closely related to 10: 5003: 2598:Sir Kenneth Pickthorn, Bt. 2590:Sir Kenneth Pickthorn, Bt. 2566:Sir Kenneth Pickthorn, Bt. 2537:. New York: Henry Schuman. 2496: 2063:Journal of Sports Sciences 2022:Cornish-Bowden A. (2012). 508:. He was President of the 477:interned. He served as an 337:Trinity College, Cambridge 208:University College, London 104:Trinity College, Cambridge 4764: 4739: 4720: 4701: 4677: 4658: 4276: 3941: 3602: 3400:Charles Scott Sherrington 3376:Frederick Gowland Hopkins 3338: 3117: 2879:Charles Scott Sherrington 2871:Frederick Gowland Hopkins 2665: 2622:Churchill Archives Centre 2594: 2576:Member of Parliament for 2574: 2562: 2555: 2348:The Physiological Society 2266:Scott, M. Gladys (1978). 2075:10.1080/02640410701701016 1811:The Journal of Physiology 1723:Bassett, D.R. Jr (2002). 1210:and oxygen debt in 1922. 1194:in 1920 in succession to 771:Michaelis-Menten kinetics 693:{\displaystyle {K_{0.5}}} 434:University College London 356:Michaelis-Menten kinetics 264:, and the grandfather of 250: 245: 241: 223: 213: 195: 181: 174: 146: 136: 109: 99: 91: 72: 50: 41: 34: 4390:Françoise Barré-Sinoussi 2887:Charles Algernon Parsons 2531:Stevenson, L.G. (1953). 2002:. London: Edward Arnold. 1579:Adventures in Biophysics 1529:Muscular Movement in Man 1461:Hill, A.V. (1924–1925). 1365:On 9 September 2015, an 460:In 1935, he served with 204:University of Manchester 122:Muscle contraction model 4942:Scientists from Bristol 4887:British Nobel laureates 4882:English Nobel laureates 2322:DC's Improbable Science 2162:"Archibald Vivian Hill" 1220:Margaret Neville Keynes 803:, no cooperativity for 383:Cambridgeshire Regiment 375:Officers Training Corps 141:Margaret Neville Keynes 4770:Nobel Prize recipients 4741:Physiology or Medicine 3171:Santiago Ramón y Cajal 3015:Geoffrey Ingram Taylor 2799:Pierre Paul Émile Roux 1983:Hastings, Max (2011). 1908:10.1098/rsnr.2010.0090 1784:10.1093/ref:odnb/31230 1685:10.1098/rsbm.1978.0005 1569:10.1098/rspb.1928.0029 1544:"Myothermic apparatus" 1454:10.1098/rspb.1924.0037 1362: 1144: 1124: 1069: 970: 950: 919: 899: 873: 853: 852:{\displaystyle h<1} 831:negative cooperativity 823: 797: 796:{\displaystyle h>1} 775:positive cooperativity 763: 743: 742:{\displaystyle v=0.5V} 714: 694: 665: 645: 625: 602: 428:In 1923, he succeeded 368:neuromuscular junction 218:Walter Morley Fletcher 4892:English physiologists 4877:English biophysicists 4668:Francis William Aston 3672:Dickinson W. Richards 3356:Julius Wagner-Jauregg 2823:Horace Tabberer Brown 2727:Alfred Russel Wallace 1519:Hill, A. V. (1926b). 1500:Hill, A. V. (1926a). 1360: 1258:John Herbert Humphrey 1208:maximal oxygen uptake 1145: 1125: 1070: 971: 951: 920: 900: 874: 854: 824: 798: 764: 744: 715: 695: 666: 646: 626: 603: 394:Ministry of Munitions 331:, he was educated at 307:. He shared the 1922 276:Archibald Vivian Hill 36:Archibald Vivian Hill 4095:Rita Levi-Montalcini 3456:Albert Szent-Györgyi 3031:Edgar Douglas Adrian 2671:Josiah Willard Gibbs 2578:Cambridge University 2253:10.1123/kr.2021-0064 2220:(7): 449–451. 1938. 2192:search.amphilsoc.org 2188:"APS Member History" 1998:Hill, A. V. (1965). 1805:Hill, A. V. (1909). 1602:Hill, A. V. (1965). 1593:Hill, A. V. (1960). 1577:Hill, A. V. (1931). 1542:Hill, A. V. (1928). 1534:Hill, A.V. (1927b). 1527:Hill, A.V. (1927a). 1408:Publications by Hill 1218:In 1913, he married 1134: 1082: 990: 960: 929: 909: 883: 863: 837: 807: 781: 753: 724: 704: 675: 655: 635: 615: 542: 483:Cambridge University 360:John Newport Langley 200:Cambridge University 4952:Royal Medal winners 4755:Otto Fritz Meyerhof 4487:William C. Campbell 4398:Elizabeth Blackburn 4235:Robert F. Furchgott 4227:Stanley B. Prusiner 4219:Rolf M. Zinkernagel 4119:George H. Hitchings 4083:Joseph L. Goldstein 4043:Bengt I. Samuelsson 3872:Earl Sutherland Jr. 2903:William Henry Bragg 2735:George William Hill 2719:Albert A. Michelson 1872:10.1021/ja02242a004 1585:Hill, A.V. (1932). 