62:
40:
568:
The closest relatives of the
Piesmatidae remain rather insufficiently determined. After the ash-grey leaf bugs were recognized as Pentatomomorpha, they were most often placed in the
48:
262:
215:
1640:
2685:
2361:
2102:
1862:
1687:
2371:
2060:
2034:
2724:
549:
543:
471:
316:
593:
510:
483:
447:
430:
420:
345:
714:
Cassis, G., & R.T. Schuh (2010) Systematic methods, fossils, and relationships within
Heteroptera (Insecta). Cladistics 26:262-280.
500:
410:
327:
2659:
533:
373:
294:
2698:
2703:
742:
2729:
2638:
2651:
690:
2783:
2711:
487:
was for some time placed in this family, but it has more recently been suggested that it is a member of the
663:(2007): An Unusual, Primitive Piesmatidae (Insecta: Heteroptera) in Cretaceous Amber from Myanmar (Burma).
2573:
2568:
2545:
17:
677:
61:
537:
but given its distribution it might more comfortably be considered closely related to the ancestor of
672:
230:
and the firm part of the wings are extensively dimpled. This resembles the similar pattern of the
735:
2755:
2498:
2750:
2690:
2536:
565:) and was initially considered to be a primitive piesmatid but this has since been rejected.
2742:
2778:
2607:
242:
8:
194:
1661:
1538:
1529:
728:
56:
2737:
2594:
1321:
1281:
1043:
562:
505:
2599:
2550:
2404:
2346:
2341:
2175:
2086:
1849:
1627:
1619:
1546:
1329:
1191:
581:
488:
179:
2399:
2366:
2305:
2209:
2204:
2141:
2116:
1857:
1710:
1551:
1515:
1505:
1487:
1423:
1078:
811:
257:
2716:
2646:
2633:
2521:
2425:
2183:
2065:
1828:
1799:
1601:
1510:
1495:
1366:
1353:
1260:
1231:
1111:
1015:
899:
858:
609:
253:
2772:
2473:
2461:
2433:
2292:
2232:
2214:
2170:
2154:
2039:
2024:
1922:
1671:
1656:
1648:
1382:
1305:
1297:
1207:
1083:
1020:
985:
946:
933:
868:
848:
515:
206:
720:
2394:
2245:
2070:
2029:
1991:
1974:
1960:
1948:
1901:
1893:
1739:
1726:
1692:
1500:
1467:
1413:
1289:
1252:
1215:
1170:
995:
920:
912:
823:
795:
352:
238:
39:
2672:
2560:
2530:
2469:
2323:
2258:
2240:
2222:
2191:
2019:
1820:
1568:
1460:
1418:
1408:
1403:
1313:
1244:
1236:
1199:
1068:
1063:
1005:
977:
960:
356:
123:
2586:
241:, and was initially taken to signify a close relationship. It is due to
2356:
2315:
2297:
2078:
1983:
1880:
1872:
1836:
1753:
1395:
1268:
1186:
1073:
1058:
1053:
1030:
938:
925:
863:
853:
832:
569:
554:
477:
458:
383:
334:
288:
235:
154:
133:
2677:
607:
604:. For some time, the Psamminae were even included in the Piesmatidae.
306:
Three subfamilies and eleven genera belong to the family
Piesmatidae:
2620:
2376:
2336:
2328:
2268:
2253:
2227:
2125:
1999:
1778:
1770:
1734:
1585:
1436:
1428:
1374:
1273:
1160:
990:
783:
771:
752:
614:
601:
597:
577:
573:
527:
363:
278:
202:
191:
183:
113:
93:
73:
2664:
2492:
2625:
2581:
2515:
2441:
2384:
2284:
2276:
2196:
2133:
2094:
2007:
1914:
1791:
1718:
1606:
1593:
1345:
1337:
1223:
1131:
1123:
1038:
1025:
1010:
1000:
789:
630:
585:
467:
338:
273:
231:
210:
2389:
2351:
2052:
2044:
1927:
1807:
1048:
558:
894:
878:
777:
765:
589:
522:
495:
462:
393:
268:
227:
223:
198:
162:
103:
83:
2465:
1947:
1458:
1144:
518:
249:
2612:
576:
analysis, with their relatives variously presumed to be the
504:
from the earliest Eocene (about 55 mya) is still a very
1292:(cottony cushion scales, giant coccids, ground pearls)
2472:. Heteroptera with Coleorrhyncha were referred to as
629:
2770:
190:. The Piesmatidae are distributed mostly in the
2081:(assassin bugs, wheel bugs, thread-legged bugs)
691:"family Piesmatidae Amyot & Serville, 1843"
481:were split off into separate genera. The genus
282:have been found on many host plants but mostly
1709:
1486:
750:
736:
809:
689:Dellapé, Pablo M.; Henry, Thomas J. (2020).
