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Patagotitan

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1211: 1111: 944: 1456: 568: 817: 5950: 5961: 877:, which he estimated at over 35 m (115 ft) in length and 65–75 tonnes (64–74 long tons; 72–83 short tons) in weight. In 2020, Campione and Evans yielded a body mass estimate of approximately 55.7 tonnes (54.8 long tons; 61.4 short tons). In 2020, Otero and Carballido published a lower mean weight estimate of 57 tonnes (56 long tons; 63 short tons) using a revised scaling equation, with a margin of error of 42.5–71.4 tonnes (41.8–70.3 long tons; 46.8–78.7 short tons); this was close to the 2017 volumetric estimate. 5968: 7362: 8094: 144: 9329: 9323: 7367: 9404: 6824: 3147: 9318: 6816: 9312: 8086: 759: 6836: 5956: 6830: 8102: 118: 9392: 1640:; the two were united by the presence of long neural spines in the front back vertebrae. These two were placed within the Lognkosauria, based on the SDLs, the tall transverse processes, and small laminae that connect the neural spine to the prezygapophyses (i.e., spinoprezygapophyseal laminae or SPPLs) in the tail vertebrae of 655:. Based on overlapping elements, these include at least three individuals from FLV1, one from FLV2, and one from FLV3 other than the holotype. MPEF-PV 3399 (from FLV1) is a second skeleton consisting of six neck vertebrae, four back vertebrae, one front tail vertebra, sixteen rear tail vertebrae, ribs, chevrons, the left 1648:
to the exclusion of other titanosaurs, where the SPPLs of the tail vertebrae were much better-developed. Notably, this emergence of this grouping corresponded to a major increase in body size as it contained the largest titanosaurs, with maximum weights increasing from 20 to 60 tonnes (22 to 66 short
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size at 40 m (131 ft) long with a weight of 77 tonnes (85 short tons). In 2017, Carballido and colleagues published a length estimate of 37 m (121 ft), and provided two weight estimates: 69 tonnes (76 short tons) using a scaling equation, and 44.2–77.6 tonnes (43.5–76.4 long tons;
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The collected sauropod fossils were organized into six partial skeletons, which likely belonged to the same species due to uniformity in their morphology and size (with all individuals differing no more than 5% in length). Although the remains all came from the same quarry, the individuals likely did
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between only one pair of vertebrae at the level of the scapular blade, which was likely a weight-bearing adaptation not seen in any other sauropod (where they were either present between all pairs or between none). Several unique features in the limbs were also likely attachment scars for muscles. In
1154:(vertebral bodies). Yet another unique characteristic was that the tops of the neural spines were concave, making them somewhat bifurcated with the tips curling forwards. On the neural spine, longitudinal laminae separated from the prespinal and postspinal laminae, which was previously only seen in 2669:
was among the most phylogenetically stable titanosaurs, with a high recuperation index of 62% (meaning that it is stable enough to not be removed due to instability in 62% of analyses) due to its firm position within the Lognkosauria. They found that the uncertainty regarding whether Lognkosauria
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are displayed in multiple museums. Research Casting International digitally scanned the specimens, which was followed by the creation of foam molds, fiberglass casts, and 3D printing. One mount is exhibited in the Museum of Paleontology Egidio Feruglio. Another is exhibited in the
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not all die at the same time. Within the 3.43-metre (11.3 ft) sediment layer containing the fossils, there are three distinct but closely-spaced levels corresponding to three burial events, which are named FLV1, FLV2, and FLV3. One skeleton from FLV3 was designated as the
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being closest to this group; unlike Carballido and colleagues, however, they recovered the Lognkosauria within the Lithostrotia. In 2019, González Riga and colleagues recovered the same arrangement, and they named the group formed by the Lognkosauria and the Rinconsauria as
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at the sides) were nearly vertical because the prezygapophyses were situated higher up on the vertebrae than the diapophyses. The PRDLs usually instead extended forward from the diapophyses in other sauropods. All of the neural spines in the back were tall and vertical in
2640:, and Lognkosauria being outside Lithostrotia; Federico Agnolin and colleagues found a similar arrangement in 2023. In 2021, Otero and colleagues recovered the same arrangement. Also in 2021, Pablo Gallina and colleagues recovered a similar arrangement, except with 536:(MPEF) in Trelew. One preliminary field expedition to the La Flecha site was carried out in late 2012 by the MPEF, followed by seven further expeditions that discovered new fossils between January 2013 and February 2015. The lead scientists on the excavation were 1085:
had short front neural spines lacking the arrow-shaped ends. Uniquely, in the first two back vertebrae, the prespinal laminae that ran alongside the front of the neural spines bulged outwards near the bottom. Also uniquely, the third back vertebra preserves a
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on the inner surface of the femur was located a third of the way from the top instead of halfway down. Uniquely among sauropods, at the outer edge of the bottom of the femur, there were a series of coarse ridges right above the articulation known as the
1130:, the front articulating surface of the first tail vertebra was flat. Also uniquely, the neural spines of the front tail vertebrae were about four to six times as wide from side to side as they were long front to back, contrasting with the condition in 935:. He attributed this conclusion to a lack of proper measurement techniques. Also, he criticized the 3D model used for the volumetric estimate as having a flat-sided torso, and noted that the neck and tail were reconstructed as being fairly long. 4274:
González Riga, B.J.; Lamanna, M.C.; Otero, A.; Ortiz David, L.D.; Kellner, A.W.A.; Ibiricu, L.M. (2019). "An overview of the appendicular skeletal anatomy of South American titanosaurian sauropods, with definition of a newly recognized clade".
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Agnolin, F.L.; Gonzalez Riga, B.J.; Aranciaga Rolando, A.M.; Rozadilla, S.; Motta, M.J.; Chimento, N.R.; Novas, F.E. (2023). "A new gigant titanosaur (Dinosauria, Sauropoda) from the Upper Cretaceous of Northwestern Patagonia, Argentina".
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Carballido, J.L.; Otero, A.; Mannion, P.D.; Salgado, L.; Moreno, A.P. (2022). "Titanosauria: A critical appraisal of its systematics and the relevance of the South American record". In Otero, A.; Carballido, J.L.; Pol, D. (eds.).
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Carballido, J.L.; Salgado, L.; Pol, D.; Canudo, J.I.; Garrido, A. (2012). "A new basal rebbachisaurid (Sauropoda, Diplodocoidea) from the Early Cretaceous of the Neuquén Basin; evolution and biogeography of the group".
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at the front; the missing fourth back vertebra presumably had a corresponding hypantrum. Hyposphene-hypantrum articulations to stiffen the vertebrae were primitively present among sauropods and lost multiple times, but
2624:, and others depending on the exclusion of some other titanosaurs from the analysis; this exclusion also affected whether Lognkosauria was inside Lithostrotia. In 2020, Carballido and colleagues found again that 679:, the left pubic bone, and the left femur. MPEF-PV 3372 (from FLV1) is a tooth. MPEF-PV 3393 (not associated with a layer) is a rear tail vertebra. MPEF-PV 3395 and MPEV-PV 3396 (both from FLV1) are left 3514: 1293:. Furthermore, in addition to a ridge that ran along the midline of the blade parallel to the bottom, there was a second ridge that ran upwards diagonally to the top of the bone, which was unique to 2674:. Instead of using Lithostrotia, they suggested that Eutitanosauria be divided into Saltasauroidea and Colossosauria. They also refined Colossosauria as being the most inclusive group that includes 1107:
is the first known sauropod where they were retained between only one pair of vertebrae. Because the shoulder blade was located at this position, this may have been a weight-bearing adaptation.
687:, while MPEF-PV 3397 (from FLV2) is a right humerus. MPEF-PV 3375 (FLV3) is a left femur while MPEF-PV 3394 (FLV1) is a right femur. MPEF-PV 3391 and MPEF-PV 3392 (both from FLV1) represent two 1382:) on the front upper portion of the humerus, there was a pair of scars above each other, with the top one being wider than tall and the bottom one being taller than wide; this was where the 1247:. Furthermore, expansion was present at both ends such that the top end of the scapula was roughly at the same level as the bottom end, which was an uncommon characteristic also seen in 1418:. However, the inward-pointing anteromedial process of the ulna was relatively slender, being 3.5 times as long as it was wide, and was also relatively horizontal with an angle of 40°; 4468:
Otero, A.; Carballido, J.L.; Salgado, L.; Ignacio Canudo, J.; Garrido, A.C. (2021). "Report of a giant titanosaur sauropod from the Upper Cretaceous of Neuquén Province, Argentina".
1521:, the bottom of the pubic "boot" was not particularly expanded. On the side of the ischium, there was also a well-developed ridge (seen to a lesser extent in the titanosauriforms 396:). Later research revised the length estimate down to 31 m (102 ft) and weight estimates down to approximately 50–57 tonnes (55–63 short tons), suggesting that 4874:
Leardi, J.M.; Pol, D. (2009). "The first crocodyliform from the Chubut Group (Chubut Province, Argentina) and its phylogenetic position within basal Mesoeucrocodylia".
1669:. Lastly, unlike the results of prior analyses, the analysis of Carballido and colleagues placed Lognkosauria outside the Lithostrotia, as separate groups within the 715: 699: 4774:
Novas, F.E.; de Valais, S.; Vickers-Rich, P.; Rich, T. (2005). "A large Cretaceous theropod from Patagonia, Argentina, and the evolution of carcharodontosaurids".
707: 3203:, was named from the fossilized wood in 2019; these trees were estimated as having been 15.3 metres (50 ft) tall. A new species of the angiosperm wood genus 3199: 719: 1434:. The radius was also fairly expanded, being 36% as broad at the top as it was long. On the inner edge of the radius, a bulging scar for the attachment of the 537: 525: 3522: 3488: 975:
had deep pleurocoels that opened from the sides into the interior of the bone; whether the neural spines of these vertebrae were horizontally expanded as in
1491:, and saltasaurids). The articulation between the pubic bones was clearly demarcated from their articulations with other bones by deflected surfaces, as in 1394:
on the outer side, there was one scar at the midpoint of the crest and one at the bottom; this was where the supracoracoideus, deltoideus clavicularis, and
1329:, in which it was strongly concave). There was a ridge running diagonally from the outer bottom corner of each sternal plate that was oriented similarly to 9493: 4753:"Estudio estratigráfico y sedimentológico de las sedimentitas portadoras de los titanosaurios gigantes del Albiano tardío de Patagonia Central, Argentina" 2658:. In 2023, Agustín Pérez Moreno and colleagues performed an analysis based on that of Gallina and colleagues, and found a similar arrangement except with 843:
Given the size of these bones, which surpass any of the previously known giant animals, the new dinosaur is the largest animal known that walked on Earth.
9508: 711: 648:, and both femora. The skeleton was chosen to be the holotype because it was the best preserved and also the one showing the most distinguishing traits. 4233:
González Riga, B.J.; Mannion, P.D.; Poropat, S.F.; Ortiz David, L.; Coria, J.P. (2018). "Osteology of the Late Cretaceous Argentinean sauropod dinosaur
3240:, meaning that the individuals were preserved where they died. Animals from other localities in the Cerro Castaño Member include the theropod dinosaurs 4594: 3462: 1539:) extending down the ischial tuberosity (a common feature among titanosaurs) down to the blade, with the combination of these features being unique to 873:
at 31 m (102 ft) in length and 50–55 tonnes (49–54 long tons; 55–61 short tons) in weight using volumetric models. This made it smaller than
695:. Another quarry was excavated about 300 metres (980 ft) west, revealing a fourth sediment layer (FLV4) with a similarly-sized sauropod skeleton. 3205: 881:
of five femora and one humerus indicates that the individuals died while they were young adults, with growth having slowed but not ceased entirely.
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In the hip girdle, the contact between the forward-pointing pubis and backward-pointing ischium was about half of the former's length (longer than
5003: 4330:"New information on the Cretaceous sauropod dinosaurs of Zhejiang Province, China: impact on Laurasian titanosauriform phylogeny and biogeography" 915:
has been estimated at 65–96.4 tonnes (71.7–106.3 short tons). In 2019, Paul noted that the articulated length of the back vertebrae was larger in
1297:. The articulating surface of the coracoid with the scapula was 80% as long front-to-back as it was tall, an uncommon characteristic shared with 3788: 2575:
in analyses including the three titanosaurs, with either one being its closest relative. In 2018, Bernardo González Riga and colleagues found
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was a particularly large and robust titanosaur. It can be distinguished from its close relatives by a suite of unique characteristics in its
3690:"Determining the largest known land animal: A critical comparison of differing methods for restoring the volume and mass of extinct animals" 3545: 3571: 4620: 3814: 1160:. At the sides of the front tail vertebrae, the transverse processes (on which the diapophyses were located) were high and thin, like 5052: 1259:
ridge was about 30% as long as the scapula. In addition to a projection at the bottom of the scapula just behind the shoulder joint (
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lived in what was then a forested region. Numerous plant fossils were found at a site about 200 metres (660 ft) away from the
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Historically, the interrelationships of titanosaurs have been highly unstable and varied between different analyses. Despite this,
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was comparable in size with other known giant titanosaurs. However, almost every measurement that could be compared was larger in
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belonged within Lithostrotia largely arose from the definition of Lithostrotia, which was based on the phylogenetically unstable
1267:), which was common among titanosaurs, there was a second one further back at the beginning of the blade, only seen otherwise in 1370:, it was about 70% as long as the femur. A number of bulges for muscle attachment on the humerus were unique characteristics of 9528: 4706:
from the Early Cretaceous of Patagonia (Chubut province, Argentina): the oldest record of angiosperm wood from South America".
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Carballido, J.L.; Scheil, M.; Knötschke, N.; Sander, P.M. (2019). "The appendicular skeleton of the dwarf macronarian sauropod
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Carballido, J.L.; Pol, D.; Otero, A.; Cerda, I.A.; Salgado, L.; Garrido, A.C.; Ramezani, J.; CĂşneo, N.R.; Krause, J.M. (2017).
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had a robust, expanded scapula, with the blade being only 4.15 times as long as it was wide at its narrowest point; except for
1032: 533: 1202:, well-developed spinodiapophyseal laminae (SDLs) ran from the neural spines down the tranverse processes to the diapophyses. 590: 416:
is one of the most-known titanosaurs, and so its interrelationships with other titanosaurs have been relatively consistent in
4217: 1315:. Like other titanosauriforms, the sternal plates were crescent-shaped with the outer edge being slightly concave (more like 606: 598: 449: 441: 5073: 1040: 634: 4752: 9543: 3288: 957: 2686:. The following phylogenetic tree shows their suggested group definitions, including only the most stable titanosaurs. 1176:, where the transverse processes of the front tail vertebrae formed wing-like shapes, they were wider at the bottom in 1048: 965: 776: 763: 642: 1120: 1088: 931:
to have been larger, he reasoned that the rest of its body would have to have been improbably large, or vice versa in
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of a new species, and was catalogued as MPEF-PV 3400. It consists of a partial skeleton lacking the skull, with three
4975: 1096: 1253: 614: 457: 9523: 1261: 1010: 689: 3878: 1148: 681: 673: 665: 481: 473: 465: 5960: 1376: 786: 518: 126: 5967: 5949: 4852: 657: 622: 4925:, a New Eucryptodiran Turtle from the Early Cretaceous of Argentina, and the Relationships of the Meiolaniidae" 1613: 800: 377: 7361: 4647:"A new cupressaceous wood from the Lower Cretaceous of central Patagonia reveals possible clonal growth habit" 4546:
PĂ©rez Moreno, A.; Otero, A.; Carballido, J.L.; Salgado, L.; Calvo, J.O. (2023). "The appendicular skeleton of
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mount. Some of the original fossils, including a femur, were also on display briefly. One is exhibited in the
9518: 9513: 5045: 900:. Wedel also criticised the polygon-based method that Carballido and colleagues used to compare the sizes of 799:(which has been moved to another exhibit). From March 2023 until January 2024, one will be displayed at the 3236:, likely related to sporadic sheet flooding. This flooding was probably too weak to transport the bones of 1579:, a ridge ran along the back of the femur between these two features, but there was no ridge in front like 388:
and land animal overall, with an estimated length of 37 m (121 ft) and an estimated weight of 69
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cautioned against the reported estimates. In blog posts, Wedel noted that based on available measurements
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48.7–85.5 short tons) using volumetric method based on 3D skeletal models. These estimates suggested that
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Gallina, P.A.; Canale, J.I.; Carballido, J.L. (2021). "The Earliest Known Titanosaur Sauropod Dinosaur".
