745:
1601:, China, near the borders of Russia and North Korea. He suffered compound fractures but managed to survive. When he sought medical attention, his story raised suspicions as Siberian tigers seldom attack humans. An investigation of the attack scene revealed that raw venison carried by the man was left untouched by the tiger. Officials suspected the man to be a poacher who provoked the attack. The following morning, tiger sightings were reported by locals along the same road, and a local TV station did an on-site coverage. The group found tiger tracks and blood spoor in the snow at the attack scene and followed them for approximately 2,500 meters, hoping to catch a glimpse of the animal. Soon, the tiger was seen ambling slowly ahead of them. As the team tried to get closer for a better camera view, the tiger suddenly turned and charged, causing the four to flee in panic. About an hour after that encounter, the tiger attacked and killed a 26-year-old woman on the same road. Authorities retrieved the body with the help of a bulldozer. By then, the tiger was found lying 20 meters away, weak and barely alive. It was successfully tranquilized and taken for examination, which revealed that the tiger was anemic and gravely injured by a poacher's snare around its neck, with the steel wire cutting deeply down to the vertebrae, severing both trachea and esophagus. Despite extensive surgery by a team of veterinarians, the tiger died of wound infection. Subsequent investigation revealed that the first victim was a poacher who set multiple snares that caught both the tiger and a deer. The man was later charged for poaching and harming
1187:, where until the beginning of the 20th century, very few wolves were sighted. Wolf numbers may have increased in the region after tigers were largely eliminated during the Russian colonisation in the late 19th century and early 20th century. This is corroborated by native inhabitants of the region claiming that they had no memory of wolves inhabiting Sikhote-Alin until the 1930s, when tiger numbers decreased. Today, wolves are considered scarce in tiger habitat, being found in scattered pockets, and usually seen travelling as loners or in small groups. First hand accounts on interactions between the two species indicate that tigers occasionally chase wolves from their kills, while wolves will scavenge from tiger kills. Tigers are not known to prey on wolves, though there are four records of tigers killing wolves without consuming them. Tigers recently released are also said to hunt wolves.
955:
1096:
759:
204:
712:
1221:
1322:, illegal deforestation and bribery of park rangers facilitated poaching of Siberian tigers. Local hunters had access to a formerly sealed off lucrative Chinese market, and this once again put the region's tiger population at risk of extinction. While improvement in the local economy has led to greater resources being invested in conservation efforts, an increase in economic activity has led to an increased rate of development and deforestation. The major obstacle in preserving the tiger is the enormous territory individual tigers require; up to 450 km (170 sq mi) is needed by a single female and more for a single male.
1562:. Numerous cases of attacks on humans were recorded in the 19th century, occurring usually in central Asia excluding Turkmenistan, Kazakhstan and the Far East. Tigers were historically rarely considered dangerous unless provoked, though in the lower reaches of the Syr-Darya, a tiger reportedly killed a woman collecting firewood and an unarmed military officer whilst passing through reed thickets. Attacks on shepherds were recorded in the lower reaches of Ili. In the Far East, during the middle and late 19th century, attacks on people were recorded. In 1867 on the Tsymukha River, tigers killed 21 men and injured 6 others. In China's
1524:, and was meant to build a Siberian tiger gene pool to ensure the genetic diversity of the tiger. The Park and its existing tiger population would be further divided into two parts, one as the protective species for genetic management and the other as the ornamental species. It was discovered that when the Heilongjiang Northeast Tiger Forest Park was founded it had only 8 tigers, but according to the current breeding rate of tigers at the park, the worldwide number of wild Siberian tigers will break through 1,000 in late 2010. South Korea expected to receive three tigers pledged for donation in 2009 by Russia in 2011.
1649:
1088:
622:
presence of authors. Their comparison with historical data indicates that up to the first half of the 20th century both male and female
Siberian tigers were on average heavier than post-1970 ones. The average historical wild male Siberian tiger weighed 215.3 kg (475 lb) and the female 137.5 kg (303 lb); the contemporary wild male Siberian tiger weighs 176.4 kg (389 lb) on average with an asymptotic limit being 222.3 kg (490 lb); a wild female weighs 117.9 kg (260 lb) on average. Historical Siberian tigers and
1386:
731:
outlines. The summer fur on the back is 15–17 mm (0.59–0.67 in) long, 30–50 mm (1.2–2.0 in) along the top of the neck, 25–35 mm (0.98–1.38 in) on the abdomen, and 14–16 mm (0.55–0.63 in) on the tail. The winter fur on the back is 40–50 mm (1.6–2.0 in), 70–110 mm (2.8–4.3 in) on the top of the neck, 70–95 mm (2.8–3.7 in) on the throat, 60–100 mm (2.4–3.9 in) on the chest and 65–105 mm (2.6–4.1 in) on the abdomen. The whiskers are 90–115 mm (3.5–4.5 in).
329:
479:
38:
1502:
687:, are very high and strong in old males, and often much more massive than usually observed in the biggest skulls of Bengal tigers. The size variation in skulls of Siberian tigers ranges from 331 to 383 mm (13.0 to 15.1 in) in nine individuals measured. A female skull is always smaller and never as heavily built and robust as that of a male. The height of the sagittal crest in its middle part reaches as much as 27 mm (1.1 in), and in its posterior part up to 46 mm (1.8 in).
278:, but partial surveys conducted after 2005 indicate that the Russian tiger population was declining. An initial census held in 2015 indicated that the Siberian tiger population had increased to 480–540 individuals in the Russian Far East, including 100 cubs. This was followed up by a more detailed census which revealed there was a total population of 562 wild Siberian tigers in Russia. As of 2014, about 35 individuals were estimated to range in the international border area between Russia and China.
59:
909:
590:
1183:) numbers, either to the point of localized extinction or to such low numbers as to make them a functionally insignificant component of the ecosystem. Wolves appear capable of escaping competitive exclusion from tigers only when human pressure decreases tiger numbers. In areas where wolves and tigers share ranges, the two species typically display a great deal of dietary overlap, resulting in intense competition. Wolf and tiger interactions are well documented in
1533:
1430:
643:
tail measures 99 cm (39 in) in males and 91 cm (36 in) in females. The longest male measured 309 cm (122 in) in total length including a tail of 101 cm (40 in) and with a chest girth of 127 cm (50 in). The longest female measured 270 cm (110 in) in total length including tail of 88 cm (35 in) and with a chest girth of 108 cm (43 in). A male captured by members of the
1890:
1570:
699:
tigers, with a maximum recorded condylobasal length of 342 mm (13.5 in). The biggest skull of a
Siberian tiger from northeast China measured 406 mm (16.0 in) in length, which is about 20–30 mm (0.79–1.18 in) more than the maximum skull lengths of tigers from the Amur region and northern India, with the exception of a skull of a northern Indian tiger from the vicinity of
581:: the northern clade comprises the Siberian and Caspian tiger populations, and the southern clade all remaining continental tiger populations. A study published in 2018 was based on 32 tiger specimens using a whole-genome sequencing for analysis. Results support six monophyletic tiger clades and indicate that the most recent common ancestor lived about 110,000 years ago.
1239:
3526:
727:
populations was brighter and more uniform than that of the Far
Eastern populations. The summer coat is coarse, while the winter coat is denser, longer, softer, and silkier. The winter fur often appears quite shaggy on the trunk and is markedly longer on the head, almost covering the ears. Siberian and Caspian tigers had the thickest fur amongst tigers.
1151:
following tiger tracks to scavenge tiger kills and to potentially prey on tigers. Despite the threat of predation, some brown bears actually benefit from the presence of tigers by appropriating tiger kills that the bears may not be able to successfully hunt themselves. Brown bears generally prefer to contest the much smaller female tigers. During
744:
2499:
Luo, S.-J.; Kim, J.-H.; Johnson, W. E.; van der Walt, J.; Martenson, J.; Yuhki, N.; Miquelle, D. G.; Uphyrkina, O.; Goodrich, J. M.; Quigley, H. B.; Tilson, R.; Brady, G.; Martelli, P.; Subramaniam, V.; McDougal, C.; Hean, S.; Huang, S.-Q.; Pan, W.; Karanth, U. K.; Sunquist, M.; Smith, J. L. D. &
924:
population estimated in 1993 reached 0.3 tigers in 100 km (39 sq mi), with a sex ratio of averaging 2.4 females per male. These density values were much lower than what had been reported for other subspecies at the time. In 2004, dramatic changes in land tenure, population density, and
726:
The fur of the
Siberian tiger is moderately thick, coarse and sparse compared to that of other felids living in the former Soviet Union. Compared to the extinct westernmost populations, the Siberian tiger's summer and winter coats contrast sharply with other subspecies. Generally, the coat of western
532:
relationships of tiger subspecies and observed a remarkable similarity between
Caspian and Siberian tigers, indicating that the Siberian tiger is the genetically closest living relative of the Caspian tiger, which strongly implies a very recent common ancestry. Based on phylogeographic analysis, they
527:
and Hebrew
University of Jerusalem collected tissue samples from 20 of 23 Caspian tiger specimens kept in museums across Eurasia. They sequenced at least one segment of five mitochondrial genes and found a low amount of variability of the mitochondrial DNA in Caspian tigers as compared to other tiger
1131:
were found in several tiger scat samples. Tigers attack black bears less often than brown bears, as the latter live in more open habitats and are not able to climb trees. In the same time period, four cases of brown bears killing female tigers and young cubs were reported, both in disputes over prey
1074:
and monitored for more than 15 months in the eastern slopes of the
Sikhote-Alin mountain range. Results of this study indicate that their distribution is closely associated with distribution of Manchurian wapiti, while distribution of wild boar was not such a strong predictor for tiger distribution.
698:
in
January 1954 had a greatest skull length of 385 mm (15.2 in), which is considerably more than the known maximum for this population and slightly exceeds that of most Siberian tigers. However, its condylobasal length was only 305 mm (12.0 in), smaller than those of the Siberian
1465:
with the
Iranian government, as conservation groups of both countries agreed on reintroducing these animals into the wild within the next five years. This issue is controversial since only 30% of such releases have been successful. In addition, the Siberian tiger is not genetically identical to the
1163:
and tigers to abstract their prey. In the
Sikhote-Alin reserve, 35% of tiger kills were stolen by bears, with tigers either departing entirely or leaving part of the kill for the bear. Some studies show that bears frequently track down tigers to usurp their kills, with occasional fatal outcomes for
730:
The whiskers and hair on the back of the head and the top of the neck are also greatly elongated. The background colour of the winter coat is generally less bright and rusty compared to that of the summer coat. Because of the winter fur's greater length, the stripes appear broader with less defined
722:
The ground colour of Siberian tigers' pelage is often very pale, especially in winter coat. However, variations within populations may be considerable. Individual variation is also found in form, length, and partly in colour, of the dark stripes, which have been described as being dark brown rather
2682:
Cho, Y. S.; Hu, L.; Hou, H.; Lee, H.; Xu, J.; Kwon, S.; Oh, S.; Kim, H. M.; Jho, S.; Kim, S.; Shin, Y. A.; Kim, B. C.; Kim, H.; Kim, C. U.; Luo, S. J.; Johnson, W. E.; Koepfli, K. P.; Schmidt-Küntzel, A.; Turner, J. A.; Marker, L.; Harper, C.; Miller, S. M.; Jacobs, W.; Bertola, L. D.; Kim, T. H.;
1861:
Kitchener, A. C.; Breitenmoser-Würsten, C.; Eizirik, E.; Gentry, A.; Werdelin, L.; Wilting, A.; Yamaguchi, N.; Abramov, A. V.; Christiansen, P.; Driscoll, C.; Duckworth, J. W.; Johnson, W.; Luo, S.-J.; Meijaard, E.; O’Donoghue, P.; Sanderson, J.; Seymour, K.; Bruford, M.; Groves, C.; Hoffmann, M.;
642:
in the Sikhote-Alin range from 178 to 208 cm (70 to 82 in) in head and body length measured in straight line, with an average of 195 cm (77 in) for males; and for females ranging from 167 to 182 cm (66 to 72 in) with an average of 174 cm (69 in). The average
553:
structure analyses clearly identified two populations separated by a development corridor in Russia. Despite their well-documented 20th century decline, the researchers failed to find evidence of a recent population bottleneck, although genetic signatures of a historical contraction were detected.
1635:
province. Park officials reported that the bus driver violated safety guidelines by leaving the vehicle to check on the condition of the bus. In September 2013, a tiger mauled a zookeeper to death at a zoo in western Germany after the worker forgot to lock a cage door during feeding time. In July
1512:
In recent years, captive breeding of tigers in China has accelerated to the point where the captive population of several tiger subspecies exceeds 4,000 animals. Three thousand specimens are reportedly held by 10–20 "significant" facilities, with the remainder scattered among some 200 facilities.
1453:
and forms a large wetland of about 8,000 km (3,100 sq mi). Until 1948, the delta was a refuge of the extinct Caspian tiger. Reintroduction of the Siberian tiger to the delta has been proposed. Large populations of wild boar inhabit the swamps of the delta. The reintroduction of the
1206:
bore evidence of predation by a Siberian tiger. The tiger apparently ambushed, pursued, and killed the lynx but only consumed it partially. This incident marks one of the first documented cases of a tiger preying on a lynx, and indicates that the tiger might have been more intent on eliminating a
863:
In 2005, the number of Amur tigers in China was estimated at 18–22, and 331–393 in the Russian Far East, comprising a breeding adult population of about 250, fewer than 100 likely to be sub-adults, more than 20 likely to be less than 3 years of age. More than 90% of the population occurred in the
1615:
in December 2007, a tiger escaped and killed a visitor, and injured two others. The animal was shot by the police. The zoo was widely criticized for maintaining only a 12.5 ft (3.8 m) fence around the tiger enclosure, while the international standard is 16 ft (4.9 m). The zoo
1581:
According to the Japanese Police Bureau in Korea, in 1928 a tiger killed one human, whereas leopards killed three, wild boars four and wolves killed 48. Six cases were recorded in 20th century Russia of unprovoked attacks leading to man-eating behaviour. Provoked attacks are however more common,
621:
In 2005, a group of Russian, American and Indian zoologists published an analysis of historical and contemporary data on body weights of wild and captive tigers, both female and male across all subspecies. The data used include weights of tigers that were older than 35 months and measured in the
1425:
Delta was suggested as a potential site for such a project. A feasibility study was initiated to investigate if the area is suitable and if such an initiative would receive support from relevant decision makers. A viable tiger population of about 100 animals would require at least 5,000 km
1353:
was founded, with the aim of providing a comprehensive picture of the ecology of the Amur tiger and the role of tigers in the Russian Far East through scientific studies. By capturing and outfitting tigers with radio collars, their social structure, land use patterns, food habits, reproduction,
1272:
and are left alone when the female leaves to hunt for food. Cubs are divided equally between sexes at birth. However, by adulthood there are usually two to four females for every male. The female cubs remain with their mothers longer, and later they establish territories close to their original
1078:
Results of a three-year study on Siberian tigers indicate that the mean interval between their kills and estimated prey consumption varied across seasons: during 2009 to 2012, three adult tigers killed prey every 7.4 days in summer and consumed a daily average of 7.89 kg (17.4 lb); in
1426:(1,900 sq mi) of large tracts of contiguous habitat with rich prey populations. Such habitat is not presently available in the delta and so cannot be provided in the short term. The proposed region is therefore unsuitable for the reintroduction, at least at this stage of development.
