1107:"Waeringopteridae" and the superfamily that includes it, "Waeringopteroidea" (named after eurypterid researcher Erik N. Kjellesvig-Waering), are not formally published clades. The names derive from a 2004 thesis by O. Erik Tetlie and are as such not actually technically valid names. Nevertheless, as phylogenetic analyses confirms the grouping as monophyletic and the names supply an easy way to refer to the group, they remain routinely used within studies centered around eurypterid taxonomy. In these studies, the names are usually within quotation marks and/or are noted to not be properly valid names that derive from a thesis. The group is also sometimes referred to as the "waeringopteroid clade" rather than as the "Waeringopteroidea" within studies to differentiate it from the properly described superfamilies, pending the formal publication of Waeringopteroidea.
1165:
135:
1996:
451:
2299:
113:
1133:, is known from outside of Laurentia and it is thus assumed that the group originated in Laurentia before they spread to modern-day Germany in the Devonian. Further fossil finds are required to properly study the distribution and evolutionary history of the group. The temporal range of the group extends from the
437:
Although confirmed by phylogenetic analyses as monophyletic, and routinely used within eurypterid taxonomy, "Waeringopteroidea" and "Waeringopteridae" are as of yet not formally published clades, the names instead deriving from a 2004 thesis by O. Erik Tetlie. Some researchers refer to the group more
468:
Although waeringopterid fossils remain rare and in most cases fragmentary and very little concrete information on the group can confidently be established due to the poor fossil record, the waeringopterid eurypterids are joined by a handful of known features shared by all genera within the group.
434:. It is thus considered likely that the group originated in Laurentia before spreading to Germany during the Devonian. As the fossil record of the group is extremely poor, little is known of the morphology of the taxa within the group.
573:
and the waeringopteroids), as inferred by O. Erik Tetlie and Markus
Poschmann in 2008, based on the results of a 2008 analysis specifically pertaining to the Adelophthalmoidea and a preceding 2004 analysis.
1251:
Dunlop, J. A., Penney, D. & Jekel, D. 2015. A summary list of fossil spiders and their relatives. In World Spider
Catalog. Natural History Museum Bern, online at
508:(the posteriormost segment of the body) was xiphous (long and pointed). The eyes of waeringopterid eurypterids were located close to the marginal rim of the
1205:
527:
reached 60 cm (24 in) in length. Though this is relatively long in comparison to modern arthropods, other members of
Diploperculata, such as the
1213:
561:. The cladogram below presents the inferred phylogenetic positions of most of the genera included in the three most derived superfamilies of the
1272:
Tetlie, O. Erik; Cuggy, Michael B. (2007-01-01). "Phylogeny of the basal swimming eurypterids (Chelicerata; Eurypterida; Eurypterina)".
1320:"A new Ordovician eurypterid (Arthropoda: Chelicerata) from southeast Turkey: Evidence for a cryptic Ordovician record of Eurypterida"
1115:
Out of the five known waeringopterid species, four are from the fossil deposits in areas that were part of the ancient continent of
2336:
2075:
2065:
1587:
414:
Waeringopterid fossils are very rare and the family only contains five known species classified in three different genera (
2326:
1360:
2027:
2042:
2286:
2070:
2012:
1503:
1230:
2060:
2050:
2004:
1386:"Cope's rule and Romer's theory: patterns of diversity and gigantism in eurypterids and Palaeozoic vertebrates"
2331:
2022:
2017:
134:
2055:
2032:
543:
The
Waeringopteridae is the only family within the superfamily Waeringopteroidea and contains the genera
493:
1688:
1580:
1433:
1178:
2321:
1995:
477:
1449:"Phylogeny and palaeoecology of the Adelophthalmoidea (Arthropoda; Chelicerata; Eurypterida)"
2302:
1573:
1331:
1256:
8:
1170:
430:, with the only species known from outside of it being a Devonian species recovered from
1335:
1507:
1468:
1420:
1297:
328:
129:
504:(abdomen). In the waeringopterids, the fifth pair of appendages were spiniferous. The
1891:
1824:
1816:
1552:
1448:
1425:
1407:
1319:
1289:
566:
1511:
1472:
1385:
1301:
442:" rather than as the superfamily "Waeringopteroidea", pending a formal publication.
