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Progalesaurus

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teeth get progressively lower and anteroposteriorly longer from front to back. The buccal surface of each post-canine is smooth. The teeth are also slightly angled so that the posterior of one tooth contacts the anterior of the next. The posterior accessory cusps of teeth 2 and 3 curve upwards towards the top of the tooth. The 4th tooth shows posterior accessory cusps as well as at least one cusp mesial to the main cusp. The 5th tooth only shows one posterior accessory cusp, but the lack of other accessory cusps is “probably due to wear.” The 6th tooth on to the 9th tooth have multiple posterior accessory cusps as well as at least one mesial cusp.
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however the number and placement of the accessory cusps are unique. The upper post-canines are poorly preserved, but the teeth that are well-enough preserved to see accessory cusps have at least one posterior to the main recurved cusp each. The bottom post-canines are extremely well preserved. The
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crossed the extinction boundary. Cynodonts may have been able to survive the mass extinction due to their burrowing behavior, as perhaps living mostly underground would have pre-conditioned these burrowers to the high levels of carbon dioxide and low levels of oxygen present during the extinction
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are long and thin, with a circular cross-section. The lower incisors are shorter than their upper counterparts. The incisors have an oval cross-section and longitudinal striations. Their upper canine’s edges are preserved well enough to conclude they lack serration, but the lower canines are not
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have been found in burrows of their own making. Burrowing probably helped Triassic cynodonts to avoid harsh above-ground conditions shortly after the Permian-Triassic extinction event. Early cynodonts have even been found in burrows with other taxa, indicating they may have cohabitated
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The maxilla forms a good portion of the side of the face and is dotted with small foramina, mostly above the canines. These foramina likely housed nerves, and were perhaps associated with whiskers. The nasal is also dotted with tiny foramina.
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The orbit faces anteriorly and is formed by the lacrimal, prefrontal, jugal, and post orbital. The post orbital notably has a deeply forked posterior margin. This characteristic is seen in some other basal cynodonts, but is widely variable.
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has remarkably large nares compared to other early cynodonts. The nares are formed externally by the premaxilla, the maxilla, and the nasal. The septomaxilla resides inside the nares, on top of the junction between maxilla and premaxilla.
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period was one of the hottest in the history of the Earth, however the climate in the areas cynodonts have been discovered was more temperate. There were high levels of carbon dioxide in the atmosphere. In the Karoo Basin, where
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Benoit, Julien; Abdala, Fernando; Van den Brandt, Marc J.; Manger, Paul R.; Rubidge, Bruce S. (2015-11-04). "Physiological implications of the abnormal absence of the parietal foramen in a late Permian cynodont (Therapsida)".
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have been found with litters far larger than modern mammals. Early cynodonts have also been preserved with juveniles, suggesting they provided parental care to their young after birth or hatching.
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The name “Progalesaurus” is derived from Greek, “pro” meaning before, “gale” meaning weasel or cat, and “saurus” meaning lizard or reptile (Sidor and Smith). The name refers to the relationship
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has been found as of yet, Sidor and Smith's findings have been included in many papers on the Permian-Triassic extinction event, cynodont diversity, and the paleoenvironment of South Africa.
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Hugot, Jean-Pierre; Gardner, Scott L; Borba, Victor; Araujo, Priscilla; Leles, Daniela; Stock Da-Rosa, Átila Augusto; Dutra, Juliana; Ferreira, Luiz Fernando; Araújo, Adauto (2014-11-13).
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ABDALA, F.; CISNEROS, J. C.; SMITH, R. M.H. (2006-10-01). "Faunal Aggregation in the Early Triassic Karoo Basin: Earliest Evidence of Shelter-Sharing Behavior Among Tetrapods?".
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Smith, Roger; Botha, Jennifer (September 2005). "The recovery of terrestrial vertebrate diversity in the South African Karoo Basin after the end-Permian extinction".
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was originally discovered by Roger M. H. Smith in 1998. The specimen was found in the Sneeuberg mountains near New Lootsberg Pass in the Karoo Basin of South Africa.
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Smith, Roger; Botha, Jennifer (2005-09-01). "The recovery of terrestrial vertebrate diversity in the South African Karoo Basin after the end-Permian extinction".
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was accompanied by "survivor fauna," and "recovery fauna." A few examples of (vertebrate) survivor fauna, creatures that crossed the extinction boundary, include
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Abdala, Fernando; Ribeiro, Ana Maria (February 2010). "Distribution and diversity patterns of Triassic cynodonts (Therapsida, Cynodontia) in Gondwana".
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has a parietal foramen, which is used for light sensing in extant taxa. Posterior to the parietal foramen the parietals are fused, forming a
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Hoffman, Eva A.; Rowe, Timothy B. (2018-08-29). "Jurassic stem-mammal perinates and the origin of mammalian reproduction and growth".
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measuring 9.35 cm in length. It likely closely resembled its early cynodont relatives, walking on four legs and covered in fur.
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still sharp enough to make any conclusions about their serration. The lower canines are slightly longer than the upper canines.
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Retallack, G.J. (2004). "Vertebrate extinction across Permian-Triassic boundary in Karoo Basin, South Africa: Reply".
4019: 2051: 1336:"Aggregations and parental care in the Early Triassic basal cynodonts Galesaurus planiceps and Thrinaxodon liorhinus" 4029: 1216:
Rubidge, Bruce S.; Sidor, Christian A. (November 2001). "Evolutionary Patterns Among Permo-Triassic Therapsids".
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and the left atlantal neural spine were recovered. Each of these elements closely resemble those of
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in what is modern South Africa. Other cynodonts and potentially even other galesaurids, like
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was discovered, there is evidence to suggest the area was a moderately damp open woodland.