1560:1928RSPSB.103..117H 1485:1926SciAm.134..224H 1472:Scientific American 1386:John Maynard Keynes 1346:British Association 1228:John Maynard Keynes 1224:John Neville Keynes 1204:Otto Fritz Meyerhof 1150:. This is called a 1123:{\displaystyle \ln} 898:{\displaystyle h=1} 822:{\displaystyle h=1} 749:, and the exponent 581: 474:William Henry Bragg 343:in the mathematics 335:and graduated from 305:operations research 4547:William Kaelin Jr. 4543:Peter J. Ratcliffe 4286:Leland H. Hartwell 4175:Richard J. Roberts 4055:Barbara McClintock 3999:Godfrey Hounsfield 3900:Nikolaas Tinbergen 3832:Marshall Nirenberg 3800:Charles B. Huggins 3589:Tadeusz Reichstein 3577:António Egas Moniz 3372:Christiaan Eijkman 2975:Thomas Hunt Morgan 2959:Henry Hallett Dale 2935:John Scott Haldane 2919:George Ellery Hale 2544:on Nobelprize.org 2324:. 31 December 2010 2268:The Academy Papers 2241:Kinesiology Review 1363: 1169:muscle contraction 1140: 1120: 1065: 966: 946: 915: 895: 869: 849: 819: 793: 759: 739: 710: 690: 661: 641: 621: 598: 567: 347:before turning to 4957:Sports scientists 4834: 4833: 4692:Jacinto Benavente 4616: 4615: 4581:Ardem Patapoutian 4430:Ralph M. Steinman 4426:Jules A. Hoffmann 4414:Robert G. Edwards 4382:Harald zur Hausen 4306:H. Robert Horvitz 4207:Eric F. Wieschaus 4143:E. Donnall Thomas 4127:J. Michael Bishop 4115:Gertrude B. Elion 3987:Hamilton O. Smith 3912:Christian de Duve 3648:Frederick Robbins 3525:Alexander Fleming 3464:Corneille Heymans 3310:Frederick Banting 3215:Allvar Gullstrand 3073: 3072: 2839:Ernest Rutherford 2658:Copley Medallists 2608: 2607: 2595:Succeeded by 2542:Archibald V. Hill 2523:978-1-61145-709-4 2503:Lusk, G. (1925). 2473:Jacoby, Charlie. 2350:. 16 January 2020 2286:www.nasonline.org 2168:. 9 February 2023 2033:978-3-527-33074-4 1790:(Subscription or 1606:. London: Arnold. 1511:978-0-8493-5494-6 1502:Muscular Activity 1382:Nicholas Humphrey 1344:President of the 1265:Honors and awards 1245:David Keynes Hill 1181:Hermann Helmholtz 1158:Muscle physiology 1143:{\displaystyle h} 969:{\displaystyle h} 918:{\displaystyle h} 872:{\displaystyle h} 762:{\displaystyle h} 713:{\displaystyle a} 664:{\displaystyle V} 644:{\displaystyle a} 624:{\displaystyle v} 596: 438:William Beveridge 333:Blundell's School 273: 272: 266:Nicholas Humphrey 258:David Keynes Hill 214:Academic advisors 176:Scientific career 61:26 September 1886 16:(Redirected from 4994: 4962:UK MPs 1935–1945 4772: 4643: 4636: 4629: 4620: 4619: 4609: 4594: 4583: 4568: 4562:Michael Houghton 4549: 4539:Gregg L. Semenza 4533: 4527:James P. Allison 4521: 4519:Michael W. Young 4505: 4503:Yoshinori Ohsumi 4497: 4481: 4465: 4463:Thomas C. Südhof 4449: 4437: 4416: 4408: 4402:Carol W. Greider 4392: 4376: 4360: 4348: 4336: 4324: 4312: 4296: 4269: 4253: 4245: 4229: 4221: 4215:Peter C. Doherty 4209: 4193: 4187:Alfred G. Gilman 4181: 4169: 4157: 4145: 4133: 4131:Harold E. Varmus 4121: 4105: 4097: 4085: 4073: 4057: 4049: 4033: 4017: 4007:Baruj Benacerraf 4001: 3989: 3973: 3957: 3934: 3918: 3902: 3886: 3874: 3866: 3850: 3834: 3824:Robert W. Holley 3818: 3802: 3790: 3774: 3762: 3746: 3730: 3728:Georg von Békésy 3722: 3710: 3698: 3696:Joshua Lederberg 3682: 3674: 3668:Werner Forssmann 3658: 3650: 3634: 3622: 3614: 3595: 3579: 3567: 3559: 3557:Bernardo Houssay 3543: 3535: 3519: 3507: 3495: 3494: 3488: 3487: 3481: 3480: 3474: 3466: 3458: 3450: 3438: 3430: 3414: 3406: 3394: 3386: 