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Neuroptera

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Powell, Erin. (2023). Defensive behaviors of the mealybug Nipaecoccus nipae (Maskell, 1893) (Hemiptera: Pseudococcidae) and the green lacewing Ceraeochrysa claveri (Navás, 1911) (Neuroptera: Chrysopidae), with videos of dorsal packet loading and mealybug ostiole function. Insecta Mundi. 1–11.
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The larvae are specialised predators, with elongated mandibles adapted for piercing and sucking. The larval body form varies between different families, depending on the nature of their prey. In general, however, they have three pairs of thoracic legs, each ending in two claws. The
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They have four wings, usually similar in size and shape, and a generalised pattern of veins. Some neuropterans have specialised sense organs in their wings, or have bristles or other structures to link their wings together during flight.
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Myskowiak, J.; Huang, D.; Azar, D.; Cai, C.; Garrouste, R.; Nel, A. (2016). "New lacewings (Insecta, Neuroptera, Osmylidae, Nymphidae) from the Lower Cretaceous Burmese amber and Crato Formation in Brazil".
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A phylogenomic analysis published in 2023 confirmed the topology of the neuropterid orders and found the relationships between the families of Neuropterida as shown in the following phylogenetic tree.
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Review of the Neropterid group orders by Engel, Winterton, and Breitkreuz (2018) included grouping of the Neuropteran families in a nested set of clades, an abandonment of the paraphyletic suborder "
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Bechly, G.; Makarkin, V. N. (2016). "A new gigantic lacewing species (Insecta: Neuroptera) from the Lower Cretaceous of Brazil confirms the occurrence of Kalligrammatidae in the Americas".
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composed of silk and soil or other debris. The pupa eventually cuts its way out of the cocoon with its mandibles, and may even move about for a short while before undergoing the
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Engel, M. S.; Grimaldi, D. A. (2008). "Diverse Neuropterida in Cretaceous amber, with particular reference to the paleofauna of Myanmar (Insecta)".
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The New Guinea Highland people claim to be able to maintain a muscular build and great stamina despite their low protein intake as a result of
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Engel, M. S.; Winterton, S. L.; Breitkreuz, L. C. (2018). "Phylogeny and evolution of Neuropterida: where have wings of lace taken us?".
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Engel, M. S. & Grimaldi, D. A., 2007: The neuropterid fauna of Dominican and Mexican amber (Neuropterida: Megaloptera, Neuroptera).
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Xu, X. X. (2014). "Electrophysiological and Behavior Responses of Chrysopa phyllochroma (Neuroptera Chrysopidae) to Plant Volatiles".
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Michael S. Engel (2005). "A remarkable kalligrammatid lacewing from the Upper Jurassic of Kazakhstan (Neuroptera: Kalligrammatidae)".
4664: 2485:"Tertiary giant lacewings (Neuroptera: Polystechotidae): Revision and description of new taxa from Western North America and Denmark" 4499: 2683:
Jones, J.R. (2019) Total‐evidence phylogeny of the owlflies (Neuroptera, Ascalaphidae) supports a new higher‐level classification.
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of insect pests has been investigated, showing that it is difficult to establish and maintain populations in fields of crops.
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to Neuroptera, and Raphidioptera as sister to this combined lineage, though these results were considered tentative. The
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Parker, S. P. (ed.), 1982: Synopsis and classification of living organisms. Vols. 1 & 2. McGrew-Hill Book Company
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Grimaldi, D. A. & Engel, M. S., 2005: Evolution of the Insects. Cambridge University Press, 2005, pages xv-755
4569: 4491: 2979: 204: 2391:"Mesozoic lacewings from China provide phylogenetic insight into evolution of the Kalligrammatidae (Neuroptera)" 2060: 4551: 1978: 3616: 2928:(a web portal that provides access to a suite of online resources that contain data on the order Neuroptera) 2936: 2541:"High morphological disparity of neuropteran larvae during the Cretaceous revealed by a new large species" 4408: 4385: 2332: 445:, which bury themselves completely out of sight and ambush prey from "pits" in the soil. Larvae of some 2972: 2922:(an online bibliography that includes data on the global scientific literature of the order Neuroptera) 1990: 583: 330:
Neuropterans are soft-bodied insects with relatively few specialized features. They have large lateral
130: 4618: 4413: 2216:"Los hemeróbidos de la Península Ibérica y Baleares (Insecta, Neuropterida, Neuroptera: Hemerobiidae)" 4631: 4579: 2702:"Improved modelling of compositional heterogeneity reconciles phylogenomic conflicts among lacewings" 569: 563: 2880: 2200: 660:
Following the end of the Cretaceous period, the diversity of neuropterans appears to have declined.
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Larvae in various families cover themselves in debris (including other insects, living and dead) as
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Molecular analysis in 2018 using mitochondrial rRNA and mitogenomic data places the Megaloptera as
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record has contributed to the understanding of the group's phylogeny. Relationships within the
2333:"The neuropterid fauna of Dominican and Mexican amber (Neuropterida, Megaloptera, Neuroptera)" 4669: 4600: 2940: 2311: 4530: 4592: 4447: 4376: 2599: 2552: 2498: 2457: 2402: 2363: 1258: 339: 2083: 1911:
are treated as a sister group to clade Eidoneuroptera formed by Neuroptera + Megaloptera.
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Neuropterida Species of the World. Lacewing Digital Library, Research Publication No. 1.
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Bibliography of the Neuropterida. Lacewing Digital Library, Research Publication No. 2.
2868: 2831: 2757: 2723: 2630: 2587: 2514: 2425: 2390: 2294: 2269: 2188: 2141: 2099: 2064: 1920: 1651: 431: 125: 1752:: moth lacewings (includes Rapismatidae and Polystoechotidae) (Early Jurassic-Present) 4587: 4434: 4245: 3857: 2854: 2823: 2815: 2761: 2727: 2635: 2617: 2568: 2430: 2299: 2237: 2174: 2127: 1994: 1759: 1186: 677: 578: 307:. During this time, several unusually large forms evolved, especially in the extinct 2835: 2718: 2701: 2518: 2103: 2068: 2019:"New lacewings (Neuroptera) from the terminal Permian and basal Triassic of Siberia" 1926:
Five species of Neuroptera are among 1681 insect species eaten by humans worldwide.
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to Recent, and the extinct lineages of the family were widespread geographically.
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origin and may be the most ancient of the Neuropteran groups. The extinct osmylid
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Adult neuropterans have four membranous wings, all about the same size, with many
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Adults of many groups are also predatory, but some do not feed, or consume only
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suitable for chewing, and lack the various adaptations found in most other
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Cai, Chen-Yang; Tihelka, Erik; Liu, Xing-YUE; Engel, Michael S. (2023).
1139: 110: 4218: 3852: 3386: 3295: 3279: 3263: 3091: 1810:: split-footed lacewings (includes Myiodactylidae) (Cretaceous-Present) 1640: 1616: 1571: 1162: 1065: 920: 810: 627: 610: 512: 509: 461: 454: 438: 263: 82: 47: 2285: 4517: 4460: 4294: 4194: 4173: 4102: 4054: 4018: 4006: 3814: 3658: 3618: 3593: 3446: 3357: 3316: 3303: 3134: 3121: 3016: 1977: 1897:
from the Late Jurassic-Early Cretaceous of China has been treated as
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Yang, Q.; Wang, Y.; Labandeira, C.C.; Shih, C.; Ren, D. (2014).
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Yan, Y.; Wang Y, Liu, X.; Winterton, S. L.; Yang, D. (2014).
