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Verbascum thapsus

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especially in the northern parts of the range, require a longer growth period and flower in their third year. Under better growing conditions, some individuals flower in the first year. Triennial individuals have been found to produce fewer seeds than biennial and annual ones. While year of flowering and size are linked to the environment, most other characteristics appear to be genetic.
313:, usually following damage. After flowering and seed release, the stem and fruits usually persist in winter, drying into dark brown, stiff structures of densely packed, ovoid-shaped, and dry seed capsules. The dried stems may persist into the following spring or even the next summer. The plant produces a shallow 309:, and reaching sizes up to 50 cm long and 14 cm across (19 inches long and 5 inches wide). They become smaller higher up the stem, and less strongly decurrent down the stem. The flowering stem is solid and 2–2.5 cm (nearly an inch) across, and occasionally branched just below the 1088:
The seeds maintain their germinative powers for decades, up to 100 years, according to some studies. Because of this, and because the plant is an extremely prolific seed bearer (each plant produces hundreds of capsules, each containing up to 700 seeds, with a total up to 180,000 or 240,000 seeds), it
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has not undergone any significant revision since Svanve Mürbeck's monographs in the 1930s, with the exception of the work of Arthur Huber-Morath, who used informal grouping in organizing the genus for the florae of Iran and Turkey to account for many intermediate species. Since Huber-Morath's groups
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application bypasses this requirement. Seeds germinate almost solely in bare soil, at temperatures between 10 and 40 °C. While they can germinate in total darkness if proper conditions are present (tests give a 35% germination rate under ideal conditions), in the wild, they in practice only do
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Great mullein most frequently grows as a colonist of bare and disturbed soil, usually on sandy or chalky ones. It grows best in dry, sandy, or gravelly soils, although it can grow in a variety of habitats, including banksides, meadows, roadsides, forest clearings, and pastures. This ability to grow
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In the 19th century, it had well over 40 different common names in English alone. Some of the more whimsical ones included "hig candlewick", "Indian rag weed", "bullicks lungwort", "Adams-rod", "hare's-beard", and "ice-leaf". Vernacular names include innumerable references to the plant's hairiness:
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of leaves in its first year of growth. The leaves are large, up to 50 cm long. The second-year plants normally produce a single unbranched stem, usually 1–2 m tall. In the eastern part of its range in China, it is, however, only reported to grow up to 1.5 m tall. The tall, pole-like
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Seeds germinate in spring and summer. Those that germinate in autumn produce plants that overwinter if they are large enough, while rosettes less than 15 cm (6 in) across die in winter. After flowering, the entire plant usually dies at the end of its second year, but some individuals,
372:, is known to occur. Flowering lasts up to three months from early to late summer (June to August in northern Europe), with flowering starting at the bottom of the spike and progressing irregularly upward; each flower opens for part of a day and only a few open at the same time around the stem. 874:"woolly mullein", "velvet mullein", or "blanket mullein", "beggar's blanket", "Moses' blanket", "poor man's blanket", "Our Lady's blanket", or "old man's blanket", and "feltwort", and so on ("flannel" is another common generic name). "Mullein" itself derives from the French word for "soft". 1097:
long after previous plants have died. Its population pattern typically consists of an ephemeral adult population followed by a long period of dormancy as seeds. Great mullein rarely establishes on new grounds without human intervention because its seeds do not disperse very far.
881:" (a name it shares with a number of other plants with tall, yellow inflorescences), and a plethora of other "X's staff" and "X's rod". The name "velvet dock" or "mullein dock" is also recorded, where "dock" is a British name applied to any broad-leaved plant. 1046:
so when exposed to light, or very close to the soil surface, which explains the plant's habitat preferences. While it can also grow in areas where some vegetation already exists, growth of the rosettes on bare soil is four to seven times more rapid.
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Roman soldiers are said to have dipped the plant stalks in grease for use as torches. Other cultures use the leaves as wicks. Native Americans and American colonists lined their shoes with leaves from the plant to keep out the cold.
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Because it cannot compete with established plants, great mullein is no longer considered a serious agricultural weed and is easily crowded out in cultivation, except in areas where vegetation is sparse to begin with, such as
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Parker, Ingrid M.; Joseph Rodriguez; Michael E. Loik (February 2003). "An Evolutionary Approach to Understanding the Biology of Invasions: Local Adaptation and General-Purpose Genotypes in the Weed Verbascum thapsus".
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is known by a variety of names. European reference books call it "great mullein". In North America, "common mullein" is used while western United States residents commonly refer to mullein as "cowboy toilet paper".
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was specified, as the practice only arose later, in the 19th century. When a lectotype (type selected amongst original material) was designated, it was assigned to specimen 242.1 of Linnaeus' herbarium, the only
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Eaton went so far as to write: "When botanists are so infatuated with wild speculation, as to tell us the mullein was introduced, they give our youngest pupils occasion to sneer at their teachers."
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are not taxonomical, Mürbeck's treatment is the most current one available, as no study has yet sought to apply genetic or molecular data extensively to the genus. In Mürbeck's classification,
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of leaves. It grows in a wide variety of habitats, but prefers well-lit, disturbed soils, where it can appear soon after the ground receives light, from long-lived seeds that persist in the
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in temperate world regions. It is a minor problem for most agricultural crops, since it is not a competitive species, being intolerant of shade from other plants and unable to survive
1347:). The plant's ability to host both pests and beneficials makes it potentially useful to maintain stable populations of insects used for biological control in other cultures, like 2009: 348:(2 mm, 0.08 in). The five stamens are of two types, with the three upper stamens being shorter, their filaments covered by yellow or whitish hairs, and having smaller 236:. It also hosts many insects, some of which can be harmful to other plants. Although individuals are easy to remove by hand, populations are difficult to eliminate permanently. 2574:
Attar, Farideh; et al. (2007). "Micromorphological studies on Verbascum (Scrophulariaceae) in Iran with emphasis on seed surface, capsule ornamentation and trichomes".
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Control of the plant, when desired, is best managed via mechanical means, such as hand pulling and hoeing, preferably followed by sowing of native plants. Animals rarely
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Sotoodeh, Arash (2014). "Verbascum oreophilum var. oreophilum and Verbascum cheiranthifolium var. asperulum (Scrophulariaceae) two new records for the flora of Iran".
1452:, are also effective, but they result in bare ground – ideal conditions for growing more mullein – and therefore require repeated application to prevent regrowth. 3381: 1445: 1232: 730: 2805: 3508: 784:
also differs from the type in having slightly larger flowers, which measure 15–30 mm wide, whereas in the type, they are 12–20 mm in diameter. Both
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populations as a whole. Goats and chickens have also been proposed to control mullein. Effective (when used with a surfactant) contact herbicides include
2605: 1340: 1302: 4356: 3914:"Numbers and types of arthropods overwintering on common mullein, Verbascum thapsus L. (Scrophulariaceae), in a central Washington fruit-growing region" 776:, the hairiness is less dense and often absent from the upper part of the anthers, while lower leaves are hardly decurrent and have longer petioles. In 3884: 4004: 1745:
87:43. Some disagree since Huber-Morath did not specifically cite sheet 242.1, and credit instead L. H. Cramer, in Dassanayake & Fosberg (1981)
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in the USA. In such ecological contexts, it crowds out native herbs and grasses; its tendency to appear after forest fires also disturbs the normal
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requires the stem to be moved by wind or animal movement; 75% of the seeds fall within 1 m of the parent plant, and 93% fall within 5 m.
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by insects during the day. While many insects visit the flowers, only some bees actually accomplish pollination. The flowering period of
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for pulmonary diseases. Leaves were smoked to attempt to treat lung ailments, a tradition that in America was rapidly transmitted to
2674: 2372: 4795: 4210: 920:. In northern Europe, it grows from sea level up to 1,850 m altitude, while in China it grows at 1,400–3,200 m altitude. 5234: 4896: 4821: 3353: 3168: 3159: 3312: 4994: 3356: 2773: 5035: 4924: 4421: 3345: 2622: 1548: 5224: 4901: 3808: 1976: 3627:
Kivilaan, A.; Robert S. Bandurski (October 1981). "The One Hundred-Year Period for Dr. Beal's Seed Viability Experiment".
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that can grow to 2 m tall or more. Its small, yellow flowers are densely grouped on a tall stem, which grows from a large
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Drury, Susan (1991). "Plants and Wart Cures in England from the Seventeenth to the Nineteenth Century: Some Examples".
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lasts from June to August in most of its range, extending to September or October in warmer climates. Visitors include
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Camzine, Scott; Bye, Robert A. (1980). "A Study Of The Medical Ethnobotany Of The Zuni Indians of New Mexico".