8093: 3181:. The plant fauna was dominated by conifers, with angiosperm wood being relatively rare. A new genus of 556:, and were specifically dated to approximately 101.62 million years in age (corresponding to the latest 143: 9538: 9382: 552:
teeth, were uncovered. The rock layers at the La Flecha site belong to the Cerro Castaño Member of the
490: 4110:"A new Cretaceous terrestrial ecosystem from Gondwana with the description of a new sauropod dinosaur" 1455: 1146:
where they were longer front to back. They were also tall, being 1.5 times taller than the underlying
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Back vertebra, showing the vertical neural spine to the right which bears the prespinal lamina on top
3906: 9533: 9365: 5140: 5038: 1609: 1126:, meaning that the front articulating surface was concave and the rear one was convex. Uniquely in 1016:, ridges of bone crossing the vertebrae, were thin and strongly developed in the back vertebrae of 7366: 1657:
represented independent increases in body size to a much lesser extent. The group of lognkosaurs,
366:, Argentina. The genus contains a single species known from at least six young adult individuals, 9498: 1395: 1383: 747: 553: 355: 9470: 487:
in the hindlimb. Among these was the presence of accessory vertebral articulations known as the
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Nunes, C.I.; Pujana, R.R.; Escapa, I.H.; Gandolfo, M.A.; CĂşneo, N.R. (2018). "A new species of
3901: 1180:. These expanded transverse processes would have resulted in greater attachment areas for the 9465: 9355: 9317: 9046: 5328: 2794: 1625: 1426:
in both aspects. The bottom end of the bone was mildly expanded, though not to the extent of
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Gaffney, E.S.; Rich, T.H.; Vickers-Rich, P.; Constantine, A.; Vacca, R.; Kool, L. (2007).
3657: 3598: 3572:"'The sheer scale is extraordinary': meet the titanosaur that dwarfs Dippy the diplodocus" 3489:"Massive titanosaur, biggest dinosaur ever found, squeezes into Museum of Natural History" 8: 6031: 3515:"Gigantic Dinosaur, 'Titanosaur,' Going on Display at American Museum of Natural History" 567: 380:
by José Carballido and colleagues. Preliminary studies and press releases suggested that
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González Riga, B.J.; Lamanna, M.C.; Ortiz David, L.D.; Calvo, J.O.; Coria, J.P. (2016).
3954: 3897: 3629: 3431: 1567:. Like other titanosauriforms, the head of the femur was angled upwards relative to the 1389: 1033: 9086: 6158: 4856: 4817: 4733: 4684: 4575: 4528: 4493: 4450: 4406: 4362: 4329: 4310: 4090: 4041: 4000: 3973: 3938: 3919: 3769: 3712: 3443: 3376: 3351: 3283: 2801: 1956: 1568: 1398:
inserted. The ulna was robust, being about 50% as broad at the top as it was long like
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fossils over a two-year period. The programme is also known by the alternative title,
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were very long, being at least five times as long as they were wide at the back, with
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from the Late Jurassic of Germany and a re-evaluation of its systematic affinities".
4367: 4314: 4302: 4213: 4131: 4094: 4046: 4028: 3978: 3879:"Giants and bizarres: body size of some southern South American Cretaceous dinosaurs" 3773: 3761: 3753: 3716: 3447: 3381: 1674: 1604: 1506: 1064: 743: 607: 599: 450: 442: 4821: 4737: 4126: 4109: 4001:"A gigantic new dinosaur from Argentina and the evolution of the sauropod hind foot" 3923: 3352:"A new giant titanosaur sheds light on body mass evolution among sauropod dinosaurs" 1041: 635: 9408: 9396: 9093: 8938: 8878: 8520: 8412: 8318: 8209: 8011: 7991: 7930: 7049: 6902: 6749: 6466: 6420: 6400: 6344: 5647: 5424: 5404: 4932: 4920: 4899: 4891: 4848: 4799: 4791: 4723: 4715: 4674: 4666: 4567: 4520: 4485: 4442: 4398: 4357: 4349: 4292: 4284: 4254: 4246: 4205: 4176: 4166: 4121: 4082: 4036: 4020: 3968: 3958: 3911: 3743: 3704: 3435: 3371: 3363: 3166: 3022: 2734: 2727: 2341: 1555: 1529: 1346:
was on average 28% as broad as it was long, making it similarly robust to those of
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and is notable for its large size. Carballido and colleagues stated in the media:
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was discovered in 2010 by a farm laborer, Aurelio Hernández, in the desert near
9237: 9152: 9034: 8979: 8929: 8741: 8734: 8680: 8652: 8607: 8580: 8455: 8420: 8362: 8289: 8202: 7966: 7845: 7817: 7796: 7781: 7703: 7676: 7494: 7478: 7444: 7287: 7086: 6958: 6784: 6523: 6502: 6451: 6443: 6295: 6288: 6195: 6094: 6082: 5890: 5862: 5699: 5670: 5663: 5609: 5523: 5509: 5493: 5468: 5298: 3193: 2866: 2757: 2747: 1809: 1796: 1763: 1670: 1505:, there was a ridge running along the side of the pubis that vanished near the 1181: 1136: 1121: 1089: 971:(neurovascular openings) on the sides. By contrast, the rear neck vertebrae of 790: 585: 4795: 4719: 4595:"Researchers Discover Fossils of Largest Dino Believed to Ever Walk the Earth" 4550:(Sauropoda: Titanosauria) from the Upper Cretaceous of Patagonia, Argentina". 4423: 1097: 1063:
had front neural spines with arrow-shaped ends, but like other members of the
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probably had 10. Several characteristics in the back vertebrae distinguished
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vertebrae, noting that the former was largely empty space. In other studies,
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Nunes, C.I.; Bodnar, J.; Escapa, I.H.; Gandolfo, M.A.; CĂşneo, N.R. (2019).
4371: 4306: 4259: 4135: 4050: 3982: 3765: 3732:"The accuracy and precision of body mass estimation in non-avian dinosaurs" 3689: 3576: 3385: 3367: 3242: 3182: 3008: 2937: 2900: 2886: 2693: 2408: 2391: 2358: 2296: 2283: 2143: 2129: 2055: 2014: 1857: 1843: 1698: 1684: 1666: 1633: 1535: 1501: 1448: 1354: 1348: 1311: 1299: 1198: 1173: 1149: 1142: 1082: 1026: 963:
on top that were horizontally flattened, and only small holes and no large
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muscle extended towards the front of the bone as a crest, as also seen in
1377: 519: 9273: 9181: 9167: 9063: 8952: 8804: 8797: 8790: 8756: 8688: 8666: 8587: 8478: 8173: 7980: 7945: 7838: 7803: 7745: 7724: 7717: 7689: 7622: 7594: 7559: 7552: 7538: 7437: 7422: 7323: 7245: 7208: 7079: 7030: 6791: 6667: 6630: 6623: 6592: 6576: 6516: 6480: 6458: 6378: 6337: 6281: 6253: 6239: 6187: 6140: 6060: 6045: 6017: 5883: 5841: 5827: 5807: 5781: 5727: 5720: 5636: 5602: 5588: 5431: 5369: 5305: 5282: 5248: 5220: 4151:"Cranial anatomy and phylogenetic position of the titanosaurian sauropod 4108:
Calvo, J.O.; Porfiri, J.D.; González-Riga, B.J.; Kellner, A.W.A. (2007).
3708: 3463:"Here's How You Squeeze the Biggest Dinosaur Into a New York City Museum" 3248: 2951: 2821: 2259: 2208: 2191: 1973: 1939: 1479: 1269: 1239: 1156: 658: 623: 43: 4804: 4728: 4679: 4353: 4297: 4202:
South American Sauropodomorph Dinosaurs: Record, Diversity and Evolution
4181: 3292:, followed the excavation, research and reconstruction of the La Flecha 1644:. The last of these characteristics were used to group lognkosaurs with 9174: 9101: 9070: 8832: 8659: 8506: 8441: 8405: 8391: 8326: 8297: 8260: 8223: 8166: 7752: 7696: 7651: 7601: 7573: 7531: 7467: 7415: 7397: 7384: 7308: 7301: 7294: 7280: 7187: 7180: 7173: 7093: 7005: 6980: 6710: 6660: 6644: 6608: 6509: 6385: 6309: 6223: 6166: 6117: 5869: 5685: 5574: 5438: 5397: 5362: 5091: 4904: 3939:"March of the Titans: The Locomotor Capabilities of Sauropod Dinosaurs" 3270: 3265: 3233: 3178: 2707: 2642: 1881: 1715: 1549: 1523: 1244: 1000:
and other titanosaurs that preserved a complete set of back vertebrae,
889: 781: 561: 541: 501: 345: 249: 201: 88: 53: 4839:, a large-toothed ceratosaur (Dinosauria: Theropoda) from Patagonia". 4232: 4024: 3748: 3731: 3282:
A BBC television programme broadcast in January 2016 and presented by
3146: 2665:
Due to its completeness, Carballido and colleagues in 2022 found that
1030:. Uniquely, the prezygodiapophyseal laminae (PRDLs, which ran between 9137: 9019: 8825: 8304: 8282: 8231: 7952: 7909: 7895: 7637: 7330: 7316: 7201: 6915: 6763: 6756: 6487: 6436: 5999: 5986: 5901: 5876: 5834: 5713: 5553: 5320: 5262: 4936: 3225: 3217: 2038: 1547:
was robust, being on average 23% as broad as it was long, similar to
1283: 1227: 878: 739: 549: 545: 400:
was of a similar size to, if not smaller than, its closest relatives
393: 363: 348: 233: 155: 93: 37: 9414: 5026:
Raising the Dinosaur Giant, PBS, with transcript near bottom of page
4467: 4273: 3998: 3546:"New mega-dinosaur at Field Museum is named Maximo, unveiled June 1" 758: 9437: 9145: 8811: 8621: 7824: 7665: 7524: 6777: 6267: 6110: 5820: 5595: 5567: 5475: 5390: 5383: 5290: 5227: 5109: 5085: 3228:
sediments of the Cerro Castaño Member, from which the specimens of
3187: 2580: 1509:(which was elongated like other titanosauriforms) at the top. Like 1185: 652: 351: 188: 175: 117: 83: 78: 63: 58: 48: 5030: 4918: 4545: 3937:
Sellers, W.I.; Margetts, L.; Coria, R.A.B.; Manning, P.L. (2013).
9457: 8181: 6585: 6429: 6316: 5774: 5007: 3174: 2600:. Also in 2019, Philip Mannion and colleagues found the group of 629: 130: 98: 73: 4107: 1071:
represents a reversal to the condition in non-titanosauriforms.
8710: 7888: 5706: 5079: 4384: 1483:) but less than half of the latter's length (common, except in 1435: 1214:
Forelimb and scapula at Museo Egidio Feruglio de Trelew, Chubut
557: 529: 165: 30: 4773: 4204:. Springer Earth System Sciences. Springer. pp. 269–298. 3789:"Did Scientists Just Unveil the Biggest Dinosaur of All Time?" 951:
The vertebrae in the front and middle portions of the neck of
4198: 3232:
were uncovered, are indicative of low-energy water flow on a
389: 9012: 5516: 4853:
10.1671/0272-4634(2004)024[0894:PAAOGS]2.0.CO;2
3936: 3409: 3221: 3177:; and leaf and flower impressions and fossilized wood from 3170: 836: 832: 698:
In 2017, José Luis Carballido, Diego Pol, Alejandro Otero,
4757:
Latin American Journal of Sedimentology and Basin Analysis
4063: 3876: 746:
for the "strength and large size" of this titanosaur. The
6674: 4701: 4644: 4328:
Mannion, P.D.; Upchurch, P.; Jin, X.; Zheng, W. (2019).
1243:, this figure was usually considerably higher among the 4510: 4327: 3877:
Mazzetta, G.V.; Christiansen, P.; Farina, R.A. (2004).
3356:
Proceedings of the Royal Society B: Biological Sciences
3349: 1188:
muscles, and therefore a more muscular tail. Also like
923:(3.67 metres (12.0 ft)), making it impossible for 888:
in 2017, science writer Riley Black and paleontologist
3410:
Otero, A.; Carballido, J.L.; PĂ©rez Moreno, A. (2020).
420:. This led to its use in a re-definition of the group 9380: 9345: 2612:
in a polytomy with other titanosaurs, which included
1059:, and the first few had expanded, arrow-shaped ends. 528:, Argentina, about 250 km (160 mi) west of 4750: 4621:"Argentine fossil biggest dinosaur ever: scientists" 4237:: implications for basal titanosaur relationships". 1386:
inserted. On the other side of the bone, behind the
750:
honours the Mayo family, owners of La Flecha ranch.