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Breitkreuz, L. C. V.; Winterton, S. L.; Engel, M. S. (2017).
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Luo, Cihang; Liu, Hua; Jarzembowski, Edmund A. (June 2022).
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Consuming the Inedible: Neglected Dimensions of Food Choice
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Yue, Bi-Song; Song, Nan; Lin, Aili; Zhao, Xincheng (2018).
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10.1660/0022-8443(2005)108[0059:ARKLFT]2.0.CO;2
505: 480: 1796:: antlions (includes Palaeoleontidae) (Cretaceous-Present) 3636: 2382: 2743: 2122:
Hoell, H. V., Doyen, J. T. & Purcell, A. H. (1998).
2081: 254:, and their relatives. The order consists of some 6,000 3579:(net-winged insects: lacewings, mantidflies, antlions) 2849:
Ramos-Elorduy, J. (2005). Maurizio G. Paoletti (ed.).
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Introduction to Insect Biology and Diversity, 2nd ed
1537:" and redefinition of Myrmeleontiformia as a clade. 358:often has adhesive discs on the last two segments. 2699: 2538: 2017:A. G. Ponomarenko & D. E. Shcherbakov (2004). 2012: 2010: 4646: 2165:. Cambridge University Press. pp. 296–302. 2044: 551:Neuropterans first appeared near the end of the 2007: 4243: 3985: 2585: 2447: 2331:Engel, Michael S.; Grimaldi, David A. (2007). 1903:within Neuroptera, while the fossil families † 1626:: pleasing lacewings (Late Cretaceous-Present) 3971: 3416: 2980: 2848: 2476: 2346: 2126:. Oxford University Press. pp. 447–450. 2048:Transactions of the Kansas Academy of Science 1672:: thorny lacewings (Early Cretaceous-Present) 4080: 2932:Illustrated database of Neuroptera (insects) 2316:: CS1 maint: multiple names: authors list ( 2160: 2146:: CS1 maint: multiple names: authors list ( 3248: 3056: 2945:Institute of Food and Agricultural Sciences 2579: 3978: 3964: 3162: 2987: 2973: 2441: 2163:The use of lacewings in biological control 2161:Senior, L. J.; McEwen, P. K. (June 2001). 1973: 1971: 1643:: beaded lacewings (Late Jurassic-Present) 1588:: spongillaflies (Late Cretaceous-Present) 258:. Neuroptera is grouped together with the 109: 2893: 2717: 2629: 2611: 2424: 2414: 2293: 2231: 2213: 1874:: silky lacewings (Late Triassic-Present) 361: 303:, and continued to diversify through the 3708:Four most speciose orders are marked in 3298:(crickets, wetas, grasshoppers, locusts) 2739: 2737: 568: 530: 517: 388: 369: 2483:Archibald, S.B.; Makarkin V.N. (2006). 1968: 296:Neuropterans first appeared during the 4647: 2117: 2115: 2113: 325: 4337: 4336: 4242: 4079: 4043: 3959: 3746: 3041: 2968: 2734: 2263: 2261: 1803:: spoonwings etc (Cretaceous-Present) 4619:F34387BF-FFE5-6755-B3E5-F943FDFB4979 4580:74fe3b22-3f4e-4ab5-9ee6-158fd6324382 4492:fc32b888-ec97-4d33-a88e-1990af11f6c9 3571:(alderflies, dobsonflies, fishflies) 1717:: brown lacewings (Jurassic-Present) 566:and a similar fauna from Australia. 322:" for their large, patterned wings. 2490:Journal of Systematic Palaeontology 2214:Monserrat, Víctor J. (2015-12-30). 2110: 1914: 1581:: osmylids (Early Jurassic-Present) 278:(snakeflies) in the unranked taxon 13: 2797: 2758:10.1146/annurev-ento-020117-043127 2258: 1558:dustywings (Late Jurassic-Present) 483:stage is enclosed in some form of 14: 4681: 2994: 2905: 2689:https://doi.org/10.1111/zsc.12382 626:is described from two fossils of 4665:Extant Permian first appearances 3930: 3081: 1370: 1337: 1297: 1257: 1214: 1172: 1138: 1026: 972: 827: 736: 708: 449:are root feeders, and larvae of 410:The larvae of most families are 129: 35: 4176:(spongeflies or spongillaflies) 2887: 2842: 2791: 2772: 2719:10.11646/palaeoentomology.6.1.8 2693: 2677: 2668: 2655: 2646: 2532: 2324: 1943: 535:One of the "butterflies of the 338:. Their mouthparts have strong 334:, and may or may not also have 2248: 2233:10.3989/graellsia.2015.v71.129 2207: 2154: 2075: 2038: 1706:: green lacewings (including † 430:, and some have been used for 1: 3501:(sawflies, wasps, ants, bees) 2781:Nova Supplementa Entomologica 2470:10.1016/j.cretres.2015.10.014 2376:10.1016/j.cretres.2015.10.029 1962: 558:, as shown by fossils of the 441:, taken to an extreme in the 282:(once known as Planipennia). 219: 3449:(cicadas, aphids, true bugs) 2613:10.1371/journal.pone.0191826 2171:10.1017/cbo9780511666117.014 663: 650:(Lower Cretaceous) periods. 526: 7: 3747: 2851:Insects: a hopeful resource 2746:Annual Review of Entomology 1528: 646:lived from the Jurassic to 10: 4686: 3639:(gnats, mosquitoes, flies) 3532:(twisted-winged parasites) 3204:(dragonflies, damselflies) 3042: 2937:Brown lacewings of Florida 2894:MacClancy, Jeremy (2007). 