872:
4693:
4700:
4713:
1090:
962:
4720:
1327:
4706:
130:
1805:
1706:
719:
417:
108:
620:
1031:, and the second was significantly enlarged indicating a caniniform larger than the one in the maxilla. The regular dentary teeth were slightly taller than those of the maxilla and bore denticles along the top half of the crown instead of the top 25%. As in the maxilla, the crowns has a medial bulge, eight to ten denticles on either side of the tip, and symmetrical
887:, preserving almost all the bone including most regions of contact with other cranial bones. The maxillae are slightly flattened, which minimises the strength of the buccal emargination, a diagnostic character of ornithischians where the maxillary tooth row and bone directly above it is inset from the outer edge of the bone. The presence of an arched
1781:. The "Upper Purbeck Beds" and a majority of the "Middle Purbeck Beds" are contained within the Durlston Formation, the oldest deposit of which is the Cinder Beds of the Stair Hole Member. The Cinder Beds have, at times, been considered the Jurassic-Cretaceous Boundary, which would result in the entire Lulworth Formation being latest Jurassic,
933:. Owen also figured a partial tooth in front of the caniniform, but it has since been lost in damage to the material. While Norman and Barrett used this as evidence for a second caniniform smaller than the one behind, Galton in 1978 and Sereno in 2012 identified only one canine present. The first post-caniniform tooth of
1687:
compared to more primitive heterodontosaurids, the function of the enlarged canines was likely the same across the clade. Based on the presence of wear along the tips of the premaxillary crown and the movement of the dentary caniniform relative to other teeth as the jaw closed, according to Sereno in
1014:
process is prominent, unlike other basal ornithischians, although the contact between the dentaries is V-shaped as in basal ornithischians. There were 11 teeth in the dentary, the first two of which were specialised compared to most ornithischians. While specialised anterior teeth were not discussed
1611:
and a prominent anterior groove leading to a foramen on the lateral surface of the dentary. All but the
Kayenta heterodontosaurid are also significantly younger than the remaining heterodontosaurids, and the presence of the three united in a clade excluding other taxa implied an unknown lineage of
1829:
There is a great deal of uncertainty as to the location of the specimens collected from the
Lulworth Formation; the only definitive way to test would be to analyse the matrix of each specimen to determine its salinity. The Purbeck Group has the most diverse ornithischian fauna of any deposit in
937:
is the largest, although only slightly taller than those following, which are all the same size. There is a round prominence along the middle of the tooth crowns, but there are no prominent ridges present on the crown. Eight to ten denticles are present on each side of the tooth crowns.
1157:. Galton published a rebuttal the following year, disagreeing with Thulborn's use of Hypsilophodontidae. As teeth in the premaxilla, one of Thulborn's diagnostic traits for the family, were a primitive feature, Galton argued they should not be used to classify ornithopods. Instead,
1692:
in 2008 that the enlarged canines of the clade were for an omnivorous diet. The hypothesis of omnivory was also supported by Norman and colleagues in 2011 under the interpretation that the canines and premaxillary teeth lacked wear from cropping vegetation. The
1785:, in age. However, despite the uncertainties about the age of the beds because of a lack of correlation through fauna or dating, it is generally accepted that the Purbeck Group is entirely earliest Cretaceous in age, with the Lulworth Formation being early
997:
processes. The dentary was deep for a basal ornithischian, its height at mid-length being 30% of the total length. The bone tapers anteriorly along its length, although the margins are subparallel under the middle of the tooth row. A row of
867:
which bears teeth along its entire length. The premaxilla bore only three teeth, lacking denticles (serrations) present in most basal ornithischians, although all teeth are subequal in size with the third being the largest.
957:
is partially complete and preserved in articulation with the maxilla of NHMUK 48210. Lack of preservation limits the anatomical details that can be identified beyond its resembling the bone known in other ornithischians.
1248:
in 2002 and supported the heterodontosaurid classification but instead referred it to the clade based on the lacking foramina on tooth-bearing bones and possessing denticles restricted to the top third of the crown.
989:, but its presence can be confirmed by the morphology of the anterior end of the known dentaries. Like in other heterodontosaurids, the predentary was only loosely articulated with the dentary in life, lacking
1256:
has been supported in further analyses including some of the first phylogenetic analyses of ornithischians, the placement of the family itself has changed over time. The family, either excluding
2373:
Catalogue of Fossil
Reptilia and Amphibia in the British Museum (Natural History). Part I. Containing the Orders Ornithosauria, Crocodilia, Dinosauria, Squamata, Rhynchocephalia and Pterosauria
527:, Owen chose to use the abbreviated form as the name for his new animal. Falconer issued a correction in 1861, specifying that he proposed the name "Sauraechmodon" instead of "Sauraechinodon".
2653:
Vertebrate paleobiodiversity of the Early
Cretaceous (Berriasian) Angeac-Charente Lagerstätte (southwestern France): implications for continental faunal turnover at the J/K boundary
1071:
based on a tall coronoid process and tapering teeth, both of which are considered widespread within ornithischians. Isolated dermal armour found in the
Purbeck Beds was referred to
492:. While the specific name honoured Beckles for his discovery of the fossils and allowing Owen to study his collection of Purbeck fossils, the generic name was derived from the
3706:
Averianov, A.O.; Martin, T.; Lopatin, A.V. (2013). "A new phylogeny for basal
Trechnotheria and Cladotheria and affinities of South American endemic Late Cretaceous mammals".
1311:
below follows the analysis by Sereno in 2012, including the validly named heterodontosaurids described at the time. The clade including the Late
Jurassic and Early Cretaceous
3836:
Butler, P.M.; Sigogneau-Russell, D.; Ensom, P.C. (2012). "Possible persistence of the morganucodontans in the Lower
Cretaceous Purbeck Limestone Group (Dorset, England)".
2744:
Barrett, P.M.; Maidment, S.C.R. (2011). "Dinosaurs of Dorset: Part III, the ornithischian dinosaurs (Dinosauria, Ornithischia) with additional comments on the sauropods".
1220:, in fact, were interpreted to possess flat wear facets on its teeth, as well as having teeth inset from the edge of the maxilla like in ornithischians more derived than
1773:
and became wetter towards the end of the
Berriasian. While the Purbeck Group was originally known as the informal Purbeck Beds, it can now be divided into the upper
1304:
considered that the placement outside
Heterodontosauridae was not because it was not within the family but instead because of the incomplete nature of its remains.
3546:
Sigogneau-Russell, D. (1999). "Réévaluation des Peramura (Mammalia, Cladotheria) sur la base de nouveaux spécimens du Crétacé inférieur d'Angleterre et du Maroc".
384:, which has been variably considered to be from the latest Jurassic or the earliest Cretaceous. Current studies accept an Early Cretaceous Berriasian age, making
1201:, known to possess enlarged canine teeth as well. However, Galton considered a fabrosaurid identity more likely as the teeth of the taxon lack flat wear facets.
2784:"Morphology, histology and identification of the 'granicones' from the Purbeck Limestone Formation (Lower Cretaceous: Berriasian) of Dorset, southern England"
572:
although not referring them to a more specific clade than Dinosauria indeterminate. The series of specimens designated as types by Owen were purchased by the
4946:
661:, were approximately 150.2-150.3 million years old, and were initially described as intermediate fabrosaur remains by their collector George Calliston in
3459:
3053:
Pol, D.; Rauhut, O.W.M.; Becerra, M. (2011). "A Middle Jurassic heterodontosaurid dinosaur from Patagonia and the evolution of heterodontosaurids".
2231:
1027:
as differing in size from following teeth in 2012. The first alveolus was extremely reduced, indicating a small peg-like first dentary tooth as in
3759:
Kielan-Jaworowska, Z.; Ensom, P.C. (1994). "Tiny plagiaulacoid multituberculate mammals from the Purbeck Limestone Formation of Dorset, England".
3014:
Butler, R.J.; Liyong, J.; Jun, C.; Godefroit, P. (2011). "The postcranial osteology and phylogenetic position of the small ornithischian dinosaur
5016:
3486:
Evans, S.E. (1998). "Lepidosaurian faunas from the Early Cretaceous: A clade in transition". In Lucas, S.G.; Kirkland, J.I.; Estep, J.W. (eds.).
2241:
1688:
2012, the primary function of the premaxillary tooth row would be for cropping vegetation. This contrasts with the earlier hypotheses of Butler
1171:
based on laterally positioned teeth, and removed from Hypsilophodontidae. Galton followed up on this classification with the naming of the genus
911:
concluded in 2002 that, based on the images of Owen prior to later damage of the fossil NHMUK 48209, a diastema was absent. Sereno reiterated in
3778:
Kielan-Jaworowska, Z.; Ensom, P.C. (1994). "Multituberculate mammals from the Upper Jurassic Purbeck Limestone Formation of southern England".
3225:
Lukashevich, E.D.; Coram, R.A.; Jarzembowski, E.A. (2001). "New true flies (Insecta: Diptera) from the Lower Cretaceous of southern England".
5080:
2679:
322:
and many other taxa from across England, while the genus name translates as "prickly tooth" in reference to the dental anatomy of the taxon.
3648:
Ensom, P.C.; Sigogneau-Russell, D. (1998). "New dryolestoid mammals from the Basal Cretaceous Purbeck Limestone Group of southern England".
2225:
638:
1990:
3457:
Evans, S.E.; Jones, M.E.H.; Matsumoto, R. (2012). "A new lizard skull from the Purbeck Limestone Group (Lower Cretaceous) of England".
2247:
1260:
as in Thulborn's 1971 study, or including the genus as in Galton's and Sereno's work, was originally considered to be a group of basal
3151:
1840:
is the only other named ornithischian from the beds and is known only from a dentary with teeth originally described as a species of
616:, a fragmentary dentary with teeth, and the only further referrals to the genus includes isolated teeth also from the Purbeck beds.
5110:
5003:
2402:
Salisbury, S.W. (2002). "Crocodilians from the Lower Cretaceous (Berriasian) Purbeck Limestone Group of Dorset, southern England".
