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1677:. Ideally winemakers want to add enough inoculum to have a viable cell population density of 5 million cells per milliliter. The exact amount of freeze-dried culture varies by manufacturer and strain of yeast but it is often around 1 gram per gallon (or 25 grams per 100 liters). Wines that could have potentially problematic fermentation (such as high sugar level late harvest or botryized wines) may have more yeast added.
349:
483:
761:
1983:
1966:
predictability of fermentation due to the high level of survival factors that cultured yeast are assured of having without necessarily needing to expose the wine to additional levels of oxygen. Winemakers using "ambient" yeasts that are resident in their winery may not have this same assurance of survival factors and may need to compensate with other winemaking techniques.
2218:, for metabolism. Additionally, Brett can produce a wide range of by-products that could influence the wine beyond just the 4-EP and 4-EG compounds previously discussed. Many of these compounds, such as the "footprints" of the 4-EP and 4-EG, will still remain in the wine even after yeast cells die and are removed by racking and sterile filtration.
225:, winemakers did not know the mechanism that somehow converted sugary grape juice into alcoholic wine. They could observe the fermentation process which was often described as "boiling", "seething" or the wine being "troubled" due to release of carbon dioxide that gave the wine a frothy, bubbling appearance. This history is preserved in the
240:", would uncover the connection between microscopic yeast cells and the process of the fermentation. It was Pasteur who discovered that yeast converted sugars in the must into alcohol and carbon dioxide, though the exact mechanisms of how the yeast would accomplish this task was not discovered till the 20th century with the
288:
strains that accentuate desirable features in wine, such as aromatic compounds, mouthfeel, and fermentation kinetics. This commercial availability of yeast strains has revolutionized the art of winemaking by allowing for more precise control over the fermentation process and the resultant wine's character.
319:) in the grape must into alcohol. The yeast accomplishes this by utilizing glucose through a series of metabolic pathways that, in the presence of oxygen, produces not only large amounts of energy for the cell but also many different intermediates that the cell needs to function. In the absence of oxygen (
1680:
Similarly, re-hydration procedures will also vary depending on the manufacturer and winery. Yeast is often inoculated in a volume of water or grape must that is 5â10 times the weight of the dry yeast. This liquid is often brought to temperature of 40 °C (104 °F) prior to the introduction of
1689:
The temperature of the starter culture is then slowly reduced, often by the graduated addition of must to get within 5â10 °C (41â50 °F) of the must that the culture will be added to. This is done to avoid the sudden cold shock that the yeast cells may experience if the starter culture was
1161:
Some distinct difference among various strains include the production of certain "off-flavor" and aromas that may be temporary (but producing a "stinky fermentation") or could stay with the wine and either have to be dealt with through other winemaking means (such as the presence of volatile sulfur
1153:
identified. Not all of the strains are suitable for winemaking and even among the strains that are, there is debate among winemakers and scientists about the actual magnitude of differences between the various strains and their potential impact on the wine. Even among strains that have demonstrated
335:
needed to keep metabolism going. It is through this process of fermentation that ethanol is released by the yeast cells as a waste product. Eventually, if the yeast cells are healthy and fermentation is allowed to run to the completion, all fermentable sugars will be used up by the yeast with only
1685:
components escape the cell. Re-hydration at lower temperatures can greatly reduce the viability of the yeast with up to 60% cell death if the yeast is re-hydrated at 15 °C (59 °F). The culture is then stirred and aerated to incorporate oxygen into the culture which the yeast uses in the
1457:
wild yeast, these genera of wild yeasts have very low tolerance to both alcohol and sulfur dioxide. They are capable of starting a fermentation and often begin this process as early as the harvest bin when clusters of grapes get slightly crushed under their own weight. Some winemakers will try to
1550:
When winemakers select a cultured yeast strain, it is largely done because the winemaker wants a predictable fermentation taken to completion by a strain that has a track record of dependability. Among the particular considerations that are often important to winemakers is a yeast's tendency to:
1334:
In winemaking, the term "wild yeast" has multiple meanings. In its most basic context, it refers to yeast that has not been introduced to the must by intentional inoculation of a cultured strain. Instead, these "wild yeasts" often come into contact with the must through their presence on harvest
287:
In modern winemaking, winemakers have the option to select from a diverse range of yeast strains, each offering distinct characteristics that influence the wine's sensory profile. These strains are readily available for purchase from specialized suppliers. Winemakers can now easily access yeast
1681:
the yeast (though some yeast strains may need temperatures below 38 °C (100 °F)) to allow the cells to disperse easily rather than clump and sink to the bottom of the container. The heat activation also allows the cells to quickly reestablish their membrane barrier before soluble
1965:
Cultured yeasts that are freeze-dried and available for inoculation of wine must are deliberately grown in commercial labs in high oxygen/low sugar conditions that favor the development of these survival factors. One of the reasons that some winemakers prefer using inoculated yeast is the
1961:
and levels of alcohol in the wine. As a waste product of its own metabolism, alcohol is actually very toxic to yeast cells. Yeast with weak survival factors and lacking sterols may succumb to these conditions before fermenting a wine to complete dryness, leaving a stuck fermentation.
1706:
In order to successfully complete a fermentation with minimum to no negative attributes being added to the wine, yeast needs to have the full assortment of its nutritional needs met. These include not only an available energy source (carbon in the form of sugars such as glucose) and
378:
and a slightly sweet taste without increasing the alcohol level of the wine, some winemakers try to intentionally favor conditions that would promote glycerol production in wine. This includes selecting yeast strains that favor glycerol production (or allowing some wild yeast like
1520:
wild yeast will have a role in beginning the fermentation of virtually every wine but for the wineries that choose to allow these yeasts to continue fermenting versus minimizing their influence do so with the intent of enhancing complexity through bio-diversity. While these
1647:
but a negative attribute for many other wines), enhancement of a wine's color or certain varietal characteristics by enzymes in the yeast cells and other metabolic products produced by the yeast, foaming and flocculation tendencies, yeasticidal properties (a trait known as
532:
available in the must or by the decomposition of dead yeast cells by other yeast that releases sulfur-containing amino acids that are further broken down by the yeast. The latter often happens with wines that sit in contact with their lees for long periods of time between
1508:
in order to encourage the sustained presence of favorable strains. But compared to inoculated yeast, these ambient yeasts hold the risk of having a more unpredictable fermentation. Not only could this unpredictability include the presence of off-flavors/aromas and higher
1166:) or leave a faulty wine. Another difference includes the "vigor" or speed of fermentation (which can also be influenced by other factors beyond yeast selection) with some yeast strains having the tendency to do "fast ferments" while others may take longer to get going.
1915:
by breaking down the cell, including the amino acids. This autolysis of the cell provides an available nitrogen source for the still-fermenting and viable yeast cells. However, this autolysis can also release sulfur-link compounds (such as the breakdown of amino acid
440:
and then to the more neutral-smelling 2,3-Butanediol. Many beer and winemakers who have a wine with too much "butteriness" will often "pitch" fresh yeast cultures into the no longer fermenting tank so that the yeast will consume the diacetyl and reduce the
1639:). These strains are tested in laboratories to determine a strain's vigor, sulfur dioxide and alcohol tolerance, production levels of acetic acid and sulfur compounds, ability to re-ferment (positive for sparkling wine but a negative attribute for sweet
1672:
and packaged for commercial use. Prior to their addition into must, these yeasts need to be re-hydrated in "starter cultures" that must be carefully monitored (particularly in regards to temperature) to ensure that the yeast cells are not killed off by
473:
acids can also be formed depending on the yeast strain. Most countries have wine laws setting the legal limit of volatile acidity, usually expressed as acetic acid, to 1200â2000 mg/L. Acetic acid can also lead to the development of the wine fault
603:
The lees left over from the secondary fermentation of sparkling wine can be seen on the bottom side of this bottle being inspected. Eventually this wine will go through riddling to collect the lees in the neck, where it will be removed prior to
1466:
with skin contact. Other winemakers may allow the wild yeasts to continue fermenting until they succumb to the toxicity of the alcohol they produce which is often between 3â5% alcohol by volume and then letting either inoculated or "ambient"
1940:
process done in the absence of oxygen, early exposure of the yeast to oxygen can be a vital component in the successful completion of that fermentation. This is because oxygen is important in the synthesis of cell "survival factors" such as
300:
In the absence of oxygen, yeast cells will take the pyruvate produced by glycolysis and reduce it into acetaldehyde which is further reduced into ethanol "recharging" the NAD+ co-enzymes that is needed for various metabolic processes of the
356:
While the production of alcohol is the most noteworthy by-product of yeast metabolism from a winemaking perspective, there are a number of other products that yeast produce that can be also influence the resulting wine. This includes
369:
needed to continue other metabolic activities. This is usually produced early in the fermentation process before the mechanisms to reduce acetaldehyde into ethanol to recharge NADH becomes the cell's primary means of maintaining
1998:
can cover the surface of a wine with a film layer that not only consumed most of the free sulfur dioxide available to protect the wine but also produces high levels of acetic acid that will contribute to volatile acidity in a
267:
were identified. The differences between the vast majority of these strains are mostly minor, though individual winemakers will develop a preference for particular strains when making certain wines or working with particular
1973:
yeasts often need a much greater exposure to oxygen in order to build up survival factors which is why many of these yeasts are often found living oxidatively as "film yeast" on the surface of wines in tanks or barrels.