2231:
1945:
1937:
1899:
1870:
1786:
1735:
1542:
1499:
1460:
1415:
1399:
1339:
1281:
1222:
803:
779:
746:
684:
1488:"A New Eurypterid From the Upper Ordovician of Manitoulin Island, Ontario, Canada"
2260:
2106:
1963:
1904:
1860:
1763:
1750:
1390:
1226:
1134:
907:
388:
28:
365:. The Waeringopteridae is the only family classified as part of the superfamily
2176:
2134:
2092:
1955:
1882:
1842:
1834:
1758:
1144:
869:
820:
629:
581:
570:
557:
532:
460:
411:, which is named in honor of eurypterid researcher Erik N. Kjellesvig-Waering.
407:
402:
370:
318:
279:
215:
32:
1464:
1285:
2315:
2183:
2155:
2148:
2120:
2099:
1852:
1791:
1781:
1773:
1727:
1717:
1709:
1665:
1629:
1556:
1547:
1526:
1411:
1343:
1293:
1020:
698:
70:
2141:
1973:
1909:
1865:
1740:
1429:
1403:
1149:
1139:
1121:
955:
883:
646:
605:
551:
545:
450:
393:
379:
310:
302:
268:
121:
2238:
2215:
2113:
1807:
1700:
1638:
1620:
729:
562:
501:
473:
374:
202:
176:
45:
476:, and other arthropods in general, waeringopterid eurypterids possessed
2204:
2190:
2169:
2162:
2127:
1968:
1656:
1647:
1596:
1003:
979:
528:
358:
189:
90:
55:
2281:
2265:
2255:
1614:
1116:
427:
398:
362:
166:
146:
95:
39:
2224:
2197:
1126:
931:
509:
455:
112:
85:
80:
65:
60:
50:
1487:
1206:"Distribution and dispersal history of Eurypterida (Chelicerata)"
1130:
497:
485:
481:
431:
100:
75:
405:) age. The name Waeringopteridae is derived from the type genus
1608:
505:
489:
384:
156:
1565:
1361:"Lille1tv - Eurypterid phylogeny and the origin of Arachnida"
291:
1504:
10.1666/0022-3360(2005)079[1166:ANECFT]2.0.CO;2
1318:
Lamsdell, James; Hoşgör, Izzet; Selden, Paul (2013-01-31).
426:). Four of these species are from the ancient continent of
1525:
Dunlop, Jason A.; Braddy, Simon J.; Tetlie, Erik (2008).
1257:
http://www.wsc.nmbe.ch/resources/fossils/Fossils16.0.pdf
515:
The waeringopterids were almost all small eurypterids.
1252:
1110:
1446:
1317:
1160:
535:, reached much larger sizes, surpassing two meters.
369:, which in turn is classified within the infraorder
275:
480:bodies and jointed appendages (limbs) covered in a
1524:
519:reached 15 cm (5.9 in) in length, while
1214:Palaeogeography, Palaeoclimatology, Palaeoecology
2313:
1447:Erik Tetlie, O; Poschmann, Markus (2008-06-01).
1383:
1203:
1581:
1485:
1384:Lamsdell, James C.; Braddy, Simon J. (2009).
1199:
1197:
1195:
1193:
391:) age and the latest known surviving member,
1271:
492:. The chelicerate body is divided into two
1588:
1574:
1190:
377:. The earliest known member of the group,
111:
1546:
1419:
1119:. Only the last known surviving species,
449:
2314:
397:, has been recovered from deposits of
383:, has been recovered from deposits of
1569:
1247:
1245:
1243:
1531:(Gross, 1933) from Overath, Germany"
1355:
1353:
1313:
1311:
1267:
1265:
1486:Stott, Tetlie; et al. (2005).