1991:Cambridge University Press 1839:(Jurassic-Late Cretaceous) 1650:: mantidflies (including † 584:Palaeopsychops marringerae 4345: 4285: 4256: 4252: 4238: 4185: 4156: 4135: 4114: 4093: 4089: 4075: 4050: 4044: 4039: 3995: 3924: 3757: 3753: 3742: 3704: 3649: 3607: 3592: 3551: 3520: 3511: 3493: 3462: 3412: 3385: 3348: 3315: 3259: 3244: 3221: 3192: 3171: 3158: 3133: 3117: 3090: 3079: 3052: 3048: 3037: 3005: 2953:Information on Neuroptera 2663:American Museum Novitates 2565:10.1017/S0016756822000176 2511:10.1017/S1477201906001817 2340:American Museum Novitates 2088:American Museum Novitates 1919:The use of Neuroptera in 1595:(Late Cretaceous-Present) 1356: 1323: 1316: 1283: 1276: 1252:(split-footed lacewings) 1243: 1236: 1200: 1190: 1166: 1156: 1132: 1125: 1115: 1063: 1048: 1041: 1020: 1013: 997: 990: 966: 956: 949: 924: 914: 907: 864: 849: 842: 821: 814: 804: 797: 778: 768: 761: 751: 730: 723: 702: 695: 564:Tunguska basin in Siberia 453:are aquatic, and feed on 216: 211: 126:Scientific classification 124: 117: 108: 23: 4297:(split-footed lacewings) 3290:(stick and leaf insects) 2926:Lacewing Digital Library 2918:Oswald, John D. (2023). 2911:Oswald, John D. (2023). 2800:Environmental Entomology 2416:10.1186/1471-2148-14-126 2395:BMC Evolutionary Biology 1987:Evolution of the Insects 1936: 735:(alderflies and allies) 3717:are paraphyletic groups 3360:(cockroaches, termites) 3124:(silverfish, firebrats) 2956:(subscription required) 2027:Paleontological Journal 653:Ithonidae are from the 642:or earlier. The family 3068:(jumping bristletails) 1933:including Neuroptera. 592: 548: 523: 522:Lifecycle of lacewings 403: 384: 362:Life cycle and ecology 4601:Paleobiology Database 2941:University of Florida 929:(pleasing lacewings) 572: 534: 521: 426:eat aphids and other 392: 377:Osmylus fulvicephalus 373: 4487:Fauna Europaea (new) 4197:(pleasing lacewings) 3670:(moths, butterflies) 1949:"Neurorthidae" is a 1850:†Brongniartiellidae) 1710:) (Jurassic-Present) 1654:) (Jurassic-Present) 595:The osmylids are of 515:neuropteran larvae. 314:, often called "the 2665:: #3587, pages 1-58 2604:2018PLoSO..1391826S 2557:2022GeoM..159..954L 2545:Geological Magazine 2503:2006JSPal...4..119A 2462:2016CrRes..58..135B 2450:Cretaceous Research 2407:2014BMCEE..14..126Y 2368:2016CrRes..59..214M 2356:Cretaceous Research 1068:(beaded lacewings) 1002:(thorny lacewings) 680:are still in flux. 491:to the adult form. 326:Anatomy and biology 4221:(beaded lacewings) 4213:(thorny lacewings) 3937:Insects portal 3908:Triadophlebioptera 2948:Featured Creatures 2687:: 06 October 2019 1921:biological control 1888:The fossil genus † 1652:Dipteromantispidae 1602:(Permian-Triassic) 1209:(silky lacewings) 1137:(brown lacewings) 971:(green lacewings) 826:(giant lacewings) 593: 573:Patterned wing of 549: 524: 432:biological control 404: 385: 240:net-winged insects 4642: 4641: 4588:Open Tree of Life 4339:Taxon identifiers 4330: 4329: 4326: 4325: 4322: 4321: 4314: 4306: 4298: 4277: 4269: 4268:(silky lacewings) 4246:Myrmeleontiformia 4234: 4233: 4230: 4229: 4222: 4214: 4211:Rhachiberothidae 4206: 4198: 4177: 4169: 4148: 4147:(brown lacewings) 4127: 4126:(green lacewings) 4106: 4071: 4070: 4058: 3953: 3952: 3920: 3919: 3916: 3915: 3858:Palaeodictyoptera 3843: 3775: 3738: 3737: 3734: 3733: 3700: 3699: 3696: 3695: 3692: 3691: 3688: 3687: 3684: 3683: 3680: 3679: 3671: 3662: 3640: 3630: 3622: 3600: 3588: 3587: 3580: 3572: 3564: 3542: 3533: 3502: 3458: 3457: 3450: 3442: 3434: 3381: 3380: 3377: 3376: 3369: 3361: 3340: 3337:Mantophasmatodea 3332: 3321: 3307: 3299: 3291: 3283: 3275: 3267: 3217: 3216: 3213: 3212: 3205: 3184: 3173:Ephemeropteroidea 3125: 3077: 3076: 3069: 2812:10.1093/ee/nvv106 2685:Zoologica Scripta 2286:10.7150/ijbs.9454 1844:Osmylopsychopidae 1831:Aetheogrammatidae 1760:Myrmeleontiformia 1609:(Middle Jurassic) 1522: 1521: 1513: 1512: 1504: 1503: 1495: 1494: 1486: 1485: 1477: 1476: 1468: 1467: 1459: 1458: 1450: 1449: 1441: 1440: 1432: 1431: 1423: 1422: 1414: 1413: 1405: 1404: 1396: 1395: 1387: 1386: 1379: 1378: 1346: 1345: 1306: 1305: 1266: 1265: 1223: 1222: 1187:Myrmeleontiformia 1171:(moth lacewings) 1104: 1103: 1095: 1094: 1086: 1085: 1077: 1076: 938: 937: 896: 895: 887: 886: 878: 877: 869:(spongillaflies) 678:Myrmeleontiformia 638:radiation in the 408: 407: 226: 225: 207: 103: 4677: 4635: 4634: 4622: 4621: 4609: 4608: 4596: 4595: 4583: 4582: 4573: 4572: 4560: 4559: 4557:NHMSYS0000841369 4547: 4546: 4534: 4533: 4521: 4520: 4508: 4507: 4495: 4494: 4482: 4481: 4469: 4468: 4456: 4455: 4443: 4442: 4430: 4429: 4417: 4416: 4404: 4403: 4394: 4393: 4381: 4380: 4379: 4366: 4365: 4364: 4334: 4333: 4312: 4304: 4296: 4275: 4267: 4254: 4253: 4240: 4239: 4220: 4212: 4204: 4196: 4175: 4167: 4166:Coniopterygidae 4158:Coniopterygoidea 4146: 4125: 4105:(moth lacewings) 4104: 4091: 4090: 4077: 4076: 4056: 4041: 4040: 3980: 3973: 3966: 3957: 3956: 3935: 3934: 3868:Permoplectoptera 3841: 3800:Diaphanopterodea 3773: 3755: 3754: 3744: 3743: 3669: 3660: 3651:Amphiesmenoptera 3638: 3632: 3628: 3620: 3610: 3605: 3604: 3599: 3596: 3578: 3570: 3562: 3540: 3531: 3518: 3517: 3514: 3500: 3491: 3490: 3448: 3440: 3433:(barklice, lice) 3432: 3424: 3414: 3413: 3410: 3409: 3367: 3359: 3338: 3330: 3329:Grylloblattodea 3319: 3305: 3297: 3289: 3281: 3273: 3265: 3257: 3256: 3253: 3246: 3245: 3242: 3241: 3203: 3182: 3169: 3168: 3160: 3159: 3156: 3155: 3123: 3115: 3114: 3085: 3067: 3054: 3053: 3050: 3049: 3039: 3038: 2989: 2982: 2975: 2966: 2965: 2957: 2900: 2899: 2891: 2885: 2884: 2878: 2874: 2872: 2864: 2846: 2840: 2839: 2806:(5): 1425–1433. 