1525:
ferment glucose and fructose into alcohol, they also have the potential to create other intermediates that could influence the aroma and flavor profile of the wine. Some of these intermediates could be positive, such as
98:
or neutral spirits to kill off the yeast cells. If fermentation is unintentionally stopped, such as when the yeasts become exhausted of available nutrients and the wine has not yet reached dryness, this is considered a
2209:
Once Brett is in a winery, it is very difficult to control even with strict hygiene and the discarding of barrels and equipment that has previously come into contact with "Bretty" wine. This is because many species of
1879:) and even the remnant of dead or extracted yeast cells such that the fermenting yeast can break down to mine for available nitrogen and nutrients. One historical winemaking tradition that is still practiced in some
513:
will produce higher amounts that contributes to the characterized "aldehydic" aromas of
Sherries. In the presence of oxygen, yeast can convert some of the ethanol presence in the wine back into acetaldehyde creating
387:
to ferment), increased oxygen exposure and aeration as well as fermenting at higher temperatures. Glycerol production is also encouraged if most available acetaldehyde is made unavailable by binding with
155:. While additions of sulfur dioxide (often added at the crusher) may limit some of the wild yeast activities, these yeasts will usually die out once the alcohol level reaches about 15% due to the
151:. These yeasts often begin the fermentation process almost as soon as the grapes are picked when the weight of the clusters in the harvest bins begin to crush the grapes, releasing the sugar-rich
2681:
1534:-like aroma. However, as with ambient yeasts, the products of these yeasts can be very unpredictable â especially in terms of the types of flavors and aromas that these yeasts can produce.
478:
which is characterized by a "nail polish remover" smell. However, small amounts of acetic acid are actually beneficial for the yeast as they use them to synthesis lipids in the cell membrane.
1343:
that have taken residence in these places over the years, sometimes being previously introduced by inoculation of prior vintages. In this context, these wild yeasts are often referred to as
352:
If a
Chardonnay has too much "buttery" diacetyl notes, winemakers may add fresh yeast to the wine to consume the diacetyl and reduce it to the more neutral-smelling fusel oil 2,3-Butanediol.
2058:
on top of the wine in the tank of barrel. Allowed to go unchecked, these yeasts can rapidly deplete the available free sulfur compounds that keeps a wine protected from oxidation and other
2089:
Growth of many unfavorable wild yeasts is generally slowed at lower cellar temperatures, so many winemakers who wish to inhibit the activities of these yeasts before the more favorable
1702:
Diammonium phosphate (or DAP) is a common additive that provides two necessary nutrients for yeast to have a healthy and sustained fermentation – nitrogen and phosphate.
1690:
added directly to the must itself which can kill up to 60% of the culture. Additionally, surviving cells exposed to cold shock tend to see an increase in hydrogen sulfide production.
1410:
of wild yeasts per berry could exist in a typical vineyard. These yeasts can be carried by air currents, birds and insects through the vineyard and even into the winery (such as by
1546:
Some winemakers favor the use of freeze-dried cultured yeast (left) and yeast nutrients (right) because of their relative predictability in beginning and completing a fermentation.
686:
Most of the benefits associated with lees contact deals with the influence on the wine of the mannoproteins released during the autolysis of the yeast cells. Composed primarily of
2082:
and shifting the pH of wine upwards to levels that make the wine prone to attack by other spoilage microbes. Commonly called "film yeast", these yeasts are distinguished from the
502: â While most of the acetaldehyde produce gets reduced to ethanol or is bound by sulfur dioxide, concentrations between 50 and 100 mg/L can remain in the wine. The
2155:), a limited amount of these compounds could be considered a positive attribute that adds to the complexity of wine. To other winemakers and with other wine styles (such as
1000:
in addition to being used in brewing and breadmaking) because of the generally reliable and positive attributes it can bring to the wine. These yeasts will usually readily
1442:, itself, is actually quite rarely found in the vineyard or on the surface freshly harvested wine grapes unless the winery frequently reintroduced winery waste (such as
1308:
1065:
2574:
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wild yeasts carries both potential benefits and risk. Some winemakers feel that the use of resident/indigenous yeast helps contribute to the unique expression of
2035:. Also any yeast can have a low tolerance to nutritional deficiencies, temperature fluctuation or extremes and excessive or low sugar levels that may lead to a
679:
and other compounds that were present in the yeast cell walls and membranes. This stirring also helps avoid the development of reductive sulfur compounds like
2147:(4-EG), that can have a wine being described as smelling like a "barnyard", "wet saddle" or "band-aid". To some winemakers and with some wine styles (such as
671:
produced in many wine regions across the globe. Typically when wines are left in contact with their lees, they are regularly stirred in order to release the
236:
was tasked by the French government to study what made some wines spoil. His work, which would later lead to
Pasteur being considered one of the "Fathers of
447: â Like glycerol, this is often formed early in fermentation. Usually found in concentrations of 500â1200 mg/L, it is a minor acid in the overall
1478:
Wineries that wish to cultivate an "in-house" ambient yeast strain will often recycle the leftover pomace of previous vintages as compost in the vineyard.
950:, very alcohol-tolerant and can grow in wines up to 18% v/v. Additionally this yeast can survive in extremely high sugar levels (as much as 60% w/w or 60
1867:
Many of these nutrients are available in the must and skins of the grapes themselves but sometimes are supplemented by winemakers with additions such as
491:
2675:
748:
necessary for the style. A small amount of sugared liquid is added to individual bottles, and the yeast is allowed to convert this to more alcohol and
1169:
Another less measurable difference that are subject to more debate and questions of winemakers preference is the influence of strain selection on the
1500:
and highly regarded estates will often tout the quality of their resident "chateau" strains. To this extent, wineries will often take the leftover
672:
1102:
While some strains of yeast may influence the sensory characteristics and aromas of young wine, these differences seem to fade as the wine ages.
414:
in the must by enzymes of the yeast. More commonly found in red wines than white but only in very small amounts between 20 and 200 mg/L.
1513:
but also the potential for a stuck fermentation if the indigenous yeast strains are not vigorous enough to fully convert all the sugars.
683:
and hydrogen sulfide that can appear if the lees layer is more than 10 cm (3.9 in) thick and undisturbed for more than a week.
461:
that can make a wine taste unbalanced and overly acidic. While acetic acid is the main volatile acid produced by yeast, trace amounts of
116:
which has been favored due to its predictable and vigorous fermentation capabilities, tolerance of relatively high levels of alcohol and
2993:
698:
as the cell wall breaks down. Not only does the release of mannoproteins impart sensory changes in the wine but they can contribute to
1623:
that have been identified and plated from wineries across the world (including notable producers from well-known wine regions such as
524: â Often produced by yeast during fermentation because of a nitrogen deficiency in the must. This can be done by a reduction of
549:
are known to produce higher levels of hydrogen sulfides than other strains, particularly if the must has some nutrient deficiencies.
2891:
1367:. Wineries that often solely rely on these "in-house" strains will sometimes market their wines as being the product of wild or
628:
and age. During the time that the wine spends in contact with the lees, a number of changes can impact the wine due to both the
1217:. Similarly, though potentially to a much smaller extent, other varieties could be influenced by hydrolytic enzymes working on
241:
1936:
meaning that they can exist in both the presence and absence of oxygen. While fermentation is traditionally thought of as an
616:. During fermentation, the first significant racking which removes the bulk of dead yeast cells is often referred to as the
272:. Some of these differences include the "vigor" or speed of fermentation, temperature tolerance, the production of volatile
2713:
2571:
1788:. A deficiency of this vitamin could lead into increase hydrogen sulfide production with off-aromas in the resulting wine.
1154:
distinctive difference when compared among young wines, these differences seem to fade and become less distinctive as the
804:
text under its asexual classification though some scientific and winemaking texts may describe specific species (such as
647:
The process of leaving the wine to spend some contact with the lees has a long history in winemaking, being known to the
2011:" and aromas that can be the by-product of some "wild yeast" fermentation such as those by species within the genera of
179:
is a species of yeast that can tolerate alcohol levels of 17â20% and is often used in fortified wine production such as
2093:
yeast kick in, will often chill their must, such as the practice of "cold soaking" the must during a pre-fermentation
2624:
Science and
Civilisation in China, Volume 6 Biology and Biological Technology, Part 5: Fermentations and Food Science
1795:
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Preparing a yeast starter culture and gradually cooling the culture down to the must temperature by adding some wine
1660:
217:
French scientist Louis
Pasteur discovered the connection between microscopic yeast and the process of fermentation.
752:. The lees are then ricked into the neck of the bottle, frozen, and expelled via pressure of the carbonated wine.
86:. This can be achieved by dropping fermentation temperatures to the point where the yeast are inactive, sterile
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835:. Other genera of yeast that can be involved in winemaking (either beneficially or as the cause of potential
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2023:
can be behind some wine faults with some strains of the yeast known to produce higher than ideal levels of
608:
When yeast cells die, they sink to the bottom of the fermentation vessel where they combine with insoluble
74:, the higher the potential alcohol level of the wine if the yeast are allowed to carry out fermentation to
1001:
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1739:) that serve as important growth and survival factors. Among the other nutritional needs of wine yeast:
1564:
Have a high sulfur dioxide tolerance but low production of sulfur compounds such as hydrogen sulfide or
1378:
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1458:"knock out" these yeasts with doses of sulfur dioxide, most often at the crusher before the grapes are
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molecules in the wine, but it would take a substantial amount of sulfur dioxide addition (far beyond
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Produce minimum foaming during fermentation which may create difficulties for cap management during
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conditions that can develop if the lees are not aerated or stirred (a process that the French call
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259:(as opposed to circular) shape of the cells. Throughout the 20th century, more than 700 different
19:
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sherry yeast that are usually welcomed by winemakers in producing the delicate fino-style wines.
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of other wines into a newly fermenting batch of wine as an additional food source for the yeast.
1072:
964:
1330:
Fruit flies are a common vector that transports ambient or "wild" yeast strains within wineries.
1296:, though the commercial flor yeast available for inoculation is often from different species of
4382:
3131:
1954:
1755:
1652:") and tolerance for nutritional deficiencies in a must that may lead to a stuck fermentation.
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1924:
that can contribute to a "stinky fermentation" or later development into various wine faults.
820:. Unless otherwise noted, this article will commonly refer to the asexual form of wine yeast.
690:
and proteins, with some glucose, mannoproteins are often bound in the cell wall of yeast with
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of the yeast cell membrane which becomes critical as the yeast becomes exposed to increasing
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Completely utilize all fermentable sugars with a predictable sugar-to-alcohol conversion rate
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on the wine, in limited amounts, as added complexity, many winemakers view the presence of
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in the 2nd century BC. Today the practice is widely associated with any red wines that are
91:
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902:), usually the most common "wild yeast" found in the vineyard. Some species are known as "
8:
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396:) to prolong glycerol production beyond just these very nascent stages of fermentation.