688:
1109:
743:
729:
661:Grimaldi, David A. & Engel, Michael S.
38:
656:
654:
652:
650:
648:
646:
625:. However a cladistic analysis rejected
209:. A common species found throughout the
708:
682:
14:
2771:
643:
226:, some 2–4 mm overall. The head,
2497:
2496:
1457:
1108:
808:
724:
2652:8b1c5ee9-1081-4d95-a531-21113a6c2c7d
2235:(jewel bugs or metallic shield bugs)
675::10.1206/0003-0082(2008)36112.0.CO;2
2047:(plant bugs, leaf bugs, grass bugs)
1994:(minute pirate bugs or flower bugs)
1651:(marsh treaders or water measurers)
24:
457:Many species were formerly in the
148:Amyot & Audinet-Serville, 1843
25:
2795:
2186:(giant shield bugs and relatives)
941:(typical treehoppers, thorn bugs)
525:might be a northern relative of
332:Nel, Waller & De Ploeg, 2004
60:
2379:(milkweed bugs, true seed bugs)
2287:(squash bugs, leaf-footed bugs)
1554:(unique-headed bugs, gnat bugs)
398:Lepelitier & Serville, 1825
168:Lepelitier & Serville, 1825
1810:(water scorpions, needle bugs)
547:was placed within the family.
309:
13:
1:
636:
222:Ash-grey leaf bugs are small
403:
7:
2436:(red bugs, cotton stainers)
415:Breddin In Schumacher, 1913
301:
10:
2800:
2505:
2457:
2424:
2314:
2267:
2153:
2124:
2115:
1982:
1973:
1943:
1913:
1892:
1871:
1848:
1819:
1790:
1769:
1752:
1705:
1670:
1639:
1618:
1584:
1567:
1537:
1528:
1482:
1478:
1453:
1394:
1365:
1263:(pupillarial palm scales)
1169:
1143:
1122:
1118:
1104:
976:
959:
911:
877:
831:
822:
818:
804:
760:
665:American Museum Novitates
561:and lived about 100 mya (
197:, with some occurring in
160:
153:
57:Scientific classification
55:
46:
37:
32:
1377:(woolly conifer aphids)
695:Lygaeoidea Species File
2300:(scentless plant bugs)
1300:(Mediterranean scales)
621:with the discovery of
508:member of the family.
2751:Paleobiology Database
2444:(bordered plant bugs)
2217:(includes ebony bugs)
1883:(creeping water bugs)
292:. The host plants of
2784:Heteroptera families
2647:Fauna Europaea (new)
2415:(ash-grey leaf bugs)
2407:(atypical seed bugs)
2073:(old world bat bugs)
2002:(bed bugs, bat bugs)
1930:(pygmy backswimmers)
1839:(velvety shore bugs)
1431:(jumping plant lice)
541:. From 2007 to 2009
243:convergent evolution
49:Parapiesma quadratum
2279:(broad-headed bugs)
1630:(velvet water bugs)
1218:(ornate pit scales)
588:, or the peculiar,
266:have been found on
195:Northern Hemisphere
1802:(giant water bugs)
1729:(spiny shore bugs)
1662:Paraphrynoveliidae
1575:(semiaquatic bugs)
1530:Enicocephalomorpha
1324:(phenacoleachiids)
1322:Phenacoleachiidae
1284:(false pit scales)
1282:Lecanodiaspididae
188:ash-grey leaf bugs
186:, commonly called
2766:
2765:
2738:Open Tree of Life
2499:Taxon identifiers
2490:
2489:
2486:
2485:
2482:
2481:
2477:
2453:
2452:
2445:
2437:
2416:
2408:
2405:Rhyparochromidae
2380:
2332:
2301:
2288:
2280:
2249:
2236:
2218:
2200:
2187:
2179:
2176:Acanthosomatidae
2163:
2145:
2137:
2111:
2110:
2098:
2090:
2089:(royal palm bugs)
2087:Thaumastocoridae
2082:
2074:
2056:
2048:
2015:
2003:
1995:
1969:
1968:
1939:
1938:
1931:
1905:
1884:
1840:
1832:
1811:
1803:
1782:
1762:
1760:(true water bugs)
1748:
1747:
1730:
1722:
1701:
1700:
1680:
1652:
1631:
1610:
1597:
1577:
1563:
1562:
1555:
1524:
1523:
1473:
1449:
1448:
1445:
1444:
1432:
1386:
1378:
1357:
1349:
1348:(giant mealybugs)
1341:
1333:
1330:Phoenicococcidae
1325:
1317:
1309:
1301:
1293:
1285:
1277:
1264:
1256:
1248:
1240:
1227:
1219:
1211:
1203:
1195:
1192:Asterolecaniidae
1179:
1153:
1135:
1100:
1099:
1096:
1095:
1092:
1091:
1044:Hypochthonellidae
1034:
969:
955:
954:
942:
929:
903:
887:
841:
563:million years ago
553:was found in mid-
466:, from which the
263:Piesma linnavouri
173:
172:
149:
27:Family of insects
16:(Redirected from
2791:
2759:
2758:
2746:
2745:
2733:
2732:
2720:
2719:
2717:NBNSYS0000160715
2707:
2706:
2694:
2693:
2681:
2680:
2668:
2667:
2655:
2654:
2642:
2641:
2629:
2628:
2616:
2615:
2603:
2602:
2590:
2589:
2577:
2576:
2564:
2563:
2554:
2553:
2541:
2540:
2539:
2526:
2525:
2524:
2494:
2493:
2464:are a different
2459:
2443:
2435:
2414:
2406:
2378:
2347:Cryptorhamphidae
2342:Colobathristidae
2330:
2299:
2286:
2278:
2247:
2234:
2216:
2199:(burrowing bugs)
2198:
2185:
2177:
2164:
2162:
2159:
2143:
2142:Termitaphididae
2135:
2122:
2121:
2096:
2088:
2080:
2072:
2054:
2046:
2012:Curalium cronini
2009:
2001:
1993:
1980:
1979:
1953:
1945:
1944:
1929:
1903:
1882:
1850:Aphelocheiroidea
1838:
1830:
1809:
1801:
1780:
1767:
1766:
1763:
1761:
1758:
1728:
1720:
1707:
1706:
1681:
1679:
1678:(water treaders)
1676:
1650:
1629:
1608:
1596:(water striders)
1595:
1582:
1581:
1578:
1576:
1573:
1553:
1552:Enicocephalidae
1547:Aenictopecheidae
1539:Enicocephaloidea
1535:
1534:
1484:
1483:
1480:
1479:
1472:
1466:
1463:
1455:
1454:
1430:
1384:
1376:
1355:
1347:
1339:
1331:
1323:
1315:
1307:
1299:
1291:
1283:
1275:
1262:
1254:
1247:(armored scales)
1246:
1238:
1225:
1217:
1209:
1201:
1193:
1180:
1178:
1175:
1154:
1152:
1149:
1133:
1120:
1119:
1106:
1105:
1032:
974:
973:
970:
968:
965:
940:
927:
901:
888:
886:
883:
842:
840:
837:
829:
828:
820:
819:
806:
805:
745:
738:
731:
722:
721:
715:
712:
706:
705:
703:
701:
686:
680:
676:
662:
658:
582:Colobathristidae
489:Thaumastocoridae
453:
436:
426:
416:
399:
389:
379:
369:
351:
333:
322:
298:remain unknown.