318: 4751:Carmona, R.P.; Umazano, A.M.; Krause, J.M. (2016). 3994: 3992: 1118:In most titanosaurs, the front tail vertebrae were 770:Skeletal mounts based on all available material of 9214: 4835:Rauhut, O.W.M. (2004). "Provenance and anatomy of 2567:has been consistently found as being grouped with 1677:shows the results of their phylogenetic analysis. 532:. This discovery was reported to P. Huerta at the 5004:Meet Patagotitan, the Biggest Dinosaur Ever Found 4194: 4192: 4148: 3345: 9485: 6741: 4943: 4267: 4226: 3989: 3343: 3341: 3339: 3337: 3335: 3333: 3331: 3329: 3327: 3325: 1067:the rear neural spines were inclined backwards; 734:one of the most completely-known members of the 4539: 2632:, with their grouping being in a polytomy with 5200: 5124: 5021:David Attenborough and the Giant Dinosaur, BBC 4378: 4189: 4057: 3730:Campione, Nicolás E.; Evans, David C. (2020). 5046: 3729: 3322: 1358:but less so than those of saltasaurids. Like 6864: 3870: 3623: 3621: 3619: 3506: 1673:and in turn the Titanosauria. The following 9494:Early Cretaceous dinosaurs of South America 8131: 4968: 4912: 4744: 4504: 4417: 4321: 4142: 3930: 3683: 3681: 3679: 3405: 3403: 3401: 3399: 3397: 3395: 847:In 2014 news reports provided estimates of 738:. The generic name combines a reference to 9509:Late Cretaceous dinosaurs of South America 5053: 5039: 4867: 4828: 4586: 3840: 3806: 3652: 3650: 3599:"Titanosaur: Life as the Biggest Dinosaur" 3537: 3173:; leaves, cones, and fossilized wood from 116: 7395: 4903: 4873: 4803: 4767: 4727: 4695: 4678: 4638: 4461: 4361: 4296: 4258: 4239:Zoological Journal of the Linnean Society 4180: 4170: 4125: 4101: 4040: 3972: 3962: 3905: 3780: 3747: 3616: 3512: 3375: 1205: 762:Reconstructed skeleton on display at the 540:and Diego Pol, with partial funding from 125:Reconstructed skeleton on display at the 5997: 4277:Anais da Academia Brasileira de CiĂŞncias 4114:Anais da Academia Brasileira de CiĂŞncias 3676: 3454: 3392: 3145: 1454: 1452:exhibited the bulge without the crest). 1209: 1109: 942: 831:was a quadrupedal herbivore with a long 815: 757: 566: 4618: 3647: 3543: 3480: 1251:. However, like other titanosaurs, the 730:. Collectively, the known remains made 651:Other specimens were designated as the 544:. More than 200 fossils, including 130 9486: 4834: 4592: 4149:Gallina, P.A.; ApesteguĂ­a, S. (2011). 3627: 534:Museum of Paleontology Egidio Feruglio 424:by Carballido and colleagues in 2022. 9419: 9418: 9344: 8118: 7382: 6851: 5984: 5187: 5072: 5034: 4976:"Attenborough and the Giant Dinosaur" 4951:"Attenborough and the Giant Dinosaur" 3857:Sauropod Vertebra Picture of the Week 3846: 3827:Sauropod Vertebra Picture of the Week 3812: 3786: 3569: 3460: 3277: 1665:was recovered as the sister group of 919:(4.47 metres (14.7 ft)) than in 884:Following the initial publication of 3687: 3630:"'Biggest dinosaur ever' discovered" 3563: 3486: 1628:by Carballido and colleagues placed 5188: 5060: 4391:Journal of Systematic Palaeontology 3289:Attenborough and the Giant Dinosaur 13: 4997: 4841:Journal of Vertebrate Paleontology 3591: 3420:Journal of Vertebrate Paleontology 1172:and other titanosaurs. Unlike the 777:American Museum of Natural History 764:American Museum of Natural History 491:hyposphene-hypantrum articulations 14: 9555: 9346:Topics in sauropodomorph research 6852: 5014: 3658:"Giant dinosaur slims down a bit" 3570:McKie, Robin (26 November 2022). 3513:Battaglia, A. (15 January 2016). 1619: 1543:. Like the humerus, the femur of 827:Like other titanosaur sauropods, 9402: 9390: 9327: 9321: 9316: 9310: 8119: 8100: 8092: 8084: 7365: 7360: 6834: 6828: 6822: 6814: 5966: 5959: 5954: 5948: 1114:Chevrons and rear tail vertebrae 927:to have had a larger torso. For 500:lived in a forested region on a 142: 41: 4612: 4127:10.1590/S0001-37652007000300013 3723: 3412:"The appendicular osteology of 3141: 787:Field Museum of Natural History 779:, where it replaced a juvenile 127:Field Museum of Natural History 7383: 1614:flexor digitorum longus muscle 806: 801:Natural History Museum, London 1: 9529:Fossil taxa described in 2017 5985: 4896:10.1016/j.cretres.2009.08.002 4671:10.1016/j.cretres.2019.02.013 4572:10.1016/j.cretres.2022.105389 4490:10.1016/j.cretres.2021.104754 4447:10.1016/j.cretres.2023.105487 4403:10.1080/14772019.2019.1683770 4289:10.1590/0001-3765201920180374 4159:Acta Palaeontologica Polonica 3697:Annals of the Carnegie Museum 3461:Black, R. (28 January 2016). 3440:10.1080/02724634.2020.1793158 3315: 640:in the shoulder girdle, both 4525:10.5710/AMGH.20.08.2020.3376 4087:10.1080/08912963.2012.672416 3964:10.1371/journal.pone.0078733 3916:10.1080/08912960410001715132 3847:Wedel, M. (10 August 2017). 3487:Mach, A. (16 January 2016). 2678:(instead of the less stable 2662:closer to the Rinconsauria. 1008:from other titanosaurs. The 938: 511: 372:, which was first announced 7: 4619:Harding, G. (17 May 2014). 4210:10.1007/978-3-030-95959-3_8 3813:Wedel, M. (9 August 2017). 3787:Black, R. (9 August 2017). 3544:Johnson, S. (11 May 2018). 3303: 3213:, was also named in 2018. 1571:on its outer top end. Like 1184:, ilio-ischiocaudalis, and 1081:, and other members of the 753: 722:named these remains as the 571:US ambassador to Argentina 10: 9560: 9544:Early Cretaceous sauropods 4593:Mohney, G. (17 May 2014). 4334:Royal Society Open Science 3849:"Some further thoughts on 3628:Morgan, J. (17 May 2014). 3298:Raising the Dinosaur Giant 1587:, and saltasaurines. Like 1094:at the rear end but not a 16:Extinct genus of dinosaurs 9427: 9351: 9340: 9308: 9210: 9119: 9045: 9007: 8998: 8962: 8928: 8846: 8780: 8729: 8720: 8647: 8638: 8616: 8605: 8540: 8531: 8146: 8142: 8127: 8114: 8081: 8047: 7999: 7990: 7862: 7684: 7675: 7584: 7519: 7508: 7477: 7410: 7406: 7391: 7378: 7358: 7270: 7227: 7218: 7120: 7111: 7048: 7013: 7004: 6910: 6901: 6890: 6875: 6860: 6847: 6811: 6737: 6709: 6693: 6684: 6575: 6419: 6368: 6248: 6237: 6206: 6153: 6149: 6138: 6093: 6012: 6008: 5993: 5980: 5946: 5900: 5815: 5806: 5747: 5738: 5646: 5619: 5562: 5551: 5492: 5463: 5452: 5414: 5357: 5353: 5342: 5272: 5215: 5211: 5196: 5183: 5151: 5133: 5120: 5068: 4929:American Museum Novitates 4796:10.1007/s00114-005-0623-3 4720:10.1163/22941932-20170206 4251:10.1093/zoolinnean/zlx103 3815:"Don't believe the hype: 3060: 3043: 3036: 3019: 3012: 2965: 2948: 2941: 2921: 2914: 2897: 2890: 2880: 2863: 2856: 2818: 2798: 2788: 2768: 2761: 2751: 2731: 2721: 2704: 2697: 2646:possibly being closer to 2446: 2429: 2422: 2405: 2388: 2381: 2355: 2338: 2331: 2324: 2307: 2300: 2280: 2273: 2256: 2249: 2205: 2188: 2181: 2164: 2157: 2140: 2133: 2123: 2052: 2035: 2028: 2011: 2004: 1987: 1970: 1953: 1936: 1919: 1902: 1895: 1878: 1871: 1854: 1847: 1827: 1820: 1813: 1793: 1786: 1760: 1743: 1736: 1729: 1712: 1695: 1688: 1636:to its closest relative, 431:and other members of the 317: 310: 139:Scientific classification 137: 124: 115: 23: 9366:List of sauropod species 3416:(Dinosauria, Sauropoda)" 3150:Life restoration of two 1396:latissimus dorsi muscles 789:, where it replaced the 386:largest known titanosaur 9524:Cerro Barcino Formation 4548:Rinconsaurus caudamirus 4235:Mendozasaurus neguyelap 1553:but somewhat less than 1384:coracobrachialis muscle 811: 700:Ignacio Alejandro Cerda 554:Cerro Barcino Formation 542:The Jurassic Foundation 356:Cerro Barcino Formation 4283:(suppl 2): e20180374. 3603:Natural History Museum 3368:10.1098/rspb.2017.1219 3284:Sir David Attenborough 3185:similar to the living 3155: 1460: 1215: 1206:Limbs and limb girdles 1115: 993:cannot be determined. 948: 845: 824: 767: 708:Alberto Carlos Garrido 580: 504:that was dominated by 9466:Paleobiology Database 9356:Sauropod neck posture 9047:Opisthocoelicaudiinae 4172:10.4202/app.2010.0011 3200:Austrocupressinoxylon 3165:specimens, including 3149: 2795:Opisthocoelicaudiinae 1626:phylogenetic analysis 1612:; this was where the 1458: 1213: 1182:caudofemoralis longus 1113: 946: 841: 823:compared with a human 819: 761: 720:Javier Marcelo Krause 628:, both plates of the 570: 471:in the forelimb, and 418:phylogenetic analyses 9519:Fossils of Argentina 9514:Cretaceous Argentina 4387:Europasaurus holgeri 4153:Bonitasaura salgadoi 3819:was not bigger than 3709:10.2992/007.085.0403 3467:Smithsonian Magazine 1387: 1375: 1260: 1252: 1147: 1119: 1095: 1087: 1047: 1046:at the front to the 1039: 1031: 1009: 964: 956: 859:was 10% larger than 688: 680: 672: 664: 656: 641: 633: 621: 613: 605: 597: 589: 538:Jose Luis Carballido 517: 488: 480: 472: 464: 456: 448: 440: 9215:Dubious titanosaurs 6032:Chinshakiangosaurus 4888:2009CrRes..30.1376L 4876:Cretaceous Research 4788:2005NW.....92..226N 4776:Naturwissenschaften 4663:2019CrRes..99..133N 4651:Cretaceous Research 4564:2023CrRes.14205389P 4552:Cretaceous Research 4482:2021CrRes.12204754O 4470:Cretaceous Research 4439:2023CrRes.14605487A 4427:Cretaceous Research 4354:10.1098/rsos.191057 4346:2019RSOS....691057M 4079:2012HBio...24..631C 4017:2016NatSR...619165G 3955:2013PLoSO...878733S 3898:2004HBio...16...71M 3688:Paul, G.S. (2019). 3519:Wall Street Journal 3432:2020JVPal..40E3158O 3414:Patagotitan mayorum 1390:deltopectoral crest 1374:. In a depression ( 1034:articular processes 728:Patagotitan mayorum 369:Patagotitan mayorum 323:Patagotitan mayorum 9087:Opisthocoelicaudia 6159:Archaeodontosaurus 4067:Historical Biology 4005:Scientific Reports 3886:Historical Biology 3736:Biological Reviews 3362:(1860): 20171219. 3278:In popular culture 3156: 2802:Opisthocoelicaudia 1957:Opisthocoelicaudia 1569:greater trochanter 1561:Opisthocoelicaudia 1461: 1428:Opisthocoelicaudia 1412:Opisthocoelicaudia 1368:Opisthocoelicaudia 1336:Opisthocoelicaudia 1216: 1116: 949: 825: 768: 716:NĂ©stor Ruben CĂşneo 581: 516:A part of a lower 9539:2014 in Argentina 9481: 9480: 9421:Taxon identifiers 9378: 9377: 9374: 9373: 9336: 9335: 9306: 9305: 9302: 9301: 9298: 9297: 9223:Bruhathkayosaurus 9206: 9205: 9202: 9201: 9198: 9197: 8924: 8923: 8842: 8841: 8764:Panamericansaurus 8601: 8600: 8597: 8596: 8110: 8109: 8079: 8078: 8075: 8074: 8071: 8070: 8067: 8066: 8043: 8042: 7511:Titanosauriformes 7504: 7503: 7374: 7373: 7356: 7355: 7352: 7351: 7348: 7347: 7344: 7343: 7340: 7339: 7152:Brachytrachelopan 7107: 7106: 7103: 7102: 6881:Haplocanthosaurus 6843: 6842: 6809: 6808: 6805: 6804: 6801: 6800: 6742:Dubious sauropods 6733: 6732: 6729: 6728: 6725: 6724: 6233: 6232: 6134: 6133: 6069:Protognathosaurus 5976: 5975: 5944: 5943: 5940: 5939: 5936: 5935: 5932: 5931: 5928: 5927: 5924: 5923: 5920: 5919: 5547: 5546: 5448: 5447: 5338: 5337: 5179: 5178: 5175: 5174: 5171: 5170: 4837:Genyodectes serus 4219:978-3-030-95958-6 4025:10.1038/srep19165 3749:10.1111/brv.12638 3525:on 5 October 2021 3310:Largest dinosaurs 3216:The fine-grained 3138: 3137: 3129: 3128: 3120: 3119: 3111: 3110: 3102: 3101: 3093: 3092: 3084: 3083: 3075: 3074: 2998: 2997: 2989: 2988: 2980: 2979: 2842: 2841: 2833: 2832: 2560: 2559: 2551: 2550: 2542: 2541: 2533: 2532: 2524: 2523: 2515: 2514: 2506: 2505: 2497: 2496: 2488: 2487: 2479: 2478: 2470: 2469: 2461: 2460: 2370: 2369: 2238: 2237: 2229: 2228: 2220: 2219: 2112: 2111: 2103: 2102: 2094: 2093: 2085: 2084: 2076: 2075: 2067: 2066: 1775: 1774: 1675:phylogenetic tree 1605:fourth trochanter 1507:obturator foramen 1065:Titanosauriformes 712:Jahandar Ramezani 586:holotype specimen 337: 336: 306: 131:Chicago, Illinois 9551: 9474: 9473: 9461: 9460: 9448: 9447: 9446: 9416: 9415: 9407: 9406: 9405: 9395: 9394: 9393: 9386: 9342: 9341: 9331: 9325: 9320: 9314: 9212: 9211: 9094:Pellegrinisaurus 9005: 9004: 8960: 8959: 8939:Bustingorrytitan 8879:Futalognkosaurus 8727: 8726: 8645: 8644: 8614: 8613: 8538: 8537: 8521:Diamantinasauria 8413:Pellegrinisaurus 8319:Jiangshanosaurus 8210:Baotianmansaurus 8144: 8143: 8140: 8139: 8129: 8128: 8116: 8115: 8105: 8104: 8097: 8096: 8089: 8088: 8012:Diamantinasaurus 7997: 7996: 7992:Diamantinasauria 7931:Jiangshanosaurus 7682: 7681: 7517: 7516: 7408: 7407: 7404: 7403: 7393: 7392: 7380: 7379: 7369: 7364: 7225: 7224: 7118: 7117: 7050:Rebbachisaurinae 7011: 7010: 6908: 6907: 6903:Rebbachisauridae 6899: 6898: 6873: 6872: 6862: 6861: 6849: 6848: 6838: 6832: 6827: 6826: 6819: 6818: 6750:Bothriospondylus 6739: 6738: 6691: 6690: 6467:Eomamenchisaurus 6421:Mamenchisauridae 6401:Lapparentosaurus 6345:Spinophorosaurus 6246: 6245: 6151: 6150: 6147: 6146: 6010: 6009: 6006: 6005: 5995: 5994: 5982: 5981: 5971: 5970: 5963: 5958: 5952: 5813: 5812: 5745: 5744: 5648:Massospondylidae 5560: 5559: 5490: 5489: 5461: 5460: 5425:Jaklapallisaurus 5405:Thecodontosaurus 5355: 5354: 5351: 5350: 5213: 5212: 5209: 5208: 5198: 5197: 5185: 5184: 5131: 5130: 5122: 5121: 5115: 5114: 5070: 5069: 5055: 5048: 5041: 5032: 5031: 4991: 4990: 4988: 4986: 4972: 4966: 4965: 4963: 4961: 4947: 4941: 4940: 4916: 4910: 4909: 4907: 4882:(6): 1376–1386. 