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Sanchez, Juan Antonio; David R. Gillespie; Robert R. McGregor (November 2003). "The effects of mullein plants (
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A synonymized checklist of the vascular flora of the United States, Canada, and Greenland (Volume 1: Checklist)
5074: 2628: 4945: 4364: 3164: 2617:(in Spanish). Vol. XIII: Plantaginaceae-Scrophulariaceae. Madrid: Real Jardín Botánico. pp. 49–97. 4321:
Turker, Arzu Ucar; N. D. Camper (October 2002). "Biological activity of common mullein, a medicinal plant".
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A given flower is open only for a single day, opening before dawn and closing in the afternoon. Flowers are
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of flowers that can occupy up to half the stem length. All parts of the plants are covered with star-shaped
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Britton, Nathaniel A.; Addison Brown (1979). "Illustrated Flora of the Northern United States and Canada".
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In the United States, it was imported very early in the 18th century and cultivated for its medicinal and
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The 1630 number in Mitch may be a typo: the beginning of the 18th century is cited in other sources.
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less than 1 mm (0.04 in) in size, marked with longitudinal ridges. A white-flowered form,
5184: 1571:. All preparations meant to be drunk have to be finely filtered to eliminate the irritating hairs. 924: 4937: 4686: 4139: 2483: 2297: 5199: 4782: 1882:. Vol. 3: Diapensiaceae to Myoporaceae. Cambridge: Cambridge University Press. p. 211. 1659: 1163: 1135: 599: 340:, the latter bright yellow and an 1.5–3 cm (0.59–1.18 in) wide. The flowers are almost 305:, with much variation in leaf shape between the upper and lower leaves on the stem, ranging from 294:. This cover is particularly thick on the leaves, giving them a silvery appearance. The species' 5143: 5061: 1122:
has been reported to consume them. Other bird species have been reported to consume the leaves (
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Ferguson, Ian Keith (1972). "V. thapsus". In Tutin, Thomas Gaskell; et al. (eds.).
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that split open by way of two valves, each capsule containing large numbers of minute, brown
167: 2314: 1314:). Useful insects are also hosted by great mullein, including predatory mites of the genera 758:(Lam.) Murb.; Mediterranean region and to 2000 metres in southwestern Austria. (syn. subsp. 5025: 4849: 4707: 4444: 4084: 3593: 3538: 3425: 2342: 2137: 1704: 1544: 1540: 1180: 1099: 1093:
for extended periods of time, and can sprout from apparently bare ground, or shortly after
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native to Europe, northern Africa, and Asia, and introduced in the Americas and Australia.
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Wilhelm, Gene (April 1974). "The mullein: Plant piscicide of the mountain folk culture".
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On flowering plants, the leaves are alternately arranged up the stem. They are thick and
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Second-year plant starting to flower, with a dead stem of the previous year, behind left
5048: 4597: 4448: 4088: 3775:, The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology. 3737:, The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornithology. 3671: 3597: 3542: 3429: 796:
has had a great many subspecies described. A recent revision led its author to maintain
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filaments and larger anthers. The plant produces small, ovoid (6 mm, 0.24 in)
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Introducing the Flowering Beauty of Glacier National Park and the Majestic High Rockies
3758:, The Birds of North America Online (A. Poole, Ed.). Ithaca: Cornell Lab of Ornitholog. 3652: 3609: 3562: 3500: 3441: 2525: 1627: 1223: 1119: 987: 848: 641: 517: 480: 341: 45: 4334: 4264: 4096: 3017:. Vol. 3. Gentianaceae to Compositae. New York: Dover Publications. p. 175. 2858: 1392: 1022: 320: 263: 32: 5020: 4668: 4487: 4398: 4338: 4268: 4218: 4180: 3977: 3928: 3804: 3712: 3644: 3554: 3492: 3437: 3324: 3043: 3018: 2942: 2941:. Vol. 1. The Northeastern States (Part 2). New York: McGraw-Hill. p. 389. 2912: 2866: 2720: 2653: 2618: 2466: 2277: 2251: 2046: 1916: 1883: 1568: 1536: 1353: 1333: 1151: 396: 295: 3672:"Habitat requirements of central European bees and the problems of partial habitats" 3504: 3445: 2960: 2784: 2529: 1054: 5156: 4673: 4452: 4330: 4295: 4260: 4092: 3969: 3800: 3636: 3601: 3546: 3484: 3433: 3360: 2686: 2583: 2556: 2517: 2450: 2384: 1702:
Due to its weedy capacities, the plant, unlike other species of the genus (such as
1650: 1249: 999: 716: 505: 438: 345: 337: 282: 229: 221: 134: 4813: 4630: 4299: 3529:): I. Latitudinal Differences in Population Dynamics and Timing of Reproduction". 1061:(with female parts maturing first), and will self-pollinate if they have not been 877:
Some names refer to the plant's size and shape: "shepherd's club(s)" or "staff", "
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describes uses of the plant for respiratory infections. It was also part of the
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Reinartz, James A. (November 1984). "Life History Variation of Common Mullein (
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Reinartz, James A. (November 1984). "Life History Variation of Common Mullein (
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to be effective, as the hair causes water to roll off the plant, much like the
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has a wide native range including Europe, northern Africa, and Asia, from the
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The Audubon Society Field Guide to North American Wildflowers, Eastern Region
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The Audubon Society field guide to North American wildflowers, eastern region
2613: 1504: 1465: 1279: 1275: 1080: 1034: 905: 457: 389: 310: 244: 183: 940: 5100: 4888: 4342: 4066: 3496: 2281: 1814: 1623: 1615: 1420:. Burning is ultimately ineffective, as it only creates new bare areas for 1417: 1322: 1307: 1192: 1105: 466: 410: 4272: 3960:
Price, W. C. (July 1940). "Comparative Host Ranges of Six Plant Viruses".
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The lectotypification is usually attributed to Arthur Huber-Morath (1971)
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were originally described as species. Due to its morphological variation,
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over the years. Introduced American populations show much less variation.
4932: 4834: 4640: 4580: 4535: 4045:. Ontario Ministry of Agriculture, Food and Rural Affairs. Archived from 3040:
Scientific and common names of 7,000 vascular plants in the United States
2005: 1845: 1672: 1619: 1552: 1449: 1370: 1366: 1316: 1295: 1188: 1094: 1077:. The hair on lower stamens may serve to provide footholds for visitors. 1062: 1042: 936: 909: 278: 5012: 3799:
The History of the British Flora, A Factual Basis for Phytogeography by
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first recommended the plant 2000 years ago, considering it useful as a
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is the type species of the genus the application of article 22 of the
4839: 4756: 4720: 3488: 3414: 3233:. IUCN Invasive Species Specialist Group. 8 July 2005. Archived from 2690: 2388: 1692: 1680: 1611: 1607: 1587: 1567:, and skin infections. An infusion of the root is also used to treat 1500: 1485: 1441: 1409: 1114:
use the hair (amongst that of various woolly plants) in making their
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In all subspecies but the type, the lower stamens are also hairy. In
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Garrett, Kimball L., Raphael, Martin G. and Dixon, Rita D. (1996).
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thought it was a native plant. In 1839, it was already reported in
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Other insects commonly found on great mullein feed exclusively on
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In book 25, Pliny describes "two principal kinds " thought to be
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The fruit of great mullein contains large numbers of minute seed.
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Banko, Paul C., Black, Jeffrey M. and Banko, Winston E. (1999).
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Harold William Rickett, ed. (1966). "The Mulleins (Verbascum)".
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California Plant Names: Latin and Greek Meanings and Derivations
1594:, and other external conditions. Topical application of various 4712: 4009:
Biological Control: A Guide to Natural Enemies in North America
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Denkschriften der Schweizerischen Naturforschenden Gesellschaft
1591: 1497: 1489: 1159: 1127: 1038: 991: 983: 944: 893: 526: 442: 349: 329: 287: 268: 4363:. American Botanical Council. February 1, 1990. Archived from 3093:
Coutts, J; Osborn, A; Edwards, A (1954). Preston, G.H. (ed.).
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Flora – Morphology, Distribution, Functional Ecology of Plants
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It has been introduced throughout the temperate world, and is
3857:. USDA Natural Resources Conservation Service. Archived from 3828:. USDA Natural Resources Conservation Service. Archived from 3015:
An illustrated flora of the northern United States and Canada
2783:. USDA Natural resources Conservation Service. Archived from 1668: 1664: 1634: 1579: 1262: 960: 948: 901: 252: 62: 3097:. Great Britain: Ward, Lock & Co., Limited. p. 255. 2250:. Philadelphia: George W. Carpenter Fund. pp. 173–174. 4507: 4111:"HOSTS – a Database of the World's Lepidopteran Hostplants" 3883:. Victoria Department of Primary Industries. Archived from 3626: 2573: 2543:
Sotoodeh, Arash; Civeyrel, L.; Attar, F. (19 March 2015). "
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degradation activated by low temperatures in the root, and
928: 361: 4217:. Vol. 3: L-R (2nd ed.). Detroit: Thomson Gale. 3200:(in Spanish). Universidad Nacional del Sur. Archived from 2715:
Schmeil, Otto; Fitschen, Jost; Seybold, Siegmund (2006).