3384:Karl Landsteiner 3378: 3366: 3358: 3350: 3348:Johannes Fibiger 3331: 3330: 3324: 3322:Willem Einthoven 3316: 3304: 3292: 3291: 3285: 3277: 3269: 3268: 3262: 3261: 3255: 3254: 3248: 3247: 3241: 3233: 3225: 3217: 3209: 3201: 3193: 3187:Élie Metchnikoff 3181: 3179:Alphonse Laveran 3173: 3161: 3153: 3145: 3137: 3129: 3100: 3093: 3086: 3077: 3076: 3066: 3058: 3055:George de Hevesy 3050: 3042: 3034: 3026: 3018: 3010: 3002: 2994: 2986: 2978: 2970: 2962: 2954: 2946: 2938: 2930: 2922: 2914: 2906: 2898: 2890: 2882: 2874: 2866: 2858: 2850: 2842: 2834: 2826: 2818: 2810: 2802: 2794: 2786: 2778: 2770: 2762: 2754: 2746: 2738: 2730: 2722: 2714: 2711:Élie Metchnikoff 2706: 2703:Dmitri Mendeleev 2698: 2690: 2682: 2674: 2651: 2644: 2637: 2628: 2627: 2570:Sir John Withers 2563:Preceded by 2553: 2552: 2549: 2538: 2527: 2508: 2490: 2489: 2487: 2485: 2470: 2464: 2463: 2461: 2459: 2449: 2443: 2442: 2440: 2438: 2423: 2417: 2416: 2414: 2412: 2403: 2395: 2386: 2385: 2383: 2381: 2366: 2360: 2359: 2357: 2355: 2340: 2334: 2333: 2331: 2329: 2314: 2308: 2303: 2297: 2296: 2294: 2292: 2282:"Archibald Hill" 2278: 2272: 2271: 2263: 2257: 2256: 2236: 2230: 2229: 2209: 2203: 2202: 2200: 2198: 2184: 2178: 2177: 2175: 2173: 2158: 2152: 2151: 2133: 2109: 2103: 2102: 2058: 2052: 2051: 2044: 2038: 2037: 2019: 2013: 2010: 2004: 2003: 1995: 1989: 1988: 1980: 1974: 1971: 1965: 1964: 1956: 1950: 1949: 1941: 1935: 1934: 1933: 1927: 1893: 1885: 1876: 1875: 1866:(9): 1361–1402. 1851: 1845: 1844: 1834: 1802: 1796: 1795: 1787: 1769: 1763: 1762: 1744: 1735:(5): 1567–1582. 1720: 1714: 1713: 1687: 1658: 1649: 1648: 1647: 1641: 1639: 1637: 1622: 1607: 1598: 1582: 1573: 1571: 1554:(723): 117–137. 1536:Living Machinery 1524: 1515: 1496: 1458: 1456: 1447:(679): 438–475. 1431: 1367:English Heritage 1316:Associate Fellow 1230:and the surgeon 1196:William Stirling 1149: 1147: 1146: 1141: 1129: 1127: 1126: 1121: 1101: 1074: 1072: 1071: 1066: 1064: 1063: 1009: 975: 973: 972: 967: 955: 953: 952: 947: 945: 944: 943: 942: 924: 922: 921: 916: 904: 902: 901: 896: 878: 876: 875: 870: 858: 856: 855: 850: 828: 826: 825: 820: 802: 800: 799: 794: 768: 766: 765: 760: 748: 746: 745: 740: 719: 717: 716: 711: 700:is the value of 699: 697: 696: 691: 689: 688: 687: 670: 668: 667: 662: 650: 648: 647: 642: 630: 628: 627: 622: 607: 605: 604: 599: 597: 595: 594: 593: 580: 575: 565: 564: 555: 462:Patrick Blackett 450:Second World War 295:, was a British 290: 224:Notable students 170: 79: 60: 58: 46: 32: 31: 21: 5002: 5001: 4997: 4996: 4995: 4993: 4992: 4991: 4837: 4836: 4835: 4830: 4768: 4760: 4751:(Great Britain) 4735: 4716: 4711:Fridtjof Nansen 4697: 4673: 4654: 4647: 4617: 4612: 4597: 4586: 4571: 4566:Charles M. Rice 4558:Harvey J. Alter 4552: 4536: 4524: 4515:Michael Rosbash 4511:Jeffrey C. Hall 4508: 4500: 4484: 4475:May-Britt Moser 4468: 4452: 4447:Shinya Yamanaka 4440: 4419: 4411: 4406:Jack W. Szostak 4395: 4379: 4374:Oliver Smithies 4363: 4351: 4339: 4327: 4322:Peter Mansfield 4315: 4310:John E. Sulston 4299: 4283: 4272: 4256: 4248: 4232: 4224: 4212: 4199:Edward B. Lewis 4196: 4184: 4172: 4160: 4148: 4136: 4124: 4108: 4103:Susumu Tonegawa 4100: 4088: 4076: 4060: 4052: 4036: 4020: 4004: 3992: 3976: 3963:Roger Guillemin 3960: 3955:Daniel Gajdusek 3951:Baruch Blumberg 3948: 3937: 3928:Renato Dulbecco 3924:David Baltimore 3921: 3905: 3892:Karl von Frisch 3889: 3877: 3869: 3853: 3837: 3821: 3812:Haldan Hartline 3805: 3793: 3777: 3765: 3749: 3744:Maurice Wilkins 3733: 3725: 3713: 3708:Arthur Kornberg 3701: 3685: 3677: 3661: 3653: 3637: 3625: 3617: 3609: 3598: 3582: 3570: 3562: 3546: 3538: 3522: 3513:Joseph Erlanger 3510: 3498: 3492: 3491: 3485: 3484: 3478: 3477: 3469: 3461: 