31:
54:
310:, in 2011, incorporated phylogenetic evidence to recircumscribe the subfamily, combining the grammitids with Mabberley's Polypodieae (recognizing a number of segregate genera). The classification of Christenhusz and Chase in 2014 vastly expanded the circumscription of Polypodiaceae, redefined Polypodioideae be what had been theretofore recognized as Polypodiaceae, including the grammitids. The tribe Polypodieae corresponded to the subfamily Polypodioideae in Christenhusz
207:
classification of 2016 (PPG I) used here, the subfamily
Polypodioideae is one of a number of subfamilies of the family Polypodiaceae which is part of the suborder Polypodiineae. In the second, the whole of the suborder is placed in a very broadly defined Polypodiaceae
332:
The following phylogram shows a likely relationship between
Polypodioideae and the other subfamilies within Polypodiaceae, based on groups described by Schuettpelz & Pryer, 2008, using the subfamily names of PPG I.
216:
is equivalent to the family
Polypodiaceae and the tribe Polypodieae to the subfamily Polypodioideae. The equivalence is shown in the following table.
774:
Maarten J. M. Christenhusz, Xian-Chun Zhang & Herald
Schneider: "A linear sequence of extant families and genera of lycophytes and ferns."
956:
303:) and the grammitid ferns, which he placed in Grammitidaceae. His tribe Polypodieae corresponded to Polypodioideae in the sense of PPG I.
917:
752:
961:
316:
The most recent circumscription of the subfamily, the PPG I classification, largely returns to that of
Christenhusz
943:
287:
Mabberley, in 2008, defined this subfamily in a very broad sense, including all of
Polypodiaceae except for the
522:
204:
178:
53:
904:
1017:
200:
895:
767:
1007:
857:
742:
181:
classification of 2016 (PPG I). Alternatively, the subfamily may be treated as the tribe
974:
8:
541:
1012:
817:
792:
718:
693:
669:
48:
40:
839:
744:
Mabberley's plant-book: a portable dictionary of plants, their classification and uses
969:
822:
748:
723:
673:
812:
804:
713:
705:
659:
909:
846:, ed. Tom A. Ranker and Christopher H. Haufler. Cambridge University Press (2008)
591:
475:
406:
355:
321:
288:
984:
880:
421:
78:
1001:
531:
384:
341:
248:
231:
174:
170:
128:
118:
826:
727:
108:
30:
930:
889:
808:
709:
611:
293:
922:
601:
581:
571:
935:
842:
Eric
Schuettpelz & Kathleen M. Pryer: "Ch. 15. Fern phylogeny" in
664:
647:
551:
166:
948:
851:
648:"A community-derived classification for extant lycophytes and ferns"
874:
299:
203:
of the subfamily were in use as of 2019. In the first, such as the
561:
791:
Christenhusz, Maarten J. M.; Chase, Mark W. (13 February 2014).
692:
Christenhusz, Maarten J. M.; Chase, Mark W. (13 February 2014).
645:
525:
classification of 2016 (PPG I) recognizes the following genera:
187:
65:
98:
88:
324:, rendering the Polypodioideae potentially paraphyletic.
784:
185:within a very broadly defined family Polypodiaceae
646:The Pteridophyte Phylogeny Group (November 2016).
790:
691:
999:
761:
833:
740:
844:Biology and Evolution of Ferns and Lycophytes
734:
793:"Trends and concepts in fern classification"
694:"Trends and concepts in fern classification"
687:
685:
683:
747:. Cambridge University Press. p. 690.
320:, but excludes the grammitids as subfamily
29:
816:
717:
680:
663:
212:, in which the subfamily Polypodioideae
1000:
641:
639:
637:
635:
633:
631:
856:
855:
652:Journal of Systematics and Evolution
173:, which is a member of the suborder
628:
13:
14:
1029:
461:"non-grammitid" = Polypodioideae
225:Christenhusz & Chase (2014)
52:
1:
840:Schuettpelz & Pryer, 2008
622:
169:belonging to the fern family
586:Humb. & Bonpl. ex Willd.
523:Pteridophyte Phylogeny Group
327:
205:Pteridophyte Phylogeny Group
179:Pteridophyte Phylogeny Group
7:
194:
10:
1034:
282:
275:Hook. & Lindl. ex Duby
864:
516:
472:
459:
452:
419:
404:
397:
382:
375:
368:
353:
346:
256:Subfamily Polypodioideae
153:
148:
49:Scientific classification
47:
37:
28:
23:
741:D. J. Mabberley (2008).