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event. Cynodont diversity increased relatively rapidly after the extinction event.
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A New Galesaurid (Therapsida: Cynodontia) From the Lower Triassic of South Africa.”
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Based on tooth shape, paleontologists believe that early cynodonts like
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The main groups of non-mammalian synapsids at Mikko's Phylogeny Archive
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Very little post-cranial skeleton is preserved in the holotype for
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The post-canine teeth are, as of now, the most distinct feature of
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was a relatively small mammal-like creature, with the skull of its
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likely had large litters, as more derived cynodonts like
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Benoit, J.; Manger, P. R.; Rubidge, B. S. (2016-05-09).
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has a dental formula of I4/3, C1/1, PC7?/9. The upper
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with its teeth setting it apart from other cynodonts.
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Sidor, Christian A.; Smith, Roger M. H. (May 2004).
1392: 1114: 416:. The crest narrows posterior to the foramen like 1079:Palaeogeography, Palaeoclimatology, Palaeoecology 4001: 933:Recovery fauna include small amphibians such as 1334:Jasinoski, Sandra C.; Abdala, Fernando (2017). 1333: 2080: 852:likely burrowed, as closely related taxa like 354:and Roger M. H. Smith in their paper titled, “ 2791: 1546: 1076: 290:is known from a single fossil of the species 1215: 1250: 1553: 1539: 1298: 1036: 992: 117: 1703: 1486: 1463: 1445: 1369: 1351: 1148: 1010: 1627: 1230:10.1146/annurev.ecolsys.32.081501.114113 1218:Annual Review of Ecology and Systematics 1110: 1108: 534:is a galesaurid, belonging to the clade 507: 338: 907:In the early Triassic of South Africa, 321:. Galesaurids appeared just before the 4002: 1489:Geological Society of America Bulletin 883:lived very soon after the devastating 3881: 3880: 3672: 2778: 2067: 1690: 1572: 1534: 1105: 453:, the foramen is relatively shallow. 1294: 1292: 1290: 1072: 1070: 1068: 1032: 1030: 988: 986: 984: 982: 980: 978: 976: 974: 972: 941:, and some archosauromorphs such as 2068: 875: 495:. The recurved main cusp resembles 420:and unlike more derived cynodonts. 13: 2779: 868:were insectivores and carnivores. 14: 4051: 1519: 1287: 1065: 1027: 969: 885:Permian-Triassic extinction event 526: 323:Permian-Triassic extinction event 3861: 3860: 3655: 3650: 2761: 2755: 2749: 2050: 2045: 1012:10.1111/j.0031-0239.2004.00378.x 138: 42: 1480: 1421: 1386: 831: 350:was first described in 2004 by 1691: 1560: 1327: 1244: 1209: 1165: 367: 360:Although no other specimen of 129:dentary in right lateral view 1: 4040:Taxa named by Christian Sidor 4035:Fossil taxa described in 2004 962: 542:belongs to the greater clade 459:does not have a fully-formed 3673: 1091:10.1016/j.palaeo.2010.01.011 466: 292:Progalesaurus lootsbergensis 7: 950: 475:is very similar to that of 309:other galesaurids, include 258:Progalesaurus lootbergensis 10: 4056: 1407:10.2110/palo.2005.p06-001r 1313:10.1016/j.crpv.2005.07.005 1051:10.1016/j.crpv.2005.07.005 418:Galesaurus and Cynosaurus, 16:Extinct genus of cynodonts 3889: 3856: 3809: 3683: 3679: 3668: 3648: 3618: 3575: 3450: 3441: 3373: 3361: 3188: 3135: 3091: 2802: 2787: 2774: 2747: 2713: 2571: 2513: 2504: 2438: 2408: 2290: 2279: 2238: 2208: 2163: 2095: 2091: 2076: 2063: 2043: 2009: 1983: 1965: 1956: 1926: 1848: 1837: 1803: 1766: 1718: 1714: 1699: 1686: 1654: 1636: 