3453: 3441: 3433: 3417: 3409: 3397: 3389: 3381: 3369: 3364:Charles Nicolle 3361: 3353: 3345: 3334: 3328: 3327: 3319: 3307: 3295: 3289: 3288: 3280: 3272: 3266: 3265: 3259: 3258: 3252: 3251: 3245: 3244: 3236: 3228: 3220: 3212: 3207:Albrecht Kossel 3204: 3196: 3184: 3176: 3164: 3156: 3148: 3140: 3132: 3124: 3113: 3104: 3074: 3069: 3061: 3053: 3045: 3037: 3029: 3021: 3013: 3007:Joseph Barcroft 3005: 2999:Robert Robinson 2997: 2989: 2981: 2973: 2965: 2957: 2949: 2941: 2933: 2925: 2917: 2911:Arthur Schuster 2909: 2901: 2893: 2885: 2877: 2869: 2863:Albert Einstein 2861: 2853: 2845: 2837: 2829: 2821: 2815:William Bayliss 2813: 2807:Hendrik Lorentz 2805: 2797: 2789: 2781: 2773: 2765: 2757: 2749: 2741: 2733: 2725: 2717: 2709: 2701: 2695:William Crookes 2693: 2685: 2677: 2669: 2661: 2655: 2613: 2604: 2600: 2586: 2581: 2572: 2568: 2524: 2499: 2494: 2493: 2483: 2481: 2471: 2467: 2457: 2455: 2451: 2450: 2446: 2436: 2434: 2424: 2420: 2410: 2408: 2396: 2389: 2379: 2377: 2367: 2363: 2353: 2351: 2342: 2341: 2337: 2327: 2325: 2316: 2315: 2311: 2304: 2300: 2290: 2288: 2280: 2279: 2275: 2264: 2260: 2237: 2233: 2211: 2210: 2206: 2196: 2194: 2186: 2185: 2181: 2171: 2169: 2160: 2159: 2155: 2110: 2106: 2059: 2055: 2046: 2045: 2041: 2034: 2020: 2016: 2011: 2007: 1996: 1992: 1981: 1977: 1972: 1968: 1957: 1953: 1942: 1938: 1928: 1886: 1879: 1852: 1848: 1803: 1799: 1789: 1771: 1770: 1766: 1721: 1717: 1659: 1652: 1642: 1635: 1633: 1623: 1619: 1614: 1512: 1422:(63): 135–142. 1410: 1394:Stephen Hawking 1378:Highgate School 1355: 1270:Officer of the 1267: 1232:Geoffrey Keynes 1216: 1160: 1135: 1132: 1131: 1097: 1083: 1080: 1079: 1059: 1055: 1005: 991: 988: 987: 961: 958: 957: 938: 937: 933: 932: 930: 927: 926: 910: 907: 906: 884: 881: 880: 864: 861: 860: 838: 835: 834: 808: 805: 804: 782: 779: 778: 754: 751: 750: 725: 722: 721: 705: 702: 701: 683: 679: 678: 676: 673: 672: 656: 653: 652: 636: 633: 632: 616: 613: 612: 589: 585: 576: 571: 566: 560: 556: 554: 543: 540: 539: 519: 442:Lord Rutherford 430:Ernest Starling 411:Douglas Hartree 407:Ralph H. Fowler 379:First World War 325: 313:mechanical work 278: 269: 234: 232:Ralph H. Fowler 230: 206: 202: 165: 160: 155: 150: 129: 124: 120: 114:Mechanical work 87: 81: 77: 68: 62: 56: 54: 37: 28: 23: 22: 15: 12: 11: 5: 5000: 4990: 4989: 4984: 4979: 4974: 4969: 4964: 4959: 4954: 4949: 4944: 4939: 4934: 4929: 4924: 4919: 4914: 4909: 4904: 4899: 4894: 4889: 4884: 4879: 4874: 4869: 4864: 4859: 4854: 4849: 4832: 4831: 4829: 4828: 4823: 4818: 4813: 4808: 4803: 4798: 4793: 4788: 4783: 4778: 4773: 4765: 4762: 4761: 4759: 4758: 4752: 4749:Archibald Hill 4745: 4743: 4737: 4736: 4734: 4733: 4726: 4724: 4718: 4717: 4715: 4714: 4707: 4705: 4699: 4698: 4696: 4695: 4688: 4686: 4675: 4674: 4672: 4671: 4664: 4662: 4656: 4655: 4646: 4645: 4638: 4631: 4623: 4614: 4613: 4611: 4610: 4603:Katalin Karikó 4595: 4584: 4569: 4550: 4534: 4522: 4506: 4498: 4482: 4466: 4459:Randy Schekman 4450: 4438: 4417: 4409: 4393: 4386:Luc Montagnier 4377: 4366:Mario Capecchi 4361: 4349: 4342:Barry Marshall 4337: 4325: 4318:Paul Lauterbur 4313: 4302:Sydney Brenner 4297: 4280: 4278: 4274: 4273: 4271: 4270: 4263:Paul Greengard 4259:Arvid Carlsson 4254: 4246: 4230: 4222: 4210: 4194: 4191:Martin Rodbell 4182: 4170: 4167:Edwin G. Krebs 4163:Edmond Fischer 4158: 4146: 4134: 4122: 4111:James W. Black 4106: 4098: 4086: 4074: 4071:César Milstein 4067:Georges Köhler 4058: 4050: 4039:Sune Bergström 4034: 4031:Torsten Wiesel 4027:David H. Hubel 4018: 4002: 3990: 3983:Daniel Nathans 3974: 3967:Andrew Schally 3958: 3945: 3943: 3939: 3938: 3936: 3935: 3919: 3903: 3887: 3880:Gerald Edelman 3875: 3867: 3864:Julius Axelrod 3851: 3848:Salvador Luria 3844:Alfred