840:
5095:
3910:
2316:
994:
990:
970:
314:
in 1861, and since their original description only additional teeth have been discovered. The specific name honours collector
3438:
1834:, and is one of few Berriasian deposits globally, but is limited almost entirely to cranial or dental material, and tracks.
3931:
3491:
1854:
known both from body fossils and from footprints. Beyond ornithischians, the Lulworth Formation also contains the theropod
2605:"Lower limits of ornithischian dinosaur body size inferred from a new Upper Jurassic heterodontosaurid from North America"
1603:
and the undescribed Kayenta heterodontosaurid all bear low-crowned teeth unlike the Gondwanan forms, as well as a lobular
5100:
2971:
772:
1651:, while all other heterodontosaurs studied formed a group at the origins of the clade. This hypothesis would reduce the
3106:
Dieudonné, P. -E.; Cruzado-Caballero, P.; Godefroit, P.; Tortosa, T. (2020). "A new phylogeny of cerapodan dinosaurs".
2929:
979:
732:
5115:
4692:
580:
and form part of the Beckles Collection, bearing specimen numbers NHMUK 48209 to 48215. Lydekker also specified that
573:
3613:(Theria, Symmetrodonta) from the Lower Cretaceous of North Africa and Europe, and a brief review of symmetrodonts".
1605:
1008:
1000:
764:
748:
509:
497:
404:, and the sediments show that the Purbeck Group was a variably lagoonal environment initially similar to the modern
4712:
3507:"Micro-computed tomography reveals a diversity of Peramuran mammals from the Purbeck Group (Berriasian) of England"
1671:
and other heterodontosaurids is poorly known; most palaeobiological aspects are based on the almost-complete genus
1017:
833:
780:
740:
1033:
951:
915:
that a diastema was present and arched based on the maxilla NHMUK 48211. Nine teeth are present in the maxilla of
3265:
2584:
2404:
943:
845:
788:
756:
4699:
2582:
Norman, D.B.; Barrett, P.M. (2002). "Ornithischian dinosaurs from the Lower Cretaceous(Berriasian) of England".
2442:
1697:
regions of the premaxilla and predentary likely were the basis for keratinous beaks as in other ornithischians.
1288:
also resolved a basal placement for heterodontosaurids, and resolved internal relationships of the family, with
388:
both the youngest and the smallest heterodontosaurid. Other dinosaurs it lived alongside include the ornithopod
3263:
Evans, S.E.; McGowan, G.J. (2002). "Lissamphibian remains from the Purbeck Limestone Group, southern England".
1177:
in 1978, and along with its description reviewed the anatomy of multiple basal ornithischian genera, including
5105:
4719:
3873:
3193:
1015:
by Galton in 1978, and were considered absent by Norman and Barrett in 2002, Sereno described the first two
3569:"New symmetrodonts (Mammalia, Theria) from the Purbeck Limestone Group, Lower Cretaceous, southern England"
2440:
Galton, P.M. (1978). "Fabrosauridae, the basal family of ornithischian dinosaurs (Reptilia: Ornithopoda)".
1296:
as well. Following additional analysis by South American palaeontologist Diego Pol and colleagues in 2011,
4705:
3295:(Brodie) (Insecta: Diptera: Rhagionidae) from the Purbeck Limestone Group (Lower Cretaceous) of England".
1228:
to be a heterodontosaurid based on the presence of canines and arched gaps in the front of the tooth row.
5090:
5085:
5070:
4663:
1591:
and the other Laurasian heterodontosaurids were excluded from the clade of Gondwanan heterodontosaurids.
503:
5125:
5120:
5075:
3903:
2292:
2204:
333:
lizard, though this was quickly revised to an intermediate ornithischian. It was referred to the clade
2349:
Monograph on the Fossil Reptilia of the Wealden and Purbeck Formations. Supplement No. V. Dinosauria (
665:. Three years later Calliston revised his description and referred the material, including jaw bones,
129:
3018:
from the Quantou Formation (Cretaceous: Aptian–Cenomanian) of Jilin Province, north-eastern China".
1280:
supported heterodontosaurids as the most basal ornithischians instead. A revised analysis by Butler
3998:
1790:
1620:
3394:, a relatively abundant pleurosternid turtle (Paracryptodira) in the Early Cretaceous of Europe".
2371:
2347:
1679:
indicate occlusion across the top of the tooth, unlike the implied jaw motion in other forms like
3347:
707:
585:
2603:
Butler, R.J.; Galton, P.M.; Porro, L.B.; Chiappe, L.M.; Henderson, D.M.; Erickson, G.M. (2009).
2288:
Monograph on the Fossil Reptilia of the Wealden and Purbeck Formations. Part V. Order Lacertilia
5047:
4971:
4094:
1628:
337:
based on general dental anatomy and incorrectly referred armour that was later identified as a
5021:
3147:
Crompton and Charig 1962: cranial anatomy, functional morphology, taxonomy, and relationships"
600:
to the freshwater "Dirt Bed", also known as the "Mammal Pit" that was excavated by Beckles in
5042:
5034:
4340:
3896:
2969:
Butler, R.J.; Upchurch, P.; Norman, D.B. (2008). "The phylogeny of ornithischian dinosaurs".
2210:
2198:
1754:
1272:. Further research, including the extensive phylogenetic analysis of British palaeontologist
1055:
was considered an intermediate, potentially ornithischian, dinosaur until it was referred by
691:
250:
1683:
where the wear facets are only on the sides of the crowns. Despite the unique jaw motion of
1647:
as the sister taxon to the taxa typically classified within Pachycephalosauria, followed by
4608:
4363:
3845:
3717:
3680:
3622:
3583:
3518:
3403:
3356:
3304:
3234:
3115:
3064:
3027:
2980:
2938:
2886:
2840:
2800:
2688:
2451:
1643:
and other heterodontosaurs were early forms of pachycephalosaurs. Their results placed the
1624:
1612:
heterodontosaurids that lasts 50 million years through the Jurassic. Missing data for this
1285:
1277:
1224:. As these were what was suggested to unite the genus with "fabrosaurs", Sereno considered
1209:
1112:
1076:
1068:
912:
686:
678:
674:
662:
601:
597:
577:
557:
531:
461:
227:
1300:
was resolved as a basal ornithischian, yet not within the family Heterodontosauridae. Pol
730:
is limited to bones of the skull, but the multiple specimens include at least some of the
8:
4640:
4215:
3708:
3055:
2791:
2652:
2526:"Taxonomy, morphology, masticatory function and phylogeny of heterodontosaurid dinosaurs"
1894:
1424:
1347:
1198:
654:
530:
Although originally described as a lacertilian by Owen, he revised his classification in
354:
203:
3849:
3721:
3684:
3626:
3587:
3522:
3407:
3360:
3308:
3238:
3119:
3068:
3031:
2984:
2942:
2890:
2844:
2804:
2692:
2455:
1793:
in Durlston Bay and has a transitional but locally obscured boundary with the overlying
871:
456:
bones of the upper and lower jaws, were found alongside shells and plant fossils in the
4925:
4652:
3818:
3741:
3372:
3320:
3287:
Mostovski, M.B.; Ross, A.J.; Szadziewski, R.; Krzeminski, W. (2003). "Redescription of
3088:
2996:
2902:
2856:
2629:
2604:
2550:
2525:
2467:
2377:
2216:
1778:
1774:
1120:
658:
609:
124:
3814:
3568:
2783:
2671:
2655:. Geodiversitas, Museum National d’Histoire Naturelle Paris, In press. ffhal-03264773f
1769:
associations. The climate of the early Purbeck Group was likely similar to the modern
941:
Little of the lacrimal, jugal, and palatine are preserved, although the margin of the
400:, which are both also fragmentary. An abundance of small mammals also lived alongside
5029:
4893:
4245:
3822:
3733:
3531:
3506:
3434:
3368:
3339:
3165:
3142:
3080:
3039:
2634:
2555:
1898:
1884:, and varieties of invertebrates. Amphibians from the Lulworth Formation include the
1536:
1273:
1241:
1230:
286:
3745:
3376:
3324:
3092:
3000:
2471:
4472:
3853:
3810:
3725:
3692:
3688:
3630:
3591:
3526:
3468:
3411:
3364:
3312:
3242:
3202:
3189:"Highly derived eutherian mammals from the earliest Cretaceous of southern Britain"
3160:
3123:
3072:
3035:
2988:
2950:
2946:
2906:
2894:
2860:
2848:
2831:
2808:
2700:
2696:
2651:
Ronan Allain, Romain Vullo, Lee Rozada, Jérémy Anquetin, Renaud Bourgeais, et al..
2624:
2616:
2545:
2535:
2459:
2325:
2122:
1636:
1108:
561:
361:, it was eventually reidentified as the most basal group of ornithischians, making
30:
3857:
3429:
Sweetman, S.C.; Evans, S.E. (2011). "Lepidosaurs (Lizards)". In Batten, D. (ed.).
3415:
3127:
773:
4915:
4620:
4488:
3983:
3958:
3668:
2715:
2307:
2286:
2005:
1972:
1836:
1750:
1060:
949:
can be identified in the fragment of the lacrimal. The ectopterygoid bone of the
908:
904:
515:
along the sides of the teeth. Owen had corresponded with British palaeontologist
507:, 'tooth', which Owen combined as "prickly tooth" to describe the anatomy of the
469:
437:
433:
3472:
2782:
Barrett, P.M.; Clarke, J.B.; Brinkman, D.B.; Chapman, S.D.; Ensom, P.C. (2002).
1942:. The Purbeck is one of the most diverse Early Cretaceous deposits globally for
1216:, and compared it to a large number of other basal ornithischians. The teeth of
1006:
are present along the margin of the buccal emargination, as in the maxilla. The
4930:
4910:
4861:
4786:
4571:
4481:
4419:
4384:
4311:
3797:
Falconer, H. (1857). "Description of Two Species of the Fossil Mammalian Genus
2173:
2155:
2069:
2033:
1966:
1911:
1847:
1816:
1153:
980:
733:
568:
as a dinosaur based on the anatomy of its teeth, describing them as similar to
429:
366:
315:
3729:
3316:
3141:
Norman, D.B.; Crompton, A.W.; Butler, R.J.; Porro, L.B.; Charig, A.J. (2011).