934:
with their daughters once the daughters reached maturity. By 2007, population density of tigers was estimated at 0.8±0.4 tigers in 100 km (39 sq mi) in the southern part of Sikhote-Alin Zapovednik, and 0.6±0.3 tigers in 100 km (39 sq mi) in the central part of the
1242:
1150:
The effect the presence of tigers has on brown bear behavior seems to vary. In the winters of 1970–1973, Yudakov and Nikolaev recorded two cases of bears showing no fear of tigers and another case of a brown bear changing path upon crossing tiger tracks. Other researchers have observed bears
662:
in 1943, reportedly measured 350 cm (140 in) "over the curves", with a tail length of about 1 m (3 ft 3 in). It weighed about 300 kg (660 lb). Dubious sources mention weights of 318 and 384 kg (701 and 847 lb) and even 408 kg (899 lb).
1241:
1246:
1245:
1240:
3288:
Qi, J.; Gu, J.; Ning, Y.; Miquelle, D. G.; Holyoak, M.; Wen, D.; Liang, X.; Liu, S.; Roberts, N.; Yang, E.; Lang, J.; Wang, F.; Li, C.; Liang, Z.; Liu, P.; Ren, Y.; Zhou, S.; Zhang, M.; Ma, J.; Chang, J. & Jiang, G. (2021). "Integrated assessments call for establishing a sustainable
1075:
Although they prey on both Siberian roe deer and sika deer, overlap of these ungulates with tigers was low. Distribution of moose was poorly associated with tiger distribution. The distribution of preferred habitat of key prey species was an accurate predictor of tiger distribution.
1247:
431:. In 2015, morphological, ecological and molecular traits of all putative tiger subspecies were analysed in a combined approach. Results support distinction of the two evolutionary groups: continental and Sunda tigers. The authors proposed recognition of only two subspecies: namely
1299:. Further exacerbating the problem is that more than 90% of the population occurred in the Sikhote Alin mountain region. Tigers rarely move across the development corridor, which separates this sub-population from the much smaller sub-population in southwest Primorye province.
1366:, which responds to all tiger-human conflicts; by continuing to enhance the large database on tiger ecology and conservation with the goal of creating a comprehensive Siberian tiger conservation plan; and training the next generation of Russian conservation biologists.
400:
of several tiger subspecies was questioned in 1999. Most putative subspecies described in the 19th and 20th centuries were distinguished on the basis of fur length and colouration, striping patterns and body size – characteristics that vary widely within populations.
1466:
Caspian tiger. Another difference is the climatic, with temperatures higher in Iran than in Siberia. Introducing exotic species into a new habitat could inflict irreversible and unknown damage. In December 2010, one of the tigers exchanged died in Eram Zoo in
1244:
617:
In the 1980s, the typical weight range of wild Siberian tigers was indicated as 180 to 306 kg (397 to 675 lb) for males and 100 to 167 kg (220 to 368 lb) for females. Exceptionally large individuals were targeted and shot by hunters.
929:
Siberian Tiger Project were detected, suggesting that when tigers are well protected from human-induced mortality for long periods, the adult female population density increases significantly. When more adult females survived, the mothers shared their
1190:
This competitive exclusion of wolves by tigers has been used by Russian conservationists to convince hunters in the Far East to tolerate the big cats, as they limit ungulate populations less than wolves, and are effective in controlling wolf numbers.
626:
were the largest ones, whereas contemporary Siberian tigers are on average lighter than Bengal tigers. The reduction of the body weight of today's Siberian tigers may be explained by concurrent causes, namely the reduced abundance of prey because of
2299:
Fauna Japonica sive Descriptio animalium, quae in itinere per Japoniam, jussu et auspiciis superiorum, qui summum in India Batava imperium tenent, suscepto, annis 1825–1830 collegit, notis, observationibus et adumbrationibus illustravit Ph. Fr. de
1147:. Predation by tigers on denned brown bears was not detected during a study carried between 1993 and 2002. Ussuri brown bears, along with the smaller black bears constitute 2.1% of the Siberian tiger's annual diet, of which 1.4% are brown bears.
1306:. Poaching of tigers and their wild prey species is considered to be driving the decline, although heavy snows in the winter of 2009 could have biased the data. In northern China’s Huang Ni He National Nature Reserve, poachers set up foremost
273:
in the Russian Far East. In 2005, there were 331–393 adult and subadult Siberian tigers in this region, with a breeding adult population of about 250 individuals. The population had been stable for more than a decade because of intensive
2851:
1369:
In August 2010, China and Russia agreed to enhance conservation and cooperation in protected areas in a transboundary area for Amur tigers. China has undertaken a series of public awareness campaigns including celebration of the first
875:
located in the vicinity of the international borders with Russia and North Korea. Camera-trap surveys carried out in the spring seasons of 2013 and 2014 revealed between 27 and 34 tigers along the China-Russian border. In April 2014,
1420:
Inspired by findings that the Amur tiger is the closest relative of the Caspian tiger, there has been discussion whether the Amur tiger could be an appropriate subspecies for reintroduction into a safe place in Central Asia. The
1354:
mortality patterns and their relation with other inhabitants of the ecosystem, including humans is studied. These data compilations will hopefully contribute toward minimizing poaching threats because of traditional hunting. The
1171:
is not specifically studied. Numerous publications on these species there are mainly episodic and survey data on this issue are collected by different authors in selected areas which do not give a complete picture of the nature.
2042:
Xiao, W.; Feng, L.; Mou, P.; Miquelle, D. G.; Hebblewhite, M.; Goldberg, J. F.; Robinson, H. S.; Zhao, X.; Zhou, B.; Wang, T. & Ge, J. (2016). "Estimating abundance and density of Amur tigers along the Sino–Russian border".
544:
Samples of 95 wild Amur tigers were collected throughout their native range to investigate questions relative to population genetic structure and demographic history. Additionally, targeted individuals from the North American
3166:
Wang, T.; Feng, L.; Mou, P.; Wu, J.; Smith, J. L.; Xiao, W.; Yang, H.; Dou, H.; Zhao, X.; Cheng, Y. & Zhou, B. (2016). "Amur tigers and leopards returning to China: direct evidence and a landscape conservation plan".
502:
makeup of the Siberian tiger and its relationship to other populations. One of the most important outcomes has been the discovery of low genetic variability in the wild population, especially when it comes to maternal or
1310:, but there is not sufficient personnel to patrol this 75 km (29 sq mi) area throughout the year. In Hunchun National Nature Reserve, poaching of ungulate species impedes recovery of the tiger population.
511:
almost completely dominates the maternal lineages of wild Siberian tigers. On the other hand, captive tigers appear to show higher mtDNA diversity. This may suggest that the subspecies has experienced a very recent
1345:
and countries with consumer markets have banned domestic trade as well. At the 14th Conference of the Parties to CITES in 2007, stronger enforcement measures were called for, as well as an end to tiger farming.
600:
The tiger is reddish-rusty, or rusty-yellow in colour, with narrow black transverse stripes. The body length is not less than 150 cm (59 in), condylobasal length of skull 250 mm (9.8 in),
4527:
3637:
1066:
was collected along the international border between Russia and China between November 2014 and April 2015; 115 scat samples of nine tigers contained foremost remains of wild boar, sika deer and roe deer.
203:
4820:
537:
less than 10,000 years ago via the Gansu−Silk Road region from eastern China, and subsequently traversed eastward to establish the Siberian tiger population in the Russian Far East. The events of the
1159:, 44 direct confrontations between bears and tigers were observed, in which bears in general were killed in 22 cases, and tigers in 12 cases. There are reports of brown bears specifically targeting
1243:
1513:
This makes China home to the second largest captive tiger population in the world, after the U.S., which in 2005 had an estimated 4,692 captive tigers. In a census conducted by the U.S.-based
4544:
774:
The Siberian tiger once inhabited much of the Korean Peninsula, Manchuria and other parts of north-eastern China, the eastern part of Siberia and the Russian Far East, perhaps as far west as
694:
had a maximum length of 297.0 to 365.8 mm (11.69 to 14.40 in), while that of females measured 195.7 to 255.5 mm (7.70 to 10.06 in). A tiger killed on the Sumbar River in
3325:
4856:
4210:
884:
dispersed westwards between 2003 and 2016. Camera trap surveys between 2013 and 2018 revealed about 55 Siberian tigers in four forested landscapes in northeastern China: Laoyeling,
566:. Overall, their results indicate the need to secure ecological connectivity between the two Russian populations to minimize loss of genetic diversity and overall susceptibility to
3372:
Yang, H.; Zhao, X.; Han, B.; Wang, T.; Mou, P.; Ge, J. & Feng, L. (2018). "Spatiotemporal patterns of Amur leopards in northeast China: Influence of tigers, prey, and humans".
2442:
Wilting, A.; Courtiol, A.; Christiansen, P.; Niedballa, J.; Scharf, A. K.; Orlando, L.; Balkenhol, N.; Hofer, H.; Kramer-Schadt, S.; Fickel, J. & Kitchener, A. C. (2015).
1164:
the tiger. A report from 1973 describes twelve known cases of brown bears killing tigers, including adult males; in all cases the tigers were subsequently eaten by the bears.
2555:
Russello, M. A.; Gladyshev, E.; Miquelle, D. & Caccone, A. (2005). "Potential genetic consequences of a recent bottleneck in the Siberian tiger of the Russian Far East".
1283:
The average lifespan for Siberian tigers ranges from 16–18 years. Wild individuals tend to live between 10–15 years, while in captivity individuals may live up to 25 years.
554:
This disparity in signal may be due to several reasons, including historical paucity in population genetic variation associated with postglacial colonisation and potential
1202:) and occasionally kill and eat them. Eurasian lynx remains have been found in the stomach contents of Siberian tigers in Russia. In March 2014, a dead lynx discovered in
4232:
2825:
Slaght, J. C.; Miquelle, D. G.; Nikolaev, I. G.; Goodrich, J. M.; Smirnov, E. N.; Traylor-Holzer, K.; Christie, S.; Arjanova, T.; Smith, J. L. D.; Karanth, K. U. (2005).
1582:
usually the result of botched attempts at capturing them. In December 1997, an injured Amur tiger attacked, killed and consumed two people. Both attacks occurred in the
1143:
When Amur tigers prey on brown bears, they usually target young and sub-adult bears, besides small female adults taken outside their dens, generally when lethargic from
293:
from Caspian tigers and living tiger populations indicate that the common ancestor of the Siberian and Caspian tigers colonized Central Asia from eastern China, via the
3649:
3496:
1897:
558:
from an extirpated Chinese population. The extent and distribution of genetic variation in captive and wild populations were similar, yet gene variants persisted
3527:"Глава 19. Взаимоотношения амурского тигра с бурым и гималайским медведями [Chapter 19. Relationship of Amur tigers with brown and Himalayan black bear]"
305:
eastward to establish the Siberian tiger population in the Russian Far East. The Caspian and Siberian tiger populations were the northernmost in mainland Asia.
2827:"Chapter 6. Who's king of the beasts? Historical and contemporary data on the body weight of wild and captive Amur tigers in comparison with other subspecies"
2593:
4907:"Mammals collected in the Maritime Province of Siberia by the Morden-Graves North Asiatic Expedition: with the description of a new hare from the Amur River"
2017:
1863:
2280:
Caroli Linnæi Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I
3989:
Gurumurthy, S.; Yu, L.; Zhang, C.; Jin, Y.; Li, W.; Zhang, X.; Fang, F. (2018). "Exploiting Data and Human Knowledge for Predicting Wildlife Poaching".
2974:
5002:
4450:
4188:
3359:
Siberian Tiger Project: Long-Term Research, Training, and Tiger-Human Conflict Mitigation in the Russian Far East. Final Report to 21st Century Tiger
3211:
4548:
4282:
2748:
Liu, Y.-C.; Sun, X.; Driscoll, C.; Miquelle, D. G.; Xu, X.; Martelli, P.; Uphyrkina, O.; Smith, J. L. D.; O’Brien, S. J. & Luo, S.-J. (2018).
541:
may have been the critical factor in the reciprocal isolation of Caspian and Siberian tigers from what was likely a single contiguous population.
2850:
Kerley, L.; Goodrich, J.; Smirnov, E.; Miquelle, D.; Nikolaev, I; Arjanova, T.; Slaght, J.; Schleyer, B.; Kuigli, H. & Hornocker, M. (2005).
900:, urine and hair was used to genetically identify 30 tigers in this region. However, only Laoyeling is thought to support a breeding population.
5010:
4800:
4250:
3670:
916:
Siberian tigers are known to travel up to 1,000 km (620 mi) over ecologically unbroken country. In 1992 and 1993, the maximum total
5300:
5216:
3959:
3571:
The Ecology of the Amur Tiger based on Long-Term Winter Observations in 1970–1973 in the Western Sector of the Central Sikhote-Alin Mountains
4864:
4205:
3872:
3759:
516:
caused by human pressure, with the founders of the captive population having been captured when genetic variability was higher in the wild.