2091:. Plant Conservation Alliance's Alien Plant Working Group 1598:-based preparations was recommended for the treatment of 970:
properties. By 1818, it had begun spreading so much that
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Journal of the Entomological Society of British Columbia
3266:. Hawaiian Ecosystems at Risk project. February 19, 2007 1622:
effects in vitro were isolated from flowers. The German
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specimen. The species had previously been designated as
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JLindquist.com: webpage with pictures of tall specimens
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A City Herbal: Lore, Legend, & Uses of Common Weeds
2719:. Wiebelsheim: Quelle & Meyer Verlag. p. 496. 2428: 2248:
The Scrophulariaceae of eastern temperate North America
1691:) that are toxic to fish, and have been widely used as 858:
W.D.J.Koch), have also been reported in North America.
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Base de Datos sobre Invasiones Biológicas en Argentina
2936: 2672: 2547:(Scrophulariaceae), a new species for Flora of Iran". 2429:
Gross, Katherine L.; Patricia A. Werner (April 1978).
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The plant has been used in an attempt to treat colds,
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region, and regionally controlled in several others).
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in a wide range of habitats has been linked to strong
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Elsevier's Dictionary of Plant Names and their Origin
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Brako, Lois; Rossman, Amy Y.; Farr, David F. (1995).
2816:. Joint Nature Conservation Committee. Archived from 2542: 1784:. The precise attribution of a third kind is unclear. 1210:
Despite not being an agricultural weed in itself, it
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variation, which has led to the plant acquiring many
3012: 2138:"Element Stewardship Abstract for Verbascum thapsus" 5215:
Plants used in traditional Native American medicine
3912:Horton, David R.; Tamera M. Lewis (December 2003). 3092: 2276:(in French). Louvain: Publications Universitaires. 2274:
Dictionnaire étymologique des noms grecs de plantes
2221:. California Invasive Plant Council. Archived from 1169: 1033:Great mullein is a biennial and generally requires 247:are made from this plant. It has been used to make 4320: 4168: 1199:(class C) and Hawaii, and the Australian state of 3464:Turker, Arzu Ucar; Ekrem Gurel (September 2005). 3463: 2865:. Vol. 2: I-Z. New York: Dover Publication. 1278:. These make the plant a potential reservoir for 1037:before it can flower. This dormancy is linked to 982:. It is now found commonly in all the states. In 5171: 3698:"Maintenance Behavior of the American Goldfinch" 3037: 2337: 2335: 537:as the correct nomenclature for this placement. 445:, though it is often assimilated to the ancient 4389: 4075:(Heteroptera: Miridae) in tomato greenhouses". 3584:): III. Differences Among Sequential Cohorts". 3042:. St. Paul, Minnesota: APS Press. p. 189. 2652:. Portland, Oregon: Timber Press. p. 588. 2603: 1408:it because of its irritating hairs, and liquid 1400:is resistant to grazing and contact herbicides. 2890: 2888: 2755:(in Catalan). Universitat de les Illes Balears 1138:). Additionally, deer and elk eat the leaves. 4215:The Gale encyclopedia of alternative medicine 2993:An Introduction to the Flora of Central Texas 2332: 2131: 2129: 2127: 2125: 2020:. World Biodiversity Database. Archived from 1191:pastures. The species is legally listed as a 807:The plant is also parent to several hybrids ( 4158: 4156: 2753:Herbario Virtual del Mediterráneo Occidental 2123: 2121: 2119: 2117: 2115: 2113: 2111: 2109: 2107: 2105: 1293:in particular. They include mullein thrips ( 1274:), and various spider mites from the family 328:Flowers are pentamerous with (usually) five 4204: 4202: 4200: 4198: 4196: 4043:control whiteflies in greenhouse tomatoes?" 2885: 2163: 2161: 2159: 2157: 884: 423: 414: 394: 4385: 4383: 4381: 4248: 4209:Hanrahan, Claire; Rebecca J. Frey (2005). 3313:"Common Mullein—the Roadside Torch Parade" 2932: 2930: 2928: 2710: 2708: 2706: 2643: 2641: 2599: 2597: 2082: 2080: 2078: 2000: 1998: 1996: 1994: 1992: 1911:Blamey, M., & Grey-Wilson, C. (1989). 1226:. It is also a potential reservoir of the 31: 4162: 4153: 3907: 3905: 3903: 3901: 3803:, Cambridge University Press, Cambridge, 2852: 2850: 2848: 2846: 2844: 2842: 2840: 2838: 2836: 2834: 2454: 2267: 2265: 2241: 2239: 2135: 2102: 2076: 2074: 2072: 2070: 2068: 2066: 2064: 2062: 2060: 2058: 2008:van der Meijden, R.; de Kort, I. (eds.). 1967: 1747:A Revised Handbook to the Flora of Ceylon 1630:in the United States and United Kingdom. 1306:(whose larva consume the seeds), and the 1002:, with scattered populations in between. 540: 4193: 4036: 3685:. London: Academic Press. pp. 1–16. 3669: 3579: 3524: 2683:Linnaean Plant Name Typification Project 2507: 2424: 2381:Linnaean Plant Name Typification Project 2219:Invasive Plants of California's Wildland 2208: 2206: 2204: 2202: 2200: 2198: 2196: 2194: 2192: 2190: 2154: 2143:. The Global Invasive Species Initiative 1963: 1961: 1907: 1905: 1903: 1901: 1899: 1875: 1518: 1391: 1130:), or to use the plant as a source when 1079: 1021: 319: 262: 4432: 4378: 4037:Ferguson, Gillian (26 September 2005). 3695: 3459: 3457: 3455: 3169:United States Department of Agriculture 3160:Germplasm Resources Information Network 3145: 3143: 2925: 2906: 2703: 2647: 2638: 2594: 2422: 2420: 2418: 2416: 2414: 2412: 2410: 2408: 2406: 2404: 2086: 1989: 1935: 1933: 1931: 1929: 1840: 1838: 1836: 1675:. The seeds contain several compounds ( 1637:, sunburn, and other skin irritations. 1514: 475:. European plants exhibit considerable 5172: 3898: 3784: 3357:Natural Resources Conservation Service 2978: 2953: 2856: 2831: 2271: 2262: 2245: 2236: 2055: 1871: 1869: 1867: 1865: 1863: 1809: 1807: 1805: 1803: 1801: 1574:Oil from the flowers was used against 1539:are manufactured from its components. 4547: 4546: 4285: 3959: 3822:"Colorado State-listed Noxious Weeds" 3304: 3125: 2187: 2035: 1958: 1896: 5124:b1c2668d-df7d-42f8-91e3-d1974580361b 4987:9fc4c943-a190-423c-917d-8a7a5e7f052b 4132: 3878: 3681:. Linnean Society Symposium Series, 3620: 3452: 3215: 3140: 2798: 2461:Reprinted in Mulligan, G.A. (1979), 2431:"The Biology of Canadian Weeds. 