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it releases alcohol and carbon dioxide (seen here as the foaming bubbles) as byproducts.
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can usually ferment a wine up to 10â11% alcohol levels before they die out. Sometimes
2101:, will not be inhibited and may even thrive during an extended period of cold soaking.
2079:
2036:
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1632:
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Quickly begin fermentation, out-competing other "wild yeasts" for nutrients in the must
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equipment, transport bins, the surface winemaking equipment and as part of the natural
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100:
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that contribute to off aromas and wine faults. Some commercial yeast strains, such as
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The process of fermentation at work on Pinot noir. As yeast consume the sugar in the
2245:"From Vines to Wines: The Complete Guide to Growing Grapes and Making Your Own Wine"
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2003:
Either directly or indirectly, wine yeast can be a culprit behind a wide variety of
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Have an alcohol-tolerance up to 15% or even higher depending on the winemaking style
1411:
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is rarely the only yeast species involved in a fermentation. Grapes brought in from
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at temperatures between 4â15 °C (39â59 °F). Though some species, such as
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1920:) which can combine with other molecules and react with alcohol to create volatile
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strain for nutrients and even inhibit it due to the high levels of acetic acid,
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that may also be produced) with the side-effect of substantially decreasing the
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that are more prevalent in particular strains. Other aromatic varieties such as
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attack. The presence of these yeasts is often identified by elevated levels of
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420: â Formed by the decomposition of amino acids by the yeast. This includes
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requires a second fermentation to occur in the bottle in order to produce the
537:. In the presence of alcohol, hydrogen sulfide can react with ethanol to form
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between 2.8 and 4. Despite its widespread use which often includes deliberate
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Second
Edition pp. 3â28 Springer Science and Business Media, New York (2010)
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can use a wide variety of carbon sources in wine and grape must, including
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whose presence in a wine may be viewed by different winemakers as either a
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78:. Sometimes winemakers will stop fermentation early in order to leave some
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2688:. International Journal of Food Microbiology 86 (2003). pp. 169â180.
2024:
2004:
1775:
1716:
1682:
1600:
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1414:). The most common wild yeasts found in the vineyard are from the genera
997:
996:(sugar mold) is favored for winemaking (for both grapes as well as other
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2131:) in their wineries as a negative influence that needs to be controlled.
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1035:) which is usually present in small amount in wines as residual sugars.
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and lees from winemaking and return them to the vineyard to be used as
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1186:
1146:
1122:
969:
791:
680:
668:
656:
429:
417:
332:
324:
202:
160:
107:
4347:
4068:
2971:
1542:
4851:
4816:
4786:
4646:
4357:
4119:
4013:
3976:
3754:
3684:
3561:
3541:
3476:
3395:
3270:
3109:
2743:
2059:
1840:
1761:
1743:
1724:
1416:
1402:
genera of yeasts that are present in the vineyard, on the surface of
1387:
1155:
1126:
1118:
1111:
1032:
1012:
894:
790:(or "imperfect" form) while others may be classified by their sexual
771:
766:
737:
609:
542:
393:
389:
226:
184:
138:
4641:
4417:
3616:
1385:èéșŽ) wild yeast with rice and various herbs, including the poisonous
1098:
786:. Therefore, some winemaking yeasts are classified by their asexual
4876:
4871:
4836:
4826:
4736:
4671:
4564:
4503:
4482:
4467:
4317:
4018:
3881:
3824:
3804:
3704:
3679:
3646:
3531:
3390:
3385:
3375:
3290:
3275:
3260:
2156:
1917:
1908:
1904:
1828:
1809:
1771:
1571:
1269:
1261:
1202:
1190:
1170:
988:
as seen under a
Differential Interference Contrast (DIC) microscope
787:
660:
579:
515:
425:
403:
358:
315:
The primary role of yeast is to convert the sugars present (namely
156:
2637:
Nan-fang ts'ao-mu chuang: a fourth century flora of
Southeast Asia
2501:
1209:
may also be influenced by yeast strains containing high levels of
1093:
195:
sugar levels. Another common yeast involved in wine production is
4846:
4766:
4751:
4696:
4609:
4589:
4544:
4529:
4513:
4487:
4452:
4412:
4402:
4352:
4297:
4174:
4159:
4078:
4058:
4023:
3956:
3556:
3516:
3496:
3461:
3420:
3345:
3104:
3099:
3094:
2942:
2748:
2215:
2055:
1950:
1885:
1863:
from a previous fermentation is added to a newly fermenting wine.
1824:
1732:
1728:
1720:
1712:
1604:
1505:
1488:
1226:
1194:
1028:
1020:
1016:
1008:
1004:
939:
860:(Teleomorphs for different species from several genera including
832:
816:
714:
of the wine as well as decrease the perception of bitterness and
687:
587:
575:
534:
529:
525:
482:
437:
411:
348:
337:
316:
256:
59:
4539:
2550:
Winery
Technology & Operations A Handbook for Small Wineries
1855:
One traditional way of providing nutrients for the yeast is the
361:
which is produced when an intermediate of the glycolysis cycle (
4886:
4811:
4741:
4721:
4716:
4676:
4387:
4312:
4307:
4292:
4269:
4254:
4229:
4219:
4189:
4179:
4169:
4033:
3998:
3988:
3946:
3941:
3921:
3901:
3854:
3829:
3819:
3669:
3536:
3526:
3506:
3481:
3410:
3285:
2907:
2870:
2043:
2032:
1921:
1890:
1832:
1767:
1501:
1447:
1428:
1277:
1182:
1121:. This sequencing helped confirm the nearly century of work by
1115:
862:
760:
571:
510:
433:
273:
148:
95:
823:
The most common yeast generally associated with winemaking is
555: â Along with acetaldehyde, this compound can react with
229:
of the word "yeast" itself which essentially means "to boil".
4621:
4427:
4377:
4332:
4322:
4287:
4199:
4073:
4008:
3951:
3814:
3789:
3784:
3769:
3749:
3734:
3714:
3699:
3606:
3596:
3591:
3551:
3546:
3521:
3491:
3456:
3451:
3415:
3400:
3335:
3205:
3119:
2691:
1799:
1785:
1764: â important for the uptake and utilization of phosphate
1336:
1142:
640:). The length of time that a wine spends on its lees (called
633:
371:
188:
51:
43:
40:
1758:
which the cell uses for transferring energy for metabolism).
1268:
well, allowing the dead yeast cells to be removed easily by
632:(or self-metabolize) of the dead yeast cells as well as the
343:
323:), the cell will continue some metabolic functions (such as
136:
are usually teeming with a variety of "wild yeast" from the
4534:
4194:
3779:
3759:
3689:
3641:
3636:
3571:
3511:
3114:
2083:
1844:
1836:
1655:
1644:
1531:
1406:
and of the grapes themselves. Anywhere from 160 to 100,000
1285:
1281:
1234:
1138:
951:
503:
192:
152:
36:
24:
1982:
251:
was first identified in late 19th century enology text as
4761:
4367:
3966:
3626:
2163:), the presence of any Brett will be considered a fault.
1903:
can assimilate nitrogen from both inorganic (ammonia and
1798:(NAD+), a co-enzyme that is important in maintaining the
1643:), development of surface film on the wine (positive for
205:
or in limited quantities as an added note of complexity.
2435:
2433:
2431:
2429:
2186:
already present in a wine that has been inoculated with
2139:(or "Brett") produces very distinctive aroma compounds,
644:) will depend on the winemaking style and type of wine.
2672:
P. Romano, C. Fiore, M. Paraggio, M. Caruso, A. Capece
2529:
pp. 97â114 Kluwer
Academic Publishers, New York (1999)
2443:
pp. 281â90 Kluwer Academic Publishers, New York (1999)
2427:
2425:
2423:
2421:
2419:
2417:
2415:
2413:
2411:
2409:
1754:(an important component of the cell membrane) and ATP (
1398:
Another use of the term "wild yeast" refers to the non-
1321:
1181:. It is believed that these wines can be influenced by
1149:. Today there are several hundred different strains of
327:) but will rely on other pathways such as reduction of
121:
2388:
2386:
2384:
2382:
2380:
2378:
2376:
2374:
2372:
2370:
2368:
1911:). As yeast cells die, enzymes within the cells begin
1668:
Pure culture yeasts that are grown in a lab are often
782:
includes classification of yeast species depending on
340:
leaving behind a negligible amount of residual sugar.
4916:
2676:
Function of yeast species and strains in wine flavour
2652:
p. 80 The Wine Appreciation Guild Third Edition 2009
2366:
2364:
2362:
2360:
2358:
2356:
2354:
2352:
2350:
2348:
1244:
some winemakers select strains (such as one known as
755:
2594:
2592:
2467:
Online Etymology Dictionary Accessed: May 31st, 2012
2406:
2323:
2321:
2319:
2317:
2315:
2313:
2311:
2309:
2307:
800:(or "Brett") that is usually referenced in wine and
3182:
2305:
2303:
2301:
2299:
2297:
2295:
2293:
2291:
2289:
2287:
612:, grape seeds, skin and pulp fragments to form the
2345:
1693:
1377:has the earliest description of winemaking using "
2589:
1859:method where the leftover grape skins and pomace
1802:balance of the cell as well as in the process of
1784: â involved in the metabolism of sugars and
794:(or "perfect" form). A common example of this is
764:Film yeast on the surface of wine in a barrel of
35:is the most important element that distinguishes
4940:
2284:
120:as well as its ability to thrive in normal wine
1977:
1907:) and organic forms (amino acids, particularly
1280:production, the surface film of yeast known as
1094:Influences of different strains on fermentation
567:) that can enhance the color of some red wines.
