434:
152:
170:
133:
349:
1606:
1621:
459:, on small stalks. The female then picks these up in her mouth, which contains special pouches for storing the sperm. She then lays her eggs, and attaches them to the sides of crevices or to moss or lichen with her mouth, smearing the sperm over them as she does so. The eggs are laid in groups of eight to twelve.
429:
Symphylans have several features linking them to early insects, such as a labium (fused second maxillae), an identical number of head segments and certain features of their legs. Each pair of legs is associated with an eversible structure, called a "coxal sac", that helps the animal absorb moisture,
299:
feeders living deep in the soil, under stones, in decaying wood, and in other moist places. They are rapid runners, can move quickly through the pores between soil particles, and are typically found from the surface down to a depth of about 50 centimetres (20 in). They
477:
releasing long threads of silk. The silk plays a role in reproduction: the male deposits up to 450 spermatophores on stalks of silk. Symphylans have also been reported releasing silk as a defense and to suspend themselves in the air.
517:
have shown conflicting results, with some supporting the
Progoneata clade, others aligning symphylans with centipedes or other arthropods, although some are weakly supported. The clade is believed to be
468:
389:, the long first maxillae, and the second pair of maxillae which are fused to form the lower lip or labium of the mouth. The antennae serve as sense organs. Disc-like
642:
1158:
For reproduction, males do not copulate with the female. Rather, they deposit 150 to 450 sperm packets (spermatophores) on top of short stalks of silk on the ground.
1053:
316:
1097:"Symphylella patrickmuelleri sp. nov. (Myriapoda: Symphyla): The oldest known Symphyla and first fossil record of Scolopendrellidae from Cretaceous Burmese amber"
368:. The body is soft and generally 2 to 10 millimetres (0.08 to 0.4 in) long, divided into two body regions: head and trunk. An exceptional size is reached in
486:
The symphylan fossil record is poorly known, with only five species recorded, all placed in living genera. The oldest records of both families are found in
455:
The genital openings are located on the fourth body segment, but the animals do not copulate. Instead, the male deposits 150 to 450 packages of sperm, or
370:
2128:
1398:
Regier, Jerome C.; Wilson, Heather M.; Shultz, Jeffrey W. (2005). "Phylogenetic analysis of
Myriapoda using three nuclear protein-coding genes".
2167:
1443:"Description of the first species of Scutigerella (Symphyla, Scutigerellidae) from China, with mitogenomic and genetic divergence analysis"
420:. Although most adult symphylans have twelve leg pairs, the first pair is absent or vestigial in some species (e.g., those in the genus
400:. The first segment is large and usually provided with a pair of legs, the last segment is slender, lacks legs, and possesses a pair of
1190:
sp. nov. (Myriapoda: Symphyla): The oldest known
Symphyla and first fossil record of Scolopendrellidae from Cretaceous Burmese amber".
2102:
494:, approximately 99 million years ago. As a result, both families are thought to have diverged before the end of the Mesozoic Era.
2141:
610:
448:
on the sides of their head, and are the only arthropods with spiracle openings on the head. These are connected to a system of
1566:
1516:
1236:
1033:
986:
594:
556:
2146:
2264:
1653:
656:
408:
throughout their life. Immature individuals have six pairs of legs on hatching, but they add an additional pair at each
1254:"Checklist of the Myriapoda in Cretaceous Burmese amber and a correction of the Myriapoda identified by Zhang (2017)"
2226:
2081:
858:
2172:
396:
The trunk comprises 15β24 segments, which are protected by overlapping dorsal plates. Ten or twelve segments bear
2094:
652:
430:
and a small stylus that may be sensory in function. Similar structures are found in the most primitive insects.
846:
718:(Myriapoda, Symphyla, Scutigerellidae), a migrating symphylan from an Amazonian black-water inundation forest"
921:
2011:
1998:
1304:
Shear, William A.; Edgecombe, Gregory D. (2010). "The geological record and phylogeny of the
Myriapoda".
1012:
647:
433:
308:
setting by consuming seeds, roots, and root hairs in cultivated soil. For example, the garden symphylan,
2016:
393:, which probably sense vibrations, are attached to the base of the antennae, as they are in centipedes.