3076:
2992:
2329:
1193:
were definitively referred by him to Fabrosauridae, although the placement of
1089:
961:
5064:
4994:
4878:
4828:
4586:
4557:
4543:
4527:
4412:
4405:
4304:
4224:
4148:
4141:
4109:
3548:
2877:
Galton, P.M. (1972). "Classification and Evolution of Ornithopod Dinosaurs".
2540:
2186:
2086:
2065:
1978:
1954:
1938:
1920:
1798:
1794:
1770:
1652:
1613:
1606:
1519:
1236:
1173:
1168:
1141:
1095:
1064:
1009:
1001:
817:
765:
749:
540:
516:
510:
493:
405:
377:
346:
65:
1051:
as a herbivorous lizard, he revised its placement to one within Dinosauria.
1018:
834:
794:
of both the upper and lower jaws. Based on proportions of the related genus
781:
741:
4920:
4871:
4814:
4800:
4793:
4748:
4564:
4519:
4447:
4426:
4329:
4203:
4195:
4171:
4125:
4060:
4047:
4010:
3919:
3737:
3634:
3595:
3246:
3207:
3188:
3084:
2829:
Thulborn, R.A. (1971). "Origins and Evolution of Ornithischian Dinosaurs".
2812:
2638:
2620:
2559:
2253:
2126:
2114:
2096:
2048:
2009:
1943:
1932:
1926:
1881:
1762:
1639:, French palaeontologist Paul-Emile Dieudonné and colleagues proposed that
1147:
1056:
1034:
952:
920:
681:
he reconsidered the material and noticed differences in tooth anatomy from
589:
549:
480:
465:
457:
357:. While the family was originally considered to be closest to more derived
311:
223:
190:
3105:
1127:
forms lacking canines (termed "fabrosaurs"), close to the Jurassic genera
944:
846:
789:
757:
4849:
4821:
4779:
4772:
4765:
4597:
4496:
4461:
4398:
4352:
4284:
4163:
4133:
4086:
4078:
2221:
2192:
2177:
2159:
2138:
2108:
2060:
1996:
1916:
1889:
1851:
1746:
1326:
1159:
1080:
896:
588:, the formal name for the historic Purbeck Beds. British palaeontologist
524:
519:, who had suggested the name "Sauraechinodon", but as the shortened form
334:
330:
40:
3340:"The turtles of the Purbeck Limestone Group of Dorset, southern England"
860:
5008:
4842:
4835:
4629:
4578:
4550:
4535:
4511:
4454:
4433:
4318:
4297:
4273:
4266:
4252:
4238:
4187:
4179:
4155:
2463:
2236:
2165:
2144:
2102:
2081:
2027:
1984:
1960:
1948:
1861:
1810:
1786:
1758:
1738:
1726:
1722:
1718:
1694:
1632:
1469:
1438:
1402:
1385:
1265:
1261:
1135:
925:
804:
625:
612:
of the Purbeck Group. Galton also referred the specimen NHMUK 48229 to
553:
489:
358:
301:
293:
85:
50:
1587:
According to the resolution of the 2012 analysis conducted by Sereno,
895:
is a topic of disagreement. While Galton and American palaeontologist
802:
would have been 62 mm (2.4 in) long, which is comparable to
107:
4807:
4440:
4117:
3871:
Sigogneau-Russell, D. (2003). "Docodonts from the British Mesozoic".
2898:
2852:
2150:
2090:
2015:
2001:
1902:
1842:
1782:
1705:
1308:
1269:
1129:
824:
650:
584:
was found in the Middle Purbeck Beds, an informal unit of the modern
474:
369:, with a ghost lineage of 50 million years of unpreserved evolution.
141:
90:
4965:
2765:
Proceedings of the Dorset Natural History and Archaeological Society
2746:
Proceedings of the Dorset Natural History and Archaeological Society
488:
for the fossils, which he considered to be part of the lizard clade
4988:
4758:
4259:
4102:
3967:
3943:
2182:
2170:
2119:
2054:
2038:
2023:
1856:
1822:
1742:
1656:
1486:
1335:
1244:
were also placed within the family. Norman and Barrett redescribed
1124:
888:
666:
646:
396:
390:
289:
174:
161:
116:
80:
75:
60:
55:
45:
3286:
2220:. Specific sites within the formation also preserve the primitive
1075:
on the basis of a stegosaurian classification by Justin Delair in
718:
4886:
3949:
2530:
2132:
2075:
2042:
1885:
1877:
1869:
1865:
1804:
1083:
453:
449:
441:
297:
95:
70:
2763:
Delair, J.B. (1959). "The Mesozoic reptiles of Dorset. Part 2".
3937:
3888:
2356:. Vol. Supp. 5. Palaeontographical Society. pp. 1–18.
1873:
1831:
1766:
1734:
642:
381:
338:
151:
1789:. The Purbeck Group is visibly underlain by the Late Jurassic
1737:. Sedimentology shows they were deposited in a fluctuation of
1107:
Classifications of many basal ornithischians were reviewed by
3669:"Monograph of the Fossil Mammalia of the Mesozoic Formations"
859:. The surface of the front of the bone along the toothrow is
816:
the smallest presumably adult heterodontosaur and one of the
619:
445:
282:
3835:
3224:
2927:, "Fabrosaurids," and the early evolution of Ornithischia".
2781:
416:
4504:
1907:
1730:
1197:
was questionable as it also bore similarities to the clade
3140:
2190:. Crocodilians from within the Lulworth deposits include
2000:
and three unnamed tooth morphologies represent the known
1753:
environments. Flora and fauna are indicative of variable
1619:
Following the earlier results of Chinese palaeontologist
1616:
may be due to the exceptionally small size of the group.
460:. These were first described in a monograph published in
349:, which was later identified as an artificial group with
3013:
2602:
3390:
Pérez-García, A. (2014). "Revision of the poorly known
1850:
is also known, along with intermediate ornithopods and
823:
The main body of both premaxillae are preserved in the
702:
In 2021 two premaxillary teeth that were attributed to
3777:
3758:
3705:
3647:
3608:
3433:. The Palaeontological Association. pp. 264–284.
1808:
Environment of the Purbeck beds including the mammals
863:
and lacks teeth, distinguishing the bone from that of
710:
of France, which is coeval with the Purbeck deposits.
637:
In the late 1970s and early 1980s, excavations of the
4947:
List of stratigraphic units with ornithischian tracks
3566:
3186:
2968:
808:
at 66 mm (2.6 in) but less than adults of
345:
was then referred to the early ornithischian family
3456:
1323:was poorly supported based on few dental features.
883:Three maxillae are preserved among the material of
428:Multiple specimens of jaw bones were discovered by
3488:Lower and Middle Cretaceous Terrestrial Ecosystems
3273:(Life and Environments in Purbeck Times): 103–119.
3220:
3218:
2592:(Life and Environments in Purbeck Times): 161–189.
2412:(Life and Environments in Purbeck Times): 121–144.
1655:of pachycephalosaurs and pull back the origins of
1208:as a member of the family Heterodontosauridae. In
641:uncovered many small ornithischian fossils in the
3870:
3545:
3187:Sweetman, S.C.; Smith, G.; Martill, D.M. (2017).
2022:The diverse mammal assemblage includes the small
831:, although fractured and crushed. A premaxillary
5062:
3282:
3280:
3052:
1252:Although a placement as a heterodontosaurid for
1139:, and more primitive than the Cretaceous genera
851:it is nested within is more similar in shape to
304:epoch. The first specimens were jaw bones named
3215:
3182:
3180:
3178:
3176:
2824:
2822:
2743:
2397:
2395:
2393:
2391:
2389:
2387:
1717:The Purbeck Group is a distinctive sequence of
1079:, although these have since been reassigned to
3982:
2739:
2737:
2735:
2733:
2365:
2363:
3904:
3864:
3699:
3602:
3539:
3428:
3277:
3099:
2964:
2962:
2960:
2918:
2916:
2872:
2870:
2680:Annual Review of Earth and Planetary Sciences
2665:
2663:
2661:
2581:
2519:
2517:
2515:
2513:
2511:
2509:
2507:
2505:
2503:
2501:
1292:being a basal genus in the family along with
3790:
3609:Sigogneau-Russell, D.; Ensom, P.C. (1998). "
3452:
3450:
3422:
3389:
3262:
3258:
3256:
3173:
2819:
2577:
2575:
2573:
2571:
2569:
2499:
2497:
2495:
2493:
2491:
2489:
2487:
2485:
2483:
2481:
2435:
2433:
2431:
2429:
2427:
2425:
2423:
2421:
2419:
2384:
973:view showing the anatomy of the tooth crowns
689:, these fossils were given their own genus,
639:Natural History Museum of Los Angeles County
365:a taxon descended from many genera from the
4058:
3803:Quarterly Journal of the Geological Society
3460:Bulletin de la Société Géologique de France
3143:"The Lower Jurassic ornithischian dinosaur
2775:
2730:
2360:
2299:
923:similar to those seen in the premaxilla of
592:narrowed down the depositional locality of
3911:
3897:
3829:
3771:
3752:
3567:Ensom, P.C.; Sigogneau-Russel, D. (2000).
