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Piper (plant)

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species are found in the Americas (about 700 species), with about 300 species from Southern Asia. There are smaller groups of species from the South Pacific (about 40 species) and Africa (about 15 species). The American, Asian, and South Pacific groups each appear to be
1350:, which were first isolated from Black Pepper, and reported to have antibiotic activities. Preliminary research reports has shown that piperine has an antibacterial activity against various bacteria such as 1671: 1297:. The diversity and ecological importance of the genus makes it a strong candidate for ecological and evolutionary studies. Most research has focused on the economically important species 1234:
effect, which has led to the banning of kava in many countries. On the other hand, the traditional preparation of the root as a calming drink appears to pose little, if any, such hazard.
884:, trade routes deteriorated and the use of pepper declined somewhat, but peppercorns, storing easily and having a high mass per volume, never ceased to be a profitable trade item. In the 1230:
risk. It has also become popular elsewhere in recent decades, and is used as a medical plant. However, pills that contain parts of the whole plant have occasionally shown a strong
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being black pepper fruit. Today, peppercorns of the three preparations (green, white and black) are one of the most widely used spices of plant origin worldwide.
2259:"Piperine treatment suppresses Helicobacter pylori toxin entry in to gastric epithelium and minimizes β-catenin mediated oncogenesis and IL-8 secretion in vitro" 1493: 947:
suppressed trade in cubeb peppercorns at the end of the 1630s to capitalize on his share of the black pepper trade. It remains a significant spice around the
418:, many of which are dominant species in their native habitat. The diversification of this taxon is of interest to understanding the evolution of plants. 720: 549: 127: 1750:
Chester F. Gorman (1971). "The Hoabinhian and after: subsistence patterns in Southeast Asia during the Late Pleistocene and Early Recent periods".
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Scott, I. M.; Jensen, H.; Nicol, R.; Lesage, L.; Bradbury, R.; Sánchez-Vindas, P.; Poveda, L.; Arnason, J. T.; Philogène, B. J. R. (August 2004).
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Sen, Sandeep; Dayanandan, Selvadurai; Davis, Thomson; Ganesan, Rengaian; Jagadish, M.R.; Mathew, P.J.; Ravikanth, Gudasalamani (September 2019).
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Sen, Sandeep; Dayanandan, Selvadurai; Davis, Thomson; Ganesan, Rengaian; Jagadish, M.R.; Mathew, P.J.; Ravikanth, Gudasalamani (September 2019).
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Culinary use of pepper plants is attested perhaps as early as 9,000 years ago. Peppercorn remains were found among the food refuse left by
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L. Sorrentino; A. Capasso; M. Schmidt (2006). "Safety of ethanolic kava extract: results of a study of chronic toxicity in rats".
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and elsewhere. Later, wars were fought by European powers, between themselves and in complex alliances and enmities with
757:– the only larger region for which comprehensive data exists – more than a dozen species are known to be on the brink of 1933:
Steven T. S. Lim; Klaus Dragull; Chung-Shih Tang; Harry C. Bittenbender; Jimmy T. Efird; Pratibha V. Nerurkar (2007).
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spice mix. This species, despite being traded more extensively in earlier times, is less common outside Africa today.
537:. This contrasts with previous theories assuming a younger radiation of the genus. An earlier potential record is of † 1935:"Effects of kava alkaloid, pipermethystine, and kavalactones on oxidative stress and cytochrome P450 in F-344 rats" 1870: 1294: 461:). A well-known and very close relative – being also part of the Piperaceae – are the radiator plants of the genus 2735: 2691: 2159:"Effects of the natural compounds embelin and piperine on the biofilm-producing property of Streptococcus mutans" 1697:
Chester F. Gorman (1969). "Hoabinhian: a pebble tool complex with early plant associations in Southeast Asia".
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species have a strong influence on their biology, making them ideal systems for research on the evolution of
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plant, called a peppercorn when it is round and pea-sized, as is usual, is distributed in the wild mainly by
17: 865:, a recipe collection complied about 400 AD, mentions "pepper" as a spice for most main dishes. In the late 1408:
Studies have been done to determine the effectiveness of piper leaves to repel different types of insects.
2208:"Inhibitory effect of piperine on Helicobacter pylori growth and adhesion to gastric adenocarcinoma cells" 2566: 2561: 2548: 1679: 2644: 2618: 2838: 2779: 2375: 2369:
M. Alejandra Jaramillo; Paul S. Manos (2001). "Phylogeny and patterns of floral diversity in the genus
2118:"Novel structural analogues of piperine as inhibitors of the NorA efflux pump of Staphylococcus aureus" 1183:
nut slices; its sap helps release the stimulating effect of these "cookies" which are widely known as
2843: 2745: 2670: 1831: 1648: 1817: 144: 2771: 2639: 1932: 1898: 795: 2833: 2613: 2491: 1939: 1227: 869:, black pepper was expensive, but was available readily enough to be used more frequently than 806: 2709: 813:. It is likely that these plants were collected from the wild rather than deliberately grown. 1427: 257: 2758: 2300:"Efficacy of Piper (Piperaceae) extracts for control of common home and garden insect pests" 1107:
border region, and elsewhere it is used to intoxicate fish which then can be easily caught.
