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Tide pool

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the intertidal as juvenile habitat, feeding or refuge ground, or as transient space between nearshore areas. Tidepools fishes can be classified as residents and non-residents (sometimes called transients or visitors). Residents are those that spend the whole lifetime in the tidepools. Non-resident species are commonly divided into two groups: secondary residents (also known as partial residents or opportunists) and transients (which can be further classified as tidal and seasonal transients). Secondary residents are species that spend only a portion of their life history in tidepools, typically during their juvenile stage, before moving on to adult subtidal habitats. Transients, on the other hand, may temporarily inhabit tidepools for various reasons such as foraging, seeking refuge, or transit. Unlike residents, transients lack specialized adaptations for intertidal life and typically occupy large tidepools for a relatively short period, ranging from a single tidal cycle to a few months. The
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Many fish species can live in tidepools. Tidepool fishes are those inhabiting the intertidal zone during part or the entirety of their life cycle, including residents displaying morphological, physiological and behavioral adaptations to withstand the fluctuating environment and non-residents that use
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Andrades, Ryan; González-Murcia, Saúl; Buser, Thaddaeus J.; Macieira, Raphael M.; Andrade, Juliana M.; Pinheiro, Hudson T.; Vilar, Ciro C.; Pimentel, Caio R.; Gasparini, João L.; Quintão, Thaís L.; Machado, Fabíola S.; Castellanos-Galindo, Gustavo; Ruiz-Campos, Gorgonio; Ojeda, F. Patricio; Martin,
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This zone is constantly covered and uncovered by water, so its inhabitants have adapted to surviving in these conditions. More plants and animals live here, compared to the high tide zone, because they are not exposed to drying conditions for so long. During low tide, anemones close up and mussels
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due to lack of water and evaporation. Many forms of the Coralline algae bring herbivores, such as mollusks "Notoacmea", to the tide pools during high tides, increasing the biomass of the area. Once low tides comes, these herbivores are exposed to carnivores in the areas, fueling the food web.
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such as fish. Seaweeds provide shelter for many animals, like sea slugs and urchins that are too fragile for other zones. These creatures can grow to larger sizes because there is more available energy and better water coverage: the water is shallow enough to allow additional sunlight for
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This area is mostly submerged and is exposed only during unusually low tide. It usually teems with life and has far more marine vegetation, especially seaweeds. Organisms in this zone do not have to be as well adapted to drying out and temperature extremes. Low tide zone organisms include
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live at very tightly constrained elevations, with tidal conditions precisely determining the exact height of an assemblage relative to sea level. The intertidal zone is periodically exposed to sun and wind, conditions that can cause barnacles to become
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is a species of fish that is named for its tide pool habitat. The Tidepool Sculpin has been found to show preferences for certain tide pools and will return to their tide pool of choice after being removed from it. This is a behavior known as
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grows in greater numbers when such competition exists; a control group with no competition produced fewer offspring than an experimental group with mussels; from this it is thought that the mussels provide protection for the developing
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Mendonça, Vanessa; Madeira, Carolina; Dias, Marta; Vermandele, Fanny; Archambault, Philippe; Dissanayake, Awantha; Canning-Clode, João; Flores, Augusto A. V.; Silva, Ana; Vinagre, Catarina (July 5, 2018). Hewitt, Judi (ed.).
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lost arms. Most species must retain an intact central part of the body to be able to regenerate, but a few can regrow from a single ray. The regeneration of these stars is possible because the vital organs are in the arms.
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increases the supply of inorganic nitrogen and phosphorus in coastal marine tide pools which allows the ecosystem the nutrients to be more productive. The shell of a California mussel is primarily composed of
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often engages in territorial fights. The white tentacles (acrorhagi), which contain stinging cells, are for fighting. The sea anemones sting each other repeatedly until one of them moves.
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The rocky shoreline exhibits distinct zones with unique characteristics. These zones are created by the tidal movements of water along the rocky shores from high to low-tide. They are:
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The intertidal or littoral zone: area between the high and low-tide marks. Can be further divided into high, mid, and low intertidal zones, which are explained below in more depth.
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The presence and abundance of flora and fauna vary between zones along the rocky shore. This is due to niche adaptations in response to the varying tides and solar exposure.
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Tide pools are often surrounded by coastal predators who feed on tide pool flora and fauna. These predators play an important role in the tide pool
2121:"The importance of form: Differences in competitive ability, resistance to consumers and environmental stress in an assemblage of coralline algae" 2695: 855:. Climate change and ocean acidification has led to a decrease in these amounts important compounds in California Mussel shells over many years. 4866: 4119: 917:. These fish crawl on the floor of tide pools using a back and forth movement of their tail fin and a rotating motion of their pectoral fins. 2160: 1205: 2187: 816: 73:. These pools typically range from a few inches to a few feet deep and a few feet across. Many of these pools exist as separate bodies of 5010: 3269: 2535: 5080: 2280: 1865:"Temporal variation of rocky intertidal resident fish assemblages - patterns and possible mechanisms with a note on sampling protocols" 1259: 1058: 769:
are known to sometimes feast on intertidal creatures at low tide. Although tide pool organisms must avoid getting washed away into the
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Paine, R.T. (December 1998). "Habitat Suitability and Local Population Persistence of the Sea Palm Postelsia Palmaeformis".
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Gibson, R. N.; Yoshiyama, R. M. (January 1, 1999), Horn, Michael H.; Martin, Karen L. M.; Chotkowski, Michael A. (eds.),
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of the sun and moon. A tidal cycle is usually about 25 hours and consists of one or two high tides and two low tides.
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which they slide across their mouth opening when not feeding. These plates also protect against predation.
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Bullard, Elizabeth M.; Torres, Ivan; Ren, Tianqi; Graeve, Olivia A.; Roy, Kaustuv (January 19, 2021).
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close their shells to keep in moisture. They reopen when the tide returns and brings them food.
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The supralittoral zone or splash zone: area above the high-tide mark, which is virtually a
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Yoshikawa, Akihiro; Goto, Ryutaro; Yasuda, Chiaki I; Asakura, Akira (July 21, 2020).
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Coralline algae "Corallinales" are predominant features of mid and low intertidal
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and other microorganism are the food sources that attract them to the tide pools.
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Rocky pool on a seashore, separated from the sea at low tide, filled with seawater
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Carvalho, Jessica; Mendonça, Vanessa; Vinagre, Catarina; Silva, Ana (June 2021).
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to break them open. Sea stars prey on mussels and are eaten by gulls themselves.
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during each high tide, which occurs once or twice daily. Organisms must survive
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environment. Occasionally gets splashed, but never gets covered by the ocean.
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US Department of Commerce, National Oceanic and Atmospheric Administration.
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are commonly found in tide pool environments. The long-wristed hermit crab (
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Tide pools in Santa Cruz, California from spray/splash zone to low tide zone
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Horn, Michael H.; Martin, Karen L. M.; Chotkowski, Michael A. (1999).
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Bressman, Noah R.; Gibb, Alice C.; Farina, Stacy C. (December 2018).
1033: 970: 939: 934: 921: 844: 744: 720: 552: 314: 201: 173: 3247: 2097: 1929: 1739: 819:") move around tide pools with tube like feet. Different species of 6018: 5847: 5837: 5812: 5782: 5762: 5742: 5712: 5586: 5515: 5046: 4874: 4553: 4465: 4412: 4367: 3865: 3860: 3687: 3602: 3461: 3286: 3079: 2889: 2435: 2430: 2355: 1608: 1263: 1115: 866: 862: 774: 594: 548: 334: 310: 221: 185: 97: 78: 63: 48: 6112: 5576: 5383: 5253: 5248: 4875: 4823: 4483: 4128: 4001: 3875: 3720: 3596: 3591: 3501: 3496: 3486: 3406: 3089: 3049: 3019: 2997: 2617: 2405: 2305: 964:. Alternatively, the mussels may prevent the growth of competing 943: 878: 869: 848: 513: 373: 357: 293: 285: 237: 209: 89: 82: 2838: 2208: 5970: 5952: 5696: 4026: 4021: 3556: 3536: 3491: 3476: 3334: 3074: 2400: 858: 740: 724: 394: 297: 225: 161:
The sublittoral or subtidal zone: area below the low-tide mark.
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reproduces clones of itself through a process of longitudinal
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to grow freely after wave action has eliminated the mussels.
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have different colors, and many are seen in tide pools. With
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Tide pools exist in the intertidal zone (the area within the
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Thomson, Donald A.; Lehner, Charles E. (April 1, 1976).
1397:. Dana Krempels, Ph.D. September 5, 2008. Archived from 288:, anemones, brown seaweed, chitons, crabs, green algae, 1483: 1375:. British Marine Life Study Society. September 5, 2008 155:
The intertidal fringe: area around the high-tide mark.