1479:
1453:Journal of Systematic Palaeontology
1274:Journal of Systematic Palaeontology
13:
1240:
1111:Distribution and paleobiogeography
14:
2348:
1440:
1350:
1308:
1262:
538:
523:reached 30 cm (12 in).
2298:
2297:
1994:
1377:
1163:
133:
43:
2287:Timeline of eurypterid research
1595:
1527:"The Early Devonian eurypterid
2337:Prehistoric arthropod families
1518:
445:
361:, an extinct group of aquatic
1:
1184:
517:Waeringopterus cumberlandicus
280:Waeringopterus cumberlandicus
1227:10.1016/j.palaeo.2007.05.011
525:Orcanopterus manitoulinensis
16:Extinct family of arthropods
7:
1156:
10:
2353:
2327:Silurian first appearances
2295:
2274:
2248:
2214:
2084:
2041:
2003:
1992:
1954:
1936:
1918:
1890:
1879:
1851:
1833:
1815:
1804:
1772:
1749:
1726:
1708:
1697:
1689:List of eurypterid genera
1681:
1674:
1603:
1465:10.1017/S1477201907002416
1286:10.1017/S1477201907002131
1179:List of eurypterid genera
1017:
1000:
993:
976:
969:
952:
945:
928:
921:
904:
897:
880:
873:
817:
800:
793:
776:
769:
743:
726:
719:
712:
695:
688:
678:
643:
626:
619:
602:
595:
585:
565:suborder of eurypterids (
334:
327:
297:
290:
274:
267:
130:Scientific classification
128:
119:
110:
23:
1548:10.1002/mmng.20020050107
1344:10.1016/j.gr.2012.04.006
496:(sections); the frontal
1492:Journal of Paleontology
1365:lille1tv.univ-lille1.fr
1204:O. Erik Tetlie (2007).
1404:10.1098/rsbl.2009.0700
465:
344:Not formally published
260:Not formally published
237:Not formally published
1529:Grossopterus overathi
1434:Supplemental material
1122:Grossopterus overathi
521:Grossopterus overathi
500:(head) and posterior
453:
440:waeringopteroid clade
2332:Devonian extinctions
512:(the "head" plate).
1336:2013GondR..23..354L
1171:Paleontology portal
1817:Onychopterelloidea
1253:http://wsc.nmbe.ch
466:
2309:
2308:
1990:
1989:
1986:
1985:
1982:
1981:
1938:Adelophthalmoidea
1920:Waeringopteroidea
1892:Carcinosomatoidea
1825:Onychopterellidae
1800:
1799:
1324:Gondwana Research
1104:
1103:
1095:
1094:
1086:
1085:
1077:
1076:
1068:
1067:
1059:
1058:
1050:
1049:
1041:
1040:
1032:
1031:
859:
858:
850:
849:
841:
840:
832:
831:
758:
757:
685:Adelophthalmoidea
667:
666:
658:
657:
592:Waeringopteroidea
567:Adelophthalmoidea
438:formally as the "
367:Waeringopteroidea
352:
351:
346:
340:
263:
262:
240:
239:
229:Waeringopteroidea
2344:
2301:
2300:
2275:Related articles
2232:Merostomichnites
1998:
1946:Adelophthalmidae
1928:Waeringopteridae
1900:Carcinosomatidae
1888:
1887:
1871:Strobilopteridae
1813:
1812:
1787:Hibbertopteridae
1736:Parastylonuridae
1706:
1705:
1679:
1678:
1590:
1583:
1576:
1567:
1566:
1561:
1560:
1550:
1522:
1516:
1515:
1498:(6): 1166–1174.