2587: 1787: 1708: 1445: 1339: 1262: 958: 1222:, kava is used to produce a calming and socializing drink somewhat similar to alcohol and 8: 1324: 1040: 838: 783: 750: 631: 1712: 1560:
Martínez, Camila; Carvalho, Mónica R.; Madriñán, Santiago; Jaramillo, Carlos A. (2015).
2805: 2626: 2456: 2392: 2275: 2258: 2234: 2207: 2183: 2158: 2093: 2068: 2049: 1805: 1769: 1732: 944: 724:) grow as sizeable, compact and attractive shrubs with tough and shiny leaves. Smaller 676: 265: 139: 2069:"Piperine, a Phytochemical Potentiator of Ciprofloxacin against Staphylococcus aureus" 915:, the fruit of other species are also important spices, many of them internationally. 2753: 2631: 2574: 2460: 2448: 2440: 2400: 2327: 2319: 2280: 2239: 2188: 2139: 2098: 2053: 2041: 2033: 1989: 1958: 1915: 1752: 1724: 1699: 1630: 1615:"The Upper Cretaceous floras of Alaska, with a description of the plant-bearing beds" 1595: 1587: 1258: 1247: 1021: 881: 715: 1982:: a Model Genus for Studies of Evolution, Chemical Ecology, and Trophic Interactions 1736: 2810: 2432: 2384: 2311: 2270: 2229: 2219: 2178: 2170: 2129: 2088: 2080: 2025: 1977: 1948: 1907: 1761: 1716: 1622: 1577: 1370: 1091: 982: 897: 772: 703: 626: 458: 438: 401: 2579: 2540: 2084: 1765: 943:), also known as tailed pepper, played a major role in the spice trade. Reputedly 2553: 2529: 1911: 1835: 1720: 1683: 1498: 1411: 1393: 1251: 1243: 1223: 1147: 1056: 953: 837:
Use of peppercorns as pungent spice is significant on an international scale. By
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the ability to withstand the chemical defences of pepper plants, for example the
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covered in tiny flowers. A somewhat less close relative is the pipevine family (
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is threatening a number of peppers. The extent of the effect of such wholesale
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is not for its looks however, but ultimately for the wide range of powerful
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International Union for the Conservation of Nature and Natural Resources
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states, in part about control of the supply of spices, perhaps the most
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distribution, and are most commonly found in the understory of lowland
542: 442: 426: 422: 404: 215: 195: 95: 60: 1773: 1566:(Piperaceae) from Colombia and diversification patterns for the genus" 816: 533:
into the multiple extant clades, coinciding with the final breakup of
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are used in cooking. West African Pepper leaves, known locally as
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in noticeable quantities) was in turn named after piperidine.
2418:"Origin and evolution of the genus Piper in Peninsular India" 2011:"Origin and evolution of the genus Piper in Peninsular India" 1676: 1530: 1522: 1226:
but without many of the negative side effects and less of an
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pepper plants. Both families have characteristic tail-shaped
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Monkey Insect Repellent | The Life of Mammals | BBC Earth
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as well as, particularly in the early 20th century, as a
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and the effect of mutualisms on biotic communities.
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or warmer regions. Pepper vines can be used much as