4871: 4865: 4864: 4832: 4826: 4825: 4807: 4771: 4765: 4764: 4748: 4742: 4741: 4731: 4699: 4693: 4692: 4682: 4642: 4636: 4635: 4633: 4631: 4616: 4610: 4609: 4607: 4605: 4590: 4584: 4583: 4543: 4537: 4536: 4508: 4502: 4501: 4465: 4459: 4458: 4421: 4415: 4414: 4382: 4376: 4375: 4365: 4325: 4319: 4318: 4300: 4271: 4265: 4264: 4262: 4230: 4224: 4223: 4196: 4187: 4186: 4184: 4174: 4146: 4140: 4139: 4129: 4105: 4099: 4098: 4061: 4055: 4054: 4044: 3996: 3987: 3986: 3976: 3966: 3934: 3928: 3927: 3909: 3883: 3874: 3868: 3867: 3865: 3863: 3844: 3838: 3837: 3835: 3833: 3810: 3804: 3803: 3801: 3799: 3784: 3778: 3777: 3751: 3742:(6): 1759–1797. 3727: 3721: 3720: 3694: 3685: 3674: 3673: 3671: 3669: 3664:. 10 August 2017 3654: 3645: 3644: 3642: 3640: 3625: 3614: 3613: 3611: 3609: 3595: 3589: 3588: 3586: 3584: 3567: 3561: 3560: 3558: 3556: 3541: 3535: 3534: 3532: 3530: 3521:. Archived from 3510: 3504: 3503: 3501: 3499: 3484: 3478: 3477: 3475: 3473: 3458: 3452: 3451: 3407: 3390: 3389: 3379: 3347: 3039: 3038: 3023:Futalognkosaurus 3015: 3014: 2944: 2943: 2917: 2916: 2893: 2892: 2883: 2882: 2859: 2858: 2791: 2790: 2764: 2763: 2754: 2753: 2735:Diamantinasaurus 2728:Diamantinasauria 2724: 2723: 2700: 2699: 2690: 2689: 2618:Futalognkosaurus 2425: 2424: 2384: 2383: 2342:Futalognkosaurus 2334: 2333: 2327: 2326: 2303: 2302: 2276: 2275: 2252: 2251: 2184: 2183: 2160: 2159: 2136: 2135: 2126: 2125: 2031: 2030: 2007: 2006: 1898: 1897: 1874: 1873: 1850: 1849: 1823: 1822: 1816: 1815: 1789: 1788: 1739: 1738: 1732: 1731: 1691: 1690: 1681: 1680: 1585:Diamantinasaurus 1556:Diamantinasaurus 1530:Jiangshanosaurus 1515:Futalognkosaurus 1469:Futalognkosaurus 1391: 1379: 1264: 1256: 1190:Futalognkosaurus 1162:Futalognkosaurus 1151: 1123: 1099: 1091: 1051: 1043: 1035: 1013: 998:Futalognkosaurus 978:Futalognkosaurus 968: 960: 910: 853: 726:of a new genus, 704:Leonardo Salgado 692: 684: 676: 668: 660: 645: 637: 625: 617: 609: 601: 593: 521: 506:coniferous trees 492: 484: 476: 468: 460: 452: 444: 320: 300: 293: 280: 264: 248: 232: 216: 147: 146: 120: 110: 40: 29:Temporal range: 21: 20: 9559: 9558: 9554: 9553: 9552: 9550: 9549: 9548: 9534:2014 in science 9484: 9483: 9482: 9477: 9469: 9464: 9456: 9451: 9442: 9441: 9436: 9423: 9413: 9403: 9401: 9391: 9389: 9381: 9379: 9370: 9361:Sauropod hiatus 9347: 9332: 9294: 9231:Campylodoniscus 9194: 9115: 9041: 8994: 8964:Nemegtosauridae 8958: 8920: 8857:Argentinosaurus 8838: 8776: 8716: 8634: 8610: 8593: 8553:Atsinganosaurus 8533:Lirainosaurinae 8527: 8514:Xianshanosaurus 8399:Normanniasaurus 8385:Narambuenatitan 8268:Dongyangosaurus 8136: 8123: 8106: 8099: 8091: 8083: 8063: 8039: 8019:Sarmientosaurus 7986: 7939:Phuwiangosaurus 7858: 7671: 7586:Brachiosauridae 7580: 7513: 7500: 7473: 7400: 7387: 7370: 7336: 7266: 7214: 7122:Dicraeosauridae 7113:Flagellicaudata 7099: 7044: 7000: 6974:Nopcsaspondylus 6966:Maraapunisaurus 6895: 6893:Diplodocimorpha 6886: 6869: 6856: 6839: 6821: 6813: 6797: 6721: 6705: 6680: 6653:Tehuelchesaurus 6571: 6531:Tienshanosaurus 6415: 6364: 6352:Tehuelchesaurus 6242: 6229: 6208:Vulcanodontidae 6202: 6143: 6130: 6089: 6039:Gongxianosaurus 6002: 5989: 5972: 5965: 5916: 5896: 5802: 5761:Jingshanosaurus 5753:Chuxiongosaurus 5740:Sauropodiformes 5734: 5657:Adeopapposaurus 5642: 5615: 5556: 5543: 5488: 5457: 5444: 5410: 5347: 5334: 5268: 5205: 5203:Sauropodomorpha 5192: 5190:Sauropodomorpha 5167: 5153:Sauropodomorpha 5147: 5141:Avemetatarsalia 5126:Avemetatarsalia 5116: 5101:Avemetatarsalia 5075: 5074: 5064: 5062:Sauropodomorpha 5059: 5017: 5000: 4998:Further reading 4995: 4994: 4984: 4982: 4974: 4973: 4969: 4959: 4957: 4949: 4948: 4944: 4917: 4913: 4872: 4868: 4833: 4829: 4772: 4768: 4749: 4745: 4704:Carlquistoxylon 4700: 4696: 4643: 4639: 4629: 4627: 4617: 4613: 4603: 4601: 4591: 4587: 4544: 4540: 4509: 4505: 4466: 4462: 4422: 4418: 4383: 4379: 4326: 4322: 4272: 4268: 4231: 4227: 4220: 4197: 4190: 4147: 4143: 4106: 4102: 4062: 4058: 3997: 3990: 3935: 3931: 3907:10.1.1.694.1650 3881: 3875: 3871: 3861: 3859: 3845: 3841: 3831: 3829: 3821:Argentinosaurus 3811: 3807: 3797: 3795: 3785: 3781: 3728: 3724: 3692: 3686: 3677: 3667: 3665: 3656: 3655: 3648: 3638: 3636: 3626: 3617: 3607: 3605: 3597: 3596: 3592: 3582: 3580: 3568: 3564: 3554: 3552: 3550:Chicago Tribune 3542: 3538: 3528: 3526: 3511: 3507: 3497: 3495: 3485: 3481: 3471: 3469: 3459: 3455: 3426:(4): e1793158. 3408: 3393: 3348: 3323: 3318: 3306: 3280: 3206:Carlquistoxylon 3144: 3139: 3130: 3121: 3112: 3103: 3094: 3085: 3076: 2999: 2990: 2981: 2843: 2834: 2652:Argentinosaurus 2630:Argentinosaurus 2614:Argentinosaurus 2593:Argentinosaurus 2569:Argentinosaurus 2561: 2552: 2543: 2534: 2525: 2516: 2507: 2498: 2489: 2480: 2471: 2462: 2433:Argentinosaurus 2371: 2239: 2230: 2221: 2113: 2104: 2095: 2086: 2077: 2068: 1776: 1638:Argentinosaurus 1622: 1610:lateral condyle 1393: 1381: 1342:The humerus of 1266: 1258: 1208: 1153: 1125: 1101: 1093: 1061:Argentinosaurus 1053: 1045: 1042:prezygapophyses 1037: 1022:Argentinosaurus 1015: 970: 962: 941: 933:Argentinosaurus 917:Argentinosaurus 913:Argentinosaurus 908: 906:Argentinosaurus 898:Argentinosaurus 875:Argentinosaurus 867:Gregory S. Paul 862:Argentinosaurus 851: 814: 809: 756: 694: 686: 678: 670: 662: 647: 639: 636:scapulocoracoid 627: 619: 611: 603: 595: 573:Marc R. Stanley 523: 514: 494: 486: 478: 470: 462: 454: 446: 429:Argentinosaurus 403:Argentinosaurus 376:and then named 360:Chubut Province 333: 326: 299: 291: 278: 262: 246: 230: 218:Sauropodomorpha 214: 141: 111: 109: 108: 107: 106: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 35: 34: 33: 27: 17: 12: 11: 5: 9557: 9547: 9546: 9541: 9536: 9531: 9526: 9521: 9516: 9511: 9506: 9501: 9499:Lithostrotians 9496: 9479: 9478: 9476: 9475: 9462: 9449: 9433: 9431: 9425: 9424: 9412: 9411: 9399: 9376: 9375: 9372: 9371: 9369: 9368: 9363: 9358: 9352: 9349: 9348: 9338: 9337: 9334: 9333: 9309: 9307: 9304: 9303: 9300: 9299: 9296: 9295: 9293: 9292: 9285: 9277: 9270: 9262: 9255: 9248: 9241: 9238:Clasmodosaurus 9234: 9227: 9218: 9216: 9208: 9207: 9204: 9203: 9200: 9199: 9196: 9195: 9193: 9192: 9185: 9178: 9171: 9164: 9157: 9153:Maxakalisaurus 9149: 9142: 9134: 9125: 9123: 9117: 9116: 9114: 9113: 9105: 9098: 9090: 9083: 9075: 9067: 9060: 9051: 9049: 9043: 9042: 9040: 9039: 9035:Quaesitosaurus 9031: 9024: 9016: 9008: 9002: 8996: 8995: 8993: 8992: 8984: 8980:Quaesitosaurus 8976: 8968: 8966: 8957: 8956: 8949: 8942: 8934: 8932: 8930:Saltasauroidea 8926: 8925: 8922: 8921: 8919: 8918: 8911: 8904: 8897: 8890: 8882: 8875: 8867: 8860: 8852: 8850: 8844: 8843: 8840: 8839: 8837: 8836: 8829: 8822: 8815: 8808: 8801: 8794: 8786: 8784: 8778: 8777: 8775: 8774: 8767: 8760: 8753: 8746: 8742:Maxakalisaurus 8738: 8735:Adamantisaurus 8730: 8724: 8718: 8717: 8715: 8714: 8707: 8699: 8692: 8685: 8681:Epachthosaurus 8677: 8670: 8663: 8656: 8653:Antarctosaurus 8648: 8642: 8636: 8635: 8633: 8632: 8625: 8617: 8611: 8608:Eutitanosauria 8606: 8603: 8602: 8599: 8598: 8595: 8594: 8592: 8591: 8584: 8581:Mansourasaurus 8577: 8570: 8563: 8556: 8549: 8541: 8535: 8529: 8528: 8526: 8525: 8517: 8510: 8503: 8496: 8489: 8482: 8475: 8468: 8460: 8456:Tastavinsaurus 8452: 8445: 8438: 8431: 8424: 8421:Petrobrasaurus 8417: 8409: 8402: 8395: 8388: 8381: 8374: 8367: 8363:Malarguesaurus 8359: 8352: 8344: 8337: 8330: 8323: 8315: 8308: 8301: 8294: 8290:Epachthosaurus 8286: 8279: 8271: 8264: 8257: 8250: 8243: 8236: 8228: 8220: 8213: 8206: 8203:Austroposeidon 8199: 8192: 8185: 8178: 8170: 8163: 8155: 8147: 8137: 8132: 8125: 8124: 8112: 8111: 8108: 8107: 8082: 8080: 8077: 8076: 8073: 8072: 8069: 8068: 8065: 8064: 8062: 8061: 8060: 8059: 8053: 8051: 8045: 8044: 8041: 8040: 8038: 8037: 8029: 8022: 8015: 8008: 8000: 7994: 7988: 7987: 7985: 7984: 7977: 7970: 7967:Tangvayosaurus 7963: 7956: 7949: 7942: 7935: 7927: 7920: 7913: 7906: 7899: 7892: 7885: 7877: 7868: 7866: 7860: 7859: 7857: 7856: 7849: 7846:Tastavinsaurus 7842: 7835: 7828: 7821: 7818:Pukyongosaurus 7814: 7807: 7800: 7797:Liubangosaurus 7793: 7785: 7782:Liaoningotitan 7778: 7771: 7763: 7756: 7749: 7742: 7735: 7728: 7721: 7714: 7707: 7704:Astrophocaudia 7700: 7693: 7685: 7679: 7677:Somphospondyli 7673: 7672: 7670: 7669: 7662: 7655: 7648: 7641: 7634: 7627: 7619: 7612: 7605: 7598: 7590: 7588: 7582: 7581: 7579: 7578: 7570: 7563: 7556: 7549: 7542: 7535: 7528: 7520: 7514: 7509: 7506: 7505: 7502: 7501: 7499: 7498: 7495:Lourinhasaurus 7491: 7483: 7481: 7479:Camarasauridae 7475: 7474: 7472: 7471: 7464: 7456: 7449: 7445:Dashanpusaurus 7441: 7434: 7426: 7419: 7411: 7401: 7396: 7389: 7388: 7376: 7375: 7372: 7371: 7359: 7357: 7354: 7353: 7350: 7349: 7346: 7345: 7342: 7341: 7338: 7337: 7335: 7334: 7327: 7320: 7313: 7305: 7298: 7291: 7288:Dinheirosaurus 7284: 7276: 7274: 7268: 7267: 7265: 7264: 7257: 7249: 7242: 7233: 7231: 7222: 7216: 7215: 7213: 7212: 7205: 7198: 7191: 7184: 7177: 7170: 7162: 7155: 7148: 7141: 7134: 7126: 7124: 7115: 7109: 7108: 7105: 7104: 7101: 7100: 7098: 7097: 7090: 7087:Rebbachisaurus 7083: 7076: 7069: 7062: 7054: 7052: 7046: 7045: 7043: 7042: 7034: 7027: 7019: 7017: 7008: 7002: 7001: 6999: 6998: 6991: 6984: 6977: 6970: 6962: 6959:Lavocatisaurus 6955: 6948: 6940: 6933: 6926: 6919: 6911: 6905: 6896: 6891: 6888: 6887: 6885: 6884: 6876: 6870: 6865: 6858: 6857: 6845: 6844: 6841: 6840: 6812: 6810: 6807: 6806: 6803: 6802: 6799: 6798: 6796: 6795: 6788: 6785:Qinlingosaurus 6781: 6774: 6767: 6760: 6753: 6745: 6743: 6735: 6734: 6731: 6730: 6727: 6726: 6723: 6722: 6720: 6719: 6718:(see below ↓ ) 6715: 6713: 6707: 6706: 6704: 6703: 6702:(see below ↓ ) 6699: 6697: 6688: 6682: 6681: 6679: 6678: 6671: 6664: 6657: 6649: 6641: 6634: 6627: 6620: 6613: 6605: 6597: 6589: 6581: 6579: 6573: 6572: 6570: 6569: 6562: 6555: 6548: 6541: 6534: 6527: 6524:Rhomaleopakhus 6520: 6513: 6506: 6503:Mamenchisaurus 6499: 6491: 6484: 6477: 6470: 6463: 6455: 6452:Chuanjiesaurus 6448: 6444:Cetiosauriscus 6440: 6433: 6425: 6423: 6417: 6416: 6414: 6413: 6405: 6397: 6389: 6382: 6374: 6372: 6366: 6365: 6363: 6362: 6355: 