2259:
1096:
497:
Despite their common name, morphological studies commonly place symphylans as more closely related to
390:
2231:
2182:
169:
17:
1646:
899:
1353:
sp. (Myriapoda: Symphyla): Extensive gene order rearrangement and evidence in favor of
Progoneata"
788:
Eberhard, S.M. & Spate (1995). "Cave
Invertebrate Survey; toward an atlas of NSW Cave Fauna".
866:
514:
310:
2208:
1938:
1143:
891:
417:
1558:
1226:
1065:
2203:
2195:
2159:
445:
151:
2218:
811:
sp. (Symphyla: Scolopendrellidae predators of arthropods and nematodes in grassland soils".
2133:
2050:
1985:
1450:
1407:
1364:
1313:
1199:
1109:
820:
449:
386:
382:
8:
1639:
1631:
1171:
1481:
1454:
1411:
1368:
1317:
1203:
1113:
824:
1976:
1625:
1283:
1076:
1025:
950:
739:
164:
1080:
832:
734:
713:
284:, but are very small, non-venomous, and only distantly related to both centipedes and
2254:
2190:
2037:
1611:
1562:
1549:
1512:
1486:
1468:
1423:
1380:
1329:
1275:
1232:
1125:
1029:
992:
982:
942:
938:
590:
552:
422:
378:
245:
1348:
1287:
954:
743:
2042:
1585:
1535:
1476:
1458:
1415:
1372:
1321:
1265:
1207:
1117:
1021:
934:
828:
770:
729:
1211:
1121:
975:
Szucsich, Nikola; Scheller, Ulf (2011). "Symphyla". In
Minelli, Alessandro (ed.).
1990:
548:
270:
241:
2003:
1419:
1376:
2089:
2076:
1463:
416:, which usually has twelve pairs of legs. This mode of development is known as
1325:
2248:
1961:
1860:
1855:
1825:
1732:
1472:
1442:
1279:
1129:
996:
487:
474:
456:
397:
226:
80:
452:
that branch through the head and the first three segments of the body only.
1865:
1840:
1830:
1801:
1490:
1427:
1384:
1347:
Gai, Yonghua; Song, Daxiang; Sun, Hongying; Yang, Qun; Zhou, Kaiya (2008).
1333:
946:
847:
https://onlinelibrary.wiley.com/doi/epdf/10.1111/j.1440-6055.1996.tb01391.x
519:
337:
141:
1397:
976:
2115:
1970:
1845:
1759:
1710:
806:
305:
301:
132:
55:
870:
348:
2029:
1893:
1835:
1811:
1722:
1694:
1526:
U. Scheller (1961). "A review of the
Australian Symphyla (Myriapoda)".
1270:
1253:
510:
491:
329:
321:
266:
118:
100:
65:
1605:
1589:
1539:
1346:
774:
2154:
2120:
2063:
1870:
1791:
1786:
1776:
1768:
1740:
1700:
1688:
1682:
1663:
506:
498:
463:
325:
292:
285:
281:
277:
211:
201:
181:
105:
49:
1932:
2107:
2068:
2024:
1955:
1880:
1820:
1745:
1620:
589:. Philadelphia, PA: Holt-Saunders International. pp. 817β818.
502:
466:
whose secretions turn into a silk-like thread. One fossil species,
336:
has been shown to be predominantly predatory, and some species are
296:
122:
95:
90:
75:
70:
60:
31:
1898:
1888:
1850:
409:
405:
365:
361:
110:
85:
922:"Trunk segment numbers and sequential segmentation in myriapods"
404:. Symphylans have been reported as living up to four years, and
1903:
1676:
1228:
Treatise on
Zoology - Anatomy, Taxonomy, Biology. The Myriapoda
413:
401:
191:
693:
sp., on establishment of pineappes in south-east
Queensland".
688:
332:. A few species are found in trees and in caves. A species of
2055:
1576:
R. J. Tillyard (1930). "The evolution of the class Insecta".
1507:
C. A. Edwards (1990). "Symphyla". In Daniel L. Dindal (ed.).
1441:
Jin, Ya-Li; Godeiro, Nerivania Nunes; Bu, Yun (2023-05-04).
