102:
1360:
143:
122:
832:"Comparative morphology of extant raptorial Mantispoidea (Neuroptera: Mantispidae, Rhachiberothidae) suggests a non-monophyletic Mantispidae and a single origin of the raptorial condition within the superfamily"
478:
455:
648:
746:
566:
489:
575:
409:
403:
382:
368:
1306:
1167:
677:
620:
560:
519:
257:
111:
667:
657:
634:
605:
594:
441:
349:
251:
245:
534:
500:
422:
687:
506:
263:
336:(beaded lacewings), they are usually considered a separate family by current authors. Another closely related group are the mantidflies (
698:
1401:
1293:
723:
777:"A new genus and species of thorny lacewing from Upper Cretaceous Kuji amber, northeastern Japan (Neuroptera, Rhachiberothidae)"
887:
1394:
142:
1430:
830:
Ardila-Camacho, Adrian; Martins, Caleb
Califre; Aspöck, Ulrike; Contreras-Ramos, Atilano (2021-06-28).
1435:
1387:
880:
1337:
1248:
101:
1420:
1332:
1324:
1311:
1280:
1375:
8:
1425:
873:
831:
807:
298:
137:
1319:
1155:
851:
812:
794:
1359:
1067:
843:
802:
784:
581:
432:
302:
283:
1196:
1075:
992:
329:
forelegs, which probably only evolved once in the common ancestor of the groups.
291:
207:
1371:
1271:
465:
847:
789:
1414:
865:
855:
798:
776:
390:
318:
59:
1220:
1183:
1175:
1096:
1054:
1046:
1025:
972:
940:
816:
704:
341:
217:
829:
1112:
1033:
1004:
337:
322:
34:
1367:
1298:
1128:
897:
733:
545:
394:
333:
294:
194:
79:
44:
1204:
1104:
1083:
1012:
964:
928:
916:
510:
469:
413:
326:
287:
174:
154:
84:
1242:
1265:
934:
737:
585:
310:
74:
69:
54:
49:
39:
1285:
1212:
314:
306:
121:
89:
64:
922:
910:
708:
345:
184:
164:
297:. The family has only 14 extant species in 4 genera found in
301:, but has a diverse fossil record extending back to the
750:
Nakamine et al. 2022 Burmese amber, Myanmar, Cenomanian
599:
Engel & Grimaldi, 2008 Burmese amber, Cenomanian
775:Nakamine, Hiroshi; Yamamoto, Shuhei (2018-12-04).
692:Aspöck, Aspöck, Johnson, Donga & Duelli, 2020
1412:
774:
1153:
895:
1395:
881:
990:
1402:
1388:
888:
874:
120:
100:
806:
788:
483:Makarkin, 2015 Burmese amber, Cenomanian
539:Whalley, 1980 Lebanese amber, Barremian
1413:
1247:
1246:
1152:
989:
953:
869:
494:Engel, 2004 Burmese amber, Cenomanian
340:), and at least some of the probably
1354:
13:
14:
1447:
614:., 2005 Lebanese amber, Barremian
554:., 2010 Lebanese amber, Barremian
528:., 2015 Burmese amber, Cenomanian
1358:
141:
32:
1086:(spongeflies or spongillaflies)
352:" also seem to be quite close.