1623: 1568: 1447:10.1186/s13071-014-0486-6 1265:10.1038/s41586-018-0441-3 1187:10.1007/s00114-015-1321-4 770: 755: 748: 728: 721: 704: 697: 673: 656: 649: 632: 625: 596: 579: 572: 565: 255: 248: 135:Scientific classification 133: 125: 116: 23: 4020:Early Triassic synapsids 382: 332:has to the more derived 4030:Fossils of South Africa 1434:Parasites & Vectors 550:, and the even broader 497:Galesaurus, Cynosaurus, 428:Galesaurus, Cynosaurus, 275:is an extinct genus of 263:Sidor & Smith, 2004 243:Sidor & Smith, 2004 1301:Comptes Rendus Palevol 1039:Comptes Rendus Palevol 4025:Triassic South Africa 3982:Paleobiology Database 1175:The Science of Nature 836:Early cynodonts like 538:. As an epicynodont, 508:Post-cranial skeleton 339:Discovery and history 305:. Close relatives of 861:interspecifically. 1353:10.7717/peerj.2875 1121:Scientific Reports 957:List of therapsids 937:Galesaurids, some 437:possesses a large 352:Christian A. Sidor 37:252.17–251.2  3997: 3996: 3969:Open Tree of Life 3883:Taxon identifiers 3874: 3873: 3852: 3851: 3848: 3847: 3664: 3663: 3646: 3645: 3642: 3641: 3614: 3613: 3610: 3609: 2770: 2769: 2745: 2744: 2741: 2740: 2737: 2736: 2733: 2732: 2413:Prozostrodontidae 2275: 2274: 2213:Probainognathidae 2059: 2058: 2041: 2040: 2037: 2036: 2033: 2032: 2029: 2028: 1833: 1832: 1771:Charassognathidae 1682: 1681: 1678: 1677: 1674: 1673: 1259:(7721): 104–108. 1133:10.1038/srep25604 828: 827: 819: 818: 810: 809: 801: 800: 792: 791: 783: 782: 686: 685: 611: 610: 303:Balfour Formation 268: 267: 244: 4047: 3990: 3989: 3977: 3976: 3964: 3963: 3951: 3950: 3938: 3937: 3925: 3924: 3923: 3910: 3909: 3908: 3878: 3877: 3864: 3863: 3796: 3786: 3776: 3766: 3756: 3746: 3736: 3726: 3716: 3706: 3696: 3681: 3680: 3670: 3669: 3659: 3654: 3448: 3447: 3371: 3370: 3364: 3191: 3138: 3096:Kuehneotheriidae 3094: 3079: 3069: 3059: 3049: 3038: 3028: 3018: 3008: 2998: 2988: 2978: 2968: 2958: 2948: 2938: 2928: 2918: 2908: 2898: 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623–636. 1034: 1025: 1024: 1014: 990: 876:Paleoenvironment 751: 750: 724: 723: 700: 699: 652: 651: 628: 627: 575: 574: 568: 567: 558: 557: 471:The mandible of 461:secondary palate 242: 235: 222: 143: 142: 121: 111: 41: 29:Temporal range: 21: 20: 4055: 4054: 4050: 4049: 4048: 4046: 4045: 4044: 4000: 3999: 3998: 3993: 3985: 3980: 3972: 3967: 3959: 3954: 3946: 3941: 3933: 3928: 3919: 3918: 3913: 3904: 3903: 3898: 3885: 3875: 3870: 3844: 3814: 3805: 3779:Panchetocynodon 3675: 3660: 3638: 3606: 3571: 3437: 3387:Hypsiprymnopsis 3357: 3184: 3131: 3087: 2821:Brachyzostrodon 2796: 2783: 2766: 2729: 2709: 2690:Xenocretosuchus 2576:Tritylodontidae 2567: 2500: 2459:Diarthrognathus 2434: 2404: 2284: 2282:Prozostrodontia 2271: 2234: 2229:Probainognathus 2204: 2159: 2085: 2083:Probainognathia 2072: 2070:Probainognathia 2055: 2025: 2011:Probainognathia 2005: 1979: 1952: 1922: 1897:Platycraniellus 1842: 1829: 1808:Procynosuchidae 1799: 1787:Charassognathus 1762: 1708: 1695: 1670: 1650: 1619: 1575: 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3665: 3662: 3661: 3649: 3647: 3644: 3643: 3640: 3639: 3637: 3636: 3635: 3634: 3625: 3623: 3616: 3615: 3612: 3611: 3608: 3607: 3605: 3604: 3597: 3590: 3582: 3580: 3573: 3572: 3570: 3569: 3565:Gondwanatheria 3561: 3554: 3547: 3540: 3533: 3526: 3519: 3512: 3504: 3497: 3489: 3482: 3475: 3467: 3464:Arboroharamiya 3460: 3451: 3445: 3439: 3438: 3436: 3435: 3428: 3421: 3413: 3405: 3402:Kalaallitkigun 3398: 3391: 3383: 3374: 3368: 3359: 3358: 3356: 3355: 3348: 3341: 3334: 3327: 3320: 3312: 3305: 3298: 3291: 3283: 3276: 3269: 3261: 3254: 3247: 3240: 3233: 3226: 3219: 3212: 3205: 3197: 3195: 3186: 3185: 3183: 3182: 3175: 3168: 3160: 3152: 3144: 3142: 3133: 3132: 3130: 3129: 3126:Kuehneotherium 3122: 3115: 3108: 3100: 3098: 3089: 3088: 3086: 3085: 3075: 3065: 3055: 3045: 3034: 3024: 