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Tide pools provide a home for many organisms such as
1779: 313:. This area is also relatively protected from large 1591:Gilliand, Sarah; Pechenik, Jan A. (December 2018). 1387: 2125:Journal of Experimental Marine Biology and Ecology 2056:Journal of Experimental Marine Biology and Ecology 1950: 1830:Journal of Experimental Marine Biology and Ecology 1918:Journal of the Fisheries Research Board of Canada 1863:Almada, VĂ­tor C.; Faria, Cláudia (June 1, 2004). 1365: 1200: 1198: 1196: 1194: 1192: 1190: 1188: 1186: 6166: 2038: 1590: 1700:, San Diego: Academic Press, pp. 264–296, 1691: 1490:Proceedings of the National Academy of Sciences 757:can dislodge mussels and draw them out to sea. 30:"Rockpool" redirects here. For other uses, see 1252: 1183: 731:. Inhabitants deal with a frequently changing 317:because of the wave action and shallow water. 5691: 5677: 4860: 4113: 3263: 2471:Stable isotope analysis in aquatic ecosystems 2181: 1823: 881:shells are impermeable, and they possess two 700: 1100:Temporary tide pool at an extreme low tide, 2536:Freshwater environmental quality parameters 1862: 1348: 1342: 5684: 5670: 5081:Latitudinal gradients in species diversity 4867: 4853: 4120: 4106: 3270: 3256: 2188: 2174: 2044: 707: 693: 545:        1888: 1663: 1527: 1509: 1415:. California Tide Pools. October 1, 2014. 1325: 1307: 4979:Predator–prey (Lotka–Volterra) equations 4618:Tritrophic interactions in plant defense 1557:Annual Review of Ecology and Systematics 887: 751:, exposure to midday sun and predators. 324: 267: 135: 38: 5011:Random generalized Lotka–Volterra model 2118: 2075: 1782:"Intertidal Fishes: life in two worlds" 1550: 1425: 615:Hydrothermal vent microbial communities 309:activity, with almost normal levels of 14: 6167: 4819:Herbivore adaptations to plant defense 2195: 1479: 1477: 1118:and create competition for resources. 5665: 4848: 4101: 3277: 3251: 2724: 2723: 2207: 2169: 2081: 1911: 1869:Reviews in Fish Biology and Fisheries 1747:Reviews in Fish Biology and Fisheries 1735: 1733: 1731: 1687: 1685: 1683: 827:, some filled with poison like with " 4834:Predator avoidance in schooling fish 2119:Padilla, Dianna K. (July 24, 1984). 1229: 1227: 1225: 1223: 1159: 1157: 1109: 777:that can vary based on the climate. 392:       300:, mussels, and sometimes even small 184:or being exposed to cold winds. Few 172:), which is submerged by the sea at 5284:Intermediate disturbance hypothesis 2799:Oceanic physical-biological process 2661:List of freshwater ecoregions (WWF) 2045:Blanchette, Carol A. (April 1996). 1570:10.1146/annurev.es.03.110172.001125 1474: 1266:. September 5, 2008. Archived from 1212:. September 5, 2008. Archived from 1133:List of British Isles rockpool life 865:live in the splash zone. Different 188:can survive such harsh conditions. 24: 5037:Ecological effects of biodiversity 3966:Integrated coastal zone management 1790:10.1016/b978-0-12-356040-7.x5000-8 1728: 1694:"13 - Intertidal Fish Communities" 1680: 25: 6211: 4373:Generalist and specialist species 2154: 1349:Andy Horton (September 5, 2008). 1220: 1154: 1088:engaged in a battle for territory 191: 92:are home to especially adaptable 5871: 5096:Occupancy–abundance relationship 4082: 4081: 3619: 3612: 3232: 3231: 3219: 3205: 2837: 2509: 2326:Colored dissolved organic matter 1912:Green, John M. (March 1, 1971). 1706:10.1016/b978-012356040-7/50014-7 1093: 1075: 1070:consuming a mussel in tide pools 1057: 1040: 1009: 674: 364: 278: 259: 256:and other intertidal organisms. 5116:Relative abundance distribution 4829:Plant defense against herbivory 4696:Competitive exclusion principle 4408:Mesopredator release hypothesis 2671:Latin America and the Caribbean 2161:Tidal swimming pools in Britain 2112: 1991: 1944: 1905: 1856: 1817: 1773: 1639: 1584: 1544: 1419: 1024:consuming a by-the-wind-sailor 955:Recent studies have shown that 920:Multiple species of Amphipods ( 4701:Consumer–resource interactions 4064:Region of freshwater influence 3153:Ecological values of mangroves 2696:North Pacific Subtropical Gyre 1426:Pfister, Catherine A. (2007). 1405: 1274: 807:Some species of sea stars can 320: 119:The Log from the Sea of Cortez 13: 1: 6200:Coastal and oceanic landforms 5547:Biological data visualization 5374:Environmental niche modelling 5101:Population viability analysis 3826:Large-scale coastal behaviour 3658:Beaches in estuaries and bays 1741:Karen L. (December 1, 2023). 1652:Journal of Crustacean Biology 1148: 1143:Sydney inter-tidal rock pools 851:which are both polymorphs of 5032:Density-dependent inhibition 3173:Marine conservation activism 3158:Fisheries and climate change 2137:10.1016/0022-0981(84)90213-2 2069:10.1016/0022-0981(95)00141-7 2024:10.1016/j.seares.2021.102035 1842:10.1016/0022-0981(76)90106-4 1309:10.1371/journal.pone.0200066 1260:"Botanical Beach Tide Pools" 938:) look similar to miniature 273:Low tide zone in a tide pool 7: 5501:Liebig's law of the minimum 5336:Resource selection function 4227:Metabolic theory of ecology 3168:Human impact on marine life 3045:Davidson Seamount § Ecology 2281:Aquatic population dynamics 1792:– via Academic Press. 1121: 892:Hermit crabs in a tide pool 352:Part of a series related to 329:The site of a tide pool in 10: 6216: 5401:Niche apportionment models 5121:Relative species abundance 4325:Primary nutritional groups 4222:List of feeding behaviours 1969:10.1016/j.zool.2018.10.003 1759:10.1007/s11160-023-09798-z 1052:at low tide in a tide pool 129: 29: 6055: 6027: 5989: 5951: 5880: 5869: 5703: 5650: 5582:Ecosystem based fisheries 5524: 5424: 5349: 5222: 5194:Interspecific competition 5159: 5086:Minimum viable population 5019: 4944:Maximum sustainable yield 4929:Intraspecific competition 4924:Effective population size 4887: 4804:Anti-predator adaptations 4789: 4668: 4595: 4552: 4474: 4441: 4338: 4315:Photosynthetic efficiency 4250: 4144: 4077: 3979: 3946: 3751: 3711: 3628: 3610: 3285: 3199: 3138: 3010: 2946: 2908: 2855: 2846: 2835: 2784:Marine primary production 2736: 2732: 2719: 2678:List of marine ecoregions 2653: 2518: 2507: 2221: 2217: 2203: 1881:10.1007/s11160-004-6750-7 791:Anthopleura elegantissima 743:content. Hazards include 32:Rockpool (disambiguation) 5863:Waste stabilization pond 5572:Ecological stoichiometry 5537:Alternative stable state 3771:Coastal biogeomorphology 3030:Coastal biogeomorphology 3025:Marine coastal ecosystem 927: 780: 735:: fluctuations in water 125: 66:that forms on the rocky 5416:Ontogenetic niche shift 5279:Ideal free distribution 5189:Ecological facilitation 4939:Malthusian growth model 4909:Consumer-resource model 4766:Paradox of the plankton 4731:Energy systems language 4451:Chemoorganoheterotrophy 4418:Optimal foraging theory 4393:Heterotrophic nutrition 2938:Paradox of the plankton 2749:Diel vertical migration 2643:Freshwater swamp forest 2361:GIS and aquatic science 2209:General components and 2004:Journal of Sea Research 1597:The Biological Bulletin 1511:10.1073/pnas.2004769118 1395:"Biology: Regeneration" 5562:Ecological forecasting 5506:Marginal value theorem 5304:Landscape epidemiology 5239:Cross-boundary subsidy 5174:Biological interaction 4524:Microbial intelligence 4212:Green world hypothesis 3668:Coastal morphodynamics 2764:Large marine ecosystem 2456:Shoaling and schooling 2049:Postelsia palmaeformis 1805:Cite journal requires 1665:10.1093/jcbiol/ruaa039 1264:British Columbia Parks 1216:on September 24, 2008. 