3331:
3134:
3007:
2957:
2913:
2867:
2756:
2707:
2658:
2341:
2339:
2308:"Note on the synonymy of the fossil genus
2280:
2278:
2276:
2274:
2272:
2270:
1339:, hypothetically restored as a dromaeosaur
919:, the first being an enlarged and slender
673:sp., an assignment supported by Galton in
106:
3641:
3560:
3530:
3498:
3479:
3447:
3383:
3253:
3206:
3164:
3152:Zoological Journal of the Linnean Society
2628:
2596:
2566:
2549:
2539:
2478:
2416:
2401:
812:at 75 mm (3.0 in) long, making
3796:
3660:
2828:
2369:
2305:
1846:. A femur and dorsal of an intermediate
1803:
1704:
1325:
1088:
960:
870:
717:
618:
415:
3046:
2672:"The Origin and Evolution of Dinosaurs"
2336:
2267:
1631:was created to unite heterodontosaurs,
843:margin of the individual bone, and the
411:
5063:
3337:
2922:
2876:
2762:
2713:
2669:
2523:
2439:
2317:Annals and Magazine of Natural History
2208:, dubious remains previously known as
1915:. Four taxa of turtles are known, the
899:interpret a diastema as present as in
604:. However, no evidence exists to link
4970:
4969:
4735:
4045:
3930:
3892:
3504:
3485:
875:Reconstruction of the known skull of
5081:Early Cretaceous dinosaurs of Europe
3666:
3492:New Mexico Museum of Natural History
2345:
2284:
1820:(center and right) and the theropod
1212:, he revised the cranial anatomy of
695:, a related but distinct taxon from
564:followed previous classification of
552:, which was within the larger clade
420:Teeth and jaw fragments referred to
3297:Journal of Systematic Palaeontology
2972:Journal of Systematic Palaeontology
1204:Sereno was the first to reclassify
300:and possibly western France in the
13:
2930:Journal of Vertebrate Paleontology
2645:
2609:Proceedings of the Royal Society B
14:
5137:
3815:10.1144/GSL.JGS.1857.013.01-02.39
1240:and an undescribed form from the
1042:
574:British Museum of Natural History
4718:
4711:
4704:
4698:
4691:
4046:
3918:
3532:10.1111/j.1475-4983.2012.01161.x
3369:10.1111/j.0031-0239.2004.00418.x
3166:10.1111/j.1096-3642.2011.00697.x
3040:10.1111/j.1475-4983.2011.01046.x
1713:on the southern coast of England
1662:
649:. These remains, collected from
444:. These fossils, including many
128:
38:
5111:Early Cretaceous ornithischians
3266:Special Papers in Palaeontology
2585:Special Papers in Palaeontology
2405:Special Papers in Palaeontology
1675:. Wear facets on the crowns of
329:was considered to be a type of
318:who discovered the material of
292:that lived during the earliest
3693:10.1080/02693445.1871.12113244
3673:The Palaeontographical Society
3494:Bulletin 14. pp. 195–200.
2951:10.1080/02724634.1991.10011386
2701:10.1146/annurev.earth.25.1.435
2004:, and fossils referred to the
1167:were united in the new family
713:
472:who also described fossils of
1:
5096:Fossil taxa described in 1861
3874:Acta Palaeontologica Polonica
3858:10.1016/j.cretres.2011.09.007
3416:10.1016/j.cretres.2014.02.015
3194:Acta Palaeontologica Polonica
3128:10.1080/08912963.2020.1793979
2260:
1659:back to the Early Jurassic.
669:and most of the hindlimb, to
408:but became wetter over time.
4736:
2443:Paläontologische Zeitschrift
818:smallest non-avian dinosaurs
7:
3473:10.2113/gssgfbull.183.6.517
1047:Owen originally classified
903:, British palaeontologists
608:to any specific bed in the
10:
5142:
5101:Taxa named by Richard Owen
2293:Palaeontographical Society
2205:Pholidosaurus purbeckensis
1700:
502:
496:
16:Extinct genus of dinosaurs
4978:
4939:
4903:
4859:
4746:
4742:
4731:
4688:
4651:
4628:
4619:
4596:
4471:
4393:
4382:
4351:
4328:
4292:
4283:
4214:
4073:
4069:
4054:
4041:
4009:
3991:
3978:
3926:
3730:10.1007/s00114-013-1028-3
3392:Dorsetochelys typocardium
3317:10.1017/S1477201903001020
3077:10.1007/s00114-011-0780-5
2993:10.1017/S1477201907002271
2330:10.1080/00222936108697425
1533:
1516:
1509:
1483:
1466:
1459:
1452:
1435:
1428:
1399:
1382:
1365:
1358:
1351:
1099:, a proposed relative of
985:bone is not preserved in
256:
249:
125:Scientific classification
123:
114:
105:
23:
5116:Ornithischians of Europe
2716:"The genera of reptiles"
2541:10.3897/zookeys.226.2840
2524:Sereno, Paul C. (2012).
2214:, and the dubious taxon
2181:; and the non-mammalian
645:Paleontological Area of
560:British palaeontologist
3431:English Wealden Fossils
3145:Heterodontosaurus tucki
2193:Goniopholis gracilidens
1709:Locality (red dot) for
708:Angeac-Charente bonebed
706:were reported from the
586:Purbeck Limestone Group
376:have been found in the
3635:10.1006/cres.1998.0115
3596:10.1006/cres.2000.0227
3293:Pseudosimulium humidum
3247:10.1006/cres.2001.0265
3208:10.4202/app.00408.2017
3016:Changchunsaurus parvus
2923:Sereno, P.C. (1991). "
2813:10.1006/cres.2002.1002
2621:10.1098/rspb.2009.1494
2019:have also been found.
1826:
1714:
1629:Heterodontosauriformes
1340:
1104:
974:
880:
726:The known material of
723:
634:
425:
424:, NHMUK 48209 to 48215
119:specimen NHMUK 48215b
5043:Paleobiology Database
3338:Milner, A.R. (2004).
2670:Sereno, P.C. (1997).
2370:Lydekker, R. (1888).
2306:Falconer, H. (1861).
2211:Goniopholis tenuidens
2199:Theriosuchus pusillus
1807:
1708:
1329:
1092:
964:
874:
721:
692:Fruitadens haagarorum
622:
548:in a clade he called
484:. Owen gave the name
419:
5106:Ornithischian genera
3505:Davis, B.M. (2012).
2154:; the non-eutherian
2085:; the non-eutherian
2064:; the non-eutherian
2037:; the non-eutherian
1893:and an intermediate
1604:
1333:being attacked by a
1103:within Fabrosauridae
1032:
1016:
1007:
999:
978:
950:
942:
844:
839:is present near the
832:
787:
779:
771:
763:
755:
747:
739:
731:
623:Life restoration of
576:(BMNH now NHMUK) in
508:
412:History of discovery
4216:Heterodontosauridae
3850:2012CrRes..33..135B
3838:Cretaceous Research
3722:2013NW....100..311A
3709:Naturwissenschaften
3685:1871MPalS..24D...5O
3627:1998CrRes..19..445S
3615:Cretaceous Research
3588:2000CrRes..21..767E
3576:Cretaceous Research
3523:2012Palgy..55..789D
3408:2014CrRes..49..152P
3396:Cretaceous Research
3361:2004Palgy..47.1441M
3309:2003JSPal...1...59M
3239:2001CrRes..22..451L
3227:Cretaceous Research
3120:2021HBio...33.2335D
3069:2011NW.....98..369P
3056:Naturwissenschaften
3032:2011Palgy..54..667B
2985:2008JSPal...6....1B
2943:1991JVPal..11..168S
2891:1972Natur.239..464G
2845:1971Natur.234...75T
2805:2002CrRes..23..279B
2792:Cretaceous Research
2714:Nopcsa, F. (1928).
2693:1997AREPS..25..435S
2456:1978PalZ...52..138G
2380:. pp. 247–248.
1425:Heterodontosaurinae
1348:Heterodontosauridae
1199:Heterodontosauridae
655:Brushy Basin Member
653:at the base of the
486:Echinodon becklesii
432:high on a cliff in
355:Heterodontosauridae
307:Echinodon becklesii
263:Echinodon becklesii
204:Heterodontosauridae
5091:Fossils of England
5086:Cretaceous England
5071:Heterodontosaurids
4926:Hypsilophodontidae
4664:Pachycephalosauria
4653:Pachycephalosauria
3289:Simulidium priscum
3108:Historical Biology
2464:10.1007/BF03006735
2378:Taylor and Francis
2312:of Professor Owen"
2217:Macellodus brodiei
1827:
1779:Lulworth Formation
1775:Durlston Formation
1715:
1627:, where the clade
1623:and colleagues in
1341:
1276:and colleagues in
1264:more derived than
1121:Hypsilophodontidae
1115:, where he placed
1105:
975:
881:
879:from Sereno (2012)
855:than more derived
724:
659:Morrison Formation
635:
610:Lulworth Formation
501:, 'hedgehog', and
426:
5126:Fossils of France
5121:Cretaceous France
5076:Berriasian genera
5058:
5057:
5030:Open Tree of Life
4972:Taxon identifiers
4963:
4962:
4959:
4958:
4955:
4954:
4894:Serendipaceratops
4727:
4726:
4686:
4685:
4682:
4681:
4678:
4677:
4674:
4673:
4378:
4377:
4374:
4373:
4246:Heterodontosaurus
4037:
4036:
4033:
4032:
4029:
4028:
3667:Owen, R. (1871).
3440:978-1-444-36711-9
2885:(5373): 464–466.
2615:(1680): 375–381.
2346:Owen, R. (1874).
2295:. pp. 31–39.
2285:Owen, R. (1861).
2123:multituberculates
2006:rhynchocephalians
1895:batrachosauroidid
1729:interbedded with
1685:Heterodontosaurus
1677:Heterodontosaurus
1673:Heterodontosaurus
1637:pachycephalosaurs
1584:
1583:
1575:
1574:
1566:
1565:
1557:
1556:
1548:
1547:
1537:Heterodontosaurus
1498:
1497:
1414:
1413:
1274:Richard J. Butler
1242:Kayenta Formation
1231:Heterodontosaurus
1093:Partial skull of
931:Heterodontosaurus
901:Heterodontosaurus
853:Heterodontosaurus
796:Heterodontosaurus
394:and the theropod
372:All specimens of
287:heterodontosaurid
274:
273:
242:E. becklesii
230:
5133:
5051:
5050:
5038:
5037:
5025:
5024:
5012:
5011:
4999:
4998:
4997:
4967:
4966:
4744:
4743:
4733:
4732:
4722:
4716:
4715:
4709:
4708:
4702:
4696:
4695:
4626:
4625:
4473:Thescelosauridae
4391:
4390:
4290:
4289:
4071:
4070:
4067:
4066:
4056:
4055:
4043:
4042:
3989:
3988:
3980:
3979:
3973:
3972:
3928:
3927:
3913:
3906:
3899:
3890:
3889:
3883:
3882:
3868:
3862:
3861:
3833:
3827:
3826:
3809:(1–2): 261–282.