6348: 6341: 6334: 6327: 6320: 6313: 6306: 6299: 6296:Dashanpusaurus 6292: 6289:Cetiosauriscus 6285: 6278: 6271: 6264: 6257: 6249: 6243: 6238: 6235: 6234: 6231: 6230: 6228: 6227: 6220: 6212: 6210: 6204: 6203: 6201: 6200: 6196:Zizhongosaurus 6192: 6184: 6177: 6170: 6163: 6154: 6144: 6139: 6136: 6135: 6132: 6131: 6129: 6128: 6121: 6114: 6107: 6099: 6097: 6095:Lessemsauridae 6091: 6090: 6088: 6087: 6083:Tuebingosaurus 6079: 6072: 6065: 6057: 6049: 6042: 6035: 6028: 6021: 6013: 6003: 5998: 5991: 5990: 5978: 5977: 5974: 5973: 5947: 5945: 5942: 5941: 5938: 5937: 5934: 5933: 5930: 5929: 5926: 5925: 5922: 5921: 5918: 5917: 5915: 5914: 5913: 5912: 5906: 5904: 5898: 5897: 5895: 5894: 5891:Sefapanosaurus 5887: 5880: 5873: 5866: 5863:Melanorosaurus 5859: 5852: 5845: 5838: 5831: 5824: 5816: 5810: 5804: 5803: 5801: 5800: 5793: 5786: 5778: 5771: 5764: 5757: 5748: 5742: 5736: 5735: 5733: 5732: 5724: 5717: 5710: 5703: 5700:Massospondylus 5696: 5689: 5682: 5674: 5671:Glacialisaurus 5667: 5664:Coloradisaurus 5660: 5652: 5650: 5644: 5643: 5641: 5640: 5633: 5625: 5623: 5617: 5616: 5614: 5613: 5610:Tuebingosaurus 5606: 5599: 5592: 5585: 5578: 5571: 5563: 5557: 5552: 5549: 5548: 5545: 5544: 5542: 5541: 5534: 5527: 5524:Pachysauriscus 5520: 5513: 5510:Gresslyosaurus 5506: 5498: 5496: 5494:Plateosauridae 5487: 5486: 5479: 5472: 5469:Plateosauravus 5464: 5458: 5453: 5450: 5449: 5446: 5445: 5443: 5442: 5435: 5428: 5420: 5418: 5412: 5411: 5409: 5408: 5401: 5394: 5387: 5380: 5373: 5366: 5358: 5348: 5343: 5340: 5339: 5336: 5335: 5333: 5332: 5325: 5317: 5309: 5302: 5299:Chromogisaurus 5295: 5287: 5278: 5276: 5270: 5269: 5267: 5266: 5259: 5252: 5245: 5238: 5231: 5224: 5216: 5206: 5201: 5194: 5193: 5181: 5180: 5177: 5176: 5173: 5172: 5169: 5168: 5166: 5165: 5164: 5163: 5157: 5155: 5149: 5148: 5146: 5145: 5144: 5143: 5134: 5128: 5118: 5117: 5113: 5112: 5103: 5094: 5088: 5082: 5066: 5065: 5058: 5057: 5050: 5043: 5035: 5029: 5028: 5023: 5016: 5015:External links 5013: 5012: 5011: 4999: 4996: 4993: 4992: 4967: 4942: 4931:(3599): 1–35. 4911: 4866: 4847:(4): 894–902. 4827: 4782:(5): 226–230. 4766: 4743: 4714:(4): 406–426. 4694: 4637: 4611: 4585: 4538: 4503: 4460: 4416: 4397:(9): 739–781. 4377: 4320: 4266: 4245:(1): 136–181. 4225: 4218: 4188: 4141: 4120:(3): 529–541. 4100: 4073:(6): 631–654. 4056: 3988: 3949:(10): e78733. 3929: 3869: 3839: 3805: 3779: 3722: 3703:(4): 335–358. 3675: 3646: 3615: 3590: 3562: 3536: 3505: 3479: 3453: 3391: 3320: 3319: 3317: 3314: 3313: 3312: 3305: 3302: 3279: 3276: 3194:Pilgerodendron 3143: 3140: 3136: 3135: 3132: 3131: 3127: 3126: 3123: 3122: 3118: 3117: 3114: 3113: 3109: 3108: 3105: 3104: 3100: 3099: 3096: 3095: 3091: 3090: 3087: 3086: 3082: 3081: 3078: 3077: 3073: 3072: 3069: 3068: 3059: 3056: 3055: 3052: 3051: 3042: 3037: 3035: 3032: 3031: 3028: 3027: 3018: 3013: 3011: 3005: 3004: 3001: 3000: 2996: 2995: 2992: 2991: 2987: 2986: 2983: 2982: 2978: 2977: 2974: 2973: 2964: 2961: 2960: 2957: 2956: 2947: 2942: 2940: 2934: 2933: 2930: 2929: 2920: 2915: 2913: 2910: 2909: 2906: 2905: 2896: 2891: 2889: 2881: 2879: 2876: 2875: 2872: 2871: 2867:Epachthosaurus 2862: 2857: 2855: 2849: 2848: 2845: 2844: 2840: 2839: 2836: 2835: 2831: 2830: 2827: 2826: 2817: 2811: 2810: 2807: 2806: 2797: 2789: 2787: 2781: 2780: 2777: 2776: 2767: 2762: 2760: 2758:Saltasauroidea 2752: 2750: 2748:Eutitanosauria 2744: 2743: 2740: 2739: 2730: 2722: 2720: 2717: 2716: 2713: 2712: 2703: 2698: 2696: 2688: 2558: 2557: 2554: 2553: 2549: 2548: 2545: 2544: 2540: 2539: 2536: 2535: 2531: 2530: 2527: 2526: 2522: 2521: 2518: 2517: 2513: 2512: 2509: 2508: 2504: 2503: 2500: 2499: 2495: 2494: 2491: 2490: 2486: 2485: 2482: 2481: 2477: 2476: 2473: 2472: 2468: 2467: 2464: 2463: 2459: 2458: 2455: 2454: 2445: 2442: 2441: 2438: 2437: 2428: 2423: 2421: 2418: 2417: 2414: 2413: 2404: 2401: 2400: 2397: 2396: 2387: 2382: 2380: 2377: 2376: 2373: 2372: 2368: 2367: 2364: 2363: 2354: 2351: 2350: 2347: 2346: 2337: 2332: 2330: 2325: 2323: 2320: 2319: 2316: 2315: 2306: 2301: 2299: 2293: 2292: 2289: 2288: 2279: 2274: 2272: 2269: 2268: 2265: 2264: 2255: 2250: 2248: 2245: 2244: 2241: 2240: 2236: 2235: 2232: 2231: 2227: 2226: 2223: 2222: 2218: 2217: 2214: 2213: 2204: 2201: 2200: 2197: 2196: 2187: 2182: 2180: 2177: 2176: 2173: 2172: 2163: 2158: 2156: 2153: 2152: 2149: 2148: 2139: 2134: 2132: 2124: 2122: 2119: 2118: 2115: 2114: 2110: 2109: 2106: 2105: 2101: 2100: 2097: 2096: 2092: 2091: 2088: 2087: 2083: 2082: 2079: 2078: 2074: 2073: 2070: 2069: 2065: 2064: 2061: 2060: 2051: 2048: 2047: 2044: 2043: 2034: 2029: 2027: 2024: 2023: 2020: 2019: 2010: 2005: 2003: 2000: 1999: 1996: 1995: 1986: 1983: 1982: 1979: 1978: 1969: 1966: 1965: 1962: 1961: 1952: 1949: 1948: 1945: 1944: 1935: 1932: 1931: 1928: 1927: 1918: 1915: 1914: 1911: 1910: 1901: 1896: 1894: 1891: 1890: 1887: 1886: 1877: 1872: 1870: 1867: 1866: 1863: 1862: 1853: 1848: 1846: 1840: 1839: 1836: 1835: 1826: 1821: 1819: 1814: 1812: 1810:Eutitanosauria 1806: 1805: 1802: 1801: 1797:Epachthosaurus 1792: 1787: 1785: 1782: 1781: 1778: 1777: 1773: 1772: 1769: 1768: 1764:Malarguesaurus 1759: 1756: 1755: 1752: 1751: 1742: 1737: 1735: 1730: 1728: 1725: 1724: 1721: 1720: 1711: 1708: 1707: 1704: 1703: 1694: 1689: 1687: 1679: 1671:Eutitanosauria 1621: 1620:Classification 1618: 1436:biceps brachii 1422:differed from 1364:Epachthosaurus 1207: 1204: 1137:Epachthosaurus 940: 937: 813: 810: 808: 805: 755: 752: 632:and the right 608:tail vertebrae 600:back vertebrae 592:neck vertebrae 513: 510: 451:tail vertebrae 344:is a genus of 335: 334: 327: 315: 314: 308: 307: 289: 285: 284: 276: 269: 268: 260: 253: 252: 244: 237: 236: 228: 221: 220: 212: 205: 204: 199: 192: 191: 186: 179: 178: 173: 169: 168: 163: 159: 158: 153: 149: 148: 135: 134: 122: 121: 113: 112: 104: 103: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 47: 42: 28: 15: 9: 6: 4: 3: 2: 9556: 9545: 9542: 9540: 9537: 9535: 9532: 9530: 9527: 9525: 9522: 9520: 9517: 9515: 9512: 9510: 9507: 9505: 9502: 9500: 9497: 9495: 9492: 9491: 9489: 9472: 9467: 9463: 9459: 9454: 9450: 9445: 9439: 9435: 9434: 9432: 9430: 9426: 9422: 9417: 9410: 9400: 9398: 9388: 9387: 9384: 9367: 9364: 9362: 9359: 9357: 9354: 9353: 9350: 9343: 9339: 9330: 9326: 9324: 9319: 9313: 9291: 9290: 9286: 9283: 9282: 9281:Mongolosaurus 9278: 9276: 9275: 9271: 9268: 9267: 9266:Macrurosaurus 9263: 9261: 9260: 9256: 9254: 9253: 9249: 9247: 9246: 9245:Hypselosaurus 9242: 9240: 9239: 9235: 9233: 9232: 9228: 9225: 9224: 9220: 9219: 9217: 9213: 9209: 9191: 9190: 9186: 9184: 9183: 9179: 9177: 9176: 9172: 9170: 9169: 9165: 9163: 9162: 9161:Neuquensaurus 9158: 9155: 9154: 9150: 9148: 9147: 9143: 9140: 9139: 9135: 9132: 9131: 9127: 9126: 9124: 9122: 9121:Saltasaurinae 9118: 9111: 9110: 9109:Zhuchengtitan 9106: 9104: 9103: 9099: 9096: 9095: 9091: 9089: 9088: 9084: 9081: 9080: 9079:Dreadnoughtus 9076: 9073: 9072: 9068: 9066: 9065: 9061: 9058: 9057: 9053: 9052: 9050: 9048: 9044: 9037: 9036: 9032: 9030: 9029: 9028:Qingxiusaurus 9025: 9022: 9021: 9017: 9015: 9014: 9010: 9009: 9006: 9003: 9001: 9000:Saltasauridae 8997: 8990: 8989: 8985: 8982: 8981: 8977: 8975: 8974: 8973:Nemegtosaurus 8970: 8969: 8967: 8965: 8961: 8955: 8954: 8950: 8948: 8947: 8943: 8941: 8940: 8936: 8935: 8933: 8931: 8927: 8917: 8916: 8912: 8910: 8909: 8905: 8903: 8902: 8898: 8896: 8895: 8894:Mendozasaurus 8891: 8888: 8887: 8883: 8881: 8880: 8876: 8873: 8872: 8871:Dzharatitanis 8868: 8866: 8865: 8861: 8859: 8858: 8854: 8853: 8851: 8849: 8845: 8835: 8834: 8830: 8828: 8827: 8823: 8821: 8820: 8819:Gondwanatitan 8816: 8814: 8813: 8809: 8807: 8806: 8802: 8800: 8799: 8795: 8793: 8792: 8788: 8787: 8785: 8783: 8779: 8773: 8772: 8768: 8766: 8765: 8761: 8759: 8758: 8754: 8752: 8751: 8750:Muyelensaurus 8747: 8744: 8743: 8739: 8737: 8736: 8732: 8731: 8728: 8725: 8723: 8719: 8713: 8712: 8708: 8705: 8704: 8700: 8698: 8697: 8693: 8691: 8690: 8686: 8683: 8682: 8678: 8676: 8675: 8674:Chucarosaurus 8671: 8669: 8668: 8664: 8662: 8661: 8657: 8655: 8654: 8650: 8649: 8646: 8643: 8641: 8640:Colossosauria 8637: 8631: 8630: 8629:Menucocelsior 8626: 8624: 8623: 8619: 8618: 8615: 8612: 8609: 8604: 8590: 8589: 8585: 8583: 8582: 8578: 8576: 8575: 8571: 8569: 8568: 8567:Lirainosaurus 8564: 8562: 8561: 8557: 8555: 8554: 8550: 8548: 8547: 8543: 8542: 8539: 8536: 8534: 8530: 8523: 8522: 8518: 8516: 8515: 8511: 8509: 8508: 8504: 8502: 8501: 8497: 8495: 8494: 8490: 8488: 8487: 8486:Trigonosaurus 8483: 8481: 8480: 8476: 8474: 8473: 8469: 8466: 8465: 8461: 8458: 8457: 8453: 8451: 8450: 8449:Ruyangosaurus 8446: 8444: 8443: 8439: 8437: 8436: 8432: 8430: 8429: 8428:Pitekunsaurus 8425: 8423: 8422: 8418: 8415: 8414: 8410: 8408: 8407: 8403: 8401: 8400: 8396: 8394: 8393: 8389: 8387: 8386: 8382: 8380: 8379: 8378:Mnyamawamtuka 8375: 8373: 8372: 8368: 8365: 8364: 8360: 8358: 8357: 8356:Magyarosaurus 8353: 8350: 8349: 8348:Ligabuesaurus 8345: 8343: 8342: 8341:Laplatasaurus 8338: 8336: 8335: 8334:Karongasaurus 8331: 8329: 8328: 8324: 8321: 8320: 8316: 8314: 8313: 8309: 8307: 8306: 8302: 8300: 8299: 8295: 8292: 8291: 8287: 8285: 8284: 8280: 8277: 8276: 8275:Dreadnoughtus 8272: 8270: 8269: 8265: 8263: 8262: 8258: 8256: 8255: 8251: 8249: 8248: 8244: 8242: 8241: 8240:Borealosaurus 8237: 8234: 8233: 8229: 8226: 8225: 8221: 8219: 8218: 8217:Barrosasaurus 8214: 8212: 8211: 8207: 8205: 8204: 8200: 8198: 8197: 8193: 8191: 8190: 8186: 8184: 8183: 8179: 8176: 8175: 8171: 8169: 8168: 8164: 8161: 8160: 8159:Aegyptosaurus 8156: 8154: 8153: 8149: 8148: 8145: 8141: 8138: 8135: 8130: 8126: 8122: 8117: 8113: 8103: 8098: 8095: 8087: 8057: 8056: 8055: 8054: 8052: 8050: 8046: 8035: 8034: 8030: 8028: 8027: 8026:Savannasaurus 8023: 8021: 8020: 8016: 8014: 8013: 8009: 8007: 8006: 8005:Australotitan 8002: 8001: 7998: 7995: 7993: 7989: 7983: 7982: 7978: 7976: 7975: 7971: 7969: 7968: 7964: 7962: 7961: 7957: 7955: 7954: 7950: 7948: 7947: 7943: 7941: 7940: 7936: 7933: 7932: 7928: 7926: 7925: 7921: 7919: 7918: 7914: 7912: 7911: 7907: 7905: 7904: 7900: 7898: 7897: 7893: 7891: 7890: 7886: 7883: 7882: 7878: 7875: 7874: 7873:Australodocus 7870: 7869: 7867: 7865: 7864:Euhelopodidae 7861: 7855: 7854: 7853:Triunfosaurus 7850: 7848: 7847: 7843: 7841: 7840: 7836: 7834: 7833: 7832:Sauroposeidon 7829: 7827: 7826: 7822: 7820: 7819: 7815: 7813: 7812: 7811:Padillasaurus 7808: 7806: 7805: 7801: 7799: 7798: 7794: 7791: 7790: 7789:Ligabuesaurus 7786: 7784: 7783: 7779: 7777: 7776: 7772: 7769: 7768: 7764: 7762: 7761: 7757: 7755: 7754: 7750: 7748: 7747: 7743: 7741: 7740: 7736: 7734: 7733: 7732:Chubutisaurus 7729: 7727: 7726: 7722: 7720: 7719: 7715: 7713: 7712: 7708: 7706: 7705: 7701: 7699: 7698: 7694: 7692: 7691: 7687: 7686: 7683: 7680: 7678: 7674: 7668: 7667: 7663: 7661: 7660: 7656: 7654: 7653: 7649: 7647: 7646: 7642: 7640: 7639: 7635: 7633: 7632: 7628: 7625: 7624: 7620: 7618: 7617: 7613: 7611: 7610: 7609:Brachiosaurus 7606: 7604: 7603: 7599: 7597: 7596: 7592: 7591: 7589: 7587: 7583: 7576: 7575: 7571: 7569: 7568: 7564: 7562: 7561: 7557: 7555: 7554: 7550: 7548: 7547: 7546:Fushanosaurus 7543: 7541: 7540: 7536: 7534: 7533: 7529: 7527: 7526: 7522: 7521: 7518: 7515: 7512: 7507: 7497: 7496: 7492: 7490: 7489: 7485: 7484: 7482: 7480: 7476: 7470: 7469: 7465: 7462: 7461: 7457: 7455: 7454: 7450: 7447: 7446: 7442: 7440: 7439: 7435: 7432: 7431: 7430:Bashunosaurus 7427: 7425: 7424: 7420: 7418: 7417: 7413: 7412: 7409: 7405: 7402: 7399: 7394: 7390: 7386: 7381: 7377: 7368: 7363: 7333: 7332: 7328: 7326: 7325: 