1166:"What is a tide pool?" 1049:Postelsia palmaeformis 893: 681:Marine life portal 346: 331:Santa Cruz, California 275: 196:The high tide zone is 141: 51: 5818:Salt evaporation pond 5567:Ecological humanities 5466:Ecological energetics 5411:Niche differentiation 5274:Habitat fragmentation 5042:Ecological extinction 4989:Small population size 4741:Feed conversion ratio 4721:Ecological succession 4653:San Francisco Estuary 4567:Ecological efficiency 4509:Microbial cooperation 4059:Physical oceanography 3183:Marine protected area 3110:Salt pannes and pools 2885:Marine larval ecology 2860:Census of Marine Life 2744:Deep scattering layer 2701:San Francisco Estuary 2666:Africa and Madagascar 2491:Underwater camouflage 2271:Aquatic biomonitoring 2211:freshwater ecosystems 1170:oceanservice.noaa.gov 949:Mytilus californianus 891: 829:Toxopnesutes pileolus 640:Van Veen grab sampler 328: 271: 139: 62:is a shallow pool of 42: 6073:Big fish–little pond 6042:Freshwater ecosystem 5976:Puddles on a surface 5592:Evolutionary ecology 5557:Ecological footprint 5552:Ecological economics 5476:Ecological threshold 5471:Ecological indicator 5341:Source–sink dynamics 5294:Land change modeling 5289:Insular biogeography 5141:Species distribution 4880:Modelling ecosystems 4539:Microbial metabolism 4378:Intraguild predation 4167:Biogeochemical cycle 4133:Modelling ecosystems 3888:Submergent coastline 3841:Marine transgression 3796:Discordant coastline 3781:Concordant coastline 2918:Marine bacteriophage 2880:Marine invertebrates 1373:"Snakelocks Anemone" 1361:on October 17, 2008. 1016:A starburst anemone 838:The presence of the 600:Deep sea communities 244:provide shelter for 6083:Constructed wetland 5642:Theoretical ecology 5617:Natural environment 5481:Ecosystem diversity 5451:Ecological collapse 5441:Bateman's principle 5396:Limiting similarity 5309:Landscape limnology 5131:Species homogeneity 4969:Population modeling 4964:Population dynamics 4781:Trophic state index 3992:Coastal engineering 2794:Ocean fertilization 2603:Trophic state index 2561:Lake stratification 2291:Aquatic respiration 2016:2021JSR...17202035C 1502:2021PNAS..11804769B 1300:2018PLoSO..1300066M 902:Pagurus longicarpus 5966:Coffee ring effect 5773:Infiltration basin 5753:Facultative lagoon 5653:Outline of ecology 5602:Industrial ecology 5597:Functional ecology 5461:Ecological deficit 5406:Niche construction 5369:Ecosystem engineer 5146:Species–area curve 5067:Introduced species 4882:: Other components 4814:Deimatic behaviour 4716:Ecological network 4648:North Pacific Gyre 4633:hydrothermal vents 4572:Ecological pyramid 4519:Microbial food web 4330:Primary production 4275:Foundation species 3961:Coastal management 3801:Emergent coastline 3472:Intertidal wetland 3375:Continental margin 3060:Intertidal wetland 3055:Intertidal ecology 2923:Marine prokaryotes 2865:Deep-sea community 2759:Iron fertilization 2682:Specific examples 2608:Upland and lowland 2526:Freshwater biology 2391:Microbial food web 2301:Aquatic toxicology 2244:Aquatic adaptation 2197:Aquatic ecosystems 1552:Connell, Joseph H. 1496:(3): e2004769118. 1401:on August 6, 2009. 1355:homepages.ed.ac.uk 1067:Pisaster ochraceus 894: 610:Intertidal ecology 585:Benthopelagic fish 506:    347: 276: 142: 83:gravitational pull 52: 6190:Coastal geography 6162: 6161: 6103:Pond of Abundance 6037:Aquatic ecosystem 5659: 5658: 5542:Balance of nature 5299:Landscape ecology 5184:Community ecology 5126:Species diversity 5062:Indicator species 5057:Gradient analysis 4934:Logistic function 4842: 4841: 4799:Animal coloration 4776:Trophic mutualism 4514:Microbial ecology 4305:Photoheterotrophs 4290:Myco-heterotrophy 4202:Ecosystem ecology 4187:Carrying capacity 4152:Abiotic component 4095: 4094: 3923:Wave-cut platform 3836:Marine regression 3380:Continental shelf 3370:Coastal waterfall 3279:Coastal geography 3245: 3244: 3226:Oceans portal 3195: 3194: 3191: 3190: 3070:Hydrothermal vent 3006: 3005: 2895:Seashore wildlife 2726:Marine ecosystems 2715: 2714: 2711: 2710: 2481:Thermal pollution 2446:Ramsar Convention 2386:Microbial ecology 2346:Fisheries science 2286:Aquatic predation 1715:978-0-12-356040-7 1698:Intertidal Fishes 1444:10.1890/06-1913.1 1270:on July 24, 2008. 1206:"NPCA Tide pools" 1110:Coastal predators 993:Calcium carbonate 853:Calcium carbonate 840:California mussel 761:pick up and drop 717: 716: 561:hydrothermal vent 106:marine biologists 16:(Redirected from 6207: 5875: 5748:Evaporation pond 5686: 5679: 5672: 5663: 5662: 5359:Ecological niche 5331:selection theory 5151:Umbrella species 5136:Species richness 5072:Invasive species 5052:Flagship species 4959:Population cycle 4954:Overexploitation 4919:Ecological yield 4869: 4862: 4855: 4846: 4845: 4751:Mesotrophic soil 4691:Climax community 4623:Marine food webs 4562:Biomagnification 4363:Chemoorganotroph 4217:Keystone species 4177:Biotic component 4122: 4115: 4108: 4099: 4098: 4085: 4084: 3846:Raised shoreline 3821:Ingression coast 3816:Graded shoreline 3791:Cuspate foreland 3623: 3616: 3567:Submarine canyon 3427:Freshwater marsh 3272: 3265: 3258: 3249: 3248: 3235: 3234: 3228: 3224: 3223: 3214: 3212:Lakes portal 3210: 3209: 3208: 3178:Marine pollution 2870:Deep-water coral 2853: 2852: 2841: 2774:Marine chemistry 2734: 2733: 2721: 2720: 2638:Freshwater marsh 2531:Freshwater biome 2513: 2229:Acoustic ecology 2219: 2218: 2205: 2204: 2190: 2183: 2176: 2167: 2166: 2149: 2148: 2116: 2110: 2109: 2092:(6): 1787–1794. 2079: 2073: 2072: 2042: 2036: 2035: 1995: 1989: 1988: 1948: 1942: 1941: 1909: 1903: 1902: 1892: 1860: 1854: 1853: 1821: 1815: 1814: 1808: 1803: 1801: 1793: 1777: 1771: 1770: 1753:(4): 1263–1290. 1737: 1726: 1725: 1724: 1722: 1689: 1678: 1677: 1667: 1643: 1637: 1636: 1588: 1582: 1581: 1548: 1542: 1541: 1531: 1513: 1481: 1472: 1471: 1438:(7): 1647–1653. 