2113:While some wine regions view the influence of
331:into ethanol (fermentation) to "recharge" the
232:In the mid-19th century, the French scientist
16:Yeasts used for alcoholic fermentation of wine
2987:
2707:
2622:Joseph Needham and Huang Hsing-Tsung (2000),
2552:pp. 67â74 The Wine Appreciation Guild (1996)
2525:B. Zoecklein, K. Fugelsang, B. Gump, F. Nury
2439:B. Zoecklein, K. Fugelsang, B. Gump, F. Nury
1871:(DAP), freeze-dried micro-nutrients (such as
1339:of a winery. Very often these are strains of
1173:flavors of certainly grape varieties such as
280:) and other compounds that may influence the
2470:
2066:, particularly acetic acid. Some strains of
1015:and metabolize glucose, sucrose, raffinose,
968:found in moist cellars that can contaminate
962:, particularly the "black yeast" species of
424:which is formed by yeast that are consuming
321:and sometimes even in the presence of oxygen
4455:and infused distilled drinks by ingredients
2521:
2519:
1046:that are involved with winemaking include:
374:balance. As glycerol contributes increased
3853:
3014:
2994:
2980:
2714:
2700:
954:) and is very resistant to sulfur dioxide.
490:are caused by special yeast native to the
1292:sherries comes from different strains of
934:, the only wine yeast that reproduced by
784:the presence or absence of a sexual phase
725:
344:Other compounds in wine produced by yeast
2516:
2175:between tanks and even nearby wineries.
2108:
2070:will metabolize acetic acid (as well as
1981:
1850:
1697:
1659:
1656:Re-hydrating freeze dried yeast cultures
1541:
1473:
1438:being the most dominant species by far.
1325:
1097:
980:
759:
598:
481:
347:
295:
212:
18:
3001:
2892:Clarification and stabilization of wine
2544:
2542:
914:that can kill off sensitive strains of
563:to create a more stable color pigment (
457: â Considered a main component of
82:and sweetness in the wine such as with
4941:
2639:, The Chinese University Press, p. 59.
2266:
2264:
2262:
2260:
2031:and volatile sulfur compounds such as
1686:synthesis of needed survival factors.
1284:used to make the distinctive style of
106:The most common yeast associated with
4450:
3852:
3181:
3013:
2975:
2695:
2642:
2626:, Cambridge University Press, p. 183.
2239:
2237:
2235:
2233:
2231:
2007:. These can include the presence of "
1927:
1794: â involved in the synthesis of
1770: â involved in the synthesis of
1570:Produce a minimum amount of residual
1492:in the wine. In wine regions such as
938:whereas most wine yeast reproduce by
486:The distinctive "aldehydic" notes of
291:
2539:
1516:It is virtually inevitable that non-
1322:Wild yeasts and natural fermentation
906:" that produce inhibitory levels of
399:Other by-products of yeast include:
247:The yeast species commonly known as
2257:
1746: â used for the production of
1537:
624:that come as the wine continues to
13:
2247:pp. 133â36 Storey Publishing 1999
2228:
1482:The use of both "ambient" and non-
756:Types of yeasts used in winemaking
169:species take over. In addition to
14:
4970:
2666:
1953:are important in maintaining the
1796:Nicotinamide adenine dinucleotide
1588:or cause bungs to pop out during
1471:strains finish the fermentation.
1042:, other species within the genus
4926:
2955:
2480:Third Edition pgs 267 & 508
2104:
976:
163:while the more alcohol tolerant
90:the wine to remove the yeast or
2629:
2616:
2562:
2274:pp. 67â73 DBQA Publishing 2005
2272:"Understanding Wine Technology"
1694:Nutritional needs of wine yeast
506:yeast strains that produce the
365:) is reduced to "recharge" the
3032:History of alcoholic beverages
2721:
2600:"The Oxford Companion to Wine"
2494:
2478:"The Oxford Companion to Wine"
2453:
2329:"The Oxford Companion to Wine"
1889:method of adding the leftover
1264:strain 505) that are known to
620:as opposed to the less coarse
436:" aroma, reducing it first to
242:EmbdenâMeyerhofâParnas pathway
159:of alcohol on the yeast cells
1:
2918:Glossary of viticulture terms
2221:
2019:. Even the common wine yeast
1110:was the first single-celled,
2923:Glossary of winemaking terms
2527:Wine Analysis and Production
2441:Wine Analysis and Production
1978:Wine faults related to yeast
1248:named after the town in the
1114:organism to have its entire
694:aroma compounds that become
7:
307:Fermentation (biochemistry)
10:
4975:
2843:Yeast assimilable nitrogen
2650:The University Wine Course
2572:Diacetyl: Homebrew Science
2125:Brettanomyces bruxellensis
1847:for healthy cell function.
1816:molecules that facilitate
1812: â involved with the
1709:yeast assimilable nitrogen
1027:cannot ferment or utilize
729:
428:, the compound that gives
304:
253:Saccharomyces ellipsoideus
208:
4901:
4460:
4446:
4280:
4089:
3866:
3862:
3848:
3657:
3431:
3195:
3191:
3177:
3077:
3024:
3020:
3009:
2951:
2933:History of the wine press
2900:
2884:
2856:
2823:Sparkling wine production
2808:
2790:
2757:
2729:
2602:Third Edition pp. 398â99
2392:K. Fugelsang, C. Edwards
2331:Third Edition pp. 778â80
2178:As a fermentation yeast,
1242:sparkling wine production
1129:in identifying different
1080:Saccharomyces pastorianus
814:sexual classification of
732:Sparkling wine production
4304:Coconut and other palms
3676:Coconut and other palms
3047:History of Bordeaux wine
2928:Wine tasting descriptors
2188:Saccharomyces cerevisiae
2021:Saccharomyces cerevisiae
1901:Saccharomyces cerevisiae
1621:Saccharomyces cerevisiae
1619:) yeasts are strains of
1440:Saccharomyces cerevisiae
1341:Saccharomyces cerevisiae
1309:Saccharomyces fermentati
1294:Saccharomyces cerevisiae
1213:enzymes that can modify
1135:Saccharomyces cerevisiae
1108:Saccharomyces cerevisiae
1066:Saccharomyces fermentati
1040:Saccharomyces cerevisiae
986:Saccharomyces cerevisiae
917:Saccharomyces cerevisiae
825:Saccharomyces cerevisiae
561:contact with grape skins
265:Saccharomyces cerevisiae
249:Saccharomyces cerevisiae
113:Saccharomyces cerevisiae
3132:Drinking establishments
2818:Malolactic fermentation
2604:Oxford University Press
2482:Oxford University Press
2333:Oxford University Press
1073:Saccharomyces paradoxus
965:Aureobasidium pullulans
594:
66:through the process of
4339:Sugarcane or molasses
3776:Sugarcane or molasses
3015:History and production
2132:
2000:
1955:selective permeability
1864:
1756:Adenosine triphosphate
1703:
1665:
1603:compaction that makes
1547:
1479:
1331:
1103:
989:
827:which is also used in
775:
726:Secondary fermentation
605:
495:
406: â Caused by the
353:
302:
218:
183:and varieties such as
28:
3062:History of Rioja wine
2202:that many strains of
2190:will out compete the
2112:
1985:
1934:facultative anaerobes
1854:
1701:
1663:
1545:
1530:, which can impart a
1477:
1453:Unlike the "ambient"
1450:) into the vineyard.
1369:natural fermentations
1329:
1315:Saccharomyces bayanus
1303:Saccharomyces beticus
1250:Champagne wine region
1193:-linked compounds by
1101:
1059:Saccharomyces beticus
1052:Saccharomyces bayanus
984:
763:
602:
485:
351:
299:
216:
176:Saccharomyces bayanus
128:from cultured stock,
22:
4773:Mammee apple flower
3052:History of Champagne
1869:diammonium phosphate
1804:ethanol fermentation
1609:fining and filtering
1595:Have high levels of
1408:colony forming units
1374:Nanfang Caomu Zhuang
1355:yeast as opposed to
1258:California Champagne
1229:derivatives such as
1087:Saccharomyces uvarum
807:Dekkera bruxellensis
72:sugars in the grapes
4823:Sugarcane/molasses
3266:Moscatel de SetĂșbal
3159:Yeast in winemaking
3147:Yeast in winemaking
3003:Alcoholic beverages
2848:Yeast in winemaking
2800:Carbonic maceration
2635:Li Hui-Lin (1979),
2548:Dr. Yair Margalit,
2171:in the transfer of
2046:several species of
2042:In the presence of
1814:secondary messenger
1611:of the wine easier.
1435:Kloeckera apiculata
931:Schizosaccharomyces
706:, help enhance the
432:and other wines a "
311:Fermentation (wine)
276:compounds (such as
33:yeast in winemaking
3057:History of Chianti
3025:History of alcohol
2838:Traditional method
2684:2015-06-06 at the
2577:2010-02-02 at the
2133:
2080:titratable acidity
2037:stuck fermentation
2001:
1928:The role of oxygen
1865:
1778:and nucleic acids.