7321: 7319: 7318: 7314: 7311: 7310: 7306: 7304: 7303: 7299: 7297: 7296: 7292: 7290: 7289: 7285: 7283: 7282: 7278: 7277: 7275: 7273: 7269: 7263: 7262: 7258: 7255: 7254: 7253:Atlantosaurus 7250: 7248: 7247: 7243: 7240: 7239: 7235: 7234: 7232: 7230: 7229:Apatosaurinae 7226: 7223: 7221: 7217: 7211: 7210: 7206: 7204: 7203: 7199: 7197: 7196: 7195:Smitanosaurus 7192: 7190: 7189: 7185: 7183: 7182: 7178: 7176: 7175: 7171: 7168: 7167: 7166:Dyslocosaurus 7163: 7161: 7160: 7159:Dicraeosaurus 7156: 7154: 7153: 7149: 7147: 7146: 7142: 7140: 7139: 7138:Amargatitanis 7135: 7133: 7132: 7128: 7127: 7125: 7123: 7119: 7116: 7114: 7110: 7096: 7095: 7091: 7089: 7088: 7084: 7082: 7081: 7077: 7075: 7074: 7073:Katepensaurus 7070: 7068: 7067: 7066:Itapeuasaurus 7063: 7061: 7060: 7059:Demandasaurus 7056: 7055: 7053: 7051: 7047: 7040: 7039: 7035: 7033: 7032: 7028: 7026: 7025: 7024:Cathartesaura 7021: 7020: 7018: 7016: 7015:Limaysaurinae 7012: 7009: 7007: 7003: 6997: 6996: 6992: 6990: 6989: 6985: 6983: 6982: 6978: 6976: 6975: 6971: 6968: 6967: 6963: 6961: 6960: 6956: 6954: 6953: 6952:Histriasaurus 6949: 6946: 6945: 6944:Dzharatitanis 6941: 6939: 6938: 6937:Comahuesaurus 6934: 6932: 6931: 6927: 6925: 6924: 6920: 6918: 6917: 6913: 6912: 6909: 6906: 6904: 6900: 6897: 6894: 6889: 6883: 6882: 6878: 6877: 6874: 6871: 6868: 6867:Diplodocoidea 6863: 6859: 6855: 6854:Diplodocoidea 6850: 6846: 6837: 6833: 6831: 6825: 6817: 6794: 6793: 6789: 6787: 6786: 6782: 6780: 6779: 6775: 6773: 6772: 6771:Gigantosaurus 6768: 6766: 6765: 6761: 6759: 6758: 6754: 6752: 6751: 6747: 6746: 6744: 6740: 6736: 6717: 6716: 6714: 6712: 6708: 6701: 6700: 6698: 6696: 6695:Diplodocoidea 6692: 6689: 6687: 6683: 6677: 6676: 6672: 6670: 6669: 6665: 6663: 6662: 6658: 6655: 6654: 6650: 6647: 6646: 6642: 6640: 6639: 6638:Narindasaurus 6635: 6633: 6632: 6628: 6626: 6625: 6621: 6619: 6618: 6617:Losillasaurus 6614: 6611: 6610: 6606: 6603: 6602: 6598: 6595: 6594: 6590: 6588: 6587: 6583: 6582: 6580: 6578: 6574: 6568: 6567: 6566:Zigongosaurus 6563: 6561: 6560: 6559:Yuanmousaurus 6556: 6554: 6553: 6552:Xinjiangtitan 6549: 6547: 6546: 6545:Wamweracaudia 6542: 6540: 6539: 6538:Tonganosaurus 6535: 6533: 6532: 6528: 6526: 6525: 6521: 6519: 6518: 6514: 6512: 6511: 6507: 6505: 6504: 6500: 6497: 6496: 6495:Klamelisaurus 6492: 6490: 6489: 6485: 6483: 6482: 6478: 6476: 6475: 6474:Huangshanlong 6471: 6469: 6468: 6464: 6461: 6460: 6456: 6454: 6453: 6449: 6446: 6445: 6441: 6439: 6438: 6434: 6432: 6431: 6427: 6426: 6424: 6422: 6418: 6411: 6410: 6406: 6403: 6402: 6398: 6395: 6394: 6393:Ferganasaurus 6390: 6388: 6387: 6383: 6381: 6380: 6376: 6375: 6373: 6371: 6370:Cetiosauridae 6367: 6361: 6360: 6356: 6354: 6353: 6349: 6347: 6346: 6342: 6340: 6339: 6335: 6333: 6332: 6328: 6326: 6325: 6321: 6319: 6318: 6314: 6312: 6311: 6307: 6305: 6304: 6300: 6298: 6297: 6293: 6291: 6290: 6286: 6284: 6283: 6279: 6277: 6276: 6272: 6270: 6269: 6265: 6263: 6262: 6258: 6256: 6255: 6251: 6250: 6247: 6244: 6241: 6236: 6226: 6225: 6221: 6219: 6218: 6217:Tazoudasaurus 6214: 6213: 6211: 6209: 6205: 6198: 6197: 6193: 6190: 6189: 6185: 6183: 6182: 6178: 6176: 6175: 6174:Ohmdenosaurus 6171: 6169: 6168: 6164: 6161: 6160: 6156: 6155: 6152: 6148: 6145: 6142: 6137: 6127: 6126: 6122: 6120: 6119: 6115: 6113: 6112: 6108: 6106: 6105: 6101: 6100: 6098: 6096: 6092: 6085: 6084: 6080: 6078: 6077: 6076:Schleitheimia 6073: 6071: 6070: 6066: 6063: 6062: 6058: 6055: 6054: 6053:Leonerasaurus 6050: 6048: 6047: 6043: 6041: 6040: 6036: 6034: 6033: 6029: 6027: 6026: 6025:Blikanasaurus 6022: 6020: 6019: 6015: 6014: 6011: 6007: 6004: 6001: 5996: 5992: 5988: 5983: 5979: 5969: 5964: 5962: 5957: 5951: 5910: 5909: 5908: 5907: 5905: 5903: 5899: 5893: 5892: 5888: 5886: 5885: 5881: 5879: 5878: 5874: 5872: 5871: 5867: 5865: 5864: 5860: 5858: 5857: 5856:Leonerasaurus 5853: 5851: 5850: 5849:Lamplughsaura 5846: 5844: 5843: 5839: 5837: 5836: 5832: 5830: 5829: 5825: 5823: 5822: 5818: 5817: 5814: 5811: 5809: 5805: 5799: 5798: 5797:Yunnanosaurus 5794: 5792: 5791: 5787: 5784: 5783: 5779: 5777: 5776: 5772: 5770: 5769: 5765: 5763: 5762: 5758: 5755: 5754: 5750: 5749: 5746: 5743: 5741: 5737: 5730: 5729: 5725: 5723: 5722: 5718: 5716: 5715: 5711: 5709: 5708: 5704: 5702: 5701: 5697: 5695: 5694: 5693:Lufengosaurus 5690: 5688: 5687: 5683: 5680: 5679: 5675: 5673: 5672: 5668: 5666: 5665: 5661: 5659: 5658: 5654: 5653: 5651: 5649: 5645: 5639: 5638: 5634: 5632: 5631: 5630:Eucnemesaurus 5627: 5626: 5624: 5622: 5621:Riojasauridae 5618: 5612: 5611: 5607: 5605: 5604: 5600: 5598: 5597: 5593: 5591: 5590: 5586: 5584: 5583: 5579: 5577: 5576: 5572: 5570: 5569: 5565: 5564: 5561: 5558: 5555: 5550: 5540: 5539: 5535: 5533: 5532: 5528: 5526: 5525: 5521: 5519: 5518: 5514: 5512: 5511: 5507: 5505: 5504: 5503:Euskelosaurus 5500: 5499: 5497: 5495: 5491: 5485: 5484: 5480: 5478: 5477: 5473: 5471: 5470: 5466: 5465: 5462: 5459: 5456: 5451: 5441: 5440: 5436: 5434: 5433: 5429: 5427: 5426: 5422: 5421: 5419: 5417: 5413: 5407: 5406: 5402: 5400: 5399: 5395: 5393: 5392: 5388: 5386: 5385: 5381: 5379: 5378: 5377:Bagualosaurus 5374: 5372: 5371: 5367: 5365: 5364: 5360: 5359: 5356: 5352: 5349: 5346: 5345:Bagualosauria 5341: 5331: 5330: 5326: 5323: 5322: 5318: 5315: 5314: 5313:Pampadromaeus 5310: 5308: 5307: 5303: 5301: 5300: 5296: 5293: 5292: 5288: 5285: 5284: 5280: 5279: 5277: 5275: 5274:Saturnaliidae 5271: 5265: 5264: 5260: 5258: 5257: 5256:Pampadromaeus 5253: 5251: 5250: 5246: 5244: 5243: 5239: 5237: 5236: 5235:Gigantoscelus 5232: 5230: 5229: 5225: 5223: 5222: 5218: 5217: 5214: 5210: 5207: 5204: 5199: 5195: 5191: 5186: 5182: 5161: 5160: 5159: 5158: 5156: 5154: 5150: 5142: 5138: 5137: 5136: 5135: 5132: 5129: 5127: 5123: 5119: 5111: 5107: 5104: 5102: 5098: 5095: 5093: 5089: 5087: 5083: 5081: 5077: 5076: 5071: 5067: 5063: 5056: 5051: 5049: 5044: 5042: 5037: 5036: 5033: 5027: 5024: 5022: 5019: 5018: 5009: 5005: 5002: 5001: 4981: 4977: 4971: 4956: 4952: 4946: 4938: 4934: 4930: 4926: 4924: 4915: 4906: 4901: 4897: 4893: 4889: 4885: 4881: 4877: 4870: 4862: 4858: 4854: 4850: 4846: 4842: 4838: 4831: 4823: 4819: 4815: 4811: 4806: 4801: 4797: 4793: 4789: 4785: 4781: 4777: 4770: 4763:(2): 127–132. 4762: 4758: 4754: 4747: 4739: 4735: 4730: 4725: 4721: 4717: 4713: 4709: 4705: 4698: 4690: 4686: 4681: 4676: 4672: 4668: 4664: 4660: 4656: 4652: 4648: 4641: 4626: 4625:NY Daily News 4622: 4615: 4600: 4596: 4589: 4581: 4577: 4573: 4569: 4565: 4561: 4557: 4553: 4549: 4542: 4534: 4530: 4526: 4522: 4518: 4514: 4507: 4499: 4495: 4491: 4487: 4483: 4479: 4475: 4471: 4464: 4456: 4452: 4448: 4444: 4440: 4436: 4432: 4428: 4420: 4412: 4408: 4404: 4400: 4396: 4392: 4388: 4381: 4373: 4369: 4364: 4359: 4355: 4351: 4347: 4343: 4340:(8): 191057. 4339: 4335: 4331: 4324: 4316: 4312: 4308: 4304: 4299: 4294: 4290: 4286: 4282: 4278: 4270: 4261: 4260:10044/1/53967 4256: 4252: 4248: 4244: 4240: 4236: 4229: 4221: 4215: 4211: 4207: 4203: 4195: 4193: 4183: 4178: 4173: 4168: 4164: 4160: 4156: 4154: 4145: 4137: 4133: 4128: 4123: 4119: 4115: 4111: 4104: 4096: 4092: 4088: 4084: 4080: 4076: 4072: 4068: 4060: 4052: 4048: 4043: 4038: 4034: 4030: 4026: 4022: 4018: 4014: 4010: 4006: 4002: 3995: 3993: 3984: 3980: 3975: 3970: 3965: 3960: 3956: 3952: 3948: 3944: 3940: 3933: 3925: 3921: 3917: 3913: 3908: 3903: 3899: 3895: 3892:(2–4): 1–13. 3891: 3887: 3880: 3873: 3858: 3854: 3852: 3843: 3828: 3824: 3822: 3818: 3809: 3794: 3790: 3783: 3775: 3771: 3767: 3763: 3759: 3755: 3750: 3745: 3741: 3737: 3733: 3726: 3718: 3714: 3710: 3706: 3702: 3698: 3691: 3684: 3682: 3680: 3663: 3659: 3653: 3651: 3635: 3631: 3624: 3622: 3620: 3604: 3600: 3594: 3579: 3578: 3573: 3566: 3551: 3547: 3540: 3524: 3520: 3516: 3509: 3494: 3490: 3483: 3468: 3464: 3457: 3449: 3445: 3441: 3437: 3433: 3429: 3425: 3421: 3417: 3415: 3406: 3404: 3402: 3400: 3398: 3396: 3387: 3383: 3378: 3373: 3369: 3365: 3361: 3357: 3353: 3346: 3344: 3342: 3340: 3338: 3336: 3334: 3332: 3330: 3328: 3326: 3321: 3311: 3308: 3307: 3301: 3299: 3295: 3291: 3290: 3285: 3275: 3273: 3272: 3267: 3263: 3262: 3261:Barcinosuchus 3258: 3257:crocodyliform 3255: 3251: 3250: 3245: 3244: 3239: 3235: 3231: 3227: 3223: 3219: 3214: 3212: 3208: 3207: 3202: 3201: 3196: 3195: 3190: 3189: 3184: 3180: 3176: 3172: 3168: 3164: 3160: 3153: 3148: 3134: 3133: 3125: 3124: 3116: 3115: 3107: 3106: 3098: 3097: 3089: 3088: 3080: 3079: 3071: 3070: 3067: 3066: 3065: 3058: 3057: 3054: 3053: 3050: 3049: 3048: 3047:Mendozasaurus 3041: 3040: 3034: 3033: 3030: 3029: 3026: 3025: 3024: 3017: 3016: 3010: 3007: 3006: 3003: 3002: 2994: 2993: 2985: 2984: 2976: 2975: 2972: 2971: 2970: 2969:Gondwanatitan 2963: 2962: 2959: 2958: 2955: 2954: 2953: 2946: 2945: 2939: 2936: 2935: 2932: 2931: 2928: 2927: 2926: 2925:Muyelensaurus 2919: 2918: 2912: 2911: 2908: 2907: 2904: 2903: 2902: 2895: 2894: 2888: 2885: 2884: 2878: 2877: 2874: 2873: 2870: 2869: 2868: 2861: 2860: 2854: 2853:Colossosauria 2851: 2850: 2847: 2846: 2838: 2837: 2829: 2828: 2825: 2824: 2823: 2816: 2815:Saltasaurinae 2813: 2812: 2809: 2808: 2805: 2804: 2803: 2796: 2793: 2792: 2786: 2785:Saltasauridae 2783: 2782: 2779: 2778: 2775: 2774: 2773: 2772:Nemegtosaurus 2766: 2765: 2759: 2756: 2755: 2749: 2746: 2745: 2742: 2741: 2738: 2737: 2736: 2729: 2726: 2725: 2719: 2718: 2715: 2714: 2711: 2710: 2709: 2702: 2701: 2695: 2692: 2691: 2687: 2685: 2681: 2680:Mendozasaurus 2677: 2673: 2668: 2663: 2661: 2657: 2653: 2649: 2645: 2644: 2639: 2635: 2631: 2628:grouped with 2627: 2623: 2622:Mendozasaurus 2619: 2615: 2611: 2607: 2603: 2599: 2598:Colossosauria 2594: 2590: 2586: 2582: 2578: 2574: 2570: 2566: 2556: 2555: 2547: 2546: 2538: 2537: 2529: 2528: 2520: 2519: 2511: 2510: 2502: 2501: 2493: 2492: 2484: 2483: 2475: 2474: 2466: 2465: 2457: 2456: 2453: 2452: 2451: 2444: 2443: 2440: 2439: 2436: 2435: 2434: 2427: 2426: 2420: 2419: 2416: 2415: 2412: 2411: 2410: 2403: 2402: 2399: 2398: 2395: 2394: 2393: 2386: 2385: 2379: 2378: 2375: 2374: 2366: 2365: 2362: 2361: 2360: 2353: 2352: 2349: 2348: 2345: 2344: 2343: 2336: 2335: 2329: 2328: 2322: 2321: 2318: 2317: 2314: 2313: 2312: 2311:Mendozasaurus 2305: 2304: 2298: 2295: 2294: 2291: 2290: 2287: 2286: 2285: 2278: 2277: 2271: 2270: 2267: 2266: 2263: 2262: 2261: 2254: 2253: 2247: 2246: 2243: 2242: 2234: 2233: 2225: 2224: 2216: 2215: 2212: 2211: 2210: 2203: 2202: 2199: 2198: 2195: 2194: 2193: 2186: 2185: 2179: 2178: 2175: 2174: 2171: 2170: 2169: 2168:Muyelensaurus 2162: 2161: 2155: 2154: 2151: 2150: 2147: 2146: 2145: 2138: 2137: 2131: 2128: 2127: 2121: 2120: 2117: 2116: 2108: 2107: 2099: 2098: 2090: 2089: 2081: 2080: 2072: 2071: 2063: 2062: 2059: 2058: 2057: 2050: 2049: 2046: 2045: 2042: 2041: 2040: 2033: 2032: 2026: 2025: 2022: 2021: 2018: 2017: 2016: 2009: 2008: 2002: 2001: 1998: 1997: 1994: 1993: 1992: 1991:Neuquensaurus 1985: 1984: 1981: 1980: 1977: 1976: 1975: 1968: 1967: 1964: 1963: 1960: 1959: 1958: 1951: 1950: 1947: 1946: 1943: 1942: 1941: 1934: 1933: 1930: 1929: 1926: 1925: 1924: 1923:Trigonosaurus 1917: 1916: 1913: 1912: 1909: 1908: 1907: 1906:Nemegtosaurus 1900: 1899: 1893: 1892: 1889: 1888: 1885: 1884: 1883: 1876: 1875: 1869: 1868: 1865: 1864: 1861: 1860: 1859: 1852: 1851: 1845: 1842: 1841: 1838: 1837: 1834: 1833: 1832: 1831:Dreadnoughtus 1825: 1824: 1818: 1817: 1811: 1808: 1807: 1804: 1803: 1800: 1799: 1798: 1791: 1790: 1784: 1783: 1780: 1779: 1771: 1770: 1767: 1766: 1765: 1758: 1757: 1754: 1753: 1750: 1749: 1748: 1747:Ruyangosaurus 1741: 1740: 1734: 1733: 1727: 1726: 1723: 1722: 1719: 1718: 