4163:
Asian big cat conservation and trade control in selected range States: evaluating implementation and effectiveness of CITES Recommendations
3688:
Goodrich, J. M.; Kerley, L. L.; Smirnov, E. N.; Miquelle, D. G.; McDonald, L.; Quigley, H. B.; Hornocker, M. G. & McDonald, T. (2008).
1652:
671:
The skull of the Siberian tiger is characterized by its large size. The facial region is very powerful and very broad in the region of the
4413:
3333:
358:
in 1844 were tiger skins with long hairs and dense coats sold in Japan, which originated in Korea, most likely from animals killed in the
3815:
4343:
1458:, which was once an important prey, is under consideration. The Ili delta is therefore considered as a suitable site for introduction.
1264:
on trees. She will spend 5 or 6 days with the male, during which she is receptive for three days. Gestation lasts from 3 to 3½ months.
817:, which crosses into Russia from China at several places in southwest Primorye. This region represents a merger zone of the East Asian
3151:
Wang, T.M.; Yang, H.T.; Xiao, W.H.; Feng, L.M.; Mou, P. & Ge, J.P. (2014). "Camera traps reveal Amur tiger breeding in NE China".
860:
are associated with the conifer forests and are near the southern limits of their distribution in the central Sikhote-Alin mountains.
963:
3458:
Miller, C. S.; Hebblewhite, M.; Petrunenko, Y. K.; Seryodkin, I. V.; DeCesare, N. J.; Goodrich, J. M. & Miquelle, D. G. (2013).
1924:
Goodrich, J.; Wibisono, H.; Miquelle, D.; Lynam, A.J.; Sanderson, E.; Chapman, S.; Gray, T.N.E.; Chanchani, P.; Harihar, A. (2022).
5305:
5138:
4797:
4042:"Relationships between humans and ungulate prey shape Amur tiger occurrence in a core protected area along the Sino-Russian border"
3934:
3495:
Seryodkin, I. V.; Kostyria, A. V.; Goodrich, J. M.; Miquelle, D. G.; Smirnov, E. N.; Kerley, L. L. & Hornocker, M. G. (2003).
5177:
4839:
3645:
3525:
Seryodkin, I. V.; Goodrich, J. M.; Kostyrya, A. V.; Schleyer, B. O.; Smirnov, E. N.; Kerley, L. L. & Miquelle, D. G. (2005).
4817:
4508:
1273:
ranges. Males, on the other hand, travel unaccompanied and range farther earlier in their lives, making them more vulnerable to
1079:
winter they killed more large-bodied prey, made kills every 5.7 days and consumed a daily average of 10.3 kg (23 lb).
609:
tooth over 26 mm (1.0 in) long. It has an extended supple body standing on rather short legs with a fairly long tail.
5285:
4227:
4166:
2155:
1566:, tigers reportedly attacked woodsmen and coachmen, and occasionally entered cabins and dragged out both adults and children.
5208:
3855:
3798:
3566:
2954:
2919:
2809:
2423:
2391:
5182:
3087:
Miquelle, D. G.; Smirnov, E. N.; Merrill, T. W.; Myslenkov, A. E.; Quigley, H.; Hornocker, M. G. & Schleyer, B. (1999).
758:
3229:
Ning, Y.; Kostyria, A.V.; Ma, J.; Chayka, M.I.; Guskov, V.Y.; Qi, J.; Sheremetyeva, I.N.; Wang, M. & Jiang, G. (2019).
1864:"A revised taxonomy of the Felidae: The final report of the Cat Classification Task Force of the IUCN Cat Specialist Group"
954:
818:
4472:
4317:
1253:
Three orphaned Siberian tigers rescued after their mothers were killed by poachers are released back to the wild in Russia
466:
In 2017, the Cat Specialist Group revised felid taxonomy and now recognizes all the tiger populations in mainland Asia as
2087:"Mitochondrial Phylogeography Illuminates the Origin of the Extinct Caspian Tiger and Its Relationship to the Amur Tiger"
1415:
3723:"Tigers and Wolves in the Russian Far East: Competitive Exclusion, Functional Redundancy, and Conservation Implications"
5310:
3573:. Institute of Biology and Soil Science, Far-Eastern Scientific Center, Academy of Sciences of the USSR. Archived from
1668:
1631:
were attacked and killed in 2010. In January 2011, a tiger attacked and killed a tour bus driver at a breeding park in
1616:
subsequently erected a taller barrier topped by an electric fence. One of the victims admitted to taunting the animal.
5020:
1095:
825:, resulting in a mosaic of forest types that vary in elevation and topography. Key habitats of the Siberian tiger are
5090:
4994:
4940:
4574:
4016:
3903:
3742:
570:
events, and support a previous study suggesting that the captive population may be a reservoir of gene variants lost
3721:
Miquelle, D. G.; Stephens, P. A.; Smirnov, E. N.; Goodrich, J. M.; Zaumyslova, O. J. & Myslenkov, A. E. (2005).
2085:
Driscoll, C. A.; Yamaguchi, N.; Bar-Gal, G. K.; Roca, A. L.; Luo, S.; Macdonald, D. W. & O'Brien, S. J. (2009).
5290:
1605:. He served two years in prison. After being released from prison, he worked in clearing the forest of old snares.
1277:
and other tigers. A Siberian tiger family comprising an adult male, a female and three cubs were recorded in 2015.
920:
of the Sikhote-Alin tiger population was estimated at 0.62 tigers in 100 km (39 sq mi). The maximum
577:
In 2013, the whole genome of the Siberian tiger was sequenced and published. Tigers in mainland Asia fall into two
5221:
4906:
3545:
3417:"Prey selection of Amur tigers in relation to the spatiotemporal overlap with prey across the Sino–Russian border"
4207:
Siberian Tiger Project: Long-Term Research, Training, and Tiger-Human Conflict Mitigation in the Russian Far East
2557:
1690:
considered the Siberian tiger as Hu Lin, the king. Since the tiger has a mark on its foreheads that looks like a
1608:
1319:
1226:
A tiger at the Rehabilitation and Reintroduction Center for Amur (Siberian) Tigers in the village of Alekseevka,
872:
690:
Female skulls range from 279.7 to 310.2 mm (11.01 to 12.21 in). The skulls of male Caspian tigers from
550:
4126:
Jo, Y. S. & Baccus, J. T. (2016). "Are large cats compatible with modern society on the Korean Peninsula?".
4089:
Goodrich, J. M.; Miquelle, D. G.; Smirnov, E. M.; Kerley, L. L.; Quigley, H. B. & Hornocker, M. G. (2010).
1514:
1493:, Russia, provided the herbivore population has reached a size warranting the introduction of large predators.
524:
373:
347:
in 1758 for the tiger. In the 19th century, several tiger specimens were collected in East Asia and described:
4985:
4983:
Amur Leopard and Tiger Alliance (ALTA) – Conserving Amur leopards and tigers in the Russian Far East and China
3231:"Dispersal of Amur tiger from spatial distribution and genetics within the eastern Changbai mountain of China"
4447:
Taming the tiger trade: China's markets for wild and captive tiger products since the 1993 domestic trade ban
1994:
1695:
4929:"Sacred Places and Masters of Hunting Luck in the Forest Worlds of the Udege People of the Russian Far East"
4445:
1637:
4977:
4183:
3037:
Cooper, D. M.; Dugmore, A. J.; Gittings, B. M.; Scharf, A. K.; Wilting, A. & Kitchener, A. C. (2016).
1268:
size is normally two or four cubs but there can be as many as six. The cubs are born blind in a sheltered
5203:
5164:
4524:
3690:"Survival rates and causes of mortality of Amur tigers on and near the Sikhote-Alin Biosphere Zapovednik"
2319:
Sitzungsberichte der Kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Classe
1812:
1156:
926:
632:
3550:(PhD) (in Russian). Vladivostok, Russia: Far Eastern National University. pp. 1–252. Archived from
871:
In August 2012, a Siberian tiger with four cubs was recorded for the first time in northeastern China's
844:, and wild boar being the most common throughout the Sikhote-Alin mountains but rare in higher altitude
711:
549:
population were sampled to assess the genetic representation found in captivity. Population genetic and
5280:
5035:
4289:
1521:
1280:
At 35 months of age, tigers are sub-adults. Males reach sexual maturity at the age of 48 to 60 months.
1220:
1107:
Following a decrease of ungulate populations from 1944 to 1959, 32 cases of Amur tigers attacking both
877:
5275:
2623:
Henry, P.; Miquelle, D.; Sugimoto, T.; McCullough, D. R.; Caccone, A. & Russello, M. A. (2009). "
2226:
1972:
1891:
A survey of tigers and leopards and prey resources in the Paektusan area, North Korea, in winter 1998
1819:
1805:
1326:
58:
46:
4762:
4257:
3112:
in the Russian Far East: the role of protected areas and landscape matrix in population persistence"
2295:"Aperçu général et spécifique sur les Mammifères qui habitent le Japon et les Iles qui en dépendent"
1948:
5295:
4887:
4505:
3615:
2237:]. Washington DC: Smithsonian Institution and the National Science Foundation. pp. 95–202.
1325:
The Siberian tiger was once common in the Korean Peninsula. It was eradicated during the period of
814:
3967:
2870:
2271:
1520:
In 1986, the Chinese government established the world's largest Siberian tiger breeding base, the
946:), but in the Changbai Mountains have been recorded more often in lower elevations than leopards.
5015:
3767:
3614:
Seryodkin, I. V.; Goodrich, J. M.; Kostyria, A. V.; Smirnov, E. N. & Miquelle, D. G. (2011).
832:
The faunal complex of the region is represented by a mixture of Asian and boreal life forms. The
355:
5104:
3991:
Compass '18. Proceedings of the 1st ACM SIGCAS Conference on Computing and Sustainable Societies
3596:
5052:
4420:
4040:
Xiao, W.; Hebblewhite, M.; Robinson, H.; Feng, L.; Zhou, B.; Mou, P.; Wang, T.; Ge, J. (2018).
1736:
1648:
1446:
880:
personnel captured a video of a tigress with cubs in inland China. The tiger population in the
857:
801:
Today, its range stretches south to north for almost 1,000 km (620 mi) the length of
245:
181:
4928:
3891:
3722:
2337:
2314:
1678:
considered the tiger a near-deity and often referred to it as "Grandfather" or "Old man". The
864:
Sikhote Alin mountain region. An unknown number of tigers survive in the reserve areas around
4569:
Vaillant, J. (2010) The Tiger: A True Story of Vengeance and Survival. Knopf Canada, Toronto
4251:
Pre-Feasibility Study on the Possible Restoration of the Caspian Tiger in the Amu Darya Delta
3642:Биоразнообразие и роль животных в экосистемах: Материалы IV Международной научной конференции
3524:
3494:
2408:
2379:
1860:
1660:
1506:
1411:
1203:
765:
538:
513:
424:
275:
4489:
4347:
3613:
2294:
5247:
5195:
5125:
5099:
3381:
3298:
3242:
3176:
3123:
3086:
3054:
3038:
2900:
2761:
2750:"Genome-wide evolutionary analysis of natural history and adaptation in the world's tigers"
2696:
2640:
2455:
2444:"Planning tiger recovery: Understanding intraspecific variation for effective conservation"
2098:
1721:
The Siberian tiger is used in heraldic symbols throughout the area where it is indigenous.
1087:
520:
402:
397:
2824:
1470:. Nevertheless, the project has its defenders, and Iran has successfully reintroduced the
8:
5028:
3687:
3616:"Intraspecific relationships between brown bears, Asiatic black bears and the Amur tiger"
2885:
2598:
1838:
1261:
885:
3547:
The ecology, behavior, management and conservation status of brown bears in Sikhote-Alin
3385:
3302:
3246:
3180:
3127:
3058:
2949:] (Nachdruck der 3. Auflage 2004 ed.). Hohenwarsleben: Westarp Wissenschaften.
2765:
2700:
2644:
2459:
2102:
1358:
has been productive in increasing local capacity to address human-tiger conflict with a
1291:
Results of genetic analysis of 95 wild Siberian tiger samples from Russia revealed that
4718:
4674:
4630:
4143:
4066:
4041:
4022:
3994:
3428:
3397:
3265:
3230:
3192:
3036:
3002:
2725:
2684:
2664:
2574:
2476:
2443:
2180:
2121:
2086:
1833:
1768:
1663:
in the 1830s. The name 'Amur tiger' was used in 1933 for Siberian tigers killed by the
1602:
1274:
1031:
983:
917:
881:
853:
845:
654:
The Siberian tiger is often considered to be the largest tiger. A wild male, killed in
482:
Phylogenetic relationship of tiger populations Note the close relationship between the
419:
53:
4836:
3921:
The Way of the Tiger : natural history and conservation of the endangered big cat
2938:
2532:
2501:
2151:
1925:
1132:
and in self-defense. Tigers mainly feed on the bear's fat deposits, such as the back,
5242:
5112:
4936:
4570:
4394:
4375:
4071:
4012:
3899:
3851:
3794:
3738:
3706:
3689:
3464:) kill rates and potential consumption rates using global positioning system collars"
3270:
3136:
3107:
2950:
2915:
2805:
2779:
2730:
2656:
2652:
2537:
2481:
2419:
2387:
2126:
2084:
2060:
1761:
1751:
1691:
1612:
1292:
1116:
1108:
1007:
991:
975:
893:
841:
837:
791:
504:
444:
440:
413:. Therefore, it was proposed to recognize only two tiger subspecies as valid, namely
369:
290:
5029:
The Amur Tiger Programme : Two Adult Tigers Tagged in the Ussuri Nature Reserve
4161:
4147:
4026:
3401:
3196:
3006:
2668:
2578:
1964:
1260:
Siberian tigers mate at any time of the year. A female signals her receptiveness by
409:
between populations in those regions is considered to have been possible during the
5117:
4888:"Distribution of the species comprised in the most numerous families of quadrupeds"
4384:
4135:
4106:
4061:
4053:
4004:
3873:"Siberian Tiger Family Caught On Film — Adult Female, Adult Male, & Three Cubs"
3730:
3701:
3475:
3438:
3389:
3306:
3260:
3250:
3212:"Siberian tiger video suggests species is returning to China, conservationists say"
3184:
3131:
3062:
2994:
2769:
2720:
2712:
2704:
2648:
2566:
2527:
2517:
2471:
2463:
2170:
2116:
2106:
2052:
1943:
1675:
1541:
1482:
1475:
1338:
1124:
826:
783:
478:
262:
250:
20:
4283:
Feasibility Study on the Possible Restoration of the Caspian Tiger in Central Asia
3574:
2826:
2622:
2498:
1714:), or a similar character meaning "Great Emperor", it is revered by the Udege and
1385:
5270:
5229:
4989:
4843:
4824:
4804:
4531:
4512:
4476:
4454:
4236:
4214:
4192:
4170:
3844:
3393:
3310:
3021:
2522:
2111:
1501:
1462:
1265:
1071:
865:
806:
648:
359:
340:
254:
164:
3497:"Denning ecology of brown bears and Asiatic black bears in the Russian Far East"
328:
4467:
4321:
3089:"Hierarchical spatial analysis of Amur tiger relationships to habitat and prey"
2998:
1923:
1788:
1715:
1563:
1490:
1486:
1471:
1407:
1227:
1168:
802:
716:
680:
602:
460:
386:
proposed by Charles Dode in 1871 was based on tiger skins from the Amur region.