1423: 1417: 1416: 1409: 1403: 1402: 1391: 1385: 1384: 1382: 1380: 1369: 1363: 1362: 1357:. Archived from 1346: 1340: 1339: 1329: 1311: 1278: 1272: 1271: 1256: 1250: 1249: 1247: 1245: 1231: 1218: 1217: 1202: 1181: 1180: 1178: 1176: 1161: 1097: 1085:Anthopleura sola 1079: 1061: 1044: 1020:Anthopleura sola 1013: 915:Homing (biology) 910:Tidepool sculpin 896:Many species of 801:Anthopleura sola 739:, salinity, and 709: 702: 695: 679: 678: 677: 559: 546: 536: 524: 507: 393: 378: 368: 367: 349: 348: 47:, west coast of 21: 6215: 6214: 6210: 6209: 6208: 6206: 6205: 6204: 6175:Bodies of water 6165: 6164: 6163: 6158: 6051: 6023: 5985: 5947: 5923:Spent fuel pool 5918:Reflecting pool 5888:Anchialine pool 5876: 5867: 5699: 5690: 5660: 5655: 5646: 5632:Systems ecology 5520: 5491:Extinction debt 5456:Ecological debt 5446:Bioluminescence 5427: 5420: 5389:marine habitats 5364:Ecological trap 5345: 5225: 5218: 5161: 5155: 5111:Rapoport's rule 5106:Priority effect 5047:Endemic species 5015: 4974:Population size 4890: 4883: 4873: 4843: 4838: 4791: 4785: 4771:Trophic cascade 4681:Bioaccumulation 4664: 4591: 4548: 4470: 4437: 4334: 4246: 4207:Ecosystem model 4140: 4126: 4096: 4091: 4073: 4049:Intertidal zone 3975: 3942: 3831:Longshore drift 3776:Coastal erosion 3747: 3707: 3643:Beach evolution 3624: 3618: 3617: 3608: 3295:Anchialine pool 3281: 3276: 3246: 3241: 3218: 3217: 3206: 3204: 3203: 3187: 3148:Coral bleaching 3134: 3115:Seagrass meadow 3012:Marine habitats 3002: 2976:Coral reef fish 2942: 2928:Marine protists 2904: 2842: 2833: 2804:Ocean turbidity 2779:Marine food web 2728: 2707: 2649: 2588:River ecosystem 2541:Freshwater fish 2514: 2505: 2311:Bioluminescence 2296:Aquatic science 2213: 2199: 2194: 2157: 2152: 2117: 2113: 2098:10.2307/1941157 2080: 2076: 2043: 2039: 1996: 1992: 1949: 1945: 1930:10.1139/f71-051 1910: 1906: 1861: 1857: 1822: 1818: 1806: 1804: 1795: 1794: 1778: 1774: 1738: 1729: 1720: 1718: 1716: 1690: 1681: 1644: 1640: 1589: 1585: 1549: 1545: 1482: 1475: 1424: 1420: 1411: 1410: 1406: 1393: 1392: 1388: 1378: 1376: 1371: 1370: 1366: 1347: 1343: 1294:(7): e0200066. 1279: 1275: 1258: 1257: 1253: 1243: 1241: 1233: 1232: 1221: 1204: 1203: 1184: 1174: 1172: 1162: 1155: 1151: 1128:Intertidal fish 1124: 1112: 1105: 1098: 1089: 1080: 1071: 1062: 1053: 1045: 1036: 1028:Velella velella 1014: 998: 930: 783: 713: 675: 673: 666: 665: 664: 660:Marine sediment 635:Epibenthic sled 629: 621: 620: 619: 574: 566: 565: 564: 563: 557: 555: 544: 541: 540: 538:seagrass meadow 534: 532: 526:mangrove forest 522: 520: 505: 501: 493: 492: 491: 467: 459: 458: 457: 438: 430: 429: 428: 404: 391: 382: 376: 365: 323: 281: 274: 262: 194: 134: 132:Intertidal zone 128: 43:A tide pool in 35: 28: 23: 22: 15: 12: 11: 5: 6213: 6203: 6202: 6197: 6192: 6187: 6185:Marine biology 6182: 6177: 6160: 6159: 6157: 6156: 6151: 6143: 6138: 6136:Water aeration 6133: 6128: 6123: 6115: 6110: 6105: 6100: 6095: 6090: 6085: 6080: 6075: 6070: 6065: 6063:Aerated lagoon 6059: 6057: 6053: 6052: 6050: 6049: 6047:Lake ecosystem 6044: 6039: 6033: 6031: 6025: 6024: 6022: 6021: 6016: 6011: 6006: 6001: 5995: 5993: 5987: 5986: 5984: 5983: 5978: 5973: 5968: 5963: 5957: 5955: 5949: 5948: 5946: 5945: 5940: 5935: 5930: 5925: 5920: 5915: 5910: 5905: 5900: 5895: 5890: 5884: 5882: 5878: 5877: 5870: 5868: 5866: 5865: 5860: 5855: 5850: 5845: 5840: 5835: 5830: 5825: 5820: 5815: 5810: 5808:Retention pond 5805: 5800: 5798:Polishing pond 5795: 5790: 5785: 5780: 5775: 5770: 5768:Immersion pond 5765: 5760: 5755: 5750: 5745: 5740: 5738:Detention pond 5735: 5730: 5725: 5720: 5718:Balancing lake 5715: 5709: 5707: 5701: 5700: 5689: 5688: 5681: 5674: 5666: 5657: 5656: 5651: 5648: 5647: 5645: 5644: 5639: 5634: 5629: 5624: 5619: 5614: 5612:Microecosystem 5609: 5604: 5599: 5594: 5589: 5584: 5579: 5574: 5569: 5564: 5559: 5554: 5549: 5544: 5539: 5534: 5528: 5526: 5522: 5521: 5519: 5518: 5513: 5511:Thorson's rule 5508: 5503: 5498: 5493: 5488: 5483: 5478: 5473: 5468: 5463: 5458: 5453: 5448: 5443: 5438: 5436:Assembly rules 5432: 5430: 5422: 5421: 5419: 5418: 5413: 5408: 5403: 5398: 5393: 5392: 5391: 5381: 5376: 5371: 5366: 5361: 5355: 5353: 5347: 5346: 5344: 5343: 5338: 5333: 5321: 5319:Patch dynamics 5316: 5314:Metapopulation 5311: 5306: 5301: 5296: 5291: 5286: 5281: 5276: 5271: 5266: 5261: 5256: 5251: 5246: 5241: 5236: 5230: 5228: 5220: 5219: 5217: 5216: 5211: 5209:Storage effect 5206: 5201: 5196: 5191: 5186: 5181: 5176: 5171: 5165: 5163: 5157: 5156: 5154: 5153: 5148: 5143: 5138: 5133: 5128: 5123: 5118: 5113: 5108: 5103: 5098: 5093: 5091:Neutral theory 5088: 5083: 5078: 5076:Native species 5069: 5064: 5059: 5054: 5049: 5044: 5039: 5034: 5029: 5023: 5021: 5017: 5016: 5014: 5013: 5008: 5007: 5006: 5001: 4991: 4986: 4981: 4976: 4971: 4966: 4961: 4956: 4951: 4949:Overpopulation 4946: 4941: 4936: 4931: 4926: 4921: 4916: 4911: 4906: 4901: 4895: 4893: 4885: 4884: 4872: 4871: 4864: 4857: 4849: 4840: 4839: 4837: 4836: 4831: 4826: 4821: 4816: 4811: 4806: 4801: 4795: 4793: 4787: 4786: 4784: 4783: 4778: 4773: 4768: 4763: 4758: 4756:Nutrient cycle 4753: 4748: 4746:Feeding frenzy 4743: 4738: 4733: 4728: 4726:Energy quality 4723: 4718: 4713: 4708: 4703: 4698: 4693: 4688: 4686:Cascade effect 4683: 4678: 4672: 4670: 4666: 4665: 4663: 4662: 4661: 4660: 4655: 4650: 4645: 4640: 4635: 4630: 4620: 4615: 4610: 4605: 4599: 4597: 4593: 4592: 4590: 4589: 4584: 4579: 4574: 4569: 4564: 4558: 4556: 4550: 4549: 4547: 4546: 4541: 4536: 4531: 4529:Microbial loop 4526: 4521: 4516: 4511: 4506: 4501: 4496: 4494:Lithoautotroph 4491: 4486: 4480: 4478: 4476:Microorganisms 4472: 4471: 4469: 4468: 4463: 4458: 4453: 4447: 4445: 4439: 4438: 4436: 4435: 4433:Prey switching 4430: 4425: 4420: 4415: 4410: 4405: 4400: 4395: 4390: 4385: 4380: 4375: 4370: 4365: 4360: 4355: 4350: 4344: 4342: 4336: 4335: 4333: 4332: 4327: 4322: 4317: 4312: 4310:Photosynthesis 4307: 4302: 4297: 4292: 4287: 4282: 4277: 4272: 4267: 4265:Chemosynthesis 4262: 4256: 4254: 4248: 4247: 4245: 4244: 4239: 4234: 4229: 4224: 4219: 4214: 4209: 4204: 4199: 4194: 4189: 4184: 4179: 4174: 4169: 4164: 4159: 4157:Abiotic stress 4154: 4148: 4146: 4142: 4141: 4125: 4124: 4117: 4110: 4102: 4093: 4092: 4090: 4089: 4078: 4075: 4074: 4072: 4071: 4066: 4061: 4056: 4051: 4046: 4045: 4044: 4039: 4034: 4029: 4024: 4019: 4014: 4009: 4004: 3994: 3989: 3983: 3981: 3977: 3976: 3974: 3973: 3968: 3963: 3958: 3952: 3950: 3944: 3943: 3941: 3940: 3935: 3930: 3925: 3920: 3915: 3910: 3905: 3900: 3895: 3890: 3885: 3880: 3879: 3878: 3868: 3863: 3858: 3853: 3848: 3843: 3838: 3833: 3828: 3823: 3818: 3813: 3808: 3803: 3798: 3793: 3788: 3783: 3778: 3773: 3768: 3763: 3757: 3755: 3749: 3748: 3746: 3745: 3744: 3743: 3738: 3728: 3723: 3717: 