3794:
3788:
3787:
3775:
3769:
3768:
3756:
3750:
3749:
3703:
3697:
3696:
3664:
3658:
3657:
3645:
3639:
3638:
3621:(3–4): 445–470.
3606:
3600:
3599:
3573:
3564:
3558:
3557:
3543:
3537:
3536:
3534:
3502:
3496:
3495:
3483:
3477:
3476:
3454:
3445:
3444:
3426:
3420:
3419:
3387:
3381:
3380:
3355:(6): 1441–1467.
3344:
3335:
3329:
3328:
3284:
3275:
3274:
3260:
3251:
3250:
3222:
3213:
3212:
3210:
3184:
3171:
3170:
3168:
3138:
3132:
3131:
3103:
3097:
3096:
3050:
3044:
3043:
3011:
3005:
3004:
2966:
2955:
2954:
2920:
2911:
2910:
2899:10.1038/239464a0
2874:
2865:
2864:
2853:10.1038/234075a0
2826:
2817:
2816:
2788:
2779:
2773:
2772:
2760:
2754:
2753:
2741:
2728:
2727:
2711:
2705:
2704:
2676:
2667:
2656:
2649:
2643:
2642:
2632:
2600:
2594:
2593:
2579:
2564:
2563:
2553:
2543:
2521:
2476:
2475:
2450:(1–2): 138–159.
2437:
2414:
2413:
2399:
2382:
2381:
2367:
2358:
2357:
2343:
2334:
2333:
2303:
2297:
2296:
2282:
1608:
1512:
1511:
1462:
1461:
1455:
1454:
1431:
1430:
1361:
1360:
1354:
1353:
1344:
1343:
1109:Richard Thulborn
1036:
1020:
1011:
1003:
982:
965:Left dentary of
954:
946:
848:
836:
791:
786:along with most
783:
775:
767:
759:
751:
743:
735:
562:Richard Lydekker
512:
506:
500:
265:
261:
222:
215:
202:
189:
133:
132:
110:
100:
37:
31:Early Cretaceous
29:Temporal range:
21:
20:
5141:
5140:
5136:
5135:
5134:
5132:
5131:
5130:
5061:
5060:
5059:
5054:
5046:
5041:
5033:
5028:
5020:
5015:
5007:
5002:
4993:
4992:
4987:
4974:
4964:
4951:
4935:
4916:Phytodinosauria
4899:
4855:
4738:
4723:
4710:
4703:
4690:
4670:
4647:
4621:Marginocephalia
4615:
4592:
4489:Changchunsaurus
4467:
4387:
4370:
4347:
4324:
4279:
4210:
4063:
4050:
4025:
4005:
3999:Avemetatarsalia
3984:Avemetatarsalia
3974:
3959:Avemetatarsalia
3933:
3932:
3922:
3917:
3887:
3886:
3869:
3865:
3834:
3830:
3801:from Purbeck".
3795:
3791:
3776:
3772:
3757:
3753:
3704:
3700:
3665:
3661:
3646:
3642:
3607:
3603:
3571:
3565:
3561:
3544:
3540:
3503:
3499:
3484:
3480:
3455:
3448:
3441:
3427:
3423:
3388:
3384:
3342:
3336:
3332:
3285:
3278:
3261:
3254:
3223:
3216:
3185:
3174:
3139:
3135:
3104:
3100:
3051:
3047:
3012:
3008:
2967:
2958:
2921:
2914:
2875:
2868:
2839:(5324): 75–78.
2827:
2820:
2786:
2780:
2776:
2761:
2757:
2742:
2731:
2720:Palaeobiologica
2712:
2708:
2674:
2668:
2659:
2650:
2646:
2601:
2597:
2580:
2567:
2522:
2479:
2438:
2417:
2400:
2385:
2368:
2361:
2344:
2337:
2304:
2300:
2283:
2268:
2263:
2174:morganucodontan
1973:Pseudosaurillus
1899:albanerpetontid
1837:Owenodon hoggii
1703:
1667:The anatomy of
1665:
1610:
1585:
1576:
1567:
1558:
1549:
1499:
1415:
1123:: derived from
1061:Scelidosaurinae
1045:
1038:
1022:
1013:
1005:
984:
969:NHMUK 48213 in
956:
948:
909:Paul M. Barrett
905:David B. Norman
850:
838:
798:, the skull of
793:
785:
777:
769:
761:
753:
745:
737:
722:Size comparison
716:
514:
470:palaeontologist
464:written by Sir
438:Isle of Purbeck
434:Durdlestone Bay
414:
270:
267:
259:
258:
245:
221:
213:
200:
187:
127:
101:
99:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
35:
34:
27:
17:
12:
11:
5:
5139:
5129:
5128:
5123:
5118:
5113:
5108:
5103:
5098:
5093:
5088:
5083:
5078:
5073:
5056:
5055:
5053:
5052:
5039:
5026:
5013:
5000:
4984:
4982:
4976:
4975:
4961:
4960:
4957:
4956:
4953:
4952:
4950:
4949:
4943:
4941:
4940:Other articles
4937:
4936:
4934:
4933:
4931:Jeholosauridae
4928:
4923:
4918:
4913:
4911:Ornithoscelida
4907:
4905:
4901:
4900:
4898:
4897:
4890:
4883:
4875:
4867:
4865:
4862:Incertae sedis
4857:
4856:
4854:
4853:
4846:
4839:
4832:
4825:
4818:
4811:
4804:
4797:
4790:
4787:Ferganocephale
4783:
4776:
4769:
4762:
4754:
4752:
4740:
4739:
4729:
4728:
4725:
4724:
4689:
4687:
4684:
4683:
4680:
4679:
4676:
4675:
4672:
4671:
4669:
4668:
4667:
4666:
4657:
4655:
4649:
4648:
4646:
4645:
4644:
4643:
4634:
4632:
4623:
4617:
4616:
4614:
4613:
4612:
4611:
4602:
4600:
4594:
4593:
4591:
4590:
4583:
4575:
4572:Thescelosaurus
4568:
4561:
4554:
4547:
4540:
4532:
4524:
4516:
4508:
4501:
4493:
4485:
4482:Albertadromeus
4477:
4475:
4469:
4468:
4466:
4465:
4458:
4451:
4444:
4437:
4430:
4423:
4420:Kulindadromeus
4416:
4409:
4402:
4394:
4388:
4385:Neornithischia
4383:
4380:
4379:
4376:
4375:
4372:
4371:
4369:
4368:
4367:
4366:
4357:
4355:
4349:
4348:
4346:
4345:
4344:
4343:
4334:
4332:
4326:
4325:
4323:
4322:
4315:
4312:Scutellosaurus
4308:
4301:
4293:
4287:
4281:
4280:
4278:
4277:
4270:
4263:
4256:
4249:
4242:
4235:
4228:
4220:
4218:
4212:
4211:
4209:
4208:
4200:
4192:
4184:
4176:
4168:
4160:
4152:
4145:
4138:
4130:
4122:
4114:
4106:
4099:
4091:
4083:
4074:
4064:
4059:
4052:
4051:
4039:
4038:
4035:
4034:
4031:
4030:
4027:
4026:
4024:
4023:
4022:
4021:
4015:
4013:
4007:
4006:
4004:
4003:
4002:
4001:
3992:
3986:
3976:
3975:
3971:
3970:
3961:
3952:
3946:
3940:
3924:
3923:
3916:
3915:
3908:
3901:
3893:
3885:
3884:
3863:
3844:(1): 135–145.
3828:
3789:
3770:
3751:
3716:(4): 311–326.
3698:
3679:(110): 1–115.
3659:
3640:
3601:
3582:(6): 767–779.
3559:
3538:
3517:(4): 789–817.
3497:
3478:
3467:(6): 517–524.
3446:
3439:
3421:
3382:
3330:
3276:
3252:
3233:(4): 451–460.
3214:
3201:(4): 657–665.
3172:
3133:
3098:
3063:(5): 369–379.
3045:
3026:(3): 667–683.
3006:
2956:
2937:(2): 168–197.
2912:
2866:
2818:
2799:(2): 279–295.
2774:
2755:
2729:
2706:
2657:
2644:
2595:
2565:
2534:(226): 1–225.