286:
270:
4740:
4696:
4652:
4608:
4586:
3993:. Menlo Park and San Jose: Association for Computing Machinery. pp. 1–8.
3188:
2774:
2749:
2685:"The tiger genome and comparative analysis with lion and snow leopard genomes"
2570:
1593:
In January 2002, a man was attacked by a tiger on a remote mountain road near
868:, on the border between China and North Korea, based on tracks and sightings.
594:
5264:
5190:
4248:
2849:
1934:
1778:
1756:
1746:
1583:
1455:
1450:
1442:
1195:
1070:
Between January 1992 and November 1994, 11 tigers were captured, fitted with
659:
529:
483:
436:
344:
282:
5169:
4389:
4370:
4111:
4090:
4008:
3480:
3459:
2235:
Mammals of the Soviet Union. Volume II, Part 2. Carnivora (Hyaenas and Cats)
2056:
908:
5075:
4398:
4318:"Russia, Iran exchange tigers for leopards but some experts express doubts"
4075:
4039:
3734:
3274:
2783:
2734:
2660:
2541:
2485:
2467:
2130:
2064:
1828:
1741:
1679:
1632:
1553:
1184:
1160:
939:
795:
672:
623:
534:
317:
266:
5234:
4784:
3326:"Population dynamics of the Amur tiger in Sikhote-Alin Zapovednik, Russia"
2315:"Revision der zur natürlichen Familie der Katzen (Feles) gehörigen Formen"
5151:
5084:
3725:. In Ray, J. C.; Redford, K. H.; Steneck, R. S. & Berger, J. (eds.).
3625:. International Association for Bear Research and Management. p. 64.
1783:
1731:
1683:
1481:
In 2005, re-introduction was planned as part of the rewilding project at
1390:
1342:
1144:
779:
589:
452:
410:
258:
140:
37:
4968:
4959:
4139:
3415:
Dou, H.; Yang, H.; Smith, J.L.; Feng, L.; Wang, T. & Ge, J. (2019).
3228:
3150:
1636:
2020, a female tiger attacked and killed a 55-year-old zookeeper at the
631:
and that the individuals were usually sick or injured and captured in a
5143:
3791:
Wildlife Science: Linking Ecological Theory and Management Applications
3551:
3332:. Cambridge: Cambridge University Press. pp. 61–70. Archived from
2708:
2418:(Second ed.). London, Burlington: Academic Press. pp. 53–84.
2184:
1888:
1773:
1517:, 2,884 tigers were documented as residing in 468 American facilities.
1307:
1100:
1039:
931:
810:
606:
567:
456:
309:
241:
120:
5156:
4545:"Devils in the Darkness: The Korean Gray Wolf was a terror for miners"
3443:
3416:
3255:
3067:
2716:
2282:(decima, reformata ed.). Holmiae: Laurentius Salvius. p. 41.
1700:
1532:
1433:
Comparison of the distribution of all tiger subspecies (1900 vs. 1990)
1429:
4894:. Vol. 1. London: Sherwood, Gilbert & Piper. pp. 85–94.
3356:
2914:. Gland, Switzerland: IUCN/SSC Cat Specialist Group. pp. 55–64.
1559:
1438:
1422:
1152:
1023:
1015:
889:
849:
695:
691:
655:
555:
533:
suggested that the ancestor of Caspian and Siberian tigers colonized
508:
428:
406:
313:
298:
110:
70:
5046:
4057:
3988:
3457:
2441:
2175:
2018:"Sex imbalance as endangered Siberian tigers show signs of recovery"
1269:
5069:
4982:
3999:
3960:"Amur Tiger Factfile - Panthera tigris altaica - ALTA Conservation"
3529:. In Miquelle, D. G.; Smirnov, E. N. & Goodrich, J. M. (eds.).
3433:
3287:
3088:
2979:(Temminck, 1844), with a remark on Temminck's mammal volume of the
1628:
1624:
1303:
971:
852:
are restricted to the southern half of the Sikhote-Alin mountains.
833:
787:
775:
628:
499:
150:
90:
3414:
3330:
Riding the Tiger. Tiger Conservation in Human-dominated Landscapes
3093:
Riding the Tiger. Tiger Conservation in Human-dominated Landscapes
3023:
Jungle trails in northern India: reminiscences of hunting in India
2384:
Riding the Tiger: Tiger Conservation in Human-Dominated Landscapes
2380:"Tiger distribution, phenotypic variation and conservation issues"
2365:
Nutzbare Tiere Ostasiens. Pelz- und Jagdtiere, Haustiere, Seetiere
4249:
Jungius, H.; Chikin, Y.; Tsaruk, O. & Pereladova, O. (2009).
1594:
1394:
959:
684:
320:
tiger", depending on the region where individuals were observed.
302:
130:
19:"Amur tiger" redirects here. For the leopard cat subspecies, see
5130:
3019:
2854:. In Miquelle, D.G.; Smirnov, E.N. & Goodrich, J.M. (eds.).
2683:
Lee, S.; Zhou, Q.; Jung, H. J.; Xu, X. & Gadhvi, P. (2013).
2554:
3720:
3623:
20th International Conference on Bear Research & Management
3371:
3328:. In Seidensticker, J.; Christie, S. & Jackson, P. (eds.).
3091:. In Seidensticker, J.; Christie, S. & Jackson, P. (eds.).
2413:
Tigers of the World: The Science, Politics and Conservation of
1705:
1687:
1569:
1467:
1059:
1051:
782:, where the Caspian tiger also reportedly occurred. During the
700:
100:
80:
2297:. In Siebold, P. F. v.; Temminck, C. J.; Schlegel, H. (eds.).
703:, which measured 413 mm (16.25 in) "over the bone".
4525:"Russia to donate three rare Siberian tigers to South Korea."
4088:
3788:
3564:
3165:
2224:
1620:
1598:
1590:
investigated both deaths, tracked down and killed the tiger.
1574:
1137:
1063:
999:
897:
822:
676:
578:
294:
238:
4344:"Iran, world, political, sport, economic news and headlines"
2886:"Two long-term resident tigers in the Ussuri Nature Reserve"
3638:"Роль бурого медведя в экосистемах Дальнего Востока России"
3531:
Tigers of Sikhote-Alin Zapovednik: Ecology and Conservation
3039:"Predicted Pleistocene–Holocene range shifts of the tiger (
2856:
Tigers in Sikhote-Alin Zapovednik: Ecology and Conservation
2834:
Tigers in Sikhote-Alin Zapovednik: Ecology and Conservation
2015:
1664:
1376:
International Forum on Tiger Conservation and Tiger Culture
1296:
1176:
1133:
1128:
1055:
1047:
498:
Several reports have been published since the 1990s on the
281:
The Siberian tiger is genetically close to the now-extinct
3898:. Chicago: University of Chicago Press. pp. 343–371.
3816:"Battle of the big cats sees tiger hunt and devour a lynx"
2832:. In D. G. Miquelle; E. N. Smirnov; J.M. Goodrich (eds.).
2406:
2382:. In Seidensticker, J.; Christie, S.; Jackson, P. (eds.).
1461:
In 2010, Russia exchanged two captive Siberian tigers for
3935:"About the Siberian tiger - Russian Geographical Society"
3533:(in Russian). Vladivostok, Russia: PSP. pp. 156–163.
2409:"What is a Tiger? Biogeography, Morphology, and Taxonomy"
1889:
Rak, K. C.; Miquelle, D. G. & Pikunov, D. G. (1998).
1437:
A second possible introduction site in Kazakhstan is the
3889:
3841:
2883:
2594:"Rare Siberian tigers face potential genetic bottleneck"
2231:
Mlekopitajuščie Sovetskogo Soiuza. Moskva: Vysšaia Škola
1082:
5016:
Amur.org.uk: Preserving leopards and tigers in the wild
4506:"S Korea to Welcome Three Siberian Tigers from Russia."
3357:
Miquelle, D.; Goodrich, J. & Seryodkin, I. (2008).
3105:
2852:"Chapter 7. Morphological indicators of the Amur tiger"
2836:(in Russian). Vladivostok, Russia: PSP. pp. 25–35.
519:
At the start of the 21st century, researchers from the
5011:
Wildlife Conservation Society's Siberian Tiger Project
2858:(in Russian). Vladivostok, Russia: PSP. pp. 1–15.
2747:
4371:"Pleistocene Park: Return of the Mammoth's Ecosystem"
3727:
Large Carnivores and the Conservation of Biodiversity
2912:
Wild Cats: Status Survey and Conservation Action Plan
2898:
1995:"Happy tigers: Siberian population continues to grow"
605:
width 180 mm (7.1 in), and length of upper
372:
in 1868 was based on a long-haired tiger skin in the
4818:"Siberian tiger attacks, kills bus driver in China."
4787:. The Associated Press, 17 January 2008. 2008-01-18.
4311:
4309:
1295:
is low, only 27–35 individuals contributed to their
4547:. English.ohmynews.com, 23 May 2007. Archived from
3567:"Hunting Behavior and Success of the Tigers' Hunts"
3289:meta-population of Amur tigers in Northeast Asia".
2041:
647:weighed 206 kg (454 lb), and the largest
4239:. Global Tiger Initiative Secretariat, Washington.
3843:
3813:
3082:
3080:
3078:
2251:Loukashkin, A. S. (1938). "The Manchurian Tiger".
829:forests with a complex composition and structure.
392:by Emil Brass in 1904 was a tiger skin from Korea.
4785:"Tiger attack victim admits taunting, police say"
4306:
4204:Miquelle, D., Goodrich, J., Seryodkin, I. (2008)
3671:"Brown Bear predation of Amur Tiger 1973 account"
3095:. UK: Cambridge University Press. pp. 71–99.
3013:
2892:
2437:
2435:
2250:
405:, tigers from different regions vary little, and
5262:
4315:
3635:
3020:Hewett, J. P. & Hewett Atkinson, L. (1938).
2845:
2843:
2502:"Phylogeography and genetic ancestry of tigers (
2312:
1659:The English name 'Siberian tiger' was coined by
1167:The relationship between the Amur tiger and the
1099:Taxidermy exhibit portraying a tiger fighting a
4892:Researches into the Physical History of Mankind
4182:Nowell, K., Bauer, H., Breitenmoser, U. (2007)
4095:) on Sikhote-Alin Biosphere Zapovednik, Russia"
3594:
3323:
3075:
2350:Proceedings of the Zoological Society of London
2292:
2220:
2218:
2216:
2214:
1949:10.2305/IUCN.UK.2022-1.RLTS.T15955A214862019.en
1884:
1882:
1880:
1862:Nowell, K.; Timmons, Z. & Tobe, S. (2017).
1046:). Siberian tigers also take smaller prey like
4885:
4091:"Spatial structure of Amur (Siberian) tigers (
2631:representation of the endangered Amur tiger".
2432:
2386:. Cambridge University Press. pp. 19–39.
2225:Heptner, V. G. & Sludskij, A. A. (1992) .
2212:
2210:
2208:
2206:
2204:
2202:
2200:
2198:
2196:
2194:
1380:China 2010 Hunchun Amur Tiger Culture Festival
1210:
912:A Siberian tiger photographed by a camera trap
836:complex is represented by seven species, with
447:, Siberian and Caspian tiger populations; and
4904:
3789:Fulbright, T. E. & Hewitt, D. G. (2007).
3565:Yudakov, A. G. & Nikolaev, I. G. (2004).
3543:
3537:
3520:
3518:
3516:
3514:
3352:
3350:
3324:Smirnov, E. N. & Miquelle, D. G. (1999).
2840:
2799:
2400:
2371:
2246:
2244:
1653:Heraldic arms of the Jewish Autonomous Oblast
5003:National Geographic Animals: Siberian Tiger
4368:
3842:Matthiessen, P. & Hornocker, M. (2001).
2968:
2966:
2933:
2931:
2818:
2269:
1877:
1302:The winter of 2006–2007 was marked by heavy
925:reproductive output in the core area of the
798:basin, before humans interrupted gene flow.
4969:IUCN/SSC Cat Specialist Group : Amur (
4933:Landscape & Culture in Northern Eurasia
4837:"Tiger kills zookeeper during feeding time"
4280:
3918:
3760:"'Putin's Tigers' are Thriving in the Wild"
3365:
3317:
3209:
2681:
2411:. In Tilson, R. & Nyhus, P. J. (eds.).
2191:
2146:
2144:
2142:
2140:
734:
5033:
4857:"Siberian tiger kills zookeeper in Zurich"
4438:
3757:
3511:
3347:
3108:"Spatial viability analysis of Amur tiger
2868:
2618:
2616:
2407:Kitchener, A. & Yamaguchi, N. (2010).
2241:
2080:
2078:
2076:
2074:
2037:
2035:
1962:
301:corridor, and then subsequently traversed
202:
36:
4791:
4411:
4388:
4242:
4110:
4065:
3998:
3705:
3479:
3442:
3432:
3264:
3254:
3135:
3066:
2972:
2963:
2937:
2928:
2773:
2724:
2531:
2521:
2475:
2377:
2362:
2174:
2150:
2120:
2110:
1992:
1947:
1558:The Siberian tiger very rarely becomes a
1341:, banning international trade. All tiger
964:Museo Civico di Storia Naturale di Milano
4542:
4125:
3890:Sunquist, M. & Sunquist, F. (2002).