3014: 3011:Paikasigudodon 3004: 2994: 2984: 2974: 2964: 2954: 2944: 2934: 2924: 2914: 2911:Erythrotherium 2904: 2894: 2884: 2874: 2864: 2854: 2844: 2834: 2824: 2814: 2803: 2797: 2794:Mammaliaformes 2792: 2785: 2784: 2781:Mammaliaformes 2772: 2771: 2768: 2767: 2748: 2746: 2743: 2742: 2739: 2738: 2735: 2734: 2731: 2730: 2728: 2727: 2726: 2725: 2719: 2717: 2715:Mammaliaformes 2711: 2710: 2708: 2707: 2700: 2693: 2686: 2679: 2672: 2665: 2658: 2651: 2644: 2637: 2630: 2627:Kayentatherium 2623: 2616: 2609: 2602: 2595: 2592:Bienotheroides 2588: 2580: 2578: 2569: 2568: 2566: 2565: 2555: 2545: 2535: 2525: 2514: 2508: 2506:Mammaliamorpha 2502: 2501: 2499: 2498: 2491: 2483: 2476: 2469: 2462: 2455: 2447: 2445: 2436: 2435: 2433: 2432: 2425: 2417: 2415: 2406: 2405: 2403: 2402: 2392: 2389:Therioherpeton 2382: 2372: 2362: 2352: 2342: 2332: 2322: 2312: 2302: 2291: 2285: 2280: 2277: 2276: 2273: 2272: 2270: 2269: 2266:Trucidocynodon 2262: 2255: 2247: 2245: 2236: 2235: 2233: 2232: 2225: 2217: 2215: 2206: 2205: 2203: 2202: 2195: 2187: 2180: 2172: 2170: 2161: 2160: 2158: 2157: 2147: 2137: 2127: 2117: 2107: 2096: 2086: 2081: 2074: 2073: 2061: 2060: 2057: 2056: 2044: 2042: 2039: 2038: 2035: 2034: 2031: 2030: 2027: 2026: 2024: 2023: 2022: 2021: 2015: 2013: 2007: 2006: 2004: 2003: 2002: 2001: 1992: 1990: 1981: 1980: 1978: 1977: 1966: 1960: 1954: 1953: 1951: 1950: 1943: 1935: 1933: 1924: 1923: 1921: 1920: 1910: 1900: 1890: 1880: 1870: 1860: 1849: 1843: 1838: 1835: 1834: 1831: 1830: 1828: 1827: 1820: 1812: 1810: 1801: 1800: 1798: 1797: 1790: 1783: 1775: 1773: 1764: 1763: 1761: 1760: 1750: 1740: 1730: 1719: 1709: 1704: 1697: 1696: 1684: 1683: 1680: 1679: 1676: 1675: 1672: 1671: 1669: 1668: 1667: 1666: 1660: 1658: 1652: 1651: 1649: 1648: 1647: 1646: 1637: 1631: 1621: 1620: 1616: 1615: 1613:Eutheriodontia 1606: 1597: 1588: 1582: 1566: 1565: 1558: 1557: 1550: 1543: 1535: 1529: 1528: 1521: 1520:External links 1518: 1515: 1514: 1479: 1420: 1401:(5): 507–512. 1385: 1326: 1307:(6): 623–636. 1286: 1243: 1224:(1): 449–480. 1208: 1164: 1104: 1064: 1026: 1005:(3): 535–556. 967: 966: 964: 961: 960: 959: 952: 949: 877: 874: 833: 830: 826: 825: 822: 821: 817: 816: 813: 812: 808: 807: 804: 803: 799: 798: 795: 794: 790: 789: 786: 785: 781: 780: 777: 776: 769: 766: 765: 762: 761: 754: 749: 747: 741: 740: 737: 736: 727: 722: 720: 717: 716: 713: 712: 703: 698: 696: 693: 692: 689: 688: 684: 683: 680: 679: 672: 669: 668: 665: 664: 655: 650: 648: 645: 644: 641: 640: 631: 626: 624: 618: 617: 614: 613: 609: 608: 605: 604: 595: 592: 591: 588: 587: 578: 573: 571: 566: 564: 556: 528: 527:Classification 525: 514:Progalesaurus. 509: 506: 468: 465: 439:zygomatic arch 414:sagittal crest 388:Progalesaurus, 384: 381: 369: 366: 340: 337: 307:Progalesaurus, 266: 265: 262: 253: 252: 246: 245: 231: 227: 226: 218: 214: 213: 208: 201: 200: 195: 188: 187: 182: 175: 174: 169: 165: 164: 159: 155: 154: 149: 145: 144: 131: 130: 123: 122: 114: 113: 105: 104: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 48: 43: 28: 15: 9: 6: 4: 3: 2: 4052: 4041: 4038: 4036: 4033: 4031: 4028: 4026: 4023: 4021: 4018: 4016: 4013: 4011: 4008: 4007: 4005: 3988: 3983: 3979: 3975: 3970: 3966: 3962: 3957: 3953: 3949: 3944: 3940: 3936: 3931: 3927: 3922: 3921:Progalesaurus 3916: 3912: 3907: 3901: 3897: 3896: 3894: 3892: 3891:Progalesaurus 3888: 3884: 3879: 3867: 3859: 3858: 3855: 3841: 3838: 3836: 3835:Symmetrodonta 3833: 3831: 3828: 3826: 3823: 3822: 3820: 3817: 3812: 3808: 3801: 3800: 3793: 3791: 3790: 3783: 3781: 3780: 3773: 3771: 3770: 3763: 3761: 3760: 3753: 3751: 3750: 3743: 3741: 3740: 3733: 3731: 3730: 3723: 3721: 3720: 3713: 3711: 3710: 3703: 3701: 3700: 3693: 3692: 3690: 3688: 3687: 3682: 3678: 3671: 3667: 3658: 3653: 3633: 3629: 3628: 3627: 3626: 3624: 3622: 3617: 3603: 3602: 3598: 3596: 3595: 3591: 3589: 3588: 3584: 3583: 3581: 