3715: 3709: 3708: 3706: 3705: 3700: 3695: 3690: 3685: 3680: 3675: 3670: 3665: 3660: 3655: 3650: 3645: 3640: 3634: 3632: 3626: 3625: 3611: 3609: 3607: 3606: 3599: 3594: 3589: 3584: 3579: 3574: 3569: 3564: 3559: 3554: 3549: 3544: 3539: 3534: 3529: 3524: 3519: 3514: 3509: 3504: 3499: 3494: 3489: 3484: 3479: 3474: 3469: 3464: 3459: 3454: 3449: 3444: 3439: 3434: 3429: 3424: 3419: 3414: 3409: 3404: 3403: 3402: 3392: 3387: 3382: 3377: 3372: 3367: 3362: 3357: 3352: 3347: 3342: 3340:Brackish marsh 3337: 3332: 3327: 3322: 3320:Barrier island 3317: 3312: 3307: 3302: 3297: 3291: 3289: 3283: 3282: 3275: 3274: 3267: 3260: 3252: 3243: 3242: 3240: 3239: 3229: 3215: 3200: 3197: 3196: 3193: 3192: 3189: 3188: 3186: 3185: 3180: 3175: 3170: 3165: 3160: 3155: 3150: 3144: 3142: 3136: 3135: 3133: 3132: 3127: 3122: 3117: 3112: 3107: 3102: 3097: 3092: 3087: 3082: 3077: 3072: 3067: 3062: 3057: 3052: 3047: 3042: 3037: 3032: 3027: 3022: 3016: 3014: 3008: 3007: 3004: 3003: 3001: 3000: 2995: 2994: 2993: 2988: 2983: 2978: 2973: 2966:Saltwater fish 2963: 2961:Marine reptile 2958: 2952: 2950: 2944: 2943: 2941: 2940: 2935: 2933:Marine viruses 2930: 2925: 2920: 2914: 2912: 2910:Microorganisms 2906: 2905: 2903: 2902: 2900:Wild fisheries 2897: 2892: 2887: 2882: 2877: 2872: 2867: 2862: 2856: 2850: 2844: 2843: 2836: 2834: 2832: 2831: 2826: 2821: 2816: 2814:Thorson's rule 2811: 2806: 2801: 2796: 2791: 2786: 2781: 2776: 2771: 2769:Marine biology 2766: 2761: 2756: 2751: 2746: 2740: 2738: 2730: 2729: 2717: 2716: 2713: 2712: 2709: 2708: 2706: 2705: 2704: 2703: 2698: 2693: 2688: 2680: 2675: 2674: 2673: 2668: 2657: 2655: 2651: 2650: 2648: 2647: 2646: 2645: 2640: 2635: 2630: 2628:Brackish marsh 2625: 2615: 2610: 2605: 2600: 2595: 2590: 2585: 2580: 2579: 2578: 2568: 2563: 2558: 2556:Lake ecosystem 2553: 2548: 2546:Hyporheic zone 2543: 2538: 2533: 2528: 2522: 2520: 2516: 2515: 2508: 2506: 2504: 2503: 2498: 2493: 2488: 2483: 2478: 2473: 2468: 2463: 2458: 2453: 2448: 2443: 2438: 2433: 2428: 2423: 2418: 2413: 2408: 2403: 2398: 2396:Microbial loop 2393: 2388: 2383: 2378: 2373: 2368: 2363: 2358: 2353: 2348: 2343: 2341:Eutrophication 2338: 2333: 2328: 2323: 2321:Cascade effect 2318: 2313: 2308: 2303: 2298: 2293: 2288: 2283: 2278: 2273: 2268: 2267: 2266: 2261: 2256: 2249:Aquatic animal 2246: 2241: 2236: 2231: 2225: 2223: 2215: 2214: 2201: 2200: 2193: 2192: 2185: 2178: 2170: 2164: 2163: 2156: 2155:External links 2153: 2151: 2150: 2131:(2): 105–127. 2111: 2074: 2037: 1990: 1943: 1924:(3): 383–389. 1904: 1875:(2): 239–250. 1855: 1816: 1807:|journal= 1772: 1727: 1714: 1679: 1638: 1609:10.1086/700188 1603:(3): 178–184. 1583: 1564:(1): 169–192. 1543: 1473: 1418: 1404: 1386: 1364: 1351:"Sea Anemones" 1341: 1273: 1251: 1219: 1182: 1152: 1150: 1147: 1146: 1145: 1140: 1135: 1130: 1123: 1120: 1111: 1108: 1107: 1106: 1099: 1092: 1090: 1082:Sea anemones, 1081: 1074: 1072: 1063: 1056: 1054: 1046: 1039: 1037: 1015: 1008: 996: 929: 926: 815:Sea urchins (" 782: 779: 715: 714: 712: 711: 704: 697: 689: 686: 685: 684: 683: 668: 667: 663: 662: 657: 652: 647: 642: 637: 631: 630: 627: 626: 623: 622: 618: 617: 612: 607: 602: 597: 592: 587: 582: 576: 575: 572: 571: 568: 567: 556: 543: 542: 533: 521: 504: 503: 502: 499: 498: 495: 494: 490: 489: 488: 487: 482: 477: 469: 468: 465: 464: 461: 460: 456: 455: 454: 453: 448: 440: 439: 436: 435: 432: 431: 427: 426: 425: 424: 419: 414: 406: 405: 402: 401: 398: 397: 370: 369: 361: 360: 354: 353: 322: 319: 307:photosynthetic 280: 277: 272: 261: 258: 254:filter feeders 193: 192:High Tide Zone 190: 163: 162: 159: 156: 153: 130:Main article: 127: 124: 114:John Steinbeck 26: 9: 6: 4: 3: 2: 6212: 6201: 6198: 6196: 6195:Natural pools 6193: 6191: 6188: 6186: 6183: 6181: 6178: 6176: 6173: 6172: 6170: 6155: 6152: 6150: 6148: 6144: 6142: 6139: 6137: 6134: 6132: 6131:Swimming hole 6129: 6127: 6124: 6122: 6120: 6116: 6114: 6111: 6109: 6106: 6104: 6101: 6099: 6096: 6094: 6091: 6089: 6086: 6084: 6081: 6079: 6078:Body of water 6076: 6074: 6071: 6069: 6066: 6064: 6061: 6060: 6058: 6054: 6048: 6045: 6043: 6040: 6038: 6035: 6034: 6032: 6030: 6026: 6020: 6017: 6015: 6014:Goldfish pond 6012: 6010: 6007: 6005: 6002: 6000: 5997: 5996: 5994: 5992: 5988: 5982: 5979: 5977: 5974: 5972: 5969: 5967: 5964: 5962: 5959: 5958: 5956: 5954: 5950: 5944: 5941: 5939: 5936: 5934: 5933:Swimming pool 5931: 5929: 5926: 5924: 5921: 5919: 5916: 5914: 5911: 5909: 5906: 5904: 5901: 5899: 5898:Infinity pool 5896: 5894: 5891: 5889: 5886: 5885: 5883: 5879: 5874: 5864: 5861: 5859: 5856: 5854: 5851: 5849: 5846: 5844: 5841: 5839: 5836: 5834: 5831: 5829: 5828:Settling pond 5826: 5824: 5823:Sediment pond 5821: 5819: 5816: 5814: 5811: 5809: 5806: 5804: 5801: 5799: 5796: 5794: 5791: 5789: 5786: 5784: 5781: 5779: 5776: 5774: 5771: 5769: 5766: 5764: 5761: 5759: 5756: 5754: 5751: 5749: 5746: 5744: 5741: 5739: 5736: 5734: 5731: 5729: 5726: 5724: 5721: 5719: 5716: 5714: 5711: 5710: 5708: 5706: 5702: 5698: 5695:, pools, and 5694: 5687: 5682: 5680: 5675: 5673: 5668: 5667: 5664: 5654: 5649: 5643: 5640: 5638: 5637:Urban ecology 5635: 5633: 5630: 5628: 5625: 5623: 5620: 5618: 5615: 5613: 5610: 5608: 5605: 5603: 5600: 5598: 5595: 5593: 5590: 5588: 5585: 5583: 5580: 5578: 5575: 5573: 5570: 5568: 5565: 5563: 5560: 5558: 5555: 5553: 5550: 5548: 5545: 5543: 5540: 5538: 5535: 5533: 5530: 5529: 5527: 5523: 5517: 5514: 5512: 5509: 5507: 5504: 5502: 5499: 5497: 5496:Kleiber's law 5494: 5492: 5489: 5487: 5484: 5482: 5479: 5477: 5474: 5472: 5469: 5467: 5464: 5462: 5459: 5457: 5454: 5452: 5449: 5447: 5444: 5442: 5439: 5437: 5434: 5433: 5431: 5429: 5423: 5417: 5414: 5412: 5409: 5407: 5404: 5402: 5399: 5397: 5394: 5390: 5387: 5386: 5385: 5382: 5380: 5377: 5375: 5372: 5370: 5367: 5365: 5362: 5360: 5357: 5356: 5354: 5352: 5348: 5342: 5339: 5337: 5334: 5332: 5330: 5326: 5322: 5320: 5317: 5315: 5312: 5310: 5307: 5305: 5302: 5300: 5297: 5295: 5292: 5290: 5287: 5285: 5282: 5280: 5277: 5275: 5272: 5270: 5269:Foster's rule 5267: 5265: 5262: 5260: 5257: 5255: 5252: 5250: 5247: 5245: 5242: 5240: 5237: 5235: 5232: 5231: 5229: 5227: 5221: 5215: 5212: 5210: 5207: 5205: 5202: 5200: 5197: 5195: 5192: 5190: 5187: 5185: 5182: 5180: 5177: 5175: 5172: 5170: 5167: 5166: 5164: 5158: 5152: 5149: 5147: 5144: 5142: 5139: 5137: 5134: 5132: 5129: 5127: 5124: 5122: 5119: 5117: 5114: 5112: 5109: 5107: 5104: 5102: 5099: 5097: 5094: 5092: 5089: 5087: 5084: 5082: 5079: 5077: 5073: 5070: 5068: 5065: 5063: 5060: 5058: 5055: 5053: 5050: 5048: 5045: 5043: 5040: 5038: 5035: 5033: 5030: 5028: 5025: 5024: 5022: 5018: 5012: 5009: 5005: 5002: 5000: 4997: 4996: 4995: 4992: 4990: 4987: 4985: 4982: 4980: 4977: 4975: 4972: 4970: 4967: 4965: 4962: 4960: 4957: 4955: 4952: 4950: 