3870:
2335:
2137:
1647:
1568:
1531:
1500:
1428:
1384:
1262:leaving urine deposits and scratch marks
1094:
1091:Siberian tiger. Frame from a camera trap
1086:
953:
907:
710:
651:male weighed 212 kg (467 lb).
638:Measurements taken by scientists of the
588:
477:
327:
4229:Global Tiger Recovery Program 2010–2022
4173:. TRAFFIC International, Cambridge, UK.
3646:Oles Honchar Dnipro National University
3488:
3361:. Bronx: Wildlife Conservation Society.
3106:Carroll, C. & Miquelle, D. (2006).
2613:
2071:
2032:
1965:"Russia Announce Tiger Census Results!"
1856:
1854:
903:
507:lineages. It seems that a single mtDNA
5263:
5021:USDA Information Resources on Tigers,
4960:IUCN/SSC Cat Specialist Group: Tiger (
4935:. Walnut Creek, CA: Left Coast Press.
4926:
4457:. TRAFFIC East Asia, Hong Kong, China.
4220:
3837:
3835:
3833:
3831:
3829:
3026:. London: Metheun and Company Limited.
2975:"Notes on Siberian long-haired tiger,
2795:
2793:
1919:
1917:
1915:
1194:Siberian tigers also compete with the
5051:
5050:
4469:Less than 3,000 Pet Tigers in America
2899:Nowell, K. & Jackson, P. (1996).
2591:
1586:valley. The anti-poaching task force
1083:Interspecific predatory relationships
5301:Siberian Tiger Re-population Project
4633:. News.sina.com.cn. 13 February 2002
4414:"Mammoth Steppes and Future Climate"
4288:(Report). WWF Russia. Archived from
4256:(Report). WWF Russia. Archived from
2016:The Siberian Times reporter (2015).
1851:
1527:
1445:. The delta is situated between the
819:temperate broadleaf and mixed forest
794:population through corridors in the
308:The Siberian tiger was also called "
265:, but currently inhabits mainly the
4419:. Science in Russia. Archived from
3826:
2790:
1935:IUCN Red List of Threatened Species
1912:
1505:A tigress with cub in captivity in
1416:Siberian Tiger Introduction Project
1362:, part of the Russian government's
949:
938:Siberian tigers share habitat with
13:
4741:"走遍中国2009年06月20日C:寻踪东北虎系列之人虎情仇(下)"
4697:"走遍中国2009年06月20日B:寻踪东北虎系列之人虎情仇(下)"
4653:"走遍中国2009年06月20日A:寻踪东北虎系列之人虎情仇(下)"
4609:"走遍中国2009年06月19日C:寻踪东北虎系列之人虎情仇(上)"
4587:"走遍中国2009年06月19日A:寻踪东北虎系列之人虎情仇(上)"
3648:. pp. 502–503. Archived from
1669:American Museum of Natural History
1207:competitor than on catching prey.
970:Prey species of the tiger include
584:
16:Tiger population in Northeast Asia
14:
5322:
4953:
4827:Associated Press, 5 January 2011.
4226:Global Tiger Initiative. (2011).
3923:. Stillwater, MN: Voyageur Press.
2884:The Amur Tiger Programme (2014).
1577:scroll from the late 18th century
1401:
1030:), and even sometimes small size
966:shows a tiger chasing a sika deer
790:, it was likely connected to the
528:subspecies. They re-assessed the
5036:"Amur tigers on 'genetic brink'"
4920:
4898:
4879:
4849:
4830:
3707:10.1111/j.1469-7998.2008.00458.x
3137:10.1111/j.1365-2664.2006.01237.x
2653:10.1111/j.1365-294X.2009.04266.x
1237:
1219:
757:
743:
261:. It once ranged throughout the
57:
5306:National symbols of South Korea
4810:
4777:
4755:
4733:
4711:
4689:
4667:
4645:
4623:
4601:
4579:
4563:
4536:
4517:
4504:People's Daily, 21 April 2011.
4498:
4494:(in Chinese). SINA Corporation.
4482:
4479:. Wildlife Times 5 (37): 12–13.
4460:
4405:
4362:
4336:
4274:
4217:. Wildlife Conservation Society
4198:
4176:
4154:
4119:
4082:
4033:
3982:
3952:
3927:
3912:
3883:
3864:
3807:
3782:
3751:
3714:
3681:
3675:International Wildlife Magazine
3663:
3629:
3607:
3588:
3558:
3451:
3408:
3281:
3222:
3216:Guardian News and Media Limited
3203:
3159:
3144:
3099:
3030:
2877:
2862:
2741:
2675:
2585:
2548:
2492:
2356:
2329:
2306:
2286:
2263:
1667:River for an exhibition in the
1496:
1332:
1320:dissolution of the Soviet Union
873:Hunchun National Nature Reserve
706:
679:prominences, especially in the
4798:Siberian tiger kills zookeeper
4320:. Payvand News. Archived from
3603:. New York: North Point Press.
2009:
1986:
1956:
1710:
1699:
1515:Feline Conservation Federation
1441:delta at the southern edge of
1313:
750:Sikhote-Alin in Primorsky Krai
525:U.S. National Cancer Institute
374:Natural History Museum, Vienna
1:
5286:Fauna of the Russian Far East
4863:. 4 July 2020. Archived from
4846:The Local, 20 September 2013.
1844:
1643:
1573:A tiger family depicted in a
980:Cervus canadensis xanthopygus
316:tiger", "Korean tiger", and "
269:mountain region in southwest
222: Range in the late 1800s
4995:World Wide Fund for Nature:
4466:Wildlife Watch Group (2011)
4444:Nowell, K., Ling, X. (2007)
3394:10.1016/j.mambio.2018.03.009
3311:10.1016/j.biocon.2021.109250
2802:Feline Behaviour and Welfare
2523:10.1371/journal.pbio.0020442
2302:. Leiden: Lugduni Batavorum.
2112:10.1371/journal.pone.0004125
813:. It also occurs within the
715:A Siberian tiger cub at the
612:
473:
211:Siberian tiger distribution
7:
4807:, TRHK News, 16 August 2010
3814:Blaszczak-Boxe, A. (2016).
3047:Diversity and Distributions
1813:Panthera tigris trinilensis
1737:Mainlaind Asian populations
1724:
1623:province and the cities of
1211:Reproduction and life cycle
1157:Sikhote-Alin Nature Reserve
323:
10:
5327:
4763:"From hunter to protector"
4675:"先后动了两次手术 我国首次抢救重伤野生东北虎纪实"
4346:. MehrNews. Archived from
3116:Journal of Applied Ecology
2999:10.1515/mamm.1967.31.4.537
1873:(Special Issue 11): 66−68.
1551:
1522:Harbin Siberian Tiger Park
1405:
1286:
878:World Wide Fund for Nature
593:Captive Siberian tiger at
18:
5311:Mammals described in 1884
5059:
4914:American Museum Novitates
4316:Khosravifard, S. (2010).
3636:Seryodkin, I. V. (2007).
3189:10.1007/s10980-015-0278-1
2871:"The Amur tiger: Ecology"
2775:10.1016/j.cub.2018.09.019
2627:population structure and
2571:10.1007/s10592-004-1860-2
2313:Fitzinger, L. J. (1868).
1973:Worldwide Fund for Nature
1820:Panthera tigris acutidens
1806:Panthera tigris soloensis
1545:
1327:Korea under Japanese rule
210:
201:
54:Scientific classification
52:
47:Leipzig Zoological Garden
44:
35:
30:
4523:Reuters, 22 April 2011.
4490:
3595:Matthiessen, P. (2000).
3460:"Estimating Amur tiger (
2804:. CABI. pp. 72–77.
2293:Temminck, C. J. (1844).
1896:(Report). Archived from
1769:Sunda island populations
735:Distribution and habitat
666:
5291:Fauna of Northeast Asia
5105:Panthera_tigris_altaica
5091:Panthera tigris altaica
5061:Panthera tigris altaica
5005:Panthera tigris altaica
4931:. In Jordan, P. (ed.).
4886:Prichard, J.C. (1836).
4743:. YouTube. 20 June 2009
4699:. YouTube. 20 June 2009
4655:. YouTube. 20 June 2009
4631:"吉林"东北虎吃人"迷雾重重:老虎胃里是空的"
4611:. YouTube. 18 June 2009
4589:. YouTube. 18 June 2009
4390:10.1126/science.1113442
4112:10.1644/09-mamm-a-293.1
4093:Panthera tigris altaica
4009:10.1145/3209811.3209879
3758:Vorontsova, M. (2015).
3481:10.1644/12-mamm-a-209.1
3462:Panthera tigris altaica
3291:Biological Conservation
3110:Panthera tigris altaica
2977:Panthera tigris altaica
2500:O'Brien, S. J. (2004).
2057:10.1111/1749-4877.12210
1546:Осторожно! Тигры рядoм!
1538:Caution! Tigers nearby!
1337:Tigers are included on
1329:between 1910 and 1945.
1123:) were recorded in the
1121:U. thibetanus ussuricus
927:Sikhote-Alin Zapovednik
490:) and Siberian (ALT or
435:comprising the Bengal,
356:Coenraad Jacob Temminck
237:is a population of the
4905:Goodwin, G.G. (1933).
4719:"吉林两男子欲捕东北虎 其中一人反被虎咬伤"
4128:Ecological Restoration
3896:Wild Cats of the World
3544:Seryodkin, I. (2006).
2800:Fraser, A. F. (2012).
2468:10.1126/sciadv.1400175
2378:Kitchener, A. (1999).
2367:. Neudamm: J. Neumann.
1942:: e.T15955A214862019.
1656:
1578:
1549:
1509:
1447:Saryesik-Atyrau Desert
1434:
1398:
1356:Siberian Tiger Project
1351:Siberian Tiger Project
1104:
1092:
967:
913:
809:east and south of the
719:
645:Siberian Tiger Project
640:Siberian Tiger Project
597:
495:
417:in mainland Asia, and
415:Panthera tigris tigris
390:Felis tigris coreensis
362:and Pisihan Mountains.
333:
246:Panthera tigris tigris
216: Modern-day range
45:Siberian tiger at the
4369:Zimov, S. A. (2005).
4195:. Cat News 47: 33–34.
4046:Ecology and Evolution
3850:. North Point Press.
3235:Ecology and Evolution
2689:Nature Communications
2592:Platt, J. R. (2009).
2558:Conservation Genetics
2270:Linnaeus, C. (1758).
1661:James Cowles Prichard
1651:
1572:
1535:
1507:DierenPark Amersfoort
1504:
1432:
1412:Species translocation
1406:Further information:
1388:
1204:Bastak Nature Reserve
1113:Ursus arctos lasiotus
1098:
1090:
957:
911:
815:Greater Xing'an Range
766:Bastak Nature Reserve
714:
592:
539:Industrial Revolution
481:
425:Greater Sunda Islands
352:Felis tigris altaicus
331:
4281:Jungius, H. (2010).
4099:Journal of Mammalogy
3964:Altaconservation.org
3919:Karanth, U. (2001).
3737:. pp. 179–207.
3468:Journal of Mammalogy
3210:Vaughan, A. (2014).
2020:. The Siberian Times
1803:Prehistoric tigers:
1686:call it "Amba". The
1103:, Vladivostok Museum
996:Naemorhedus caudatus
944:P. pardus orientalis
904:Ecology and behavior
521:University of Oxford
276:conservation efforts
5034:Walker, M. (2009).
4765:. Chinadaily.com.cn
4185:Cats at CITES COP14
4140:10.3368/er.34.3.173
3386:2018MamBi..92..120Y
3303:2021BCons.26109250Q
3247:2019EcoEv...9.2415N
3181:2016LaEco..31..491W
3128:2006JApEc..43.1056C
3059:2016DivDi..22.1199C
2869:WCS Russia (2015).
2766:2018CBio...28E3840L
2701:2013NatCo...4.2433C
2645:2009MolEc..18.3173H
2599:Scientific American
2460:2015SciA....1E0175W
2103:2009PLoSO...4.4125D
2045:Integrative Zoology
1963:WWF Russia (2015).
1839:Holocene extinction
1360:Tiger Response Team
1032:Asiatic black bears
988:Moschus moschiferus
886:Zhangguangcai Range
332:Siberian tiger face
4988:2017-03-18 at the
4978:21st Century Tiger
4927:Sasaki, S (2011).
4842:2020-09-15 at the
4823:2015-02-06 at the
4816:Shahid, A. (2011)
4803:2012-08-18 at the
4721:. News.sina.com.cn
4677:. News.sina.com.cn
4530:2019-10-01 at the
4511:2019-10-01 at the
4475:2020-10-03 at the
4453:2012-01-17 at the
4426:on 29 October 2013
4412:Zimov, S. (2007).
4324:on 15 October 2020
4235:2011-08-26 at the
4213:2012-04-25 at the
4191:2020-08-07 at the
4169:2020-10-03 at the
4160:Nowell, K. (2007)
3846:Tigers In The Snow
3694:Journal of Zoology
3677:. 20 October 2009.
3644:. Denpropetrovsk:
3601:Tigers in the Snow
2973:Mazák, V. (1967).
2709:10.1038/ncomms3433
2363:Brass, E. (1904).
1993:Hance, J. (2015).
1900:on 5 December 2020
1657:
1603:endangered species
1579:
1550:
1510:
1489:basin in northern
1435:
1399:
1389:Siberian Tiger at
1374:in July 2010, and
1117:Ussuri black bears
1105:
1093:
1012:Capreolus pygargus
984:Siberian musk deer
968:
918:population density
914:
882:Changbai Mountains
854:Siberian musk deer
846:spruce-fir forests
805:and into southern
720:
633:conflict situation
598:
514:genetic bottleneck
496:
334:
5281:Mammals of Russia
5258:
5257:
5243:Open Tree of Life
5053:Taxon identifiers
4543:Neff, R. (2007).
4383:(5723): 796–798.
3871:Ayre, J. (2015).