3578: 3577:Hahnodontidae 3574: 3567: 3566: 3562: 3560: 3559: 3555: 3553: 3552: 3548: 3546: 3545: 3541: 3539: 3538: 3537:Sineleutherus 3534: 3532: 3531: 3527: 3525: 3524: 3520: 3518: 3517: 3513: 3510: 3509: 3505: 3503: 3502: 3498: 3495: 3494: 3490: 3488: 3487: 3483: 3481: 3480: 3476: 3473: 3472: 3468: 3466: 3465: 3461: 3458: 3457: 3453: 3452: 3449: 3446: 3444: 3440: 3434: 3433: 3429: 3427: 3426: 3422: 3419: 3418: 3414: 3411: 3410: 3406: 3404: 3403: 3399: 3397: 3396: 3392: 3389: 3388: 3384: 3381: 3380: 3376: 3375: 3372: 3369: 3367: 3360: 3354: 3353: 3349: 3347: 3346: 3345:Tashkumyrodon 3342: 3340: 3339: 3335: 3333: 3332: 3331:Sibirotherium 3328: 3326: 3325: 3324:Peraiocynodon 3321: 3318: 3317: 3313: 3311: 3310: 3306: 3304: 3303: 3299: 3297: 3296: 3292: 3289: 3288: 3284: 3282: 3281: 3280:Hutegotherium 3277: 3275: 3274: 3270: 3267: 3266: 3262: 3260: 3259: 3255: 3253: 3252: 3248: 3246: 3245: 3241: 3239: 3238: 3234: 3232: 3231: 3227: 3225: 3224: 3223:Cyrtlatherium 3220: 3218: 3217: 3213: 3211: 3210: 3206: 3204: 3203: 3199: 3198: 3196: 3194: 3187: 3181: 3180: 3176: 3174: 3173: 3169: 3166: 3165: 3161: 3158: 3157: 3153: 3151: 3150: 3146: 3145: 3143: 3141: 3140:Shuotheriidae 3134: 3128: 3127: 3123: 3121: 3120: 3116: 3114: 3113: 3109: 3107: 3106: 3102: 3101: 3099: 3097: 3090: 3084: 3083: 3076: 3074: 3073: 3066: 3064: 3063: 3056: 3054: 3053: 3046: 3043: 3042: 3035: 3033: 3032: 3025: 3023: 3022: 3015: 3013: 3012: 3005: 3003: 3002: 2995: 2993: 2992: 2985: 2983: 2982: 2981:Megazostrodon 2975: 2973: 2972: 2971:Indozostrodon 2965: 2963: 2962: 2955: 2953: 2952: 2945: 2943: 2942: 2941:Hallautherium 2935: 2933: 2932: 2925: 2923: 2922: 2915: 2913: 2912: 2905: 2903: 2902: 2895: 2893: 2892: 2885: 2883: 2882: 2875: 2873: 2872: 2865: 2863: 2862: 2855: 2853: 2852: 2851:Cifellilestes 2845: 2843: 2842: 2835: 2833: 2832: 2825: 2823: 2822: 2815: 2813: 2812: 2805: 2804: 2801: 2798: 2795: 2790: 2786: 2782: 2777: 2773: 2764: 2760: 2758: 2752: 2723: 2722: 2721: 2720: 2718: 2716: 2712: 2706: 2705: 2701: 2699: 2698: 2694: 2692: 2691: 2687: 2685: 2684: 2680: 2678: 2677: 2676:Stereognathus 2673: 2671: 2670: 2666: 2664: 2663: 2659: 2657: 2656: 2652: 2650: 2649: 2645: 2643: 2642: 2638: 2636: 2635: 2631: 2629: 2628: 2624: 2622: 2621: 2617: 2615: 2614: 2610: 2608: 2607: 2603: 2601: 2600: 2596: 2594: 2593: 2589: 2587: 2586: 2582: 2581: 2579: 2577: 2570: 2564: 2563: 2556: 2554: 2553: 2546: 2544: 2543: 2536: 2534: 2533: 2526: 2524: 2523: 2522:Adelobasileus 2516: 2515: 2512: 2509: 2507: 2503: 2497: 2496: 2492: 2489: 2488: 2484: 2482: 2481: 2477: 2475: 2474: 2470: 2468: 2467: 2463: 2461: 2460: 2456: 2454: 2453: 2449: 2448: 2446: 2444: 2437: 2431: 2430: 2429:Pseudotherium 2426: 2424: 2423: 2419: 2418: 2416: 2414: 2407: 2401: 2400: 2393: 2391: 2390: 2383: 2381: 2380: 2373: 2371: 2370: 2363: 2361: 2360: 2353: 2351: 2350: 2343: 2341: 2340: 2333: 2331: 2330: 2323: 2321: 2320: 2313: 2311: 2310: 2303: 2301: 2300: 2299:Alemoatherium 2293: 2292: 2289: 2286: 2283: 2278: 2268: 2267: 2263: 2261: 2260: 2256: 2254: 2253: 2249: 2248: 2246: 2244: 2237: 2231: 2230: 2226: 2224: 2223: 2219: 2218: 2216: 2214: 2207: 2201: 2200: 2196: 2193: 2192: 2188: 2186: 2185: 2181: 2179: 2178: 2174: 2173: 2171: 2169: 2162: 2156: 2155: 2148: 2146: 2145: 2138: 2136: 2135: 2128: 2126: 2125: 2118: 2116: 2115: 2114:Candelariodon 2108: 2106: 2105: 2098: 2097: 2094: 2090: 2087: 2084: 2079: 2075: 2071: 2066: 2062: 2053: 2048: 2019: 2018: 2017: 2016: 2014: 2012: 2008: 2000: 1996: 1995: 1994: 1993: 1991: 1989: 1982: 1976: 1975: 1968: 1967: 1964: 1961: 1959: 1955: 1949: 1948: 1947:Progalesaurus 1944: 1942: 1941: 1937: 1936: 1934: 1932: 1925: 1919: 1918: 1911: 1909: 1908: 1901: 1899: 1898: 1891: 1889: 1888: 1881: 1879: 1878: 1877:Nanictosaurus 1871: 1869: 1868: 1861: 1859: 1858: 1851: 1850: 1847: 1844: 1841: 1840:Epicynodontia 1836: 1826: 