4947: 4945: 4942: 4940: 4937: 4935: 4932: 4930: 4927: 4925: 4922: 4920: 4917: 4915: 4912: 4910: 4907: 4905: 4902: 4900: 4897: 4896: 4894: 4892: 4886: 4881: 4877: 4870: 4865: 4863: 4858: 4856: 4851: 4850: 4847: 4835: 4832: 4830: 4827: 4825: 4822: 4820: 4817: 4815: 4812: 4810: 4807: 4805: 4802: 4800: 4797: 4796: 4794: 4788: 4782: 4779: 4777: 4774: 4772: 4769: 4767: 4764: 4762: 4759: 4757: 4754: 4752: 4749: 4747: 4744: 4742: 4739: 4737: 4734: 4732: 4729: 4727: 4724: 4722: 4719: 4717: 4714: 4712: 4709: 4707: 4704: 4702: 4699: 4697: 4694: 4692: 4689: 4687: 4684: 4682: 4679: 4677: 4674: 4673: 4671: 4667: 4659: 4656: 4654: 4651: 4649: 4646: 4644: 4641: 4639: 4636: 4634: 4631: 4629: 4626: 4625: 4624: 4621: 4619: 4616: 4614: 4611: 4609: 4606: 4604: 4601: 4600: 4598: 4594: 4588: 4587:Trophic level 4585: 4583: 4580: 4578: 4575: 4573: 4570: 4568: 4565: 4563: 4560: 4559: 4557: 4555: 4551: 4545: 4544:Phage ecology 4542: 4540: 4537: 4535: 4534:Microbial mat 4532: 4530: 4527: 4525: 4522: 4520: 4517: 4515: 4512: 4510: 4507: 4505: 4502: 4500: 4497: 4495: 4492: 4490: 4489:Bacteriophage 4487: 4485: 4482: 4481: 4479: 4477: 4473: 4467: 4464: 4462: 4459: 4457: 4456:Decomposition 4454: 4452: 4449: 4448: 4446: 4444: 4440: 4434: 4431: 4429: 4426: 4424: 4421: 4419: 4416: 4414: 4411: 4409: 4406: 4404: 4403:Mesopredators 4401: 4399: 4396: 4394: 4391: 4389: 4386: 4384: 4381: 4379: 4376: 4374: 4371: 4369: 4366: 4364: 4361: 4359: 4356: 4354: 4351: 4349: 4348:Apex predator 4346: 4345: 4343: 4341: 4337: 4331: 4328: 4326: 4323: 4321: 4318: 4316: 4313: 4311: 4308: 4306: 4303: 4301: 4298: 4296: 4293: 4291: 4288: 4286: 4283: 4281: 4278: 4276: 4273: 4271: 4268: 4266: 4263: 4261: 4258: 4257: 4255: 4253: 4249: 4243: 4240: 4238: 4235: 4233: 4230: 4228: 4225: 4223: 4220: 4218: 4215: 4213: 4210: 4208: 4205: 4203: 4200: 4198: 4195: 4193: 4190: 4188: 4185: 4183: 4182:Biotic stress 4180: 4178: 4175: 4173: 4170: 4168: 4165: 4163: 4160: 4158: 4155: 4153: 4150: 4149: 4147: 4143: 4138: 4134: 4130: 4123: 4118: 4116: 4111: 4109: 4104: 4103: 4100: 4088: 4080: 4079: 4076: 4070: 4067: 4065: 4062: 4060: 4057: 4055: 4054:Littoral zone 4052: 4050: 4047: 4043: 4040: 4038: 4035: 4033: 4030: 4028: 4025: 4023: 4020: 4018: 4015: 4013: 4010: 4008: 4005: 4003: 4000: 3999: 3998: 3995: 3993: 3990: 3988: 3987:Bulkhead line 3985: 3984: 3982: 3978: 3972: 3969: 3967: 3964: 3962: 3959: 3957: 3954: 3953: 3951: 3949: 3945: 3939: 3936: 3934: 3931: 3929: 3928:Wave shoaling 3926: 3924: 3921: 3919: 3916: 3914: 3911: 3909: 3906: 3904: 3903:Surge channel 3901: 3899: 3896: 3894: 3891: 3889: 3886: 3884: 3881: 3877: 3874: 3873: 3872: 3869: 3867: 3864: 3862: 3859: 3857: 3854: 3852: 3849: 3847: 3844: 3842: 3839: 3837: 3834: 3832: 3829: 3827: 3824: 3822: 3819: 3817: 3814: 3812: 3809: 3807: 3804: 3802: 3799: 3797: 3794: 3792: 3789: 3787: 3784: 3782: 3779: 3777: 3774: 3772: 3769: 3767: 3766:Cliffed coast 3764: 3762: 3759: 3758: 3756: 3754: 3750: 3742: 3739: 3737: 3734: 3733: 3732: 3729: 3727: 3724: 3722: 3719: 3718: 3716: 3714: 3710: 3704: 3701: 3699: 3696: 3694: 3693:Shingle beach 3691: 3689: 3686: 3684: 3681: 3679: 3676: 3674: 3671: 3669: 3666: 3664: 3661: 3659: 3656: 3654: 3651: 3649: 3646: 3644: 3641: 3639: 3636: 3635: 3633: 3631: 3627: 3622: 3615: 3605: 3604: 3600: 3598: 3595: 3593: 3590: 3588: 3585: 3583: 3580: 3578: 3575: 3573: 3570: 3568: 3565: 3563: 3560: 3558: 3555: 3553: 3550: 3548: 3545: 3543: 3540: 3538: 3535: 3533: 3530: 3528: 3525: 3523: 3520: 3518: 3515: 3513: 3510: 3508: 3505: 3503: 3500: 3498: 3495: 3493: 3490: 3488: 3485: 3483: 3480: 3478: 3475: 3473: 3470: 3468: 3465: 3463: 3460: 3458: 3455: 3453: 3450: 3448: 3445: 3443: 3440: 3438: 3435: 3433: 3430: 3428: 3425: 3423: 3420: 3418: 3415: 3413: 3410: 3408: 3405: 3401: 3398: 3397: 3396: 3393: 3391: 3388: 3386: 3383: 3381: 3378: 3376: 3373: 3371: 3368: 3366: 3365:Coastal plain 3363: 3361: 3358: 3356: 3353: 3351: 3348: 3346: 3343: 3341: 3338: 3336: 3333: 3331: 3328: 3326: 3323: 3321: 3318: 3316: 3313: 3311: 3308: 3306: 3303: 3301: 3298: 3296: 3293: 3292: 3290: 3288: 3284: 3280: 3273: 3268: 3266: 3261: 3259: 3254: 3253: 3250: 3238: 3230: 3227: 3222: 3216: 3213: 3202: 3201: 3198: 3184: 3181: 3179: 3176: 3174: 3171: 3169: 3166: 3164: 3161: 3159: 3156: 3154: 3151: 3149: 3146: 3145: 3143: 3141: 3137: 3131: 3128: 3126: 3123: 3121: 3120:Sponge ground 3118: 3116: 3113: 3111: 3108: 3106: 3103: 3101: 3098: 3096: 3093: 3091: 3088: 3086: 3085:Marine biomes 3083: 3081: 3078: 3076: 3073: 3071: 3068: 3066: 3063: 3061: 3058: 3056: 3053: 3051: 3048: 3046: 3043: 3041: 3038: 3036: 3033: 3031: 3028: 3026: 3023: 3021: 3018: 3017: 3015: 3013: 3009: 2999: 2996: 2992: 2989: 2987: 2986:Demersal fish 2984: 2982: 2981:Deep-sea fish 2979: 2977: 2974: 2972: 2969: 2968: 2967: 2964: 2962: 2959: 2957: 2956:Marine mammal 2954: 2953: 2951: 2949: 2945: 2939: 2936: 2934: 2931: 2929: 2926: 2924: 2921: 2919: 2916: 2915: 2913: 2911: 2907: 2901: 2898: 2896: 2893: 2891: 2888: 2886: 2883: 2881: 2878: 2876: 2873: 2871: 2868: 2866: 2863: 2861: 2858: 2857: 2854: 2851: 2849: 2845: 2840: 2830: 2827: 2825: 2822: 2820: 2817: 2815: 2812: 2810: 2807: 2805: 2802: 2800: 2797: 2795: 2792: 2790: 2787: 2785: 2782: 2780: 2777: 2775: 2772: 2770: 2767: 2765: 2762: 2760: 2757: 2755: 2752: 2750: 2747: 2745: 2742: 2741: 2739: 2735: 2731: 2727: 2722: 2718: 2702: 2699: 2697: 2694: 2692: 2689: 2687: 2684: 2683: 2681: 2679: 2676: 2672: 2669: 2667: 2664: 2663: 2662: 2659: 2658: 2656: 2652: 2644: 2641: 2639: 2636: 2634: 2631: 2629: 2626: 2624: 2621: 2620: 2619: 2616: 2614: 2611: 2609: 2606: 2604: 2601: 2599: 2596: 2594: 2591: 2589: 2586: 2584: 2581: 2577: 2574: 2573: 2572: 2569: 2567: 2564: 2562: 2559: 2557: 2554: 2552: 2549: 2547: 2544: 2542: 2539: 2537: 2534: 2532: 2529: 2527: 2524: 2523: 2521: 2517: 2512: 2502: 2499: 2497: 2494: 2492: 2489: 2487: 2486:Trophic level 2484: 2482: 2479: 2477: 2474: 2472: 2469: 2467: 2464: 2462: 2459: 2457: 2454: 2452: 2451:Sediment trap 2449: 2447: 2444: 2442: 2439: 2437: 2434: 2432: 2429: 2427: 2426:Phytoplankton 2424: 2422: 2419: 2417: 2414: 2412: 2409: 2407: 2404: 2402: 2399: 2397: 2394: 2392: 2389: 2387: 2384: 2382: 2379: 2377: 2374: 2372: 2369: 2367: 2364: 2362: 2359: 2357: 2354: 2352: 2349: 2347: 2344: 2342: 2339: 2337: 2334: 2332: 2329: 2327: 2324: 2322: 2319: 2317: 2314: 2312: 2309: 2307: 2304: 2302: 2299: 2297: 2294: 2292: 2289: 2287: 2284: 2282: 2279: 2277: 2276:Aquatic plant 2274: 2272: 2269: 2265: 2262: 2260: 2257: 2255: 2252: 2251: 2250: 2247: 2245: 2242: 2240: 2239:Anoxic waters 2237: 2235: 2232: 2230: 2227: 2226: 2224: 2220: 2216: 2212: 2206: 2202: 2198: 2191: 2186: 2184: 2179: 2177: 2172: 2171: 2168: 2162: 2159: 2158: 2146: 2142: 2138: 2134: 2130: 2126: 2122: 