2477:
2415:
2383:
2359:
2335:
2298:
2265:
2264:
2262:
2259:
2232:Pseudosimulium
2156:eutriconodonts
2070:Spalacotherium
2034:Durlstotherium
1967:Paramacellodus
1944:lepidosaurians
1912:Sunnybatrachus
1848:hadrosauriform
1817:Durlstotherium
1791:Portland Group
1777:and the lower
1702:
1699:
1664:
1661:
1582:
1581:
1578:
1577:
1573:
1572:
1569:
1568:
1564:
1563:
1560:
1559:
1555:
1554:
1551:
1550:
1546:
1545:
1542:
1541:
1532:
1529:
1528:
1525:
1524:
1515:
1510:
1508:
1505:
1504:
1501:
1500:
1496:
1495:
1492:
1491:
1482:
1479:
1478:
1475:
1474:
1465:
1460:
1458:
1453:
1451:
1448:
1447:
1444:
1443:
1434:
1429:
1427:
1421:
1420:
1417:
1416:
1412:
1411:
1408:
1407:
1398:
1395:
1394:
1391:
1390:
1381:
1378:
1377:
1374:
1373:
1364:
1359:
1357:
1352:
1350:
1342:
1154:Thescelosaurus
1044:
1043:Classification
1041:
715:
712:
677:, although in
430:Samuel Beckles
413:
410:
367:Early Jurassic
353:reassigned to
316:Samuel Beckles
272:
271:
268:
254:
253:
247:
246:
238:
236:
232:
231:
211:
207:
206:
198:
194:
193:
185:
178:
177:
172:
165:
164:
159:
155:
154:
149:
145:
144:
139:
135:
134:
121:
120:
112:
111:
103:
102:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
39:
28:
15:
9:
6:
4:
3:
2:
5138:
5127:
5124:
5122:
5119:
5117:
5114:
5112:
5109:
5107:
5104:
5102:
5099:
5097:
5094:
5092:
5089:
5087:
5084:
5082:
5079:
5077:
5074:
5072:
5069:
5068:
5066:
5049:
5044:
5040:
5036:
5031:
5027:
5023:
5018:
5014:
5010:
5005:
5001:
4996:
4990:
4986:
4985:
4983:
4981:
4977:
4973:
4968:
4948:
4945:
4944:
4942:
4938:
4932:
4929:
4927:
4924:
4922:
4919:
4917:
4914:
4912:
4909:
4908:
4906:
4902:
4896:
4895:
4891:
4889:
4888:
4884:
4881:
4880:
4879:Dravidosaurus
4876:
4874:
4873:
4869:
4868:
4866:
4864:
4863:
4858:
4852:
4851:
4847:
4845:
4844:
4840:
4838:
4837:
4833:
4831:
4830:
4829:Syngonosaurus
4826:
4824:
4823:
4819:
4817:
4816:
4812:
4810:
4809:
4805:
4803:
4802:
4798:
4796:
4795:
4791:
4789:
4788:
4784:
4782:
4781:
4777:
4775:
4774:
4770:
4768:
4767:
4763:
4761:
4760:
4756:
4755:
4753:
4751:
4750:
4745:
4741:
4734:
4730:
4721:
4717:
4714:
4707:
4701:
4694:
4665:
4661:
4660:
4659:
4658:
4656:
4654:
4650:
4642:
4638:
4637:
4636:
4635:
4633:
4631:
4627:
4624:
4622:
4618:
4610:
4606:
4605:
4604:
4603:
4601:
4599:
4595:
4589:
4588:
4587:Zephyrosaurus
4584:
4581:
4580:
4576:
4574:
4573:
4569:
4567:
4566:
4562:
4560:
4559:
4558:Oryctodromeus
4555:
4553:
4552:
4548:
4546:
4545:
4544:Nevadadromeus
4541:
4538:
4537:
4533:
4530:
4529:
4528:Koreanosaurus
4525:
4522:
4521:
4517:
4514:
4513:
4509:
4507:
4506:
4502:
4499:
4498:
4494:
4491:
4490:
4486:
4484:
4483:
4479:
4478:
4476:
4474:
4470:
4464:
4463:
4459:
4457:
4456:
4452:
4450:
4449:
4445:
4443:
4442:
4438:
4436:
4435:
4431:
4429:
4428:
4424:
4422:
4421:
4417:
4415:
4414:
4413:Hypsilophodon
4410:
4408:
4407:
4406:Hexinlusaurus
4403:
4401:
4400:
4396:
4395:
4392:
4389:
4386:
4381:
4365:
4361:
4360:
4359:
4358:
4356:
4354:
4350:
4342:
4338:
4337:
4336:
4335:
4333:
4331:
4327:
4321:
4320:
4316:
4314:
4313:
4309:
4307:
4306:
4305:Scelidosaurus
4302:
4300:
4299:
4295:
4294:
4291:
4288:
4286:
4282:
4276:
4275:
4271:
4269:
4268:
4264:
4262:
4261:
4257:
4255:
4254:
4250:
4248:
4247:
4243:
4241:
4240:
4236:
4234:
4233:
4229:
4227:
4226:
4225:Abrictosaurus
4222:
4221:
4219:
4217:
4213:
4206:
4205:
4201:
4198:
4197:
4193:
4190:
4189:
4185:
4182:
4181:
4177:
4174:
4173:
4169:
4166:
4165:
4161:
4158:
4157:
4153:
4151:
4150:
4149:Lesothosaurus
4146:
4144:
4143:
4142:Laquintasaura
4139:
4136:
4135:
4131:
4128:
4127:
4123:
4120:
4119:
4115:
4112:
4111:
4110:Eucoelophysis
4107:
4105:
4104:
4100:
4097:
4096:
4092:
4089:
4088:
4084:
4081:
4080:
4076:
4075:
4072:
4068:
4065:
4062:
4057:
4053:
4049:
4044:
4040:
4019:
4018:
4017:
4016:
4014:
4012:
4008:
4000:
3996:
3995:
3994:
3993:
3990:
3987:
3985:
3981:
3977:
3969:
3965:
3962:
3960:
3956:
3953:
3951:
3947:
3945:
3941:
3939:
3935:
3934:
3929:
3925:
3921:
3914:
3909:
3907:
3902:
3900:
3895:
3894:
3891:
3881:(3): 357–374.
3880:
3876:
3875:
3867:
3859:
3855:
3851:
3847:
3843:
3839:
3832:
3824:
3820:
3816:
3812:
3808:
3804:
3800:
3793:
3785:
3781:
3780:Palaeontology
3774:
3766:
3762:
3761:Palaeontology
3755:
3747:
3743:
3739:
3735:
3731:
3727:
3723:
3719:
3715:
3711:
3710:
3702:
3694:
3690:
3686:
3682:
3678:
3674:
3670:
3663:
3655:
3651:
3650:Palaeontology
3644:
3636:
3632:
3628:
3624:
3620:
3616:
3612:
3605:
3597:
3593:
3589:
3585:
3581:
3577:
3570:
3563:
3555:
3551:
3550:
3549:Geodiversitas
3542:
3533:
3528:
3524:
3520:
3516:
3512:
3511:Palaeontology
3508:
3501:
3493:
3489:
3482:
3474:
3470:
3466:
3462:
3461:
3453:
3451:
3442:
3436:
3432:
3425:
3417:
3413:
3409:
3405:
3401:
3397:
3393:
3386:
3378:
3374:
3370:
3366:
3362:
3358:
3354:
3350:
3349:
3348:Palaeontology
3341:
3334:
3326:
3322:
3318:
3314:
3310:
3306:
3302:
3298:
3294:
3291:Westwood and
3290:
3283:
3281:
3272:
3268:
3267:
3259:
3257:
3248:
3244:
3240:
3236:
3232:
3228:
3221:
3219:
3209:
3204:
3200:
3196:
3195:
3190:
3183:
3181:
3179:
3177:
3167:
3162:
3158:
3154:
3153:
3148:
3146:
3137:
3129:
3125:
3121:
3117:
3113:
3109:
3102:
3094:
3090:
3086:
3082:
3078:
3074:
3070:
3066:
3062:
3058:
3057:
3049:
3041:
3037:
3033:
3029:
3025:
3021:
3020:Palaeontology
3017:
3010:
3002:
2998:
2994:
2990:
2986:
2982:
2978:
2974:
2973:
2965:
2963:
2961:
2952:
2948:
2944:
2940:
2936:
2932:
2931:
2926:
2925:Lesothosaurus
2919:
2917:
2908:
2904:
2900:
2896:
2892:
2888:
2884:
2880:
2873:
2871:
2862:
2858:
2854:
2850:
2846:
2842:
2838:
2834:
2833:
2825:
2823:
2814:
2810:
2806:
2802:
2798:
2794:
2793:
2785:
2778:
2770:
2766:
2759:
2751:
2747:
2740:
2738:
2736:
2734:
2725:
2721:
2717:
2710:
2702:
2698:
2694:
2690:
2686:
2682:
2681:
2673:
2666:
2664:
2662:
2654:
2648:
2640:
2636:
2631:
2626:
2622:
2618:
2614:
2610:
2606:
2599:
2591:
2587:
2586:
2578:
2576:
2574:
2572:
2570:
2561:
2557:
2552:
2547:
2542:
2537:
2533:
2532:
2527:
2520:
2518:
2516:
2514:
2512:
2510:
2508:
2506:
2504:
2502:
2500:
2498:
2496:
2494:
2492:
2490:
2488:
2486:
2484:
2482:
2473:
2469:
2465:
2461:
2457:
2453:
2449:
2445:
2444:
2436:
2434:
2432:
2430:
2428:
2426:
2424:
2422:
2420:
2411:
2407:
2406:
2398:
2396:
2394:
2392:
2390:
2388:
2379:
2375:
2374:
2366:
2364:
2355:
2354:
2350:
2342:
2340:
2331:
2327:
2323:
2319:
2318:
2313:
2311:
2302:
2294:
2290:
2289:
2281:
2279:
2277:
2275:
2273:
2271:
2266:
2258:
2256:
2255:
2250:
2249:
2244:
2243:
2242:Eoptychoptera
2238:
2234:
2233:
2228:
2227:
2223:
2219:
2218:
2213:
2212:
2207:
2206:
2201:
2200:
2195:
2194:
2189:
2188:
2187:Peraiocynodon
2184:
2180:
2179:
2175:
2172:
2168:
2167:
2162:
2161:
2157:
2153:
2152:
2147:
2146:
2141:
2140:
2135:
2134:
2129:
2128:
2124:
2121:
2117:
2116:
2111:
2110:
2105:
2104:
2099:
2098:
2093:
2092:
2088:
2084:
2083:
2078:
2077:
2072:
2071:
2067:
2066:symmetrodonts
2063:
2062:
2057:
2056:
2051:
2050:
2045:
2044:
2040:
2036:
2035:
2030:
2029:
2025:
2020:
2018:
2017:
2012:
2011:
2007:
2003:
1999:
1998:
1993:
1992:
1987:
1986:
1981:
1980:
1979:Parasaurillus
1975:
1974:
1969:
1968:
1963:
1962:
1957:
1956:
1955:Dorsetisaurus
1951:
1950:
1946:. The genera
1945:
1941:
1940:
1939:Pleurosternon
1935:
1934:
1929:
1928:
1923:
1922:
1921:Dorsetochelys
1918:
1914:
1913:
1909:
1905:
1904:
1900:
1896:
1892:
1891:
1887:
1883:
1879:
1875:
1871:
1867:
1863:
1859:
1858:
1853:
1849:
1845:
1844:
1839:
1838:
1833:
1825:
1824:
1819:
1818:
1813:
1812:
1806:
1802:
1800:
1799:Peveril Point
1796:
1795:Wealden Group
1792:
1788:
1784:
1780:
1776:
1772:
1771:Mediterranean
1768:
1764:
1760:
1756:
1752:
1748:
1744:
1740:
1736:
1732:
1728:
1724:
1720:
1712:
1707:
1698:
1696:
1691:
1686:
1682:
1678:
1674:
1670:
1663:Palaeobiology
1660:
1658:
1654:
1653:ghost lineage
1650:
1646:
1642:
1638:
1634:
1630:
1626:
1622:
1617:
1615:
1614:ghost lineage
1609:
1602:
1598:
1594:
1590:
1580:
1579:
1571:
1570:
1562:
1561:
1553:
1552:
1544:
1543:
1540:
1539:
1538:
1531:
1530:
1527:
1526:
1523:
1522:
1521:
1520:Abrictosaurus
1514:
1513:
1507:
1506:
1503:
1502:
1494:
1493:
1490:
1489:
1488:
1481:
1480:
1477:
1476:
1473:
1472:
1471:
1464:
1463:
1457:
1456:
1450:
1449:
1446:
1445:
1442:
1441:
1440:
1433:
1432:
1426:
1423:
1422:
1419:
1418:
1410:
1409:
1406:
1405:
1404:
1397:
1396:
1393:
1392:
1389:
1388:
1387:
1380:
1379:
1376:
1375:
1372:
1371:
1370:
1363:
1362:
1356:
1355:
1349:
1346:
1345:
1338:
1337:
1332:
1328:
1324:
1322:
1318:
1314:
1310:
1305:
1303:
1299:
1295:
1294:Abrictosaurus
1291:
1287:
1283:
1279:
1275:
1271:
1267:
1263:
1259:
1255:
1250:
1247:
1243:
1239:
1238:
1237:Abrictosaurus
1233:
1232:
1227:
1223:
1222:Lesothosaurus
1219:
1215:
1214:Lesothosaurus
1211:
1207:
1202:
1200:
1196:
1192:
1188:
1184:
1180:
1176:
1175:
1174:Lesothosaurus
1170:
1169:Fabrosauridae
1166:
1162:
1161:
1156:
1155:
1150:
1149:
1144:
1143:
1142:Hypsilophodon
1138:
1137:
1132:
1131:
1126:
1122:
1118:
1114:
1110:
1102:
1098:
1097:
1096:Lesothosaurus
1091:
1087:
1085:
1082:
1078:
1074:
1070:
1066:
1065:Stegosauridae
1062:
1059:to the clade
1058:
1054:
1050:
1040:
1037:
1030:
1026:
1021:
1012:
1004:
996:
992:
988:
983:
972:
968:
963:
959:
955:
947:
939:
936:
932:
928:
927:
922:
918:
914:
910:
906:
902:
898:
894:
890:
886:
878:
873:
869:
866:
865:Lesothosaurus
862:
858:
857:Hypsilophodon
854:
849:
842:
837:
830:
826:
821:
819:
815:
811:
807:
806:
801:
797:
792:
784:
776:
774:ectopterygoid
768:
760:
752:
744:
736:
729:
720:
711:
709:
705:
700:
698:
694:
693:
688:
684:
680:
676:
672:
668:
664:
660:
656:
652:
648:
644:
640:
632:
629:, originally
628:
627:
621:
617:
615:
611:
607:
603:
599:
595:
591:
587:
583:
579:
575:
571:
570:Scelidosaurus
567:
563:
559:
555:
551:
547:
543:
542:
541:Scelidosaurus
537:
533:
528:
526:
522:
518:
517:Hugh Falconer
513:
505:
499:
495:
494:Ancient Greek
491:
487:
483:
482:
477:
476:
471:
467:
463:
459:
455:
451:
447:
443:
439:
435:
431:
423:
418:
409:
407:
406:Mediterranean
403:
399:
398:
393:
392:
387:
383:
379:
378:Purbeck Group
375:
370:
368:
364:
360:
356:
352:
348:
347:Fabrosauridae
344:
340:
336:
332:
328:
323:
321:
317:
313:
309:
308:
303:
299:
295:
291:
288:
284:
280:
279:
266:
264:
255:
252:
251:Binomial name
248:
244:
243:
237:
234:
233:
229:
225:
220:
219:
212:
209:
208:
205:
199:
196:
195:
192:
186:
183:
180:
179:
176:
173:
170:
167:
166:
163:
160:
157:
156:
153:
150:
147:
146:
143:
140:
137:
136:
131:
126:
122:
118:
113:
109:
104:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
32:
26:
22:
19:
4979:
4921:Silesauridae
4892:
4885:
4877:
4872:Ajkaceratops
4870:
4860:
4848:
4841:
4834:
4827:
4820:
4815:Notoceratops
4813:
4806:
4801:Gongbusaurus
4799:
4794:Geranosaurus
4792:
4785:
4778:
4771:
4764:
4757:
4749:Nomina dubia
4747:
4697:
4585:
4577:
4570:
4565:Parksosaurus
4563:
4556:
4549:
4542:
4534:
4526:
4520:Jeholosaurus
4518:
4510:
4503:
4495:
4487:
4480:
4460:
4453:
4448:Sanxiasaurus
4446:
4439:
4432:
4427:Minimocursor
4425:
4418:
4411:
4404:
4397:
4341:Ankylosauria
4330:Ankylosauria
4317:
4310:
4303:
4296:
4272:
4265:
4258:
4251:
4244:
4237:
4231:
4230:
4223:
4204:Technosaurus
4202:
4196:Soumyasaurus
4194:
4186:
4178:
4172:Pisanosaurus
4170:
4162:
4154:
4147:
4140:
4132:
4126:Ignotosaurus
4124:
4116:
4108:
4101:
4093:
4085:
4077:
4061:Ornithischia
4048:Ornithischia
4011:Ornithischia
3963:
3954:
3920:Ornithischia
3878:
3872:
3866:
3841:
3837:
3831:
3806:
3802:
3798:
3792:
3786:(1): 95–126.
3783:
3779:
3773:
3764:
3760:
3754:
3713:
3707:
3701:
3676:
3672:
3662:
3653:
3649:
3643:
3618:
3614:
3610:
3604:
3579:
3575:
3562:
3556:(1): 93–127.
3553:
3547:
3541:
3514:
3510:
3500:
3487:
3481:
3464:
3458:
3430:
3424:
3399:
3395:
3391:
3385:
3352:
3346:
3333:
3303:(1): 59–64.
3300:
3296:
3292:
3288:
3270:
3264:
3230:
3226:
3198:
3192:
3156:
3150:
3144:
3136:
3114:(10): 1–21.
3111:
3107:
3101:
3060:
3054:
3048:
3023:
3019:
3015:
3009:
2976:
2970:
2934:
2928:
2924:
2882:
2878:
2836:
2830:
2796:
2790:
2777:
2768:
2764:
2758:
2749:
2745:
2723:
2719:
2709:
2684:
2678:
2647:
2612:
2608:
2598:
2589:
2583:
2529:
2447:
2441:
2409:
2403:
2372:
2352:
2348:
2321:
2315:
2309:
2301:
2287:
2254:Eucorethrina
2252:
2246:
2240:
2230:
2224:
2215:
2209:
2203:
2197:
2191:
2185:
2176:
2164:
2158:
2149:
2143:
2137:
2131:
2127:Albionbaatar
2125:
2115:Phascolestes
2113:
2107:
2101:
2097:Amblotherium
2095:
2089:
2087:dryolestoids
2080:
2074:
2068:
2059:
2053:
2049:Peramuroides
2047:
2041:
2032:
2026:
2021:
2014:
2010:Homoeosaurus
2008:
1995:
1989:
1983:
1977:
1971:
1965:
1959:
1953:
1947:
1937:
1933:Hylaeochelys
1931:
1927:Helochelydra
1925:
1919:
1910:
1901:
1888:
1882:crocodilians
1855:
1841:
1835:
1828:
1821:
1815:
1809:
1716:
1710:
1689:
1684:
1680:
1676:
1672:
1668:
1666:
1648:
1644:
1640:
1633:ceratopsians
1618:
1600:
1596:
1592:
1588:
1586:
1535:
1534:
1518:
1517:
1485:
1484:
1468:
1467:
1437:
1436:
1401:
1400:
1384:
1383:
1368:
1367:
1366:
1334:
1330:
1320:
1316:
1312:
1306:
1301:
1297:
1293:
1289:
1281:
1257:
1253:
1251:
1245:
1235:
1229:
1225:
1221:
1217:
1213:
1205:
1203:
1194:
1190:
1186:
1182:
1178:
1172:
1164:
1158:
1152:
1148:Parksosaurus
1146:
1140:
1134:
1128:
1116:
1106:
1100:
1094:
1072:
1057:Franz Nopcsa
1052:
1048:
1046:
1028:
1024:
986:
976:
966:
940:
934:
930:
924:
916:
900:
892:
884:
882:
876:
864:
856:
852:
828:
827:specimen of
822:
813:
809:
803:
799:
795:
727:
725:
703:
701:
696:
690:
682:
670:
636:
630:
624:
613:
605:
593:
590:Peter Galton
581:
569:
565:
550:Prionodontia
545:
539:
535:
529:
520:
485:
481:Megalosaurus
479:
473:
468:, a British
466:Richard Owen
458:Purbeck Beds
440:in southern
427:
421:
401:
395:
389:
385:
373:
371:
362:
350:
342:
326:
325:Originally,
324:
319:
312:Richard Owen
306:
305:
296:of southern
277:
276:
275:
262:
257:
241:
240:
217:
216:
191:Ornithischia
181:
168:
24:
18:
4850:Trimucrodon
4822:Siluosaurus
4780:Fabrosaurus
4773:Craspedodon
4766:Bienosaurus
4609:Ornithopoda
4598:Ornithopoda
4497:Changmiania
4462:Yandusaurus
4399:Agilisaurus
4364:Stegosauria
4353:Stegosauria
4285:Thyreophora
4164:Lutungutali
4134:Kwanasaurus
4087:Asilisaurus
4079:Amanasaurus
3767:(1): 17–31.