28. 2401: 2312: 2212: 1977:University of California at Berkeley 1926: 1913:Flora of Britain and Northern Europe 1833: 1150:have been recorded from part of the 823:). All occur in Eurasia, and three, 4521:Microphotographies of great mullein 3851:"Hawaii State-listed Noxious Weeds" 3696:Coutlee, Ellen L. (December 1963). 3677:. In Matheson; et al. (eds.). 2965:Texas Parks & Wildlife Magazine 1860: 1798: 1511:, although quantities are unknown. 1029:grows best with little competition. 429:) for an unspecified herb from the 352:, while the lower two stamens have 13: 5230:Introduced plants of South America 4475: 4002: 2717:Flora von Deutschland, 93. Auflage 2136:Hoshovsky, Marc C. (August 2001). 1523:"Verbasci flos": dried flowers of 1396:Because of ample irritating hair, 1179:semidesertic areas of the eastern 14: 5246: 4501: 3470:L.): Recent Advances in Research" 3310: 3264:Pacific Island Ecosystems at Risk 2939:Wild Flowers of the United States 2490:. Northern Ontario Plant Database 2443:Canadian Journal of Plant Science 2353:. Institute for Systematic Botany 2246:Pennel, Francis Whittier (1935). 2167: 2004: 1724:They are all hairy in subspecies 1460: 5075:urn:lsid:ipni.org:names:770183-1 4213:. In Jacqueline L. Longe (ed.). 4175:. Ash Tree Publishing. pp.  4071:) on the population dynamics of 3789:. Glacier Park, Inc. p. 42. 3438:10.1046/j.1523-1739.2003.02019.x 3231:Global Invasive Species Database 3137:, retrieved on November 6, 2009. 2911:. New York: Knopf. p. 798. 2604:Benedí Gonzalez, Carles (2009). 2351:Atlas of Florida vascular Plants 2303:, retrieved on November 6, 2009. 1649:Mullein may be cultivated as an 1248:hosts insects from 29 different 1170:Agricultural impacts and control 1141: 811:). Of these, the most common is 49: 4486:. Amsterdam: Elsevier Science. 4426: 4411: 4349: 4314: 4279: 4242: 4103: 4060: 4030: 3996: 3953: 3872: 3843: 3814: 3793: 3778: 3761: 3740: 3723: 3689: 3663: 3573: 3518: 3408: 3374: 3338: 3321:Weed Science Society of America 3277: 3248: 3182: 3113: 3101: 3086: 3068: 3056: 3031: 3006: 2900: 2766: 2733: 2673:Natural History Museum (2022). 2666: 2567: 2536: 2501: 2476: 2371:Natural History Museum (2022). 2364: 2306: 2288: 2043:New RHS Dictionary of Gardening 1770: 1761: 1752: 1735: 1718: 1663:), great mullein was linked to 1252:. Most of the pests found were 959:. It has also been reported in 861: 768:(Willk.) Nyman; Spain, endemic. 5235:Flora naturalised in Australia 4393:; Olmstead, Nancy C. (1985) . 3405:, retrieved December 29, 2006. 3095:The Complete Book of Gardening 2018:Interactive Flora of NW Europe 1432:usually have little effect on 1203:(regionally prohibited in the 762:(Scrad.) Bonnier & Layens) 258: 1: 4335:10.1016/S0378-8741(02)00186-1 4300:10.1080/0015587X.1991.9715809 4265:10.1016/S0378-8741(80)81017-8 4097:10.1016/S1049-9644(03)00116-6 4005:"Campylomma verbasci (Meyer)" 3292:Japan Ministry of Environment 3165:Agricultural Research Service 2463:The Biology of Canadian Weeds 2174:Weedy Wildflowers of Illinois 1848:. Drugs.com. 23 November 2020 1791: 1640: 741:The three usually recognized 4323:Journal of Ethnopharmacology 4252:Journal of Ethnopharmacology 2907:Niering, William A. (1979). 2749:(Schrad.) Bonnier et layens" 1708:), is not often cultivated. 1559:of powdered root applied to 1555:, however, use the plant in 7: 5225:Taxa named by Carl Linnaeus 4361:The Commission E Monographs 2588:10.1016/j.flora.2006.04.001 1472:flowers and leaves include 986:, it is most common in the 375: 10: 5251: 4481: 3962:American Journal of Botany 3629:American Journal of Botany 3119: 3107: 3062: 2894: 2857:Grieve, Margaret (1982) . 2685:. Natural History Museum. 2561:10.11646/phytotaxa.203.1.8 2522:10.11646/phytotaxa.178.3.6 2383:. Natural History Museum. 1915:. Hodder & Stoughton, 1448:. Ground herbicides, like 1258:Frankliniella occidentalis 1017: 521:(wavy-leaved mullein). As 503:alongside species such as 239:Although commonly used in 4555: 4163:Silverman, Maida (1977). 3785:Reiner, Ralph E. (1969). 3769:White-headed Woodpecker ( 3317:Intriguing World of Weeds 2545:Verbascum shahsavarensis 509:(black or dark mullein), 415: 173: 166: 46:Scientific classification 44: 39: 30: 23: 5220:Plants described in 1753 3679:The Conservation of Bees 2986:"BIO 406D-Native Plants" 2041:Huxley, A., ed. (1992). 1821:. CABI. 20 November 2018 1711: 1214:a number of insects and 885:Distribution and habitat 324:A closeup of the flowers 5195:Flora of temperate Asia 3811:, 1975 edition page 318 3670:Westrich, Paul (1996). 1653:. As for many plants, ( 1455: 1369:species, including the 1164:Ipswichian interglacial 1162:from some parts of the 1136:white-headed woodpecker 409:had been first used by 388:was first described by 4482:Watts, Donald (2000). 4397:. Knopf. p. 653. 4115:Natural History Museum 3586:The Journal of Ecology 3531:The Journal of Ecology 2995:. Michael Gruenstaeudl 2648:Kartesz, John (1994). 2298:Den virtuella Floran: 2176:. Illinois Wildflowers 1970:"Verbascum thapsus L." 1968:Wetherwax, M. (1993). 1531:Although long used in 1528: 1401: 1289:species in general or 1085: 1030: 541:Subspecies and hybrids 424: 395: 382:botanical nomenclature 336:, and a five-petalled 325: 307:oblong to oblanceolate 298:is 2n = 36. 281:plant that produces a 271: 267:A stemless rosette in 5190:Flora of North Africa 4513:The type specimen of 4142:. Plants For A Future 3771:Picoides albolarvatus 3477:Phytotherapy Research 3190:"Ficha de la Especie 2313:Charters, Michael L. 2087:Remaley, Tom (1998). 1527:as used in herbal tea 1522: 1446:sulfurometuron-methyl 1395: 1254:western flower thrips 1233:Erysiphum cichoraceum 1228:cucumber mosaic virus 1185:ecological succession 1083: 1025: 600:V. pulverulentum 515:(white mullein), and 323: 286:stems end in a dense 266: 3418:Conservation Biology 3390:NatureServe Explorer 2787:on November 19, 2010 1973:Jepson Manual online 1657:described it in his 1515:Traditional medicine 1266:species such as the 802:V. crassifolium 698:W.D.J.Koch, may be a 241:traditional medicine 4449:1974GeoRv..64..235W 4436:Geographical Review 4391:Niering, William A. 4259:(4): 365–388, 378. 4140:"Verbascum thapsus" 4089:2003BiolC..28..313S 4049:on January 27, 2006 4003:Higbee, Bradley S. 3942:on October 16, 2010 3733:Branta sandvicensis 3598:1984JEcol..72..927R 3543:1984JEcol..72..897R 3430:2003ConBi..17...59P 3363:on October 16, 2010 3346:"Verbascum thapsus" 2814:Wildlife Statistics 2272:Carnoy, A. (1959). 