624:. A few species, commonly called "ant pipers" (e.g. 445:, is most closely related to the lizardtail family ( 2256: 2205: 1418:rubbing the piper leaves on them to repel insects. 1441:; the affinity of the African species is unclear. 1384:The significant secondary metabolites of kava are 477:and the common name "pepper" are derived from the 2163:Journal of Traditional and Complementary Medicine 1850: 1749: 1696: 1242:The genus contains species suitable for studying 1131:is sometimes used in herbalism, and long pepper ( 2825: 396:, is an economically and ecologically important 753:on the genus is unknown, but in the forests of 1830: 1555: 1553: 718:, while other species, like lacquered pepper ( 2257:Tharmalingam, N.; Park, M.; LEE, M.H (2016). 2206:Tharmalingam, N.; Kim, SH.; Park, M. (2014). 1612: 1366:toxin entry to gastric epithelial cells. The 1218:region, where it has been widely spread as a 2364: 2362: 1789:Apicius: Cookery and Dining in Imperial Rome 951:region today, however. West African pepper ( 1550: 1400:compound in this plant's stems and leaves. 1015:) is used to wrap meat for grilling in the 583:; one species – the Japanese Pepper ( 563: 2263:American Journal of Translational Research 1988:. Boston, MA: Kluwer Academic Publishers. 1025:) is used raw or cooked as a vegetable in 116: 2359: 2274: 2233: 2223: 2182: 2133: 2092: 1952: 1837:Theatrum Botanicum: the Theater of Plants 1581: 789: 1677:2007 IUCN Red List of Threatened Species 1267: 1167:species has gained large-scale use as a 1050: 815: 2156: 981:, are used as a flavoring vegetable in 927:spice internationally; it has a rather 923:), is possibly the second-most popular 888:, international traders were nicknamed 14: 2826: 1613:Hollick, Arthur; Martin, G.C. (1930). 794:For other spices called "pepper", see 525:appears to be nested within the clade 2490: 2489: 2425:Molecular Phylogenetics and Evolution 2122:Journal of Antimicrobial Chemotherapy 2115: 2073:Antimicrobial Agents and Chemotherapy 2018:Molecular Phylogenetics and Evolution 2746:e889a822-477b-46af-97f6-6e3478c6f6bb 2066: 1403: 1143:and (together with black pepper) of 24: 1007:have major importance in cooking: 786:found particularly in the fruits. 778:The most significant human use of 25: 2855: 2468: 2157:Dwivedi, D.; et al. (2016). 1976:L. A. Dyer; A. N. Palmer (2004). 1873:. Earthcare. 2008. Archived from 1476:, but other sources keep them in 2785:urn:lsid:ipni.org:names:326982-2 2475:The families of flowering plants 1295:ecology and evolutionary biology 1139:, where it was an ingredient of 935:is used as the fruits are tiny. 911:Due to the wide distribution of 690:. The latter can be significant 143: 48: 2338: 2291: 2250: 2199: 2150: 2116:Kumar, A.; et al. (2008). 2109: 2067:Khan, IA.; et al. (2006). 2060: 2002: 1969: 1926: 1889: 1863: 1844: 1373:is named after the former, and 1278:, extracted from black pepper ( 961:, and is sometimes used in the 2304:Journal of Economic Entomology 1824: 1780: 1743: 1690: 1665: 1641: 1606: 1511: 1358:, and gastric cancer pathogen 1046: 410:It contains about 1,000–2,000 13: 1: 2085:10.1128/aac.50.2.810-812.2006 1766:10.1080/00438243.1971.9979482 1544: 1320:The obligate and facultative 1237: 1060: 829:Green (pickled unripe fruits) 765:species (e.g. spiked pepper, 702:Many pepper plants make good 2212:Infectious Agents and Cancer 1912:10.1016/j.phymed.2006.01.006 1721:10.1126/science.163.3868.671 1396:is suspected to be the main 999:) has a variety of uses. In 827:) corns, from left to right: 556:, although this affinity to 496: 421:Pepper plants belong to the 7: 2437:10.1016/j.ympev.2019.05.033 2175:10.1016/j.jtcme.2014.11.025 2030:10.1016/j.ympev.2019.05.033 1537:were also employed by many. 1444:Some species are sometimes 1342:found in pepper plants are 1135:) is similarly employed in 1003:, leaves of two species of 931:-like "heat" and the whole 833:Black (dried unripe fruits) 761:. On the other hand, other 10: 2860: 2376:American Journal of Botany 1857:Encyclopedia of Asian Food 1851:Charmaine Solomon (1998). 