147:
65:
64:
42:
30:
29:
21:
2799:
2798:
2794:
2793:
2792:
2790:
2789:
2788:
2769:
2768:
2767:
2762:
2754:
2749:
2741:
2736:
2728:
2723:
2715:
2710:
2702:
2697:
2689:
2684:
2676:
2671:
2663:
2658:
2650:
2645:
2637:
2632:
2624:
2619:
2611:
2606:
2598:
2593:
2585:
2580:
2572:
2567:
2559:
2557:
2549:
2544:
2535:
2534:
2529:
2520:
2519:
2514:
2501:
2491:
2478:
2449:
2420:
2400:Pachygronthidae
2367:Heterogastridae
2310:
2306:Stenocephalidae
2263:
2210:Parastrachiidae
2205:Thaumastellidae
2184:Tessaratomidae
2160:
2158:
2157:
2149:
2117:Pentatomomorpha
2107:
1965:
1952:(Coleorrhyncha)
1951:
1950:
1949:Peloridiomorpha
1935:
1909:
1888:
1867:
1858:Aphelocheiridae
1844:
1829:Gelastocoridae
1815:
1800:Belostomatidae
1786:
1781:(water boatmen)
1759:
1757:
1756:
1744:
1711:Leptopodomorpha
1697:
1677:
1675:
1674:
1666:
1635:
1614:
1574:
1572:
1571:
1559:
1520:
1516:Stemmocryptidae
1506:Hypsipterygidae
1488:Dipsocoromorpha
1474:
1470:
1464:
1441:
1424:Phacopteronidae
1390:
1361:
1356:(stictococcids)
1354:Stictococcidae
1338:Pseudococcidae
1316:(ensign scales)
1261:Halimococcidae
1177:(scale insects)
1176:
1174:
1173:
1165:
1150:
1148:
1147:
1139:
1114:
1088:
1079:Tettigometridae
966:
964:
963:
951:
907:
902:(hairy cicadas)
900:Tettigarctidae
884:
882:
881:
873:
838:
836:
835:
814:
812:Auchenorrhyncha
800:
756:
749:
719:
718:
713:
709:
699:
697:
687:
683:
671:
660:
659:
644:
639:
613:or placed in a
451:
434:
424:
414:
406:
397:
387:
377:
367:
349:
331:
320:
312:
304:
258:Caryophyllaceae
252:sap, mostly of
216:Piesma cinereum
169:
166:
146:
59:
28:
23:
22:
15:
12:
11:
5:
2797:
2787:
2786:
2781:
2764:
2763:
2761:
2760:
2747:
2734:
2721:
2708:
2695:
2682:
2669:
2656:
2643:
2634:Fauna Europaea
2630:
2617:
2604:
2591:
2578:
2565:
2555:
2542:
2527:
2511:
2509:
2503:
2502:
2488:
2487:
2484:
2483:
2480:
2479:
2458:
2455:
2454:
2451:
2450:
2448:
2447:
2439:
2434:Pyrrhocoridae
2430:
2428:
2426:Pyrrhocoroidea
2422:
2421:
2419:
2418:
2410:
2402:
2397:
2392:
2387:
2382:
2374:
2369:
2364:
2359:
2354:
2349:
2344:
2339:
2334:
2326:
2320:
2318:
2312:
2311:
2309:
2308:
2303:
2295:
2290:
2282:
2273:
2271:
2265:
2264:
2262:
2261:
2256:
2251:
2243:
2238:
2233:Scutelleridae
2230:
2225:
2220:
2215:Thyreocoridae
2212:
2207:
2202:
2194:
2189:
2181:
2173:
2167:
2165:
2151:
2150:
2148:
2147:
2144:(termite bugs)
2139:
2130:
2128:
2119:
2113:
2112:
2109:
2108:
2106:
2105:
2100:
2092:
2084:
2076:
2068:
2066:Plokiophilidae
2063:
2058:
2050:
2042:
2037:
2032:
2027:
2022:
2017:
2005:
1997:
1988:
1986:
1977:
1971:
1970:
1967:
1966:
1964:
1963:
1957:
1955:
1941:
1940:
1937:
1936:
1934:
1933:
1925:
1919:
1917:
1911:
1910:
1908:
1907:
1904:(backswimmers)
1898:
1896:
1890:
1889:
1887:
1886:
1877:
1875:
1869:
1868:
1866:
1865:
1860:
1854:
1852:
1846:
1845:
1843:
1842:
1834:
1825:
1823:
1817:
1816:
1814:
1813:
1805:
1796:
1794:
1788:
1787:
1785:
1784:
1775:
1773:
1764:
1750:
1749:
1746:
1745:
1743:
1742:
1737:
1732:
1724:
1715:
1713:
1703:
1702:
1699:
1698:
1696:
1695:
1690:
1684:
1682:
1668:
1667:
1665:
1664:
1659:
1654:
1649:Hydrometridae
1645:
1643:
1641:Hydrometroidea
1637:
1636:
1634:
1633:
1624:
1622:
1616:
1615:
1613:
1612:
1604:
1602:Hermatobatidae
1599:
1590:
1588:
1579:
1565:
1564:
1561:
1560:
1558:
1557:
1549:
1543:
1541:
1532:
1526:
1525:
1522:
1521:
1519:
1518:
1513:
1511:Schizopteridae
1508:
1503:
1498:
1496:Ceratocombidae
1492:
1490:
1476:
1475:
1451:
1450:
1447:
1446:
1443:
1442:
1440:
1439:
1434:
1426:
1421:
1416:
1411:
1406:
1400:
1398:
1392:
1391:
1389:
1388:
1385:(phylloxerans)
1383:Phylloxeridae
1380:
1371:
1369:
1367:Phylloxeroidea
1363:
1362:
1360:
1359:
1351:
1343:
1335:
1327:
1319:
1311:
1308:(giant scales)
1306:Monophlebidae
1303:
1298:Micrococcidae
1295:
1287:
1279:
1271:
1266:
1258:
1250:
1242:
1237:Dactylopiidae
1234:
1232:Conchaspididae
1229:
1221:
1213:
1210:(carayonemids)
1208:Carayonemidae
1205:
1197:
1189:
1183:
1181:
1167:
1166:
1164:
1163:
1157:
1155:
1141:
1140:
1138:
1137:
1128:
1126:
1116:
1115:
1112:Sternorrhyncha
1102:
1101:
1098:
1097:
1094:
1093:
1090:
1089:
1087:
1086:
1081:
1076:
1071:
1066:
1061:
1056:
1051:
1046:
1041:
1036:
1033:(lanternflies)
1028:
1023:
1018:
1016:Dictyopharidae
1013:
1008:
1003:
998:
993:
988:
982:
980:
971:
967:(planthoppers)
957:
956:
953:
952:
950:
949:
944:
936:
931:
923:
917:
915:
909:
908:
906:
905:
897:
891:
889:
875:
874:
872:
871:
866:
861:
859:Clastopteridae
856:
851:
845:
843:
826:
816:
815:
802:
801:
799:
798:
792:
786:
780:
774:
768:
761:
758:
757:
748:
747:
740:
733:
725:
717:
716:
707:
681:
641:
640:
638:
635:
610:incertae sedis
494:There are two
455:
454:
452:Kormilev, 1969
443:
442:
438:
437:
435:Bergroth, 1921
427:
417:
405:
402:
401:
400:
390:
388:Pericart, 1974
380:
370:
360:
341:
323:
321:Pericart, 1974
311:
308:
303:
300:
254:Chenopodiaceae
171:
170:
167:
158:
157:
151:
150:
141:
137:
136:
131:
127:
126:
121:
117:
116:
111:
107:
106:
101:
97:
96:
91:
87:
86:
81:
77:
76:
71:
67:
66:
53:
52:
44:
43:
35:
34:
26:
9:
6:
4:
3:
2:
2796:
2785:
2782:
2780:
2777:
2776:
2774:
2757:
2752:
2748:
2744:
2739:
2735:
2731:
2726:
2722:
2718:
2713:
2709:
2705:
2700:
2696:
2692:
2687:
2683:
2679:
2674:
2670:
2666:
2661:
2657:
2653:
2648:
2644:
2640:
2635:
2631:
2627:
2622:
2618:
2614:
2609:
2605:
2601:
2596:
2592:
2588:
2583:
2579:
2575:
2570:
2566:
2562:
2556:
2552:
2547:
2543:
2538:
2532:
2528:
2523:
2517:
2513:
2512:
2510:
2508:
2504:
2500:
2495:
2475:
2474:Prosorrhyncha
2471:
2467:
2463:
2462:Coleorrhyncha
2456:
2446:
2440:
2438:
2432:
2431:
2429:
2427:
2423:
2417:
2411:
2409:
2403:
2401:
2398:
2396:
2393:
2391:
2388:
2386:
2383:
2381:
2375:
2373:
2370:
2368:
2365:
2363:
2362:Henicocoridae
2360:
2358:
2355:
2353:
2350:
2348:
2345:
2343:
2340:
2338:
2335:
2333:
2327:
2325:
2322:
2321:
2319:
2317:
2313:
2307:
2304:
2302:
2296:
2294:
2293:Hyocephalidae
2291:
2289:
2283:
2281:
2275:
2274:
2272:
2270:
2266:
2260:
2257:
2255:
2252:
2250:
2246:Pentatomidae
2244:
2242:
2239:
2237:
2231:
2229:
2226:
2224:
2221:
2219:
2213:
2211:
2208:
2206:
2203:
2201:
2195:
2193:
2190:
2188:
2182:
2180:
2178:(shield bugs)
2174:
2172:
2171:Urostylididae
2169:
2168:
2166:
2161:(shield bugs)
2156:
2155:Pentatomoidea
2152:
2146:
2140:
2138:
2132:
2131:
2129:
2127:
2123:
2120:
2118:
2114:
2104:
2103:Velocipedidae
2101:
2099:
2093:
2091:
2085:
2083:
2077:
2075:
2071:Polyctenidae
2069:
2067:
2064:
2062:
2059:
2057:
2055:(damsel bugs)
2051:
2049:
2043:
2041:
2040:Microphysidae
2038:
2036:
2033:
2031:
2028:
2026:
2025:Lasiochilidae
2023:
2021:
2018:
2016:
2013:
2006:
2004:
1998:
1996:
1992:Anthocoridae
1990:
1989:
1987:
1985:
1981:
1978:
1976:
1972:
1962:
1959:
1958:
1956:
1954:
1946:
1942:
1932:
1926:
1924:
1923:Helotrephidae
1921:
1920:
1918:
1916:
1912:
1906:
1902:Notonectidae
1900:
1899:
1897:
1895:
1894:Notonectoidea
1891:
1885:
1879:
1878:
1876:
1874:
1870:
1864:
1863:Potamocoridae
1861:
1859:
1856:
1855:
1853:
1851:
1847:
1841:
1835:
1833:
1827:
1826:
1824:
1822:
1818:
1812:
1806:
1804:
1798:
1797:
1795:
1793:
1789:
1783:
1777:
1776:
1774:
1772:
1768:
1765:
1755:
1751:
1741:
1738:
1736:
1733:
1731:
1727:Leptopodidae
1725:
1723:
1717:
1716:
1714:
1712:
1708:
1704:
1694:
1691:
1689:
1688:Madeoveliidae
1686:
1685:
1683:
1673:
1672:Mesovelioidea
1669:
1663:
1660:
1658:
1657:Macroveliidae
1655:
1653:
1647:
1646:
1644:
1642:
1638:
1632:
1626:
1625:
1623:
1621:
1617:
1611:
1609:(riffle bugs)
1605:
1603:
1600:
1598:
1592:
1591:
1589:
1587:
1583:
1580:
1570:
1566:
1556:
1550:
1548:
1545:
1544:
1542:
1540:
1536:
1533:
1531:
1527:
1517:
1514:
1512:
1509:
1507:
1504:
1502:
1499:
1497:
1494:
1493:
1491:
1489:
1485:
1481:
1477:
1469:
1468:Coleorrhyncha
1462:
1456:
1452:
1438:
1435:
1433:
1427:
1425:
1422:
1420:
1417:
1415:
1412:
1410:
1407:
1405:
1402:
1401:
1399:
1397:
1393:
1387:
1381:
1379:
1373:
1372:
1370:
1368:
1364:
1358:
1352:
1350:
1344:
1342:
1336:
1334:
1332:(palm scales)
1328:
1326:
1320:
1318:
1312:
1310:
1304:
1302:
1296:
1294:
1290:Margarodidae
1288:
1286:
1280:
1278:
1272:
1270:
1267:
1265:
1259:
1257:
1255:(felt scales)
1253:Eriococcidae
1251:
1249:
1243:
1241:
1235:
1233:
1230:
1228:
1226:(soft scales)
1222:
1220:
1216:Cerococcidae
1214:
1212:
1206:
1204:
1198:
1196:
1190:
1188:
1185:
1184:
1182:
1172:
1168:
1162:
1159:
1158:
1156:
1146:
1142:
1136:
1130:
1129:
1127:
1125:
1121:
1117:
1113:
1107:
1103:
1085:
1084:Tropiduchidae
1082:
1080:
1077:
1075:
1072:
1070:
1067:
1065:
1062:
1060:
1057:
1055:
1052:
1050:
1047:
1045:
1042:
1040:
1037:
1035:
1029:
1027:
1024:
1022:
1021:Eurybrachidae
1019:
1017:
1014:
1012:
1009:
1007:
1004:
1002:
999:
997:
994:
992:
989:
987:
986:Acanaloniidae
984:
983:
981:
979:
975:
972:
962:
961:Fulgoromorpha
958:
948:
947:Myerslopiidae
945:
943:
937:
935:
934:Melizoderidae
932:
930:
928:(leafhoppers)
926:Cicadellidae
924:
922:
919:
918:
916:
914:
910:
904:
898:
896:
893:
892:
890:
880:
876:
870:
869:Machaerotidae
867:
865:
862:
860:
857:
855:
852:
850:
849:Aphrophoridae
847:
846:
844:
839:(froghoppers)
834:
830:
827:
825:
821:
817:
813:
807:
803:
797:
793:
791:
787:
785:
781:
779:
775:
773:
769:
767:
763:
762:
759:
754:
746:
741:
739:
734:
732:
727:
726:
723:
711:
696:
692:
685:
679:
674:
669:
666:
657:
655:
653:
651:
649:
647:
642:
634:
632:
628:
624:
620:
616:
612:
611:
605:
603:
599:
595:
591:
587:
583:
579:
575:
571:
566:
564:
560:
556:
552:
551:
546:
545:
540:
536:
535:
530:
529:
524:
520:
517:
513:
512:
507:
503:
502:
497:
492:
490:
486:
485:
480:
479:
474:
473:
469:
465:
464:
460:
450:
449:
445:
444:
440:
439:
433:
432:
428:
423:
422:
418:
413:
412:
408:
407:
396:
395:
391:
386:
385:
381:
376:
375:
371:
366:
365:
361:
358:
354:
348:
347:
342:
340:
336:
330:
329:
324:
319:
318:
314:
313:
307:
299:
297:
296:
291:
290:
285:
281:
280:
275:
271:
270:
265:
264:
259:
255:
251:
248:They feed on
246:
244:
240:
237:
233:
229:
225:
220:
218:
217:
212:
208:
207:South America
204:
200:
196:
193:
189:
185:
181:
177:
165:
164:
159:
156:
152:
145:
142:
139:
138:
135:
132:
130:Superfamily:
129:
128:
125:
122:
119:
118:
115:
112:
109:
108:
105:
102:
99:
98:
95:
92:
89:
88:
85:
82:
79:
78:
75:
72:
69:
68:
63:
58:
54:
51:
50:
45:
41:
36:
31:
19:
2506:
2413:Piesmatidae
2412:
2395:Oxycarenidae
2372:Idiostolidae
2331:(stilt bugs)
2248:(stink bugs)
2061:Pachynomidae
2035:Medocostidae
2030:Lyctocoridae
2011:
1975:Cimicomorpha
1961:Peloridiidae
1740:Aepophilidae
1721:(shore bugs)
1693:Mesoveliidae
1501:Dipsocoridae
1414:Carsidaridae
1314:Ortheziidae
1276:(lac scales)
1245:Diaspididae
1239:(cochineals)
1202:(beesoniids)
1200:Beesoniidae
1194:(pit scales)
1134:(whiteflies)
1132:Aleyrodidae
1124:Aleyrodoidea
996:Achilixiidae
939:Membracidae
921:Aetalionidae
913:Membracoidea
824:Cicadomorpha
796:Paraneoptera
794:Superorder:
788:Infraclass:
710:
698:. Retrieved
694:
684:
678:PDF fulltext
667:
664:
626:
622:
619:Piesmatoidea
618:
617:superfamily
608:
606:
567:
548:
542:
538:
532:
526:
509:
499:
493:
482:
476:
470:
461:
456:
446:
429:
425:Slater, 1970
419:
409:
392:
382:
372:
362:
353:Baltic amber
344:
326:
315:
305:
293:
287:
283:
277:
267:
261:
247:
239:Cimicomorpha
221:
214:
187:
175:
174:
161:
143:
47:
33:Piesmatidae
2779:Piesmatidae
2673:iNaturalist
2551:PIESMATIDAE
2537:Piesmatidae
2531:Wikispecies
2507:Piesmatidae
2470:Heteroptera
2324:Artheneidae
2298:Rhopalidae
2259:Megarididae
2241:Plataspidae
2223:Lestoniidae
2192:Dinidoridae
2136:(flat bugs)
2097:(lace bugs)
2079:Reduviidae
2020:Joppeicidae
2008:Curaliidae
1881:Naucoridae
1873:Naucoroidea
1837:Ochteridae
1831:(toad bugs)
1821:Ochteroidea
1569:Gerromorpha
1461:Heteroptera
1419:Homotomidae
1409:Calophyidae
1404:Aphalaridae
1340:(mealybugs)
1069:Nogodinidae
1064:Meenoplidae
1031:Fulgoridae
1006:Delphacidae
978:Fulgoroidea
833:Cercopoidea
627:Cretopiesma
623:Cretopiesma
557:amber from
550:Cretopiesma
544:Cretopiesma
514:, found in
441:Thaicorniae
378:Drake, 1948
368:Drake, 1924
357:Rovno amber
350:Popov, 2001
310:Piesmatinae
178:is a small
176:Piesmatidae
144:Piesmatidae
124:Heteroptera
18:Piesmatinae
2773:Categories
2377:Lygaeidae
2357:Geocoridae
2329:Berytidae
2316:Lygaeoidea
2000:Cimicidae