2215:"Verbascum thapsus" 2213:Pitcairn, Michael. 2089:"Verbascum thapsus" 1941:"Verbascum thapsus" 1610:, hemorrhoids, and 1365:. A number of pest 1349:Campylomma verbasci 1345:Campylomma verbasci 1268:tarnished plant bug 1195:in the US state of 1008:phenotype variation 916:, and south to the 752:; type, widespread. 731:V. pyramidatum 710:V. × semialbum 550: 512:Verbascum lychnitis 495:is placed in sect. 380:For the purpose of 4077:Biological Control 4013:Cornell University 3861:on October 8, 2010 3754:Loxioides bailleui 2961:"Beyond the River" 2677:Verbascum × thapsi 2456:10.4141/cjps78-062 1705:V. phoeniceum 1660:Naturalis Historia 1628:National Formulary 1614:, amongst others. 1529: 1402: 1224:beneficial insects 1120:American goldfinch 1108:bees of the genus 1086: 1031: 988:Maritime Provinces 790:V. t. crassifolium 782:V. t. crassifolium 774:V. t. crassifolium 756:V. t. crassifolium 621:V. phlomoides 545: 518:Verbascum sinuatum 344:, with very short 326: 272: 209:, is a species of 5167: 5166: 5021:Open Tree of Life 4977:Verbascum~thapsus 4700:Verbascum_thapsus 4661:verbascum-thapsus 4587:Verbascum thapsus 4557:Verbascum thapsus 4549:Taxon identifiers 4515:Verbascum thapsus 4073:Dicyphus hesperus 4069:Verbascum thapsus 4041:Dicyphus hesperus 3887:on March 15, 2011 3582:Verbascum Thapsus 3527:Verbascum Thapsus 3468:Verbascum thapsus 3466:"Common Mullein ( 3384:Verbascum thapsus 3258:Verbascum thapsus 3225:Verbascum thapsus 3192:Verbascum thapsus 3153:Verbascum thapsus 3134:Verbascum thapsus 2808:Verbascum thapsus 2743:Verbascum thapsus 2624:978-84-00-08747-0 2486:Verbascum thapsus 2437:V. blattaria 2433:Verbascum thapsus 2375:Verbascum thapsus 2345:Verbascum thapsus 2300:Verbascum thapsus 2014:(Mullein, Great)" 2012:Verbascum thapsus 1819:(common mullein)" 1817:Verbascum thapsus 1430:Cucullia verbasci 1389:as a host plant. 1361:), a predator of 1354:Dicyphus hesperus 1341:mullein plant bug 1337:Orius tristicolor 1334:minute pirate bug 1312:Cucullia verbasci 1303:Gymnaetron tetrum 1218:, including both 1152:Cromer Forest Bed 943:, North America, 939:, tropical Asia, 798:V. giganteum 739: 738: 684:V. lychnitis 663:V. blattaria 579:V. speciosum 548:Verbascum thapsus 523:Verbascum thapsus 397:Species Plantarum 386:Verbascum thapsus 296:chromosome number 194:Verbascum thapsus 190: 189: 177:Verbascum thapsus 25:Verbascum thapsus 5242: 5210:Medicinal plants 5160: 5159: 5147: 5146: 5137: 5136: 5127: 5126: 5114: 5113: 5104: 5103: 5091: 5090: 5078: 5077: 5065: 5064: 5052: 5051: 5039: 5038: 5029: 5028: 5016: 5015: 5003: 5002: 4990: 4989: 4980: 4979: 4967: 4966: 4954: 4953: 4951:NBNSYS0000004046 4941: 4940: 4928: 4927: 4915: 4914: 4905: 4904: 4892: 4891: 4879: 4878: 4866: 4865: 4853: 4852: 4843: 4842: 4830: 4829: 4817: 4816: 4804: 4803: 4791: 4790: 4778: 4777: 4765: 4764: 4752: 4751: 4742: 4741: 4729: 4728: 4716: 4715: 4703: 4702: 4690: 4689: 4677: 4676: 4664: 4663: 4654: 4653: 4644: 4643: 4634: 4633: 4621: 4620: 4611: 4610: 4601: 4600: 4591: 4590: 4589: 4576: 4575: 4574: 4544: 4543: 4497: 4469: 4468: 4430: 4424: 4415: 4409: 4408: 4387: 4376: 4375: 4373: 4372: 4357:"Mullein flower" 4353: 4347: 4346: 4329:(2–3): 117–125. 4318: 4312: 4311: 4283: 4277: 4276: 4246: 4240: 4239: 4237: 4236: 4231:on July 15, 2009 4227:. Archived from 4206: 4191: 4190: 4174: 4160: 4151: 4150: 4148: 4147: 4136: 4130: 4129: 4127: 4126: 4121:on June 27, 2008 4117:. Archived from 4107: 4101: 4100: 4064: 4058: 4057: 4055: 4054: 4034: 4028: 4027: 4025: 4024: 4015:. Archived from 4000: 3994: 3993: 3957: 3951: 3950: 3948: 3947: 3941: 3935:. Archived from 3918: 3909: 3896: 3895: 3893: 3892: 3876: 3870: 3869: 3867: 3866: 3847: 3841: 3840: 3838: 3837: 3818: 3812: 3801:Sir Harry Godwin 3797: 3791: 3790: 3782: 3776: 3765: 3759: 3744: 3738: 3731:Hawaiian Goose ( 3727: 3721: 3720: 3702: 3693: 3687: 3686: 3676: 3667: 3661: 3660: 3635:(9): 1290–1292. 3624: 3618: 3617: 3577: 3571: 3570: 3522: 3516: 3515: 3513: 3507:. Archived from 3489:10.1002/ptr.1653 3474: 3461: 3450: 3449: 3412: 3406: 3404: 3402: 3401: 3392:. Archived from 3378: 3372: 3371: 3369: 3368: 3359:. Archived from 3342: 3336: 3335: 3333: 3332: 3323:. Archived from 3311:Mitch, Larry W. 3308: 3302: 3301: 3299: 3298: 3289: 3281: 3275: 3274: 3272: 3271: 3252: 3246: 3245: 3243: 3242: 3219: 3213: 3212: 3210: 3209: 3186: 3180: 3179: 3177: 3175: 3147: 3138: 3132:Flora Europaea: 3129: 3123: 3117: 3111: 3105: 3099: 3098: 3090: 3084: 3083: 3072: 3066: 3060: 3054: 3053: 3035: 3029: 3028: 3010: 3004: 3003: 3001: 3000: 2990: 2982: 2976: 2975: 2973: 2972: 2957: 2951: 2950: 2934: 2923: 2922: 2904: 2898: 2892: 2883: 2882: 2880: 2879: 2859:"Mullein, Great" 2854: 2829: 2828: 2826: 2825: 2802: 2796: 2795: 2793: 2792: 2770: 2764: 2763: 2761: 2760: 2737: 2731: 2730: 2712: 2701: 2700: 2698: 2697: 2691:10.5519/qwv6u7j5 2670: 2664: 2663: 2645: 2636: 2635: 2633: 2627:. Archived from 2610: 2601: 2592: 2591: 2571: 2565: 2564: 2540: 2534: 2533: 2505: 2499: 2498: 2496: 2495: 2480: 2474: 2460: 2458: 2426: 2399: 2398: 2396: 2395: 2389:10.5519/qwv6u7j5 2368: 2362: 2361: 2359: 2358: 2339: 2330: 2329: 2327: 2326: 2310: 2304: 2296: 2292: 2286: 2285: 2269: 2260: 2259: 2243: 2234: 2233: 2231: 2230: 2210: 2185: 2184: 2182: 2181: 2165: 2152: 2151: 2149: 2148: 2142: 2133: 2100: 2099: 2097: 2096: 2084: 2053: 2039: 2033: 2032: 2030: 2029: 2002: 1987: 1986: 1984: 1983: 1965: 1956: 1955: 1953: 1952: 1937: 1924: 1909: 1894: 1893: 1873: 1858: 1857: 1855: 1853: 1842: 1831: 1830: 1828: 1826: 1811: 1785: 1782:V. sinuatum 1774: 1768: 1765: 1759: 1756: 1750: 1739: 1733: 1722: 1651:ornamental plant 1375:Papaipema nebris 1244:. A study found 1055:self-fecundating 1000:British Columbia 978:and in 1876, in 900:east to western 642:V. virgatum 551: 544: 506:Verbascum nigrum 486:The taxonomy of 456:At the time, no 427: 420: 419: 404:specific epithet 400: 179: 135:Scrophulariaceae 54: 53: 35: 21: 20: 16:Species of plant 5250: 5249: 5245: 5244: 5243: 5241: 5240: 5239: 5185:Flora of Europe 5170: 5169: 5168: 5163: 5155: 5150: 5142: 5140: 5132: 5130: 5122: 5117: 5109: 5107: 5099: 5094: 5086: 5081: 5073: 5068: 5060: 5055: 5047: 5042: 5034: 5032: 5024: 5019: 5011: 5008:Observation.org 5006: 4998: 4993: 4985: 4983: 4975: 4970: 4962: 4957: 4949: 4944: 4936: 4931: 4923: 4918: 4910: 4909:MichiganFlora: 4908: 4900: 4895: 4887: 4882: 4874: 4869: 4861: 4856: 4848: 4846: 4838: 4833: 4825: 4820: 4812: 4807: 4799: 4794: 4786: 4781: 4773: 4768: 4760: 4755: 