756:
689:
D. A. H. Murray & D. Smith (1983). "Effect of Symphyla,
1841:
Malacostraca (woodlice, shrimps, crayfish, lobsters, crabs)
1054:
BIOLOGICAL SCIENCE FUNDAMENTALS AND SYSTEMATICS - Volum III
790:
A Report Prepared Under NSW Heritage Assistance Program NEP
1661:
1303:
1299:
1297:
787:
757:
S. Clark & P. Greenslade (1996). "Review of Tasmanian
711:
670:
H. Boyle (1981). "Symphyla control in young plant cane".
426:), so adults in some species have only eleven leg pairs.
1866:
Branchiopoda (fairy, tadpole, clam shrimps, water fleas)
1010:
Minelli, Alessandro; Golovatch, Sergei I. (2013-01-01),
1578:
Papers and Proceedings of the Royal Society of Tasmania
1294:
1173:
Structure, composition and properties of spun products
437:
Life stages of symphylans: eggs, juvenile, and adult
1601:
807:
D. E. Walter, J. C. Moore & S. Loring (1989). "
1548:
1095:
1011:
920:
1391:
584:
2246:
1009:
974:
863:Integrated Pest Management on Peppermint-IPMP3.0
640:
1555:Synopsis and Classification of Living Organisms
159:Scollopendrellid symphylan (Scolopendrellidae)
1575:
1340:
481:
374:, which attains lengths of 12-13 mm (0.5 in).
1647:
1506:
1251:
1185:
1093:
1020:, Waltham: Academic Press, pp. 421β432,
1018:Encyclopedia of Biodiversity (Second Edition)
538:
536:
534:
288:. More than 200 species are known worldwide.
1746:Arachnida (spiders, scorpions, ticks, mites)
1440:
1252:Wesener, Thomas; Moritz, Leif (2018-12-17).
1094:Moritz, Leif; Wesener, Thomas (2018-04-01).
636:
634:
632:
630:
1546:
1525:
1218:
1654:
1640:
695:Queensland Journal of Agricultural Science
542:
531:
150:
131:
1480:
1462:
1269:
733:
627:
669:
432:
347:
1224:
1186:Moritz, Leif; Wesener, Thomas (2017). "
1169:
714:"On the natural history and ecology of
381:, a postantennal organ, three pairs of
14:
2247:
1349:"The complete mitochondrial genome of
761:Bagnall (Symphyla: Scutigerellidae)".
1937:
1936:
1846:Thecostraca (barnacles and relatives)
1635:
1511:. New York: Wiley. pp. 891β910.
1400:Molecular Phylogenetics and Evolution
1357:Molecular Phylogenetics and Evolution
1306:Arthropod Structure & Development
918:
444:Symphylans breathe through a pair of
314:can be a pest of crops. A species of
304:, but can do considerable harm in an
2183:3843e723-e6df-4c5e-9cfb-11dd9c86476d
2095:c90fd375-4c43-46ac-94c5-25503beb90ef
580:
578:
576:
574:
572:
570:
568:
981:. Leiden: Brill. pp. 445β466.
364:myriapods without eyes and without
24:
1899:Diplura (two-pronged bristletails)
1557:. New York: McGraw-Hill. pp.
1500:
1026:10.1016/b978-0-12-384719-5.00208-2
712:J. Adis & U. Scheller (1984).
25:
2276:
1861:Cephalocarida (horseshoe shrimps)
1597:
919:Fusco, Giuseppe (December 2005).
565:
1619:
1604:
939:10.1111/j.1525-142X.2005.05064.x
672:Cane Growers' Quarterly Bulletin
168:
53:
1434:
1245:
1179:
1163:
1136:
1087:
1058:
1047:
1003:
968:
912:
884:
851:
839:
800:
641:Penny Greenslade (2002-03-31).