1570:American Journal of Botany 1497:, a plant from the family 1425: 1421: 1179:) leaves are used to wrap 1123:properties. Black pepper ( 1019:region, while wild betel ( 793: 560:is not entirely reliable. 2498: 2481:Piper species in Thailand 1649:"Yukok_Koyukuk_2_Geology" 1095:is used medically by the 1033:; The stems and roots of 880:As Europe moved into the 697: 646:, but small fruit-eating 271: 264: 254: 249: 140:Scientific classification 138: 124: 115: 34: 1533:and large quantities of 1504: 845:including black pepper ( 831:White (dried ripe seeds) 564:Distribution and ecology 1377:(which is not found in 1115:appears to have strong 1039:are used as a spice in 993:or Mexican pepperleaf ( 957:), is commonly used in 841:, there was a vigorous 796:Pepper (disambiguation) 728:, like Celebes pepper ( 501:The earliest fossil of 2225:10.1186/1750-9378-9-43 1940:Toxicological Sciences 1432:The largest number of 1414:have been recorded by 1283: 1067: 973:Not only the seeds of 834: 790:As spice and vegetable 414:of shrubs, herbs, and 2316:10.1093/jee/97.4.1390 1954:10.1093/toxsci/kfm035 1871:"Tropical root crops" 1428:List of Piper species 1340:secondary metabolites 1271: 1054: 989:-influenced cooking, 819: 1859:. Periplus Editions. 1494:Lacistema aggregatum 1489:P. fasciculatum 1483:The species called " 1356:Streptococcus mutans 1263:evolutionary biology 1194:Conversely, another 959:West African cuisine 892:("pepper-sacks") in 1840:. London: T. Cotes. 1713:1969Sci...163..671G 1583:10.3732/ajb.1400427 1360:Helicobacter pylori 1317:hot spot in India. 1303:P. methysticum 1206:), is used for its 1204:P. methysticum 1077:) has been used in 1041:Bangladeshi cuisine 1022:P. sarmentosum 839:classical antiquity 784:secondary compounds 751:habitat destruction 694:to pepper growers. 638:. The fruit of the 620:or almost as small 608:species are either 2135:10.1093/jac/dkn088 1682:2018-05-08 at the 1619:Professional Paper 1562:"A Late Cretaceous 1313:originated in the 1284: 1156:(rejuvenating and 1092:P. darienense 1068: 1057:Kava club in Tonga 945:Philip IV of Spain 835: 775:in certain areas. 721:P. magnificum 716:temperate climates 677:Homidiana subpicta 310:Trel. & Stehlé 125:Lacquered pepper ( 2839:Pantropical flora 2821: 2820: 2754:Open Tree of Life 2492:Taxon identifiers 1995:978-0-306-48498-8 1753:World Archaeology 1707:(3868): 671–673. 1653:clamp.ibcas.ac.cn 1404:Repelling insects 1259:community ecology 1248:molecular biology 954:P. guineense 882:Early Middle Ages 593:and southernmost 381: 380: 375: 367: 359: 351: 347:Pleiostachyopiper 343: 335: 327: 319: 311: 303: 295: 287: 279: 245: 16:(Redirected from 2851: 2844:Piperales genera 2814: 2813: 2801: 2800: 2788: 2787: 2775: 2774: 2762: 2761: 2749: 2748: 2739: 2738: 2726: 2725: 2713: 2712: 2700: 2699: 2687: 2686: 2674: 2673: 2661: 2660: 2648: 2647: 2635: 2634: 2622: 2621: 2609: 2608: 2596: 2595: 2583: 2582: 2570: 2569: 2557: 2556: 2544: 2543: 2534: 2533: 2532: 2519: 2518: 2517: 2487: 2486: 2464: 2422: 2409: 2408: 2366: 2357: 2356: 2355: 2354: 2342: 2336: 2335: 2310:(4): 1390–1403. 2295: 2289: 2288: 2278: 2254: 2248: 2247: 2237: 2227: 2203: 2197: 2196: 2186: 2154: 2148: 2147: 2137: 2128:(6): 1270–1276. 2113: 2107: 2106: 2096: 2064: 2058: 2057: 2015: 2006: 2000: 1999: 1987: 1973: 1967: 1966: 1956: 1930: 1924: 1923: 1893: 1887: 1886: 1884: 1882: 1867: 1861: 1860: 1848: 1842: 1841: 1828: 1822: 1821: 1815: 1811: 1809: 1801: 1799: 1797: 1784: 1778: 1777: 1747: 1741: 1740: 1694: 1688: 1669: 1663: 1662: 1660: 1659: 1645: 1639: 1638: 1610: 1604: 1603: 1585: 1557: 1538: 1515: 1485:Piper aggregatum 1448:into the genera 1412:Capuchin monkeys 1371:functional group 1301:(black pepper), 1293:for research in 1252:natural products 1214:effects. In the 1141:Triphala Guggulu 1065: 1062: 898:Hanseatic League 773:invasive species 