Hershey 3835: 3819: 3803: 3791: 3780:François Jacob 3775: 3763: 3747: 3731: 3723: 3711: 3699: 3683: 3675: 3664:André Cournand 3659: 3651: 3635: 3623: 3620:Selman Waksman 3615: 3606: 3604: 3600: 3599: 3597: 3596: 3585:Edward Kendall 3580: 3568: 3560: 3544: 3541:Hermann Muller 3536: 3520: 3517:Herbert Gasser 3508: 3496: 3489: 3482: 3475: 3472:Gerhard Domagk 3467: 3459: 3451: 3439: 3431: 3428:William Murphy 3420:George Whipple 3415: 3407: 3395: 3387: 3379: 3367: 3359: 3351: 3342: 3340: 3336: 3335: 3333: 3332: 3325: 3317: 3305: 3298:Archibald Hill 3293: 3286: 3278: 3270: 3263: 3256: 3249: 3242: 3234: 3231:Charles Richet 3226: 3218: 3210: 3202: 3194: 3182: 3174: 3162: 3154: 3146: 3138: 3130: 3121: 3119: 3115: 3114: 3103: 3102: 3095: 3088: 3080: 3071: 3070: 3068: 3067: 3063:James Chadwick 3059: 3051: 3047:Archibald Hill 3043: 3035: 3027: 3019: 3011: 3003: 2995: 2987: 2979: 2971: 2963: 2955: 2947: 2939: 2931: 2927:Theobald Smith 2923: 2915: 2907: 2899: 2891: 2883: 2875: 2867: 2859: 2851: 2843: 2835: 2827: 2819: 2811: 2803: 2795: 2787: 2779: 2771: 2763: 2755: 2747: 2743:Francis Galton 2739: 2731: 2723: 2715: 2707: 2699: 2691: 2683: 2675: 2666: 2663: 2662: 2654: 2653: 2646: 2639: 2631: 2625: 2624: 2616:The Papers of 2612: 2611:External links 2609: 2606: 2605: 2596: 2593: 2573: 2564: 2560: 2559: 2551: 2550: 2539: 2528: 2522: 2509: 2498: 2495: 2492: 2491: 2465: 2444: 2418: 2387: 2375:Ham & High 2361: 2335: 2309: 2298: 2273: 2258: 2231: 2204: 2179: 2153: 2124:(5): 591–603. 2104: 2069:(4): 365–400. 2053: 2039: 2032: 2014: 2005: 1990: 1975: 1966: 1951: 1936: 1902:(4): 393–410. 1877: 1846: 1817:(5): 361–373. 1797: 1764: 1715: 1650: 1631:NobelPrize.org 1616: 1615: 1613: 1610: 1609: 1608: 1599: 1590: 1583: 1574: 1539: 1532: 1525: 1516: 1510: 1497: 1479:(4): 224–225. 1466: 1459: 1432: 1409: 1406: 1354: 1351: 1350: 1349: 1342: 1336: 1330: 1323: 1313: 1306: 1299: 1288: 1282: 1277:Fellow of the 1275: 1266: 1263: 1262: 1261: 1254: 1248: 1242: 1215: 1212: 1159: 1156: 1139: 1119: 1116: 1113: 1110: 1107: 1104: 1100: 1096: 1093: 1090: 1087: 1076: 1075: 1062: 1058: 1054: 1051: 1048: 1045: 1042: 1039: 1036: 1033: 1030: 1027: 1024: 1021: 1018: 1015: 1012: 1008: 1004: 1001: 998: 995: 965: 941: 936: 914: 894: 891: 888: 868: 848: 845: 842: 818: 815: 812: 792: 789: 786: 758: 738: 735: 732: 729: 709: 686: 682: 660: 651:of substrate, 640: 620: 609: 608: 592: 588: 584: 579: 574: 570: 563: 559: 553: 550: 547: 518: 515: 419:James Crowther 324: 321: 271: 270: 251: 248: 247: 243: 242: 239: 238: 225: 221: 220: 215: 211: 210: 197: 193: 192: 183: 179: 178: 172: 171: 162:Actonian Prize 148: 144: 143: 138: 134: 133: 111: 110:Known for 107: 106: 101: 97: 96: 93: 89: 88: 82: 80:(aged 90) 74: 70: 69: 63: 52: 48: 47: 39: 38: 35: 26: 9: 6: 4: 3: 2: 4999: 4988: 4985: 4983: 4980: 4978: 4975: 4973: 4970: 4968: 4965: 4963: 4960: 4958: 4955: 4953: 4950: 4948: 4945: 4943: 4940: 4938: 4935: 4933: 4930: 4928: 4925: 4923: 4920: 4918: 4915: 4913: 4910: 4908: 4907:Keynes family 4905: 4903: 4900: 4898: 4895: 4893: 4890: 4888: 4885: 4883: 4880: 4878: 4875: 4873: 4870: 4868: 4865: 4863: 4860: 4858: 4855: 4853: 4850: 4848: 4845: 4844: 4842: 4827: 4824: 4822: 4819: 4817: 4814: 4812: 4809: 4807: 4804: 4802: 4799: 4797: 4794: 4792: 4789: 4787: 4784: 4782: 4779: 4777: 4774: 4771: 4767: 4766: 4763: 4756: 4753: 4750: 4747: 4746: 4744: 4742: 4738: 4731: 4728: 4727: 4725: 4723: 4719: 4712: 4709: 4708: 4706: 4704: 4700: 4693: 4690: 4689: 4687: 4684: 4680: 4676: 4669: 4666: 4665: 4663: 4661: 4657: 4652: 4644: 4639: 4637: 4632: 4630: 4625: 4624: 4621: 4608: 4607:Drew Weissman 4604: 4600: 4596: 4593: 4589: 4585: 4582: 4578: 4574: 4570: 4567: 4563: 4559: 4555: 4551: 4548: 