1704:
1666:
1645:some Sherry styles
1641:late-harvest wines
1548:
1480:
1332:
1104:
990:
776:
736:The production of
606:
496:
354:
336:the unfermentable
303:
292:Role in winemaking
219:
191:harvested at high
101:stuck fermentation
58:of the fruit into
29:
4949:Food microbiology
4914:
4913:
4897:
4896:
4600:Chocolate liqueur
4561:Blackthorn shrub
4442:
4441:
4438:
4437:
4394:Various starches
4150:Tennessee whiskey
3844:
3843:
3840:
3839:
3811:Various starches
3173:
3172:
3169:
3168:
3067:History of Sherry
2969:
2968:
2739:Late harvest wine
2598:J. Robinson (ed)
2476:J. Robinson (ed)
2394:Wine Microbiology
2327:J. Robinson (ed)
1823:Trace amounts of
1719:or YAN) but also
1432:with the species
1388:Gelsemium elegans
1137:that are used in
947:Zygosaccharomyces
704:protein stability
663:, sparkling wine
651:and described by
565:pyranoanthocyanin
492:Jerez wine region
48:absence of oxygen
4966:
4931:
4930:
4929:
4922:
4521:Herbs de Majorca
4495:Anisado Mallorca
4448:
4447:
4250:American whiskey
4241:Multiple grains
4130:Buckwheat whisky
3864:
3863:
3850:
3849:
3623:Multiple grains
3193:
3192:
3184:Fermented drinks
3179:
3178:
3085:Alcohol industry
3022:
3021:
3011:
3010:
2996:
2989:
2982:
2973:
2972:
2960:
2959:
2716:
2709:
2702:
2693:
2692:
2660:
2646:
2640:
2633:
2627:
2620:
2614:
2596:
2587:
2568:Brewing Science
2566:
2560:
2546:
2537:
2523:
2514:
2513:
2511:
2509:
2498:
2492:
2474:
2468:
2457:
2451:
2437:
2404:
2390:
2343:
2325:
2282:
2268:
2255:
2241:
2123:species such as
2064:volatile acidity
1986:Film yeast like
1959:osmotic pressure
1782:Pantothenic acid
1566:dimethyl sulfide
1538:Inoculated yeast
1511:volatile acidity
1260:, also known as
1185:produced by the
1164:hydrogen sulfide
992:The yeast genus
774:region of France
657:barrel fermented
539:ethyl mercaptans
522:Hydrogen sulfide
459:volatile acidity
363:dihydroxyacetone
278:hydrogen sulfide
221:For most of the
4974:
4973:
4969:
4968:
4967:
4965:
4964:
4963:
4954:Saccharomycetes
4939:
4938:
4937:
4927:
4925:
4917:
4915:
4910:
4893:
4862:CrĂšme de cassis
4857:CrĂšme de banane
4843:Various fruits
4712:CrĂšme de menthe
4473:CrĂšme de Noyaux
4456:
4434:
4276:
4260:Canadian whisky
4234:Kaoliang liquor
4140:Bourbon whiskey
4105:Japanese whisky
4085:
4054:Marillenschnaps
4030:Various fruits
3858:
3836:
3665:Agave americana
3653:
3427:
3382:Various fruits
3187:
3165:
3073:
3042:History of wine
3037:History of beer
3016:
3005:
3000:
2970:
2965:
2962:Wine portal
2954:
2947:
2938:History of wine
2896:
2880:
2852:
2804:
2786:
2767:Deacidification
2753:
2725:
2720:
2686:Wayback Machine
2669:
2664:
2663:
2647:
2643:
2634:
2630:
2621:
2617:
2597:
2590:
2579:Wayback Machine
2567:
2563:
2547:
2540:
2524:
2517:
2507:
2505:
2500:
2499:
2495:
2475:
2471:
2459:Douglas Harper
2458:
2454:
2438:
2407:
2391:
2346:
2326:
2285:
2269:
2258:
2242:
2229:
2224:
2145:4-Ethylguaiacol
2135:The wine yeast
2107:
2076:isoamyl acetate
1980:
1930:
1883:regions is the
1696:
1658:
1615:Inoculated (or
1540:
1371:. The (c. 304)
1324:
1175:Sauvignon blanc
1162:compounds like
1096:
1038:In addition to
979:
924:Saccharomycodes
758:
742:sparkling wines
734:
728:
677:polysaccharides
597:
559:extracted from
449:acidity of wine
346:
313:
294:
270:grape varieties
223:history of wine
211:
80:residual sugars
17:
12:
11:
5:
4972:
4962:
4961:
4956:
4951:
4936:
4935:
4912:
4911:
4909:
4908:
4902:
4899:
4898:
4895:
4894:
4892:
4891:
4890:
4889:
4881:
4880:
4879:
4874:
4869:
4864:
4859:
4854:
4849:
4841:
4840:
4839:
4831:
4830:
4829:
4821:
4820:
4819:
4814:
4806:
4805:
4804:
4799:
4794:
4789:
4781:
4780:
4779:
4771:
4770:
4769:
4764:
4756:
4755:
4754:
4749:
4744:
4739:
4734:
4726:
4725:
4724:
4719:
4714:
4709:
4704:
4699:
4691:
4690:
4689:
4681:
4680:
4679:
4674:
4666:
4665:
4664:
4659:
4651:
4650:
4649:
4644:
4636:
4635:
4634:
4626:
4625:
4624:
4614:
4613:
4612:
4604:
4603:
4602:
4594:
4593:
4592:
4586:Chili peppers
4584:
4583:
4582:
4577:
4575:Cherry Heering
4569:
4568:
4567:
4559:
4558:
4557:
4549:
4548:
4547:
4542:
4537:
4532:
4527:
4526:
4525:
4524:
4523:
4511:
4506:
4501:
4500:
4499:
4498:
4497:
4485:
4477:
4476:
4475:
4470:
4461:
4458:
4457:
4444:
4443:
4440:
4439:
4436:
4435:
4433:
4432:
4431:
4430:
4425:
4420:
4415:
4410:
4405:
4400:
4392:
4391:
4390:
4385:
4380:
4375:
4373:Seco Herrerano
4370:
4365:
4360:
4355:
4350:
4345:
4337:
4336:
4335:
4327:
4326:
4325:
4320:
4315:
4310:
4302:
4301:
4300:
4295:
4284:
4282:
4278:
4277:
4275:
4274:
4273:
4272:
4267:
4262:
4257:
4252:
4247:
4239:
4238:
4237:
4224:
4223:
4222:
4217:
4212:
4204:
4203:
4202:
4197:
4192:
4187:
4185:Mixiang Baijiu
4182:
4177:
4172:
4167:
4162:
4154:
4153:
4152:
4147:
4142:
4134:
4133:
4132:
4124:
4123:
4122:
4114:
4113:
4112:
4107:
4102:
4093:
4091:
4087:
4086:
4084:
4083:
4082:
4081:
4076:
4071:
4066:
4061:
4056:
4051:
4046:
4041:
4036:
4028:
4027:
4026:
4021:
4016:
4011:
4006:
4001:
3996:
3986:
3985:
3984:
3979:
3971:
3970:
3969:
3961:
3960:
3959:
3954:
3949:
3944:
3939:
3934:
3926:
3925:
3924:
3916:
3915:
3914:
3906:
3905:
3904:
3896:
3895:
3894:
3886:
3885:
3884:
3879:
3870:
3868:
3860:
3859:
3857:by ingredients
3846:
3845:
3842:
3841:
3838:
3837:
3835:
3834:
3833:
3832:
3827:
3822:
3817:
3809:
3808:
3807:
3802:
3794:
3793:
3792:
3787:
3782:
3774:
3773:
3772:
3764:
3763:
3762:
3757:
3752:
3747:
3739:
3738:
3737:
3729:
3728:
3727:
3719:
3718:
3717:
3709:
3708:
3707:
3702:
3697:
3692:
3687:
3682:
3674:
3673:
3672:
3661:
3659:
3655:
3654:
3652:
3651:
3650:
3649:
3644:
3639:
3634:
3629:
3621:
3620:
3619:
3611:
3610:
3609:
3601:
3600:
3599:
3594:
3589:
3584:
3579:
3574:
3569:
3564:
3559:
3554:
3549:
3544:
3539:
3534:
3529:
3524:
3519:
3514:
3509:
3504:
3499:
3494:
3486:
3485:
3484:
3479:
3471:
3470:
3469:
3464:
3459:
3454:
3446:
3445:
3444:
3435:
3433:
3429:
3428:
3426:
3425:
3424:
3423:
3418:
3413:
3408:
3403:
3398:
3393:
3388:
3380:
3379:
3378:
3370:
3369:
3368:
3360:
3359:
3358:
3350:
3349:
3348:
3340:
3339:
3338:
3330:
3329:
3328:
3320:
3319:
3318:
3310:
3309:
3308:
3300:
3299:
3298:
3293:
3288:
3283:
3278:
3273:
3268:
3263:
3258:
3253:
3245:
3244:
3243:
3235:
3234:
3233:
3225:
3224:
3223:
3218:
3210:
3209:
3208:
3199:
3197:
3189:
3188:
3186:by ingredients
3175:
3174:
3171:
3170:
3167:
3166:
3164:
3163:
3162:
3161:
3151:
3150:
3149:
3144:
3134:
3129:
3124:
3123:
3122:
3117:
3112:
3107:
3102:
3092:
3087:
3081:
3079:
3075:
3074:
3072:
3071:
3070:
3069:
3064:
3059:
3054:
3049:
3039:
3034:
3028:
3026:
3018:
3017:
3007:
3006:
2999:
2998:
2991:
2984:
2976:
2967:
2966:
2952:
2949:
2948:
2946:
2945:
2940:
2935:
2930:
2925:
2920:
2915:
2910:
2904:
2902:
2898:
2897:
2895:
2894:
2888:
2886:
2882:
2881:
2879:
2878:
2873:
2868:
2862:
2860:
2854:
2853:
2851:
2850:
2845:
2840:
2835:
2830:
2828:Sugars in wine
2825:
2820:
2814:
2812:
2806:
2805:
2803:
2802:
2796:
2794:
2788:
2787:
2785:
2784:
2779:
2777:Chaptalization
2774:
2769:
2763:
2761:
2755:
2754:
2752:
2751:
2746:
2741:
2735:
2733:
2727:
2726:
2719:
2718:
2711:
2704:
2696:
2690:
2689:
2668:
2667:External links