4747: 4745: 4737: 4732: 4724: 4719: 4711: 4706: 4698: 4693: 4685: 4680: 4672: 4667: 4659: 4657: 4649: 4647: 4639: 4637: 4629: 4624: 4616: 4614: 4606: 4604: 4596: 4594: 4585: 4584: 4579: 4570: 4569: 4564: 4551: 4504: 4494: 4478: 4476:Further reading 4473: 4472: 4431: 4427: 4416: 4412: 4405: 4388: 4379: 4370: 4368: 4355: 4354: 4350: 4319: 4315: 4284: 4280: 4247: 4243: 4234: 4232: 4225: 4207: 4194: 4187: 4161: 4154: 4145: 4143: 4138: 4137: 4133: 4124: 4122: 4109: 4108: 4104: 4065: 4061: 4052: 4050: 4035: 4031: 4022: 4020: 4001: 3997: 3974:10.2307/2437088 3958: 3954: 3945: 3943: 3939: 3916: 3910: 3899: 3890: 3888: 3881:"Great Mullein" 3879:Faithful, Ian. 3877: 3873: 3864: 3862: 3855:PLANTS Database 3849: 3848: 3844: 3835: 3833: 3832:on July 2, 2009 3826:PLANTS Database 3820: 3819: 3815: 3798: 3794: 3783: 3779: 3766: 3762: 3746:Banko, Paul C. 3745: 3741: 3728: 3724: 3705:Wilson Bulletin 3700: 3694: 3690: 3674: 3668: 3664: 3641:10.2307/2443054 3625: 3621: 3606:10.2307/2259541 3578: 3574: 3551:10.2307/2259539 3523: 3519: 3511: 3472: 3462: 3453: 3413: 3409: 3399: 3397: 3380: 3379: 3375: 3366: 3364: 3350:PLANTS Database 3344: 3343: 3339: 3330: 3328: 3327:on July 2, 2009 3309: 3305: 3296: 3294: 3287: 3283: 3282: 3278: 3269: 3267: 3254: 3253: 3249: 3240: 3238: 3221: 3220: 3216: 3207: 3205: 3188: 3187: 3183: 3173: 3171: 3149: 3148: 3141: 3130: 3126: 3122:, pp. 302, 634. 3118: 3114: 3106: 3102: 3091: 3087: 3074: 3073: 3069: 3065:, pp. 108, 369. 3061: 3057: 3050: 3036: 3032: 3025: 3011: 3007: 2998: 2996: 2988: 2984: 2983: 2979: 2970: 2968: 2959: 2958: 2954: 2935: 2926: 2919: 2905: 2901: 2893: 2886: 2877: 2875: 2873: 2863:A Modern Herbal 2855: 2832: 2823: 2821: 2806:"Great Mullein 2804: 2803: 2799: 2790: 2788: 2772: 2771: 2767: 2758: 2756: 2739: 2738: 2734: 2727: 2713: 2704: 2695: 2693: 2671: 2667: 2660: 2646: 2639: 2631: 2625: 2608: 2602: 2595: 2572: 2568: 2541: 2537: 2506: 2502: 2493: 2491: 2482: 2481: 2477: 2427: 2402: 2393: 2391: 2369: 2365: 2356: 2354: 2341: 2340: 2333: 2324: 2322: 2315:"Plant name: T" 2311: 2307: 2294: 2293: 2289: 2270: 2263: 2244: 2237: 2228: 2226: 2211: 2188: 2179: 2177: 2170:"Great Mullein" 2166: 2155: 2146: 2144: 2140: 2134: 2103: 2094: 2092: 2085: 2056: 2040: 2036: 2027: 2025: 2003: 1990: 1981: 1979: 1966: 1959: 1950: 1948: 1939: 1938: 1927: 1910: 1897: 1890: 1874: 1861: 1851: 1849: 1844: 1843: 1834: 1824: 1822: 1813: 1812: 1799: 1794: 1789: 1788: 1778:V. thapsus 1775: 1771: 1766: 1762: 1757: 1753: 1740: 1736: 1726:V. crassifolium 1723: 1719: 1714: 1655:Pliny the Elder 1643: 1618:compounds with 1596:V. thapsus 1549:Native American 1533:herbal medicine 1517: 1478:polysaccharides 1463: 1458: 1434:V. thapsus 1398:V. thapsus 1387:V. thapsus 1383:Strymon melinus 1379:gray hairstreak 1291:V. thapsus 1246:V. thapsus 1172: 1144: 1100:Seed dispersion 1089:remains in the 1067:V. thapsus 1035:winter dormancy 1027:V. thapsus 1020: 904:, north to the 887: 867:V. thapsus 864: 804:into synonymy. 794:V. thapsus 786:V. t. giganteum 778:V. t. giganteum 766:V. t. giganteum 699: 697: 559:parent species 558: 543: 493:V. thapsus 463:V. thapsus 378: 366:V. thapsus 332:, a five-lobed 261: 203:greater mullein 186: 181: 175: 162: 159:V. thapsus 48: 17: 12: 11: 5: 5248: 5238: 5237: 5232: 5227: 5222: 5217: 5212: 5207: 5202: 5200:Flora of Spain 5197: 5192: 5187: 5182: 5165: 5164: 5162: 5161: 5157:wfo-0000419130 5148: 5138: 5134:common-mullein 5128: 5115: 5105: 5092: 5079: 5066: 5053: 5040: 5030: 5017: 5004: 4991: 4981: 4968: 4955: 4942: 4929: 4916: 4906: 4893: 4880: 4867: 4854: 4844: 4831: 4818: 4805: 4792: 4779: 4766: 4753: 4743: 4730: 4717: 4704: 4691: 4678: 4665: 4655: 4645: 4635: 4622: 4612: 4602: 4592: 4577: 4561: 4559: 4553: 4552: 4539: 4538: 4533: 4523: 4518: 4510: 4503: 4502:External links 4500: 4499: 4498: 4492: 4477: 4474: 4471: 4470: 4457:10.2307/213812 4443:(2): 235–252. 4425: 4410: 4403: 4377: 4348: 4313: 4278: 4241: 4223: 4192: 4185: 4152: 4131: 4102: 4083:(3): 313–319. 4059: 4029: 3995: 3968:(7): 530–541. 3952: 3897: 3871: 3842: 3813: 3792: 3777: 3760: 3739: 3722: 3711:(4): 342–357. 3688: 3662: 3619: 3592:(3): 927–936. 3572: 3537:(3): 897–912. 3517: 3514:on 2015-02-13. 3483:(9): 733–739. 3451: 3407: 3373: 3337: 3303: 3276: 3247: 3214: 3181: 3139: 3124: 3112: 3100: 3085: 3076:"mullein (n.)" 3067: 3055: 3048: 3030: 3023: 3005: 2977: 2967:. Dale Weisman 2952: 2924: 2917: 2899: 2897:, pp. 633–634. 2884: 2871: 2830: 2797: 2781:PLANTS Profile 2765: 2732: 2725: 2702: 2665: 2658: 2637: 2634:on 2011-05-13. 2623: 2593: 2582:(2): 169–175. 2566: 2535: 2516:(3): 205–210. 2500: 2475: 2473:, pp. 320–331. 2449:(2): 401–413. 2400: 2363: 2331: 2321:. Calflora.net 2305: 2287: 2261: 2235: 2186: 2153: 2101: 2054: 2034: 1988: 1957: 1945:Flora of China 1925: 1895: 1888: 1879:Flora Europaea 1859: 1832: 1796: 1795: 1793: 1790: 1787: 1786: 1769: 1760: 1751: 1734: 1716: 1715: 1713: 1710: 1642: 1639: 1569:athlete's foot 1516: 1513: 1505:essential oils 1466:Phytochemicals 1462: 1461:Phytochemicals 1459: 1457: 1454: 1242:Texas root rot 1238:powdery mildew 1236:(the cucurbit 1205:West Gippsland 1171: 1168: 1156:West Wittering 1154:series and at 1143: 1140: 1126:) or flowers ( 1124:Hawaiian goose 1091:soil seed bank 1019: 1016: 898:Canary Islands 886: 883: 863: 860: 821:V. nigrum 770: 769: 763: 753: 737: 736: 734: 727: 723: 722: 720: 717:V. nigrum 713: 706: 705: 702:nomen ambiguum 687: 680: 669: 668: 666: 659: 648: 647: 645: 638: 627: 626: 624: 617: 606: 605: 603: 596: 585: 584: 582: 575: 572:duernsteinense 564: 563: 560: 555: 542: 539: 437:, near modern 433:settlement of 377: 374: 279:dicotyledonous 260: 257: 245:approved drugs 226:soil seed bank 218:biennial plant 216:It is a hairy 207:common mullein 188: 187: 182: 171: 170: 164: 163: 156: 154: 150: 149: 142: 138: 137: 132: 128: 127: 122: 118: 117: 112: 105: 104: 99: 92: 91: 86: 79: 78: 73: 66: 65: 60: 56: 55: 42: 41: 37: 36: 28: 27: 15: 9: 6: 4: 3: 2: 5247: 5236: 5233: 5231: 5228: 5226: 5223: 5221: 5218: 5216: 5213: 5211: 5208: 5206: 5203: 5201: 5198: 5196: 5193: 5191: 5188: 5186: 5183: 5181: 5178: 5177: 5175: 5158: 5153: 5149: 5145: 5139: 5135: 5129: 5125: 5120: 5116: 5112: 5106: 5102: 5097: 5093: 5089: 5084: 5080: 5076: 5071: 5067: 5063: 5058: 5054: 5050: 5045: 5041: 5037: 5031: 5027: 5022: 5018: 5014: 5009: 5005: 5001: 4996: 4992: 4988: 4982: 4978: 4973: 4969: 4965: 4960: 4956: 4952: 4947: 4943: 4939: 4934: 4930: 4926: 4921: 4917: 4913: 4907: 4903: 4898: 4894: 4890: 4885: 4881: 4877: 4872: 4868: 4864: 4859: 4855: 4851: 4845: 4841: 4836: 4832: 4828: 4823: 4819: 4815: 4810: 4806: 4802: 4797: 4793: 4789: 4784: 4780: 4776: 4771: 4767: 4763: 4758: 4754: 4750: 4744: 4740: 4735: 4731: 4727: 4722: 4718: 4714: 4709: 4705: 4701: 4696: 4692: 4688: 4683: 4679: 4675: 4670: 4666: 4662: 4656: 4652: 4646: 4642: 4636: 4632: 4627: 4623: 4619: 4613: 4609: 4603: 4599: 4593: 4588: 4582: 4578: 4573: 4567: 4563: 4562: 4560: 4558: 4554: 4550: 4545: 4541: 4537: 4534: 4531: 4527: 4526:Seeds picture 4524: 4522: 4519: 4517: 4516: 4511: 4509: 4508:Verbascum.org 4506: 4505: 4495: 4493:0-444-50356-0 4489: 4485: 4480: 4479: 4466: 4462: 4458: 4454: 4450: 4446: 4442: 4438: 4437: 4429: 4423: 4419: 4414: 4406: 4404:0-394-50432-1 4400: 4396: 4392: 4386: 4384: 4382: 4367:on 2006-05-11 4366: 4362: 4358: 4352: 4344: 4340: 4336: 4332: 4328: 4324: 4317: 4309: 4305: 4301: 4297: 4294:(1): 97–100. 