320:has been recorded as a pest of
27:Class of many-legged arthropods
781:
750:
705:
682:
663:
653:Australian National University
603:
343:
13:
1:
1528:Australian Journal of Zoology
1212:10.1016/j.cretres.2017.11.018
1122:10.1016/j.cretres.2017.11.018
833:10.1016/S0031-4056(24)00190-2
735:10.1016/S0031-4056(23)05818-3
525:
377:The head has long, segmented
356:sp., showing their small size
1553:. In Sybil P. Parker (ed.).
1225:Minelli, Alessandro (2011).
7:
2265:Taxa named by John A. Ryder
1831:Pentastomida (tongue worms)
1741:Xiphosura (horseshoe crabs)
1420:10.1016/j.ympev.2004.09.005
1377:10.1016/j.ympev.2008.08.010
1188:Symphylella patrickmuelleri
1016:, in Levin, Simon A (ed.),
927:Evolution & Development
648:Australian Faunal Directory
482:Fossil record and evolution
469:Symphylella patrickmuelleri
302:consume decaying vegetation
10:
2281:
1464:10.3897/zookeys.1157.99686
585:Barnes, Robert D. (1982).
29:
1945:
1920:
1879:
1809:
1800:
1767:
1758:
1731:
1723:Pycnogonida (sea spiders)
1718:
1709:
1671:
1326:10.1016/j.asd.2009.11.002
611:"ITIS - Report: Symphyla"
239:
234:
165:Scientific classification
163:
158:
149:
139:
130:
41:
1894:Collembola (springtails)
1821:Ostracoda (seed shrimps)
1170:Schulze, L. (Jan 1975),
1148:Encyclopedia of Arkansas
900:Encyclopedia of Arkansas
30:Not to be confused with
1925:are paraphyletic groups
978:The Myriapoda. Volume 1
867:Oregon State University
545:Entomology, 3rd Edition
473:was found preserved in
439:Scutigerella immaculata
352:Several individuals of
311:Scutigerella immaculata
291:Symphyla are primarily
1836:Branchiura (fish lice)
1792:Diplopoda (millipedes)
1777:Chilopoda (centipedes)
1231:. BRILL. p. 459.
1081:10.7591/j.ctvn1tb6g.14
441:
357:
280:. Symphylans resemble
145:sp. (Scutigerellidae)
2204:Paleobiology Database
763:Invertebrate Taxonomy
462:Symphylans also have
436:
351:
2090:Fauna Europaea (new)
1666:classes by subphylum
1547:U. Scheller (1982).
845:Scheller, U. (1996)
587:Invertebrate Zoology
360:Symphyla are small,
265:, are soil-dwelling
1889:Protura (coneheads)
1455:2023ZooK.1157..145J
1412:2005MolPE..34..147R
1369:2008MolPE..49..574G
1318:2010ArtSD..39..174S
1204:2018CrRes..84..258M
1192:Cretaceous Research
1114:2018CrRes..84..258M
1102:Cretaceous Research
859:"Garden Symphylans"
825:1989Pedob..33..113W
716:Hanseniella arborea
543:C. Gillott (2005).
391:organs of TΓΆmΓΆsvΓ‘ry
1509:Soil Biology Guide
1271:10.15560/14.6.1131
442:
358:
2260:Arthropod classes
2242:
2241:
2191:Open Tree of Life
1939:Taxon identifiers
1930:
1929:
1916:
1915:
1912:
1911:
1904:Insecta (insects)
1754:
1753:
1612:Arthropods portal
1590:10.26749/IXAJ2535
1568:978-0-07-079031-5
1540:10.1071/ZO9610140
1518:978-0-471-04551-9
1449:(1157): 145β161.