2665:
2662:
2661:
2641:
2628:
2615:
2588:
2586:November, 2002
2561:
2538:
2515:
2493:
2469:
2452:
2405:
2344:
2283:
2256:
2226:
2225:
2223:
2220:
2106:
2103:
1979:
1976:
1929:
1926:
1849:
1848:
1821:
1807:
1792:Nicotinic acid
1789:
1779:
1765:
1759:
1695:
1692:
1657:
1654:
1637:Barossa Valley
1613:
1612:
1593:
1582:
1568:
1562:
1559:
1556:
1539:
1536:
1462:or allowed to
1365:cultured yeast
1323:
1320:
1223:norisoprenoids
1199:GewĂŒrztraminer
1095:
1092:
1091:
1090:
1083:
1076:
1069:
1062:
1055:
978:
975:
974:
973:
955:
943:
927:
920:
891:
853:
757:
754:
750:carbon dioxide
730:Main article:
727:
724:
653:Cato the Elder
649:Ancient Romans
596:
593:
592:
591:
568:
550:
547:Montrachet 522
519:
480:
479:
452:
442:
422:2,3-Butanediol
415:
345:
342:
293:
290:
210:
207:
118:sulfur dioxide
64:carbon dioxide
15:
9:
6:
4:
3:
2:
4971:
4960:
4957:
4955:
4952:
4950:
4947:
4946:
4944:
4934:
4924:
4923:
4920:
4907:
4904:
4903:
4900:
4888:
4885:
4884:
4882:
4878:
4875:
4873:
4870:
4868:
4865:
4863:
4860:
4858:
4855:
4853:
4850:
4848:
4845:
4844:
4842:
4838:
4835:
4834:
4832:
4828:
4825:
4824:
4822:
4818:
4815:
4813:
4810:
4809:
4807:
4803:
4800:
4798:
4797:Grand Marnier
4795:
4793:
4790:
4788:
4785:
4784:
4782:
4778:
4775:
4774:
4772:
4768:
4765:
4763:
4760:
4759:
4757:
4753:
4750:
4748:
4745:
4743:
4740:
4738:
4735:
4733:
4730:
4729:
4727:
4723:
4720:
4718:
4715:
4713:
4710:
4708:
4705:
4703:
4700:
4698:
4695:
4694:
4692:
4688:
4685:
4684:
4682:
4678:
4675:
4673:
4670:
4669:
4667:
4663:
4660:
4658:
4657:Cream liqueur
4655:
4654:
4652:
4648:
4645:
4643:
4640:
4639:
4637:
4633:
4630:
4629:
4627:
4623:
4620:
4619:
4618:
4615:
4611:
4608:
4607:
4605:
4601:
4598:
4597:
4595:
4591:
4588:
4587:
4585:
4581:
4578:
4576:
4573:
4572:
4570:
4566:
4563:
4562:
4560:
4556:
4553:
4552:
4550:
4546:
4543:
4541:
4538:
4536:
4533:
4531:
4528:
4522:
4519:
4518:
4517:
4516:
4515:
4512:
4510:
4507:
4505:
4502:
4496:
4493:
4492:
4491:
4490:
4489:
4486:
4484:
4481:
4480:
4478:
4474:
4471:
4469:
4466:
4465:
4463:
4462:
4459:
4454:
4449:
4445:
4429:
4426:
4424:
4421:
4419:
4416:
4414:
4411:
4409:
4406:
4404:
4401:
4399:
4396:
4395:
4393:
4389:
4386:
4384:
4381:
4379:
4376:
4374:
4371:
4369:
4366:
4364:
4361:
4359:
4356:
4354:
4351:
4349:
4346:
4344:
4341:
4340:
4338:
4334:
4331:
4330:
4328:
4324:
4321:
4319:
4316:
4314:
4311:
4309:
4306:
4305:
4303:
4299:
4296:
4294:
4291:
4290:
4289:
4286:
4285:
4283:
4279:
4271:
4268:
4266:
4263:
4261:
4258:
4256:
4253:
4251:
4248:
4246:
4243:
4242:
4240:
4235:
4231:
4228:
4227:
4225:
4221:
4218:
4216:
4213:
4211:
4208:
4207:
4205:
4201:
4198:
4196:
4193:
4191:
4188:
4186:
4183:
4181:
4178:
4176:
4173:
4171:
4168:
4166:
4163:
4161:
4158:
4157:
4155:
4151:
4148:
4146:
4143:
4141:
4138:
4137:
4135:
4131:
4128:
4127:
4125:
4121:
4118:
4117:
4115:
4111:
4110:Scotch whisky
4108:
4106:
4103:
4101:
4100:Irish whiskey
4098:
4097:
4095:
4094:
4092:
4088:
4080:
4077:
4075:
4072:
4070:
4067:
4065:
4062:
4060:
4057:
4055:
4052:
4050:
4047:
4045:
4042:
4040:
4037:
4035:
4032:
4031:
4029:
4025:
4022:
4020:
4017:
4015:
4012:
4010:
4007:
4005:
4002:
4000:
3997:
3995:
3992:
3991:
3990:
3987:
3983:
3980:
3978:
3975:
3974:
3972:
3968:
3965:
3964:
3962:
3958:
3955:
3953:
3950:
3948:
3945:
3943:
3940:
3938:
3935:
3933:
3930:
3929:
3927:
3923:
3920:
3919:
3917:
3913:
3910:
3909:
3907:
3903:
3900:
3899:
3897:
3893:
3890:
3889:
3888:Cashew apple
3887:
3883:
3880:
3878:
3875:
3874:
3872:
3871:
3869:
3865:
3861:
3856:
3851:
3847:
3831:
3828:
3826:
3823:
3821:
3818:
3816:
3813:
3812:
3810:
3806:
3803:
3801:
3800:Fermented tea
3798:
3797:
3795:
3791:
3788:
3786:
3783:
3781:
3778:
3777:
3775:
3771:
3768:
3767:
3765:
3761:
3758:
3756:
3753:
3751:
3748:
3746:
3743:
3742:
3740:
3736:
3733:
3732:
3730:
3726:
3723:
3722:
3720:
3716:
3713:
3712:
3710:
3706:
3703:
3701:
3698:
3696:
3693:
3691:
3688:
3686:
3683:
3681:
3678:
3677:
3675:
3671:
3668:
3667:
3666:
3663:
3662:
3660:
3656:
3648:
3645:
3643:
3640:
3638:
3635:
3633:
3630:
3628:
3625:
3624:
3622:
3618:
3615:
3614:
3612:
3608:
3605:
3604:
3602:
3598:
3595:
3593:
3590:
3588:
3585:
3583:
3582:Shaoxing wine
3580:
3578:
3575:
3573:
3570:
3568:
3565:
3563:
3560:
3558:
3555:
3553:
3550:
3548:
3545:
3543:
3540:
3538:
3535:
3533:
3530:
3528:
3525:
3523:
3520:
3518:
3515:
3513:
3510:
3508:
3505:
3503:
3500:
3498:
3495:
3493:
3490:
3489:
3487:
3483:
3480:
3478:
3475:
3474:
3472:
3468:
3465:
3463:
3460:
3458:
3455:
3453:
3450:
3449:
3447:
3443:
3440:
3439:
3437:
3436:
3434:
3430:
3422:
3419:
3417:
3414:
3412:
3409:
3407:
3404:
3402:
3399:
3397:
3394:
3392:
3389:
3387:
3384:
3383:
3381:
3377:
3374:
3373:
3372:Prickly pear
3371:
3367:
3364:
3363:
3361:
3357:
3354:
3353:
3351:
3347:
3344:
3343:
3341:
3337:
3334:
3333:
3331:
3327:
3324:
3323:
3321:
3317:
3314:
3313:
3311:
3307:
3304:
3303:
3301:
3297:
3294:
3292:
3289:
3287:
3284:
3282:
3279:
3277:
3274:
3272:
3269:
3267:
3264:
3262:
3259:
3257:
3254:
3252:
3249:
3248:
3246:
3242:
3239:
3238:
3236:
3232:
3229:
3228:
3226:
3222:
3219:
3217:
3214:
3213:
3211:
3207:
3204:
3203:
3201:
3200:
3198:
3194:
3190:
3185:
3180:
3176:
3160:
3157:
3156:
3155:
3152:
3148:
3145:
3143:
3140:
3139:
3138:
3135:
3133:
3130:
3128:
3125:
3121:
3118:
3116:
3113:
3111:
3108:
3106:
3103:
3101:
3098:
3097:
3096:
3093:
3091:
3088:
3086:
3083:
3082:
3080:
3076:
3068:
3065:
3063:
3060:
3058:
3055:
3053:
3050:
3048:
3045:
3044:
3043:
3040:
3038:
3035:
3033:
3030:
3029:
3027:
3023:
3019:
3012:
3008:
3004:
2997:
2992:
2990:
2985:
2983:
2978:
2977:
2974:
2964:
2963:
2958:
2950:
2944:
2941:
2939:
2936:
2934:
2931:
2929:
2926:
2924:
2921:
2919:
2916:
2914:
2911:
2909:
2906:
2905:
2903:
2899:
2893:
2890:
2889:
2887:
2883:
2877:
2874:
2872:
2869:
2867:
2864:
2863:
2861:
2859:
2855:
2849:
2846:
2844:
2841:
2839:
2836:
2834:
2831:
2829:
2826:
2824:
2821:
2819:
2816:
2815:
2813:
2811:
2807:
2801:
2798:
2797:
2795:
2793:
2789:
2783:
2780:
2778:
2775:
2773:
2770:
2768:
2765:
2764:
2762:
2760:
2756:
2750:
2747:
2745:
2742:
2740:
2737:
2736:
2734:
2732:
2728:
2724:
2717:
2712:
2710:
2705:
2703:
2698:
2697:
2694:
2687:
2683:
2679:
2677:
2671:
2670:
2659:
2655:
2651:
2645:
2638:
2632:
2625:
2619:
2613:
2609:
2605:
2601:
2595:
2593:
2585:
2584:Brew Magazine
2582:
2580:
2576:
2573:
2565:
2559:
2555:
2551:
2545:
2543:
2536:
2532:
2528:
2522:
2520:
2503:
2497:
2491:
2487:
2483:
2479:
2473:
2466:
2464:
2456:
2450:
2446:
2442:
2436:
2434:
2432:
2430:
2428:
2426:
2424:
2422:
2420:
2418:
2416:
2414:
2412:
2410:
2403:
2399:
2395:
2389:
2387:
2385:
2383:
2381:
2379:
2377:
2375:
2373:
2371:
2369:
2367:
2365:
2363:
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2359:
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2308:
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2300:
2298:
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2292:
2290:
2288:
2281:
2277:
2273:
2267:
2265:
2263:
2261:
2254:
2250:
2246:
2240:
2238:
2236:
2234:
2232:
2227:
2219:
2217:
2213:
2212:Brettanomyces
2207:
2206:can produce.