627:Piper cenocladum 589:) from southern 577:tropical forests 550:Kaltag Formation 507:Piper margaritae 459:Aristolochiaceae 373: 365: 357: 355:Pleistachyopiper 349: 341: 333: 325: 317: 309: 301: 293: 285: 277: 275:Anderssoniopiper 241: 148: 147: 120: 110: 47: 40:Temporal range: 32: 31: 21: 2859: 2858: 2854: 2853: 2852: 2850: 2849: 2848: 2824: 2823: 2822: 2817: 2809: 2804: 2796: 2791: 2783: 2778: 2770: 2765: 2757: 2752: 2744: 2742: 2734: 2729: 2721: 2716: 2708: 2703: 2695: 2690: 2682: 2677: 2669: 2664: 2656: 2651: 2643: 2638: 2630: 2625: 2617: 2612: 2604: 2599: 2591: 2586: 2578: 2573: 2565: 2560: 2552: 2547: 2539: 2537: 2528: 2527: 2522: 2513: 2512: 2507: 2494: 2471: 2420: 2412: 2389:10.2307/2657072 2373:(Piperaceae)". 2367: 2360: 2352: 2350: 2344: 2343: 2339: 2296: 2292: 2255: 2251: 2204: 2200: 2155: 2151: 2114: 2110: 2065: 2061: 2013: 2007: 2003: 1996: 1985: 1974: 1970: 1931: 1927: 1894: 1890: 1880: 1878: 1877:on 19 July 2008 1869: 1868: 1864: 1849: 1845: 1829: 1825: 1813: 1812: 1803: 1802: 1795: 1793: 1786: 1785: 1781: 1748: 1744: 1695: 1691: 1684:Wayback Machine 1674:(IUCN) (2007): 1670: 1666: 1657: 1655: 1647: 1646: 1642: 1611: 1607: 1558: 1551: 1547: 1542: 1541: 1520:generally uses 1516: 1512: 1507: 1499:Lacistemataceae 1491:" are actually 1430: 1424: 1406: 1394:Pipermethystine 1244:natural history 1240: 1224:benzodiazepines 1111:, often called 1063: 1049: 996:P. auritum 843:trade of spices 832: 830: 828: 799: 792: 747:primary forests 731:P. ornatum 706:for gardens in 700: 616:; some grow as 571:species have a 566: 539:Piper arcuatile 511:Late Cretaceous 499: 485:, denoting the 472:scientific name 256:1000–2000; see 240: 142: 111: 109: 108: 103: 98: 93: 88: 83: 78: 73: 68: 63: 58: 53: 42: 41: 38: 28: 27:Genus of plants 23: 22: 15: 12: 11: 5: 2857: 2847: 2846: 2841: 2836: 2819: 2818: 2816: 2815: 2811:wfo-4000029808 2802: 2789: 2776: 2763: 2750: 2740: 2727: 2714: 2701: 2688: 2675: 2662: 2649: 2636: 2623: 2610: 2597: 2584: 2571: 2558: 2545: 2535: 2520: 2504: 2502: 2496: 2495: 2484: 2483: 2478: 2470: 2469:External links 2467: 2466: 2465: 2411: 2410: 2383:(4): 706–716. 2358: 2337: 2290: 2249: 2198: 2149: 2108: 2079:(2): 810–812. 2059: 2001: 1994: 1968: 1947:(1): 214–221. 1925: 1906:(8): 542–549. 1888: 1862: 1843: 1832:John Parkinson 1823: 1814:|website= 1779: 1760:(3): 300–320. 1742: 1689: 1687:. IUCN, Gland. 1664: 1640: 1605: 1576:(2): 273–289. 1548: 1546: 1543: 1540: 1539: 1509: 1508: 1506: 1503: 1423: 1420: 1405: 1402: 1362:and decreased 1327:found in some 1239: 1236: 1133:P. longum 1125:P. nigrum 1048: 1045: 1001:Southeast Asia 941:P. cubeba 921:P. longum 848:P. nigrum 791: 788: 741:Unsustainable 699: 696: 565: 562: 498: 495: 455:inflorescences 379: 378: 377: 376: 368: 360: 352: 344: 336: 328: 320: 312: 304: 296: 288: 280: 269: 268: 262: 261: 252: 251: 247: 246: 233: 229: 228: 223: 219: 218: 213: 209: 208: 203: 199: 198: 193: 186: 185: 180: 173: 172: 167: 160: 159: 154: 150: 149: 136: 135: 122: 121: 113: 112: 104: 99: 94: 89: 84: 79: 74: 69: 64: 59: 54: 49: 39: 26: 9: 6: 4: 3: 2: 2856: 2845: 2842: 2840: 2837: 2835: 2834:Piper (plant) 2832: 2831: 2829: 2812: 2807: 2803: 2799: 2794: 2790: 2786: 2781: 2777: 2773: 2768: 2764: 2760: 2755: 2751: 2747: 2741: 2737: 2732: 2728: 2724: 2719: 2715: 2711: 2706: 2702: 2698: 2693: 2689: 2685: 2680: 2676: 2672: 2667: 2663: 2659: 2654: 2650: 2646: 2641: 2637: 2633: 2628: 2624: 2620: 2615: 2611: 2607: 2602: 2598: 2594: 2589: 2585: 2581: 2576: 2572: 2568: 2563: 2559: 2555: 2550: 2546: 2542: 2536: 2531: 2525: 2521: 2516: 2510: 2506: 2505: 2503: 2501: 2497: 2493: 2488: 2482: 2479: 2476: 2473: 2472: 2462: 2458: 2454: 2450: 2446: 2442: 2438: 2434: 2430: 2426: 2419: 2414: 2413: 2406: 2402: 2398: 2394: 2390: 2386: 2382: 2378: 2377: 2372: 2365: 2363: 2349: 2348: 2341: 2333: 2329: 2325: 2321: 2317: 2313: 2309: 2305: 2301: 2294: 2286: 2282: 2277: 2272: 2269:(8): 885–98. 