2115: 2107: 2103: 2099: 2095: 2091: 2087: 2086: 2078: 2070: 2066: 2062: 2058: 2057: 2052: 2050: 2041: 2033: 2029: 2025: 2021: 2017: 2013: 2009: 2005: 2001: 1994: 1986: 1982: 1978: 1974: 1970: 1966: 1962: 1958: 1954: 1947: 1939: 1935: 1931: 1927: 1923: 1919: 1915: 1908: 1900: 1896: 1891: 1890:10400.12/1509 1886: 1882: 1878: 1874: 1870: 1866: 1859: 1851: 1847: 1843: 1839: 1835: 1831: 1827: 1820: 1812: 1799: 1791: 1787: 1783: 1776: 1768: 1764: 1760: 1756: 1752: 1748: 1744: 1736: 1734: 1732: 1717: 1711: 1707: 1703: 1699: 1695: 1688: 1686: 1684: 1675: 1671: 1666: 1661: 1657: 1653: 1649: 1642: 1634: 1630: 1626: 1622: 1618: 1614: 1610: 1606: 1602: 1598: 1594: 1587: 1579: 1575: 1571: 1567: 1563: 1559: 1558: 1553: 1547: 1539: 1535: 1530: 1525: 1521: 1517: 1512: 1507: 1503: 1499: 1495: 1491: 1487: 1480: 1478: 1469: 1465: 1461: 1457: 1453: 1449: 1445: 1441: 1437: 1433: 1429: 1422: 1414: 1413:"Sea Urchins" 1408: 1400: 1396: 1390: 1374: 1368: 1360: 1356: 1352: 1345: 1337: 1333: 1328: 1323: 1319: 1315: 1310: 1305: 1301: 1297: 1293: 1289: 1285: 1277: 1269: 1265: 1261: 1255: 1240: 1236: 1230: 1228: 1226: 1224: 1215: 1211: 1207: 1201: 1199: 1197: 1195: 1193: 1191: 1189: 1187: 1171: 1167: 1160: 1158: 1153: 1144: 1141: 1139: 1136: 1134: 1131: 1129: 1126: 1125: 1119: 1117: 1103: 1096: 1091: 1087: 1086: 1078: 1073: 1069: 1068: 1060: 1055: 1051: 1050: 1043: 1038: 1035: 1031: 1029: 1023: 1021: 1012: 1007: 1006: 1005: 1002: 994: 990: 985: 983: 979: 978: 973: 972: 967: 963: 958: 953: 951: 950: 945: 941: 937: 936: 925: 923: 918: 916: 911: 905: 903: 899: 890: 886: 884: 880: 876: 871: 868: 864: 860: 856: 854: 850: 846: 841: 836: 834: 830: 826: 822: 818: 813: 810: 805: 803: 802: 797: 793: 792: 788: 778: 776: 772: 768: 764: 760: 756: 752: 750: 746: 742: 738: 734: 730: 726: 722: 710: 705: 703: 698: 696: 691: 690: 688: 687: 682: 672: 671: 670: 669: 661: 658: 656: 655:Demersal zone 653: 651: 648: 646: 645:Bioirrigation 643: 641: 638: 636: 633: 632: 625: 624: 616: 613: 611: 608: 606: 605:Demersal fish 603: 601: 598: 596: 593: 591: 590:Bottom feeder 588: 586: 583: 581: 578: 577: 570: 569: 562: 554: 550: 539: 531: 527: 519: 515: 511: 497: 496: 486: 483: 481: 478: 476: 473: 472: 471: 470: 463: 462: 452: 449: 447: 444: 443: 442: 441: 434: 433: 423: 420: 418: 415: 413: 410: 409: 408: 407: 400: 399: 396: 389: 385: 384:Benthopelagic 381: 375: 372: 371: 363: 362: 359: 356: 355: 351: 350: 344: 340: 336: 332: 327: 318: 316: 312: 308: 303: 299: 295: 291: 287: 279:Low Tide Zone 270: 266: 260:Mid Tide Zone 257: 255: 251: 247: 243: 239: 235: 231: 227: 223: 219: 215: 214:invertebrates 211: 207: 203: 199: 189: 187: 183: 179: 175: 171: 166: 160: 157: 154: 151: 147: 146: 145: 138: 133: 123: 121: 120: 115: 111: 110:philosophical 108:, as well as 107: 103: 99: 95: 91: 86: 84: 80: 76: 72: 69: 65: 61: 57: 50: 46: 41: 37: 33: 19: 6147:Water Lilies 6146: 6141:Water garden 6121:(M C Escher) 6118: 6068:Bakki shower 5937: 5903:Natural pool 5803:Raceway pond 5733:Cooling pond 5723:Ballast pond 5622:Regime shift 5607:Macroecology 5328: 5324: 5264:Edge effects 5234:Biogeography 5179:Commensalism 5027:Biodiversity 4904:Allee effect 4657: 4643:kelp forests 4596:Example webs 4461:Detritivores 4300:Organotrophs 4280:Kinetotrophs 4232:Productivity 3918:Volcanic arc 3806:Feeder bluff 3713:River mouths 3678:Raised beach 3673:Pocket beach 3601: 3581: 3572:Tidal island 3562:Strand plain 3507:Natural arch 3140:Conservation 3129: 2991:Pelagic fish 2971:Coastal fish 2875:Marine fungi 2613:Water garden 2496:Water column 2441:Productivity 2416:Pelagic zone 2376:Macrobenthos 2366:Hydrobiology 2336:Ecohydrology 2128: 2124: 2114: 2089: 2083: 2077: 2060: 2054: 2048: 2040: 2007: 2003: 1993: 1960: 1956: 1946: 1921: 1917: 1907: 1872: 1868: 1858: 1833: 1829: 1819: 1798:cite journal 1775: 1750: 1746: 1721:December 18, 1719:, retrieved 1697: 1655: 1651: 1641: 1600: 1596: 1586: 1561: 1555: 1546: 1493: 1489: 1435: 1431: 1421: 1407: 1399:the original 1389: 1379:September 6, 1377:. Retrieved 1367: 1359:the original 1354: 1344: 1291: 1287: 1276: 1268:the original 1254: 1244:February 26, 1242:. Retrieved 1238: 1214:the original 1175:February 26, 1173:. Retrieved 1169: 1113: 1102:Kachemak Bay 1083: 1065: 1047: 1025: 1017: 986: 981: 975: 969: 962:gametophytes 956: 954: 947: 933: 931: 919: 906: 895: 857: 837: 814: 806: 799: 789: 784: 753: 718: 650:Bioturbation 580:Benthic fish 518:bivalve reef 509: 485:hyperbenthos 451:phytobenthos 422:microbenthos 412:macrobenthos 380:Benthic zone 358:Benthic life 339:sea anemones 282: 263: 250:hermit crabs 218:sea anemones 195: 167: 164: 143: 117: 87: 59: 55: 53: 36: 6093:Hydric soil 5981:Seep puddle 5943:Vernal pool 5928:Stream pool 5913:Plunge pool 5778:Kettle pond 5758:Garden pond 5259:Disturbance 5162:interaction 4984:Recruitment 4914:Depensation 4706:Copiotrophs 4577:Energy flow 4499:Lithotrophy 4443:Decomposers 4423:Planktivore 4398:Insectivore 4388:Heterotroph 4353:Bacterivore 4320:Phototrophs 4270:Chemotrophs 4242:Restoration 4192:Competition 4069:River plume 3883:Steep coast 3856:Rocky shore 3851:Rip current 3731:River delta 3703:Wash margin 3698:Storm beach 3688:Shell beach 3653:Beach wrack 3648:Beach ridge 3638:Beach cusps 3587:Tied island 3577:Tidal marsh 3300:Archipelago 3125:Sponge reef 3100:Rocky shore 3095:Oyster reef 3065:Kelp forest 2948:Vertebrates 2848:Marine life 2824:Viral shunt 2789:Marine snow 2691:Maharashtra 2598:Stream pool 2501:Zooplankton 2421:Photic zone 2381:Meiobenthos 2234:Algal bloom 2063:(1): 1–14. 1836:(1): 1–29. 1239:www.nps.gov 1138:Rocky shore 1001:desiccation 980:, allowing 977:Halosaccion 932:Sea palms ( 898:Hermit crab 787:sea anemone 767:Black bears 763:sea urchins 737:temperature 733:environment 573:Communities 475:endobenthos 466:By location 417:meiobenthos 343:sea sponges 321:Marine life 302:vertebrates 246:nudibranchs 234:green algae 202:wave action 176:and during 170:tidal range 150:terrestrial 112:essayists: 102:naturalists 6169:Categories 6108:Pond liner 6098:Phytotelma 6029:Ecosystems 5999:Beaver dam 5908:Ocean pool 5893:Brine pool 5858:Waste pond 5833:Solar pond 5627:Sexecology 5204:Parasitism 5169:Antibiosis 5004:Resistance 4999:Resilience 4889:Population 4809:Camouflage 4761:Oligotroph 4676:Ascendency 4638:intertidal 4628:cold seeps 4582:Food chain 4383:Herbivores 4358:Carnivores 4285:Mixotrophs 4260:Autotrophs 4139:components 3997:Grain size 3971:Submersion 3948:Management 3933:Wind fetch 3893:Surf break 3811:Flat coast 3741:regressive 3527:Salt marsh 3385:Coral reef 3105:Salt marsh 3040:Coral reef 2829:Whale fall 2809:Photophore 2686:Everglades 2654:Ecoregions 2593:Stream bed 2566:Macrophyte 2519:Freshwater 2351:Food chain 2264:Water bird 2010:: 102035. 