4293: 4289: 4282: 4274: 4270: 4266: 4262: 4258: 4254: 4253: 4245: 4230: 4226: 4224:0-7876-7427-3 4220: 4216: 4212: 4205: 4203: 4201: 4199: 4197: 4188: 4186:1-888123-00-1 4182: 4178: 4173: 4172: 4166: 4159: 4157: 4141: 4135: 4120: 4116: 4112: 4106: 4098: 4094: 4090: 4086: 4082: 4078: 4074: 4070: 4063: 4048: 4044: 4042: 4033: 4019:on 2012-04-25 4018: 4014: 4010: 4006: 3999: 3991: 3987: 3983: 3979: 3975: 3971: 3967: 3963: 3956: 3938: 3934: 3930: 3926: 3922: 3915: 3908: 3906: 3904: 3902: 3886: 3882: 3875: 3860: 3856: 3852: 3846: 3831: 3827: 3823: 3817: 3810: 3809:0 521 20254 X 3806: 3802: 3796: 3788: 3781: 3774: 3772: 3764: 3757: 3755: 3749: 3743: 3736: 3734: 3726: 3718: 3714: 3710: 3706: 3699: 3692: 3684: 3680: 3673: 3666: 3658: 3654: 3650: 3646: 3642: 3638: 3634: 3630: 3623: 3615: 3611: 3607: 3603: 3599: 3595: 3591: 3587: 3583: 3576: 3568: 3564: 3560: 3556: 3552: 3548: 3544: 3540: 3536: 3532: 3528: 3521: 3510: 3506: 3502: 3498: 3494: 3490: 3486: 3482: 3478: 3471: 3469: 3460: 3458: 3456: 3447: 3443: 3439: 3435: 3431: 3427: 3423: 3419: 3411: 3396:on 2008-05-11 3395: 3391: 3387: 3385: 3377: 3362: 3358: 3355: 3351: 3347: 3341: 3326: 3322: 3318: 3314: 3307: 3293: 3286: 3280: 3265: 3261: 3259: 3251: 3237:on 2009-07-15 3236: 3232: 3228: 3226: 3218: 3204:on 2009-07-15 3203: 3199: 3195: 3193: 3185: 3170: 3166: 3162: 3161: 3156: 3154: 3146: 3144: 3136: 3135: 3128: 3121: 3116: 3109: 3104: 3096: 3089: 3081: 3077: 3071: 3064: 3059: 3051: 3049:0-89054-171-X 3045: 3041: 3034: 3026: 3024:0-486-22644-1 3020: 3016: 3009: 2994: 2987: 2981: 2966: 2962: 2956: 2948: 2944: 2940: 2933: 2931: 2929: 2920: 2918:0-394-50432-1 2914: 2910: 2903: 2896: 2891: 2889: 2874: 2872:0-486-22799-5 2868: 2864: 2860: 2853: 2851: 2849: 2847: 2845: 2843: 2841: 2839: 2837: 2835: 2820:on 2007-09-28 2819: 2815: 2811: 2809: 2801: 2786: 2782: 2779: 2777: 2769: 2754: 2750: 2748: 2744: 2736: 2728: 2726:3-494-01413-2 2722: 2718: 2711: 2709: 2707: 2692: 2688: 2684: 2680: 2678: 2669: 2661: 2659:0-88192-204-8 2655: 2651: 2644: 2642: 2630: 2626: 2620: 2616: 2615: 2614:Flora Iberica 2607: 2600: 2598: 2589: 2585: 2581: 2577: 2570: 2562: 2558: 2554: 2550: 2546: 2539: 2531: 2527: 2523: 2519: 2515: 2511: 2504: 2489: 2487: 2479: 2472: 2471:0-662-10668-7 2468: 2464: 2457: 2452: 2448: 2444: 2440: 2438: 2434: 2425: 2423: 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Macmillan. 2044: 2038: 2024:on 2012-06-06 2023: 2019: 2015: 2013: 2007: 2001: 1999: 1997: 1995: 1993: 1978: 1974: 1971: 1964: 1962: 1947:. eFloras.org 1946: 1942: 1936: 1934: 1932: 1930: 1922: 1921:0-340-40170-2 1918: 1914: 1908: 1906: 1904: 1902: 1900: 1891: 1889:0-521-08489-X 1885: 1881: 1880: 1872: 1870: 1868: 1866: 1864: 1847: 1841: 1839: 1837: 1820: 1818: 1810: 1808: 1806: 1804: 1802: 1797: 1783: 1779: 1773: 1764: 1755: 1748: 1744: 1738: 1731: 1727: 1721: 1717: 1709: 1707: 1706: 1700: 1698: 1694: 1690: 1686: 1682: 1678: 1674: 1670: 1666: 1662: 1661: 1656: 1652: 1647: 1638: 1636: 1631: 1629: 1625: 1621: 1617: 1613: 1609: 1605: 1601: 1597: 1593: 1589: 1585: 1581: 1577: 1572: 1570: 1566: 1562: 1558: 1554: 1551:peoples. The 1550: 1546: 1545:folk medicine 1542: 1538: 1534: 1526: 1521: 1512: 1510: 1506: 1502: 1499: 1495: 1491: 1487: 1483: 1479: 1475: 1471: 1467: 1453: 1451: 1447: 1443: 1439: 1435: 1431: 1427: 1423: 1419: 1415: 1411: 1407: 1399: 1394: 1390: 1388: 1384: 1380: 1376: 1372: 1368: 1364: 1360: 1356: 1355: 1350: 1346: 1342: 1338: 1335: 1331: 1330: 1325: 1324: 1319: 1318: 1313: 1309: 1305: 1304: 1299: 1297: 1292: 1288: 1283: 1281: 1280:overwintering 1277: 1276:Tetranychidae 1273: 1272:L. lineolaris 1269: 1265: 1264: 1259: 1255: 1251: 1247: 1243: 1239: 1235: 1234: 1229: 1225: 1221: 1217: 1213: 1208: 1206: 1202: 1198: 1194: 1190: 1186: 1182: 1181:Sierra Nevada 1178: 1167: 1165: 1161: 1157: 1153: 1149: 1142:Fossil record 1139: 1137: 1134:for insects ( 1133: 1129: 1125: 1121: 1117: 1113: 1112: 1107: 1103: 1101: 1096: 1092: 1082: 1078: 1076: 1072: 1068: 1064: 1060: 1056: 1051: 1047: 1044: 1040: 1036: 1028: 1024: 1015: 1013: 1009: 1003: 1001: 997: 993: 990:and southern 989: 985: 981: 977: 973: 969: 964: 962: 958: 954: 950: 946: 942: 938: 934: 930: 926: 921: 919: 915: 911: 907: 906:British Isles 903: 899: 895: 891: 882: 880: 875: 871: 868: 859: 857: 854: ×  853: 850: 846: 843: ×  842: 838: 835: ×  834: 830: 827: ×  826: 822: 818: 815: ×  814: 810: 805: 803: 799: 795: 791: 787: 783: 779: 775: 767: 764: 761: 757: 754: 751: 750:V. t. thapsus 748: 747: 746: 744: 735: 733: 732: 728: 725: 724: 721: 719: 718: 714: 711: 708: 707: 704: 703: 696: 693: ×  692: 688: 686: 685: 681: 678: 675: ×  674: 671: 670: 667: 665: 664: 660: 657: 654: ×  653: 650: 649: 646: 644: 643: 639: 636: 633: ×  632: 629: 628: 625: 623: 622: 618: 615: 612: ×  611: 608: 607: 604: 602: 601: 597: 594: 591: ×  590: 587: 586: 583: 581: 580: 576: 573: 570: ×  569: 566: 565: 561: 556: 553: 552: 549: 538: 536: 532: 528: 524: 520: 519: 514: 513: 508: 507: 502: 498: 497:Bothrospermae 494: 489: 484: 482: 478: 474: 473: 468: 464: 459: 458:type specimen 454: 452: 448: 444: 440: 436: 432: 431:Ancient Greek 428: 426: 418: 412: 408: 405: 401: 399: 398: 391: 390:Carl Linnaeus 387: 383: 373: 371: 367: 363: 359: 355: 351: 347: 343: 339: 335: 331: 322: 318: 316: 312: 311:inflorescence 308: 304: 299: 297: 293: 289: 284: 280: 276: 270: 265: 256: 254: 250: 246: 242: 237: 235: 231: 227: 223: 219: 214: 212: 208: 204: 200: 199:great mullein 196: 195: 185: 180: 178: 172: 169: 168:Binomial name 165: 161: 160: 155: 152: 151: 148: 147: 143: 140: 139: 136: 133: 130: 129: 126: 123: 120: 119: 116: 113: 110: 107: 106: 103: 100: 97: 94: 93: 90: 87: 84: 81: 80: 77: 76:Tracheophytes 74: 71: 68: 67: 64: 61: 58: 57: 52: 47: 43: 38: 34: 29: 26: 22: 19: 4556: 4540: 4514: 4483: 4440: 4434: 4428: 4413: 4394: 4369:. 