1483:
1477:
1476:
1444:
1438:
1437:
1423:
1381:
1375:
1374:
1372:
1371:
1357:
1348:
1347:
1315:
1306:
1305:
1269:
1260:
1249:
1238:
1237:
1235:
1229:. Archived from
1221:(3–4): 557–574.
1210:
1201:
1173:
1168:
1167:
1166:
996:
995:
972:
971:
948:
947:
924:
923:
900:
899:
876:
875:
804:Parahughmilleria
796:
795:
780:Nanahughmilleria
772:
771:
747:Pittsfordipterus
722:
721:
715:
714:
691:
690:
681:
680:
622:
621:
598:
597:
588:
587:
578:
577:
373:in the suborder
355:Waeringopteridae
342:
338:
277:
258:
255:
252:Waeringopteridae
250:
235:
232:
227:
214:
201:
188:
138:
137:
115:
105:
42:
27:Temporal range:
24:Waeringopteridae
21:
20:
2352:
2351:
2347:
2346:
2345:
2343:
2342:
2341:
2312:
2311:
2310:
2305:
2291:
2270:
2261:Chasmataspidida
2244:
2210:
2107:Campylocephalus
2080:
2037:
1999:
1978:
1964:Hughmilleriidae
1950:
1932:
1914:
1905:Megalograptidae
1881:
1875:
1861:Dolichopteridae
1847:
1835:Moselopteroidea
1829:
1806:
1796:
1782:Drepanopteridae
1768:
1764:Hardieopteridae
1751:Kokomopteroidea
1745:
1722:
1699:
1693:
1670:
1599:
1594:
1564:
1523:
1519:
1484:
1480:
1445:
1441:
1391:Biology Letters
1382:
1378:
1369:
1367:
1359:
1358:
1351:
1316:
1309:
1270:
1263:
1255:, version 16.0
1250:
1241:
1233:
1208:
1202:
1191:
1187:
1169:
1164:
1162:
1159:
1135:Late Ordovician
1113:
1105:
1096:
1087:
1078:
1069:
1060:
1051:
1042:
1033:
908:Herefordopterus
860:
851:
842:
833:
759:
668:
659:
541:
533:carcinosomatids
472:Like all other
448:
389:Late Ordovician
357:is a family of
341:
337:Orcanopteridae
286:
283:
257:
254:
248:
234:
231:
225:
212:
199:
186:
132:
106:
104:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
53:
48:
37:
36:
29:Late Ordovician
25:
17:
12:
11:
5:
2350:
2340:
2339:
2334:
2329:
2324:
2322:Diploperculata
2307:
2306:
2296:
2293:
2292:
2290:
2289:
2284:
2278:
2276:
2272:
2271:
2269:
2268:
2263:
2258:
2252:
2250:
2249:Related groups
2246:
2245:
2243:
2242:
2235:
2228:
2220:
2218:
2212:
2211:
2209:
2208:
2201:
2194:
2187:
2180:
2177:Onychopterella
2173:
2166:
2159:
2152:
2145:
2138:
2135:Hibbertopterus
2131:
2124:
2117:
2110:
2103:
2096:
2093:Adelophthalmus
2088:
2086:
2085:Notable genera
2082:
2081:
2079:
2078:
2073:
2068:
2063:
2058:
2053:
2047:
2045:
2039:
2038:
2036:
2035:
2030:
2025:
2020:
2015:
2009:
2007:
2001:
2000:
1993:
1991:
1988:
1987:
1984:
1983:
1980:
1979:
1977:
1976:
1971:
1966:
1960:
1958:
1956:Pterygotioidea
1952:
1951:
1949:
1948:
1942:
1940:
1934:
1933:
1931:
1930:
1924:
1922:
1916:
1915:
1913:
1912:
1907:
1902:
1896:
1894:
1885:
1883:Diploperculata
1877:
1876:
1874:
1873:
1868:
1863:
1857:
1855:
1849:
1848:
1846:
1845:
1843:Moselopteridae
1839:
1837:
1831:
1830:
1828:
1827:
1821:
1819:
1810:
1802:
1801:
1798:
1797:
1795:
1794:
1789:
1784:
1778:
1776:
1770:
1769:
1767:
1766:
1761:
1759:Kokomopteridae
1755:
1753:
1747:
1746:
1744:
1743:
1738:
1732:
1730:
1724:
1723:
1721:
1720:
1714:
1712:
1710:Rhenopteroidea
1703:
1695:
1694:
1692:
1691:
1685:
1683:
1676:
1672:
1671:
1669:
1668:
1659:
1650:
1641:
1632:
1623:
1617:
1611:
1604:
1601:
1600:
1593:
1592:
1585:
1578:
1570:
1563:
1562:
1517:
1478:
1459:(2): 237–249.