2795: 2789: 2788: 2776: 2770: 2769: 2741: 2732: 2731: 2721: 2706:Palaeoentomology 2697: 2691: 2681: 2675: 2672: 2666: 2659: 2653: 2650: 2644: 2643: 2633: 2615: 2583: 2577: 2576: 2536: 2530: 2529: 2527: 2525: 2480: 2474: 2473: 2445: 2439: 2438: 2428: 2418: 2386: 2380: 2379: 2350: 2344: 2343: 2337: 2328: 2322: 2321: 2315: 2307: 2297: 2265: 2256: 2252: 2246: 2245: 2235: 2211: 2205: 2204: 2198: 2194: 2192: 2184: 2158: 2152: 2151: 2145: 2137: 2119: 2108: 2107: 2079: 2073: 2072: 2042: 2036: 2035: 2034:(S2): S197–S203. 2023: 2014: 2005: 2004: 1983:Michael S. Engel 1975: 1956: 1947: 1915:In human culture 1896: 1837:Kalligrammatidae 1670:Rhachiberothidae 1549:Coniopterygoidea 1374: 1358: 1357: 1341: 1325: 1324: 1319: 1318: 1301: 1285: 1284: 1279: 1278: 1261: 1245: 1244: 1239: 1238: 1218: 1202: 1201: 1193: 1192: 1176: 1159: 1158: 1142: 1128: 1127: 1118: 1117: 1044: 1043: 1030: 1016: 1015: 1000:Rhachiberothidae 993: 992: 976: 959: 958: 952: 951: 917: 916: 910: 909: 845: 844: 831: 817: 816: 807: 806: 800: 799: 775:Coniopterygoidea 771: 770: 764: 763: 754: 753: 740: 726: 725: 712: 698: 697: 688: 687: 644:Kalligrammatidae 640:Early Cretaceous 601:Early Cretaceous 545:Kalligrammatidae 366: 365: 312:Kalligrammatidae 199: 134: 133: 113: 98: 97: 34: 27:Temporal range: 21: 20: 16:Order of insects 4685: 4684: 4680: 4679: 4678: 4676: 4675: 4674: 4645: 4644: 4643: 4638: 4630: 4625: 4617: 4612: 4604: 4599: 4591: 4586: 4578: 4576: 4568: 4563: 4555: 4550: 4542: 4537: 4529: 4524: 4516: 4511: 4503: 4498: 4490: 4485: 4477: 4472: 4464: 4459: 4451: 4446: 4438: 4433: 4425: 4420: 4412: 4407: 4399: 4397: 4389: 4384: 4375: 4374: 4369: 4360: 4359: 4354: 4341: 4331: 4318: 4303:Myrmeleontidae 4287:Myrmeleontoidea 4281: 4248: 4226: 4181: 4152: 4131: 4110: 4085: 4083:Hemerobiiformia 4067: 4046: 4035: 3991: 3984: 3954: 3949: 3929: 3912: 3878:Protelytroptera 3810:Eudiaphanoptera 3795:Coxoplectoptera 3790:Carbotriplurida 3785:Campylopteridae 3762:Aethiocarenodea 3749: 3730: 3676: 3645: 3621:(scorpionflies) 3608: 3597: 3595: 3584: 3547: 3512: 3507: 3485: 3483: 3481: 3479: 3477: 3475: 3473: 3471: 3469: 3467: 3465: 3454: 3404: 3402: 3400: 3398: 3396: 3394: 3392: 3390: 3388: 3373: 3344: 3318: 3311: 3306:(angel insects) 3249: 3236: 3234: 3232: 3230: 3228: 3226: 3224: 3209: 3188: 3150: 3148: 3146: 3144: 3142: 3140: 3138: 3136: 3129: 3109: 3107: 3105: 3103: 3101: 3099: 3097: 3095: 3093: 3086: 3073: 3044: 3033: 3001: 2993: 2955: 2908: 2903: 2892: 2888: 2876: 2875: 2866: 2865: 2861: 2847: 2843: 2796: 2792: 2777: 2773: 2742: 2735: 2698: 2694: 2682: 2678: 2673: 2669: 2660: 2656: 2651: 2647: 2598:(1): e0191826. 2584: 2580: 2537: 2533: 2523: 2521: 2481: 2477: 2446: 2442: 2387: 2383: 2351: 2347: 2335: 2329: 2325: 2309: 2308: 2266: 2259: 2253: 2249: 2212: 2208: 2196: 2195: 2186: 2185: 2181: 2159: 2155: 2139: 2138: 2134: 2120: 2111: 2080: 2076: 2043: 2039: 2021: 2015: 2008: 2001: 1976: 1969: 1965: 1960: 1959: 1948: 1944: 1939: 1917: 1905:Permoberothidae 1894: 1866:Prohemerobiidae 1768:Myrmeleontoidea 1708:Mesochrysopidae 1698:Ascalochrysidae 1556:Coniopterygidae 1535:Hemerobiiformia 1531: 1525: 1523: 1514: 1505: 1496: 1487: 1478: 1469: 1460: 1451: 1442: 1433: 1424: 1415: 1406: 1397: 1388: 1233:Myrmeleontoidea 1105: 1096: 1087: 1078: 939: 897: 888: 879: 781:Coniopterygidae 683: 666: 541:Sophogramma lii 529: 364: 346:insect groups. 328: 242:, includes the 198: 128: 104: 96: 95: 90: 85: 80: 75: 70: 65: 60: 55: 50: 45: 40: 29: 28: 25: 17: 12: 11: 5: 4683: 4673: 4672: 4667: 4662: 4657: 4640: 4639: 4637: 4636: 4623: 4610: 4597: 4584: 4574: 4561: 4548: 4535: 4522: 4509: 4496: 4483: 4474:Fauna Europaea 4470: 4457: 4444: 4431: 4418: 4405: 4395: 4382: 4367: 4351: 4349: 4343: 4342: 4328: 4327: 4324: 4323: 4320: 4319: 4317: 4316: 4308: 4300: 4291: 4289: 4283: 4282: 4280: 4279: 4271: 4262: 4260: 4250: 4249: 4236: 4235: 4232: 4231: 4228: 4227: 4225: 4224: 4216: 4208: 4200: 4191: 4189: 4183: 4182: 4180: 4179: 4171: 4162: 4160: 4154: 4153: 4151: 4150: 4141: 4139: 4133: 4132: 4130: 4129: 4120: 4118: 4112: 4111: 4109: 4108: 4099: 4097: 4087: 4086: 4073: 4072: 4069: 4068: 4066: 4065: 4060: 4051: 4048: 4047: 4037: 4036: 4034: 4033: 4027: 4021: 4015: 4009: 4003: 3996: 3993: 3992: 3983: 3982: 3975: 3968: 3960: 3951: 3950: 3948: 3947: 3940: 3925: 3922: 3921: 3918: 3917: 3914: 3913: 3911: 3910: 3905: 3900: 3895: 3893:Protozygoptera 3890: 3888:Protorthoptera 3885: 3883:Protephemerida 3880: 3875: 3873:Protanisoptera 3870: 3865: 3860: 3855: 3850: 3845: 3842:(griffinflies) 3840:Meganisoptera 3837: 3832: 3827: 3822: 3820:Glosselytrodea 3817: 3812: 3807: 3802: 3797: 3792: 3787: 3782: 3777: 3769: 3764: 3758: 3751: 3750: 3740: 3739: 3736: 3735: 3732: 3731: 3729: 3728: 3718: 3712: 3705: 3702: 3701: 3698: 3697: 3694: 3693: 3690: 3689: 3686: 3685: 3682: 3681: 3678: 3677: 3675: 3674: 3664: 3655: 3653: 3647: 3646: 3644: 3643: 3633: 3613: 3611: 3602: 3590: 3589: 3586: 3585: 3583: 3582: 3574: 