823:
768:
505:Nakamine & Yamamoto, 2018
1:
761:
325:members of the group possess
1374:. You can help Knowledge by
7:
408:McKellar & Engel, 2009
355:
348:forms collectively called "
321:. Like the closely related
10:
1452:
1353:
509:, Japan, Late Cretaceous (
1255:
1195:
1166:
1162:
1148:
1095:
1066:
1045:
1024:
1003:
999:
985:
960:
954:
949:
905:
848:10.11646/zootaxa.4992.1.1
790:10.3897/zookeys.802.28754
662:Aspöck & Aspöck, 1997
642:Burmese amber, Cenomanian
628:Burmese amber, Cenomanian
449:Burmese amber, Cenomanian
242:
237:
138:Scientific classification
136:
128:
119:
108:
99:
23:
1207:(split-footed lacewings)
115:holotype in Oise amber
131:Uranoberotha chariessa
1370:related article is a
1333:Paleobiology Database
332:Long included in the
1107:(pleasing lacewings)
639:Nakamine et al, 2020
625:Nakamine et al, 2020
569:, France, Cenomanian
468:, Early Cretaceous (
446:Nakamine et al, 2020
435:, France, Cenomanian
387:Nakamine et al, 2020
309:(then part of North
1431:Neuroptera families
584:, Late Cretaceous (
412:, Late Cretaceous (
393:, Late Cretaceous (
278:, sometimes called
1131:(beaded lacewings)
1123:(thorny lacewings)
736:, France, Eocene (
479:Creagroparaberotha
456:Chimerhachiberotha
299:Sub-Saharan Africa
129:Reconstruction of
1383:
1382:
1348:
1347:
1320:Open Tree of Life
1249:Taxon identifiers
1240:
1239:
1236:
1235:
1232:
1231:
1224:
1216:
1208:
1187:
1179:
1178:(silky lacewings)
1156:Myrmeleontiformia
1144:
1143:
1140:
1139:
1132:
1124:
1121:Rhachiberothidae
1116:
1108:
1087:
1079:
1058:
1057:(brown lacewings)
1037:
1036:(green lacewings)
1016:
981:
980:
968:
741:
711:
693:
683:
673:
663:
653:
643:
640:
629:
626:
615:
600:
589:
570:
555:
540:
529:
514:
495:
484:
473:
450:
447:
436:
417:
398:
388:
377:
364:
361:Rhachiberothidae
273:
272:
233:
30:Barremian–Present
16:Family of insects
1443:
1436:Neuroptera stubs
1404:
1397:
1390:
1362:
1355:
1341:
1340:
1328:
1327:
1315:
1314:
1302:
1301:
1289:
1288:
1276:
1275:
1274:
1257:Rhachiberothidae
1244:
1243:
1222:
1214:
1206:
1185:
1177:
1164:
1163:
1150:
1149:
1130:
1122:
1114:
1106:
1085:
1077:
1076:Coniopterygidae
1068:Coniopterygoidea
1056:
1035:
1015:(moth lacewings)
1014:
1001:
1000:
987:
986:
966:
951:
950:
890:
883:
876:
867:
866:
860:
859:
827:
821:
820:
810:
792:
783:(802): 109–120.
772:
727:
702:
691:
681:
671:
661:
651:
649:Rhachiberothinae
641:
638:
627:
624:
609:
598:
582:New Jersey amber
580:Grimaldi, 2000
579:
564:
549:
538:
523:
504:
493:
482:
459:
448:
445:
433:Charentese amber
426:
407:
389:
386:
371:
362:
303:Early Cretaceous
280:thorny lacewings
276:Rhachiberothidae
231:
228:Rhachiberothidae
146:
145:
124:
104:
94:
31:
27:Temporal range:
24:Rhachiberothidae
21:
20:
1451:
1450:
1446:
1445:
1444:
1442:
1441:
1440:
1411:
1410:
1409:
1408:
1351:
1349:
1344:
1336:
1331:
1323:
1318:
1310:
1305:
1297:
1292:
1284:
1279:
1270:
1269:
1264:
1251:
1241:
1228:
1213:Myrmeleontidae
1197:Myrmeleontoidea
1191:
1158:
1136:
1091:
1062:
1041:
1020:
995:
993:Hemerobiiformia
977:
956:
945:
901:
894:
864:
863:
828:
824:
773:
769:
764:
758:
747:Paradoxoberotha
567:Bezonnais amber
565:SchlĂĽter, 1978
490:Eorhachiberotha
358:
267:
261:
255:
249:
230:
140:
95:
93:
92:
87:
82:
77:
72:
67:
62:
57:
52:
47:
42:
37:
29:
28:
25:
17:
12:
11:
5:
1449:
1439:
1438:
1433:
1428:
1423:
1407:
1406:
1399:
1392:
1384:
1381:
1380:
1363:
1346:
1345:
1343:
1342:
1329:
1316:
1303:
1290:
1277:
1261:
1259:
1253:
1252:
1238:
1237:
1234:
1233:
1230:
1229:
1227:
1226:
1218:
1210:
1201:
1199:
1193:
1192:
1190:
1189:
1181:
1172:
1170:
1160:
1159:
1146:
1145:
1142:
1141:
1138:
1137:
1135:
1134:
1126:
1118:
1110:
1101:
1099:
1093:
1092:
1090:
1089:
1081:
1072:
1070:
1064:
1063:
1061:
1060:
1051:
1049:
1043:
1042:
1040:
1039:
1030:
1028:
1022:
1021:
1019:
1018:
1009:
1007:
997:
996:
983:
982:
979:
978:
976:
975:
970:
961:
958:
957:
947:
946:
944:
943:
937:
931:
925:
919:
913:
906:
903:
902:
893:
892:
885:
878:
870:
862:
861:
822:
766:
765:
763:
760:
756:
755:
754:
753:
752:
751:
742:
717:Incertae sedis
714:
713:
712:
694:
684:
674:
664:
646:
645:
644:
630:
616:
601:
590:
576:Rhachibermissa
571:
556:
550:Petrulevicius
541:
530:
515:
496:
485:
474:
466:Lebanese amber
451:
437:
418:
410:Canadian amber
404:Albertoberotha
399:
383:Acanthoberotha
369:Paraberothinae
357:
354:
288:winged insects
271:
270:
240:
239:
235:
234:
225:
221:
220:
215:
211:
210:
205:
198:
197:
192:
188:
187:
182:
178:
177:
172:
168:
167:
162:
158:
157:
152:
148:
147:
134:
133:
126:
125:
117:
116:
106:
105:
97:
96:
88:
83:
78:
73:
68:
63:
58:
53:
48:
43:
38:
33:
26:
15:
9:
6:
4:
3:
2:
1448:
1437:
1434:
1432:
1429:
1427:
1424:
1422:
1419:
1418:
1416:
1405:
1400:
1398:
1393:
1391:
1386:
1385:
1379:
1377:
1373:
1369:
1364:
1361:
1357:
1356:
1352:
1339:
1334:
1330:
1326:
1321:
1317:
1313:
1308:
1304:
1300:
1295:
1291:
1287:
1282:
1278:
1273:
1267:
1263:
1262:
1260:
1258:
1254:
1250:
1245:
1225:
1221:Ascalaphidae
1219:
1217:
1211:
1209:
1203:
1202:
1200:
1198:
1194:
1188:
1184:Nemopteridae
1182:
1180:
1176:Psychopsidae
1174:
1173:
1171:
1169:
1168:Nemopteroidea
1165:
1161:
1157:
1151:
1147:
1133:
1127:
1125:
1119:
1117:
1115:(mantidflies)
1111:
1109:
1103:
1102:
1100:
1098:
1094:
1088:
1082:
1080:
1074:
1073:
1071:
1069:
1065:
1059:
1055:Hemerobiidae
1053:
1052:
1050:
1048:
1047:Hemerobioidea
1044:
1038:
1032:
1031:
1029:
1027:
1023:
1017:
1011:
1010:
1008:
1006:
1002:
998:
994:
988:
984:
974:
971:
969:
963:
962:
959:
952:
948:
942:
938:
936:
932:
930:
926:
924:
920:
918:
914:
912:
908:
907:
904:
899:
891:
886:
884:
879:
877:
872:
871:
868:
857:
853:
849:
845:
841:
837:
833:
826:
818:
814:
809:
804:
800:
796:
791:
786:
782:
778:
771:
767:
759:
749:
748:
743:
739:
735:
731:
726:
725:
720:
719:
718:
715:
710:
706:
701:
700:
695:
690:
689:
685:
680:
679:
678:Rhachiberotha
675:
670:
669:
665:
660:
659:
655:
654:
650:
647:
637:
636:
631:
623:
622:
621:Stygioberotha
617:
613:
608:
607:
602:
597:
596:
591:
587:
583:
578:
577:
572:
568:
563:
562:
561:Retinoberotha
557:
553:
548:
547:
542:
537:
536:
531:
527:
522:
521:
520:Micromantispa
516:
512:
508:
503:
502:
497:
492:
491:
486:
481:
480:
475:
471:
467:
463:
458:
457:
452:
444:
443:
438:
434:
430:
425:
424:
419:
415:
411:
406:
405:
400:
396:
392:
391:Burmese amber
385:
384:
379:
378:
375:
370:
366:
365:
360:
359:
353:
351:
347:
343:
339:
335:
330:
328:
324:
320:
319:North America
316:
312:
308:
304:
300:
296:
293:
289:
285:
281:
277:
269:
268:and see text