1439:
1398:(2): 265–269.
1376:
1349:
1330:(1): 354–366.
1307:
1280:(3): 345–356.
1261:
1239:
1236:on 2011-07-18.
1188:
1186:
1183:
1182:
1181:
1175:
1174:
1158:
1155:
1145:Early Devonian
1112:
1109:
1102:
1101:
1098:
1097:
1093:
1092:
1089:
1088:
1084:
1083:
1080:
1079:
1075:
1074:
1071:
1070:
1066:
1065:
1062:
1061:
1057:
1056:
1053:
1052:
1048:
1047:
1044:
1043:
1039:
1038:
1035:
1034:
1030:
1029:
1026:
1025:
1016:
1013:
1012:
1009:
1008:
999:
994:
992:
989:
988:
985:
984:
975:
970:
968:
965:
964:
961:
960:
951:
946:
944:
941:
940:
937:
936:
927:
922:
920:
917:
916:
913:
912:
903:
898:
896:
893:
892:
889:
888:
879:
874:
872:
870:Pterygotioidea
866:
865:
862:
861:
857:
856:
853:
852:
848:
847:
844:
843:
839:
838:
835:
834:
830:
829:
826:
825:
821:Adelophthalmus
816:
813:
812:
809:
808:
799:
794:
792:
789:
788:
785:
784:
775:
770:
768:
765:
764:
761:
760:
756:
755:
752:
751:
742:
739:
738:
735:
734:
725:
720:
718:
713:
711:
708:
707:
704:
703:
694:
689:
687:
679:
677:
674:
673:
670:
669:
665:
664:
661:
660:
656:
655:
652:
651:
642:
639:
638:
635:
634:
630:Waeringopterus
625:
620:
618:
615:
614:
611:
610:
601:
596:
594:
586:
584:
582:Diploperculata
576:
571:Pterygotioidea
558:Waeringopterus
540:
539:Classification
537:
461:Waeringopterus
447:
444:
416:Waeringopterus
408:Waeringopterus
403:Early Devonian
371:Diploperculata
350:
349:
348:
347:
332:
331:
325:
324:
323:
322:
319:Waeringopterus
314:
306:
295:
294:
288:
287:
284:
272:
271:
265:
264:
246:
242:
241:
223:
219:
218:
216:Diploperculata
210:
206:
205:
197:
193:
192:
184:
180:
179:
174:
170:
169:
164:
160:
159:
154:
150:
149:
144:
140:
139:
126:
125:
117:
116:
108:
107:
99:
94:
89:
84:
79:
74:
69:
64:
59:
54:
49:
44:
33:Early Devonian
26:
15:
9:
6:
4:
3:
2:
2349:
2338:
2335:
2333:
2330:
2328:
2325:
2323:
2320:
2319:
2317:
2304:
2294:
2288:
2285:
2283:
2280:
2279:
2277:
2273:
2267:
2264:
2262:
2259:
2257:
2254:
2253:
2251:
2247:
2241:
2240:
2236:
2234:
2233:
2229:
2227:
2226:
2222:
2221:
2219:
2217:
2213:
2207:
2206:
2202:
2200:
2199:
2195:
2193:
2192:
2188:
2186:
2185:
2184:Pentecopterus
2181:
2179:
2178:
2174:
2172:
2171:
2167:
2165:
2164:
2160:
2158:
2157:
2156:Megalograptus
2153:
2151:
2150:
2149:Jaekelopterus
2146:
2144:
2143:
2139:
2137:
2136:
2132:
2130:
2129:
2125:
2123:
2122:
2121:Drepanopterus
2118:
2116:
2115:
2111:
2109:
2108:
2104:
2102:
2101:
2100:Brachyopterus
2097:
2095:
2094:
2090:
2089:
2087:
2083:
2077:
2076:South America
2074:
2072:
2069:
2067:
2066:North America
2064:
2062:
2059:
2057:
2054:
2052:
2049:
2048:
2046:
2044:
2040:
2034:
2031:
2029:
2028:Carboniferous
2026:
2024:
2021:
2019:
2016:
2014:
2011:
2010:
2008:
2006:
2005:Geochronology
2002:
1997:
1975:
1972:
1970:
1967:
1965:
1962:
1961:
1959:
1957:
1953:
1947:
1944:
1943:
1941:
1939:
1935:
1929:
1926:
1925:
1923:
1921:
1917:
1911:
1908:
1906:
1903:
1901:
1898:
1897:
1895:
1893:
1889:
1886:
1884:
1878:
1872:
1869:
1867:
1864:
1862:
1859:
1858:
1856:
1854:
1853:Eurypteroidea
1850:
1844:
1841:
1840:
1838:
1836:
1832:
1826:
1823:
1822:
1820:
1818:
1814:
1811:
1809:
1803:
1793:
1792:Mycteroptidae
1790:
1788:
1785:
1783:
1780:
1779:
1777:
1775:
1774:Mycteropoidea
1771:
1765:
1762:
1760:
1757:
1756:
1754:
1752:
1748:
1742:
1739:
1737:
1734:
1733:
1731:
1729:
1728:Stylonuroidea
1725:
1719:
1718:Rhenopteridae
1716:
1715:
1713:
1711:
1707:
1704:
1702:
1696:
1690:
1687:
1686:
1684:
1680:
1677:
1673:
1667:
1666:Sclerophorata
1663:
1660:
1658:
1654:
1651:
1649:
1645:
1642:
1640:
1636:
1633:
1631:
1630:Euchelicerata
1627:
1624:
1622:
1618:
1616:
1612:
1610:
1606:
1605:
1602:
1598:
1591:
1586:
1584:
1579:
1577:
1572:
1571:
1568:
1558:
1554:
1549:
1544:
1541:(1): 93–104.
1540:
1536:
1535:Fossil Record
1532:
1530:
1521:
1513:
1509:
1505:
1501:
1497:
1493:
1489:
1482:
1474:
1470:
1466:
1462:
1458:
1454:
1450:
1443:
1435:
1431:
1427:
1422:
1417:
1413:
1409:
1405:
1401:
1397:
1393:
1392:
1387:
1380:
1366:
1362:
1356:
1354:
1345:
1341:
1337:
1333:
1329:
1325:
1321:
1314:
1312:
1303:
1299:
1295:
1291:
1287:
1283:
1279:
1275:
1268:
1266:
1258:
1254:
1248:
1246:
1244:
1232:
1228:
1224:
1220:
1216:
1215:
1207:
1200:
1198:
1196:
1194:
1189:
1180:
1177:
1176:
1172:
1161:
1154:
1152:
1151:
1146:
1142:
1141:
1136:
1132:
1128:
1124:
1123:
1118:
1108:
1100:
1099:
1091:
1090:
1082:
1081:
1073:
1072:
1064:
1063:
1055:
1054:
1046:
1045:
1037:
1036:
1028:
1027:
1024:
1023:
1022:
1021:Jaekelopterus
1015:
1014:
1011:
1010:
1007:
1006:
1005:
998:
997:
991:
990:
987:
986:
983:
982:
981:
974:
973:
967:
966:
963:
962:
959:
958:
957:
950:
949:
943:
942:
939:
938:
935:
934:
933:
926:
925:
919:
918:
915:
914:
911:
910:
909:
902:
901:
895:
894:
891:
890:
887:
886:
885:
878:
877:
871:
868:
867:
864:
863:
855:
854:
846:
845:
837:
836:
828:
827:
824:
823:
822:
815:
814:
811:
810:
807:
806:
805:
798:
797:
791:
790:
787:
786:
783:
782:
781:
774:
773:
767:
766:
763:
762:
754:
753:
750:
749:
748:
741:
740:
737:
736:
733:
732:
731:
724:
723:
717:
716:
710:
709:
706:
705:
702:
701:
700:
699:Eysyslopterus
693:
692:
686:
683:
682:
676:
675:
672:
671:
663:
662:
654:
653:
650:
649:
648:
641:
640:
637:
636:
633:
632:
631:
624:
623:
617:
616:
613:
612:
609:
608:
607:
600:
599:
593:
590:
589:
583:
580:
579:
575:
572:
568:
564:
560:
559:
554:
553:
548:
547:
536:
534:
530:
526:
522:
518:
513:
511:
507:
503:
499:
495:
491:
487:
483:
479:
475:
470:
463:
462:
457:
452:
443:
441:
435:
433:
429:
425:
421:
417:
412:
410:
409:
404:
400:
396:
395:
390:
386:
382:
381:
376:
372:
368:
364:
360:
356:
345:
336:
335:
333:
330:
326:
321:
320:
315:
313:
312:
307:
305:
304:
299:
298:
296:
293:
289:
282:
281:
273:
270:
266:
261:
253:
247:
244:
243:
238:
230:
224:
222:Superfamily:
221:
220:
217:
211:
208:
207:
204:
198:
195:
194:
191:
185:
182:
181:
178:
175:
172:
171:
168:
165:
162:
161:
158:
155:
152:
151:
148:
145:
142:
141:
136:
131:
127:
124:
123:
118:
114:
109:
102:
97:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
41:
38:449–407
34:
30:
22:
19:
2237:
2230:
2223:
2203:
2196:
2189:
2182:
2175:
2168:
2161:
2154:
2147:
2142:Hughmilleria
2140:
2133:
2126:
2119:
2112:
2105:
2098:
2091:
1974:Pterygotidae
1927:
1919:
1910:Mixopteridae
1866:Eurypteridae
1741:Stylonuridae
1661:
1652:
1643:
1634:
1625:
1538:
1534:
1528:
1520:
1495:
1491:
1481:
1456:
1452:
1442:
1395:
1389:
1379:
1368:. Retrieved
1364:
1327:
1323:
1277:
1273:
1231:the original
1218:
1212:
1150:Grossopterus
1148:
1140:Orcanopterus
1138:
1120:
1114:
1106:
1019:
1018:
1002:
1001:
978:
977:
956:Erettopterus
954:
953:
930:
929:
906:
905:
884:Hughmilleria
882:
881:
819:
818:
802:
801:
778:
777:
745:
744:
728:
727:
697:
696:
647:Grossopterus
645:
644:
628:
627:
606:Orcanopterus
604:
603:
591:
556:
552:Orcanopterus
550:
546:Grossopterus
544:
542:
524:
520:
516:
514:
484:composed of
474:chelicerates
471:
467:
459:
439:
436:
424:Grossopterus
423:
420:Orcanopterus
419:
415:
413:
406:
394:Grossopterus
392:
380:Orcanopterus
378:
366:
354:
353:
343:
339:Tetlie, 2004
317:
311:Orcanopterus
309:
303:Grossopterus
301:
285:Swartz, 1923
278:
269:Type species
259:
256:Tetlie, 2004
251:
236:
233:Tetlie, 2004
228:
209:Infraorder:
122:Orcanopterus
120:
18:
2239:Palmichnium
2216:Ichnogenera
2114:Carcinosoma