2205:
2204:Brettanomyces
2201:
2200:octanoic acid
2197:
2196:decanoic acid
2193:
2192:Saccharomyces
2189:
2185:
2184:Brettanomyces
2181:
2180:Brettanomyces
2176:
2174:
2173:Brettanomyces
2170:
2166:
2162:
2158:
2154:
2150:
2146:
2142:
2141:4-Ethylphenol
2138:
2137:Brettanomyces
2130:
2126:
2122:
2121:Brettanomyces
2118:
2117:
2116:Brettanomyces
2111:
2105:Brettanomyces
2102:
2100:
2099:Brettanomyces
2096:
2092:
2091:Saccharomyces
2087:
2085:
2081:
2077:
2073:
2072:ethyl acetate
2069:
2065:
2061:
2057:
2054:can create a
2053:
2049:
2045:
2040:
2038:
2034:
2030:
2026:
2022:
2018:
2014:
2010:
2006:
1997:
1993:
1989:
1984:
1975:
1972:
1971:Saccharomyces
1967:
1963:
1960:
1956:
1952:
1948:
1944:
1939:
1935:
1925:
1923:
1919:
1914:
1910:
1906:
1902:
1898:
1896:
1892:
1888:
1887:
1882:
1878:
1874:
1870:
1862:
1858:
1853:
1846:
1842:
1838:
1834:
1830:
1826:
1822:
1819:
1818:cell division
1815:
1811:
1808:
1805:
1801:
1797:
1793:
1790:
1787:
1783:
1780:
1777:
1773:
1769:
1766:
1763:
1760:
1757:
1753:
1752:phospholipids
1749:
1748:nucleic acids
1745:
1742:
1741:
1740:
1738:
1734:
1730:
1726:
1722:
1718:
1714:
1710:
1700:
1691:
1687:
1684:
1678:
1676:
1671:
1662:
1653:
1651:
1646:
1642:
1638:
1634:
1630:
1626:
1622:
1618:
1617:pure cultured
1610:
1606:
1602:
1598:
1594:
1592:fermentation.
1591:
1587:
1583:
1581:
1577:
1573:
1569:
1567:
1563:
1560:
1557:
1554:
1553:
1552:
1544:
1535:
1533:
1529:
1528:phenylethanol
1524:
1523:Saccharomyces
1519:
1518:Saccharomyces
1514:
1512:
1507:
1503:
1499:
1495:
1491:
1490:
1485:
1484:Saccharomyces
1476:
1472:
1470:
1469:Saccharomyces
1465:
1461:
1456:
1455:Saccharomyces
1451:
1449:
1445:
1441:
1437:
1436:
1431:
1430:
1425:
1424:
1419:
1418:
1413:
1409:
1405:
1401:
1400:Saccharomyces
1396:
1394:
1390:
1389:
1384:
1380:
1376:
1375:
1370:
1366:
1362:
1358:
1354:
1350:
1346:
1342:
1338:
1328:
1319:
1317:
1316:
1311:
1310:
1305:
1304:
1299:
1298:Saccharomyces
1295:
1291:
1287:
1283:
1279:
1275:
1271:
1267:
1263:
1259:
1255:
1251:
1247:
1243:
1238:
1236:
1232:
1228:
1224:
1220:
1216:
1212:
1208:
1204:
1200:
1196:
1192:
1188:
1184:
1180:
1176:
1172:
1167:
1165:
1159:
1157:
1152:
1151:S. cerevisiae
1148:
1144:
1140:
1136:
1132:
1128:
1124:
1120:
1117:
1113:
1109:
1100:
1089:
1088:
1084:
1082:
1081:
1077:
1075:
1074:
1070:
1068:
1067:
1063:
1061:
1060:
1056:
1054:
1053:
1049:
1048:
1047:
1045:
1044:Saccharomyces
1041:
1036:
1034:
1030:
1026:
1025:Saccharomyces
1022:
1018:
1014:
1010:
1006:
1003:
999:
995:
994:Saccharomyces
987:
983:
977:Saccharomyces
971:
967:
966:
961:
960:
959:Aureobasidium
956:
953:
949:
948:
944:
941:
937:
933:
932:
928:
926:
925:
921:
919:
918:
913:
909:
908:ethyl acetate
905:
901:
900:Hanseniaspora
897:
896:
892:
889:
888:
887:Kluyveromyces
883:
882:
877:
876:
871:
870:
869:Metschnikowia
865:
864:
859:
858:
854:
851:
847:
846:
845:Brettanomyces
842:
841:
840:
838:
834:
830:
826:
821:
819:
818:
813:
809:
808:
803:
799:
798:
797:Brettanomyces
793:
789:
785:
781:
773:
769:
768:
762:
753:
751:
747:
743:
739:
733:
723:
721:
717:
713:
709:
705:
701:
697:
693:
689:
684:
682:
678:
674:
673:mannoproteins
670:
666:
662:
658:
654:
650:
645:
643:
639:
635:
631:
627:
623:
619:
615:
611:
601:
589:
585:
581:
577:
573:
569:
566:
562:
558:
554:
551:
548:
544:
540:
536:
531:
527:
523:
520:
517:
512:
509:
505:
501:
498:
497:
493:
489:
484:
477:
476:ethyl acetate
472:
468:
464:
460:
456:
453:
450:
446:
445:Succinic acid
443:
439:
435:
431:
427:
423:
419:
416:
413:
409:
408:demethylation
405:
402:
401:
400:
397:
395:
391:
386:
385:Metschnikowia
382:
377:
373:
368:
364:
360:
350:
341:
339:
334:
330:
326:
322:
318:
312:
308:
298:
289:
285:
284:of the wine.
283:
279:
275:
271:
266:
262:
258:
254:
250:
245:
243:
239:
235:
234:Louis Pasteur
230:
228:
224:
215:
206:
204:
200:
199:
198:Brettanomyces
194:
190:
186:
182:
178:
177:
172:
171:S. cerevisiae
168:
167:
166:Saccharomyces
162:
158:
154:
150:
147:
146:
141:
140:
135:
131:
130:S. cerevisiae
127:
123:
119:
115:
114:
109:
104:
102:
97:
93:
92:fortification
89:
85:
84:dessert wines
81:
77:
73:
69:
65:
61:
57:
54:converts the
53:
49:
45:
42:
38:
34:
26:
21:
4145:Corn whiskey
4049:Himbeergeist
3902:Kirschwasser
3664:
3362:Pomegranate
3256:Marsala wine
3251:Madeira wine
3158:
3146:
3142:Simple syrup
3137:Fermentation
3127:Distillation
2953:
2847:
2810:Fermentation
2673:
2649:
2644:
2636:
2631:
2623:
2618:
2599:
2583:
2569:
2564:
2549:
2526:
2506:. Retrieved
2504:. Scott Labs
2502:"Wine Yeast"
2496:
2477:
2472:
2460:
2455:
2440:
2393:
2328:
2271:
2244:
2211:
2208:
2203:
2191:
2187:
2183:
2179:
2177:
2172:
2136:
2134:
2128:
2124:
2120:
2114:
2098:
2090:
2088:
2067:
2056:film surface
2051:
2047:
2041:
2029:acetaldehyde
2020:
2016:
2012:
2002:
1995:
1991:
1987:
1970:
1968:
1964:
1931:
1900:
1899:
1884:
1881:Italian wine
1876:
1872:
1866:
1860:
1856:
1705:
1688:
1679:
1670:freeze dried
1667:
1650:Killer yeast
1620:
1616:
1614:
1597:flocculation
1580:acetaldehyde
1549:
1522:
1517:
1515:
1487:
1483:
1481:
1468:
1454:
1452:
1439:
1433:
1427:
1421:
1415:
1399:
1397:
1392:
1386:
1382:
1379:herb ferment
1372:
1368:
1364:
1360:
1356:
1352:
1348:
1344:
1340:
1333:
1313:
1307:
1301:
1297:
1293:
1274:disgorgement
1257:
1245:
1239:
1215:monoterpenes
1211:glycosidases
1168:
1160:
1150:
1134:
1107:
1105:
1085:
1078:
1071:
1064:
1057:
1050:
1043:
1039:
1037:
1024:
993:
991:
985:
963:
957:
945:
929:
922:
915:
904:killer yeast
899:
898:(Teleomorph
893:
885:
879:
875:Issatchenkia
873:
867:
861:
855:
849:
848:(Teleomorph
843:
829:bread making
824:
822:
815:
810:) under its
805:
795:
777:
765:
735:
685:
646:
641:
637:
621:
617:
607:
557:anthocyanins
553:Pyruvic acid
546:
508:Spanish wine
500:Acetaldehyde
488:Sherry wines
398:
394:legal limits
384:
380:
355:
329:acetaldehyde
314:
286:
264:
252:
248:
246:
238:Microbiology
231:
220:
196:
174:
170:
164:
143:
137:
129:
111:
105:
68:fermentation
32:
31:The role of
30:
4808:Star anise
4702:Bénédictine
4662:Irish cream
4398:Aguardiente
4343:Aguardiente
4215:Rye whiskey
3725:Ginger wine
3442:Barley wine
3406:Mulled wine
3356:Plum jerkum
3326:Lychee wine
3316:Longan wine
3241:Bokbunja-ju
3231:Bignay wine
3221:Banana wine
3216:Banana beer
3090:Bathtub gin
2913:Wine bottle
2885:Other steps
2876:Wine cellar
2833:SĂŒssreserve
2167:are common
2165:Fruit flies
2143:(4-EP) and
2025:acetic acid
2009:off flavors
2005:wine faults
1932:Yeasts are
1776:fatty acids
1717:amino acids
1683:cytoplasmic
1633:Napa Valley
1576:acetic acid
1412:fruit flies
1231:polyphenols
1189:of certain
1123:mycologists
1023:. However,
998:fruit wines
972:in barrels.