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Retrieved 1816: 1781: 1777: 1772: 1763: 1754: 1746: 1742: 1737: 1730:V. giganteum 1729: 1725: 1720: 1703: 1701: 1673:evil spirits 1658: 1648: 1644: 1632: 1624:Commission E 1616:Glycyrrhizin 1595: 1573: 1530: 1524: 1469: 1464: 1433: 1429: 1425: 1418:lotus effect 1403: 1397: 1386: 1385:), also use 1382: 1374: 1352: 1348: 1344: 1336: 1327: 1323:Typhlodromus 1321: 1315: 1311: 1308:mullein moth 1301: 1294: 1290: 1286: 1284: 1271: 1261: 1257: 1245: 1231: 1209: 1193:noxious weed 1173: 1147: 1145: 1109: 1104: 1095:forest fires 1087: 1066: 1052: 1048: 1032: 1026: 1014:capacities. 1010:rather than 1004: 965: 922: 889: 888: 876: 872: 866: 865: 862:Common names 855: 851: 844: 840: 839:Franch. and 836: 832: 828: 824: 820: 816: 812: 808: 806: 801: 797: 793: 789: 785: 781: 777: 773: 771: 765: 759: 755: 749: 740: 729: 715: 709: 700: 694: 690: 682: 676: 672: 661: 655: 651: 640: 634: 630: 619: 613: 609: 598: 592: 588: 577: 571: 567: 554:Hybrid name 547: 534: 530: 529:gives sect. 522: 516: 510: 504: 500: 496: 492: 487: 485: 477:phenotypical 470: 467:type species 462: 455: 422: 411:Theophrastus 406: 393: 392:in his 1753 385: 379: 369: 365: 327: 300: 274: 273: 238: 215: 206: 202: 198: 193: 192: 191: 176: 174: 158: 157: 145: 108: 95: 82: 69: 24: 18: 5049:kew-2453334 4933:NatureServe 4835:iNaturalist 4581:Wikispecies 3174:November 6, 2606:"Verbascum" 2347:: citation" 1620:bactericide 1553:Zuni people 1541:Dioscorides 1450:tebuthiuron 1424:to occupy. 1414:surfactants 1371:stalk borer 1367:Lepidoptera 1317:Galendromus 1296:Haplothrips 1177:Californian 1059:protogynous 1043:gibberellin 937:New Zealand 925:established 910:Scandinavia 879:Aaron's rod 837:pterocaulon 656:pterocaulon 546:Hybrids of 501:Fasciculata 259:Description 89:Angiosperms 5205:Demulcents 5174:Categories 5141:WisFlora: 5044:Plant List 4648:Calflora: 4532:collection 4371:2009-11-06 4235:2006-11-30 4146:2011-03-16 4125:2009-11-06 4053:2009-11-06 4023:2011-12-21 3946:2009-11-06 3891:2009-11-06 3865:2009-11-06 3836:2009-11-06 3400:2009-11-06 3367:2009-11-06 3331:2009-11-06 3297:2009-11-06 3270:2009-11-06 3241:2009-11-06 3208:2009-11-06 3080:Etymonline 2999:2010-06-26 2971:2010-06-26 2878:2009-11-06 2824:2009-11-06 2791:2007-03-07 2759:2009-11-06 2745:L. subsp. 2696:2007-05-14 2494:2009-11-06 2394:2009-11-06 2357:2009-11-06 2325:2009-11-06 2229:2009-11-06 2180:2009-11-06 2147:2009-11-06 2095:2009-11-06 2028:2011-11-20 1982:2009-11-06 1951:2009-11-06 1852:7 February 1792:References 1681:glycosides 1641:Other uses 1612:chilblains 1608:carbuncles 1525:V. thapsus 1501:glycosides 1486:flavonoids 1470:V. thapsus 1438:glyphosate 1410:herbicides 1363:whiteflies 1339:, and the 1329:Amblyseius 1189:overgrazed 1148:V. thapsus 1106:Megachilid 1075:hoverflies 1063:pollinated 1012:adaptation 980:California 972:Amos Eaton 953:Hispaniola 941:La Réunion 890:V. thapsus 819:Chaub. (× 743:subspecies 334:calyx tube 275:V. thapsus 5180:Verbascum 4788:200021301 4775:200021301 4757:FloraBase 4658:Cal-IPC: 4528:from the 4211:"Mullein" 4165:"Mullein" 3982:0002-9122 3933:0071-0733 3927:: 79–86. 3717:0043-5643 3649:0002-9122 3559:0022-0477 2810:Linnaeus" 2776:Verbascum 2555:(1): 76. 2549:Phytotaxa 2510:Phytotaxa 2488:synonymy" 2006:Stace, C. 1846:"Mullein" 1693:piscicide 1588:frostbite 1557:poultices 1442:triclopyr 1426:G. tetrum 1422:seedlings 1287:Verbascum 1146:Seeds of 1111:Anthidium 1073:bees and 968:piscicide 957:Argentina 933:Australia 918:Himalayas 831:Fritsch, 817:semialbum 809:see table 800:but sink 635:lemaitrei 535:Verbascum 533:subsect. 531:Verbascum 499:subsect. 488:Verbascum 472:Verbascum 370:candicans 303:decurrent 292:trichomes 153:Species: 146:Verbascum 59:Kingdom: 5119:VicFlora 5108:VASCAN: 5101:29200424 5096:Tropicos 4972:NSWFlora 4938:2.155675 4876:11196336 4863:770183-1 4695:eFloraSA 4638:BioLib: 4566:Wikidata 4343:12241986 4288:Folklore 3752:Palila ( 3750:(2002). 3505:11961736 3497:16222647 3446:18289261 2947:66017920 2747:montanum 2530:86267331 1689:rotenone 1685:coumarin 1677:saponins 1584:earaches 1576:catarrhs 1509:rotenone 1482:mucilage 1474:saponins 1412:require 1298:verbasci 1250:families 1216:diseases 1201:Victoria 1197:Colorado 1166:layers. 1132:foraging 1071:halictid 976:Michigan 760:montanum 658:Franch. 616:Fritsch 593:godronii 481:synonyms 449:city of 447:Tunisian 439:Syracuse 376:Taxonomy 358:capsules 354:glabrous 346:pedicels 230:invasive 184:Linnaeus 131:Family: 125:Lamiales 115:Asterids 102:Eudicots 4920:MoBotPF 4801:3171949 4682:Ecocrop 4572:Q160095 4445:Bibcode 4422:English 4308:1260360 4273:6893476 4085:Bibcode 3990:2437088 3657:2443054 3614:2259541 3594:Bibcode 3567:2259539 3539:Bibcode 3426:Bibcode 2435:L. and 2282:3284108 1825:13 July 1697:fishing 1665:witches 1494:iridoid 1490:tannins 1359:Miridae 1282:pests. 1018:Ecology 996:Ontario 914:Siberia 856:spurium 829:kerneri 712:Chaub. 695:spurium 637:Boreau 614:kerneri 595:Boreau 574:Teyber 451:Thapsus 435:Thapsos 425:Thapsos 407:thapsus 350:anthers 342:sessile 338:corolla 315:taproot 283:rosette 253:torches 234:tilling 222:rosette 211:mullein 141:Genus: 121:Order: 63:Plantae 5057:PLANTS 4984:NZOR: 4925:287011 4749:vertha 4746:FEIS: 4739:182859 4713:578530 4631:110231 4615:APDB: 4595:AoFP: 4490:  4465:213812 4463:  4401:  4341:  4306:  4271:  4221:  4183:  4177:99–104 4113:. The 3988:  3980:  3931:  3807:  3748:et al. 3715:  3655:  3647:  3612:  3565:  3557:  3503:  3495:  3444:  3046:  3021:  2945:  2915:  2869:  2723:  2656:  2621:  2528:  2469:  2280:  2256:852625 2254:  2049:  1919:  1886:  1749:3:389. 1669:curses 1592:eczema 1580:colics 1565:rashes 1535:, no 1503:, and 1498:lignin 1377:) and 1332:, the 1326:, and 1240:) and 1160:Sussex 1128:palila 1039:starch 998:, and 992:Quebec 984:Canada 955:, and 945:Hawaii 912:, and 894:Azores 845:thapsi 677:thapsi 562:Notes 527:ICNafp 443:Sicily 402:. 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Index


Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Eudicots
Asterids
Lamiales
Scrophulariaceae
Verbascum
Binomial name
Linnaeus
mullein
biennial plant
rosette
soil seed bank
invasive
tilling
traditional medicine
approved drugs
dyes
torches

Hawaii
dicotyledonous
rosette
spike
trichomes
chromosome number

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