4544: 4540: 4535: 4532: 4528: 4523: 4520: 4516: 4512: 4507: 4504: 4499: 4496: 4492: 4491:Satoshi Ōmura 4488: 4483: 4480: 4476: 4472: 4467: 4464: 4460: 4456: 4455:James Rothman 4451: 4448: 4444: 4439: 4435: 4431: 4427: 4423: 4422:Bruce Beutler 4418: 4415: 4410: 4407: 4403: 4399: 4394: 4391: 4387: 4383: 4378: 4375: 4371: 4367: 4362: 4359: 4355: 4350: 4347: 4343: 4338: 4335: 4334:Linda B. Buck 4331: 4326: 4323: 4319: 4314: 4311: 4307: 4303: 4298: 4295: 4291: 4287: 4282: 4281: 4279: 4275: 4268: 4264: 4260: 4255: 4252: 4251:Günter Blobel 4247: 4244: 4240: 4239:Louis Ignarro 4236: 4231: 4228: 4223: 4220: 4216: 4211: 4208: 4204: 4200: 4195: 4192: 4188: 4183: 4180: 4179:Phillip Sharp 4176: 4171: 4168: 4164: 4159: 4156: 4152: 4147: 4144: 4140: 4139:Joseph Murray 4135: 4132: 4128: 4123: 4120: 4116: 4112: 4107: 4104: 4099: 4096: 4092: 4091:Stanley Cohen 4087: 4084: 4080: 4079:Michael Brown 4075: 4072: 4068: 4064: 4059: 4056: 4051: 4048: 4044: 4040: 4035: 4032: 4028: 4024: 4019: 4016: 4012: 4008: 4003: 4000: 3996: 3995:Allan Cormack 3991: 3988: 3984: 3980: 3975: 3972: 3971:Rosalyn Yalow 3968: 3964: 3959: 3956: 3952: 3947: 3946: 3944: 3940: 3933: 3929: 3925: 3920: 3917: 3916:George Palade 3913: 3909: 3908:Albert Claude 3904: 3901: 3897: 3896:Konrad Lorenz 3893: 3888: 3885: 3884:Rodney Porter 3881: 3876: 3873: 3868: 3865: 3861: 3860:Ulf von Euler 3857: 3852: 3849: 3845: 3841: 3836: 3833: 3829: 3825: 3820: 3817: 3813: 3809: 3808:Ragnar Granit 3804: 3801: 3797: 3792: 3789: 3788:Jacques Monod 3785: 3781: 3776: 3773: 3769: 3764: 3761: 3760:Andrew Huxley 3757: 3753: 3748: 3745: 3741: 3737: 3736:Francis Crick 3732: 3729: 3724: 3721: 3720:Peter Medawar 3717: 3712: 3709: 3705: 3700: 3697: 3693: 3689: 3688:George Beadle 3684: 3681: 3676: 3673: 3669: 3665: 3660: 3657: 3656:Hugo Theorell 3652: 3649: 3645: 3644:Thomas Weller 3641: 3636: 3633: 3632:Fritz Lipmann 3629: 3624: 3621: 3616: 3613: 3608: 3607: 3605: 3601: 3594: 3590: 3586: 3581: 3578: 3574: 3569: 3566: 3561: 3558: 3554: 3550: 3545: 3542: 3537: 3534: 3533:Howard Florey 3530: 3526: 3521: 3518: 3514: 3509: 3506: 3502: 3497: 3490: 3483: 3476: 3473: 3468: 3465: 3460: 3457: 3452: 3449: 3445: 3440: 3437: 3432: 3429: 3425: 3421: 3416: 3413: 3412:Thomas Morgan 3408: 3405: 3401: 3396: 3393: 3388: 3385: 3380: 3377: 3373: 3368: 3365: 3360: 3357: 3352: 3349: 3344: 3343: 3341: 3337: 3326: 3323: 3318: 3315: 3311: 3306: 3303: 3302:Otto Meyerhof 3299: 3294: 3287: 3284: 3279: 3276: 3271: 3264: 3257: 3250: 3243: 3240: 3239:Róbert Bárány 3235: 3232: 3227: 3224: 3223:Alexis Carrel 3219: 3216: 3211: 3208: 3203: 3200: 3195: 3192: 3188: 3183: 3180: 3175: 3172: 3168: 3167:Camillo Golgi 3163: 3160: 3155: 3152: 3147: 3144: 3139: 3136: 3131: 3128: 3123: 3122: 3120: 3116: 3112: 3108: 3101: 3096: 3094: 3089: 3087: 3082: 3081: 3078: 3064: 3060: 3056: 3052: 3048: 3044: 3040: 3036: 3032: 3028: 3024: 3020: 3016: 3012: 3008: 3004: 3000: 2996: 2992: 2988: 2984: 2983:Paul Langevin 2980: 2976: 2972: 2968: 2964: 2960: 2956: 2952: 2948: 2944: 2940: 2936: 2932: 2928: 2924: 2920: 2916: 2912: 2908: 2904: 2900: 2896: 2892: 2888: 2884: 2880: 2876: 2872: 2868: 2864: 2860: 2856: 2852: 2848: 2844: 2840: 2836: 2832: 2831:Joseph Larmor 2828: 2824: 2820: 2816: 2812: 2808: 2804: 2800: 2796: 2792: 2788: 2784: 2780: 2776: 2775:J. J. Thomson 2772: 2768: 2767:Ray Lankester 2764: 2760: 2756: 2752: 2751:George Darwin 2748: 2744: 2740: 2736: 2732: 2728: 2724: 2720: 2716: 2712: 2708: 2704: 2700: 2696: 2692: 2688: 2684: 2680: 2679:Joseph Lister 2676: 2672: 2668: 2667: 2664: 2659: 2652: 2647: 2645: 2640: 2638: 2633: 2632: 2629: 2623: 2619: 2615: 2614: 2603: 2602:Wilson Harris 2599: 2592: 2591: 2585: 2580: 2579: 2571: 2567: 2561: 