3656:(1): 35–55.
3402:: 152–162.
3159:: 182–279.
2979:(1): 1–40.
2687:: 435–489.
2324:(46): 341.
2237:nematoceran
2222:snipe flies
2178:Purbeckodon
2160:Trioracodon
2139:Gerhardodon
2109:Chunnelodon
2061:Kouriogenys
1997:Parviraptor
1917:cryptodires
1890:Apricosiren
1886:salamanders
1852:ankylosaurs
1755:terrestrial
1747:hypersaline
1725:and shelly
1266:ankylosaurs
1262:ornithopods
1187:Fabrosaurus
1160:Fabrosaurus
897:Paul Sereno
714:Description
685:proper. In
596:further in
525:preoccupied
448:as well as
359:ornithopods
335:Stegosauria
331:herbivorous
115:Dentary of
5065:Categories
4904:Other taxa
4843:Tichosteus
4836:Tatisaurus
4641:Ceratopsia
4630:Ceratopsia
4579:Yueosaurus
4551:Orodromeus
4536:Nanosaurus
4455:Xiaosaurus
4434:Nanosaurus
4319:Yuxisaurus
4298:Emausaurus
4274:Tianyulong
4267:Pegomastax
4253:Lycorhinus
4239:Fruitadens
4188:Silesaurus
4180:Sacisaurus
4156:Lewisuchus
4020:see below↓
3968:Dinosauria
3950:Sauropsida
3799:Plagiaulax
3611:Thereuodon
2752:: 145–163.
2726:: 163–188.
2261:References
2235:, and the
2226:Simulidium
2169:, the non-
2166:Triconodon
2145:Plagiaulax
2118:; the non-
2103:Dorsetodon
2082:Thereuodon
2039:peramurans
2028:Durlstodon
2024:eutherians
1985:Purbicella
1961:Durotrigia
1949:Becklesius
1862:amphibians
1811:Durlstodon
1787:Berriasian
1759:lacustrine
1749:and quasi-
1739:freshwater
1727:limestones
1723:sandstones
1719:evaporites
1695:edentulous
1649:Tianyulong
1601:Tianyulong
1597:Fruitadens
1470:Pegomastax
1439:Lycorhinus
1403:Tianyulong
1386:Fruitadens
1321:Tianyulong
1317:Fruitadens
1270:stegosaurs
1189:. All but
1183:Nanosaurus
1136:Nanosaurus
1029:Lycorhinus
981:predentary
926:Lycorhinus
810:Fruitadens
805:Tianyulong
734:premaxilla
651:sandstones
626:Fruitadens
554:Dinosauria
511:serrations
490:Lacertilia
302:Berriasian
294:Cretaceous
269:Owen, 1861
175:Dinosauria
36:Berriasian
4980:Echinodon
4808:Laosaurus
4441:Phyllodon
4232:Echinodon
4118:Gamatavus
3936:Kingdom:
3823:128645686
2351:Iguanodon
2310:Echinodon
2171:mammalian
2151:Sunnyodon
2120:eutherian
2091:Achyrodon
2016:Opisthias
2002:squamates
1991:Saurillus
1903:Celtedens
1843:Iguanodon
1783:Tithonian
1711:Echinodon
1681:Echinodon
1669:Echinodon
1645:Echinodon
1641:Echinodon
1593:Echinodon
1589:Echinodon
1369:Echinodon
1331:Echinodon
1313:Echinodon
1309:cladogram
1298:Echinodon
1290:Echinodon
1258:Echinodon
1254:Echinodon
1246:Echinodon
1226:Echinodon
1218:Echinodon
1206:Echinodon
1195:Echinodon
1191:Echinodon
1179:Echinodon
1165:Echinodon
1130:Laosaurus
1117:Echinodon
1101:Echinodon
1081:solemydid
1073:Echinodon
1053:Echinodon
1049:Echinodon
1025:Echinodon
987:Echinodon
967:Echinodon
935:Echinodon
917:Echinodon
893:Echinodon
885:Echinodon
877:Echinodon
829:Echinodon
825:lectotype
814:Echinodon
800:Echinodon
728:Echinodon
704:Echinodon
697:Echinodon
683:Echinodon
671:Echinodon
667:vertebrae
631:Echinodon
614:Echinodon
606:Echinodon
594:Echinodon
582:Echinodon
566:Echinodon
546:Iguanodon
536:Echinodon
534:to group
521:Echinodon
475:Iguanodon
422:Echinodon
402:Echinodon
386:Echinodon
374:Echinodon
363:Echinodon
351:Echinodon
343:Echinodon
327:Echinodon
320:Echinodon
278:Echinodon
235:Species:
218:Echinodon
148:Kingdom:
142:Eukaryota
25:Echinodon
4989:Wikidata
4759:Alocodon
4737:See also
4260:Manidens
4103:Eocursor
4095:Diodorus
3944:Chordata
3942:Phylum:
3938:Animalia
3746:18504005
3738:23494201
3377:86775565
3325:55161519
3093:22636871
3085:21452054
3001:86728076
2771:: 52–90.
2639:19846460
2560:23166462
2472:84613826
2248:Brodilka
2183:docodont
2055:Magnimus
1906:and the
1857:Nuthetes
1823:Nuthetes
1814:(left),
1767:lagoonal
1743:brackish
1657:cerapods
1607:cingulum
1487:Manidens
1336:Nuthetes
1125:Triassic
1010:coronoid
1002:foramina
889:diastema
861:textured
841:anterior
766:palatine
750:lacrimal
647:Colorado
523:was not
397:Nuthetes
391:Owenodon
290:dinosaur
197:Family:
162:Chordata
158:Phylum:
152:Animalia
138:Domain:
117:paratype
5035:4127061
5022:1285235
5009:4823157
4995:Q134292
4887:Jakapil
3948:Class:
3846:Bibcode
3718:Bibcode
3681:Bibcode
3623:Bibcode
3584:Bibcode
3519:Bibcode
3404:Bibcode
3357:Bibcode
3305:Bibcode
3235:Bibcode
3116:Bibcode
3065:Bibcode
3028:Bibcode
2981:Bibcode
2939:Bibcode
2907:4196759
2887:Bibcode
2861:4193262
2841:Bibcode
2801:Bibcode
2689:Bibcode
2630:2842649
2551:3491919
2531:ZooKeys
2452:Bibcode
2133:Bolodon
2076:Tinodon
2043:Peramus
1878:mammals
1870:lizards
1866:turtles
1721:, thin
1701:Ecology
1621:Xu Xing
1119:within
1084:turtles
1063:within
1019:alveoli
995:ventral
991:lateral
835:foramen
782:dentary
742:maxilla
657:of the
454:dentary
450:maxilla
442:England
436:on the
310:by Sir
298:England
260:†
239:†
210:Genus:
3821:
3744:
3736:
3437:
3375:
3323:
3091:
3083:
2999:
2905:
2879:Nature
2859:
2832:Nature
2637:
2627:
2558:
2548:
2470:
2239:flies
1897:, the
1874:snakes
1832:Dorset
1763:saline
1751:marine
1735:shales
1690:et al.
1302:et al.
1282:et al.
1035:enamel
971:medial
953:palate
921:canine
643:Fruita
498:εχινος
382:Dorset
339:turtle
5048:38717
5017:IRMNG
3964:Clade
3955:Clade
3819:S2CID
3742:S2CID
3572:(PDF)
3373:S2CID
3343:(PDF)
3321:S2CID
3089:S2CID
2997:S2CID
2903:S2CID
2857:S2CID
2787:(PDF)
2675:(PDF)
2468:S2CID
945:orbit
847:fossa
790:teeth
758:jugal
556:. In
538:with
446:teeth
283:genus
281:is a
182:Clade
169:Clade
5004:GBIF
4662:see
4639:see
4607:see
4512:Haya
4505:Fona
4362:see
4339:see
3997:see
3734:PMID
3435:ISBN
3081:PMID
2635:PMID
2556:PMID
2251:and
2229:and
2163:and
2148:and
2112:and
2079:and
2058:and
2031:and
2013:and
1936:and
1908:frog
1880:and
1765:and
1733:and
1731:marl
1635:and
1625:2006
1319:and
1307:The
1286:2011
1278:2008
1268:and
1210:1991
1185:and
1163:and
1151:and
1133:and
1113:1971
1077:1959
1069:1928
993:and
977:The
929:and
913:2012
907:and
778:and
687:2009
679:2006
675:2002
663:1984
602:1857
598:1978
578:1876
558:1888
544:and
532:1874
504:ὀδών
478:and
462:1861
452:and
341:'s.
228:1861
224:Owen
41:PreꞒ
3854:doi
3811:doi
3726:doi
3714:100
3689:doi
3631:doi
3592:doi
3527:doi
3469:doi
3465:183
3412:doi
3365:doi
3313:doi
3243:doi
3203:doi
3161:doi
3157:162
3124:doi
3073:doi
3036:doi
2989:doi
2947:doi
2895:doi
2883:239
2849:doi
2837:234
2809:doi
2750:132
2697:doi
2625:PMC
2617:doi
2613:277
2546:PMC
2536:doi
2460:doi
2326:doi
1797:at
1284:in
1111:in
1067:in
1023:of
891:in
633:sp.
380:of
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