3566: 3561:Raphidioptera 3557: 3555: 3549: 3548: 3546: 3545: 3535: 3526: 3524: 3515: 3513:Neuropteroidea 3509: 3508: 3506: 3505: 3494: 3488: 3460: 3459: 3456: 3455: 3453: 3452: 3444: 3436: 3427: 3425: 3407: 3383: 3382: 3379: 3378: 3375: 3374: 3372: 3371: 3363: 3354: 3352: 3346: 3345: 3343: 3342: 3334: 3331:(ice-crawlers) 3325: 3323: 3313: 3312: 3310: 3309: 3301: 3293: 3285: 3277: 3269: 3260: 3254: 3239: 3219: 3218: 3215: 3214: 3211: 3210: 3208: 3207: 3198: 3196: 3190: 3189: 3187: 3186: 3181:Ephemeroptera 3177: 3175: 3166: 3153: 3131: 3130: 3128: 3127: 3118: 3112: 3088: 3087: 3080: 3078: 3075: 3074: 3072: 3071: 3066:Archaeognatha 3062: 3060: 3046: 3045: 3035: 3034: 3032: 3031: 3025: 3019: 3013: 3006: 3003: 3002: 2992: 2991: 2984: 2977: 2969: 2963: 2962: 2960:Web of Science 2950: 2934: 2929: 2923: 2916: 2907: 2906:External links 2904: 2902: 2901: 2886: 2860:978-1578083398 2859: 2841: 2790: 2771: 2733: 2692: 2676: 2667: 2654: 2645: 2578: 2551:(6): 954–962. 2531: 2497:(2): 119–155. 2475: 2440: 2381: 2345: 2323: 2280:(8): 895–908. 2274:Int J Biol Sci 2257: 2247: 2222:(in Spanish). 2206: 2180:978-0511666117 2179: 2153: 2132: 2109: 2096:10.1206/3890.1 2090:(3890): 1–44. 2074: 2037: 2006: 1999: 1979:David Grimaldi 1966: 1964: 1961: 1958: 1957: 1941: 1940: 1938: 1935: 1931:eating insects 1916: 1913: 1900:incertae sedis 1891:Mesohemerobius 1886: 1885: 1884: 1883: 1882: 1881: 1880: 1879: 1878: 1877: 1876: 1875: 1868: 1862: 1859:Grammosmylidae 1851: 1840: 1833: 1819: 1818: 1817: 1811: 1804: 1797: 1794:Myrmeleontidae 1790: 1783: 1755: 1754: 1753: 1732: 1731: 1730: 1728:Solenoptilidae 1724: 1718: 1711: 1700: 1675: 1674: 1673: 1666: 1659:Mesoberothidae 1655: 1644: 1629: 1628: 1627: 1612: 1611: 1610: 1607:Saucrosmylidae 1603: 1600:Archeosmylidae 1596: 1589: 1582: 1561: 1560: 1559: 1530: 1527: 1520: 1519: 1516: 1515: 1511: 1510: 1507: 1506: 1502: 1501: 1498: 1497: 1493: 1492: 1489: 1488: 1484: 1483: 1480: 1479: 1475: 1474: 1471: 1470: 1466: 1465: 1462: 1461: 1457: 1456: 1453: 1452: 1448: 1447: 1444: 1443: 1439: 1438: 1435: 1434: 1430: 1429: 1426: 1425: 1421: 1420: 1417: 1416: 1412: 1411: 1408: 1407: 1403: 1402: 1399: 1398: 1394: 1393: 1390: 1389: 1385: 1384: 1381: 1380: 1377: 1376: 1367: 1355: 1352: 1351: 1348: 1347: 1344: 1343: 1334: 1330:Myrmeleontidae 1322: 1317: 1315: 1312: 1311: 1308: 1307: 1304: 1303: 1294: 1282: 1277: 1275: 1272: 1271: 1268: 1267: 1264: 1263: 1254: 1242: 1237: 1235: 1229: 1228: 1225: 1224: 1221: 1220: 1211: 1199: 1191: 1189: 1183: 1182: 1179: 1178: 1165: 1157: 1155: 1149: 1148: 1145: 1144: 1131: 1126: 1124: 1116: 1114: 1111: 1110: 1107: 1106: 1102: 1101: 1098: 1097: 1093: 1092: 1089: 1088: 1084: 1083: 1080: 1079: 1075: 1074: 1071: 1070: 1062: 1059: 1058: 1055: 1054: 1047: 1042: 1040: 1037: 1036: 1033: 1032: 1025:(mantidflies) 1019: 1014: 1012: 1009: 1008: 1005: 1004: 996: 991: 989: 983: 982: 979: 978: 965: 957: 955: 950: 948: 945: 944: 941: 940: 936: 935: 932: 931: 923: 915: 913: 908: 906: 903: 902: 899: 898: 894: 893: 890: 889: 885: 884: 881: 880: 876: 875: 872: 871: 863: 860: 859: 856: 855: 848: 843: 841: 838: 837: 834: 833: 820: 815: 813: 805: 803: 798: 796: 790: 789: 786: 785: 777: 769: 767: 762: 760: 752: 750: 747: 746: 743: 742: 729: 724: 722: 719: 718: 715: 714: 701: 696: 694: 686: 665: 662: 528: 525: 424:coniopterygids 406: 405: 386: 363: 360: 327: 324: 224: 223: 214: 213: 209: 208: 193: 189: 188: 183: 176: 175: 170: 166: 165: 160: 156: 155: 150: 146: 145: 140: 136: 135: 122: 121: 119:Green lacewing 115: 114: 106: 105: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 41: 36: 26: 15: 9: 6: 4: 3: 2: 4682: 4671: 4668: 4666: 4663: 4661: 4660:Insect orders 4658: 4656: 4653: 4652: 4650: 4633: 4628: 4624: 4620: 4615: 4611: 4607: 4602: 4598: 4594: 4589: 4585: 4581: 4575: 4571: 4566: 4562: 4558: 4553: 4549: 4545: 4540: 4536: 4532: 4527: 4523: 4519: 4514: 4510: 4506: 4501: 4497: 4493: 4488: 4484: 4480: 4475: 4471: 4467: 4462: 4458: 4454: 4449: 4445: 4441: 4436: 4432: 4428: 4423: 4419: 4415: 4410: 4406: 4402: 4396: 4392: 4387: 4383: 4378: 4372: 4368: 4363: 4357: 4353: 4352: 4350: 4348: 4344: 4340: 4335: 4315: 4311:Ascalaphidae 4309: 4307: 4301: 4299: 4293: 4292: 4290: 4288: 4284: 4278: 4274:Nemopteridae 4272: 4270: 4266:Psychopsidae 4264: 4263: 4261: 4259: 4258:Nemopteroidea 4255: 4251: 4247: 4241: 4237: 4223: 4217: 4215: 4209: 4207: 4205:(mantidflies) 4201: 4199: 4193: 4192: 4190: 4188: 4184: 4178: 4172: 4170: 4164: 4163: 4161: 4159: 4155: 4149: 4145:Hemerobiidae 4143: 4142: 4140: 4138: 4137:Hemerobioidea 4134: 4128: 4122: 4121: 4119: 4117: 4113: 4107: 4101: 4100: 4098: 4096: 4092: 4088: 4084: 4078: 4074: 4064: 4061: 4059: 4053: 4052: 4049: 4042: 4038: 4032: 4028: 4026: 4022: 4020: 4016: 4014: 4010: 4008: 4004: 4002: 3998: 3997: 3994: 3989: 3981: 3976: 3974: 3969: 3967: 3962: 3961: 3958: 3946: 3945: 3941: 3939: 3938: 3933: 3927: 3926: 3923: 3909: 3906: 3904: 3901: 3899: 3896: 3894: 3891: 3889: 3886: 3884: 3881: 3879: 3876: 3874: 3871: 3869: 3866: 3864: 3861: 3859: 3856: 3854: 3851: 3849: 3848:Megasecoptera 3846: 3844: 3838: 3836: 3833: 