2558: 2554: 2548: 2543: 2540: 2536: 2535: 2529: 2525: 2519: 2515: 2510: 2506: 2501: 2500: 2480: 2476: 2469: 2454: 2448: 2433: 2429: 2422: 2407: 2402: 2394: 2392: 2376: 2372: 2365: 2349: 2345: 2339: 2323: 2319: 2313: 2307: 2302: 2287: 2283: 2277: 2269: 2262: 2254: 2250: 2246: 2242: 2235: 2227: 2223: 2219: 2215: 2208: 2193: 2189: 2183: 2167: 2163: 2157: 2149: 2145: 2141: 2137: 2132: 2127: 2123: 2119: 2115: 2108: 2100: 2096: 2092: 2088: 2084: 2080: 2076: 2072: 2068: 2064: 2057: 2049: 2043: 2035: 2029: 2025: 2018: 2009: 2001: 1994: 1986: 1979: 1970: 1962: 1955: 1947: 1940: 1932: 1925: 1921: 1917: 1913: 1909: 1905: 1901: 1897: 1892: 1884: 1882: 1873: 1869: 1865: 1861: 1857: 1850: 1842: 1838: 1833: 1828: 1824: 1820: 1816: 1812: 1808: 1801: 1793: 1785: 1781: 1777: 1776: 1768: 1760: 1756: 1752: 1748: 1743: 1738: 1734: 1730: 1726: 1719: 1711: 1707: 1703: 1699: 1695: 1691: 1686: 1681: 1677: 1673: 1672: 1667: 1663: 1662:Katz, Bernard 1657: 1655: 1646: 1632: 1628: 1621: 1617: 1605: 1600: 1596: 1591: 1588: 1584: 1580: 1575: 1570: 1565: 1561: 1557: 1553: 1549: 1545: 1540: 1537: 1533: 1530: 1526: 1522: 1517: 1513: 1507: 1503: 1498: 1494: 1490: 1486: 1482: 1478: 1474: 1473: 1467: 1464: 1460: 1455: 1450: 1446: 1442: 1438: 1433: 1429: 1425: 1421: 1417: 1412: 1411: 1405: 1402: 1399: 1395: 1391: 1390:Sigmund Freud 1387: 1383: 1379: 1375: 1371: 1368: 1359: 1347: 1343: 1340: 1337: 1334: 1331: 1328: 1324: 1321: 1317: 1314: 1311: 1307: 1304: 1300: 1297: 1293: 1289: 1286: 1283: 1280: 1279:Royal Society 1276: 1273: 1269: 1268: 1259: 1255: 1252: 1249: 1246: 1243: 1240: 1237: 1236: 1235: 1233: 1229: 1225: 1221: 1214:Personal life 1211: 1209: 1205: 1201: 1197: 1193: 1188: 1186: 1182: 1177: 1175: 1170: 1166: 1155: 1153: 1137: 1111: 1108: 1105: 1098: 1094: 1088: 1085: 1060: 1056: 1052: 1049: 1046: 1043: 1040: 1037: 1034: 1031: 1028: 1019: 1016: 1013: 1006: 1002: 996: 993: 986: 985: 984: 981: 979: 963: 956:rather than 934: 912: 892: 889: 886: 866: 846: 843: 840: 832: 816: 813: 810: 790: 787: 784: 776: 772: 756: 736: 733: 730: 727: 707: 684: 680: 658: 638: 618: 590: 586: 582: 577: 572: 568: 561: 557: 551: 548: 545: 538: 537: 536: 534: 530: 526: 525: 524:Hill equation 514: 511: 507: 503: 498: 494: 492: 487: 484: 480: 475: 471: 467: 463: 458: 455: 451: 447: 443: 439: 435: 431: 426: 424: 420: 416: 412: 408: 404: 400: 395: 391: 390:Horace Darwin 386: 384: 380: 376: 371: 369: 365: 361: 357: 352: 350: 346: 342: 338: 334: 330: 320: 318: 314: 310: 306: 302: 298: 294: 289: 285: 281: 277: 267: 263: 259: 255: 249: 244: 240: 237: 233: 229: 226: 222: 219: 216: 212: 209: 205: 201: 198: 194: 191: 187: 184: 180: 177: 173: 168: 163: 158: 153: 149: 145: 142: 139: 135: 132: 128: 123: 119: 115: 112: 108: 105: 102: 98: 94: 90: 85: 75: 71: 66: 53: 49: 45: 40: 33: 30: 19: 4800: 4748: 4592:Svante Pääbo 4577:David Julius 4531:Tasuku Honjo 4479:Edvard Moser 4471:John O'Keefe 4434:posthumously 4433: 4370:Martin Evans 4346:Robin Warren 4330:Richard Axel 4277:2001–present 4155:Bert Sakmann 4023:Roger Sperry 4015:George Snell 4011:Jean Dausset 3979:Werner Arber 3932:Howard Temin 3856:Bernard Katz 3840:Max Delbrück 3796:Francis Rous 3772:Feodor Lynen 3768:Konrad Bloch 3756:Alan Hodgkin 3740:James Watson 3716:Frank Burnet 3704:Severo Ochoa 3692:Edward Tatum 3680:Daniel Bovet 3593:Philip Hench 3505:Edward Doisy 3436:Hans Spemann 3424:George Minot 3404:Edgar Adrian 3392:Otto Warburg 3314:John Macleod 3297: 3283:August Krogh 3275:Jules Bordet 3191:Paul Ehrlich 3143:Niels Finsen 3127:Emil Behring 3046: 3023:Oswald Avery 2991:Thomas Lewis 2951:Arthur Evans 2687:Eduard Suess 2588: 2575: 2533: 2513: 2504: 2482:. 