3857:978-0-86547-596-0
3800:978-0-8493-7487-6
3770:on August 7, 2020
3444:10.2981/wlb.00508
3374:Mammalian Biology
3256:10.1002/ece3.4832
3169:Landscape Ecology
3068:10.1111/ddi.12484
3053:(11): 1199–1211.
2956:978-3-89432-759-0
2921:978-2-8317-0045-8
2905:(Linnaeus, 1758)"
2811:978-1-84593-926-7
2760:(23): 3840–3849.
2639:(15): 3173–3184.
2633:Molecular Ecology
2425:978-0-08-094751-8
2393:978-0-521-64835-6
2336:Dode, C. (1871).
2253:The China Journal
2167:Mammalian Species
1762:South China tiger
1752:Indochinese tiger
1732:Tiger populations
1692:Chinese character
1613:San Francisco Zoo
1528:Attacks on humans
1293:genetic diversity
1248:
1008:Siberian roe deer
992:long-tailed goral
976:Manchurian wapiti
894:Khingan Mountains
842:Siberian roe deer
838:Manchurian wapiti
792:South China tiger
505:mitochondrial DNA
370:Leopold Fitzinger
366:Tigris longipilis
291:mitochondrial DNA
271:Primorye Province
227:
226:
182:P. t. tigris
171:
5318:
5276:Mammals of China
5251:
5250:
5238:
5237:
5225:
5224:
5212:
5211:
5199:
5198:
5186:
5185:
5173:
5172:
5160:
5159:
5147:
5146:
5134:
5133:
5121:
5120:
5108:
5107:
5095:
5094:
5093:
5080:
5079:
5078:
5048:
5047:
5043:
4947:
4946:
4924:
4918:
4917:
4911:
4902:
4896:
4895:
4883:
4877:
4876:
4874:
4872:
4853:
4847:
4834:
4828:
4814:
4808:
4795:
4789:
4788:
4781:
4775:
4774:
4772:
4770:
4759:
4753:
4752:
4750:
4748:
4737:
4731:
4730:
4728:
4726:
4715:
4709:
4708:
4706:
4704:
4693:
4687:
4686:
4684:
4682:
4671:
4665:
4664:
4662:
4660:
4649:
4643:
4642:
4640:
4638:
4627:
4621:
4620:
4618:
4616:
4605:
4599:
4598:
4596:
4594:
4583:
4577:
4567:
4561:
4560:
4558:
4556:
4540:
4534:
4521:
4515:
4502:
4496:
4495:
4486:
4480:
4464:
4458:
4442:
4436:
4435:
4433:
4431:
4425:
4418:
4409:
4403:
4402:
4392:
4366:
4360:
4359:
4357:
4355:
4340:
4334:
4333:
4331:
4329:
4313:
4304:
4303:
4301:
4300:
4294:
4287:
4278:
4272:
4271:
4269:
4268:
4262:
4255:
4246:
4240:
4224:
4218:
4202:
4196:
4180:
4174:
4158:
4152:
4151:
4123:
4117:
4116:
4114:
4086:
4080:
4079:
4069:
4037:
4031:
4030:
4002:
3986:
3980:
3979:
3977:
3975:
3966:. Archived from
3956:
3950:
3949:
3947:
3945:
3931:
3925:
3924:
3916:
3910:
3909:
3887:
3881:
3880:
3868:
3862:
3861:
3849:
3839:
3824:
3823:
3811:
3805:
3804:
3786:
3780:
3779:
3777:
3775:
3766:. Archived from
3755:
3749:
3748:
3731:Washington, D.C.
3718:
3712:
3711:
3709:
3685:
3679:
3678:
3667:
3661:
3660:
3658:
3657:
3633:
3627:
3626:
3620:
3611:
3605:
3604:
3592:
3586:
3585:
3583:
3582:
3562:
3556:
3555:
3541:
3535:
3534:
3522:
3509:
3508:
3492:
3486:
3485:
3483:
3455:
3449:
3448:
3446:
3436:
3421:Wildlife Biology
3412:
3406:
3405:
3369:
3363:
3362:
3354:
3345:
3344:
3342:
3341:
3321:
3315:
3314:
3285:
3279:
3278:
3268:
3258:
3241:(5): 2415–2424.
3226:
3220:
3219:
3207:
3201:
3200:
3163:
3157:
3156:
3148:
3142:
3141:
3139:
3122:(6): 1056–1068.
3103:
3097:
3096:
3084:
3073:
3072:
3070:
3034:
3028:
3027:
3017:
3011:
3010:
2970:
2961:
2960:
2935:
2926:
2925:
2909:
2896:
2890:
2889:
2881:
2875:
2874:
2866:
2860:
2859:
2847:
2838:
2837:
2831:
2822:
2816:
2815:
2797:
2788:
2787:
2777:
2745:
2739:
2738:
2728:
2679:
2673:
2672:
2620:
2611:
2610:
2608:
2606:
2589:
2583:
2582:
2552:
2546:
2545:
2535:
2525:
2496:
2490:
2489:
2479:
2448:Science Advances
2439:
2430:
2429:
2404:
2398:
2397:
2375:
2369:
2368:
2360:
2354:
2353:
2333:
2327:
2326:
2310:
2304:
2303:
2290:
2284:
2283:
2267:
2261:
2260:
2248:
2239:
2238:
2222:
2189:
2188:
2178:
2164:
2148:
2135:
2134:
2124:
2114:
2082:
2069:
2068:
2039:
2030:
2029:
2027:
2025:
2013:
2007:
2006:
2004:
2002:
1990:
1984:
1983:
1981:
1979:
1969:tigers.panda.org
1960:
1954:
1953:
1951:
1921:
1910:
1909:
1907:
1905:
1895:
1886:
1875:
1874:
1868:
1858:
1712:
1703:
1676:Tungusic peoples
1640:in Switzerland.
1588:Inspection Tiger
1547:
1483:Pleistocene Park
1476:Caspian red deer
1463:Persian leopards
1372:Global Tiger Day
1364:Inspection Tiger
1339:CITES Appendix I
1250:
1249:
1223:
1155:research in the
1125:Russian Far East
1036:Ursus thibetanus
950:Hunting and diet
935:protected area.
784:late Pleistocene
778:and the area of
761:
747:
427:and possibly in
289:study comparing
263:Korean Peninsula
251:Russian Far East
221:
215:
206:
197:
185:
170:
168:
62:
61:
40:
28:
27:
21:Amur leopard cat
5326:
5325:
5321:
5320:
5319:
5317:
5316:
5315:
5296:Northeast China
5261:
5260:
5259:
5254:
5246:
5241:
5233:
5230:Observation.org
5228:
5220:
5215:
5207:
5202:
5194:
5189:
5181:
5176:
5168:
5163:
5155:
5150:
5142:
5137:
5129:
5124:
5116:
5111:
5103:
5098:
5089:
5088:
5083:
5074:
5073:
5068:
5055:
5023:Panthera tigris
4990:Wayback Machine
4962:Panthera tigris
4956:
4951:
4950:
4943:
4925:
4921:
4909:
4903:
4899:
4884:
4880:
4870:
4868:
4867:on July 5, 2020
4855:
4854:
4850:
4844:Wayback Machine
4835:
4831:
4825:Wayback Machine
4815:
4811:
4805:Wayback Machine
4796:
4792:
4783:
4782:
4778:
4768:
4766:
4761:
4760:
4756:
4746:
4744:
4739:
4738:
4734:
4724:
4722:
4717:
4716:
4712:
4702:
4700:
4695:
4694:
4690:
4680:
4678:
4673:
4672:
4668:
4658:
4656:
4651:
4650:
4646:
4636:
4634:
4629:
4628:
4624:
4614:
4612:
4607:
4606:
4602:
4592:
4590:
4585:
4584:
4580:
4568:
4564:
4554:
4552:
4551:on 5 March 2016
4541:
4537:
4532:Wayback Machine
4522:
4518:
4513:Wayback Machine
4503:
4499:
4492:
4488:
4487:
4483:
4477:Wayback Machine
4465:
4461:
4455:Wayback Machine
4443:
4439:
4429:
4427:
4423:
4416:
4410:
4406:
4367:
4363:
4353:
4351:
4342:
4341:
4337:
4327:
4325:
4314:
4307:
4298:
4296:
4292:
4285:
4279:
4275:
4266:
4264:
4260:
4253:
4247:
4243:
4237:Wayback Machine
4225:
4221:
4215:Wayback Machine
4203:
4199:
4193:Wayback Machine
4181:
4177:
4171:Wayback Machine
4159:
4155:
4124:
4120:
4087:
4083:
4058:10.1002/ece3.46
4038:
4034:
4019:
3987:
3983:
3973:
3971:
3958:
3957:
3953:
3943:
3941:
3933:
3932:
3928:
3917:
3913:
3906:
3888:
3884:
3869:
3865:
3858:
3840:
3827:
3812:
3808:
3801:
3787:
3783:
3773:
3771:
3756:
3752:
3745:
3719:
3715:
3686:
3682:
3669:
3668:
3664:
3655:
3653:
3634:
3630:
3618:
3612:
3608:
3593:
3589:
3580:
3578:
3563:
3559:
3542:
3538:
3523:
3512:
3493:
3489:
3456:
3452:
3413:
3409:
3370:
3366:
3355:
3348:
3339:
3337:
3322:
3318:
3286:
3282:
3227:
3223:
3208:
3204:
3164:
3160:
3149:
3145:
3104:
3100:
3085:
3076:
3041:Panthera tigris
3035:
3031:
3018:
3014:
2971:
2964:
2957:
2936:
2929:
2922:
2907:
2903:Panthera tigris
2897:
2893:
2882:
2878:
2867:
2863:
2848:
2841:
2829:
2823:
2819:
2812:
2798:
2791:
2754:Current Biology
2746:
2742:
2680:
2676:
2621:
2614:
2604:
2602:
2590:
2586:
2553:
2549:
2504:Panthera tigris
2497:
2493:
2454:(5): e1400175.
2440:
2433:
2426:
2415:Panthera tigris
2405:
2401:
2394:
2376:
2372:
2361:
2357:
2352:(May): 480–481.
2334:
2330:
2311:
2307:
2291:
2287:
2268:
2264:
2249:
2242:
2223:
2192:
2176:10.2307/3504004
2162:
2158:Panthera tigris
2149:
2138:
2083:
2072:
2040:
2033:
2023:
2021:
2014:
2010:
2000:
1998:
1991:
1987:
1977:
1975:
1961:
1957:
1928:Panthera tigris
1922:
1913:
1903:
1901:
1893:
1887:
1878:
1866:
1859:
1852:
1847:
1834:Bergmann's rule
1825:
1797:
1727:
1646:
1556:
1530:
1499:
1418:
1404:
1382:in August 2010.
1335:
1316:
1289:
1258:
1257:
1256:
1255:
1254:
1251:
1238:
1233:
1232:
1231:
1224:
1213:
1175:Tigers depress
1085:
952:
906:
866:Baekdu Mountain
807:Khabarovsk Krai
772:
771:
770:
769:
768:
762:
753:
752:
751:
748:
737:
709:
669:
629:illegal hunting
615:
587:
585:Characteristics
562:that were lost
476:
451:comprising the
403:Morphologically
341:scientific name
326:
287:phylogeographic
285:. Results of a
255:Northeast China
223:
219:
217:
213:
193:
179:
169:
162:
56:
31:Siberian tiger
24:
17:
12:
11:
5:
5324:
5314:
5313:
5308:
5303:
5298:
5293:
5288:
5283:
5278:
5273:
5256:
5255:
5253:
5252:
5239:
5226:
5213:
5200:
5187:
5174:
5161:
5148:
5135:
5122:
5109:
5096:
5081:
5065:
5063:
5057:
5056:
5045:
5044:
5040:BBC Earth News
5031:
5026:
5018:
5013:
5008:
5000:
4992:
4980:
4975:
4966:
4955:
4954:External links
4952:
4949:
4948:
4941:
4919:
4897:
4878:
4848:
4829:
4809:
4790:
4776:
4754:
4732:
4710:
4688:
4666:
4644:
4622:
4600:
4578:
4562:
4535:
4516:
4497:
4491:黑龙江东北虎林园将打造千虎园
4481:
4459:
4437:
4404:
4361:
4335:
4305:
4273:
4241:
4219:
4197:
4175:
4153:
4134:(3): 173–183.
4118:
4105:(3): 737–748.
4081:
4052:(4): 529–545.
4032:
4017:
3981:
3951:
3926:
3911:
3904:
3882:
3863:
3856:
3825:
3806:
3799:
3781:
3750:
3743:
3713:
3680:
3662:
3628:
3606:
3587:
3557:
3554:on 2013-12-24.
3536:
3510:
3487:
3474:(4): 845–855.
3450:
3407:
3364:
3346:
3316:
3297:(12): 109250.
3280:
3221:
3202:
3175:(3): 491−503.
3158:
3143:
3098:
3074:
3029:
3012:
2993:(4): 537–573.
2981:Fauna Japonica
2962:
2955:
2927:
2920:
2891:
2876:
2861:
2839:
2817:
2810:
2789:
2740:
2674:
2612:
2584:
2565:(5): 707–713.
2547:
2491:
2431:
2424:
2399:
2392:
2370:
2355:
2328:
2305:
2285:
2262:
2240:
2190:
2136:
2070:
2051:(4): 322−332.