2268: 2264: 2260: 2253: 2245: 2241: 2236: 2231: 2226: 2221: 2217: 2213: 2209: 2202: 2194: 2190: 2185: 2180: 2176: 2172: 2168: 2164: 2160: 2153: 2145: 2141: 2136: 2131: 2127: 2123: 2119: 2112: 2104: 2100: 2095: 2090: 2086: 2082: 2078: 2074: 2070: 2063: 2055: 2051: 2047: 2043: 2039: 2035: 2031: 2027: 2023: 2019: 2012: 2005: 1997: 1991: 1984: 1983: 1979: 1972: 1964: 1960: 1955: 1950: 1946: 1942: 1941: 1936: 1929: 1921: 1917: 1913: 1909: 1905: 1901: 1900: 1899:Phytomedicine 1892: 1876: 1872: 1866: 1858: 1854: 1847: 1839: 1838: 1833: 1827: 1819: 1807: 1791: 1790: 1783: 1775: 1771: 1767: 1763: 1759: 1755: 1754: 1746: 1738: 1734: 1730: 1726: 1722: 1718: 1714: 1710: 1706: 1702: 1701: 1693: 1686: 1685: 1681: 1678: 1673: 1668: 1654: 1650: 1644: 1636: 1632: 1628: 1627:10.3133/pp159 1624: 1620: 1616: 1609: 1601: 1597: 1593: 1589: 1584: 1579: 1575: 1571: 1567: 1565: 1556: 1554: 1549: 1536: 1532: 1529:instead; raw 1528: 1525: 1524: 1519: 1514: 1510: 1502: 1500: 1496: 1495: 1490: 1486: 1481: 1479: 1475: 1471: 1470:Trianaeopiper 1467: 1463: 1459: 1455: 1451: 1447: 1442: 1440: 1435: 1429: 1419: 1417: 1413: 1409: 1401: 1399: 1395: 1391: 1387: 1382: 1380: 1376: 1372: 1369: 1365: 1361: 1357: 1353: 1349: 1345: 1341: 1336: 1334: 1330: 1326: 1323: 1318: 1316: 1315:Western Ghats 1312: 1308: 1307:P. betle 1304: 1300: 1296: 1292: 1288: 1281: 1277: 1274: 1270: 1266: 1264: 1260: 1256: 1253: 1249: 1245: 1235: 1233: 1229: 1225: 1221: 1217: 1213: 1209: 1205: 1201: 1197: 1192: 1190: 1186: 1182: 1178: 1174: 1170: 1166: 1161: 1159: 1155: 1154: 1149: 1146: 1142: 1138: 1134: 1130: 1129:essential oil 1126: 1122: 1118: 1114: 1110: 1109:Spiked pepper 1106: 1102: 1098: 1094: 1093: 1088: 1084: 1080: 1079:folk medicine 1076: 1072: 1058: 1053: 1044: 1042: 1038: 1037: 1032: 1028: 1024: 1023: 1018: 1014: 1013:P. lolot 1010: 1006: 1002: 998: 997: 992: 988: 984: 980: 976: 971: 969: 968: 964: 960: 956: 955: 950: 946: 942: 938: 934: 933:inflorescence 930: 926: 922: 918: 914: 909: 907: 903: 899: 896:towns of the 895: 891: 887: 883: 878: 876: 872: 868: 864: 863: 858: 854: 850: 849: 844: 840: 826: 822: 818: 814: 812: 808: 804: 797: 787: 785: 781: 776: 774: 770: 769: 764: 760: 756: 752: 748: 744: 739: 737: 733: 732: 727: 723: 722: 717: 713: 709: 705: 695: 693: 689: 688: 684:of the genus 683: 679: 678: 674: 671: 667: 663: 659: 658: 654:of the genus 653: 649: 645: 641: 637: 633: 630:), live in a 629: 628: 623: 619: 615: 611: 607: 602: 600: 596: 592: 588: 587: 582: 581:cloud forests 578: 574: 570: 561: 559: 555: 551: 548: 544: 540: 536: 532: 528: 524: 523:P. margaritae 520: 516: 515:Maastrichtian 512: 508: 504: 494: 492: 488: 484: 480: 476: 473: 468: 466: 465: 460: 456: 452: 448: 444: 440: 436: 432: 428: 424: 419: 417: 413: 408: 406: 403: 399: 395: 391: 390:pepper plants 387: 386: 372: 371:Trianaeopiper 369: 364: 361: 356: 353: 348: 345: 340: 337: 332: 329: 324: 323:Lindeniopiper 321: 316: 313: 308: 305: 300: 297: 292: 289: 284: 281: 276: 273: 272: 270: 267: 263: 260: 259: 253: 248: 244: 239: 238: 234: 231: 230: 227: 224: 221: 220: 217: 214: 211: 210: 207: 204: 201: 200: 197: 194: 191: 188: 187: 184: 181: 178: 175: 174: 171: 170:Tracheophytes 168: 165: 162: 161: 158: 155: 152: 151: 146: 141: 137: 134: 133:inflorescence 130: 129: 128:P. magnificum 123: 119: 114: 107: 102: 97: 92: 87: 82: 77: 72: 67: 62: 57: 52: 45: 44:Maastrichtian 37: 33: 30: 19: 18:Piper (genus) 2499: 2428: 2424: 2380: 2374: 2370: 2351:, retrieved 2346: 2340: 2307: 2303: 2293: 2266: 2262: 2252: 2215: 2211: 2201: 2169:(1): 57–61. 2166: 2162: 2152: 2125: 2121: 2111: 2076: 2072: 2062: 2021: 2017: 2004: 1981: 1978: 1971: 1944: 1938: 1928: 1903: 1897: 1891: 1879:. Retrieved 1875:the original 1865: 1856: 1846: 1836: 1826: 1794:. Retrieved 1792:. 