1825: 1821: 1819: 1818: 1817:Procynosuchus 1814: 1813: 1811: 1809: 1802: 1796: 1795: 1791: 1789: 1788: 1784: 1782: 1781: 1777: 1776: 1774: 1772: 1765: 1759: 1758: 1751: 1749: 1748: 1741: 1739: 1738: 1731: 1729: 1728: 1721: 1720: 1717: 1713: 1710: 1707: 1702: 1698: 1694: 1689: 1685: 1664: 1663: 1662: 1661: 1659: 1657: 1653: 1645: 1641: 1640: 1639: 1638: 1635: 1632: 1630: 1626: 1622: 1614: 1610: 1607: 1605: 1601: 1598: 1596: 1592: 1589: 1587: 1583: 1581: 1577: 1576: 1571: 1567: 1563: 1556: 1551: 1549: 1544: 1542: 1537: 1536: 1533: 1527: 1524: 1523: 1510: 1506: 1502: 1498: 1494: 1490: 1483: 1475: 1471: 1466: 1461: 1457: 1453: 1448: 1443: 1439: 1435: 1431: 1424: 1416: 1412: 1408: 1404: 1400: 1396: 1389: 1381: 1377: 1372: 1367: 1363: 1359: 1354: 1349: 1345: 1341: 1337: 1330: 1322: 1318: 1314: 1310: 1306: 1302: 1295: 1293: 1291: 1282: 1278: 1274: 1270: 1266: 1262: 1258: 1254: 1247: 1239: 1235: 1231: 1227: 1223: 1219: 1212: 1204: 1200: 1196: 1192: 1188: 1184: 1181:(11–12): 69. 1180: 1176: 1168: 1160: 1156: 1151: 1146: 1142: 1138: 1134: 1130: 1126: 1122: 1118: 1111: 1109: 1100: 1096: 1092: 1088: 1084: 1080: 1073: 1071: 1069: 1060: 1056: 1052: 1048: 1044: 1040: 1033: 1031: 1022: 1018: 1013: 1008: 1004: 1000: 999:Palaeontology 996: 989: 987: 985: 983: 981: 979: 977: 975: 973: 968: 958: 955: 954: 948: 946: 945: 944:Proterosuchus 940: 936: 932: 930: 925: 923: 919: 915: 910: 909:Progalesaurus 905: 903: 902:Progalesaurus 898: 893: 890: 886: 882: 881:Progalesaurus 873: 871: 867: 866:Progalesaurus 862: 859: 855: 851: 850:Progalesaurus 847: 845: 844: 843:Tritylodontid 839: 838:Progalesaurus 824: 823: 815: 814: 806: 805: 797: 796: 788: 787: 779: 778: 775: 774: 768: 767: 764: 763: 760: 759: 753: 752: 746: 743: 742: 739: 738: 735: 734: 733: 726: 725: 719: 718: 715: 714: 711: 710: 709: 702: 701: 695: 694: 691: 690: 682: 681: 678: 677: 676:Progalesaurus 671: 670: 667: 666: 663: 662: 661: 654: 653: 647: 646: 643: 642: 639: 638: 637: 630: 629: 623: 622:Epicynodontia 620: 619: 616: 615: 607: 606: 603: 602: 601: 600:Procynosuchus 594: 593: 590: 589: 586: 585: 584: 577: 576: 570: 569: 563: 560: 559: 555: 553: 549: 545: 541: 540:Progalesaurus 537: 536:Epicynodontia 533: 532:Progalesaurus 524: 523: 519: 515: 505: 502: 501:Probelesodon, 498: 494: 493:Progalesaurus 489: 486: 482: 481:Progalesaurus 478: 474: 473:Progalesaurus 464: 462: 458: 457:Progalesaurus 454: 452: 451: 446: 445: 440: 436: 434: 429: 425: 424:Progalesaurus 421: 419: 415: 411: 410:Progalesaurus 407: 403: 399: 396: 394: 389: 380: 378: 374: 373:Progalesaurus 365: 363: 362:Progalesaurus 358: 357: 353: 349: 348:Progalesaurus 345: 344:Progalesaurus 336: 335: 331: 330:Progalesaurus 326: 324: 320: 319: 314: 313: 308: 304: 300: 298: 293: 289: 288:Progalesaurus 285: 281: 278: 274: 273: 272:Progalesaurus 260: 259: 254: 251: 247: 240: 239: 238:Progalesaurus 232: 229: 228: 225: 219: 216: 215: 212: 209: 206: 203: 202: 199: 196: 193: 190: 189: 186: 183: 180: 177: 176: 173: 170: 167: 166: 163: 160: 157: 156: 153: 150: 147: 146: 141: 136: 132: 128: 127:Progalesaurus 124: 120: 115: 101: 96: 91: 86: 81: 76: 71: 66: 61: 56: 51: 46: 40: 32: 26: 25:Progalesaurus 22: 19: 3890: 3840:Triconodonta 3816:Polyphyletic 3811:Paraphyletic 3797: 3787: 3777: 3767: 3757: 3747: 3737: 3727: 3717: 3707: 3699:Cistecynodon 3697: 3684: 3599: 3592: 3585: 3563: 3556: 3549: 3542: 3535: 3528: 3521: 3514: 3506: 3501:Maiopatagium 3499: 3491: 3486:Cryoharamiya 3484: 3477: 3469: 3462: 3454: 3443:Euharamiyida 3430: 3423: 3415: 3407: 3400: 3393: 3385: 3377: 3350: 3343: 3336: 3329: 3322: 3314: 3309:Microdocodon 3307: 3300: 3295:Khorotherium 3293: 3285: 3278: 3271: 3263: 3256: 3249: 3242: 3235: 3228: 3221: 3216:Castorocauda 3214: 3207: 3202:Agilodocodon 3200: 3177: 3172:Pseudotribos 3170: 3162: 3154: 3147: 3124: 3117: 3110: 3103: 3080: 3070: 3060: 3050: 3039: 3029: 