2031:
2008:
1997:. Mongabay.com
1985:
1955:
1911:
1876:
1849:
1848:
1846:
1843:
1842:
1841:
1836:
1831:
1826:
1824:
1823:
1816:
1809:
1800:
1796:
1795:
1794:
1793:
1792:
1791:
1789:Sumatran tiger
1786:
1781:
1776:
1766:
1765:
1764:
1759:
1754:
1749:
1744:
1728:
1726:
1723:
1716:Chinese people
1645:
1642:
1619:Zookeepers in
1609:In an incident
1599:Jilin province
1564:Jilin Province
1529:
1526:
1498:
1495:
1472:Persian onager
1408:Reintroduction
1403:
1402:Reintroduction
1400:
1334:
1331:
1315:
1312:
1288:
1285:
1252:
1236:
1235:
1234:
1228:Primorsky Krai
1225:
1218:
1217:
1216:
1215:
1214:
1212:
1209:
1169:Himalayan bear
1127:, and hair of
1084:
1081:
951:
948:
905:
902:
803:Primorsky Krai
763:
756:
755:
754:
749:
742:
741:
740:
739:
738:
736:
733:
717:Pittsburgh Zoo
708:
705:
681:sagittal crest
668:
665:
649:radio-collared
614:
611:
586:
583:
475:
472:
461:Sumatran tiger
449:P. t. sondaica
420:P. t. sondaica
394:
393:
387:
377:
363:
325:
322:
249:native to the
231:Siberian tiger
225:
224:
218:
212:
208:
207:
199:
198:
195:Siberian tiger
191:
187:
186:
177:
173:
172:
160:
156:
155:
148:
144:
143:
138:
134:
133:
128:
124:
123:
118:
114:
113:
108:
104:
103:
98:
94:
93:
88:
84:
83:
78:
74:
73:
68:
64:
63:
50:
49:
42:
41:
33:
32:
15:
9:
6:
4:
3:
2:
5323:
5312:
5309:
5307:
5304:
5302:
5299:
5297:
5294:
5292:
5289:
5287:
5284:
5282:
5279:
5277:
5274:
5272:
5269:
5268:
5266:
5249:
5244:
5240:
5236:
5231:
5227:
5223:
5218:
5214:
5210:
5205:
5201:
5197:
5192:
5188:
5184:
5179:
5175:
5171:
5166:
5162:
5158:
5153:
5149:
5145:
5140:
5136:
5132:
5127:
5123:
5119:
5114:
5110:
5106:
5101:
5097:
5092:
5086:
5082:
5077:
5071:
5067:
5066:
5064:
5062:
5058:
5054:
5049:
5041:
5037:
5032:
5030:
5027:
5025:
5024:
5019:
5017:
5014:
5012:
5009:
5007:
5006:
5001:
4999:
4998:
4993:
4991:
4987:
4984:
4981:
4979:
4976:
4974:
4972:
4971:P. t. altaica
4967:
4965:
4963:
4958:
4957:
4944:
4942:9781315425658
4938:
4934:
4930:
4923:
4915:
4908:
4901:
4893:
4889:
4882:
4866:
4862:
4858:
4852:
4845:
4841:
4838:
4833:
4826:
4822:
4819:
4813:
4806:
4802:
4799:
4794:
4786:
4780:
4764:
4758:
4742:
4736:
4720:
4714:
4698:
4692:
4676:
4670:
4654:
4648:
4632:
4626:
4610:
4604:
4588:
4582:
4576:
4575:0-307-26893-4
4572:
4566:
4550:
4546:
4539:
4533:
4529:
4526:
4520:
4514:
4510:
4507:
4501:
4493:
4485:
4478:
4474:
4471:
4470:
4463:
4456:
4452:
4449:
4448:
4441:
4422:
4415:
4408:
4400:
4396:
4391:
4386:
4382:
4378:
4377:
4372:
4365:
4350:on 2011-07-14
4349:
4345:
4339:
4323:
4319:
4312:
4310:
4295:on 2016-03-04
4291:
4284:
4277:
4263:on 2016-10-22
4259:
4252:
4245:
4238:
4234:
4231:
4230:
4223:
4216:
4212:
4209:
4208:
4201:
4194:
4190:
4187:
4186:
4179:
4172:
4168:
4165:
4164:
4157:
4149:
4145:
4141:
4137:
4133:
4129:
4122:
4113:
4108:
4104:
4100:
4096:
4094:
4085:
4077:
4073:
4068:
4063:
4059:
4055:
4051:
4047:
4043:
4036:
4028:
4024:
4020:
4018:9781450358163
4014:
4010:
4006:
4001:
3996:
3992:
3985:
3970:on 2018-01-09
3969:
3965:
3961:
3955:
3940:
3936:
3930:
3922:
3915:
3907:
3905:9780226518237
3901:
3897:
3893:
3886:
3878:
3874:
3867:
3859:
3853:
3848:
3847:
3838:
3836:
3834:
3832:
3830:
3821:
3820:New Scientist
3817:
3810:
3802:
3796:
3793:. CRC Press.
3792:
3785:
3769:
3765:
3761:
3754:
3746:
3744:9781597266093
3740:
3736:
3732:
3728:
3724:
3717:
3708:
3703:
3699:
3695:
3691:
3684:
3676:
3672:
3666:
3652:on 2011-08-17
3651:
3647:
3643:
3639:
3632:
3624:
3617:
3610:
3602:
3598:
3591:
3577:on 2020-02-19
3576:
3572:
3568:
3561:
3553:
3549:
3548:
3540:
3532:
3528:
3521:
3519:
3517:
3515:
3506:
3502:
3498:
3491:
3482:
3477:
3473:
3469:
3465:
3463:
3454:
3445:
3440:
3435:
3430:
3426:
3422:
3418:
3411:
3403:
3399:
3395:
3391:
3387:
3383:
3379:
3375:
3368:
3360:
3353:
3351:
3336:on 2012-04-06
3335:
3331:
3327:
3320:
3312:
3308:
3304:
3300:
3296:
3292:
3284:
3276:
3272:
3267:
3262:
3257:
3252:
3248:
3244:
3240:
3236:
3232:
3225:
3217:
3213:
3206:
3198:
3194:
3190:
3186:
3182:
3178:
3174:
3170:
3162:
3154:
3147:
3138:
3133:
3129:
3125:
3121:
3117:
3113:
3111:
3102:
3094:
3090:
3083:
3081:
3079:
3069:
3064:
3060:
3056:
3052:
3048:
3044:
3042:
3033:
3025:
3024:
3016:
3008:
3004:
3000:
2996:
2992:
2988:
2984:
2982:
2978:
2969:
2967:
2958:
2952:
2948:
2944:
2940:
2934:
2932:
2923:
2917:
2913:
2906:
2904:
2895:
2887:
2880:
2872:
2865:
2857:
2853:
2846:
2844:
2835:
2828:
2821:
2813:
2807:
2803:
2796:
2794:
2785:
2781:
2776:
2771:
2767:
2763:
2759:
2755:
2751:
2744:
2736:
2732:
2727:
2722:
2718:
2714:
2710:
2706:
2702:
2698:
2694:
2690:
2686:
2678:
2670:
2666:
2662:
2658:
2654:
2650:
2646:
2642:
2638:
2634:
2630:
2626:
2619:
2617:
2601:
2600:
2595:
2588:
2580:
2576:
2572:
2568:
2564:
2560:
2559:
2551:
2543:
2539:
2534:
2529:
2524:
2519:
2515:
2511:
2507:
2505:
2495:
2487:
2483:
2478:
2473:
2469:
2465:
2461:
2457:
2453:
2449:
2445:
2438:
2436:
2427:
2421:
2417:
2414:
2410:
2403:
2395:
2389:
2385:
2381:
2374:
2366:
2359:
2351:
2347:
2345:
2341:
2332:
2324:
2320:
2316:
2309:
2301:
2296:
2289:
2281:
2277:
2275:
2266:
2259:(3): 127–133.
2258:
2254:
2247:
2245:
2236:
2232:
2228:
2221:
2219:
2217:
2215:
2213:
2211:
2209:
2207:
2205:
2203:
2201:
2199:
2197:
2195:
2186:
2182:
2177:
2172:
2168:
2161:
2159:
2153:
2147:
2145:
2143:
2141:
2132:
2128:
2123:
2118:
2113:
2108:
2104:
2100:
2096:
2092:
2088:
2081:
2079:
2077:
2075:
2066:
2062:
2058:
2054:
2050:
2046:
2038:
2036:
2019:
2012:
1996:
1989:
1974:
1970:
1966:
1959:
1950:
1945:
1941:
1937:
1936:
1931:
1929:
1920:
1918:
1916:
1899:
1892:
1885:
1883:
1881:
1872:
1865:
1857:
1855:
1850:
1840:
1837:
1835:
1832:
1830:
1827:
1822:
1821:
1817:
1815:
1814:
1810:
1808:
1807:
1802:
1801:
1799:
1798:
1790:
1787:
1785:
1782:
1780:
1779:Bornean tiger
1777:
1775:
1772:
1771:
1770:
1767:
1763:
1760:
1758:
1757:Malayan tiger
1755:
1753:
1750:
1748:
1747:Caspian tiger
1745:
1743:
1740:
1739:
1738:
1735:
1734:
1733:
1730:
1729:
1722:
1719:
1717:
1713:
1707:
1702:
1697:
1693:
1689:
1685:
1681:
1677:
1672:
1670:
1666:
1662:
1654:
1650:
1641:
1639:
1634:
1630:
1626:
1622:
1617:
1614:
1610:
1606:
1604:
1600:
1596:
1591:
1589:
1585:
1576:
1571:
1567:
1565:
1561:
1555:
1543:
1539:
1534:
1525:
1523:
1518:
1516:
1508:
1503:
1494:
1492:
1488:
1484:
1479:
1477:
1473:
1469:
1464:
1459:
1457:
1452:
1451:Taukum Desert
1448:
1444:
1443:Lake Balkhash
1440:
1431:
1427:
1424:
1417:
1413:
1409:
1396:
1392:
1387:
1383:
1381:
1377:
1373:
1367:
1365:
1361:
1357:
1352:
1349:In 1992, the
1347:
1344:
1340:
1330:
1328:
1323:
1321:
1311:
1309:
1305:
1300:
1298:
1294:
1284:
1281:
1278:
1276:
1271:
1267:
1263:
1229:
1222:
1208:
1205:
1201:
1197:
1196:Eurasian lynx
1192:
1188:
1186:
1182:
1178:
1173:
1170:
1165:
1162:
1161:Amur leopards
1158:
1154:
1148:
1146:
1141:
1139:
1135:
1130:
1126:
1122:
1118:
1114:
1110:
1102:
1097:
1089:
1080:
1076:
1073:
1072:radio-collars
1068:
1065:
1061:
1057:
1053:
1049:
1045:
1041:
1037:
1033:
1029:
1025:
1021:
1020:Cervus nippon
1017:
1013:
1009:
1005:
1001:
997:
993:
989:
985:
981:
977:
973:
965:
961:
956:
947:
945:
941:
940:Amur leopards
936:
933:
928:
923:
919:
910:
901:
899:
895:
891:
887:
883:
879:
874:
869:
867:
861:
859:
855:
851:
847:
843:
839:
835:
830:
828:
824:
820:
816:
812:
808:
804:
799:
797:
793:
789:
785:
781:
777:
767:
760:
746:
732:
728:
724:
718:
713:
704:
702:
697:
693:
688:
686:
682:
678:
674:
664:
661:
660:Sungari River
657:
652:
650:
646:
641:
636:
635:with people.
634:
630:
625:
624:Bengal tigers
619:
610:
608:
604:
596:
591:
582:
580:
575:
573:
569:
565:
561:
557:
552:
548:
542:
540:
536:
531:
526:
522:
517:
515:
510:
506:
501:
493:
492:P. t. altaica
489:
488:P. t. virgata
485:
480:
471:
469:
464:
463:populations.
462:
458:
454:
450:
446:
442:
438:
434:
430:
426:
422:
421:
416:
412:
408:
404:
399:
391:
388:
385:
381:
378:
375:
371:
367:
364:
361:
357:
353:
350:
349:
348:
346:
345:Carl Linnaeus
342:
338:
330:
321:
319:
315:
311:
306:
304:
300:
296:
292:
288:
284:
283:Caspian tiger
279:
277:
272:
268:
264:
260:
257:and possibly
256:
252:
248:
247:
243:
240:
236:
232:
209:
205:
200:
196:
192:
189:
188:
184:
183:
178:
175:
174:
167:
166:
161:
158:
157:
154:
153:
149:
146:
145:
142:
139:
136:
135:
132:
129:
126:
125:
122:
119:
116:
115:
112:
109:
106:
105:
102:
99:
96:
95:
92:
89:
86:
85:
82:
79:
76:
75:
72:
69:
66:
65:
60:
55:
51:
48:
43:
39:
34:
29:
26:
22:
5060:
5039:
5022:
5004:
4996:
4970:
4961:
4932:
4922:
4916:(681): 1–18.
4913:
4900:
4891:
4881:
4869:. Retrieved
4865:the original
4860:
4851:
4832:
4812:
4793:
4779:
4767:. Retrieved
4757:
4745:. Retrieved
4735:
4723:. Retrieved
4713:
4701:. Retrieved
4691:
4679:. Retrieved
4669:
4657:. Retrieved
4647:
4635:. Retrieved
4625:
4613:. Retrieved
4603:
4591:. Retrieved
4581:
4565:
4553:. Retrieved
4549:the original
4538:
4519:
4500:
4484:
4468:
4462:
4446:
4440:
4428:. Retrieved
4421:the original
4407:
4380:
4374:
4364:
4352:. Retrieved
4348:the original
4338:
4326:. Retrieved
4322:the original
4297:. Retrieved
4290:the original
4276:
4265:. Retrieved
4258:the original
4244:
4228:
4222:
4206:
4200:
4184:
4178:
4162:
4156:
4131:
4127:
4121:
4102:
4098:
4092:
4084:
4049:
4045:
4035:
3990:
3984:
3972:. Retrieved
3968:the original
3963:
3954:
3942:. Retrieved
3938:
3929:
3920:
3914:
3895:
3885:
3876:
3866:
3845:
3819:
3809:
3790:
3784:
3772:. Retrieved
3768:the original
3763:
3753:
3735:Island Press
3726:
3716:
3697:
3693:
3683:
3674:
3665:
3654:. Retrieved
3650:the original
3641:
3631:
3622:
3609:
3600:
3590:
3579:. Retrieved
3575:the original
3570:
3560:
3552:the original
3546:
3539:
3530:
3504:
3500:
3490:
3471:
3467:
3461:
3453:
3424:
3420:
3410:
3377:
3373:
3367:
3358:
3338:. Retrieved
3334:the original
3329:
3319:
3294:
3290:
3283:
3238:
3234:
3224:
3215:
3205:
3172:
3168:
3161:
3155:(61): 18−19.
3152:
3146:
3119:
3115:
3109:
3101:
3092:
3050:
3046:
3040:
3032:
3022:
3015:
2990:
2986:
2980:
2976:
2946:
2942:
2911:
2902:
2894:
2879:
2864:
2855:
2833:
2820:
2801:
2757:
2753:
2743:
2692:
2688:
2677:
2636:
2632:
2628:
2624:
2603:. Retrieved
2597:
2587:
2562:
2556:
2550:
2516:(12): e442.