3831: 3828: 3826: 3823: 3821: 3818: 3816: 3813: 3811: 3808: 3806: 3803: 3801: 3798: 3796: 3793: 3791: 3788: 3786: 3783: 3781: 3778: 3776: 3770: 3768: 3765: 3763: 3760: 3759: 3756: 3752: 3745: 3741: 3727: 3725: 3719: 3716: 3713: 3711: 3707: 3706: 3703: 3673: 3672: 3665: 3663: 3661:(caddisflies) 3657: 3656: 3654: 3652: 3648: 3642: 3641: 3634: 3631: 3627:Siphonaptera 3624: 3623: 3615: 3614: 3612: 3606: 3603: 3601: 3598:(Mecopterida) 3591: 3581: 3575: 3573: 3567: 3565: 3559: 3558: 3556: 3554: 3550: 3544: 3543: 3536: 3534: 3530:Strepsiptera 3528: 3527: 3525: 3523: 3519: 3516: 3510: 3504: 3503: 3496: 3495: 3492: 3489: 3487: 3461: 3451: 3445: 3443: 3439:Thysanoptera 3437: 3435: 3429: 3428: 3426: 3421: 3420: 3415: 3411: 3408: 3406: 3384: 3370: 3364: 3362: 3356: 3355: 3353: 3351: 3347: 3341: 3335: 3333: 3327: 3326: 3324: 3322: 3314: 3308: 3302: 3300: 3294: 3292: 3286: 3284: 3282:(webspinners) 3278: 3276: 3270: 3268: 3262: 3261: 3258: 3255: 3252: 3247: 3243: 3240: 3238: 3220: 3206: 3200: 3199: 3197: 3195: 3191: 3185: 3179: 3178: 3176: 3174: 3170: 3167: 3165: 3161: 3157: 3154: 3152: 3132: 3126: 3120: 3119: 3116: 3113: 3111: 3089: 3084: 3070: 3064: 3063: 3061: 3059: 3055: 3051: 3047: 3040: 3036: 3030: 3026: 3024: 3020: 3018: 3014: 3012: 3008: 3007: 3004: 3000: 2997: 2990: 2985: 2983: 2978: 2976: 2971: 2970: 2967: 2961: 2954: 2951: 2949: 2946: 2942: 2938: 2935: 2933: 2930: 2927: 2924: 2921: 2917: 2914: 2910: 2909: 2897: 2890: 2882: 2870: 2862: 2856: 2852: 2845: 2837: 2833: 2829: 2825: 2821: 2817: 2813: 2809: 2805: 2801: 2794: 2786: 2782: 2775: 2767: 2763: 2759: 2755: 2751: 2747: 2740: 2738: 2729: 2725: 2720: 2715: 2711: 2707: 2703: 2696: 2690: 2686: 2680: 2671: 2664: 2658: 2649: 2641: 2637: 2632: 2627: 2623: 2619: 2614: 2609: 2605: 2601: 2597: 2593: 2589: 2582: 2574: 2570: 2566: 2562: 2558: 2554: 2550: 2546: 2542: 2535: 2520: 2516: 2512: 2508: 2504: 2500: 2496: 2492: 2491: 2486: 2479: 2471: 2467: 2463: 2459: 2455: 2451: 2444: 2436: 2432: 2427: 2422: 2417: 2412: 2408: 2404: 2400: 2396: 2392: 2385: 2377: 2373: 2369: 2365: 2361: 2357: 2349: 2342:(3587): 1–58. 2341: 2334: 2327: 2319: 2313: 2305: 2301: 2296: 2291: 2287: 2283: 2279: 2275: 2271: 2264: 2262: 2251: 2243: 2239: 2234: 2229: 2225: 2221: 2217: 2210: 2202: 2190: 2182: 2176: 2172: 2168: 2164: 2157: 2149: 2143: 2135: 2133:0-19-510033-6 2129: 2125: 2118: 2116: 2114: 2105: 2101: 2097: 2093: 2089: 2085: 2078: 2070: 2066: 2062: 2058: 2054: 2050: 2049: 2041: 2033: 2029: 2028: 2020: 2013: 2011: 2002: 2000:0-521-82149-5 1996: 1992: 1988: 1984: 1980: 1974: 1972: 1967: 1954: 1953: 1946: 1942: 1934: 1932: 1927: 1924: 1922: 1912: 1910: 1909:Permithonidae 1906: 1902: 1901: 1893: 1892: 1873: 1869: 1867: 1863: 1860: 1856: 1855:Panfiloviidae 1852: 1849: 1845: 1841: 1838: 1834: 1832: 1828: 1827: 1826: 1825: 1824:Psychopsoidea 1820: 1816: 1815:Rafaelianidae 1812: 1809: 1805: 1802: 1798: 1795: 1791: 1788: 1787:Babinskaiidae 1784: 1781: 1777: 1776: 1774: 1773:Nemopteroidea 1770: 1769: 1764: 1763: 1762: 1761: 1756: 1751: 1747: 1746: 1745: 1744: 1739: 1738: 1737: 1736:Geoneuroptera 1733: 1729: 1725: 1723: 1719: 1716: 1712: 1709: 1705: 1701: 1699: 1695: 1694: 1692: 1688: 1687: 1686:Hemerobioidea 1682: 1681: 1680: 1679:Neoneuroptera 1676: 1671: 1667: 1664: 1660: 1656: 1653: 1649: 1645: 1642: 1638: 1637: 1636: 1635: 1630: 1625: 1621: 1620: 1619: 1618: 1613: 1608: 1604: 1601: 1597: 1594: 1590: 1587: 1583: 1580: 1576: 1575: 1574: 1573: 1568: 1567: 1566: 1562: 1557: 1553: 1552: 1551: 1550: 1545: 1544: 1543: 1542: 1538: 1536: 1526: 1518: 1517: 1509: 1508: 1500: 1499: 1491: 1490: 1482: 1481: 1473: 1472: 1464: 1463: 1455: 1454: 1446: 1445: 1437: 1436: 1428: 1427: 1419: 1418: 1410: 1409: 1401: 1400: 1392: 1391: 1383: 1382: 1375: 1373: 1368: 1366: 1364: 1360: 1359: 1354: 1353: 1350: 1349: 1342: 1340: 1335: 1333: 1331: 1327: 1326: 1321: 1320: 1314: 1313: 1310: 1309: 1302: 1300: 1295: 1293: 1292:(spoonwings) 1291: 1287: 1286: 1281: 1280: 1274: 1273: 1270: 1269: 1262: 1260: 1255: 1253: 1251: 1247: 1246: 1241: 1240: 1234: 1231: 1230: 1227: 1226: 1219: 1217: 1212: 1210: 1208: 1204: 1203: 1198: 1197:Psychopsoidea 1195: 1194: 1188: 1185: 1184: 1181: 1180: 1177: 1175: 1170: 1164: 1161: 1160: 1154: 1153:Geoneuroptera 1151: 1150: 1147: 1146: 1143: 1141: 1136: 1130: 1129: 1123: 1122:Neoneuroptera 1120: 1119: 1113: 1112: 1109: 1108: 1100: 1099: 1091: 1090: 1082: 1081: 1073: 1072: 1069: 1067: 1061: 1060: 1057: 1056: 1053: 1052: 1046: 1045: 1039: 1038: 1035: 1034: 1031: 1029: 1024: 1018: 1017: 1011: 1010: 1007: 1006: 1003: 1001: 995: 994: 988: 985: 984: 981: 980: 977: 975: 970: 964: 961: 960: 954: 953: 947: 946: 943: 942: 934: 933: 930: 928: 922: 919: 918: 912: 911: 905: 904: 901: 900: 892: 891: 883: 882: 874: 873: 870: 868: 862: 861: 858: 857: 854: 853: 847: 846: 840: 839: 836: 835: 832: 830: 825: 819: 818: 812: 809: 808: 802: 801: 795: 792: 791: 788: 787: 784: 783:(dustywings) 782: 776: 773: 772: 766: 765: 759: 756: 755: 749: 748: 745: 744: 741: 739: 734: 728: 727: 721: 720: 717: 716: 713: 711: 707:(snakeflies) 706: 705:Raphidioptera 700: 699: 693: 