2009-08-19 1788: 1782: 1757: 1751: 1745: 1704: 1698: 1692: 1675: 1667: 1656:. Retrieved 1652: 1643: 1618: 1608: 1573: 1569: 1563: 1521: 1517: 1513: 1492: 1488: 1484: 1482: 1477: 1473: 1469: 1466:Sarcorhachis 1465: 1461: 1457: 1453: 1449: 1443: 1439:monophyletic 1433: 1431: 1410: 1407: 1390:flavokawains 1386:kavalactones 1383: 1378: 1363: 1359: 1355: 1351: 1337: 1328: 1319: 1310: 1306: 1305:(kava), and 1302: 1298: 1286: 1285: 1280:Piper nigrum 1279: 1273:Crystallized 1241: 1203: 1195: 1193: 1184: 1176: 1164: 1162: 1160:) purposes. 1151: 1144: 1140: 1132: 1124: 1117:disinfectant 1112: 1090: 1074: 1069: 1034: 1031:Thai cuisine 1020: 1012: 1004: 994: 990: 978: 974: 972: 965: 963:East African 952: 949:Indian Ocean 940: 924: 920: 912: 910: 902:Indian Ocean 890:Pfeffersäcke 889: 879: 867:Roman Empire 860: 846: 836: 825:Piper nigrum 824: 821:Black pepper 805:artisans at 800: 779: 777: 766: 762: 745:of tropical 740: 729: 719: 701: 685: 682:flea beetles 675: 664:, some have 655: 639: 625: 605: 603: 584: 568: 567: 557: 538: 530: 526: 522: 506: 502: 500: 490: 482: 474: 469: 462: 429:but neither 425:, which are 420: 409: 394:pepper vines 393: 389: 384: 383: 382: 370: 362: 354: 346: 338: 330: 322: 314: 306: 298: 290: 282: 274: 255: 236: 235: 189: 176: 163: 126: 35: 29: 2679:iNaturalist 2524:Wikispecies 2431:: 102–113. 2024:: 102–113. 1796:December 9, 1462:Arctottonia 1450:Pothomorphe 1398:hepatotoxic 1291:model genus 1232:hepatotoxic 1220:canoe plant 1158:detoxifying 1150:, used for 1097:Kuna people 1089:flavoring. 1064: 2005 1047:As medicine 1036:Piper chaba 917:Long pepper 886:Middle Ages 807:Spirit Cave 708:subtropical 704:ornamentals 573:pantropical 487:long pepper 447:Saururaceae 427:angiosperms 363:Pothomorphe 283:Arctottonia 226:Piperoideae 222:Subfamily: 183:Angiosperms 2828:Categories 2353:2023-09-25 1658:2023-06-19 1545:References 1527:fish sauce 1454:Macropiper 1446:segregated 1426:See also: 1375:piperazine 1368:piperidine 1338:Important 1325:mutualists 1238:In science 1212:euphoriant 1208:depressant 1181:betel palm 1121:antibiotic 991:hoja santa 985:stews. In 906:archetypal 853:South Asia 803:Hoabinhian 768:P. aduncum 759:extinction 736:pot plants 670:sematurine 662:herbivores 610:herbaceous 586:P. kadsura 543:Cenomanian 527:Schilleria 451:amphibious 443:Piperaceae 423:magnoliids 405:Piperaceae 331:Macropiper 315:Lepianthes 307:Discipiper 216:Piperaceae 196:Magnoliids 2461:167218493 2445:1055-7903 2324:0022-0493 2218:(9): 43. 2054:167218493 2038:1055-7903 1853:"Cha plu" 1816:ignored ( 1806:cite book 1635:2330-7102 1592:0002-9122 1416:BBC Earth 1379:P. nigrum 1364:H. pylori 1352:S. aureus 1348:chavicine 1333:symbioses 1311:P. nigrum 1299:P. nigrum 1255:chemistry 1228:addiction 1198:species, 1169:stimulant 1087:cigarette 1083:herbalism 1075:P. cubeba 1017:Indochina 632:mutualism 547:Santonian 541:from the 509:from the 497:Evolution 491:P. longum 464:Peperomia 206:Piperales 153:Kingdom: 46:- present 2798:40009192 2793:Tropicos 2509:Wikidata 2453:31132521 2405:11302858 2332:15384353 2285:27158376 2244:25584066 2193:26870681 2144:18334493 2103:16436753 2046:31132521 1963:17329236 1920:16904878 1834:(1640). 1737:34052655 1729:17742735 1680:Archived 1600:25667080 1474:Zippelia 1344:piperine 1276:piperine 1177:P. betle 1153:rasayana 1137:Ayurveda 1105:Colombia 983:Nigerian 811:Thailand 680:or some 657:Carollia 535:Gondwana 519:Colombia 479:Sanskrit 437:. Their 435:eudicots 431:monocots 291:Artanthe 266:Synonyms 250:Species 212:Family: 2710:1298322 2697:32042-1 2658:3075433 2515:Q159536 2397:2657072 2276:4846933 2235:4290101 2184:4738039 2094:1366922 1881:15 July 1709:Bibcode 1700:Science 1518:Apicius 1487:" and " 1458:Ottonia 1422:Species 1216:Pacific 1145:Trikatu 1099:of the 987:Mexican 967:berbere 862:Apicius 851:) from 755:Ecuador 743:logging 726:species 666:evolved 650:– e.g. 648:mammals 505:is of † 483:pippali 412:species 400:in the 342:Spreng. 