2513:
2510:PLOS Biology
2509:
2503:
2494:
2451:
2447:
2416:
2412:
2402:
2383:
2373:
2364:
2358:
2349:
2343:
2340:Felis tigris
2339:
2331:
2322:
2318:
2308:
2298:
2288:
2279:
2274:Felis tigris
2273:
2265:
2256:
2252:
2234:
2230:
2169:(152): 1–8.
2166:
2157:
2097:(1): e4125.
2094:
2090:
2048:
2044:
2022:. Retrieved
2011:
1999:. Retrieved
1988:
1976:. Retrieved
1968:
1958:
1939:
1933:
1927:
1902:. Retrieved
1898:the original
1870:
1829:Amur leopard
1818:
1811:
1804:
1742:Bengal tiger
1720:
1709:
1694:for 'King' (
1673:
1658:
1633:Heilongjiang
1618:
1607:
1592:
1587:
1580:
1557:
1554:Tiger attack
1537:
1519:
1511:
1497:In captivity
1487:Kolyma River
1480:
1460:
1456:Bukhara deer
1436:
1419:
1379:
1375:
1371:
1368:
1363:
1359:
1355:
1350:
1348:
1343:range states
1336:
1333:Conservation
1324:
1317:
1301:
1290:
1282:
1279:
1259:
1199:
1193:
1189:
1185:Sikhote-Alin
1180:
1174:
1166:
1149:
1142:
1120:
1112:
1109:Ussuri brown
1106:
1077:
1069:
1044:Ursus arctos
1043:
1035:
1027:
1019:
1011:
1003:
995:
987:
979:
969:
943:
937:
921:
915:
870:
862:
831:
800:
796:Yellow River
773:
729:
725:
723:than black.
721:
707:Fur and coat
689:
670:
653:
644:
639:
637:
620:
616:
599:
576:
571:
563:
559:
546:
543:
535:Central Asia
530:phylogenetic
518:
497:
491:
487:
468:P. t. tigris
467:
465:
448:
433:P. t. tigris
432:
418:
414:
395:
389:
383:
380:Felis tigris
379:
368:proposed by
365:
354:proposed by
351:
343:proposed by
337:Felis tigris
336:
335:
307:
280:
267:Sikhote-Alin
244:
234:
230:
228:
194:
190:Population:
180:
176:Subspecies:
163:
151:
25:
5152:iNaturalist
5085:Wikispecies
3877:Planet Save
3764:Ecology.com
3427:(1): 1–11.
3380:: 120–128.
2024:18 December
1784:Javan tiger
1684:Nani people
1584:Bikin River
1536:Sign warns
1391:Yerevan Zoo
1314:In the past
1308:snare traps
1181:Canis lupus
1145:hibernation
1040:brown bears
1004:Alces alces
932:home ranges
892:and Lesser
827:Korean pine
780:Lake Baikal
764:A tiger in
685:occipitalis
683:and crista
595:Münster Zoo
445:South China
441:Indochinese
411:Pleistocene
259:North Korea
141:Pantherinae
137:Subfamily:
5265:Categories
4997:Amur tiger
4299:2014-10-03
4267:2011-10-06
4000:1805.05356
3700:(4): 323.
3656:2015-07-15
3581:2014-09-17
3434:1810.11195
3340:2018-03-31
2717:2263/32583
2605:4 November
2325:: 421–519.
1845:References
1774:Bali tiger
1644:In culture
1638:Zürich Zoo
1552:See also:
1318:After the
1101:brown bear
1028:Sus scrofa
858:Amur moose
811:Amur River
607:carnassial
568:stochastic
314:Manchurian
242:subspecies
235:Amur tiger
121:Feliformia
117:Suborder:
3974:5 January
3944:5 January
3507:(2): 159.
2947:The Tiger
2943:Der Tiger
2939:Mazák, V.
2344:amurensis
2152:Mazák, V.
1655:in Russia
1560:man-eater
1439:Ili River
1423:Amu-Darya
1200:Lynx lynx
1153:telemetry
1058:and even
1024:wild boar
1016:sika deer
972:ungulates
890:Wandashan
850:Sika deer
696:Kopet Dag
692:Turkestan
656:Manchuria
613:Body size
603:zygomatic
556:gene flow
509:haplotype
494:) tigers.
474:Phylogeny
429:Sundaland
407:gene flow
384:amurensis
312:tiger", "
299:Silk Road
165:P. tigris
159:Species:
111:Carnivora
77:Kingdom:
71:Eukaryota
5209:14000261
5070:Wikidata
4986:Archived
4840:Archived
4821:Archived
4801:Archived
4769:6 August
4747:6 August
4725:6 August
4703:6 August
4681:6 August
4659:6 August
4637:6 August
4615:6 August
4593:6 August
4555:6 August
4528:Archived
4509:Archived
4473:Archived
4451:Archived
4399:15879196
4354:6 August
4328:6 August
4233:Archived
4211:Archived
4189:Archived
4167:Archived
4148:88992035
4076:22393520
4027:21682090
3402:90802525
3275:30891189
3197:10597364
3153:Cat News
3007:85177441
2987:Mammalia
2941:(1983).
2901:"Tiger,
2784:30482605
2735:24045858
2695:: 2433.
2669:25766120
2661:19555412
2579:37492591
2542:15583716
2486:26601191
2154:(1981).
2131:19142238
2091:PLOS ONE
2065:27136188
1871:Cat News
1725:See also
1629:Shenzhen
1625:Shanghai
1449:and the
1304:poaching
1275:poachers
1230:, Russia
974:such as
834:ungulate
821:and the
788:Holocene
776:Mongolia
551:Bayesian
486:(PTV or
398:validity
339:was the
324:Taxonomy
318:Ussurian
152:Panthera
127:Family:
101:Mammalia
91:Chordata
87:Phylum:
81:Animalia
67:Domain:
5144:5219420
5131:1253183
4861:Reuters
4376:Science
4067:3287338
3892:"Tiger"
3774:June 7,
3382:Bibcode
3299:Bibcode
3266:6405893
3243:Bibcode
3177:Bibcode
3124:Bibcode
3055:Bibcode
2762:Bibcode
2726:3778509
2697:Bibcode
2641:Bibcode
2629:ex situ
2625:In situ
2477:4640610
2456:Bibcode
2342:, var.
2300:Siebold
2227:"Tiger"
2185:3504004
2122:2624500
2099:Bibcode
2001:13 June
1978:June 7,
1904:16 June
1696:Chinese
1611:at the
1595:Hunchun
1542:Russian
1491:Yakutia
1485:in the
1395:Armenia
1287:Threats
1052:rabbits
962:at the
960:diorama
673:canines
658:by the
572:in situ
564:in situ
560:ex situ
547:ex situ
500:genetic
484:Caspian
437:Malayan
423:in the
303:Siberia
147:Genus:
131:Felidae
107:Order:
97:Class:
5271:Tigers
5235:156823
5183:726472
5170:121116
5157:130796
5076:Q69581
4939:
4871:4 July
4573:
4397:
4146:
4074:
4064:
4025:
4015:
3939:Rgo.ru
3902:
3854:
3797:
3741:
3400:
3273:
3263:
3195:
3005:
2953:
2918:
2808:
2782:
2733:
2723:
2667:
2659:
2577:
2540:
2533:534810
2530:
2484:
2474:
2422:
2390:
2183:
2129:
2119:
2063:
1708::
1706:pinyin
1698::
1688:Manchu
1575:Korean
1468:Tehran
1414:, and
1266:Litter
1115:) and
1060:salmon
1038:) and
1014:) and
701:Nagina
675:. The
579:clades
220:
214:
5248:19403
5222:74533
5196:15956
5118:5K5M2
4910:(PDF)
4430:5 May
4424:(PDF)
4417:(PDF)
4293:(PDF)
4286:(PDF)
4261:(PDF)
4254:(PDF)
4144:S2CID
4023:S2CID
3995:arXiv
3619:(PDF)
3597:"One"
3501:Ursus
3429:arXiv
3398:S2CID
3193:S2CID
3003:S2CID
2945:[
2908:(PDF)
2830:(PDF)
2665:S2CID
2575:S2CID
2233:[
2181:JSTOR
2163:(PDF)
1894:(PDF)
1867:(PDF)
1680:Udege
1621:Anhui
1297:genes
1138:groin
1129:bears
1056:pikas
1048:hares
1000:moose
922:adult
898:Feces
823:taiga
677:skull
667:Skull
453:Javan
382:var.
360:Altai
295:Gansu
239:tiger
5217:NCBI
5191:IUCN
5178:ITIS
5139:GBIF
4937:ISBN
4873:2020
4771:2011
4749:2011
4727:2011
4705:2011
4683:2011
4661:2011
4639:2011
4617:2011
4595:2011
4571:ISBN
4557:2011
4432:2013
4395:PMID
4356:2011
4330:2011
4072:PMID
4013:ISBN
3976:2018
3946:2018
3900:ISBN
3852:ISBN
3795:ISBN
3776:2015
3739:ISBN
3425:2019
3271:PMID
2951:ISBN
2916:ISBN
2806:ISBN
2780:PMID
2731:PMID
2657:PMID
2607:2014
2538:PMID
2482:PMID
2420:ISBN
2388:ISBN
2127:PMID
2061:PMID
2026:2015
2003:2015
1980:2015
1940:2022
1906:2012
1711:Wáng
1682:and
1674:The
1665:Amur
1627:and
1474:and
1378:and
1177:wolf
1136:and
1134:hams
1064:Scat
856:and
786:and
459:and
457:Bali
396:The
310:Amur
229:The
5204:MSW
5165:ISC
5126:EoL
5113:CoL
5100:ADW
4385:doi
4381:308
4136:doi
4107:doi
4062:PMC
4054:doi
4005:doi
3702:doi
3698:276
3476:doi
3439:doi
3390:doi
3307:doi
3295:261
3261:PMC
3251:doi
3185:doi
3132:doi
3063:doi
2995:doi
2770:doi
2721:PMC
2713:hdl
2705:doi
2649:doi
2567:doi
2528:PMC
2518:doi
2472:PMC
2464:doi
2171:doi
2117:PMC
2107:doi
2053:doi
1944:doi
1597:in
1270:den
1022:),
1006:),
998:),
990:),
982:),
233:or
5267::
5245::
5232::
5219::
5206::
5193::
5180::
5167::
5154::
5141::
5128::
5115::
5102::
5087::
5072::
5038:.
4912:.
4890:.
4859:.
4393:.
4379:.
4373:.
4308:^
4142:.
4132:34
4130:.
4103:91
4101:.
4097:.
4070:.
4060:.
4048:.
4044:.
4021:.
4011:.
4003:.
3962:.
3937:.
3894:.
3875:.
3828:^
3818:.
3762:.
3733::
3729:.
3696:.
3692:.
3673:.
3640:.
3621:.
3599:.
3569:.
3513:^
3505:14
3503:.
3499:.
3472:94
3470:.
3466:.
3437:.
3423:.
3419:.
3396:.
3388:.
3378:92
3376:.
3349:^
3305:.
3293:.
3269:.
3259:.
3249:.
3237:.
3233:.
3214:.
3191:.
3183:.
3173:31
3171:.
3130:.
3120:43
3118:.
3114:.
3077:^
3061:.
3051:22
3049:.
3045:.
3043:)"
3001:.
2991:31
2989:.
2985:.
2965:^
2930:^
2910:.
2842:^
2792:^
2778:.
2768:.
2758:28
2756:.
2752:.
2729:.
2719:.
2711:.
2703:.
2691:.
2687:.
2663:.
2655:.
2647:.
2637:18
2635:.
2615:^
2596:.
2573:.
2561:.
2536:.
2526:.
2512:.
2508:.
2506:)"
2480:.
2470:.
2462:.
2452:11
2450:.
2446:.
2434:^
2348:.
2323:58
2321:.
2317:.
2278:.
2257:28
2255:.
2243:^
2229:.
2193:^
2179:.
2165:.
2139:^
2125:.
2115:.
2105:.
2093:.
2089:.
2073:^
2059:.
2049:11
2047:.
2034:^
1971:.
1967:.
1938:.
1932:.
1914:^
1879:^
1869:.
1853:^
1718:.
1704:;
1671:.
1548:).
1544::
1478:.
1410:,
1393:,
1140:.
1062:.
1054:,
1050:,
958:A
896:.
888:,
848:.
840:,
574:.
523:,
470:.
455:,
443:,
439:,
253:,
5042:.
4973:)
4964:)
4945:.
4875:.
4773:.
4751:.
4729:.
4707:.
4685:.
4663:.
4641:.
4619:.
4597:.
4559:.
4434:.
4401:.
4387::
4358:.
4332:.
4302:.
4270:.
4150:.
4138::
4115:.
4109::
4078:.
4056::
4050:1
4029:.
4007::
3997::
3978:.
3948:.
3908:.
3879:.
3860:.
3822:.
3803:.
3778:.
3747:.
3710:.
3704::
3659:.
3584:.
3484:.
3478::
3447:.
3441::
3431::
3404:.
3392::
3384::
3343:.
3313:.
3309::
3301::
3277:.
3253::
3245::
3239:9
3218:.
3199:.
3187::
3179::
3140:.
3134::
3126::
3071:.
3065::
3057::
3009:.
2997::
2983:"
2959:.
2924:.
2888:.
2873:.
2814:.
2786:.
2772::
2764::
2737:.
2715::
2707::
2699::
2693:4
2671:.
2651::
2643::
2609:.
2581:.
2569::
2563:5
2544:.
2520::
2514:2
2488:.
2466::
2458::
2428:.
2396:.
2346:"
2338:"
2276:"
2272:"
2187:.
2173::
2160:"
2156:"
2133:.
2109::
2101::
2095:4
2067:.
2055::
2028:.
2005:.
1982:.
1952:.
1946::
1930:"
1926:"
1908:.
1701:王
1540:(
1397:.
1198:(
1179:(
1119:(
1111:(
1042:(
1034:(
1026:(
1018:(
1010:(
1002:(
994:(
986:(
978:(
942:(
376:.
297:−
23:.
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.