3019: 3009: 2999: 2991:Morganucodon 2989: 2979: 2969: 2959: 2949: 2939: 2929: 2919: 2909: 2899: 2889: 2881:Dinnetherium 2879: 2871:Dianoconodon 2869: 2859: 2849: 2839: 2831:Bridetherium 2829: 2819: 2809: 2754: 2702: 2697:Yuanotherium 2695: 2688: 2681: 2674: 2667: 2660: 2653: 2646: 2639: 2632: 2625: 2618: 2613:Dinnebitodon 2611: 2604: 2597: 2590: 2585:Bienotherium 2583: 2560: 2550: 2540: 2530: 2520: 2493: 2485: 2480:Pachygenelus 2478: 2473:Irajatherium 2471: 2466:Elliotherium 2464: 2457: 2450: 2427: 2422:Prozostrodon 2420: 2397: 2387: 2377: 2367: 2357: 2347: 2339:Microconodon 2337: 2327: 2319:Dromatherium 2317: 2307: 2297: 2264: 2257: 2250: 2243:Ecteniniidae 2227: 2220: 2197: 2189: 2182: 2175: 2152: 2144:Protheriodon 2142: 2132: 2122: 2112: 2104:Agudotherium 2102: 1974:Kataigidodon 1972: 1958:Eucynodontia 1946: 1945: 1938: 1931:Galesauridae 1915: 1905: 1895: 1885: 1875: 1865: 1855: 1822: 1815: 1792: 1785: 1778: 1757:Uralocynodon 1755: 1745: 1735: 1725: 1608: 1599: 1590: 1492: 1488: 1482: 1437: 1433: 1423: 1398: 1394: 1388: 1343: 1339: 1329: 1304: 1300: 1256: 1252: 1246: 1221: 1217: 1211: 1178: 1174: 1167: 1127:(1): 25604. 1124: 1120: 1082: 1078: 1042: 1038: 1002: 998: 942: 935:Micropholis, 934: 927: 922:Moschorhinus 918:Tetracynodon 914:Lystrosaurus 912: 908: 906: 901: 894: 888: 880: 879: 865: 863: 857: 853: 849: 848: 841: 837: 835: 832:Paleobiology 771: 756: 745:Eucynodontia 730: 729: 706: 705: 675: 674: 658: 657: 634: 633: 598: 597: 581: 580: 539: 531: 530: 522:Thrinaxodon. 521: 513: 511: 500: 496: 492: 490: 480: 476: 472: 470: 456: 455: 448: 442: 431: 427: 423: 422: 417: 409: 408: 404: 400: 391: 387: 386: 372: 371: 361: 359: 355: 347: 343: 342: 333: 329: 327: 316: 310: 306: 297:Lystrosaurus 296: 291: 287: 271: 270: 269: 257: 256: 250:Type species 237: 236: 224:Galesauridae 204: 191: 178: 126: 24: 18: 4015:Induan life 3915:Wikispecies 3799:Saurodesmus 3587:Cifelliodon 3551:Woodeatonia 3544:Vilevolodon 3523:Sharypovoia 3493:Kermackodon 3456:Allostaffia 3425:Theroteinus 3395:Haramiyavia 3352:Tegotherium 3338:Simpsonodon 3316:Paritatodon 3265:Gondtherium 3251:Dsungarodon 3209:Borealestes 3179:Shuotherium 3164:Paritatodon 3149:Feredocodon 3112:Kotatherium 3072:Wareolestes 3041:Sinoconodon 3021:Purbeckodon 2961:Indotherium 2951:Helvetiodon 2931:Hadrocodium 2921:Gondwanadon 2901:Eozostrodon 2891:Dyskritodon 2811:Bocaconodon 2669:Shartegodon 2655:Oligokyphus 2648:Nuurtherium 2641:Montirictus 2620:Fossiomanus 2606:Dianzhongia 2599:Bocatherium 2552:Sinoconodon 2495:Tritheledon 2369:Rewaconodon 2329:Inditherium 2222:Bonacynodon 2184:Chiniquodon 2154:Tessellatia 1999:Cynognathia 1988:Cynognathia 1907:Thrinaxodon 1887:Nythosaurus 1737:Nanocynodon 1495:(9): 1295. 889:Cynosaurus, 854:Thrinaxodon 758:Cynognathia 708:Thrinaxodon 477:Galesaurus, 444:Cynognathus 433:Thrinaxodon 368:Description 334:Galesaurus. 4004:Categories 3759:Meurthodon 3479:Butlerodon 3471:Baidabatyr 3409:Kollikodon 3379:Avashishta 3366:Haramiyida 3302:Krusatodon 3273:Haldanodon 3244:Docofossor 3230:Dobunnodon 3105:Fluctuodon 3052:Storchodon 3031:Rosierodon 2841:Cherwellia 2724:see below↓ 2704:Yunnanodon 2683:Tritylodon 2662:Polistodon 2542:Brasilodon 2487:Riograndia 2452:Chaliminia 2309:Deccanodon 2259:Ecteninion 2252:Diegocanis 2199:Riojanodon 2191:Cromptodon 2124:Charruodon 2020:see below↓ 1940:Galesaurus 1867:Cynosaurus 1857:Bolotridon 1794:Nshimbodon 1747:Madysaurus 1706:Cynodontia 1693:Cynodontia 1665:see below↓ 1656:Cynodontia 1604:Therapsida 1562:Cynodontia 1440:(1): 486. 963:References 895:The early 870:Coprolites 858:Galesaurus 660:Galesaurus 636:Cynosaurus 562:Cynodontia 544:Cynodontia 450:Diademodon 393:Galesaurus 318:Cynosaurus 312:Galesaurus 277:galesaurid 211:Cynodontia 198:Therapsida 4010:Cynodonts 3594:Denisodon 3508:Megaconus 3193:Docodonta 3082:Woutersia 3062:Stylidens 3001:Paceyodon 2399:Tricuspes 2349:Polonodon 1917:Vetusodon 1780:Abdalodon 1644:Synapsida 1629:Synapsida 1595:Synapsida 1578:Kingdom: 1509:0016-7606 1456:1756-3305 1415:0883-1351 1362:2167-8359 1346:: e2875. 