1984:Cimicoidea
1779:Corixidae
1771:Corixoidea
1754:Nepomorpha
1429:Psyllidae
1396:Psylloidea
1375:Adelgidae
1274:Kerriidae
1269:Kermesidae
1187:Aclerdidae
1145:Aphidoidea
1074:Ricaniidae
1059:Lophopidae
1054:Kinnaridae
879:Cicadoidea
864:Epipygidae
854:Cercopidae
782:Subclass:
772:Arthropoda
637:References
570:Lygaeoidea
555:Cretaceous
478:Parapiesma
472:Afropiesma
459:type genus
384:Parapiesma
337:, France,
335:Oise amber
317:Afropiesma
289:Proteaceae
236:infraorder
155:Type genus
134:Lygaeoidea
120:Suborder:
94:Arthropoda
2442:Largidae
2337:Blissidae
2285:Coreidae
2277:Alydidae
2269:Coreoidea
2254:Canopidae
2228:Phloeidae
2197:Cydnidae
2134:Aradidae
2126:Aradoidea
2095:Tingidae
1735:Omaniidae
1719:Saldidae
1628:Hebridae
1620:Hebroidea
1607:Veliidae
1594:Gerridae
1586:Gerroidea
1459:Suborder
1437:Triozidae
1346:Putoidae
1224:Coccidae
1171:Coccoidea
1161:Aphididae
1110:Suborder
991:Achilidae
895:Cicadidae
885:(cicadas)
810:Suborder
784:Pterygota
764:Kingdom:
753:Hemiptera
700:29 August
615:monotypic
602:Lygaeidae
598:subfamily
594:Psamminae
578:Berytidae
574:cladistic
572:based on
528:Mcateella
521:from the
511:Heissiana
484:Thaicoris
468:subgenera
448:Thaicoris
431:Sympeplus
421:Saxicoris
404:Psamminae
364:Mcateella
346:Heissiana
279:Mcateella
245:however.
203:Australia
192:temperate
184:true bugs
114:Hemiptera
80:Kingdom:
74:Eukaryota
2582:BugGuide
2558:BioLib:
2522:Q1950799
2516:Wikidata
2385:Malcidae
2053:Nabidae
2045:Miridae
1928:Pleidae
1915:Pleoidea
1808:Nepidae
1792:Nepoidea
1151:(aphids)
1039:Gengidae
1026:Flatidae
1011:Derbidae
1001:Cixiidae
790:Neoptera
770:Phylum:
766:Animalia
755:families
670:: 1-17.
631:Aradidae
586:Malcidae
539:Eopiesma
501:Eopiesma
498:genera.
411:Psammium
359:, Eocene
339:Ypresian
328:Eopiesma
302:Taxonomy
274:Fabaceae
232:Tingidae
211:Americas
140:Family:
90:Phylum:
84:Animalia
70:Domain:
2390:Ninidae
2352:Cymidae
1049:Issidae
778:Insecta
776:Class:
751:Extant
600:of the
559:Myanmar
234:of the
224:insects
110:Order:
104:Insecta
100:Class:
2756:218539
2743:568266
2730:236392
2704:107629
2691:100726
2678:343428
2626:1PIESF
2587:247450
2574:478458
2460:Note:
592:-like
590:beetle
534:Miespa
523:Eocene
516:Baltic
496:fossil
463:Piesma
394:Piesma
374:Miespa
295:Miespa
284:Acacia
269:Acacia
228:thorax
199:Africa
180:family
163:Piesma
2686:IRMNG
2639:12841
2600:8NKPC
2561:17128
2468:from
2466:clade
1465:(with
519:amber
506:basal
250:plant
2725:NCBI
2699:ITIS
2665:9651
2660:GBIF
2621:EPPO
2569:BOLD
702:2020
668:3611
596:, a
584:and
531:and
475:and
286:and
256:and
205:and
2712:NBN
2613:625
2608:EoL
2595:CoL
2546:AFD
673:DOI
276:).
213:is
182:of
2775::
2753::
2740::
2727::
2714::
2701::
2688::
2675::
2662::
2649::
2636::
2623::
2610::
2597::
2584::
2571::
2548::
2533::
2518::
693:.
645:^
633:.
580:,
491:.
355:,
343:â€
325:â€
260:.
219:.
201:,
2476:.
2014:)
2010:(
1471:)
744:e
737:t
730:v
704:.
272:(
20:)
Text is available under the Creative Commons Attribution-ShareAlike License. Additional terms may apply.