266:
265:
260:
259:
258:Rhachiberotha
254:
253:
248:
247:
241:
236:
229:
226:
223:
222:
219:
216:
214:Superfamily:
213:
212:
209:
206:
203:
200:
199:
196:
193:
190:
189:
186:
183:
180:
179:
176:
173:
170:
169:
166:
163:
160:
159:
156:
153:
150:
149:
144:
139:
135:
132:
127:
123:
118:
114:
113:
112:Oisea celinea
107:
103:
98:
91:
86:
81:
76:
71:
66:
61:
56:
51:
46:
41:
36:
22:
19:
1421:Mantispoidea
1376:expanding it
1365:
1350:
1256:
1186:(spoonwings)
1120:
1113:Mantispidae
1097:Mantispoidea
1078:(dustywings)
1034:Chrysopidae
1026:Chrysopoidea
973:Nevrorthidae
941:Holometabola
939:Superorder:
933:Infraclass:
839:
835:
825:
780:
770:
757:
745:
729:
722:
716:
705:Baltic amber
703:Engel, 2004
697:
686:
682:Tjeder, 1959
676:
672:Tjeder, 1959
668:Mucroberotha
666:
658:Hoelzeliella
656:
652:Tjeder, 1959
635:Uranoberotha
633:
619:
611:
606:Spinoberotha
604:
595:Scoloberotha
593:
574:
559:
551:
544:
533:
525:
518:
499:
488:
477:
461:
454:
442:Astioberotha
440:
428:
421:
402:
381:
373:
363:Tjeder, 1959
350:Mesithonidae
342:paraphyletic
331:
279:
275:
274:
262:
256:
252:Mucroberotha
250:
246:Hoelzeliella
244:
243:
232:Tjeder, 1959
227:
218:Mantispoidea
208:Euneuroptera
201:
130:
110:
18:
1129:Berothidae
1005:Ithonioidea
842:(1): 1–89.
535:Paraberotha
501:Kujiberotha
423:Alboberotha
338:Mantispidae
323:Mantispidae
1426:Neuroptera
1415:Categories
1368:Neuroptera
1223:(owlflies)
1215:(antlions)
1205:Nymphidae
1105:Dilaridae
1084:Sisyridae
1013:Ithonidae
967:(osmylids)
965:Osmylidae
927:Subclass:
917:Arthropoda
898:Neuroptera
762:References
734:Oise amber
688:Rhachiella
546:Raptorapax
507:Kuji amber
395:Cenomanian
334:Berothidae
295:Neuroptera
264:Rhachiella
195:Neuroptera
175:Arthropoda
1154:Suborder
991:Suborder
929:Pterygota
909:Kingdom:
856:1175-5334
799:1313-2970
511:Santonian
470:Barremian
464:., 2005
414:Campanian
344:group of
327:raptorial
161:Kingdom:
155:Eukaryota
1286:10642973
1272:Q7320015
1266:Wikidata
935:Neoptera
915:Phylum:
911:Animalia
900:families
817:30568531
738:Ypresian
732:., 2005
699:Whalfera
586:Turonian
431:., 2005
356:Taxonomy
311:Gondwana
282:, are a
224:Family:
171:Phylum:
165:Animalia
151:Domain:
1325:6290586
1299:8518137
923:Insecta
921:Class:
896:Extant
836:Zootaxa
808:6290048
781:ZooKeys
376:., 2005
315:Eurasia
307:Lebanon
290:in the
238:Genera
191:Order:
185:Insecta
181:Class:
1338:193525
1312:121955
854:
815:
805:
797:
709:Eocene
346:fossil
284:family
1366:This
1307:IRMNG
955:Basal
730:et al
724:Oisea
612:et al
552:et al
526:et al
462:et al
429:et al
374:et al
292:order
202:Clade
1372:stub
1294:GBIF
852:ISSN
840:4992
813:PMID
795:ISSN
728:Nel
610:Nel
524:Shi
460:Nel
427:Nel
372:Nel
317:and
109:The
35:Preęž’
1281:EoL
844:doi
803:PMC
785:doi
313:),
305:in
286:of
1417::
1335::
1322::
1309::
1296::
1283::
1268::
850:.
838:.
834:.
811:.
801:.
793:.
779:.
707:,
204::
85:Pg
1403:e
1396:t
1389:v
1378:.
889:e
882:t
875:v
858:.
846::
819:.
787::
744:â€
740:)
721:â€
696:â€
632:â€
618:â€
603:â€
592:â€
588:)
573:â€
558:â€
543:â€
532:â€
517:â€
513:)
498:â€
487:â€
476:â€
472:)
453:â€
439:â€
420:â€
416:)
401:â€
397:)
380:â€
367:â€
90:N
80:K
75:J
70:T
65:P
60:C
55:D
50:S
45:O
40:ęž’
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