339:Ottonia 299:Chavica 232:Genus: 202:Order: 157:Plantae 2767:PLANTS 2759:283089 2743:NZOR: 2645:125531 2619:125531 2567:148718 2541:193742 2538:APDB: 2459:  2451:  2443:  2403:  2395:  2330:  2322:  2283:  2273:  2242:  2232:  2191:  2181:  2142:  2101:  2091:  2052:  2044:  2036:  1992:  1961:  1918:  1774:124120 1772:  1735:  1727:  1633:  1598:  1590:  1472:, and 1261:, and 1113:matico 1101:Panama 894:German 859:. The 857:Europe 698:Usages 618:shrubs 439:family 416:lianas 402:family 388:, the 2772:PIPER 2736:13215 2723:18301 2705:IRMNG 2684:83584 2632:Piper 2627:FoAO2 2606:1PIPG 2593:52541 2554:78830 2530:Piper 2500:Piper 2457:S2CID 2421:(PDF) 2393:JSTOR 2371:Piper 2050:S2CID 2014:(PDF) 1986:(PDF) 1980:Piper 1770:JSTOR 1733:S2CID 1564:Piper 1535:herbs 1531:brine 1523:garum 1505:Notes 1478:Piper 1434:Piper 1329:Piper 1289:is a 1287:Piper 1196:Piper 1189:India 1173:Betel 1165:Piper 1148:pills 1071:Cubeb 1055:At a 1027:Malay 1009:lolot 1005:Piper 979:uziza 975:Piper 937:Cubeb 929:chili 925:Piper 913:Piper 875:sugar 780:Piper 763:Piper 692:pests 687:Lanka 644:birds 640:Piper 634:with 622:trees 614:vines 606:Piper 604:Most 599:frost 595:Korea 591:Japan 569:Piper 558:Piper 554:Yukon 531:Piper 517:) of 503:Piper 481:term 475:Piper 398:genus 385:Piper 374:Trel. 358:Trel. 350:Trel. 326:Trel. 286:Trel. 278:Trel. 237:Piper 190:Clade 177:Clade 164:Clade 36:Piper 2780:POWO 2731:NCBI 2718:ITIS 2692:IPNI 2671:9421 2666:GRIN 2653:GBIF 2601:EPPO 2580:6QRC 2562:BOLD 2549:APNI 2449:PMID 2441:ISSN 2401:PMID 2328:PMID 2320:ISSN 2281:PMID 2240:PMID 2189:PMID 2140:PMID 2099:PMID 2042:PMID 2034:ISSN 1990:ISBN 1959:PMID 1916:PMID 1883:2008 1818:help 1798:2018 1725:PMID 1631:ISSN 1596:PMID 1588:ISSN 1388:and 1346:and 1210:and 1200:kava 1163:One 1119:and 1081:and 1029:and 871:salt 673:moth 652:bats 636:ants 470:The 433:nor 366:Miq. 334:Miq. 318:Raf. 302:Miq. 294:Miq. 258:list 51:PreꞒ 2806:WFO 2640:FoC 2614:FNA 2588:EoL 2575:CoL 2433:doi 2429:138 2385:doi 2312:doi 2271:PMC 2230:PMC 2220:doi 2179:PMC 2171:doi 2130:doi 2089:PMC 2081:doi 2026:doi 2022:138 1949:doi 1908:doi 1762:doi 1717:doi 1705:163 1623:doi 1578:doi 1574:102 1322:ant 1187:in 1185:pan 873:or 855:to 714:in 712:ivy 612:or 552:of 545:to 493:). 392:or 2830:: 2808:: 2795:: 2782:: 2769:: 2756:: 2733:: 2720:: 2707:: 2694:: 2681:: 2668:: 2655:: 2642:: 2629:: 2616:: 2603:: 2590:: 2577:: 2564:: 2551:: 2526:: 2511:: 2455:. 2447:. 2439:. 2427:. 2423:. 2399:. 2391:. 2381:88 2379:. 2361:^ 2326:. 2318:. 2308:97 2306:. 2302:. 2279:. 2267:15 2265:. 2261:. 2238:. 2228:. 2216:16 2214:. 2210:. 2187:. 2177:. 2165:. 2161:. 2138:. 2126:61 2124:. 2120:. 2097:. 2087:. 2077:50 2075:. 2071:. 2048:. 2040:. 2032:. 2020:. 2016:. 1957:. 1945:97 1943:. 1937:. 1914:. 1904:13 1902:. 1855:. 1810:: 1808:}} 1804:{{ 1768:. 1756:. 1731:. 1723:. 1715:. 1703:. 1651:. 1629:. 1621:. 1617:. 1594:. 1586:. 1572:. 1568:. 1552:^ 1501:. 1480:. 1468:, 1464:, 1460:, 1456:, 1452:, 1392:. 1354:, 1265:. 1257:, 1250:, 1246:, 1191:. 1171:. 1127:) 1061:c. 1043:. 877:. 809:, 738:. 521:. 467:. 441:, 407:. 243:L. 192:: 179:: 166:: 131:) 101:Pg 2463:. 2435:: 2407:. 2387:: 2334:. 2314:: 2287:. 2246:. 2222:: 2195:. 2173:: 2167:6 2146:. 2132:: 2105:. 2083:: 2056:. 2028:: 1998:. 1965:. 1951:: 1922:. 1910:: 1885:. 1820:) 1800:. 1776:. 1764:: 1758:2 1739:. 1719:: 1711:: 1661:. 1637:. 1625:: 1602:. 1580:: 1282:) 1202:( 1175:( 1103:- 1073:( 1066:) 1059:( 1011:( 939:( 919:( 823:( 798:. 513:( 489:( 106:N 96:K 91:J 86:T 81:P 76:C 71:D 66:S 61:O 56:Ꞓ 20:)

Index

Piper (genus)
Maastrichtian
PreꞒ

O
S
D
C
P
T
J
K
Pg
N

P. magnificum
inflorescence
Scientific classification
Edit this classification
Plantae
Tracheophytes
Angiosperms
Magnoliids
Piperales
Piperaceae
Piperoideae
Piper
L.
list
Synonyms

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