1238:978-90-04-15611-1
1073:Arthropod Anatomy
1066:"X: The Symphyla"
1035:978-0-12-384720-1
988:978-90-04-18826-6
896:Entry: Symphylans
775:10.1071/IT9960189
643:"Class: Symphyla"
596:978-0-03-056747-6
558:978-1-4020-3182-3
515:Molecular studies
371:Hanseniella magna
276:in the subphylum
259:garden centipedes
252:
251:
246:Scolopendrellidae
230:
125:
16:(Redirected from
2272:
2235:
2234:
2222:
2221:
2212:
2211:
2199:
2198:
2186:
2185:
2176:
2175:
2163:
2162:
2150:
2149:
2137:
2136:
2124:
2123:
2111:
2110:
2098:
2097:
2085:
2084:
2072:
2071:
2059:
2058:
2046:
2045:
2033:
2032:
2020:
2019:
2007:
2006:
1994:
1993:
1981:
1980:
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1933:
1807:
1806:
1765:
1764:
1716:
1715:
1656:
1649:
1642:
1633:
1632:
1624:Data related to
1623:
1614:
1609:
1608:
1593:
1572:
1552:
1543:
1522:
1495:
1494:
1484:
1466:
1438:
1432:
1431:
1395:
1389:
1388:
1344:
1338:
1337:
1312:(2β3): 174β190.
1301:
1292:
1291:
1273:
1264:(6): 1131β1140.
1249:
1243:
1242:
1222:
1216:
1215:
1183:
1177:
1176:
1167:
1161:
1160:
1155:
1154:
1140:
1134:
1133:
1099:
1091:
1085:
1084:
1070:
1062:
1056:
1051:
1045:
1044:
1043:
1042:
1015:
1007:
1001:
1000:
972:
966:
965:
963:
961:
924:
916:
910:
909:
907:
906:
888:
882:
881:
879:
878:
869:. Archived from
855:
849:
843:
837:
836:
804:
798:
797:
785:
779:
778:
754:
748:
747:
737:
709:
703:
702:
686:
680:
679:
667:
661:
660:
655:. Archived from
638:
625:
624:
622:
621:
607:
601:
600:
582:
563:
562:
540:
490:from the middle
418:hemianamorphosis
412:until the adult
263:pseudocentipedes
257:, also known as
225:
173:
172:
154:
135:
116:
115:
52:
45:Temporal range:
39:
38:
21:
2280:
2279:
2275:
2274:
2273:
2271:
2270:
2269:
2245:
2244:
2243:
2238:
2230:
2225:
2217:
2215:
2207:
2202:
2194:
2189:
2181:
2179:
2171:
2166:
2158:
2153:
2145:
2140:
2132:
2127:
2119:
2114:
2106:
2101:
2093:
2088:
2080:
2075:
2067:
2062:
2054:
2049:
2041:
2036:
2028:
2023:
2015:
2010:
2002:
1997:
1989:
1984:
1975:
1974:
1969:
1960:
1959:
1954:
1941:
1931:
1926:
1908:
1875:
1796:
1750:
1727:
1705:
1667:
1660:
1610:
1603:
1600:
1569:
1519:
1503:
1501:Further reading
1498:
1439:
1435:
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1341:
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1239:
1223:
1219:
1184:
1180:
1168:
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1152:
1150:
1142:
1141:
1137:
1092:
1088:
1068:
1064:
1063:
1059:
1052:
1048:
1040:
1038:
1036:
1008:
1004:
989:
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969:
959:
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904:
902:
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885:
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755:
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710:
706:
687:
683:
668:
664:
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628:
619:
617:
609:
608:
604:
597:
583:
566:
559:
549:Springer Verlag
541:
532:
528:
509:, in the clade
484:
346:
244:
242:Scutigerellidae
224:
167:
126:
114:
113:
108:
103:
98:
93:
88:
83:
78:
73:
68:
63:
58:
47:
46:
43:
35:
28:
23:
22:
15:
12:
11:
5:
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2240:
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2223:
2213:
2200:
2187:
2177:
2164:
2151:
2138:
2125:
2112:
2099:
2086:
2077:Fauna Europaea
2073:
2060:
2047:
2034:
2021:
2008:
1995:
1982:
1967:
1951:
1949:
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1942:
1928:
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1598:External links
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1534:(1): 140β171.
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659:on 2015-09-24.