912:acetic acid
881:Torulaspora
839:) include:
837:wine faults
812:sporulating
802:viticulture
746:carbonation
716:astringency
696:volatilized
692:hydrophobic
667:as well as
455:Acetic acid
367:NADH enzyme
255:due to the
126:inoculation
70:. The more
4959:Winemaking
4943:Categories
4867:Limoncello
4802:Triple sec
4777:Eau créole
4747:Irish Mist
4732:BÀrenjÀger
4687:Frangelico
4617:Cloudberry
4596:Chocolate
4580:Maraschino
4126:Buckwheat
4039:Eau de vie
3342:Pineapple
3306:Duhat wine
3302:Java plum
3296:White wine
3154:Winemaking
3078:Production
2792:Maceration
2782:Wine press
2772:Destemming
2723:Winemaking
2658:0932664695
2612:0198609906
2558:0932664660
2535:0834217015
2490:0198609906
2449:0834217015
2402:0387333495
2341:0198609906
2280:1891267914
2253:1580171052
2222:References
2149:Pinot noir
2095:maceration
1947:lanosterol
1943:ergosterol
1913:autolyzing
1861:(pictured)
1737:riboflavin
1675:cold shock
1586:maceration
1498:classified
1404:grapevines
1357:inoculated
1349:indigenous
1290:manzanilla
1266:flocculate
1219:aliphatics
1187:hydrolysis
1147:winemaking
1127:enologists
1112:eukaryotic
970:aging wine
792:teleomorph
681:mercaptans
669:Chardonnay
618:gross lees
543:disulfides
430:Chardonnay
418:Fusel oils
333:co-enzymes
325:glycolysis
305:See also:
257:elliptical
203:wine fault
161:physiology
108:winemaking
4852:Cedratine
4817:Sassolino
4787:Cointreau
4707:BrennivĂn
4683:Hazelnut
4647:Tia Maria
4606:Cinnamon
4590:Pertsivka
4555:Bierlikör
4358:Desi daru
4120:Bierbrand
4014:Tsikoudia
3977:Slivovitz
3877:Applejack
3755:Kabarawan
3731:Galangal
3685:Palm wine
3562:Rice wine
3542:Makgeolli
3477:Oshikundu
3396:Hippocras
3281:Rosé wine
3271:Port wine
3237:Bokbunja
3110:Lautering
2744:Noble rot
2648:M. Baldy
2508:23 August
2243:Jeff Cox
2159:from the
2060:microbial
2013:Kloeckera
1969:Wild non-
1938:anaerobic
1893:from the
1841:manganese
1762:Potassium
1744:Phosphate
1731:(such as
1725:magnesium
1723:(such as
1417:Kloeckera
1156:wines age
1119:sequenced
1106:In 1996,
1033:arabinose
1031:(such as
1013:raffinose
895:Kloeckera
772:Jura wine
770:from the
767:Vin jaune
740:and many
738:Champagne
712:mouthfeel
665:Champagne
638:bĂątonnage
634:reductive
630:autolysis
622:fine lees
610:tartrates
471:propionic
390:bisulfite
381:Kloeckera
227:etymology
185:Zinfandel
139:Kloeckera
88:filtering
46:. In the
4877:Sloe gin
4872:Schnapps
4837:Licor 43
4833:Vanilla
4827:Charanda
4758:Juniper
4737:Drambuie
4672:Advocaat
4628:Coconut
4565:Patxaran
4504:Anisette
4483:Absinthe
4468:Amaretto
4453:Liqueurs
4318:Lambanog
4226:Sorghum
4165:Cheongju
4019:Tsipouro
3963:Juniper
3937:Armagnac
3882:Calvados
3825:Parakari
3805:Kombucha
3705:Tunggang
3680:Bahalina
3647:Huangjiu
3613:Sorghum
3567:RÆ°á»Łu cáș§n
3532:Gwaha-ju
3467:TesgĂŒino
3391:Dubonnet
3386:Conditum
3376:Colonche
3291:Vermouth
3276:Red wine
3261:Mistelle
2759:Pressing
2682:Archived
2575:Archived
2270:D. Bird
2157:Riesling
2153:Burgundy
2129:pictured
1992:pictured
1949:. These
1918:cysteine
1909:arginine
1905:ammonium
1895:pressing
1829:chlorine
1810:Inositol
1772:proteins
1729:vitamins
1721:minerals
1635:and the
1629:Burgundy
1625:Bordeaux
1572:pyruvate
1494:Bordeaux
1464:macerate
1361:selected
1270:riddling
1262:UC-Davis
1203:Riesling
1191:cysteine
1179:SĂ©millon
1171:varietal
1029:pentoses
788:anamorph
780:taxonomy
700:tartrate
661:Muscadet
604:corking.
580:lactones
570:Various
535:rackings
530:sulfites
526:sulfates
516:oxidized
426:diacetyl
404:Methanol
359:glycerol
157:toxicity
4883:Walnut
4847:Campari
4792:Curaçao
4783:Orange
4767:Jenever
4752:Krupnik
4697:Aquavit
4638:Coffee
4610:Tentura
4571:Cherry
4545:Sambuca
4530:Mastika
4514:Hierbas
4488:Anisado
4464:Almond
4413:Horilka
4403:Akvavit
4353:Clairin
4348:Cachaça
4298:Tequila
4175:Lao-Lao
4160:Awamori
4096:Barley
4090:Cereals
4079:Schnaps
4069:PĂĄlinka
4059:Nalewka
4024:Zivania
3957:Zivania
3898:Cherry
3855:Liquors
3721:Ginger
3557:Pangasi
3517:Choujiu
3497:Amazake
3473:Millet
3462:Tejuino
3438:Barley
3432:Cereals
3421:Sangria
3346:Tepache
3322:Lychee
3312:Longan
3227:Bignay
3212:Banana
3105:Malting
3100:Brewery
3095:Brewing
2943:Terroir
2901:Related
2749:Vintage
2731:Harvest
2216:ethanol
2048:Candida
2017:Candida
1988:Candida
1951:sterols
1886:ripasso
1873:Go-Ferm
1857:ripasso
1825:calcium
1806:itself.
1733:thiamin
1713:ammonia
1605:racking
1506:compost
1489:terroir
1460:pressed
1423:Candida
1353:natural
1345:ambient
1246:Ăpernay
1233:in the
1227:benzene
1195:enzymes
1131:strains
1021:ethanol
1017:maltose
1009:sucrose
1005:glucose
1002:ferment
940:budding
936:fission
857:Candida
850:Dekkera
833:brewing
817:Dekkera
720:tannins
688:mannose
642:sur lie
588:acetals
584:phenols
576:ketones
518:aromas.
463:butyric
438:acetoin
434:buttery
412:pectins
338:pentose
317:glucose
261:strains
209:History
145:Candida
134:harvest
76:dryness
60:alcohol
4919:Portal
4887:Nocino
4812:Pastis
4742:Glayva
4728:Honey
4722:Unicum
4717:Metaxa
4693:Herbs
4677:Eggnog
4653:Cream
4642:KahlĂșa
4632:Malibu
4479:Anise
4423:ShĆchĆ«
4418:PoitĂn
4408:Bangla
4388:Tharra
4329:Dairy
4313:Laksoy
4308:Arrack
4293:Mezcal
4270:Whisky
4265:ShĆchĆ«
4255:Baijiu
4230:Baijiu
4220:Starka
4190:Shochu
4180:Lihing
4170:Cholai
4136:Maize
4034:Brandy
3999:Grappa
3994:Chacha
3989:Pomace
3947:Cognac
3942:Brandy
3928:Grape
3922:Boukha
3908:Dates
3873:Apple
3830:Ibwatu
3820:Chicha
3766:Sugar
3741:Honey
3711:Dairy
3670:Pulque
3617:PendhÄ
3537:Hariya
3527:Dansul
3507:Beopju
3482:Tongba
3411:Nabidh
3286:Sherry
3247:Grape
3202:Apple
2908:Winery
2871:Solera
2656:
2610:
2556:
2533:
2488:
2447:
2400:
2339:
2278:
2251:
2169:vector
2068:Pichia
2052:Pichia
2044:oxygen
2033:thiols
1996:Pichia
1994:) and
1922:thiols
1891:pomace
1877:Ferm-K
1833:copper
1786:lipids
1768:Biotin
1727:) and
1590:barrel
1502:pomace
1448:pomace
1429:Pichia
1278:Sherry
1254:France
1225:, and
1207:Muscat
1183:thiols
1116:genome
863:Pichia
778:Yeast
626:settle
572:esters
511:Sherry
467:formic
441:aroma.
301:yeast.
274:sulfur
149:genera
96:brandy
56:sugars
4933:Drink
4622:Lakka
4551:Beer
4428:Vodka
4378:Sulai
4363:Guaro
4333:Arkhi
4323:Sotol
4288:Agave
4281:Other
4200:Sulai
4156:Rice
4116:Beer
4074:Rakia
4044:Geist
4009:Orujo
3982:ÈuicÄ
3973:Plum
3952:Pisco
3912:Araqi
3867:Fruit
3815:Cauim
3790:Palek
3785:Intus
3770:Kilju
3750:Byais
3735:Byais
3715:Kumis
3700:Tuhak
3658:Other
3607:Kvass
3597:Tapuy
3592:Tapai
3587:Sonti
3552:Mirin
3547:Mijiu
3522:Chuak
3492:Agkud
3488:Rice
3457:Pozol
3452:Mageu
3448:Corn
3416:Pruno
3401:Jabol
3366:Rimon
3352:Plum
3336:Perry
3332:Pear
3206:Cider
3196:Fruit
3120:Yeast
2858:Aging
2606:2006
2484:2006
2463:Yeast
2335:2006
2161:Mosel
2151:from
1999:wine.
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1383:cÇoqĆ«
1337:flora
1276:. In
1143:bread
372:redox
282:aroma
189:Syrah
181:ports
94:with
52:yeast
44:juice
41:fruit
39:from
4906:List
4668:Egg
4540:Rakı
4535:Ouzo
4509:Arak
4383:Sura
4210:Korn
4206:Rye
4195:Soju
4064:Oghi
4004:Marc
3932:Arak
3918:Fig
3892:Feni
3796:Tea
3780:Basi
3760:Mead
3745:Bais
3695:TubĂą
3690:Tuak
3642:Boza
3637:Beer
3603:Rye
3577:Sato
3572:Sake
3512:Brem
3115:Wort
2654:ISBN
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2486:ISBN
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2198:and
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2074:and
2050:and
2015:and
1945:and
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1843:and
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1735:and
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3967:Gin
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3627:Ale
3502:Apo
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