1808:Eurypterina
1701:Stylonurina
1639:Prosomapoda
1621:Chelicerata
1619:Subphylum:
1597:Eurypterids
1125:, from the
730:Bassipterus
563:Eurypterina
529:pterygotids
502:opisthosoma
446:Description
375:Eurypterina
359:eurypterids
203:Eurypterina
190:Eurypterida
177:Chelicerata
173:Subphylum:
2316:Categories
2205:Stylonurus
2191:Pterygotus
2170:Mixopterus
2163:Megarachne
2128:Eurypterus
2013:Ordovician
1969:Slimonidae
1880:Infraorder
1657:Dekatriata
1648:Planaterga
1615:Arthropoda
1370:2018-04-02
1185:References
1004:Acutiramus
980:Pterygotus
363:arthropods
196:Suborder:
167:Arthropoda
2282:Metastoma
2266:Xiphosura
2256:Arachnida
2043:Geography
1607:Kingdom:
1557:1860-1014
1412:1744-9561
1294:1477-2019
1143:) to the
1117:Laurentia
478:segmented
456:carapaces
428:Laurentia
399:Siegenian
153:Kingdom:
147:Eukaryota
2303:Category
2225:Arcuites
2198:Slimonia
2023:Devonian
2018:Silurian
1805:Suborder
1698:Suborder
1675:Taxonomy
1613:Phylum:
1609:Animalia
1512:55014027
1473:59488956
1430:19828493
1302:88413267
1157:See also
1127:Devonian
932:Slimonia
510:carapace
486:proteins
329:Synonyms
245:Family:
163:Phylum:
157:Animalia
143:Domain:
2071:Oceania
2033:Permian
1421:2865068
1332:Bibcode
1131:Germany
531:or the
498:prosoma
494:tagmata
482:cuticle
454:Fossil
432:Germany
183:Order:
2061:Europe
2051:Africa
1682:Genera
1555:
1510:
1471:
1428:
1418:
1410:
1300:
1292:
1259:(PDF).
506:telson
490:chitin
385:Katian
292:Genera
1662:Clade
1653:Clade
1644:Clade
1635:Clade
1626:Clade
1508:S2CID
1469:S2CID
1298:S2CID
1234:(PDF)
1209:(PDF)
2056:Asia
1553:ISSN
1426:PMID
1408:ISSN
1290:ISSN
555:and
488:and
422:and
46:PreꞒ
1543:doi
1500:doi
1461:doi
1416:PMC
1400:doi
1340:doi
1282:doi
1223:doi
1219:252
1153:).
1129:of
458:of
2318::
1664::
1655::
1646::
1637::
1628::
1551:.
1537:.
1533:.
1506:.
1496:79
1494:.
1490:.
1467:.
1455:.
1451:.
1432:.
1424:.
1414:.
1406:.
1394:.
1388:.
1363:.
1352:^
1338:.
1328:23
1326:.
1322:.
1310:^
1296:.
1288:.
1276:.
1264:^
1242:^
1217:.
1211:.
1192:^
569:,
549:,
418:,
96:Pg
40:Ma
35:,
31:-
1589:e
1582:t
1575:v
1559:.
1545::
1539:5
1514:.
1502::
1475:.
1463::
1457:6
1436:.
1402::
1396:6
1373:.
1346:.
1342::
1334::
1304:.
1284::
1278:5
1225::
1147:(
1137:(
464:.
401:(
387:(
316:†
308:†
300:†
276:†
249:†
226:†
213:†
200:†
187:†
101:N
91:K
86:J
81:T
76:P
71:C
66:D
61:S
56:O
51:Ꞓ
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