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1802:Pancrustacea
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1151:. Retrieved
1147:
1144:"Symphylans"
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1017:
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930:
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903:. Retrieved
895:
892:"Symphylans"
886:
875:. Retrieved
871:the original
862:
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813:Pedobiologia
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646:
618:. Retrieved
615:www.itis.gov
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520:monophyletic
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354:Scutigerella
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338:saprophagous
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306:agricultural
290:
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258:
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253:
240:
221:
142:Scutigerella
140:
36:
2116:iNaturalist
1971:Wikispecies
1760:Mandibulata
1711:Chelicerata
1695:Protostomia
1693:(unranked)
1687:(unranked)
1681:Subkingdom
1351:Symphylella
1198:: 258β263.
1108:: 258β263.
1013:"Myriapods"
809:Symphylella
759:Hanseniella
691:Hanseniella
423:Symphylella
344:Description
334:Symphylella
317:Hanseniella
208:Subphylum:
2249:Categories
1664:Arthropoda
1550:"Symphyla"
1258:Check List
1153:2023-05-02
1041:2022-02-28
905:2021-05-29
877:2007-07-02
701:: 121β123.
678:: 115β116.
620:2024-02-28
526:References
511:Progoneata
507:centipedes
499:millipedes
492:Cretaceous
464:spinnerets
383:mouthparts
330:Queensland
326:pineapples
322:sugar cane
293:herbivores
286:millipedes
282:centipedes
267:arthropods
255:Symphylans
202:Arthropoda
119:Cretaceous
48:99β0
2155:MilliBase
1871:Remipedia
1812:Crustacea
1787:Pauropoda
1769:Myriapoda
1701:Ecdysozoa
1689:Bilateria
1683:Eumetazoa
1473:1313-2970
1280:1809-127X
1130:0195-6671
997:812207443
960:25 August
728:: 35β41.
505:than the
503:pauropods
446:spiracles
387:mandibles
278:Myriapoda
235:Families
212:Myriapoda
188:Kingdom:
182:Eukaryota
18:Symphylan
2255:Symphyla
2025:BugGuide
2004:Symphyla
1991:Symphyla
1977:Symphyla
1956:Wikidata
1947:Symphyla
1881:Hexapoda
1851:Copepoda
1782:Symphyla
1677:Animalia
1675:Kingdom
1626:Symphyla
1584:: 1β89.
1491:37215163
1482:10193434
1428:15579388
1385:18782622
1334:19944188
1288:92289899
955:21401688
947:16336414
744:89444581
450:tracheae
379:antennae
297:detritus
274:Symphyla
222:Symphyla
198:Phylum:
192:Animalia
178:Domain:
42:Symphyla
32:Symphyta
2196:1054098
1962:Q244147
1662:Extant
1559:688β689
1451:Bibcode
1447:ZooKeys
1408:Bibcode
1365:Bibcode
1314:Bibcode
1200:Bibcode
1110:Bibcode
821:Bibcode
366:pigment
362:cryptic
269:of the
218:Class:
123:Present
2232:394515
2219:582676
2216:uBio:
2209:294419
2180:NZOR:
2160:394515
2147:154408
2069:1SYMPC
1923:italic
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796:: 765.
742:
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414:instar
229:, 1880
2227:WoRMS
2173:61986
2129:IRMNG
2121:84638
2082:11622
2043:6226W
2030:16496
2017:80390
1284:S2CID
1077:JSTOR
1069:(PDF)
951:S2CID
740:S2CID
410:moult
406:moult
402:cerci
271:class
227:Ryder
2168:NCBI
2142:ITIS
2134:1461
2103:GBIF
2064:EPPO
2056:6560
2012:BOLD
1582:1930
1563:ISBN
1513:ISBN
1487:PMID
1469:ISSN
1424:PMID
1381:PMID
1330:PMID
1276:ISSN
1233:ISBN
1126:ISSN
1030:ISBN
993:OCLC
983:ISBN
962:2020
943:PMID
591:ISBN
553:ISBN
501:and
398:legs
324:and
295:and
117:Mid
56:Preκ
2108:849
2051:EoL
2038:CoL
1999:AFD
1986:ADW
1586:doi
1536:doi
1477:PMC
1459:doi
1416:doi
1373:doi
1322:doi
1266:doi
1208:doi
1118:doi
1022:doi
935:doi
829:doi
771:doi
730:doi
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