690: 689: 685: 681: 679: 675: 671: 661: 658: 656: 651: 649: 645: 641: 637: 633: 632:Burmese amber 629: 625: 624: 619: 616: 615:middle Eocene 612: 608: 607: 602: 598: 590: 586: 585: 580: 576: 571: 567: 565: 561: 560:Permithonidae 557: 554: 546: 542: 538: 533: 520: 516: 514: 511: 507: 503: 499: 497: 492: 490: 486: 482: 478: 477:holometabolic 473: 471: 467: 463: 459: 456: 452: 448: 444: 440: 435: 433: 429: 425: 421: 417: 413: 402: 398: 397: 391: 387: 383: 379: 378: 372: 368: 367: 359: 357: 351: 347: 345: 344:holometabolan 341: 337: 333: 332:compound eyes 323: 321: 317: 313: 310: 306: 302: 299: 294: 292: 291:metamorphosis 288: 283: 281: 277: 276:Raphidioptera 273: 269: 265: 261: 257: 253: 249: 245: 241: 237: 234: 231: 222: 221: 215: 210: 206: 202: 197: 194: 191: 190: 187: 184: 181: 178: 177: 174: 171: 168: 167: 164: 161: 158: 157: 154: 151: 148: 147: 144: 141: 138: 137: 132: 127: 123: 120: 116: 112: 107: 101: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 44: 39: 33: 22: 19: 4670:Neuropterida 4346: 4276:(spoonwings) 4203:Mantispidae 4187:Mantispoidea 4168:(dustywings) 4124:Chrysopidae 4116:Chrysopoidea 4063:Nevrorthidae 4031:Holometabola 4029:Superorder: 4023:Infraclass: 3987: 3942: 3928: 3898:Syntonoptera 3780:Caloneurodea 3772:Blattoptera 3723: 3714: 3709: 3668:Lepidoptera 3666: 3659:Trichoptera 3635: 3617: 3576: 3569:Megaloptera 3563:(snakeflies) 3553:Neuropterida 3537: 3522:Coleopterida 3499:Hymenoptera 3497: 3419:Paraneoptera 3417: 3339:(gladiators) 3288:Phasmatodea 3266:(stoneflies) 3251:Polyneoptera 3194:Odonatoptera 3058:Monocondylia 3023:Pancrustacea 3021:(unranked): 2947: 2895: 2889: 2850: 2844: 2803: 2799: 2793: 2784: 2780: 2774: 2749: 2745: 2709: 2705: 2695: 2684: 2679: 2670: 2662: 2657: 2648: 2595: 2591: 2581: 2548: 2544: 2534: 2522:. Retrieved 2494: 2488: 2478: 2453: 2449: 2443: 2398: 2394: 2384: 2359: 2355: 2348: 2339: 2326: 2312:cite journal 2277: 2273: 2250: 2223: 2219: 2209: 2162: 2156: 2123: 2087: 2077: 2055:(1): 59–62. 2052: 2046: 2040: 2031: 2025: 1986: 1950: 1945: 1928: 1925: 1918: 1898: 1889: 1887: 1872:Psychopsidae 1822:Superfamily 1821: 1801:Nemopteridae 1789:(Cretaceous) 1780:Ascalaphidae 1766:Superfamily 1765: 1758: 1741:Superfamily 1740: 1735: 1715:Hemerobiidae 1691:Chrysopoidea 1684:Superfamily 1683: 1678: 1665:) (Triassic) 1663:Mesithonidae 1661:(including † 1634:Mantispoidea 1632:Superfamily 1631: 1615:Superfamily 1614: 1593:Nevrorthidae 1570:Superfamily 1569: 1565:Euneuroptera 1564: 1547:Superfamily 1546: 1540: 1539: 1532: 1524: 1369: 1363:Ascalaphidae 1361: 1336: 1328: 1296: 1290:Nemopteridae 1288: 1256: 1248: 1213: 1207:Psychopsidae 1205: 1167: 1135:Hemerobiidae 1133: 1064: 1051:Symphrasinae 1049: 1021: 998: 987:Mantispoidea 967: 963:Chrysopoidea 925: 865: 852:Nevrorthidae 850: 822: 794:Euneuroptera 779: 757: 731: 703: 692:Neuropterida 682: 667: 659: 652: 621: 620:. The genus 618:Baltic amber 604: 594: 582: 550: 540: 500: 493: 479:orders, the 475:As in other 474: 436: 428:pest insects 409: 394: 375: 352: 348: 329: 305:Mesozoic era 295: 284: 280:Neuropterida 239: 235: 227: 217: 195: 186:Neuropterida 179: 18: 4513:iNaturalist 4371:Wikispecies 4219:Berothidae 4095:Ithonioidea 3944:Wikispecies 3903:Titanoptera 3835:Lapeyriidae 3830:Hypoperlida 3805:Eoblattodea 3774:(roachoids) 3767:Archodonata 3609:Antliophora 3577:Neuroptera 3539:Coleoptera 3350:Dictyoptera 3320:(Xenonomia) 3296:Orthoptera 3280:Embioptera 3272:Dermaptera 3264:Plecoptera 3164:Palaeoptera 3027:Subphylum: 2898:. Berghahn. 2877:|work= 2752:: 531–551. 2524:January 27, 2456:: 135–140. 2362:: 214–227. 2197:|work= 1743:Ithonioidea 1722:Osmylitidae 1704:Chrysopidae 1648:Mantispidae 1365:(owlflies) 1332:(antlions) 1023:Mantispidae 969:Chrysopidae 733:Megaloptera 636:crown group 634:, implying 606:Protosmylus 508:, and some 420:hemerobiids 316:butterflies 272:dobsonflies 260:Megaloptera 248:mantidflies 30:299–0  4655:Neuroptera 4649:Categories 4391:Neuroptera 4377:Neuroptera 4347:Neuroptera 4313:(owlflies) 4305:(antlions) 4295:Nymphidae 4195:Dilaridae 4174:Sisyridae 4103:Ithonidae 4057:(osmylids) 4055:Osmylidae 4017:Subclass: 4007:Arthropoda 3988:Neuroptera 3853:Miomoptera 3619:Mecoptera 3447:Hemiptera 3368:(mantises) 3358:Blattodea 3304:Zoraptera 3183:(mayflies) 3122:Zygentoma 3017:Arthropoda 2401:(1): 126. 2226:(2): 026. 