1717: 1710: 1709: 1706: 1705: 1702: 1701: 1700: 1693: 1692: 1686: 1683: 1682: 1678: 1676: 1672: 1668: 1664: 1660: 1656: 1652: 1651:Dreadnoughtus 1647: 1643: 1639: 1635: 1631: 1627: 1617: 1615: 1611: 1606: 1602: 1598: 1594: 1593:Neuquensaurus 1590: 1586: 1582: 1578: 1574: 1573:Neuquensaurus 1570: 1566: 1562: 1558: 1557: 1552: 1551: 1546: 1542: 1538: 1537: 1532: 1531: 1526: 1525: 1520: 1516: 1512: 1508: 1504: 1503: 1498: 1494: 1490: 1486: 1482: 1481: 1476: 1475: 1474:Muyelensaurus 1470: 1466: 1465:Dreadnoughtus 1457: 1453: 1451: 1450: 1445: 1441: 1437: 1433: 1429: 1425: 1424:Neuquensaurus 1421: 1417: 1413: 1409: 1408:Neuquensaurus 1405: 1401: 1400:Dreadnoughtus 1397: 1392: 1385: 1380: 1373: 1369: 1365: 1361: 1360:Dreadnoughtus 1357: 1356: 1351: 1350: 1345: 1340: 1338: 1337: 1332: 1331:Dreadnoughtus 1328: 1327:Saltasauridae 1324: 1323: 1322:Savannasaurus 1318: 1317:Dreadnoughtus 1314: 1313: 1308: 1307: 1306:Ruyangosaurus 1302: 1301: 1296: 1292: 1291: 1290:Sauroposeidon 1286: 1285: 1280: 1276: 1275:Dreadnoughtus 1272: 1271: 1265: 1257: 1250: 1246: 1242: 1241: 1236: 1235: 1234:Neuquensaurus 1230: 1229: 1224: 1220: 1212: 1203: 1201: 1200: 1195: 1194:Mendozasaurus 1191: 1187: 1183: 1179: 1175: 1171: 1170:Dreadnoughtus 1167: 1166:Mendozasaurus 1163: 1159: 1158: 1152: 1145: 1144: 1139: 1138: 1133: 1132:Dreadnoughtus 1129: 1124: 1112: 1108: 1106: 1100: 1092: 1084: 1080: 1076: 1075: 1074:Dreadnoughtus 1070: 1066: 1062: 1058: 1052: 1044: 1036: 1029: 1028: 1023: 1019: 1014: 1007: 1003: 999: 994: 992: 991: 990:Mendozasaurus 986: 985: 984:Ligabuesaurus 980: 979: 974: 969: 961: 959:neural spines 954: 945: 936: 934: 930: 926: 922: 918: 914: 907: 903: 899: 895: 891: 887: 882: 880: 876: 872: 868: 864: 863: 858: 850: 844: 840: 838: 834: 830: 822: 818: 804: 802: 798: 797: 796:Tyrannosaurus 792: 788: 784: 783: 778: 773: 765: 760: 751: 749: 748:specific name 745: 742:with a Greek 741: 737: 733: 729: 725: 721: 717: 713: 709: 705: 701: 696: 693: 685: 677: 669: 661: 654: 649: 646: 638: 631: 626: 618: 610: 602: 594: 587: 578: 574: 569: 565: 563: 559: 555: 551: 548:bones and 57 547: 543: 539: 535: 531: 527: 522: 509: 507: 503: 499: 493: 485: 477: 469: 461: 453: 445: 438: 434: 430: 425: 423: 422:Colossosauria 419: 415: 411: 410: 405: 404: 399: 395: 391: 387: 383: 379: 375: 371: 370: 365: 361: 357: 353: 350: 347: 346:titanosaurian 343: 342: 331: 325: 324: 316: 313: 309: 304: 298: 297: 290: 287: 286: 283: 277: 274: 271: 270: 267: 261: 258: 255: 254: 251: 245: 242: 239: 238: 235: 229: 226: 223: 222: 219: 213: 210: 207: 206: 203: 200: 197: 194: 193: 190: 187: 184: 181: 180: 177: 174: 171: 170: 167: 164: 161: 160: 157: 154: 151: 150: 145: 140: 136: 132: 128: 123: 119: 114: 100: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 39: 32: 26: 22: 19: 9428: 9315: 9289:Titanosaurus 9287: 9279: 9272: 9264: 9259:Loricosaurus 9257: 9252:Iuticosaurus 9250: 9243: 9236: 9229: 9221: 9189:Yamanasaurus 9187: 9180: 9173: 9166: 9159: 9151: 9144: 9136: 9130:Abditosaurus 9128: 9107: 9100: 9092: 9085: 9077: 9069: 9062: 9056:Abditosaurus 9054: 9033: 9026: 9018: 9011: 8988:Tapuiasaurus 8986: 8978: 8971: 8951: 8946:Titanomachya 8944: 8937: 8915:Puertasaurus 8913: 8907: 8906: 8901:Notocolossus 8899: 8892: 8886:Jiangxititan 8884: 8877: 8869: 8864:Drusilasaura 8862: 8855: 8848:Lognkosauria 8831: 8824: 8817: 8810: 8803: 8796: 8789: 8782:Aeolosaurini 8771:Rinconsaurus 8769: 8762: 8755: 8748: 8740: 8733: 8722:Rinconsauria 8709: 8703:Tengrisaurus 8701: 8696:Quetecsaurus 8694: 8687: 8679: 8672: 8665: 8658: 8651: 8627: 8620: 8586: 8579: 8574:Lohuecotitan 8572: 8565: 8560:Garrigatitan 8558: 8551: 8546:Ampelosaurus 8544: 8519: 8512: 8505: 8500:Uberabatitan 8498: 8493:Sonidosaurus 8491: 8484: 8477: 8470: 8464:Tengrisaurus 8462: 8454: 8447: 8440: 8435:Rapetosaurus 8433: 8426: 8419: 8411: 8404: 8397: 8390: 8383: 8376: 8371:Malawisaurus 8369: 8361: 8354: 8346: 8339: 8332: 8325: 8317: 8312:Huabeisaurus 8310: 8303: 8296: 8288: 8281: 8273: 8266: 8259: 8254:Choconsaurus 8252: 8247:Brasilotitan 8245: 8238: 8230: 8222: 8215: 8208: 8201: 8196:Atacamatitan 8194: 8189:Argyrosaurus 8187: 8180: 8172: 8165: 8157: 8152:Abdarainurus 8150: 8134:Titanosauria 8121:Titanosauria 8090: 8049:Titanosauria 8033:Wintonotitan 8031: 8024: 8017: 8010: 8003: 7979: 7974:Yongjinglong 7972: 7965: 7960:Tambatitanis 7958: 7951: 7944: 7937: 7929: 7924:Huanghetitan 7922: 7917:Huabeisaurus 7915: 7908: 7903:Gannansaurus 7901: 7894: 7887: 7881:Chiayusaurus 7879: 7871: 7851: 7844: 7837: 7830: 7823: 7816: 7809: 7802: 7795: 7787: 7780: 7775:Jiutaisaurus 7773: 7767:Jiangxititan 7765: 7760:Garumbatitan 7758: 7751: 7744: 7739:Dongbeititan 7737: 7730: 7723: 7716: 7711:Austrosaurus 7709: 7702: 7695: 7688: 7664: 7659:Venenosaurus 7657: 7650: 7645:Sonorasaurus 7643: 7636: 7631:Giraffatitan 7629: 7621: 7616:Cedarosaurus 7614: 7607: 7600: 7593: 7572: 7567:Pelorosaurus 7565: 7558: 7551: 7544: 7537: 7530: 7523: 7493: 7488:Camarasaurus 7486: 7466: 7460:Haestasaurus 7458: 7453:Europasaurus 7451: 7443: 7436: 7428: 7421: 7414: 7329: 7322: 7315: 7307: 7300: 7293: 7286: 7279: 7272:Diplodocinae 7261:Brontosaurus 7259: 7251: 7244: 7238:Amphicoelias 7236: 7220:Diplodocidae 7207: 7200: 7193: 7186: 7179: 7172: 7164: 7157: 7150: 7145:Bajadasaurus 7143: 7136: 7131:Amargasaurus 7129: 7092: 7085: 7078: 7071: 7064: 7057: 7038:Rayososaurus 7036: 7029: 7022: 6995:Zapalasaurus 6993: 6988:Xenoposeidon 6986: 6979: 6972: 6964: 6957: 6950: 6942: 6935: 6930:Campananeyen 6928: 6923:Amazonsaurus 6921: 6914: 6879: 6820: 6790: 6783: 6776: 6769: 6762: 6755: 6748: 6686:Neosauropoda 6673: 6666: 6659: 6651: 6643: 6636: 6629: 6622: 6615: 6607: 6601:Haestasaurus 6599: 6591: 6584: 6564: 6557: 6550: 6543: 6536: 6529: 6522: 6515: 6508: 6501: 6493: 6486: 6479: 6472: 6465: 6457: 6450: 6442: 6435: 6428: 6409:Patagosaurus 6407: 6399: 6391: 6384: 6377: 6359:Volkheimeria 6357: 6350: 6343: 6336: 6331:Perijasaurus 6329: 6324:Nebulasaurus 6322: 6315: 6308: 6303:Dystrophaeus 6301: 6294: 6287: 6280: 6275:Barapasaurus 6273: 6266: 6261:Asiatosaurus 6259: 6252: 6222: 6215: 6194: 6186: 6181:Rhoetosaurus 6179: 6172: 6165: 6157: 6125:Lessemsaurus 6123: 6116: 6109: 6104:Antetonitrus 6102: 6081: 6074: 6067: 6059: 6051: 6044: 6037: 6030: 6023: 6016: 5953: 5889: 5882: 5875: 5868: 5861: 5854: 5847: 5840: 5833: 5826: 5819: 5795: 5790:Yizhousaurus 5788: 5780: 5773: 5766: 5759: 5751: 5726: 5719: 5712: 5705: 5698: 5691: 5684: 5678:Ignavusaurus 5676: 5669: 5662: 5655: 5635: 5628: 5608: 5601: 5594: 5587: 5582:Ignavusaurus 5580: 5573: 5566: 5538:Yimenosaurus 5536: 5531:Plateosaurus 5529: 5522: 5515: 5508: 5501: 5483:Xixiposaurus 5481: 5474: 5467: 5455:Plateosauria 5437: 5430: 5423: 5416:Unaysauridae 5403: 5396: 5389: 5382: 5375: 5368: 5361: 5327: 5319: 5311: 5304: 5297: 5289: 5281: 5261: 5254: 5247: 5242:Guaibasaurus 5240: 5233: 5226: 5219: 5105: 5096: 4983:. Retrieved 4979: 4970: 4958:. Retrieved 4954: 4945: 4928: 4922: 4914: 4879: 4875: 4869: 4844: 4840: 4836: 4830: 4805:11336/103474 4779: 4775: 4769: 4760: 4756: 4746: 4729:11336/117578 4711: 4708:IAWA Journal 4707: 4703: 4697: 4680:11336/127834 4654: 4650: 4640: 4628:. Retrieved 4624: 4614: 4602:. Retrieved 4598: 4588: 4555: 4551: 4547: 4541: 4519:(1): 35–51. 4516: 4512: 4506: 4473: 4469: 4463: 4430: 4426: 4419: 4394: 4390: 4386: 4380: 4337: 4333: 4323: 4298:11336/106658 4280: 4276: 4269: 4242: 4238: 4234: 4228: 4201: 4182:11336/192915 4165:(1): 45–60. 4162: 4158: 4152: 4144: 4117: 4113: 4103: 4070: 4066: 4059: 4008: 4004: 3946: 3942: 3932: 3889: 3885: 3872: 3860:. Retrieved 3856: 3850: 3842: 3830:. Retrieved 3826: 3820: 3816: 3808: 3796:. Retrieved 3792: 3782: 3739: 3735: 3725: 3700: 3696: 3666:. Retrieved 3661: 3637:. Retrieved 3633: 3606:. Retrieved 3602: 3593: 3581:. Retrieved 3577:The Guardian 3575: 3565: 3553:. Retrieved 3549: 3539: 3527:. Retrieved 3523:the original 3518: 3508: 3496:. Retrieved 3492: 3482: 3470:. Retrieved 3466: 3456: 3423: 3419: 3413: 3359: 3355: 3297: 3293: 3287: 3281: 3269: 3259: 3247: 3243:Tyrannotitan 3241: 3237: 3229: 3215: 3210: 3204: 3198: 3192: 3186: 3162: 3158: 3157: 3151: 3142:Paleoecology 3063: 3062: 3061: 3045: 3044: 3021: 3020: 3009:Lognkosauria 2967: 2966: 2950: 2949: 2938:Aeolosaurini 2923: 2922: 2901:Rinconsaurus 2899: 2898: 2887:Rinconsauria 2865: 2864: 2820: 2819: 2800: 2799: 2770: 2769: 2733: 2732: 2706: 2705: 2694:Titanosauria 2683: 2679: 2675: 2672:Malawisaurus 2671: 2666: 2664: 2659: 2656:Puertasaurus 2655: 2651: 2647: 2641: 2638:Drusilasaura 2637: 2634:Puertasaurus 2633: 2629: 2625: 2621: 2617: 2613: 2610:Notocolossus 2609: 2606:Puertasaurus 2605: 2601: 2592: 2589:Notocolossus 2588: 2585:Puertasaurus 2584: 2576: 2573:Puertasaurus 2572: 2568: 2564: 2562: 2449: 2448: 2447: 2431: 2430: 2409:Drusilasaura 2407: 2406: 2392:Puertasaurus 2390: 2389: 2359:Quetecsaurus 2357: 2356: 2340: 2339: 2309: 2308: 2297:Lognkosauria 2284:Notocolossus 2282: 2281: 2258: 2257: 2207: 2206: 2190: 2189: 2166: 2165: 2144:Rinconsaurus 2142: 2141: 2130:Rinconsauria 2056:Tapuiasaurus 2054: 2053: 2037: 2036: 2015:Rapetosaurus 2013: 2012: 1989: 1988: 1972: 1971: 1955: 1954: 1938: 1937: 1921: 1920: 1904: 1903: 1880: 1879: 1858:Malawisaurus 1856: 1855: 1844:Lithostrotia 1829: 1828: 1795: 1794: 1762: 1761: 1745: 1744: 1714: 1713: 1699:Wintonotitan 1697: 1696: 1685:Titanosauria 1667:Rinconsauria 1662: 1659:Notocolossus 1658: 1654: 1650: 1646:Notocolossus 1645: 1641: 1637: 1634:sister group 1629: 1623: 1600: 1597:Rapetosaurus 1596: 1592: 1588: 1584: 1580: 1577:Rapetosaurus 1576: 1572: 1564: 1560: 1554: 1548: 1544: 1540: 1536:Venenosaurus 1534: 1528: 1522: 1518: 1514: 1510: 1502:Uberabatitan 1500: 1496: 1492: 1489:Malawisaurus 1488: 1484: 1478: 1472: 1468: 1464: 1462: 1449:Haestasaurus 1447: 1444:Giraffatitan 1443: 1439: 1431: 1427: 1423: 1419: 1415: 1411: 1407: 1403: 1399: 1371: 1367: 1363: 1359: 1355:Rapetosaurus 1353: 1349:Notocolossus 1347: 1343: 1341: 1334: 1330: 1320: 1316: 1312:Tapuiasaurus 1310: 1304: 1300:Giraffatitan 1298: 1294: 1288: 1282: 1279:Drusilasaura 1278: 1274: 1268: 1248: 1238: 1232: 1226: 1223:Drusilasaura 1222: 1218: 1217: 1199:Drusilasaura 1197: 1193: 1189: 1177: 1174:Diplodocidae 1169: 1165: 1161: 1155: 1143:Malawisaurus 1141: 1135: 1131: 1127: 1117: 1104: 1083:Lognkosauria 1079:Puertasaurus 1078: 1072: 1068: 1060: 1056: 1027:Puertasaurus 1025: 1021: 1017: 1005: 1001: 997: 995: 988: 982: 976: 972: 952: 950: 932: 928: 924: 920: 916: 912: 905: 901: 897: 893: 885: 883: 874: 870: 860: 856: 848: 846: 842: 828: 826: 820: 794: 793:specimen of 780: 771: 769: 736:Titanosauria 731: 727: 724:type species 697: 650: 604:, six front 582: 576: 515: 497: 436: 433:Lognkosauria 428: 426: 413: 409:Puertasaurus 407: 401: 397: 381: 368: 367: 340: 339: 338: 329: 322: 321: 312:Type species 302: 295: 294: 282:Lognkosauria 272: 266:Titanosauria 256: 240: 224: 208: 195: 182: 36:101.62  24: 18: 9504:Albian life 9429:Patagotitan 9274:Microcoelus 9182:Saltasaurus 9168:Paralititan 9064:Alamosaurus 8953:Udelartitan 8908:Patagotitan 8805:Bravasaurus 8798:Arrudatitan 8791:Aeolosaurus 8757:Overosaurus 8689:Jainosaurus 8667:Bonitasaura 8588:Paludititan 8479:Traukutitan 8174:Angolatitan 7981:Yunmenglong 7946:Qiaowanlong 7839:Sibirotitan 7804:Oceanotitan 7746:Europatitan 7725:Brontomerus 7718:Brohisaurus 7690:Angolatitan 7623:Galvesaurus 7595:Abydosaurus 7560:Ornithopsis 7553:Fusuisaurus 7539:Eucamerotus 7438:Daanosaurus 7423:Aragosaurus 7324:Supersaurus 7246:Apatosaurus 7209:Tharosaurus 7080:Nigersaurus 7031:Limaysaurus 6792:Ultrasaurus 6668:Turiasaurus 6631:Moabosaurus 6624:Mierasaurus 6593:Bellusaurus 6577:Turiasauria 6517:Qijianglong 6481:Hudiesaurus 6459:Datousaurus 6379:Cetiosaurus 6338:Shunosaurus 6282:Bellusaurus 6254:Algoasaurus 6240:Eusauropoda 6188:Sanpasaurus 6141:Gravisauria 6061:Pulanesaura 6046:Isanosaurus 6018:Amygdalodon 5884:Pulanesaura 5842:Irisosaurus 5828:Anchisaurus 5808:Anchisauria 5782:Xingxiulong 5728:Xingxiulong 5721:Sarahsaurus 5637:Riojasaurus 5603:Sarahsaurus 5589:Kholumolumo 5432:Macrocollum 5370:Asylosaurus 5306:Nhandumirim 5283:Buriolestes 5249:Mbiresaurus 5221:Buriolestes 4980:BBC iPlayer 4905:11336/90736 4657:: 133–148. 