781:: 7-54. (18 Feb. 2011)
269:J.Presl & C.Presl
252:J.Presl & C.Presl
241:J.Presl & C.Presl
239:Family Polypodiaceae
474:"grammitid" ferns =
199:Two very different
809:10.1093/aob/mct299
710:10.1093/aob/mct299
273:Tribe Polypodieae
41:Polypodium vulgare
16:Subfamily of ferns
1018:Plant subfamilies
995:
994:
970:Open Tree of Life
858:Taxon identifiers
754:978-0-521-82071-4
665:10.1111/jse.12229
617:
607:
597:
587:
577:
567:
557:
547:
537:
513:
512:
504:
503:
495:
494:
486:
485:
441:
440:
432:
431:
280:
279:
276:
270:
259:
253:
242:
236:
160:
159:
144:
38:Common polypody,
1025:
988:
987:
978:
977:
965:
964:
952:
951:
939:
938:
926:
925:
913:
912:
900:
899:
898:
885:
884:
883:
853:
852:
847:
837:
831:
830:
820:
797:Annals of Botany
788:
782:
765:
759:
758:
738:
732:
731:
721:
698:Annals of Botany
689:
678:
677:
667:
643:
615:
605:
595:
585:
575:
565:
555:
545:
535:
455:
454:
400:
399:
378:
377:
371:
370:
349:
348:
337:
336:
274:
268:
257:
251:
240:
234:
219:
218:
201:circumscriptions
142:
57:
56:
33:
21:
20:
1033:
1032:
1028:
1027:
1026:
1024:
1023:
1022:
998:
997:
996:
991:
983:
981:
973:
968:
960:
955:
947:
942:
934:
929:
921:
916:
908:
903:
894:
893:
888:
879:
878:
873:
860:
850:
838:
834:
789:
785:
766:
762:
755:
739:
735:
690:
681:
644:
629:
625:
620:
592:Pleurosoriopsis
519:
514:
505:
496:
487:
476:Grammitidoideae
442:
433:
407:Platycerioideae
356:Loxogrammoideae
330:
322:Grammitidoideae
289:Platycerioideae
285:
197:
141:
51:
24:Polypodioideae
17:
12:
11:
5:
1031:
1021:
1020:
1015:
1010:
993:
992:
990:
989:
979:
966:
953:
940:
927:
914:
901:
896:Polypodioideae
886:
870:
868:
866:Polypodioideae
862:
861:
849:
848:
832:
803:(4): 571–594.
783:
760:
753:
733:
704:(4): 571–594.
679:
658:(6): 563–603.
626:
624:
621:
619:
618:
608:
598:
588:
578:
568:
558:
548:
538:
527:
518:
515:
511:
510:
507:
506:
502:
501:
498:
497:
493:
492:
489:
488:
484:
483:
480:
479:
471:
468:
467:
464:
463:
458:
453:
451:
448:
447:
444:
443:
439:
438:
435:
434:
430:
429:
426:
425:
422:Microsoroideae
418:
415:
414:
411:
410:
403:
398:
396:
393:
392:
389:
388:
381:
376:
374:
369:
367:
364:
363:
360:
359:
352:
347:
345:
335:
329:
326:
284:
281:
278:
277:
271:
266:Polypodioideae
261:
260:
254:
244:
243:
237:
227:
226:
223:
196:
193:
163:Polypodioideae
158:
157:
151:
150:
146:
145:
143:B.K.Nayar 1970
139:Polypodioideae
136:
132:
131:
126:
122:
121:
116:
112:
111:
106:
102:
101:
99:Polypodiopsida
96:
92:
91:
89:Polypodiophyta
86:
82:
81:
76:
69:
68:
63:
59:
58:
45:
44:
35:
34:
26:
25:
15:
9:
6:
4:
3:
2:
1030:
1019:
1016:
1014:
1011:
1009:
1008:Polypodiaceae
1006:
1005:
1003:
986:
980:
976:
971:
967:
963:
958:
954:
950:
945:
941:
937:
932:
928:
924:
919:
915:
911:
906:
902:
897:
891:
887:
882:
876:
872:
871:
869:
867:
863:
859:
854:
845:
841:
836:
828:
824:
819:
814:
810:
806:
802:
798:
794:
787:
780:
777:
773:
771:
768:Christenhusz
764:
756:
750:
746:
745:
737:
729:
725:
720:
715:
711:
707:
703:
699:
695:
688:
686:
684:
675:
671:
666:
661:
657:
653:
649:
642:
640:
638:
636:
634:
632:
627:
614:
613:
609:
604:
603:
599:
594:
593:
589:
584:
583:
579:
576:(R.Br.) J.Sm.