1658:(4): 488. 1149:References 1064:Sea star, 989:tide pools 940:palm trees 875:desiccated 817:Echinoidia 809:regenerate 530:coral reef 500:By habitat 480:epibenthos 446:zoobenthos 216:, such as 174:high tides 88:Tide pool 68:intertidal 45:Porto Covo 6088:Full pond 6009:Fish pond 6004:Duck pond 5961:Bird bath 5938:Tide pool 5843:Stew pond 5793:Mill pond 5788:Melt pond 5532:Allometry 5486:Emergence 5214:Symbiosis 5199:Mutualism 4994:Stability 4899:Abundance 4711:Dominance 4669:Processes 4658:tide pool 4554:Food webs 4428:Predation 4413:Omnivores 4340:Consumers 4295:Mycotroph 4252:Producers 4197:Ecosystem 4162:Behaviour 3956:Accretion 3938:Wind wave 3898:Surf zone 3753:Processes 3726:Mouth bar 3683:Recession 3663:Beachrock 3582:Tide pool 3512:Peninsula 3400:cliff-top 3287:Landforms 3130:Tide pool 3035:Cold seep 2819:Upwelling 2583:Rheotaxis 2576:Fish pond 2551:Limnology 2476:Substrate 2461:Siltation 2331:Dead zone 2145:0022-0981 2032:234822803 1963:: 20–28. 1938:0015-296X 1899:1573-5184 1850:0022-0981 1767:1573-5184 1674:0278-0372 1617:0006-3185 1520:0027-8424 1452:0012-9658 1318:1932-6203 1034:hydrozoan 982:Postelsia 971:Corallina 957:Postelsia 935:Postelsia 922:Amphipoda 863:barnacles 845:Aragonite 775:food webs 747:, strong 721:sea stars 553:cold seep 510:tide pool 508:Shallow: 335:sea stars 315:predators 240:. Marine 186:organisms 116:wrote in 60:rock pool 56:tide pool 6019:Koi pond 5848:Tailings 5838:Stepwell 5813:Sag pond 5783:Log pond 5763:Ice pond 5743:Dew pond 5713:Ash pond 5587:Endolith 5516:Xerosere 5428:networks 5244:Ecocline 4790:Defense, 4466:Detritus 4368:Foraging 4237:Resource 4087:Category 3913:Undertow 3866:Sea foam 3861:Sea cave 3761:Blowhole 3603:Windwatt 3462:Headland 3310:Avulsion 3237:Category 3163:HERMIONE 3080:Mangrove 2890:Seagrass 2436:Pleuston 2431:Plankton 2411:Particle 2356:Food web 1985:54486846 1977:30502824 1633:58602447 1625:30624115 1538:33431664 1468:17645011 1460:27651282 1336:29975745 1288:PLOS ONE 1122:See also 1116:food web 1104:, Alaska 968:such as 867:barnacle 749:currents 595:Deep sea 549:seamount 388:coupling 333:showing 311:salinity 290:hydroids 222:sea star 206:currents 98:salinity 90:habitats 79:low tide 77:only at 64:seawater 49:Portugal 18:Rockpool 6149:(Monet) 6113:Ponding 6056:Related 5953:Puddles 5697:puddles 5577:Ecopath 5384:Habitat 5254:Ecotype 5249:Ecotone 5226:ecology 5224:Spatial 5160:Species 5020:Species 4891:ecology 4876:Ecology 4824:Mimicry 4792:counter 4736:f-ratio 4484:Archaea 4172:Biomass 4145:General 4137:Trophic 4129:Ecology 4037:shingle 4017:granule 4002:boulder 3980:Related 3876:peresyp 3786:Current 3721:Debouch 3630:Beaches 3597:Waituna 3592:Tombolo 3502:Mudflat 3497:Machair 3487:Isthmus 3407:Estuary 3350:Channel 3090:Mudflat 3050:Estuary 3020:Bay mud 2998:Seabird 2754:f-ratio 2737:General 2618:Wetland 2406:Neuston 2371:Hypoxia 2316:Biomass 2306:Benthos 2222:General 2106:1941157 2085:Ecology 2012:Bibcode 1957:Zoology 1578:2096846 1529:7826377 1498:Bibcode 1432:Ecology 1327:6033428 1296:Bibcode 1032:a blue 944:thallus 879:calcite 870:species 859:Lichens 849:Calcite 796:fission 725:mussels 628:Related 514:bay mud 437:By type 403:By size 374:Benthos 298:limpets 294:isopods 286:abalone 238:mussels 226:chitons 210:seaweed 198:flooded 94:animals 6126:Spring 6119:Puddle 5971:Puddle 4608:Rivers 4504:Marine 4027:pebble 4022:gravel 4012:cobble 3557:Strait 3537:Skerry 3492:Lagoon 3477:Island 3432:Fundus 3335:Bodden 3075:Lagoon 2401:Nekton 2259:Mammal 2254:Insect 2143:  2104:  2030:  1983:  1975:  1936:  1897:  1848:  1765:  1712:  1672:  1631:  1623:  1615:  1576:  1536:  1526:  1518:  1466:  1458:  1450:  1334:  1324:  1316:  883:plates 825:spines 821:urchin 741:oxygen 547:Deep: 395:Seabed 341:, and 236:, and 178:storms 6180:Tides 5991:Biome 5881:Pools 5705:Ponds 5693:Ponds 5525:Other 5426:Other 5379:Guild 5351:Niche 4603:Lakes 3908:Swash 3871:Shoal 3552:Stack 3542:Sound 3532:Shoal 3482:Islet 3467:Inlet 3457:Hapua 3422:Fjord 3417:Fjard 3412:Firth 3360:Coast 3355:Cliff 3330:Bight 3305:Atoll 2466:Spawn 2102:JSTOR 2028:S2CID 1981:S2CID 1629:S2CID 1574:JSTOR 1456:JSTOR 995:(CaCO 966:algae 928:Flora 833:Algae 781:Fauna 771:ocean 759:Gulls 755:Waves 745:waves 729:clams 242:algae 230:crabs 126:Zones 75:water 71:shore 6154:Well 5853:Tarn 5728:Beel 4613:Soil 4042:silt 4032:sand 4007:clay 3736:mega 3547:Spit 3517:Reef 3447:Gulf 3395:Dune 3390:Cove 3345:Cape 3315:Ayre 2571:Pond 2141:ISSN 1973:PMID 1934:ISSN 1895:ISSN 1846:ISSN 1811:help 1763:ISSN 1723:2023 1710:ISBN 1670:ISSN 1621:PMID 1613:ISSN 1534:PMID 1516:ISSN 1464:PMID 1448:ISSN 1381:2008 1332:PMID 1314:ISSN 1246:2023 1210:NPCA 1177:2023 861:and 847:and 785:The 727:and 248:and 212:and 104:and 3522:Ria 3452:Gut 3442:Geo 3437:Gat 3325:Bay 2633:Fen 2623:Bog 2133:doi 2094:doi 2065:doi 2061:197 2020:doi 2008:172 1965:doi 1961:131 1926:doi 1885:hdl 1877:doi 1838:doi 1786:doi 1755:doi 1702:doi 1660:doi 1605:doi 1601:235 1566:doi 1524:PMC 1506:doi 1494:118 1440:doi 1322:PMC 1304:doi 974:or 182:sun 58:or 6171:: 5074:/ 4878:: 4135:: 4131:: 2139:. 2129:79 2127:. 2123:. 2100:. 2090:69 2088:. 2059:. 2053:. 2026:. 2018:. 2006:. 2002:. 1979:. 1971:. 1959:. 1955:. 1932:. 1922:28 1920:. 1916:. 1893:. 1883:. 1873:14 1871:. 1867:. 1844:. 1834:22 1832:. 1828:. 1802:: 1800:}} 1796:{{ 1784:. 1761:. 1751:33 1749:. 1745:. 1730:^ 1708:, 1696:, 1682:^ 1668:. 1656:40 1654:. 1650:. 1627:. 1619:. 1611:. 1599:. 1595:. 1572:. 1560:. 1532:. 1522:. 1514:. 1504:. 1492:. 1488:. 1476:^ 1462:. 1454:. 1446:. 1436:88 1434:. 1430:. 1353:. 1330:. 1320:. 1312:. 1302:. 1292:13 1290:. 1286:. 1262:. 1237:. 1222:^ 1208:. 1185:^ 1168:. 1156:^ 1030:), 991:. 952:. 723:, 551:• 528:• 516:• 512:• 390:) 337:, 296:, 292:, 232:, 228:, 224:, 220:, 204:, 54:A 5685:e 5678:t 5671:v 5329:K 5327:/ 5325:r 4868:e 4861:t 4854:v 4121:e 4114:t 4107:v 3271:e 3264:t 3257:v 2189:e 2182:t 2175:v 2147:. 2135:: 2108:. 2096:: 2071:. 2067:: 2051:" 2034:. 2022:: 2014:: 1987:. 1967:: 1940:. 1928:: 1901:. 1887:: 1879:: 1852:. 1840:: 1813:) 1809:( 1788:: 1769:. 1757:: 1704:: 1676:. 1662:: 1635:. 1607:: 1580:. 1568:: 1562:3 1540:. 1508:: 1500:: 1470:. 1442:: 1383:. 1338:. 1306:: 1298:: 1248:. 1179:. 1026:( 1022:) 1018:( 997:3 708:e 701:t 694:v 386:( 345:. 34:. 20:)

Index

Rockpool
Rockpool (disambiguation)

Porto Covo
Portugal
seawater
intertidal
shore
water
low tide
gravitational pull
habitats
animals
salinity
naturalists
marine biologists
philosophical
John Steinbeck
The Log from the Sea of Cortez
Intertidal zone

terrestrial
tidal range
high tides
storms
sun
organisms
flooded
wave action
currents

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