4513:Ameghiniana 3851:Patagotitan 3817:Patagotitan 3793:Smithsonian 3583:26 November 3294:Patagotitan 3254:peirosaurid 3249:Genyodectes 3238:Patagotitan 3230:Patagotitan 3211:C. australe 3179:angiosperms 3163:Patagotitan 3159:Patagotitan 3152:Patagotitan 3064:Patagotitan 2952:Aeolosaurus 2822:Saltasaurus 2684:Saltasaurus 2676:Patagotitan 2667:Patagotitan 2648:Patagotitan 2626:Patagotitan 2602:Patagotitan 2577:Patagotitan 2565:Patagotitan 2450:Patagotitan 2260:Bonitasaura 2209:Overosaurus 2192:Aeolosaurus 1974:Saltasaurus 1940:Alamosaurus 1663:Bonitasaura 1655:Alamosaurus 1642:Patagotitan 1630:Patagotitan 1624:In 2017, a 1601:Saltasaurus 1589:Bonitasaura 1581:Alamosaurus 1565:Saltasaurus 1545:Patagotitan 1541:Patagotitan 1519:Alamosaurus 1517:but unlike 1493:Alamosaurus 1480:Qiaowanlong 1432:Saltasaurus 1420:Patagotitan 1416:Saltasaurus 1372:Patagotitan 1344:Patagotitan 1295:Patagotitan 1270:Alamosaurus 1240:Saltasaurus 1219:Patagotitan 1178:Patagotitan 1168:but unlike 1157:Bonitasaura 1128:Patagotitan 1105:Patagotitan 1069:Patagotitan 1057:Patagotitan 1050:diapophyses 1038:, from the 1018:Patagotitan 1006:Patagotitan 1002:Patagotitan 973:Patagotitan 967:pleurocoels 953:Patagotitan 929:Patagotitan 925:Patagotitan 921:Patagotitan 902:Patagotitan 894:Patagotitan 886:Patagotitan 871:Patagotitan 857:Patagotitan 849:Patagotitan 829:Patagotitan 807:Description 772:Patagotitan 732:Patagotitan 644:pubic bones 577:Patagotitan 560:age of the 498:Patagotitan 437:Patagotitan 414:Patagotitan 398:Patagotitan 382:Patagotitan 341:Patagotitan 328:Carballido 301:Carballido 296:Patagotitan 25:Patagotitan 9488:Categories 9175:Rocasaurus 9102:Qunkasaura 9071:Baurutitan 8833:Shingopana 8660:Baalsaurus 8507:Volgatitan 8442:Rukwatitan 8406:Nullotitan 8392:Ninjatitan 8327:Kaijutitan 8298:Gandititan 8261:Daxiatitan 8224:Baurutitan 8167:Andesaurus 8058:see below↓ 7753:Fukuititan 7697:Arkharavia 7652:Soriatitan 7602:Atlasaurus 7574:Rugocaudia 7532:Duriatitan 7468:Yuzhoulong 7416:Abrosaurus 7398:Macronaria 7385:Macronaria 7309:Kaatedocus 7302:Galeamopus 7295:Diplodocus 7281:Barosaurus 7188:Pilmatueia 7181:Lingwulong 7174:Kaatedocus 7094:Tataouinea 7006:Khebbashia 6981:Sidersaura 6711:Macronaria 6661:Tendaguria 6645:Oplosaurus 6609:Janenschia 6510:Omeisaurus 6386:Chebsaurus 6310:Janenschia 6224:Vulcanodon 6167:Kotasaurus 6118:Ledumahadi 5911:see below↓ 5870:Meroktenos 5686:Leyesaurus 5575:Gyposaurus 5439:Unaysaurus 5398:Pantydraco 5363:Arcusaurus 5329:Saturnalia 5162:see below↓ 5110:Dinosauria 5092:Sauropsida 4923:Chubutemys 4558:: 105389. 4476:: 104754. 4433:: 105487. 3316:References 3271:Chubutemys 3266:cryptodire 3264:, and the 3234:floodplain 2708:Andesaurus 2682:) but not 2660:Ninjatitan 2643:Ninjatitan 2579:to form a 1882:Baurutitan 1716:Andesaurus 1616:inserted. 1550:Rocasaurus 1524:Andesaurus 1245:Macronaria 1122:procoelous 1090:hyposphene 890:Matt Wedel 869:estimated 865:. In 2019 821:P. mayorum 782:Barosaurus 766:, New York 562:Cretaceous 502:floodplain 394:short tons 250:Macronaria 202:Saurischia 189:Dinosauria 9444:Q17103736 9409:Argentina 9397:Dinosaurs 9138:Bonatitan 9020:Isisaurus 8826:Punatitan 8305:Hamititan 8283:Elaltitan 8232:Bonatitan 7953:Silutitan 7910:Gobititan 7896:Euhelopus 7638:Lusotitan 7331:Tornieria 7317:Leinkupal 7202:Suuwassea 6916:Agustinia 6764:Dinodocus 6757:Cardiodon 6488:Jingiella 6437:Anhuilong 6000:Sauropoda 5987:Sauropoda 5902:Sauropoda 5877:Mussaurus 5835:Camelotia 5714:Pradhania 5554:Massopoda 5321:Panphagia 5263:Panphagia 5078:Kingdom: 4937:2246/5895 4861:131178611 4689:135268573 4580:252799392 4533:226680080 4498:233582290 4455:256559829 4411:213155599 4315:198493761 4095:130423764 4033:2045-2322 4011:: 19165. 3902:CiteSeerX 3862:8 January 3832:8 January 3798:8 January 3774:221404013 3758:1469-185X 3717:210840060 3639:8 January 3448:225012747 3226:siltstone 3224:-bearing 3218:sandstone 2039:Isisaurus 1511:Elaltitan 1497:Elaltitan 1485:Isisaurus 1440:Elaltitan 1404:Isisaurus 1325:than the 1284:Elaltitan 1249:Isisaurus 1228:Isisaurus 1098:hypantrum 1020:, unlike 939:Vertebrae 879:Histology 740:Patagonia 653:paratypes 526:La Flecha 512:Discovery 412:. Still, 364:Patagonia 354:from the 234:Sauropoda 162:Kingdom: 156:Eukaryota 9458:47458376 9438:Wikidata 9146:Ibirania 8812:Caieiria 8622:Inawentu 7825:Ruixinia 7666:Vouivria 7525:Astrodon 6778:Neosodon 6268:Bagualia 6111:Ingentia 5821:Aardonyx 5768:Qianlong 5596:Musankwa 5568:Gryponyx 5476:Ruehleia 5391:Nambalia 5384:Efraasia 5291:Eoraptor 5228:Eoraptor 5086:Chordata 5084:Phylum: 5080:Animalia 4822:24015414 4814:15834691 4738:92383900 4599:ABC News 4372:31598266 4307:31340217 4136:17768539 4051:26777391 3983:24348896 3943:PLOS ONE 3924:56028251 3766:32869488 3662:BBC News 3634:BBC News 3386:28794222 3304:See also 3188:Fitzroya 3175:conifers 3167:pinnules 2581:polytomy 1255:acromion 1186:spinalis 754:Exhibits 616:chevrons 612:, three 550:theropod 546:sauropod 459:scapulae 384:was the 352:dinosaur 349:sauropod 176:Chordata 172:Phylum: 166:Animalia 152:Domain: 9383:Portals 8182:Arackar 6586:Amanzia 6430:Analong 6317:Jobaria 5775:Seitaad 5090:Class: 5008:Ed Yong 4884:Bibcode 4784:Bibcode 4659:Bibcode 4560:Bibcode 4478:Bibcode 4435:Bibcode 4363:6731702 4342:Bibcode 4075:Bibcode 4042:4725985 4013:Bibcode 3974:3864407 3951:Bibcode 3894:Bibcode 3668:24 July 3555:24 July 3428:Bibcode 3377:5563814 3268:turtle 3183:cypress 3154:at dawn 2591:, with 1649:tons). 1632:as the 1499:. Like 1263:glenoid 1012:laminae 691:fibulae 671:, both 630:sternum 378:in 2017 374:in 2014 288:Genus: 133:, U.S. 105:↓ 9471:358623 8711:Vahiny 8472:Tiamat 7889:Erketu 5707:Ngwevu 5010:, 2017 4985:4 July 4960:4 July 4859:  4820:  4812:  4736:  4687:  4630:17 May 4604:17 May 4578:  4531:  4496:  4453:  4409:  4370:  4360:  4313:  4305:  4216:  4134:  4093:  4049:  4039:  4031:  3981:  3971:  3922:  3904:  3772:  3764:  3756:  3715:  3608:4 July 3529:2 July 3498:2 July 3472:3 July 3446:  3384:  3374:  3252:, the 2608:, and 1661:, and 1603:, the 1599:, and 1563:, and 1533:, and 1477:, and 1414:, and 1366:, and 1309:, and 1287:, and 1237:, and 1196:, and 1150:centra 1140:, and 987:, and 852:'s 683:humeri 675:ischia 667:radius 596:, six 558:Albian 530:Trelew 496:life, 483:femora 475:ischia 467:humeri 390:tonnes 332:, 2017 330:et al. 305:, 2017 303:et al. 31:Albian 5106:Clade 5097:Clade 4857:S2CID 4818:S2CID 4734:S2CID 4685:S2CID 4576:S2CID 4529:S2CID 4494:S2CID 4451:S2CID 4407:S2CID 4311:S2CID 4091:S2CID 3920:S2CID 3882:(PDF) 3770:S2CID 3713:S2CID 3693:(PDF) 3444:S2CID 3171:ferns 3169:from 2654:than 2583:with 1459:Femur 1378:fossa 996:Like 909:' 744:Titan 579:bones 575:with 520:femur 427:Like 273:Clade 257:Clade 241:Clade 225:Clade 209:Clade 196:Clade 183:Clade 9013:Igai 5517:Issi 5139:see 4987:2023 4962:2023 4810:PMID 4632:2014 4606:2014 4368:PMID 4303:PMID 4214:ISBN 4132:PMID 4047:PMID 4029:ISSN 3979:PMID 3890:2004 3864:2019 3834:2019 3800:2019 3762:PMID 3754:ISSN 3670:2022 3641:2019 3610:2023 3585:2022 3557:2022 3531:2022 3500:2022 3474:2022 3382:PMID 3246:and 3222:tuff 3220:and 3191:and 2650:and 2636:and 2587:and 2571:and 1653:and 1575:and 1513:and 1495:and 1446:and 1430:and 1352:and 1333:and 1319:and 1164:and 1024:and 904:and 837:tail 835:and 833:neck 812:Size 718:and 663:and 659:ulna 624:ribs 479:and 463:and 447:and 443:back 406:and 392:(76 44:Preęž’ 9453:EoL 6675:Zby 5006:by 4955:BBC 4933:hdl 4900:hdl 4892:doi 4849:doi 4800:hdl 4792:doi 4724:hdl 4716:doi 4675:hdl 4667:doi 4568:doi 4556:142 4521:doi 4486:doi 4474:122 4443:doi 4431:146 4399:doi 4358:PMC 4350:doi 4293:hdl 4285:doi 4255:hdl 4247:doi 4243:184 4206:doi 4177:hdl 4167:doi 4122:doi 4083:doi 4037:PMC 4021:doi 3969:PMC 3959:doi 3912:doi 3744:doi 3705:doi 3493:PBS 3436:doi 3372:PMC 3364:doi 3360:284 791:Sue 564:). 358:in 9490:: 9468:: 9455:: 9440:: 5108:: 5099:: 4978:. 4953:. 4927:. 4898:. 4890:. 4880:30 4878:. 4855:. 4845:24 4843:. 4816:. 4808:. 4798:. 4790:. 4780:92 4778:. 4761:23 4759:. 4755:. 4732:. 4722:. 4712:39 4710:. 4683:. 4673:. 4665:. 4655:99 4653:. 4649:. 4623:. 4597:. 4574:. 4566:. 4554:. 4527:. 4517:58 4515:. 4492:. 4484:. 4472:. 4449:. 4441:. 4429:. 4405:. 4395:18 4393:. 4366:. 4356:. 4348:. 4336:. 4332:. 4309:. 4301:. 4291:. 4281:91 4279:. 4253:. 4241:. 4212:. 4191:^ 4175:. 4163:56 4161:. 4157:. 4130:. 4118:79 4116:. 4112:. 4089:. 4081:. 4071:24 4069:. 4045:. 4035:. 4027:. 4019:. 4007:. 4003:. 3991:^ 3977:. 3967:. 3957:. 3945:. 3941:. 3918:. 3910:. 3900:. 3888:. 3884:. 3855:. 3825:. 3791:. 3768:. 3760:. 3752:. 3740:95 3738:. 3734:. 3711:. 3701:85 3699:. 3695:. 3678:^ 3660:. 3649:^ 3632:. 3618:^ 3601:. 3574:. 3548:. 3517:. 3491:. 3465:. 3442:. 3434:. 3424:40 3422:. 3418:. 3394:^ 3380:. 3370:. 3358:. 3354:. 3324:^ 3300:. 3286:, 3274:. 3209:, 3197:, 2620:, 2616:, 2604:, 1595:, 1591:, 1583:, 1559:, 1527:, 1487:, 1471:, 1467:, 1410:, 1406:, 1402:, 1362:, 1339:. 1303:, 1281:, 1277:, 1273:, 1231:, 1225:, 1192:, 1134:, 1077:, 981:, 803:. 714:, 710:, 706:, 702:, 620:, 508:. 455:, 435:, 362:, 275:: 259:: 243:: 227:: 211:: 198:: 185:: 129:, 94:Pg 38:Ma 9385:: 9284:? 9269:? 9226:? 9156:? 9141:? 9133:? 9112:? 9097:? 9082:? 9074:? 9059:? 9038:? 9023:? 8991:? 8983:? 8889:? 8874:? 8745:? 8706:? 8684:? 8524:? 8467:? 8459:? 8416:? 8366:? 8351:? 8322:? 8293:? 8278:? 8235:? 8227:? 8177:? 8162:? 8036:? 7934:? 7884:? 7876:? 7792:? 7770:? 7626:? 7577:? 7463:? 7448:? 7433:? 7312:? 7256:? 7241:? 7169:? 7041:? 6969:? 6947:? 6656:? 6648:? 6612:? 6604:? 6596:? 6498:? 6462:? 6447:? 6412:? 6404:? 6396:? 6199:? 6191:? 6162:? 6086:? 6064:? 6056:? 5785:? 5756:? 5731:? 5681:? 5324:? 5316:? 5294:? 5286:? 5054:e 5047:t 5040:v 4989:. 4964:. 4939:. 4935:: 4921:" 4908:. 4902:: 4894:: 4886:: 4863:. 4851:: 4824:. 4802:: 4794:: 4786:: 4740:. 4726:: 4718:: 4691:. 4677:: 4669:: 4661:: 4634:. 4608:. 4582:. 4570:: 4562:: 4535:. 4523:: 4500:. 4488:: 4480:: 4457:. 4445:: 4437:: 4413:. 4401:: 4374:. 4352:: 4344:: 4338:6 4317:. 4295:: 4287:: 4263:. 4257:: 4249:: 4222:. 4208:: 4185:. 4179:: 4169:: 4155:" 4138:. 4124:: 4097:. 4085:: 4077:: 4053:. 4023:: 4015:: 4009:6 3985:. 3961:: 3953:: 3947:8 3926:. 3914:: 3896:: 3866:. 3853:" 3836:. 3823:" 3802:. 3776:. 3746:: 3719:. 3707:: 3672:. 3643:. 3612:. 3587:. 3559:. 3533:. 3502:. 3476:. 3450:. 3438:: 3430:: 3388:. 3366:: 1442:( 319:† 292:† 279:† 263:† 247:† 231:† 215:† 99:N 89:K 84:J 79:T 74:P 69:C 64:D 59:S 54:O 49:ęž’

Index

Albian
Ma
Preęž’
ęž’
O
S
D
C
P
T
J
K
Pg
N

Field Museum of Natural History
Chicago, Illinois
Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Dinosauria
Saurischia
Sauropodomorpha
Sauropoda
Macronaria
Titanosauria
Lognkosauria
Patagotitan

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