574:
573:
569:
564:
563:
559:
554:
553:
549:
544:
543:
539:
534:
533:
532:Campyloneurum
529:
528:
526:
524:
509:
508:
500:
499:
491:
490:
482:
481:
478:
477:
470:
469:
466:
465:
462:
457:
456:
450:
449:
446:
445:
437:
436:
428:
427:
424:
423:
417:
416:
413:
412:
409:
408:
402:
401:
395:
394:
391:
390:
387:
386:
385:Drynarioideae
380:
379:
373:
372:
366:
365:
362:
361:
358:
357:
351:
350:
344:
343:
342:Polypodiaceae
339:
338:
334:
325:
323:
319:
314:
313:
309:
306:Christenhusz
304:
302:
301:
296:
295:
290:
272:
267:
263:
262:
255:
250:
249:Polypodiaceae
246:
245:
238:
233:
232:Polypodiineae
229:
228:
224:
221:
220:
217:
215:
211:
206:
202:
192:
190:
189:
184:
180:
176:
175:Polypodiineae
172:
171:Polypodiaceae
168:
164:
156:
152:
147:
140:
137:
134:
133:
130:
129:Polypodiaceae
127:
124:
123:
120:
119:Polypodiineae
117:
114:
113:
110:
107:
104:
103:
100:
97:
94:
93:
90:
87:
84:
83:
80:
79:Tracheophytes
77:
74:
71:
70:
67:
64:
61:
60:
55:
50:
46:
43:
42:
36:
32:
27:
22:
19:
865:
843:
835:
800:
796:
786:
778:
775:
769:
763:
743:
736:
701:
697:
655:
651:
610:
600:
590:
580:
570:
560:
550:
540:
530:
520:
473:
460:
420:
405:
383:
354:
340:
331:
317:
315:
311:
307:
305:
298:
292:
286:
265:
213:
209:
198:
186:
182:
162:
161:
154:
138:
109:Polypodiales
72:
39:
18:
931:iNaturalist
890:Wikispecies
612:Serpocaulon
542:Microgramma
294:Platycerium
183:Polypodieae
135:Subfamily:
1002:Categories
949:17450180-1
623:References
606:L. Perhaps
602:Polypodium
582:Pleopeltis
572:Phlebodium
566:M.G.Price.
264:Subfamily
214:sensu lato
210:sensu lato
188:sensu lato
155:See text.
115:Suborder:
85:Division:
1013:Epiphytes
776:Phytotaxa
552:Niphidium
328:Phylogeny
258:B.K.Nayar
230:Suborder
167:subfamily
62:Kingdom:
982:VASCAN:
881:Q2417265
875:Wikidata
827:24532607
728:24532607
674:39980610
300:Pyrrosia
195:Taxonomy
125:Family:
975:7045578
962:2017695
923:4939709
818:3936591
719:3936591
616:A.R.Sm.
562:Pecluma
546:C.Presl
536:C.Presl
283:History
247:Family
235:Dumort.
177:in the
149:Genera
105:Order:
95:Class:
66:Plantae
936:332945
910:222674
825:
815:
770:et al.
751:
726:
716:
672:
517:Genera
318:et al.
312:et al.
308:et al.
670:S2CID
596:Fomin
556:J.Sm.
222:PPG I
165:is a
73:Clade
957:NCBI
944:IPNI
918:GBIF
905:APNI
823:PMID
772:2011
749:ISBN
724:PMID
521:The
297:and
985:375
813:PMC
805:doi
801:113
714:PMC
706:doi
702:113
660:doi
1004::
972::
959::
946::
933::
920::
907::
892::
877::
821:.
811:.
799:.
795:.
779:19
722:.
712:.
700:.
696:.
682:^
668:.
656:54
654:.
650:.
630:^
191:.
75::
829:.
807::
757:.
730:.
708::
676:.
662::
291:(
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
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