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Hardy 2847:Horace Lamb 2791:James Dewar 2783:Ivan Pavlov 2759:Felix Klein 2660:(1901–1950) 2247:(1): 6–25. 1370:Blue plaque 1353:Blue plaque 1320:Kinesiology 1165:Magnus Blix 720:that gives 533:haemoglobin 479:independent 228:Te-Pei Feng 157:Royal Medal 92:Nationality 76:3 June 1977 4841:Categories 4730:Niels Bohr 4679:Literature 4294:Paul Nurse 3628:Hans Krebs 3553:Gerty Cori 3501:Henrik Dam 3448:Otto Loewi 3444:Henry Dale 2967:Niels Bohr 2895:Max Planck 2618:A. V. Hill 1794:required.) 1678:: 71–149. 1612:References 1398:Ralph Kohn 1239:Polly Hill 1185:biophysics 349:physiology 301:biophysics 293:A. V. Hill 254:Polly Hill 190:biophysics 186:Physiology 127:biophysics 57:1886-09-26 18:A. V. Hill 4757:(Germany) 4732:(Denmark) 4670:(England) 4660:Chemistry 4653:laureates 4495:Tu Youyou 4047:John Vane 3942:1976–2000 3603:1951–1975 3549:Carl Cori 3339:1926–1950 3118:1901–1925 3107:Laureates 2458:29 August 2437:29 August 2411:8 October 2380:29 August 2354:29 August 2328:29 August 2140:0195-9131 2083:0264-0414 1916:0035-9149 1636:29 August 1152:Hill plot 1109:− 1089:⁡ 1053:⁡ 1044:− 1038:⁡ 1017:− 997:⁡ 399:equations 392:from the 339:as third 323:Biography 125:Founding 100:Education 86:, England 84:Cambridge 67:, England 4713:(Norway) 4290:Tim Hunt 2620:held at 2099:33768722 2091:18228167 1924:22332470 1841:16992989 1759:14704104 1751:12381740 1710:46444782 1702:11615743 1664:(1978). 1428:20263725 1374:Highgate 464:and Sir 364:receptor 341:wrangler 327:Born in 4722:Physics 4694:(Spain) 3109:of the 2582:1940 – 2497:Sources 2148:9140894 1832:1533665 1556:Bibcode 1481:Bibcode 978:protein 425:(OBE). 329:Bristol 317:muscles 118:muscles 95:British 65:Bristol 4537:2019: 4525:2018: 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1780:doi 1737:doi 1680:doi 1564:doi 1552:103 1489:doi 1477:224 1449:doi 1401:FRS 1294:on 1176:. 1061:0.5 925:or 734:0.5 685:0.5 573:0.5 531:to 315:in 288:FRS 284:OBE 116:in 4843:: 4605:/ 4601:: 4590:: 4579:/ 4575:: 4564:/ 4560:/ 4556:: 4545:/ 4541:/ 4529:/ 4517:/ 4513:/ 4493:/ 4489:/ 4477:/ 4473:/ 4461:/ 4457:/ 4445:/ 4428:/ 4424:/ 4404:/ 4400:/ 4388:/ 4384:/ 4372:/ 4368:/ 4356:/ 4344:/ 4332:/ 4320:/ 4308:/ 4304:/ 4292:/ 4288:/ 4265:/ 4261:/ 4241:/ 4237:/ 4217:/ 4205:/ 4201:/ 4189:/ 4177:/ 4165:/ 4153:/ 4141:/ 4129:/ 4117:/ 4113:/ 4093:/ 4081:/ 4069:/ 4065:/ 4045:/ 4041:/ 4029:/ 4025:/ 4013:/ 4009:/ 3997:/ 3985:/ 3981:/ 3969:/ 3965:/ 3953:/ 3930:/ 3926:/ 3914:/ 3910:/ 3898:/ 3894:/ 3882:/ 3862:/ 3858:/ 3846:/ 3842:/ 3830:/ 3826:/ 3814:/ 3810:/ 3798:/ 3786:/ 3782:/ 3770:/ 3758:/ 3754:/ 3742:/ 3738:/ 3718:/ 3706:/ 3694:/ 3690:/ 3670:/ 3666:/ 3646:/ 3642:/ 3630:/ 3591:/ 3587:/ 3575:/ 3555:/ 3551:/ 3531:/ 3527:/ 3515:/ 3503:/ 3446:/ 3426:/ 3422:/ 3402:/ 3374:/ 3312:/ 3300:/ 3189:/ 3169:/ 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1099:/ 1095:v 1092:[ 1057:K 1047:h 1041:a 1032:h 1029:= 1026:] 1023:) 1020:v 1014:V 1011:( 1007:/ 1003:v 1000:[ 964:h 940:H 935:n 913:h 893:1 890:= 887:h 867:h 847:1 841:h 817:1 814:= 811:h 791:1 785:h 757:h 737:V 731:= 728:v 708:a 681:K 659:V 639:a 619:v 591:h 587:a 583:+ 578:h 569:K 562:h 558:a 552:V 549:= 546:v 268:. 59:) 55:( 20:)

Index

A. V. Hill

Bristol
Cambridge
Trinity College, Cambridge
Mechanical work
muscles
Muscle contraction model
biophysics
Hill equation (biochemistry)
Margaret Neville Keynes
Nobel Prize in Physiology or Medicine
Royal Medal
Actonian Prize
Copley Medal
Physiology
biophysics
Cambridge University
University of Manchester
University College, London
Walter Morley Fletcher
Te-Pei Feng
Ralph H. Fowler
Bernard Katz
Polly Hill
David Keynes Hill
Maurice Hill
Nicholas Humphrey
CH
OBE

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