1321:1631-0683 1273:0028-0836 1238:0066-4162 1195:0028-1042 1141:2045-2322 1099:0031-0182 1059:1631-0683 1021:0031-0239 548:Synapsida 467:Dentition 185:Synapsida 158:Kingdom: 152:Eukaryota 3935:24177067 3906:Q2122571 3900:Wikidata 3866:Category 3769:Mitredon 3739:Kunminia 3729:"Hahnia" 3709:Eoraetia 3674:See also 3632:Mammalia 3621:Mammalia 3601:Hahnodon 3558:Xianshou 3530:Shenshou 3432:Thomasia 3287:Itatodon 3258:Ergetiis 3156:Itatodon 3119:Kuehneon 2861:Delsatia 2634:Lufengia 2562:Tikiodon 1586:Chordata 1584:Phylum: 1580:Animalia 1474:25495824 1380:28097072 1281:30158701 1203:26538062 1159:27157809 951:See also 897:Triassic 485:incisors 377:holotype 284:Triassic 280:cynodont 217:Family: 172:Chordata 168:Phylum: 162:Animalia 148:Domain: 3974:4127889 3961:1171785 3948:4817626 3749:Lepagia 3237:Docodon 2177:Aleodon 2134:Lumkuia 1824:Sludica 1465:4236488 1395:PALAIOS 1371:5228509 1150:4860582 552:Amniota 518:scapula 426:, like 301:of the 230:Genus: 106:↓ 3987:171915 3818:groups 3719:Gaumia 3619:Crown- 3516:Qishou 1727:Dvinia 1507:  1472:  1462:  1454:  1413:  1378:  1368:  1360:  1319:  1279:  1271:  1253:Nature 1236:  1201:  1193:  1157:  1147:  1139:  1097:  1057:  1019:  583:Dvinia 31:Induan 3956:IRMNG 1609:Clade 1600:Clade 1591:Clade 1340:PeerJ 390:like 383:Skull 205:Clade 192:Clade 179:Clade 3943:GBIF 3630:see 3417:Mojo 1997:see 1642:see 1505:ISSN 1470:PMID 1452:ISSN 1411:ISSN 1376:PMID 1358:ISSN 1317:ISSN 1277:PMID 1269:ISSN 1234:ISSN 1199:PMID 1191:ISSN 1155:PMID 1137:ISSN 1095:ISSN 1055:ISSN 1017:ISSN 926:and 856:and 499:and 430:and 315:and 45:PreꞒ 3930:EoL 1497:doi 1493:116 1460:PMC 1442:doi 1403:doi 1366:PMC 1348:doi 1309:doi 1261:doi 1257:561 1226:doi 1183:doi 1179:102 1145:PMC 1129:doi 1087:doi 1083:286 1047:doi 1007:doi 447:or 4006:: 3984:: 3971:: 3958:: 3945:: 3932:: 3917:: 3902:: 1611:: 1602:: 1593:: 1503:. 1491:. 1468:. 1458:. 1450:. 1436:. 1432:. 1409:. 1399:21 1397:. 1374:. 1364:. 1356:. 1342:. 1338:. 1315:. 1303:. 1289:^ 1275:. 1267:. 1255:. 1232:. 1222:32 1220:. 1197:. 1189:. 1177:. 1153:. 1143:. 1135:. 1123:. 1119:. 1107:^ 1093:. 1081:. 1067:^ 1053:. 1041:. 1029:^ 1015:. 1003:47 1001:. 997:. 971:^ 947:. 920:, 916:, 554:. 286:. 207:: 194:: 181:: 95:Pg 39:Ma 3813:/ 3802:? 3795:† 3785:† 3775:† 3765:† 3755:† 3745:† 3735:† 3725:† 3715:† 3705:† 3695:† 3579:? 3568:? 3511:? 3496:? 3474:? 3459:? 3420:? 3412:? 3390:? 3382:? 3363:† 3319:? 3290:? 3268:? 3190:† 3167:? 3159:? 3137:† 3093:† 3078:† 3068:† 3058:† 3048:† 3044:? 3037:† 3027:† 3017:† 3007:† 2997:† 2987:† 2977:† 2967:† 2957:† 2947:† 2937:† 2927:† 2917:† 2907:† 2897:† 2887:† 2877:† 2867:† 2857:† 2847:† 2837:† 2827:† 2817:† 2807:† 2573:† 2558:† 2548:† 2538:† 2528:† 2518:† 2490:? 2440:† 2410:† 2395:† 2385:† 2375:† 2365:† 2355:† 2345:† 2335:† 2325:† 2315:† 2305:† 2295:† 2240:† 2210:† 2194:? 2165:† 2150:† 2140:† 2130:† 2120:† 2110:† 2100:† 1985:† 1970:† 1928:† 1913:† 1903:† 1893:† 1883:† 1873:† 1863:† 1853:† 1805:† 1768:† 1753:† 1743:† 1733:† 1723:† 1554:e 1547:t 1540:v 1511:. 1499:: 1476:. 1444:: 1438:7 1417:. 1405:: 1382:. 1350:: 1344:5 1323:. 1311:: 1305:4 1283:. 1263:: 1240:. 1228:: 1205:. 1185:: 1161:. 1131:: 1125:6 1101:. 1089:: 1061:. 1049:: 1043:4 1023:. 1009:: 931:. 924:, 435:, 395:, 234:† 221:† 100:N 90:K 85:J 80:T 75:P 70:C 65:D 60:S 55:O 50:Ꞓ 34:~

Index

Induan
Ma
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

Scientific classification
Edit this classification
Eukaryota
Animalia
Chordata
Synapsida
Therapsida
Cynodontia
Galesauridae
Progalesaurus
Type species
galesaurid
cynodont
Triassic
Lystrosaurus Assemblage Zone

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