1963:References 1895:Ping, 1928 1782:: owlflies 1641:Berothidae 1617:Dilaroidea 1572:Osmyloidea 1541:Neuroptera 1163:Ithonoidea 1066:Berothidae 921:Dilaroidea 811:Osmyloidea 758:Neuroptera 628:Cenomanian 611:fossilized 513:parasitize 510:lake flies 472:egg sacs. 462:mantispids 455:freshwater 439:camouflage 416:chrysopids 264:alderflies 236:Neuroptera 196:Neuroptera 163:Arthropoda 24:Neuroptera 4244:Suborder 4081:Suborder 4019:Pterygota 3999:Kingdom: 3825:Heraridea 3815:Geroptera 3720:Based on 3594:Panorpida 3541:(beetles) 3431:Psocodea 3366:Mantodea 3317:Notoptera 3274:(earwigs) 3009:Kingdom: 2879:ignored ( 2869:cite book 2820:0046-225X 2728:257245897 2622:1932-6203 2573:0016-7568 2242:1989-953X 2220:Graellsia 2199:ignored ( 2189:cite book 2142:cite book 1808:Nymphidae 1750:Ithonidae 1624:Dilaridae 1586:Sisyridae 1579:Osmylidae 1250:Nymphidae 1169:Ithonidae 927:Dilaridae 867:Sisyridae 824:Osmylidae 664:Phylogeny 623:Burmaleon 589:Ithonidae 581:) fossil 575:Paleogene 562:from the 527:Evolution 466:parasites 451:Sisyridae 447:Ithonidae 443:ant lions 412:predators 401:Sisyridae 393:Larva of 382:Osmylidae 374:Larva of 340:mandibles 268:fishflies 244:lacewings 149:Kingdom: 143:Eukaryota 102:to recent 4422:BugGuide 4398:BioLib: 4356:Wikidata 4025:Neoptera 4005:Phylum: 4001:Animalia 3990:families 3863:Paoliida 3637:Diptera 3441:(thrips) 3202:Odonata 3029:Hexapoda 3015:Phylum: 3011:Animalia 2836:46558266 2828:26314008 2766:29324039 2640:29381758 2592:PLOS ONE 2519:55970660 2435:24912379 2304:25170303 2104:55878344 2069:86240200 1985:(2005). 1864:Family † 1853:Family † 1842:Family † 1835:Family † 1829:Family † 1813:Family † 1785:Family † 1726:Family † 1720:Family † 1696:Family † 1657:Family † 1605:Family † 1598:Family † 1529:Taxonomy 655:Jurassic 597:Jurassic 537:Jurassic 460:. A few 320:Jurassic 252:antlions 220:Taxonomy 201:Linnaeus 159:Phylum: 153:Animalia 139:Domain: 4362:Q156438 4013:Insecta 4011:Class: 3986:Extant 3748:Extinct 3722:Sasaki 3629:(fleas) 3423: * 2939:on the 2787:: 1–86. 2631:5790268 2600:Bibcode 2553:Bibcode 2499:Bibcode 2458:Bibcode 2426:4113026 2403:Bibcode 2364:Bibcode 2295:4147223 1870:Family 1806:Family 1799:Family 1792:Family 1778:Family 1748:Family 1713:Family 1702:Family 1668:Family 1646:Family 1639:Family 1622:Family 1591:Family 1584:Family 1577:Family 1554:family 553:Permian 502:Beetles 458:sponges 414:. Many 356:abdomen 318:of the 298:Permian 256:species 212:clades 192:Order: 173:Insecta 169:Class: 100:Permian 4632:989447 4606:115551 4577:NZOR: 4544:114999 4466:1NEURO 3726:(2013) 3724:et al. 3715:Italic 3043:Extant 2999:orders 2996:Insect 2857:  2834:  2826:  2818:  2764:  2726:  2638:  2628:  2620:  2571:  2517:  2433:  2423:  2302:  2292:  2240:  2177:  2130:  2102:  2067:  1997:  1981:& 1952:lapsus 1857:(syn † 1757:Clade 1734:Clade 1689:(inc. 1677:Clade 1563:Clade 674:fossil 670:sister 648:Aptian 556:period 496:nectar 485:cocoon 470:spider 396:Sisyra 336:ocelli 309:family 301:period 274:) and 270:, and 230:insect 4627:WoRMS 4614:Plazi 4593:81974 4531:10490 4526:IRMNG 4518:48763 4479:11815 4401:16845 4045:Basal 2832:S2CID 2724:S2CID 2515:S2CID 2336:(PDF) 2100:S2CID 2065:S2CID 2022:(PDF) 1937:Notes 1907:and † 1771:(syn 506:wasps 489:moult 481:pupal 399:sp., 287:veins 238:, or 233:order 180:Clade 4570:7516 4565:NCBI 4539:ITIS 4505:1501 4500:GBIF 4461:EPPO 4409:BOLD 3710:bold 2881:help 2855:ISBN 2824:PMID 2816:ISSN 2762:PMID 2636:PMID 2618:ISSN 2569:ISSN 2526:2010 2431:PMID 2318:link 2300:PMID 2238:ISSN 2201:help 2175:ISBN 2148:link 2128:ISBN 1995:ISBN 630:age 577:(49 464:are 422:and 228:The 218:See 205:1758 38:PreꞒ 4552:NBN 4453:937 4448:EoL 4440:3ND 4435:CoL 4414:107 4386:AFD 2958:at 2808:doi 2754:doi 2714:doi 2626:PMC 2608:doi 2561:doi 2549:159 2507:doi 2466:doi 2421:PMC 2411:doi 2372:doi 2290:PMC 2282:doi 2228:doi 2167:doi 2092:doi 2057:doi 2053:108 1848:syn 613:in 609:is 599:or 579:mya 539:", 468:of 4651:: 4629:: 4616:: 4603:: 4590:: 4567:: 4554:: 4541:: 4528:: 4515:: 4502:: 4489:: 4476:: 4463:: 4450:: 4437:: 4427:61 4424:: 4411:: 4388:: 4373:: 4358:: 3625:+ 2943:/ 2873:: 2871:}} 2867:{{ 2830:. 2822:. 2814:. 2804:44 2802:. 2785:20 2783:. 2760:. 2750:63 2748:. 2736:^ 2722:. 2712:. 2708:. 2704:. 2634:. 2624:. 2616:. 2606:. 2596:13 2594:. 2590:. 2567:. 2559:. 2547:. 2543:. 2513:. 2505:. 2493:. 2487:. 2464:. 2454:58 2452:. 2429:. 2419:. 2409:. 2399:14 2397:. 2393:. 2370:. 2360:59 2358:. 2338:. 2314:}} 2310:{{ 2298:. 2288:. 2278:10 2276:. 2272:. 2260:^ 2236:. 2224:71 2218:. 2193:: 2191:}} 2187:{{ 2173:. 2144:}} 2140:{{ 2112:^ 2098:. 2086:. 2063:. 2051:. 2032:38 2030:. 2024:. 2009:^ 1993:. 1989:. 1970:^ 1775:) 1693:) 504:, 498:. 418:, 380:, 293:. 266:, 250:, 246:, 203:, 182:: 88:Pg 32:Ma 3979:e 3972:t 3965:v 3486:a 3484:l 3482:o 3480:b 3478:a 3476:t 3474:e 3472:m 3470:o 3468:l 3466:o 3464:H 3405:a 3403:l 3401:o 3399:b 3397:a 3395:t 3393:e 3391:m 3389:u 3387:E 3237:a 3235:r 3233:e 3231:t 3229:p 3227:o 3225:e 3223:N 3151:a 3149:t 3147:o 3145:g 3143:y 3141:r 3139:e 3137:t 3135:P 3110:a 3108:i 3106:l 3104:y 3102:d 3100:n 3098:o 3096:c 3094:i 3092:D 2988:e 2981:t 2974:v 2883:) 2863:. 2838:. 2810:: 2768:. 2756:: 2730:. 2716:: 2710:6 2642:. 2610:: 2602:: 2575:. 2563:: 2555:: 2528:. 2509:: 2501:: 2495:4 2472:. 2468:: 2460:: 2437:. 2413:: 2405:: 2378:. 2374:: 2366:: 2320:) 2306:. 2284:: 2244:. 2230:: 2203:) 2183:. 2169:: 2150:) 2136:. 2106:. 2094:: 2071:. 2059:: 2003:. 1955:. 1861:) 1846:( 591:) 587:( 547:) 543:( 262:( 93:N 83:K 78:J 73:T 68:P 63:C 58:D 53:S 48:O 43:Ꞓ

Index

Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N
Permian

Green lacewing
Scientific classification
Edit this classification
Eukaryota
Animalia
Arthropoda
Insecta
Neuropterida
Neuroptera
Linnaeus
1758
Taxonomy
insect
order
lacewings

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